628 lines
76 KiB
XML
628 lines
76 KiB
XML
<document id="B53DC896F26E5E1CD41D6E70354F545D" ID-CLB-Dataset="299409" ID-DOI="10.5281/zenodo.11067293" ID-GBIF-Dataset="9235db45-04c4-4570-8987-cb84765763a4" ID-ISSN="0068-547X" ID-Zenodo-Dep="11067293" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="karina" IM.treatmentCitations_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1714052207303" checkinUser="felipe" docAuthor="Kavanaugh, David H., Cueva-Dabkoski, Mollie & Liang, Hongbin" docDate="2023" docId="644DE316FF80FFA9D34FDA46FC726554" docLanguage="en" docName="ProcCalifAcadSci.67.20.451-491.pdf" docOrigin="Proceedings of the California Academy of Sciences 67 (20)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Tetragonoderus elegans Andrewes" docType="treatment" docVersion="5" lastPageNumber="468" masterDocId="98749B6EFF8FFFB8D334D867FFCA6027" masterDocTitle="Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, western Yunnan Province, China: Species of the Tribe Cyclosomini Laporte, 1934 (Coleoptera: Carabidae), with Descriptions of Two New Species." masterLastPageNumber="491" masterPageNumber="451" pageNumber="466" updateTime="1719954462017" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="56F5C180614E285E2373B7C5CB89BB31">
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<mods:title id="9D78BEEA3F04953193D3D2AEA31BAB0D">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, western Yunnan Province, China: Species of the Tribe Cyclosomini Laporte, 1934 (Coleoptera: Carabidae), with Descriptions of Two New Species.</mods:title>
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<mods:name id="A6AAB2AC211E08294B4865C4701FE903" type="personal">
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<mods:roleTerm id="AE8948CE8A42A6879F45DEFD51F89A8C">Author</mods:roleTerm>
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<mods:namePart id="6DE94F28C15AA3B5FEE51B6755F1048F">Kavanaugh, David H.</mods:namePart>
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<mods:affiliation id="4F668A0D0AC1402FDC7415C28A257B40">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118.</mods:affiliation>
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</mods:name>
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<mods:name id="3151EFB46373F8D04CC4056DA01BA4A7" type="personal">
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</mods:role>
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<mods:namePart id="7C7FB60A853B27CE2FDD911721CBA223">Cueva-Dabkoski, Mollie</mods:namePart>
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<mods:affiliation id="607BAF35202B0E89BDC29235BFF19B66">Student Science Fellow, California Academy of Sciences, San Francisco, and & 2355 Pearl Street, Santa Monica, CA 90405.</mods:affiliation>
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<mods:name id="AE86B034DC8FDE381DB70A70C14E49E8" type="personal">
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<mods:roleTerm id="6D5A00FC363488FF23A3F84492B65C7C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="50A30035A190703C3EE7DD31873A5D1B">Liang, Hongbin</mods:namePart>
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<mods:affiliation id="189851998449566C9D061F2A615F236B">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="C8CA2D7BF79D535B95D735973D90D46C">text</mods:typeOfResource>
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<mods:title id="78CB29DA911DF05E5BA6B459ABCF2DB0">Proceedings of the California Academy of Sciences</mods:title>
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<mods:part id="8085327E8661496FCAA715E8BB270D1D">
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<mods:date id="31A8DF8FEBE8D8372532BA226ECE27CE">2023</mods:date>
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<mods:number id="D265092DA129321684566D21491E2D56">2023-02-15</mods:number>
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<mods:detail id="A6DA27E4E225F291517A1514A755D929" type="volume">
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<mods:number id="FC2DED56D6933F2B308289BB6B771204">67</mods:number>
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<mods:identifier id="96A664A5E520073EDA695C09C6AB332A" type="DOI">10.5281/zenodo.11067293</mods:identifier>
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<treatment id="644DE316FF80FFA9D34FDA46FC726554" ID-DOI="http://doi.org/10.5281/zenodo.11067323" ID-GBIF-Taxon="234412275" ID-Zenodo-Dep="11067323" LSID="urn:lsid:plazi:treatment:644DE316FF80FFA9D34FDA46FC726554" httpUri="http://treatment.plazi.org/id/644DE316FF80FFA9D34FDA46FC726554" lastPageId="17" lastPageNumber="468" pageId="15" pageNumber="466">
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<subSubSection id="A4FE018BFF80FFB7D34FDA46FD2B627F" pageId="15" pageNumber="466" type="nomenclature">
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<paragraph id="EC5B5200FF80FFB7D34FDA46FE33621E" blockId="15.[123,737,545,600]" box="[123,505,545,569]" pageId="15" pageNumber="466">
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<heading id="B713E56CFF80FFB7D34FDA46FE33621E" box="[123,505,545,569]" fontSize="10" level="2" pageId="15" pageNumber="466" reason="4">
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<taxonomicName id="2BE42983FF80FFB7D34FDA46FE33621E" ID-CoL="7C2YQ" authority="Andrewes" authorityName="Andrewes" box="[123,505,545,569]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="elegans">
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<emphasis id="DE908E12FF80FFB7D34FDA46FE33621E" bold="true" box="[123,505,545,569]" pageId="15" pageNumber="466">
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<emphasis id="DE908E12FF80FFB7D34FDA46FE4B621E" bold="true" box="[123,385,545,569]" italics="true" pageId="15" pageNumber="466">Tetragonoderus elegans</emphasis>
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Andrewes
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="EC5B5200FF80FFB7D34FDA25FD2B627F" blockId="15.[123,737,545,600]" box="[123,737,578,600]" pageId="15" pageNumber="466">
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<figureCitation id="74DF4E85FF80FFB7D34FDA25FF1C627F" box="[123,214,578,600]" captionStart="FIGURE" captionStartId="5.[686,698,839,858]" captionTargetBox="[666,1158,216,826]" captionTargetId="figure-473@5.[652,1176,216,826]" captionText="FIGURE 5. Left hind tibial medial spur and tarsomere 1, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.12628237" httpUri="https://zenodo.org/record/12628237/files/figure.png" pageId="15" pageNumber="466">Figures 5</figureCitation>
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, 6, 10, 13C, 14C, 15C, 16C, 17C, 19A-B, 21, and 23.
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</paragraph>
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</subSubSection>
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<subSubSection id="A4FE018BFF80FFB7D34FDA0BFD84629A" pageId="15" pageNumber="466" type="reference_group">
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<paragraph id="EC5B5200FF80FFB7D34FDA0BFD026287" blockId="15.[123,1174,620,701]" pageId="15" pageNumber="466">
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<treatmentCitationGroup id="CCF4752EFF80FFB7D34FDA0BFD026287" pageId="15" pageNumber="466">
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<treatmentCitation id="6D457411FF80FFB7D34FDA0BFDF462A5" author="ANDREWES, H. E." box="[123,574,620,642]" page="524" pageId="15" pageNumber="466" year="1931">
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<taxonomicName id="2BE42983FF80FFB7D34FDA0BFDF462A5" ID-CoL="7C2YQ" authority="Andrewes, 1931: 524" authorityName="Andrewes" authorityPageNumber="524" authorityYear="1931" box="[123,574,620,642]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="elegans">
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<emphasis id="DE908E12FF80FFB7D34FDA0BFEAF62A5" box="[123,357,620,642]" italics="true" pageId="15" pageNumber="466">Tetragonoderus elegans</emphasis>
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<bibRefCitation id="88752FF1FF80FFB7D25BDA0BFDF462A5" author="ANDREWES, H. E." box="[367,574,620,642]" pageId="15" pageNumber="466" pagination="513 - 528" refId="ref15875" refString="ANDREWES, H. E. 1931. Papers on Oriental Carabidae. XXV. Annals and Magazine of Natural History (Ser. 10) 7: 513 - 528." type="journal article" year="1931">Andrewes, 1931:524</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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<typeStatus id="335FECA2FF80FFB7D178DA0BFD0762A5" box="[588,717,620,642]" pageId="15" pageNumber="466" type="holotype">HOLOTYPE</typeStatus>
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, a female, in NHMUK. Type locality:
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<collectingCountry id="94F31290FF80FFB7D76ADA0BFB5862A5" box="[1118,1170,620,642]" name="India" pageId="15" pageNumber="466">India</collectingCountry>
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,
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<collectingRegion id="2E209CE2FF80FFB7D3AFDAEDFEDE6287" box="[155,276,650,672]" country="India" name="Uttarakhand" pageId="15" pageNumber="466">Uttarakhand</collectingRegion>
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, Dehradun, Bindal River.
