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<document id="F1C433C9A6F27796CC8D310A2DD63B72" ID-CLB-Dataset="299409" ID-DOI="10.5281/zenodo.11067293" ID-GBIF-Dataset="9235db45-04c4-4570-8987-cb84765763a4" ID-ISSN="0068-547X" ID-Zenodo-Dep="11067293" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="karina" IM.treatmentCitations_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1714052207303" checkinUser="felipe" docAuthor="Kavanaugh, David H., Cueva-Dabkoski, Mollie &amp; Liang, Hongbin" docDate="2023" docId="644DE316FF88FFB0D25EDAAEFB446304" docLanguage="en" docName="ProcCalifAcadSci.67.20.451-491.pdf" docOrigin="Proceedings of the California Academy of Sciences 67 (20)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, western Yunnan Province, China: Species of the Tribe Cyclosomini Laporte, 1934 (Coleoptera: Carabidae), with Descriptions of Two New Species." docType="key" docVersion="2" lastPageNumber="459" masterDocId="98749B6EFF8FFFB8D334D867FFCA6027" masterDocTitle="Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, western Yunnan Province, China: Species of the Tribe Cyclosomini Laporte, 1934 (Coleoptera: Carabidae), with Descriptions of Two New Species." masterLastPageNumber="491" masterPageNumber="451" pageNumber="458" updateTime="1719949330493" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="0672CB942B4FC036B95F7F9B0520352B">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, western Yunnan Province, China: Species of the Tribe Cyclosomini Laporte, 1934 (Coleoptera: Carabidae), with Descriptions of Two New Species.</mods:title>
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<mods:namePart id="64BA7014D2FF06D7F50B8BE3B7385887">Kavanaugh, David H.</mods:namePart>
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<mods:namePart id="0A7AB3547AFD3242BF4CFE5DFD4AD766">Cueva-Dabkoski, Mollie</mods:namePart>
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<mods:namePart id="B2D5DE31862F26DA7BAEC75199BBB22E">Liang, Hongbin</mods:namePart>
<mods:affiliation id="B0853A42B5B116E3052F9505C048402A">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China.</mods:affiliation>
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Key for identification of adult cyclosomine species of the Gaoligong Shan region of 
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1 Size larger, BL = 
<quantity id="2B1CFFE5FF88FFBFD263DB44FE6B631E" box="[343,417,803,825]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="7" pageNumber="458" unit="mm" value="6.8">6.8 mm</quantity>
or more; pronotum (
<figureCitation id="74DF4E85FF88FFBFD159DB44FD0E631E" box="[621,708,803,825]" captionStart="FIGURE" captionStartId="30.[155,167,1512,1531]" captionTargetBox="[140,1157,216,1497]" captionTargetId="figure-11@30.[140,1157,216,1497]" captionTargetPageId="30" captionText="FIGURE 13. Pronota of cyclosomines, dorsal aspect.A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (CASENT 1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov., (CASENT1039394, holotype, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT 1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067317" httpUri="https://zenodo.org/record/11067317/files/figure.png" pageId="7" pageNumber="458">Fig. 13A</figureCitation>
) very broad, trapezoidal, almost as wide as elytra at humeral angles, distinctly widest at base; elytra broadly rounded apically, elytral color pattern as in 
<figureCitation id="74DF4E85FF88FFBFD39FDB3AFF366354" box="[171,252,861,883]" captionStart="FIGURE" captionStartId="10.[155,167,1587,1606]" captionTargetBox="[126,1180,216,1575]" captionTargetId="figure-67@10.[120,1180,216,1575]" captionTargetPageId="10" captionText="FIGURE 7. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. A. Holotype, dorsal aspect, scale line = 1.0 mm (CASENT 1015370, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Labels associated with holotype; C. Map showing locality record (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067301" httpUri="https://zenodo.org/record/11067301/files/figure.png" pageId="7" pageNumber="458">Fig. 7A</figureCitation>
; front tarsomeres 1 to 3 markedly expanded laterally (
<figureCitation id="74DF4E85FF88FFBFD019DB3AFC596354" box="[813,915,861,883]" captionStart="FIGURE" captionStartId="31.[155,167,1512,1531]" captionTargetBox="[253,1040,238,1497]" captionTargetId="figure-15@31.[253,1044,216,1497]" captionTargetPageId="31" captionText="FIGURE 14. Male front tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067319" httpUri="https://zenodo.org/record/11067319/files/figure.png" pageId="7" pageNumber="458">Figs. 14A</figureCitation>
, 
<figureCitation id="74DF4E85FF88FFBFD095DB3AFC016354" box="[929,971,861,883]" captionStart="FIGURE" captionStartId="32.[155,167,1524,1543]" captionTargetBox="[279,1026,226,1510]" captionTargetId="figure-11@32.[269,1026,216,1510]" captionTargetPageId="32" captionText="FIGURE 15. Male front tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067321" httpUri="https://zenodo.org/record/11067321/files/figure.png" pageId="7" pageNumber="458">15A</figureCitation>
) in both males and females................... 