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<treatmentCitation id="6D457411FF80FFB7D125DAEDFD0E6287" author="CSIKI, E." box="[529,708,650,672]" page="1296" pageId="15" pageNumber="466" year="1932">
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<bibRefCitation id="88752FF1FF80FFB7D125DAEDFD0E6287" author="CSIKI, E." box="[529,708,650,672]" pageId="15" pageNumber="466" pagination="1279 - 1598" refId="ref16350" refString="CSIKI, E. 1932. Coleopterorum catalogus. Pars 124. Carabidae: Harpalinae VII. W. Junk, Berlin, pp. 1279 - 1598." type="book chapter" year="1932">Csiki (1932:1296)</bibRefCitation>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="EC5B5200FF80FFB7D34FDAC0FD84629A" blockId="15.[123,1174,620,701]" box="[123,590,679,701]" pageId="15" pageNumber="466">
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<treatmentCitationGroup id="CCF4752EFF80FFB7D34FDAC0FD84629A" box="[123,590,679,701]" pageId="15" pageNumber="466">
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<taxonomicName id="2BE42983FF80FFB7D34FDAC0FE52629A" ID-CoL="32T7K" authority="(Andrewes)" baseAuthorityName="Andrewes" box="[123,408,679,701]" class="Insecta" family="Carabidae" genus="Cyclicus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="elegans">
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<emphasis id="DE908E12FF80FFB7D34FDAC0FED7629A" box="[123,285,679,701]" italics="true" pageId="15" pageNumber="466">Cyclicus elegans</emphasis>
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(Andrewes)
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</taxonomicName>
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,
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<treatmentCitation id="6D457411FF80FFB7D296DACFFD80629A" author="LORENZ, W." box="[418,586,679,701]" page="453" pageId="15" pageNumber="466" year="2005">
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<bibRefCitation id="88752FF1FF80FFB7D296DACFFD80629A" author="LORENZ, W." box="[418,586,679,701]" pageId="15" pageNumber="466" refId="ref17529" refString="LORENZ, W. 2005. Systematic List of Extant Ground Beetles of the World (Insecta Coleoptera " Geadephaga ": Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second edition. W. Lorenz, Tutzing, 530 pp." type="book" year="2005">Lorenz 2005:453</bibRefCitation>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="A4FE018BFF80FFA9D39FDAB5FC726554" lastPageId="17" lastPageNumber="468" pageId="15" pageNumber="466" type="diagnosis">
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<paragraph id="EC5B5200FF80FFB7D39FDAB5FBAD6456" blockId="15.[123,1174,721,1726]" pageId="15" pageNumber="466">
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<emphasis id="DE908E12FF80FFB7D39FDAB5FED562CE" bold="true" box="[171,287,722,745]" pageId="15" pageNumber="466">Diagnosis.</emphasis>
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Adults of
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<taxonomicName id="2BE42983FF80FFB7D2A5DAB5FE3762CE" box="[401,509,721,745]" class="Insecta" family="Carabidae" genus="Cyclicus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="elegans">
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<emphasis id="DE908E12FF80FFB7D2A5DAB5FE3762CE" box="[401,509,721,745]" italics="true" pageId="15" pageNumber="466">T. elegans</emphasis>
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</taxonomicName>
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can be distinguished from those of other
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<emphasis id="DE908E12FF80FFB7D089DAB6FB8B62CE" box="[957,1089,721,745]" italics="true" pageId="15" pageNumber="466">cyclosomine</emphasis>
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species in the study area by the following combination of character states: Body size medium for genus, BL males =
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<quantity id="2B1CFFE5FF80FFB7D3CBDB75FE53630D" box="[255,409,786,810]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" metricValueMax="5.2" metricValueMin="4.4" pageId="15" pageNumber="466" unit="mm" value="4.8" valueMax="5.2" valueMin="4.4">4.4 to 5.2 mm</quantity>
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, females
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<quantity id="2B1CFFE5FF80FFB7D2CBDB75FD50630D" box="[511,666,786,810]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" metricValueMax="5.6" metricValueMin="4.6" pageId="15" pageNumber="466" unit="mm" value="5.1" valueMax="5.6" valueMin="4.6">4.6 to 5.6 mm</quantity>
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; pronotum dark, black or piceous, with distinct greenish or bronze metallic reflection, slightly narrowed basally (ratio PWM/PWB =1.13 to 1.20); elytral color pattern as in
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<figureCitation id="74DF4E85FF80FFB7D2A1DB33FE32634B" box="[405,504,852,876]" captionStart="FIGURE" captionStartId="16.[155,167,1603,1622]" captionTargetBox="[120,1176,216,1581]" captionTargetId="figure-60@16.[120,1176,216,1590]" captionTargetPageId="16" captionText="FIGURE 10. Tetragonoderus elegans Andrewes. A. Dorsal aspect, scale line = 1.0 mm (CASENT1039409, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); B. Map showing locality records (black circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067305" httpUri="https://zenodo.org/record/11067305/files/figure.png" pageId="15" pageNumber="466">Fig. 10A</figureCitation>
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, with dark areas more expansive; connection between medial portion of the basal dark band and humeral portion broader, at least one-quarter length of the humeral portion; middle band with portion between anterior and posterior dark edges distinctly darker (orange-brown) than remainder of pale elytral areas; femora pale tan; pronotum widest at or anterior to middle, slightly narrowed basally (ratio PWM/PWB =1.13 to 1.20); elytra slightly to distinctly and obliquely truncate apically; front tarsomeres 1 to 3 without lateral expansions (
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<figureCitation id="74DF4E85FF80FFB7D754DB90FF5F6417" captionStart="FIGURE" captionStartId="31.[155,167,1512,1531]" captionTargetBox="[253,1040,238,1497]" captionTargetId="figure-15@31.[253,1044,216,1497]" captionTargetPageId="31" captionText="FIGURE 14. Male front tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067319" httpUri="https://zenodo.org/record/11067319/files/figure.png" pageId="15" pageNumber="466">Figs. 14</figureCitation>
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CA, 15C); male with middle tarsomeres 1 to 4 (
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<figureCitation id="74DF4E85FF80FFB7D191DC7FFCCD6417" box="[677,775,1048,1072]" captionStart="FIGURE" captionStartId="33.[155,167,1521,1540]" captionTargetBox="[215,1075,216,1495]" captionTargetId="figure-133@33.[215,1081,216,1509]" captionTargetPageId="33" captionText="FIGURE 16. Male middle tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067325" httpUri="https://zenodo.org/record/11067325/files/figure.png" pageId="15" pageNumber="466">Fig. 16C</figureCitation>
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) distinctly wider than in female and with pads of adhesive setae ventrally (
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<figureCitation id="74DF4E85FF80FFB7D123DC5FFDB36477" box="[535,633,1080,1104]" captionStart="FIGURE" captionStartId="34.[155,167,1521,1540]" captionTargetBox="[264,1031,216,1505]" captionTargetId="figure-129@34.[244,1053,216,1509]" captionTargetPageId="34" captionText="FIGURE 17. Male middle tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067327" httpUri="https://zenodo.org/record/11067327/files/figure.png" pageId="15" pageNumber="466">Fig. 17C</figureCitation>
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); median lobe of male genitalia (
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<figureCitation id="74DF4E85FF80FFB7D0E9DC5FFB9C6477" box="[989,1110,1080,1104]" captionStart="FIGURE" captionStartId="36.[155,167,1545,1564]" captionTargetBox="[120,1164,216,1527]" captionTargetId="figure-83@36.[120,1176,216,1542]" captionTargetPageId="36" captionText="FIGURE 19. Male genitalia, Tetragonoderus spp. A, B. T. elegans Andrewes (CASENT1039498, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); C, D. T. punctatus (Wiedemann) (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E, F. T. microthorax Jian and Tian (Jamiri-Sessa, Arunachal, India). A, C, E. Dorsal aspect; B, D, F. Left lateral aspect; scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067331" httpUri="https://zenodo.org/record/11067331/files/figure.png" pageId="15" pageNumber="466">Fig. 19A,B</figureCitation>
|
|
) with apical lamella long and bent ventrally, internal sac with several large and conspicuous spines.
|
|
</paragraph>
|
|
<paragraph id="EC5B5200FF80FFB7D39FDC1CFB5C64D3" blockId="15.[123,1174,721,1726]" pageId="15" pageNumber="466">
|
|
In the study area, specimens of
|
|
<taxonomicName id="2BE42983FF80FFB7D2CEDC1CFDAF64B5" box="[506,613,1146,1170]" class="Insecta" family="Carabidae" genus="Cyclicus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="elegans">
|
|
<emphasis id="DE908E12FF80FFB7D2CEDC1CFDAF64B5" box="[506,613,1146,1170]" italics="true" pageId="15" pageNumber="466">T. elegans</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<figureCitation id="74DF4E85FF80FFB7D140DC1DFD1F64B5" box="[628,725,1146,1170]" captionStart="FIGURE" captionStartId="16.[155,167,1603,1622]" captionTargetBox="[120,1176,216,1581]" captionTargetId="figure-60@16.[120,1176,216,1590]" captionTargetPageId="16" captionText="FIGURE 10. Tetragonoderus elegans Andrewes. A. Dorsal aspect, scale line = 1.0 mm (CASENT1039409, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); B. Map showing locality records (black circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067305" httpUri="https://zenodo.org/record/11067305/files/figure.png" pageId="15" pageNumber="466">Fig. 10A</figureCitation>
|
|
) might be confused with those of
|
|
<taxonomicName id="2BE42983FF80FFB7D70BDC1CFF656494" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="punctatus">
|
|
<emphasis id="DE908E12FF80FFB7D70BDC1CFF656494" italics="true" pageId="15" pageNumber="466">T. punctatus</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<figureCitation id="74DF4E85FF80FFB7D3F4DCFDFEE96495" box="[192,291,1178,1202]" captionStart="FIGURE" captionStartId="22.[155,167,1587,1606]" captionTargetBox="[128,1176,216,1576]" captionTargetId="figure-59@22.[120,1176,216,1576]" captionTargetPageId="22" captionText="FIGURE 12. Tetragonoderus punctatus (Wiedemann). A. Dorsal aspect, scale line = 1.0 mm (CASENT1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); B. Map showing locality record (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067313" httpUri="https://zenodo.org/record/11067313/files/figure.png" pageId="15" pageNumber="466">Fig. 12A</figureCitation>
|
|
) and also those of
|
|
<taxonomicName id="2BE42983FF80FFB7D2CDDCFBFD526495" authorityName="Jian & Tian" authorityYear="2009" box="[505,664,1178,1203]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="microthorax">
|
|
<emphasis id="DE908E12FF80FFB7D2CDDCFBFD526495" box="[505,664,1178,1203]" italics="true" pageId="15" pageNumber="466">T. microthorax</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<figureCitation id="74DF4E85FF80FFB7D19DDCFDFD366495" box="[681,764,1178,1202]" captionStart="FIGURE" captionStartId="37.[155,167,1091,1110]" captionTargetBox="[384,875,236,1075]" captionTargetId="figure-14@37.[384,912,216,1079]" captionTargetPageId="37" captionText="FIGURE 20. Tetragonoderus microthorax Jian and Tian, dorsal habitus of holotype (Jinghong, Yunnan, China); scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11067333" httpUri="https://zenodo.org/record/11067333/files/figure.png" pageId="15" pageNumber="466">Fig. 20</figureCitation>
|
|
), although the latter has not yet been recorded from the study area. Refer to the key for features distinguishing members of these three species and the Diagnosis section for
|
|
<taxonomicName id="2BE42983FF80FFB7D13CDCBAFD4064D3" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[520,650,1245,1268]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="punctatus">
|
|
<emphasis id="DE908E12FF80FFB7D13CDCBAFD4064D3" box="[520,650,1245,1268]" italics="true" pageId="15" pageNumber="466">T. punctatus</emphasis>
|
|
</taxonomicName>
|
|
below for further discussion of these differences.
|
|
</paragraph>
|
|
<paragraph id="EC5B5200FF80FFB7D39FDC9AFB5C663C" blockId="15.[123,1174,721,1726]" pageId="15" pageNumber="466">
|
|
<emphasis id="DE908E12FF80FFB7D39FDC9AFE5C6533" bold="true" box="[171,406,1277,1300]" pageId="15" pageNumber="466">Habitat distribution.</emphasis>
|
|
Within the study area, members of this species have been found only on open sandy shores of medium- to large-sized rivers, where they remain buried in the sand or under cover during daylight hours and are active on the sand surface in moist areas at night. A few specimens have been collected during the day from under small stones and drift debris in the same habitat. At night, these beetles are active in upper beach areas, where zones of slightly moist and dry sand meet (
|
|
<figureCitation id="74DF4E85FF80FFB7D3C1DDC7FE8F659F" box="[245,325,1440,1464]" captionStart="FIGURE" captionStartId="38.[155,167,1623,1642]" captionTargetBox="[136,1161,216,1607]" captionTargetId="figure-10@38.[136,1161,216,1607]" captionTargetPageId="38" captionText="FIGURE 21. Photographs of habitat for cyclosomine species, Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov., Tetragonoderus elegans Andrewes, T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov., and T. punctatus Wiedemann at Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China. A. lower river bank; B. sandy upper bank with sparse vegetation. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.11067337" httpUri="https://zenodo.org/record/11067337/files/figure.png" pageId="15" pageNumber="466">Fig. 21</figureCitation>
|
|
). Members of this species were found at night together with
|
|
<taxonomicName id="2BE42983FF80FFB7D0F9DDC6FB85659F" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[973,1103,1441,1464]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="punctatus">
|
|
<emphasis id="DE908E12FF80FFB7D0F9DDC6FB85659F" box="[973,1103,1441,1464]" italics="true" pageId="15" pageNumber="466">T. punctatus</emphasis>
|
|
</taxonomicName>
|
|
adults at two localities along the Longchuan River and in daytime under stones in the same habitat with
|
|
<taxonomicName id="2BE42983FF80FFB7D34FDD85FEC265DE" authorityName="Kavanaugh and Cueva-Dabkoski" authorityYear="2023" box="[123,264,1505,1529]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="parviculus">
|
|
<emphasis id="DE908E12FF80FFB7D34FDD85FEC265DE" box="[123,264,1505,1529]" italics="true" pageId="15" pageNumber="466">T. parviculus</emphasis>
|
|
</taxonomicName>
|
|
adults. Within the Gaoligong
|
|
<collectingRegion id="2E209CE2FF80FFB7D165DD86FD1C65DE" box="[593,726,1505,1529]" country="Myanmar" name="Shan" pageId="15" pageNumber="466">Shan region</collectingRegion>
|
|
, this species occurs at relatively low elevations, below
|
|
<quantity id="2B1CFFE5FF80FFB7D22ADE65FEBB663D" box="[286,369,1538,1562]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="466" unit="m" value="2000.0">2000 m</quantity>
|
|
, with our records documenting its occurrence in the
|
|
<quantity id="2B1CFFE5FF80FFB7D090DE65FB87663D" box="[932,1101,1538,1562]" metricMagnitude="3" metricUnit="m" metricValue="1.5375" metricValueMax="1.89" metricValueMin="1.185" pageId="15" pageNumber="466" unit="m" value="1537.5" valueMax="1890.0" valueMin="1185.0">1185 to 1890 m</quantity>
|
|
range.
|
|
</paragraph>
|
|
<paragraph id="EC5B5200FF80FFA9D39FDE44FE916274" blockId="15.[123,1174,721,1726]" lastBlockId="17.[123,1174,219,1396]" lastPageId="17" lastPageNumber="468" pageId="15" pageNumber="466">
|
|
<emphasis id="DE908E12FF80FFB7D39FDE44FD18661D" bold="true" box="[171,722,1571,1594]" pageId="15" pageNumber="466">Geographical distribution within the Gaoligong</emphasis>
|
|
<emphasis id="DE908E12FF80FFB7D1EFDE44FCD6661D" bold="true" box="[731,796,1571,1594]" pageId="15" pageNumber="466">Shan.</emphasis>
|
|
<figureCitation id="74DF4E85FF80FFB7D011DE45FC42661D" box="[805,904,1570,1594]" captionStart="FIGURE" captionStartId="16.[155,167,1603,1622]" captionTargetBox="[120,1176,216,1581]" captionTargetId="figure-60@16.[120,1176,216,1590]" captionTargetPageId="16" captionText="FIGURE 10. Tetragonoderus elegans Andrewes. A. Dorsal aspect, scale line = 1.0 mm (CASENT1039409, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); B. Map showing locality records (black circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067305" httpUri="https://zenodo.org/record/11067305/files/figure.png" pageId="15" pageNumber="466">Fig. 10B</figureCitation>
|
|
.
|
|
<materialsCitation id="5C8C585DFF80FFB7D0A0DE44FCDC66BB" ID-GBIF-Occurrence="4902827319" collectingDate="2002-10-13" collectionCode="CAS" collectorName="We & D. H. Kavanaugh & P. E. Marek & H. B. Liang & D. Z. Dong" county="Fugong County" elevation="1185" location="Nu Jiang" municipality="Shangpa Township" pageId="15" pageNumber="466" specimenCount="215" specimenCount-female="66" specimenCount-male="42" stateProvince="Shan">
|
|
We examined a total of
|
|
<specimenCount id="FAE29989FF80FFB7D34FDE24FED3667C" box="[123,281,1603,1627]" count="107" pageId="15" pageNumber="466" type="generic">107 specimens</specimenCount>
|
|
(
|
|
<specimenCount id="FAE29989FF80FFB7D21DDE24FE41667C" box="[297,395,1603,1627]" count="42" pageId="15" pageNumber="466" type="male">42 males</specimenCount>
|
|
and
|
|
<specimenCount id="FAE29989FF80FFB7D2F5DE24FDF2667C" box="[449,568,1603,1627]" count="65" pageId="15" pageNumber="466" type="female">65 females</specimenCount>
|
|
) from the following localities:
|
|
<emphasis id="DE908E12FF80FFB7D0BEDE23FB88667B" bold="true" box="[906,1090,1604,1628]" italics="true" pageId="15" pageNumber="466">
|
|
<collectingCounty id="053A2A8CFF80FFB7D0BEDE23FBF1667B" box="[906,1083,1604,1628]" pageId="15" pageNumber="466">Fugong County</collectingCounty>
|
|
:
|
|
</emphasis>
|
|
<collectingMunicipality id="0C3FC87AFF80FFB7D77EDE24FEC2665B" pageId="15" pageNumber="466">Shangpa Township</collectingMunicipality>
|
|
(west bank of
|
|
<location id="E93B04DBFF80FFB7D28FDE02FDE9665B" LSID="urn:lsid:plazi:treatment:644DE316FF80FFA9D34FDA46FC726554:E93B04DBFF80FFB7D28FDE02FDE9665B" box="[443,547,1636,1660]" county="Fugong County" municipality="Shangpa Township" name="Nu Jiang" pageId="15" pageNumber="466" stateProvince="Shan">Nu Jiang</location>
|
|
, 26.90668°/98.86339°,
|
|
<quantity id="2B1CFFE5FF80FFB7D01EDE03FC4B665B" box="[810,897,1636,1660]" metricMagnitude="3" metricUnit="m" metricValue="1.185" pageId="15" pageNumber="466" unit="m" value="1185.0">
|
|
<elevation id="67C9B533FF80FFB7D01EDE03FC4B665B" box="[810,897,1636,1660]" metricMagnitude="3" metricUnit="m" metricValue="1.185" pageId="15" pageNumber="466" unit="m" value="1185.0">1185 m</elevation>
|
|
</quantity>
|
|
,
|
|
<date id="985A74C0FF80FFB7D0A5DE03FB98665B" box="[913,1106,1636,1660]" pageId="15" pageNumber="466" value="2002-10-13">
|
|
<collectingDate id="881E8D28FF80FFB7D0A5DE03FB98665B" box="[913,1106,1636,1660]" pageId="15" pageNumber="466" value="2002-10-13">13 October 2002</collectingDate>
|
|
</date>
|
|
,
|
|
<collectorName id="411137D6FF80FFB7D756DE02FF3D66BB" pageId="15" pageNumber="466">D.H. Kavanaugh</collectorName>
|
|
,
|
|
<collectorName id="411137D6FF80FFB7D236DEE2FEB166BB" box="[258,379,1668,1692]" pageId="15" pageNumber="466">P.E. Marek</collectorName>
|
|
,
|
|
<collectorName id="411137D6FF80FFB7D2B3DEE2FE3566BB" box="[391,511,1668,1692]" pageId="15" pageNumber="466">H.B. Liang</collectorName>
|
|
&
|
|
<collectorName id="411137D6FF80FFB7D117DEE2FD5C66BA" box="[547,662,1669,1693]" pageId="15" pageNumber="466">D.Z. Dong</collectorName>
|
|
collectors [
|
|
</materialsCitation>
|
|