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<taxonomicName id="2BE42983FF88FFBFD2D5DB1DFBFC63B7" authority="Kavanaugh and Cueva-Dabkoski" authorityName="Kavanaugh and Cueva-Dabkoski" authorityYear="2023" box="[481,1078,890,912]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="acutangulus" status="sp. nov.">
<emphasis id="DE908E12FF88FFBFD2D5DB1DFD1363A8" bold="true" box="[481,729,890,911]" italics="true" pageId="7" pageNumber="458">Cyclosomus acutangulus</emphasis>
Kavanaugh and Cueva-Dabkoski
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, 
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1’ Size smaller, BL = less than 6.0 mm; pronotum (
<figureCitation id="74DF4E85FF88FFBFD14ADBF0FD06638A" box="[638,716,919,941]" captionStart="FIGURE" captionStartId="30.[155,167,1512,1531]" captionTargetBox="[140,1157,216,1497]" captionTargetId="figure-11@30.[140,1157,216,1497]" captionTargetPageId="30" captionText="FIGURE 13. Pronota of cyclosomines, dorsal aspect.A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (CASENT 1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov., (CASENT1039394, holotype, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT 1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067317" httpUri="https://zenodo.org/record/11067317/files/figure.png" pageId="7" pageNumber="458">Figs. 13</figureCitation>
B-D) narrower, widest at or anterior to middle; elytra slightly to distinctly and obliquely truncate apically, elytral color pattern varied; front tarsomeres 1 to 3 without distinct lateral expansions (
<figureCitation id="74DF4E85FF88FFBFD1BDDBB7FD2863C1" box="[649,738,976,998]" captionStart="FIGURE" captionStartId="31.[155,167,1512,1531]" captionTargetBox="[253,1040,238,1497]" captionTargetId="figure-15@31.[253,1044,216,1497]" captionTargetPageId="31" captionText="FIGURE 14. Male front tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067319" httpUri="https://zenodo.org/record/11067319/files/figure.png" pageId="7" pageNumber="458">Figs. 14</figureCitation>
B-F), broader in males than in females............................................................... 
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spp. 2
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2 (1’) Femora dark black or piceous; elytra dark without pale markings in basal half, with or without pale markings in apical half (
<figureCitation id="74DF4E85FF88FFBFD2A3DC54FE27646E" box="[407,493,1075,1097]" captionStart="FIGURE" captionStartId="13.[155,167,1592,1611]" captionTargetBox="[128,1175,226,1577]" captionTargetId="figure-56@13.[120,1176,216,1581]" captionTargetPageId="13" captionText="FIGURE 9. Tetragonoderus arcuatus (Dejean). A. Dorsal aspect, scale line = 1.0 mm (CASENT1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); B. Map showing locality record (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067303" httpUri="https://zenodo.org/record/11067303/files/figure.png" pageId="7" pageNumber="458">Figs. 9A</figureCitation>
, 
<figureCitation id="74DF4E85FF88FFBFD2CCDC54FDEA646E" box="[504,544,1075,1097]" captionStart="FIGURE" captionStartId="19.[155,167,1597,1616]" captionTargetBox="[123,1176,216,1583]" captionTargetId="figure-73@19.[120,1176,216,1584]" captionTargetPageId="19" captionText="FIGURE 11. Tetragonoderus parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. A. Holotype, dorsal aspect, scale line = 1.0 mm (CASENT1039396, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); B. Labels associated with holotype; C. Map showing locality records (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067309" httpUri="https://zenodo.org/record/11067309/files/figure.png" pageId="7" pageNumber="458">11A</figureCitation>
)................................................. 3
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2’ Femora pale tan; elytra with pale areas in both basal and apical halves (
<figureCitation id="74DF4E85FF88FFBFD06BDC28FC0B6442" box="[863,961,1103,1125]" captionStart="FIGURE" captionStartId="16.[155,167,1603,1622]" captionTargetBox="[120,1176,216,1581]" captionTargetId="figure-60@16.[120,1176,216,1590]" captionTargetPageId="16" captionText="FIGURE 10. Tetragonoderus elegans Andrewes. A. Dorsal aspect, scale line = 1.0 mm (CASENT1039409, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); B. Map showing locality records (black circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067305" httpUri="https://zenodo.org/record/11067305/files/figure.png" pageId="7" pageNumber="458">Figs. 10A</figureCitation>
, 
<figureCitation id="74DF4E85FF88FFBFD0F8DC28FC3D6442" box="[972,1015,1103,1125]" captionStart="FIGURE" captionStartId="22.[155,167,1587,1606]" captionTargetBox="[128,1176,216,1576]" captionTargetId="figure-59@22.[120,1176,216,1576]" captionTargetPageId="22" captionText="FIGURE 12. Tetragonoderus punctatus (Wiedemann). A. Dorsal aspect, scale line = 1.0 mm (CASENT1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); B. Map showing locality record (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067313" httpUri="https://zenodo.org/record/11067313/files/figure.png" pageId="7" pageNumber="458">12A</figureCitation>
, 
<figureCitation id="74DF4E85FF88FFBFD736DC28FBD46442" box="[1026,1054,1103,1125]" captionStart="FIGURE" captionStartId="37.[155,167,1091,1110]" captionTargetBox="[384,875,236,1075]" captionTargetId="figure-14@37.[384,912,216,1079]" captionTargetPageId="37" captionText="FIGURE 20. Tetragonoderus microthorax Jian and Tian, dorsal habitus of holotype (Jinghong, Yunnan, China); scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11067333" httpUri="https://zenodo.org/record/11067333/files/figure.png" pageId="7" pageNumber="458">20</figureCitation>
)........ 4
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3 (2’) Size smaller, BL = 
<quantity id="2B1CFFE5FF88FFBFD247DC1FFE7764A9" box="[371,445,1144,1166]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="7" pageNumber="458" unit="mm" value="4.5">4.5 mm</quantity>
or less; elytra with distinct transverse subapical band of pale spots extended from interval 4 to interval 8 (
<figureCitation id="74DF4E85FF88FFBFD2D0DCF2FDF7648C" box="[484,573,1173,1195]" captionStart="FIGURE" captionStartId="19.[155,167,1597,1616]" captionTargetBox="[123,1176,216,1583]" captionTargetId="figure-73@19.[120,1176,216,1584]" captionTargetPageId="19" captionText="FIGURE 11. Tetragonoderus parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. A. Holotype, dorsal aspect, scale line = 1.0 mm (CASENT1039396, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); B. Labels associated with holotype; C. Map showing locality records (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067309" httpUri="https://zenodo.org/record/11067309/files/figure.png" pageId="7" pageNumber="458">Fig. 11A</figureCitation>
), faintly present also on interval 
<quantity id="2B1CFFE5FF88FFBFD049DCF2FC69648C" box="[893,931,1173,1195]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="7" pageNumber="458" unit="in" value="3.0">3 in</quantity>
some individuals, posterior discal setiferous puncture on interval 3 inserted more posteriad, at level of pale transverse subapical band; dorsal surfaces shiny, with distinct aeneous-greenish metallic reflection, of similar sheen throughout, elytral microsculpture comprised of more or less regularly transverse meshes, except their orientation slightly distorted in or near discal setal pore punctures........................................................... 