<materialsCitation id="5C8C585DFF80FFB7D023DEE3FC0166BB" ID-GBIF-Occurrence="4902827326" box="[791,971,1668,1692]" collectingDate="2002-10-13" collectionCode="CAS" collectorName="We & D. H. Kavanaugh & P. E. Marek & H. B. Liang & D. Z. Dong" county="Fugong County" elevation="1185" location="Nu Jiang" municipality="Shangpa Township" pageId="15" pageNumber="466" specimenCount="215" specimenCount-female="66" specimenCount-male="42" stateProvince="Shan">
|
|
<specimenCount id="FAE29989FF80FFB7D023DEE3FCBD66BB" box="[791,887,1668,1692]" count="1" pageId="15" pageNumber="466" type="female">1 female</specimenCount>
|
|
;
|
|
<collectionCode id="8AF5CAC5FF80FFB7D0B6DEE3FC7666BB" box="[898,956,1668,1692]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="15" pageNumber="466" type="Museum">CAS</collectionCode>
|
|
])
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="5C8C585DFF80FFA9D0E2DEE2FD0760D4" ID-GBIF-Occurrence="4902827336" collectingDate="2002-10-08" collectingDateMax="2002-10-09" collectingDateMin="2002-10-08" collectionCode="CAS, IOZ" collectorName="Nu Jiang & D. H. Kavanaugh & P. E. Marek & Liang" county="Gongshan County" elevation="1430" lastPageId="17" lastPageNumber="468" location="Dashada" municipality="Cikai Township" pageId="15" pageNumber="466" specimenCount="19" specimenCount-female="13" specimenCount-male="6" stateProvince="Shan">
|
|
<emphasis id="DE908E12FF80FFB7D0E2DEE2FF5D6699" bold="true" italics="true" pageId="15" pageNumber="466">
|
|
<collectingCounty id="053A2A8CFF80FFB7D0E2DEE2FF5A6699" pageId="15" pageNumber="466">Gongshan County</collectingCounty>
|
|
:
|
|
</emphasis>
|
|
<collectingMunicipality id="0C3FC87AFF80FFB7D3AADEC2FE82669A" box="[158,328,1701,1725]" pageId="15" pageNumber="466">Cikai Township</collectingMunicipality>
|
|
(
|
|
<collectorName id="411137D6FF80FFB7D262DEC2FE72669A" box="[342,440,1701,1725]" pageId="15" pageNumber="466">Nu Jiang</collectorName>
|
|
at
|
|
<location id="E93B04DBFF80FFB7D2EEDEC2FDF3669A" LSID="urn:lsid:plazi:treatment:644DE316FF80FFA9D34FDA46FC726554:E93B04DBFF80FFB7D2EEDEC2FDF3669A" box="[474,569,1701,1725]" county="Gongshan County" municipality="Cikai Township" name="Dashada" pageId="15" pageNumber="466" stateProvince="Shan">Dashada</location>
|
|
, 27.73845°/98.67092°,
|
|
<quantity id="2B1CFFE5FF80FFB7D003DEC2FC40669A" box="[823,906,1701,1725]" metricMagnitude="3" metricUnit="m" metricValue="1.43" pageId="15" pageNumber="466" unit="m" value="1430.0">
|
|
<elevation id="67C9B533FF80FFB7D003DEC2FC40669A" box="[823,906,1701,1725]" metricMagnitude="3" metricUnit="m" metricValue="1.43" pageId="15" pageNumber="466" unit="m" value="1430.0">1430 m</elevation>
|
|
</quantity>
|
|
,
|
|
<date id="985A74C0FF80FFB7D0A2DEC2FB9C669A" box="[918,1110,1701,1725]" pageId="15" pageNumber="466" value="2002-10-08" valueMax="2002-10-09" valueMin="2002-10-08">
|
|
<collectingDate id="881E8D28FF80FFB7D0A2DEC2FB9C669A" box="[918,1110,1701,1725]" pageId="15" pageNumber="466" value="2002-10-08" valueMax="2002-10-09" valueMin="2002-10-08">8-9 October 2002</collectingDate>
|
|
</date>
|
|
,
|
|
<collectorName id="411137D6FF80FFA9D756DEC1FF3D60D4" lastPageId="17" lastPageNumber="468" pageId="15" pageNumber="466">D.H. Kavanaugh</collectorName>
|
|
,
|
|
<collectorName id="411137D6FF9EFFA9D23FD8BCFE4060D4" box="[267,394,219,243]" pageId="17" pageNumber="468">P.E. Marek</collectorName>
|
|
& H.B.
|
|
<collectorName id="411137D6FF9EFFA9D134D8BCFDF460D4" box="[512,574,219,243]" pageId="17" pageNumber="468">Liang</collectorName>
|
|
collectors [
|
|
</materialsCitation>
|
|
<materialsCitation id="5C8C585DFF9EFFA9D1F9D8BCFB5B60D4" ID-GBIF-Occurrence="4902827322" box="[717,1169,219,243]" collectingDate="2002-10-08" collectingDateMax="2002-10-09" collectingDateMin="2002-10-08" collectionCode="CAS, IOZ" collectorName="Nu Jiang & D. H. Kavanaugh & P. E. Marek & Liang" county="Gongshan County" elevation="1430" location="Dashada" municipality="Cikai Township" pageId="17" pageNumber="468" specimenCount="19" specimenCount-female="13" specimenCount-male="6" stateProvince="Shan">
|
|
<specimenCount id="FAE29989FF9EFFA9D1F9D8BCFCE060D4" box="[717,810,219,243]" count="6" pageId="17" pageNumber="468" type="male">6 males</specimenCount>
|
|
and
|
|
<specimenCount id="FAE29989FF9EFFA9D045D8BCFC3B60D4" box="[881,1009,219,243]" count="13" pageId="17" pageNumber="468" type="female">13 females</specimenCount>
|
|
;
|
|
<collectionCode id="8AF5CAC5FF9EFFA9D731D8BCFBF160D4" box="[1029,1083,219,243]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="17" pageNumber="468" type="Museum">CAS</collectionCode>
|
|
,
|
|
<collectionCode id="8AF5CAC5FF9EFFA9D764D8BCFB4960D4" box="[1104,1155,219,243]" pageId="17" pageNumber="468">IOZ</collectionCode>
|
|
])
|
|
</materialsCitation>
|
|
;
|
|
<materialsCitation id="5C8C585DFF9EFFA9D34FD89CFEA16114" ID-GBIF-Occurrence="4902827339" collectingDate="2004-10-30" collectingDateMax="2021-05-30" collectingDateMin="2004-10-30" collectionCode="CAS, IOZ" collectorName="H. B. Liang & Silalong He & D. H. Kavanaugh & G. Tang & D. Z. Dong & Kongdang & H. B Liang & Y. Xu & Pukawang & N, H. B Liang" county="Dulongjiang Township" elevation="1355" location="Mulangdang" municipality="Bapo" pageId="17" pageNumber="468" specimenCount="33" specimenCount-female="20" specimenCount-male="13" stateProvince="Shan">
|
|
<collectingCounty id="053A2A8CFF9EFFA9D34FD89CFEB96134" box="[123,371,251,275]" pageId="17" pageNumber="468">Dulongjiang Township</collectingCounty>
|
|
(
|
|
<collectingMunicipality id="0C3FC87AFF9EFFA9D2B2D89CFE086134" box="[390,450,251,275]" pageId="17" pageNumber="468">Bapo</collectingMunicipality>
|
|
,
|
|
<location id="E93B04DBFF9EFFA9D2E5D89CFD916134" LSID="urn:lsid:plazi:treatment:644DE316FF80FFA9D34FDA46FC726554:E93B04DBFF9EFFA9D2E5D89CFD916134" box="[465,603,251,275]" county="Dulongjiang Township" municipality="Bapo" name="Mulangdang" pageId="17" pageNumber="468" stateProvince="Shan">Mulangdang</location>
|
|
, 27.75256°/98.34745°,
|
|
<quantity id="2B1CFFE5FF9EFFA9D06BD89CFC7C6134" box="[863,950,251,275]" metricMagnitude="3" metricUnit="m" metricValue="1.355" pageId="17" pageNumber="468" unit="m" value="1355.0">
|
|
<elevation id="67C9B533FF9EFFA9D06BD89CFC7C6134" box="[863,950,251,275]" metricMagnitude="3" metricUnit="m" metricValue="1.355" pageId="17" pageNumber="468" unit="m" value="1355.0">1355 m</elevation>
|
|
</quantity>
|
|
,
|
|
<date id="985A74C0FF9EFFA9D0F2D89CFB5B6134" box="[966,1169,251,275]" pageId="17" pageNumber="468" value="2004-11-04">
|
|
<collectingDate id="881E8D28FF9EFFA9D0F2D89CFB5B6134" box="[966,1169,251,275]" pageId="17" pageNumber="468" value="2004-11-04">4 November 2004</collectingDate>
|
|
</date>
|
|
,
|
|
<collectorName id="411137D6FF9EFFA9D34FD97BFF3E6114" box="[123,244,283,307]" pageId="17" pageNumber="468">H.B. Liang</collectorName>
|
|
collector [
|
|
</materialsCitation>
|
|
<materialsCitation id="5C8C585DFF9EFFA9D25FD97CFEAD6154" ID-GBIF-Occurrence="4902827323" collectingDate="2004-10-30" collectingDateMax="2021-05-30" collectingDateMin="2004-10-30" collectionCode="CAS, IOZ" collectorName="H. B. Liang & Silalong He & D. H. Kavanaugh & G. Tang & D. Z. Dong & Kongdang & H. B Liang & Y. Xu & Pukawang & N, H. B Liang" county="Dulongjiang Township" elevation="1355" location="Mulangdang" municipality="Bapo" pageId="17" pageNumber="468" specimenCount="33" specimenCount-female="20" specimenCount-male="13" stateProvince="Shan">
|
|
<specimenCount id="FAE29989FF9EFFA9D25FD97CFE0A6114" box="[363,448,283,307]" count="3" pageId="17" pageNumber="468" type="male">3 males</specimenCount>
|
|
and
|
|
<specimenCount id="FAE29989FF9EFFA9D2CDD97CFDBB6114" box="[505,625,283,307]" count="11" pageId="17" pageNumber="468" type="female">11 females</specimenCount>
|
|
;
|
|
<collectionCode id="8AF5CAC5FF9EFFA9D14BD97CFD7F6114" box="[639,693,283,307]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="17" pageNumber="468" type="Museum">CAS</collectionCode>
|
|
,
|
|
<collectionCode id="8AF5CAC5FF9EFFA9D1F6D97CFD3E6114" box="[706,756,283,307]" pageId="17" pageNumber="468">IOZ</collectionCode>
|
|
]), (south edge of
|
|
<location id="E93B04DBFF9EFFA9D082D97CFB5C6114" LSID="urn:lsid:plazi:treatment:644DE316FF80FFA9D34FDA46FC726554:E93B04DBFF9EFFA9D082D97CFB5C6114" box="[950,1174,283,307]" county="Dulongjiang Township" municipality="Bapo" name="Dizhengdang village" pageId="17" pageNumber="468" stateProvince="Shan">Dizhengdang village</location>
|
|
along
|
|
<collectorName id="411137D6FF9EFFA9D389D95CFE8A6174" box="[189,320,315,339]" pageId="17" pageNumber="468">Silalong He</collectorName>
|
|
, 28.07654°/98.32603°,
|
|
<quantity id="2B1CFFE5FF9EFFA9D10AD95CFD586174" box="[574,658,315,339]" metricMagnitude="3" metricUnit="m" metricValue="1.89" pageId="17" pageNumber="468" unit="m" value="1890.0">
|
|
<elevation id="67C9B533FF9EFFA9D10AD95CFD586174" box="[574,658,315,339]" metricMagnitude="3" metricUnit="m" metricValue="1.89" pageId="17" pageNumber="468" unit="m" value="1890.0">1890 m</elevation>
|
|
</quantity>
|
|
,
|
|
<date id="985A74C0FF9EFFA9D1AAD95CFC9F6174" box="[670,853,315,339]" pageId="17" pageNumber="468" value="2004-10-30">
|
|
<collectingDate id="881E8D28FF9EFFA9D1AAD95CFC9F6174" box="[670,853,315,339]" pageId="17" pageNumber="468" value="2004-10-30">30 October 2004</collectingDate>
|
|
</date>
|
|
,
|
|
<collectorName id="411137D6FF9EFFA9D055D95BFBD26174" box="[865,1048,315,339]" pageId="17" pageNumber="468">D.H. Kavanaugh</collectorName>
|
|
,
|
|
<collectorName id="411137D6FF9EFFA9D717D95CFBB36173" box="[1059,1145,315,340]" pageId="17" pageNumber="468">G. Tang</collectorName>
|
|
&
|
|
<collectorName id="411137D6FF9EFFA9D34FD93BFF266153" box="[123,236,348,372]" pageId="17" pageNumber="468">D.Z. Dong</collectorName>
|
|
collectors [
|
|
</materialsCitation>
|
|
<materialsCitation id="5C8C585DFF9EFFA9D253D93CFD6A61B4" ID-GBIF-Occurrence="4902827338" collectingDate="2004-10-30" collectingDateMax="2021-05-30" collectingDateMin="2004-10-30" collectionCode="CAS, IOZ" collectorName="H. B. Liang & Silalong He & D. H. Kavanaugh & G. Tang & D. Z. Dong & Kongdang & H. B Liang & Y. Xu & Pukawang & N, H. B Liang" county="Dulongjiang Township" elevation="1355" location="Mulangdang" municipality="Bapo" pageId="17" pageNumber="468" specimenCount="33" specimenCount-female="20" specimenCount-male="13" stateProvince="Shan">
|
|
<specimenCount id="FAE29989FF9EFFA9D253D93CFE706154" box="[359,442,347,371]" count="2" pageId="17" pageNumber="468" type="male">2 males</specimenCount>
|
|
and
|
|
<specimenCount id="FAE29989FF9EFFA9D2D8D93CFD9F6154" box="[492,597,347,371]" count="3" pageId="17" pageNumber="468" type="female">3 females</specimenCount>
|
|
;
|
|
<collectionCode id="8AF5CAC5FF9EFFA9D16BD93CFD5F6154" box="[607,661,347,371]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="17" pageNumber="468" type="Museum">CAS</collectionCode>
|
|
,
|
|
<collectionCode id="8AF5CAC5FF9EFFA9D194D93CFD186154" box="[672,722,347,371]" pageId="17" pageNumber="468">IOZ</collectionCode>
|
|
]), (
|
|
<collectorName id="411137D6FF9EFFA9D1C6D93CFCAF6154" box="[754,869,347,371]" pageId="17" pageNumber="468">Kongdang</collectorName>
|
|
, 27.89791°/98.33843°,
|
|
<quantity id="2B1CFFE5FF9EFFA9D754D93CFF5861B4" metricMagnitude="3" metricUnit="m" metricValue="1.581" pageId="17" pageNumber="468" unit="m" value="1581.0">
|
|
<elevation id="67C9B533FF9EFFA9D754D93CFF5861B4" metricMagnitude="3" metricUnit="m" metricValue="1.581" pageId="17" pageNumber="468" unit="m" value="1581.0">1581 m</elevation>
|
|
</quantity>
|
|
,
|
|
<date id="985A74C0FF9EFFA9D3A9D91CFEE761B4" box="[157,301,379,404]" pageId="17" pageNumber="468" value="2021-05-30">
|
|
<collectingDate id="881E8D28FF9EFFA9D3A9D91CFEE761B4" box="[157,301,379,404]" pageId="17" pageNumber="468" value="2021-05-30">30 May 2021</collectingDate>
|
|
</date>
|
|
,
|
|
<collectorName id="411137D6FF9EFFA9D20CD91BFE6261B4" box="[312,424,379,403]" pageId="17" pageNumber="468">H.B Liang</collectorName>
|
|
&
|
|
<collectorName id="411137D6FF9EFFA9D2FDD91BFDCD61B4" box="[457,519,380,403]" pageId="17" pageNumber="468">Y. Xu</collectorName>
|
|
Y. collectors [
|
|
</materialsCitation>
|
|
<materialsCitation id="5C8C585DFF9EFFA9D194D91CFC6E6194" ID-GBIF-Occurrence="4902827321" collectingDate="2004-10-30" collectingDateMax="2021-05-30" collectingDateMin="2004-10-30" collectionCode="CAS, IOZ" collectorName="H. B. Liang & Silalong He & D. H. Kavanaugh & G. Tang & D. Z. Dong & Kongdang & H. B Liang & Y. Xu & Pukawang & N, H. B Liang" county="Dulongjiang Township" elevation="1355" location="Mulangdang" municipality="Bapo" pageId="17" pageNumber="468" specimenCount="33" specimenCount-female="20" specimenCount-male="13" stateProvince="Shan">
|
|
<specimenCount id="FAE29989FF9EFFA9D194D91CFD3F61B4" box="[672,757,379,403]" count="8" pageId="17" pageNumber="468" type="male">8 males</specimenCount>
|
|
,
|
|
<specimenCount id="FAE29989FF9EFFA9D034D91CFCA061B4" box="[768,874,379,403]" count="5" pageId="17" pageNumber="468" type="female">5 females</specimenCount>
|
|
,
|
|
<collectionCode id="8AF5CAC5FF9EFFA9D040D91CFC6C61B4" box="[884,934,379,403]" pageId="17" pageNumber="468">IOZ</collectionCode>
|
|
]), (
|
|
<collectorName id="411137D6FF9EFFA9D0F3D91CFBF061B4" box="[967,1082,379,403]" pageId="17" pageNumber="468">Pukawang</collectorName>
|
|
, on road / 27.84016°/98.32233,
|
|
<quantity id="2B1CFFE5FF9EFFA9D25BD9FCFE0B6194" box="[367,449,411,435]" metricMagnitude="3" metricUnit="m" metricValue="1.458" pageId="17" pageNumber="468" unit="m" value="1458.0">
|
|
<elevation id="67C9B533FF9EFFA9D25BD9FCFE0B6194" box="[367,449,411,435]" metricMagnitude="3" metricUnit="m" metricValue="1.458" pageId="17" pageNumber="468" unit="m" value="1458.0">1458 m</elevation>
|
|
</quantity>
|
|
,
|
|
<date id="985A74C0FF9EFFA9D2FFD9FCFDFC6194" box="[459,566,411,435]" pageId="17" pageNumber="468" value="2021-05-29">
|
|
<collectingDate id="881E8D28FF9EFFA9D2FFD9FCFDFC6194" box="[459,566,411,435]" pageId="17" pageNumber="468" value="2021-05-29">2021.5.29</collectingDate>
|
|
</date>
|
|
<collectorName id="411137D6FF9EFFA9D10FD9FBFD016194" box="[571,715,411,435]" pageId="17" pageNumber="468">N, H.B Liang</collectorName>
|
|
&
|
|
<collectorName id="411137D6FF9EFFA9D1DED9FBFCE26194" box="[746,808,412,435]" pageId="17" pageNumber="468">Y. Xu</collectorName>
|
|
collectors [
|
|
</materialsCitation>
|
|
<materialsCitation id="5C8C585DFF9EFFA9D090D9FCFB876194" ID-GBIF-Occurrence="4902827306" box="[932,1101,411,435]" collectingDate="2004-10-30" collectingDateMax="2021-05-30" collectingDateMin="2004-10-30" collectionCode="CAS, IOZ" collectorName="H. B. Liang & Silalong He & D. H. Kavanaugh & G. Tang & D. Z. Dong & Kongdang & H. B Liang & Y. Xu & Pukawang & N, H. B Liang" county="Dulongjiang Township" elevation="1355" location="Mulangdang" municipality="Bapo" pageId="17" pageNumber="468" specimenCount="33" specimenCount-female="20" specimenCount-male="13" stateProvince="Shan">
|
|
<specimenCount id="FAE29989FF9EFFA9D090D9FCFBC86194" box="[932,1026,411,435]" count="1" pageId="17" pageNumber="468" type="female">1 female</specimenCount>
|
|
,
|
|
<collectionCode id="8AF5CAC5FF9EFFA9D738D9FCFBF56194" box="[1036,1087,411,435]" pageId="17" pageNumber="468">IOZ</collectionCode>
|
|
])