<emphasis id="DE908E12FF88FFBFD2FBDD42FB44651C" bold="true" box="[463,1166,1317,1339]" pageId="7" pageNumber="458">
<taxonomicName id="2BE42983FF88FFBFD2FBDD42FBFC651C" authority="Kavanaugh and Cueva-Dabkoski" authorityName="Kavanaugh and Cueva-Dabkoski" authorityYear="2023" box="[463,1078,1317,1339]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="parviculus" status="sp. nov.">
<emphasis id="DE908E12FF88FFBFD2FBDD42FD13651D" bold="true" box="[463,729,1317,1338]" italics="true" pageId="7" pageNumber="458">Tetragonoderus parviculus</emphasis>
Kavanaugh and Cueva-Dabkoski
</taxonomicName>
, 
<taxonomicNameLabel id="C5A33369FF88FFBFD775DD41FB44651C" box="[1089,1166,1318,1339]" pageId="7" pageNumber="458" rank="species">sp. nov.</taxonomicNameLabel>
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<keyLead id="5715F135FF88FFBFD34FDD25FB456622" pageId="7" pageNumber="458">
3’ Size larger, BL = 
<quantity id="2B1CFFE5FF88FFBFD263DD25FE6A657F" box="[343,416,1346,1368]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="7" pageNumber="458" unit="mm" value="4.9">4.9 mm</quantity>
or more; elytra (
<figureCitation id="74DF4E85FF88FFBFD171DD25FD5B657F" box="[581,657,1346,1368]" captionStart="FIGURE" captionStartId="13.[155,167,1592,1611]" captionTargetBox="[128,1175,226,1577]" captionTargetId="figure-56@13.[120,1176,216,1581]" captionTargetPageId="13" captionText="FIGURE 9. Tetragonoderus arcuatus (Dejean). A. Dorsal aspect, scale line = 1.0 mm (CASENT1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); B. Map showing locality record (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067303" httpUri="https://zenodo.org/record/11067303/files/figure.png" pageId="7" pageNumber="458">Fig. 9A</figureCitation>
) with only faint pale markings in region of subapical band, most evident on intervals 7 and 8, posterior discal setiferous puncture on interval 3 inserted distinctly anterior to level of pale subapical markings (i.e., closer to anterior discal setiferous pore); dorsal surfaces duller, with distinct bronze/copper metallic reflection, sericeous (silky) and mottled in posterior half, with elytral microsculpture comprised of a complex pattern of transverse, longitudinal, and oblique patches of elongate meshes, most distorted near discal and umbilicate setal pore punctures.................................................................. 
<taxonomicName id="2BE42983FF88FFBFD067DD88FB456622" authority="Dejean" authorityName="Dejean" authorityYear="1829" box="[851,1167,1519,1541]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="arcuatus">
<emphasis id="DE908E12FF88FFBFD067DD88FB8E6622" box="[851,1092,1519,1541]" italics="true" pageId="7" pageNumber="458">Tetragonoderus arcuatus</emphasis>
Dejean
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</keyStep>
<keyStep id="57104AA5FF88FFB0D34FDE70FB4561D3" lastPageId="8" lastPageNumber="459" pageId="7" pageNumber="458">
<paragraph id="EC5B5200FF88FFB0D34FDE70FB446129" blockId="7.[123,1174,1559,1725]" lastBlockId="8.[123,1191,219,500]" lastPageId="8" lastPageNumber="459" pageId="7" pageNumber="458">
<keyLead id="5715F135FF88FFB0D34FDE70FB446129" lastPageId="8" lastPageNumber="459" pageId="7" pageNumber="458">
4 (3’) Elytral color pattern as in 
<figureCitation id="74DF4E85FF88FFBFD28CDE70FDDA660A" box="[440,528,1559,1581]" captionStart="FIGURE" captionStartId="16.[155,167,1603,1622]" captionTargetBox="[120,1176,216,1581]" captionTargetId="figure-60@16.[120,1176,216,1590]" captionTargetPageId="16" captionText="FIGURE 10. Tetragonoderus elegans Andrewes. A. Dorsal aspect, scale line = 1.0 mm (CASENT1039409, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); B. Map showing locality records (black circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067305" httpUri="https://zenodo.org/record/11067305/files/figure.png" pageId="7" pageNumber="458">Fig. 10A</figureCitation>
, with dark areas more expansive; connection between medial portion of the basal dark band and humeral portion broader, at least one-quarter length of the humeral portion; middle band with portion between anterior and posterior dark edges distinctly darker (orange-brown) than remainder of pale elytral areas; pronotum dark, black or piceous, with distinct greenish or bronze metallic reflection, slightly narrowed basally (ratio PWM/PWB =1.13 to 1.20); male with middle tarsomeres 1 to 4 (
<figureCitation id="74DF4E85FF88FFBFD250DEC0FE76669A" box="[356,444,1703,1725]" captionStart="FIGURE" captionStartId="33.[155,167,1521,1540]" captionTargetBox="[215,1075,216,1495]" captionTargetId="figure-133@33.[215,1081,216,1509]" captionTargetPageId="33" captionText="FIGURE 16. Male middle tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067325" httpUri="https://zenodo.org/record/11067325/files/figure.png" pageId="7" pageNumber="458">Fig. 16C</figureCitation>
) distinctly wider than in female and with pads of adhesive setae ventrally (
<figureCitation id="74DF4E85FF87FFB0D386D8BCFEC060D6" box="[178,266,219,241]" captionStart="FIGURE" captionStartId="34.[155,167,1521,1540]" captionTargetBox="[264,1031,216,1505]" captionTargetId="figure-129@34.[244,1053,216,1509]" captionTargetPageId="34" captionText="FIGURE 17. Male middle tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067327" httpUri="https://zenodo.org/record/11067327/files/figure.png" pageId="8" pageNumber="459">Fig. 17C</figureCitation>
); median lobe of male genitalia (
<figureCitation id="74DF4E85FF87FFB0D17ED8BCFD0A60D6" box="[586,704,219,241]" captionStart="FIGURE" captionStartId="36.[155,167,1545,1564]" captionTargetBox="[120,1164,216,1527]" captionTargetId="figure-83@36.[120,1176,216,1542]" captionTargetPageId="36" captionText="FIGURE 19. Male genitalia, Tetragonoderus spp. A, B. T. elegans Andrewes (CASENT1039498, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); C, D. T. punctatus (Wiedemann) (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E, F. T. microthorax Jian and Tian (Jamiri-Sessa, Arunachal, India). A, C, E. Dorsal aspect; B, D, F. Left lateral aspect; scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067331" httpUri="https://zenodo.org/record/11067331/files/figure.png" pageId="8" pageNumber="459">Figs. 19A,B</figureCitation>
) with conspicuous large spines on the internal sac....................................................... 
<taxonomicName id="2BE42983FF87FFB0D074D89FFB446129" authority="Andrewes" authorityName="Andrewes" box="[832,1166,248,270]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="DE908E12FF87FFB0D074D89FFBED6129" box="[832,1063,248,270]" italics="true" pageId="8" pageNumber="459">Tetragonoderus elegans</emphasis>
Andrewes
</taxonomicName>
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<paragraph id="EC5B5200FF87FFB0D34FD973FB4561D3" blockId="8.[123,1191,219,500]" pageId="8" pageNumber="459">
<keyLead id="5715F135FF87FFB0D34FD973FB4561D3" pageId="8" pageNumber="459">
4’ Elytral color pattern as in 
<figureCitation id="74DF4E85FF87FFB0D290D973FDCE610D" box="[420,516,276,298]" captionStart="FIGURE" captionStartId="22.[155,167,1587,1606]" captionTargetBox="[128,1176,216,1576]" captionTargetId="figure-59@22.[120,1176,216,1576]" captionTargetPageId="22" captionText="FIGURE 12. Tetragonoderus punctatus (Wiedemann). A. Dorsal aspect, scale line = 1.0 mm (CASENT1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); B. Map showing locality record (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067313" httpUri="https://zenodo.org/record/11067313/files/figure.png" pageId="8" pageNumber="459">Figs. 12A</figureCitation>
, 
<figureCitation id="74DF4E85FF87FFB0D13AD973FDE2610D" box="[526,552,276,298]" captionStart="FIGURE" captionStartId="37.[155,167,1091,1110]" captionTargetBox="[384,875,236,1075]" captionTargetId="figure-14@37.[384,912,216,1079]" captionTargetPageId="37" captionText="FIGURE 20. Tetragonoderus microthorax Jian and Tian, dorsal habitus of holotype (Jinghong, Yunnan, China); scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11067333" httpUri="https://zenodo.org/record/11067333/files/figure.png" pageId="8" pageNumber="459">20</figureCitation>
, with dark areas less expansive; connection between medial portion of the basal macula and humeral macula absent or very narrow, less than one-fifth the length of humeral macula; middle discal band with portion between anterior and posterior dark edges not or only very slightly darker than remainder of pale elytral areas; pronotum piceous or rufopiceous, with or without faint greenish metallic reflection, narrowed basally or not; male with middle mesotarsomeres (