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="5C8C585DFF9EFFA9D763D9FCFF326234" ID-GBIF-Occurrence="4902827304" collectingDate="2006-06-03" collectionCode="R, CAS, IOZ" collectorName="D. H. Kavanaugh & L. Brett & H. B. Liang & D. Z. Dong" county="Tengchong County" elevation="1230" location="Longchuan Jiang" municipality="Wuhe Township" pageId="17" pageNumber="468" specimenCount="65" specimenCount-female="35" specimenCount-male="30" stateProvince="Shan">
|
|
<emphasis id="DE908E12FF9EFFA9D763D9FCFEED61F4" bold="true" italics="true" pageId="17" pageNumber="468">
|
|
<collectingCounty id="053A2A8CFF9EFFA9D763D9FCFEEA61F4" pageId="17" pageNumber="468">Tengchong County</collectingCounty>
|
|
:
|
|
</emphasis>
|
|
<collectingMunicipality id="0C3FC87AFF9EFFA9D202D9DCFE2761F4" box="[310,493,443,467]" pageId="17" pageNumber="468">Wuhe Township</collectingMunicipality>
|
|
(
|
|
<location id="E93B04DBFF9EFFA9D137D9DCFD0E61F4" LSID="urn:lsid:plazi:treatment:644DE316FF80FFA9D34FDA46FC726554:E93B04DBFF9EFFA9D137D9DCFD0E61F4" box="[515,708,443,467]" county="Tengchong County" municipality="Wuhe Township" name="Longchuan Jiang" pageId="17" pageNumber="468" stateProvince="Shan">Longchuan Jiang</location>
|
|
just below bridge at
|
|
<location id="E93B04DBFF9EFFA9D0E5D9DCFB5B61F4" LSID="urn:lsid:plazi:treatment:644DE316FF80FFA9D34FDA46FC726554:E93B04DBFF9EFFA9D0E5D9DCFB5B61F4" box="[977,1169,443,467]" county="Tengchong County" municipality="Wuhe Township" name="Menglian village" pageId="17" pageNumber="468" stateProvince="Shan">Menglian village</location>
|
|
, 24.89176°/98.67551°,
|
|
<quantity id="2B1CFFE5FF9EFFA9D258D9BCFE7461D4" box="[364,446,475,499]" metricMagnitude="3" metricUnit="m" metricValue="1.23" pageId="17" pageNumber="468" unit="m" value="1230.0">
|
|
<elevation id="67C9B533FF9EFFA9D258D9BCFE7461D4" box="[364,446,475,499]" metricMagnitude="3" metricUnit="m" metricValue="1.23" pageId="17" pageNumber="468" unit="m" value="1230.0">1230 m</elevation>
|
|
</quantity>
|
|
,
|
|
<date id="985A74C0FF9EFFA9D2FDD9BCFD8161D4" box="[457,587,475,499]" pageId="17" pageNumber="468" value="2006-06-03">
|
|
<collectingDate id="881E8D28FF9EFFA9D2FDD9BCFD8161D4" box="[457,587,475,499]" pageId="17" pageNumber="468" value="2006-06-03">3 June 2006</collectingDate>
|
|
</date>
|
|
,
|
|
<collectorName id="411137D6FF9EFFA9D161D9BBFCC161D4" box="[597,779,475,499]" pageId="17" pageNumber="468">D.H. Kavanaugh</collectorName>
|
|
,
|
|
<collectionCode id="8AF5CAC5FF9EFFA9D021D9BBFCE061D4" box="[789,810,476,499]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="17" pageNumber="468">R</collectionCode>
|
|
.
|
|
<collectorName id="411137D6FF9EFFA9D019D9BBFC4861D4" box="[813,898,476,499]" pageId="17" pageNumber="468">L. Brett</collectorName>
|
|
,
|
|
<collectorName id="411137D6FF9EFFA9D0B8D9BBFBC861D4" box="[908,1026,475,499]" pageId="17" pageNumber="468">H.B. Liang</collectorName>
|
|
&
|
|
<collectorName id="411137D6FF9EFFA9D717D9BBFB5F61D3" box="[1059,1173,476,500]" pageId="17" pageNumber="468">D.Z. Dong</collectorName>
|
|
collectors [
|
|
</materialsCitation>
|
|
<materialsCitation id="5C8C585DFF9EFFA9D3C3D99CFD0A6234" ID-GBIF-Occurrence="4902827312" box="[247,704,507,531]" collectingDate="2006-06-03" collectionCode="R, CAS, IOZ" collectorName="D. H. Kavanaugh & L. Brett & H. B. Liang & D. Z. Dong" county="Tengchong County" elevation="1230" location="Longchuan Jiang" municipality="Wuhe Township" pageId="17" pageNumber="468" specimenCount="65" specimenCount-female="35" specimenCount-male="30" stateProvince="Shan">
|
|
<specimenCount id="FAE29989FF9EFFA9D3C3D99CFEAB6234" box="[247,353,507,531]" count="30" pageId="17" pageNumber="468" type="male">30 males</specimenCount>
|
|
and
|
|
<specimenCount id="FAE29989FF9EFFA9D290D99CFDE96234" box="[420,547,507,531]" count="35" pageId="17" pageNumber="468" type="female">35 females</specimenCount>
|
|
;
|
|
<collectionCode id="8AF5CAC5FF9EFFA9D102D99CFDA66234" box="[566,620,507,531]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="17" pageNumber="468" type="Museum">CAS</collectionCode>
|
|
,
|
|
<collectionCode id="8AF5CAC5FF9EFFA9D14BD99CFD7B6234" box="[639,689,507,531]" pageId="17" pageNumber="468">IOZ</collectionCode>
|
|
])
|
|
</materialsCitation>
|
|
,
|
|
<materialsCitation id="5C8C585DFF9EFFA9D1E6D99CFEAC6214" ID-GBIF-Occurrence="4902827337" collectorName="Longjiang Bridge" location="Longchuan Jiang" pageId="17" pageNumber="468" specimenCount="1" stateProvince="Shan">
|
|
(
|
|
<location id="E93B04DBFF9EFFA9D1EED99CFC506234" LSID="urn:lsid:plazi:treatment:644DE316FF80FFA9D34FDA46FC726554:E93B04DBFF9EFFA9D1EED99CFC506234" box="[730,922,507,531]" name="Longchuan Jiang" pageId="17" pageNumber="468" stateProvince="Shan">Longchuan Jiang</location>
|
|
at
|
|
<collectorName id="411137D6FF9EFFA9D0FED99CFB5B6234" box="[970,1169,507,531]" pageId="17" pageNumber="468">Longjiang Bridge</collectorName>
|
|
, 24.89889°/98.66667°,
|
|
</materialsCitation>
|
|
<materialsCitation id="5C8C585DFF9EFFA9D25ADA7CFBC06214" ID-GBIF-Occurrence="4902827328" box="[366,1034,539,563]" collectingDate="2003-10-28" collectionCode="CAS, IOZ" collectorName="H. B. Liang & Shi" elevation="1215" pageId="17" pageNumber="468" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Shan">
|
|
<quantity id="2B1CFFE5FF9EFFA9D25ADA7CFE086214" box="[366,450,539,563]" metricMagnitude="3" metricUnit="m" metricValue="1.215" pageId="17" pageNumber="468" unit="m" value="1215.0">
|
|
<elevation id="67C9B533FF9EFFA9D25ADA7CFE086214" box="[366,450,539,563]" metricMagnitude="3" metricUnit="m" metricValue="1.215" pageId="17" pageNumber="468" unit="m" value="1215.0">1215 m</elevation>
|
|
</quantity>
|
|
,
|
|
<date id="985A74C0FF9EFFA9D2F9DA7CFD4F6214" box="[461,645,539,563]" pageId="17" pageNumber="468" value="2003-10-28">
|
|
<collectingDate id="881E8D28FF9EFFA9D2F9DA7CFD4F6214" box="[461,645,539,563]" pageId="17" pageNumber="468" value="2003-10-28">28 October 2003</collectingDate>
|
|
</date>
|
|
,
|
|
<collectorName id="411137D6FF9EFFA9D1A5DA7BFCC36214" box="[657,777,539,563]" pageId="17" pageNumber="468">H.B. Liang</collectorName>
|
|
& X.C.
|
|
<collectorName id="411137D6FF9EFFA9D053DA7CFC416214" box="[871,907,539,563]" pageId="17" pageNumber="468">Shi</collectorName>
|
|
collectors [
|
|
</materialsCitation>
|
|
<materialsCitation id="5C8C585DFF9EFFA9D73EDA7CFE916274" ID-GBIF-Occurrence="4902827330" collectingDate="2003-10-28" collectionCode="CAS, IOZ" collectorName="H. B. Liang & Shi" elevation="1215" pageId="17" pageNumber="468" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Shan">
|
|
<specimenCount id="FAE29989FF9EFFA9D73EDA7CFB996214" box="[1034,1107,539,563]" count="1" pageId="17" pageNumber="468" type="male">1 male</specimenCount>
|
|
and
|
|
<specimenCount id="FAE29989FF9EFFA9D7BCDA7CFF1B6274" count="2" pageId="17" pageNumber="468" type="female">2 females</specimenCount>
|
|
;
|
|
<collectionCode id="8AF5CAC5FF9EFFA9D3E8DA5CFED86274" box="[220,274,571,595]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="17" pageNumber="468" type="Museum">CAS</collectionCode>
|
|
,
|
|
<collectionCode id="8AF5CAC5FF9EFFA9D22ADA5CFE9A6274" box="[286,336,571,595]" pageId="17" pageNumber="468">IOZ</collectionCode>
|
|
].
|
|
</materialsCitation>
|
|
</paragraph>
|
|
<caption id="B89B0288FF9FFFA8D3AFDE24FC5166AF" ID-DOI="http://doi.org/10.5281/zenodo.11067305" ID-Zenodo-Dep="11067305" httpUri="https://zenodo.org/record/11067305/files/figure.png" pageId="16" pageNumber="467" startId="16.[155,167,1603,1622]" targetBox="[120,1176,216,1581]" targetPageId="16" targetType="figure">
|
|
<paragraph id="EC5B5200FF9FFFA8D3AFDE24FC5166AF" blockId="16.[123,1174,1602,1673]" pageId="16" pageNumber="467">
|
|
<smallCapsWord id="EABDC4DCFF9FFFA8D3AFDE24FF126672" baselines="1617,1617" box="[155,216,1603,1622]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="16" pageNumber="467">FIGURE</smallCapsWord>
|
|
10.
|
|
<taxonomicName id="2BE42983FF9FFFA8D234DE25FD816672" authority="Andrewes. A." authorityName="Andrewes. A." box="[256,587,1602,1621]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="species" species="elegans">
|
|
<emphasis id="DE908E12FF9FFFA8D234DE25FE046672" box="[256,462,1602,1621]" italics="true" pageId="16" pageNumber="467">Tetragonoderus elegans</emphasis>
|
|
Andrewes. A.
|
|
</taxonomicName>
|
|
Dorsal aspect, scale line = 1.0 mm (CASENT1039409, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); B. Map showing locality records (black circles) for this species in the Gaoligong Shan region, scale line = 100 km.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="EC5B5200FF9EFFA9D39FDA3CFF3A6334" blockId="17.[123,1174,219,1396]" pageId="17" pageNumber="468">
|
|
Members of this species were collected from the northern to the southern parts of the study area (Core Areas 1, 2, 3 and 6), on both sides of mountain range in the north, only on the eastern side in the central part (Core Area 3) and only on the western side in the southern part (Core Area 6). This distribution pattern is most likely an artifact of inadequate sampling on the western slope of the mountain range, much of which is in
|
|
<collectingCountry id="94F31290FF9EFFA9D150DABBFD0762D3" box="[612,717,732,756]" name="Myanmar" pageId="17" pageNumber="468">Myanmar</collectingCountry>
|
|
, and in the small southeastern part of the study area.
|
|
</paragraph>
|
|
<paragraph id="EC5B5200FF9EFFA9D39FDB7BFDE86454" blockId="17.[123,1174,219,1396]" pageId="17" pageNumber="468">
|
|
<emphasis id="DE908E12FF9EFFA9D39FDB7BFDF86314" bold="true" box="[171,562,796,819]" pageId="17" pageNumber="468">Overall geographical distribution.</emphasis>
|
|
(
|
|
<figureCitation id="74DF4E85FF9EFFA9D176DB7CFD5E6314" box="[578,660,795,819]" captionStart="FIGURE" captionStartId="40.[155,167,997,1016]" captionTargetBox="[120,1176,216,987]" captionTargetId="figure-10@40.[120,1176,216,987]" captionTargetPageId="40" captionText="FIGURE 23. Map showing approximate known overall geographical distributions of cyclosomine species occurring in the Gaoligong Shan. a (blue) = Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov.; b (yellow) = Tetragonoderus arcuatus Dejean; c (red) = T. elegans Andrewes; d (white dot) = T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov.; e (orange) = T. punctatus Wiedemann. Modified from Wikimedia Commons, World Atlas of the World, at URL: http://upload.wikimedia.org/wikipedia/commons/8/8f/Whole_world_-_land_and_oceans_12000.jpg. Scale line = 500 km." figureDoi="http://doi.org/10.5281/zenodo.11067341" httpUri="https://zenodo.org/record/11067341/files/figure.png" pageId="17" pageNumber="468">Fig. 23</figureCitation>
|
|
).
|
|
<collectingMunicipality id="0C3FC87AFF9EFFA9D19CDB7CFD1D6314" box="[680,727,795,819]" pageId="17" pageNumber="468">This</collectingMunicipality>
|
|
species has been known previously only from a few localities on the southern slope of the
|
|
<location id="E93B04DBFF9EFFA9D196DB5CFCC06374" LSID="urn:lsid:plazi:treatment:644DE316FF80FFA9D34FDA46FC726554:E93B04DBFF9EFFA9D196DB5CFCC06374" box="[674,778,827,851]" country="India" county="Medog County" latitude="29.3519" longLatPrecision="6" longitude="95.3417" municipality="This" name="Himalaya" pageId="17" pageNumber="468" stateProvince="Uttarakhand">Himalaya</location>
|
|
in northcentral
|
|
<collectingCountry id="94F31290FF9EFFA9D08EDB5CFC3B6374" box="[954,1009,827,851]" name="India" pageId="17" pageNumber="468">India</collectingCountry>
|
|
(
|
|
<collectingRegion id="2E209CE2FF9EFFA9D734DB5CFB416374" box="[1024,1163,827,851]" country="India" name="Uttarakhand" pageId="17" pageNumber="468">Uttarakhand</collectingRegion>
|
|
). Records of its occurrence in the study area represent the first for
|
|
<collectingCountry id="94F31290FF9EFFA9D009DB3CFCB76354" box="[829,893,859,883]" name="China" pageId="17" pageNumber="468">China</collectingCountry>
|
|
and the easternmost limit of its known distribution.
|
|
<materialsCitation id="5C8C585DFF9EFFA9D2A6DB1CFC0B6394" ID-GBIF-Occurrence="4902827333" collectingDate="2011-07-04" collectingDateMax="2015-08-30" collectingDateMin="2011-07-04" collectionCode="IOZ" collectorName="H. B. Liang & Km & Xia Zayu & Y. Liu & It & Himalayan" country="India" county="Medog County" elevation="707" latitude="29.3519" location="Himalaya" longLatPrecision="6" longitude="95.3417" municipality="This" pageId="17" pageNumber="468" specimenCount="24" specimenCount-female="9" specimenCount-male="15" stateProvince="Uttarakhand">
|
|
Other new records from
|
|
<collectingCountry id="94F31290FF9EFFA9D1AFDB1CFD1163B4" box="[667,731,891,915]" name="China" pageId="17" pageNumber="468">China</collectingCountry>
|
|
are within
|
|
<collectingRegion id="2E209CE2FF9EFFA9D061DB1CFC4663B4" box="[853,908,891,915]" country="China" name="Xizang" pageId="17" pageNumber="468">Tibet</collectingRegion>
|
|
(
|
|
<emphasis id="DE908E12FF9EFFA9D0A9DB1BFB8363B4" bold="true" box="[925,1097,892,916]" pageId="17" pageNumber="468">
|
|
<emphasis id="DE908E12FF9EFFA9D0A9DB1BFB8A63B3" bold="true" box="[925,1088,892,916]" italics="true" pageId="17" pageNumber="468">
|
|
<collectingCounty id="053A2A8CFF9EFFA9D0A9DB1BFB8A63B3" box="[925,1088,892,916]" pageId="17" pageNumber="468">Medog County</collectingCounty>
|
|
</emphasis>
|
|
:
|
|
</emphasis>
|
|
<location id="E93B04DBFF9EFFA9D764DB1CFF5E6394" LSID="urn:lsid:plazi:treatment:644DE316FF80FFA9D34FDA46FC726554:E93B04DBFF9EFFA9D764DB1CFF5E6394" country="India" county="Medog County" latitude="29.3519" longLatPrecision="6" longitude="95.3417" municipality="This" name="Ximohe" pageId="17" pageNumber="468" stateProvince="Uttarakhand">Ximohe</location>
|
|
bridge,
|
|
<geoCoordinate id="89D034C7FF9EFFA9D3C4DBFCFE836394" box="[240,329,923,947]" degrees="29.3519" direction="north" orientation="latitude" pageId="17" pageNumber="468" precision="5" value="29.3519">29.3519</geoCoordinate>
|
|
/
|
|
<geoCoordinate id="89D034C7FF9EFFA9D266DBFCFE616394" box="[338,427,923,947]" degrees="95.3417" direction="east" orientation="longitude" pageId="17" pageNumber="468" precision="5" value="95.3417">95.3417</geoCoordinate>
|
|
°,
|
|
<quantity id="2B1CFFE5FF9EFFA9D28BDBFCFDCD6394" box="[447,519,923,947]" metricMagnitude="2" metricUnit="m" metricValue="7.07" pageId="17" pageNumber="468" unit="m" value="707.0">
|
|
<elevation id="67C9B533FF9EFFA9D28BDBFCFDCD6394" box="[447,519,923,947]" metricMagnitude="2" metricUnit="m" metricValue="7.07" pageId="17" pageNumber="468" unit="m" value="707.0">707 m</elevation>
|
|
</quantity>
|
|
,
|
|
<date id="985A74C0FF9EFFA9D120DBFCFD0E6394" box="[532,708,923,947]" pageId="17" pageNumber="468" value="2015-08-25">
|
|
<collectingDate id="881E8D28FF9EFFA9D120DBFCFD0E6394" box="[532,708,923,947]" pageId="17" pageNumber="468" value="2015-08-25">25 August 2015</collectingDate>
|
|
</date>
|
|
,
|
|
<collectorName id="411137D6FF9EFFA9D1E5DBFBFC806394" box="[721,842,923,947]" pageId="17" pageNumber="468">H.B. Liang</collectorName>
|
|
collector [
|
|
</materialsCitation>
|
|
<materialsCitation id="5C8C585DFF9EFFA9D0F5DBFCFF2363D4" ID-GBIF-Occurrence="4902827309" collectingDate="2011-07-04" collectingDateMax="2015-08-30" collectingDateMin="2011-07-04" collectionCode="IOZ" collectorName="H. B. Liang & Km & Xia Zayu & Y. Liu & It & Himalayan" country="India" county="Medog County" elevation="707" latitude="29.3519" location="Himalaya" longLatPrecision="6" longitude="95.3417" municipality="This" pageId="17" pageNumber="468" specimenCount="24" specimenCount-female="9" specimenCount-male="15" stateProvince="Uttarakhand">
|
|
<specimenCount id="FAE29989FF9EFFA9D0F5DBFCFBD36394" box="[961,1049,923,947]" count="5" pageId="17" pageNumber="468" type="male">5 males</specimenCount>