<figureCitation id="74DF4E85FF87FFB0D751D9EFFF2C619C" captionStart="FIGURE" captionStartId="33.[155,167,1521,1540]" captionTargetBox="[215,1075,216,1495]" captionTargetId="figure-133@33.[215,1081,216,1509]" captionTargetPageId="33" captionText="FIGURE 16. Male middle tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067325" httpUri="https://zenodo.org/record/11067325/files/figure.png" pageId="8" pageNumber="459">Figs. 16E,F</figureCitation>
) only slightly wider than in female and with only tarsomeres 1 to 3 with pads of adhesive setae ventrally (i.e., tarsomere 4 without adhesive setae ventrally) (
<figureCitation id="74DF4E85FF87FFB0D1C1D9A5FCA361FF" box="[757,873,450,472]" captionStart="FIGURE" captionStartId="34.[155,167,1521,1540]" captionTargetBox="[264,1031,216,1505]" captionTargetId="figure-129@34.[244,1053,216,1509]" captionTargetPageId="34" captionText="FIGURE 17. Male middle tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067327" httpUri="https://zenodo.org/record/11067327/files/figure.png" pageId="8" pageNumber="459">Figs. 17E,F</figureCitation>
); median lobe of male genitalia (
<figureCitation id="74DF4E85FF87FFB0D386D9B9FF3561D3" box="[178,255,478,500]" captionStart="FIGURE" captionStartId="36.[155,167,1545,1564]" captionTargetBox="[120,1164,216,1527]" captionTargetId="figure-83@36.[120,1176,216,1542]" captionTargetPageId="36" captionText="FIGURE 19. Male genitalia, Tetragonoderus spp. A, B. T. elegans Andrewes (CASENT1039498, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); C, D. T. punctatus (Wiedemann) (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E, F. T. microthorax Jian and Tian (Jamiri-Sessa, Arunachal, India). A, C, E. Dorsal aspect; B, D, F. Left lateral aspect; scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067331" httpUri="https://zenodo.org/record/11067331/files/figure.png" pageId="8" pageNumber="459">Figs. 19</figureCitation>
C-F) without spines on the internal sac........................................... 5
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<paragraph id="EC5B5200FF87FFB0D34FDA6FFB4462B6" blockId="8.[123,1174,520,803]" pageId="8" pageNumber="459">
<keyLead id="5715F135FF87FFB0D34FDA6FFB4462B6" pageId="8" pageNumber="459">
5 (4’) Pronotum (
<figureCitation id="74DF4E85FF87FFB0D217DA6FFEBD6239" box="[291,375,520,542]" captionStart="FIGURE" captionStartId="30.[155,167,1512,1531]" captionTargetBox="[140,1157,216,1497]" captionTargetId="figure-11@30.[140,1157,216,1497]" captionTargetPageId="30" captionText="FIGURE 13. Pronota of cyclosomines, dorsal aspect.A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (CASENT 1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov., (CASENT1039394, holotype, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT 1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067317" httpUri="https://zenodo.org/record/11067317/files/figure.png" pageId="8" pageNumber="459">Fig. 13E</figureCitation>
) slightly shorter and wider (ratio PWM/PL = 1.52 to 1.70), widest near mid-length and posterior to insertion of midlateral seta, not or only very slightly narrowed basally (ratio PWB/PWA = 1.15 to 1.25), lateral margins slightly and evenly convex or straight or very slightly sinuate anterior to basal angles; elytra intervals flat and striae moderately impressed; median lobe of male genitalia (
<figureCitation id="74DF4E85FF87FFB0D751DA39FF2162B6" captionStart="FIGURE" captionStartId="36.[155,167,1545,1564]" captionTargetBox="[120,1164,216,1527]" captionTargetId="figure-83@36.[120,1176,216,1542]" captionTargetPageId="36" captionText="FIGURE 19. Male genitalia, Tetragonoderus spp. A, B. T. elegans Andrewes (CASENT1039498, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); C, D. T. punctatus (Wiedemann) (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E, F. T. microthorax Jian and Tian (Jamiri-Sessa, Arunachal, India). A, C, E. Dorsal aspect; B, D, F. Left lateral aspect; scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067331" httpUri="https://zenodo.org/record/11067331/files/figure.png" pageId="8" pageNumber="459">Figs. 19C,D</figureCitation>
) with apical lamella slightly shorter and broader...... 
<taxonomicName id="2BE42983FF87FFB0D03EDA1CFB4462B6" authority="(Wiedemann)" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[778,1166,635,657]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="punctatus">
<emphasis id="DE908E12FF87FFB0D03EDA1CFBCF62B6" box="[778,1029,635,657]" italics="true" pageId="8" pageNumber="459">Tetragonoderus punctatus</emphasis>
(Wiedemann)
</taxonomicName>
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<paragraph id="EC5B5200FF87FFB0D34FDAFFFB446304" blockId="8.[123,1174,520,803]" pageId="8" pageNumber="459">
<keyLead id="5715F135FF87FFB0D34FDAFFFB446304" pageId="8" pageNumber="459">
5’ Pronotum (
<figureCitation id="74DF4E85FF87FFB0D222DAFFFEA26289" box="[278,360,664,686]" captionStart="FIGURE" captionStartId="30.[155,167,1512,1531]" captionTargetBox="[140,1157,216,1497]" captionTargetId="figure-11@30.[140,1157,216,1497]" captionTargetPageId="30" captionText="FIGURE 13. Pronota of cyclosomines, dorsal aspect.A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (CASENT 1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov., (CASENT1039394, holotype, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT 1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067317" httpUri="https://zenodo.org/record/11067317/files/figure.png" pageId="8" pageNumber="459">Fig. 13F</figureCitation>
) slightly longer and narrower (ratio PWM/PL = 1.48 to 1.57), widest anterior to mid-length at or near insertion of midlateral seta, slightly to moderately narrowed basally (ratio PWB/PWA = 1.04 to 1.14), lateral margins slightly to moderately sinuate anterior to basal angles; elytra intervals slightly convex and striae deeply impressed; median lobe of male genitalia (
<figureCitation id="74DF4E85FF87FFB0D091DA89FBD36323" box="[933,1049,750,772]" captionStart="FIGURE" captionStartId="36.[155,167,1545,1564]" captionTargetBox="[120,1164,216,1527]" captionTargetId="figure-83@36.[120,1176,216,1542]" captionTargetPageId="36" captionText="FIGURE 19. Male genitalia, Tetragonoderus spp. A, B. T. elegans Andrewes (CASENT1039498, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); C, D. T. punctatus (Wiedemann) (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E, F. T. microthorax Jian and Tian (Jamiri-Sessa, Arunachal, India). A, C, E. Dorsal aspect; B, D, F. Left lateral aspect; scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067331" httpUri="https://zenodo.org/record/11067331/files/figure.png" pageId="8" pageNumber="459">Figs. 19E,F</figureCitation>
) with apical lamella slightly longer and narrower................... [
<taxonomicName id="2BE42983FF87FFB0D1CFDB6AFB406304" authority="Jian &amp; Tian" authorityName="Jian &amp; Tian" authorityYear="2009" box="[763,1162,781,803]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="microthorax">
<emphasis id="DE908E12FF87FFB0D1CFDB6AFBC56304" box="[763,1039,781,803]" italics="true" pageId="8" pageNumber="459">Tetragonoderus microthorax</emphasis>
Jian &amp; Tian
</taxonomicName>
]
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>