|
|
,
|
|
<specimenCount id="FAE29989FF9EFFA9D712DBFCFB586394" box="[1062,1170,923,947]" count="5" pageId="17" pageNumber="468" type="female">5 females</specimenCount>
|
|
,
|
|
<collectionCode id="8AF5CAC5FF9EFFA9D34FDBDCFF6763F4" box="[123,173,955,979]" pageId="17" pageNumber="468">IOZ</collectionCode>
|
|
]; Zha-Mo road
|
|
<collectorName id="411137D6FF9EFFA9D26CDBDBFE4A63F4" box="[344,384,956,979]" pageId="17" pageNumber="468">Km</collectorName>
|
|
62,
|
|
<collectingMethod id="35A52A17FF9EFFA9D287DBDCFDD263F4" box="[435,536,955,979]" pageId="17" pageNumber="468">light trap</collectingMethod>
|
|
, 29.7086°/95.5775°,
|
|
<quantity id="2B1CFFE5FF9EFFA9D1CADBDCFC9963F4" box="[766,851,955,979]" metricMagnitude="3" metricUnit="m" metricValue="2.787" pageId="17" pageNumber="468" unit="m" value="2787.0">
|
|
<elevation id="67C9B533FF9EFFA9D1CADBDCFC9963F4" box="[766,851,955,979]" metricMagnitude="3" metricUnit="m" metricValue="2.787" pageId="17" pageNumber="468" unit="m" value="2787.0">2787 m</elevation>
|
|
</quantity>
|
|
,
|
|
<date id="985A74C0FF9EFFA9D054DBDCFBDA63F4" box="[864,1040,955,979]" pageId="17" pageNumber="468" value="2015-08-30">
|
|
<collectingDate id="881E8D28FF9EFFA9D054DBDCFBDA63F4" box="[864,1040,955,979]" pageId="17" pageNumber="468" value="2015-08-30">30 August 2015</collectingDate>
|
|
</date>
|
|
,
|
|
<collectorName id="411137D6FF9EFFA9D728DBDBFB5F63F4" box="[1052,1173,955,979]" pageId="17" pageNumber="468">H.B. Liang</collectorName>
|
|
collector [
|
|
</materialsCitation>
|
|
<materialsCitation id="5C8C585DFF9EFFA9D3DEDBBCFED86434" ID-GBIF-Occurrence="4902827305" collectingDate="2011-07-04" collectingDateMax="2015-08-30" collectingDateMin="2011-07-04" collectionCode="IOZ" collectorName="H. B. Liang & Km & Xia Zayu & Y. Liu & It & Himalayan" country="India" county="Medog County" elevation="707" latitude="29.3519" location="Himalaya" longLatPrecision="6" longitude="95.3417" municipality="This" pageId="17" pageNumber="468" specimenCount="24" specimenCount-female="9" specimenCount-male="15" stateProvince="Uttarakhand">
|
|
<specimenCount id="FAE29989FF9EFFA9D3DEDBBCFEFF63D4" box="[234,309,987,1011]" count="1" pageId="17" pageNumber="468" type="male">1 male</specimenCount>
|
|
,
|
|
<collectionCode id="8AF5CAC5FF9EFFA9D276DBBCFEBE63D4" box="[322,372,987,1011]" pageId="17" pageNumber="468">IOZ</collectionCode>
|
|
].
|
|
<emphasis id="DE908E12FF9EFFA9D2B2DBBBFDEA63D3" bold="true" box="[390,544,988,1012]" italics="true" pageId="17" pageNumber="468">
|
|
<location id="E93B04DBFF9EFFA9D2B2DBBBFDD363D3" LSID="urn:lsid:plazi:treatment:644DE316FF80FFA9D34FDA46FC726554:E93B04DBFF9EFFA9D2B2DBBBFDD363D3" box="[390,537,988,1012]" country="India" county="Medog County" latitude="29.3519" longLatPrecision="6" longitude="95.3417" municipality="This" name="Zayu County" pageId="17" pageNumber="468" stateProvince="Uttarakhand">Zayu County</location>
|
|
:
|
|
</emphasis>
|
|
<collectorName id="411137D6FF9EFFA9D11CDBBCFD5A63D3" box="[552,656,987,1012]" pageId="17" pageNumber="468">Xia Zayu</collectorName>
|
|
, 28.50760°/97.00965°,
|
|
<quantity id="2B1CFFE5FF9EFFA9D0A5DBBCFC2F63D4" box="[913,997,987,1011]" metricMagnitude="3" metricUnit="m" metricValue="1.6509999999999998" pageId="17" pageNumber="468" unit="m" value="1651.0">
|
|
<elevation id="67C9B533FF9EFFA9D0A5DBBCFC2F63D4" box="[913,997,987,1011]" metricMagnitude="3" metricUnit="m" metricValue="1.6509999999999998" pageId="17" pageNumber="468" unit="m" value="1651.0">1651 m</elevation>
|
|
</quantity>
|
|
,
|
|
<date id="985A74C0FF9EFFA9D0C6DBBCFBBE63D4" box="[1010,1140,987,1011]" pageId="17" pageNumber="468" value="2011-07-04">
|
|
<collectingDate id="881E8D28FF9EFFA9D0C6DBBCFBBE63D4" box="[1010,1140,987,1011]" pageId="17" pageNumber="468" value="2011-07-04">4 July 2011</collectingDate>
|
|
</date>
|
|
,
|
|
<collectorName id="411137D6FF9EFFA9D74BDBBBFF6A6434" pageId="17" pageNumber="468">Y. Liu</collectorName>
|
|
collector [
|
|
</materialsCitation>
|
|
<materialsCitation id="5C8C585DFF9EFFA9D227DB9CFDFA6434" ID-GBIF-Occurrence="4902827324" box="[275,560,1019,1043]" collectingDate="2011-07-04" collectingDateMax="2015-08-30" collectingDateMin="2011-07-04" collectionCode="IOZ" collectorName="H. B. Liang & Km & Xia Zayu & Y. Liu & It & Himalayan" country="India" county="Medog County" elevation="707" latitude="29.3519" location="Himalaya" longLatPrecision="6" longitude="95.3417" municipality="This" pageId="17" pageNumber="468" specimenCount="24" specimenCount-female="9" specimenCount-male="15" stateProvince="Uttarakhand">
|
|
<specimenCount id="FAE29989FF9EFFA9D227DB9CFEA06434" box="[275,362,1019,1043]" count="9" pageId="17" pageNumber="468" type="male">9 males</specimenCount>
|
|
,
|
|
<specimenCount id="FAE29989FF9EFFA9D241DB9CFE156434" box="[373,479,1019,1043]" count="4" pageId="17" pageNumber="468" type="female">4 females</specimenCount>
|
|
,
|
|
<collectionCode id="8AF5CAC5FF9EFFA9D2DEDB9CFDD76434" box="[490,541,1019,1043]" pageId="17" pageNumber="468">IOZ</collectionCode>
|
|
]).
|
|
</materialsCitation>
|
|
<collectorName id="411137D6FF9EFFA9D102DB9BFD8C6434" box="[566,582,1020,1043]" pageId="17" pageNumber="468">It</collectorName>
|
|
is likely that
|
|
<taxonomicName id="2BE42983FF9EFFA9D1EEDB9BFC8C6434" box="[730,838,1019,1043]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="elegans">
|
|
<emphasis id="DE908E12FF9EFFA9D1EEDB9BFC8C6434" box="[730,838,1019,1043]" italics="true" pageId="17" pageNumber="468">T. elegans</emphasis>
|
|
</taxonomicName>
|
|
occurs also in suitable habitats in the intervening region, including northern
|
|
<collectingCountry id="94F31290FF9EFFA9D147DC7BFD116413" box="[627,731,1052,1076]" name="Myanmar" pageId="17" pageNumber="468">Myanmar</collectingCountry>
|
|
(
|
|
<collectingRegion id="2E209CE2FF9EFFA9D1D9DC7CFCB56414" box="[749,895,1051,1075]" country="Myanmar" name="Kachin" pageId="17" pageNumber="468">Kachin State</collectingRegion>
|
|
), northeasternmost
|
|
<collectingCountry id="94F31290FF9EFFA9D76CDC7CFB5B6414" box="[1112,1169,1051,1075]" name="India" pageId="17" pageNumber="468">India</collectingCountry>
|
|
,
|
|
<collectingCountry id="94F31290FF9EFFA9D34FDC5CFF016474" box="[123,203,1083,1107]" name="Bhutan" pageId="17" pageNumber="468">Bhutan</collectingCountry>
|
|
, and
|
|
<collectingCountry id="94F31290FF9EFFA9D235DC5CFE896474" box="[257,323,1083,1107]" name="Nepal" pageId="17" pageNumber="468">Nepal</collectingCountry>
|
|
, at elevations between 400 and
|
|
<quantity id="2B1CFFE5FF9EFFA9D1A6DC5CFD286474" box="[658,738,1083,1107]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="468" unit="m" value="2000.0">
|
|
<elevation id="67C9B533FF9EFFA9D1A6DC5CFD286474" box="[658,738,1083,1107]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="468" unit="m" value="2000.0">2000 m</elevation>
|
|
</quantity>
|
|
along rivers draining the southern slopes of the
|
|
<collectorName id="411137D6FF9EFFA9D3F4DC3CFEFF6454" box="[192,309,1115,1139]" pageId="17" pageNumber="468">Himalayan</collectorName>
|
|
ranges in these areas.
|
|
</paragraph>
|
|
<paragraph id="EC5B5200FF9EFFA9D39FDC1BFC726554" blockId="17.[123,1174,219,1396]" pageId="17" pageNumber="468">
|
|
<emphasis id="DE908E12FF9EFFA9D39FDC1BFCA664B4" bold="true" box="[171,876,1148,1172]" pageId="17" pageNumber="468">
|
|
Geographical relationships with other
|
|
<taxonomicName id="2BE42983FF9EFFA9D157DC1BFCA664B4" box="[611,876,1148,1172]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="undetermined">
|
|
<emphasis id="DE908E12FF9EFFA9D157DC1BFCC564B3" bold="true" box="[611,783,1148,1172]" italics="true" pageId="17" pageNumber="468">Tetragonoderus</emphasis>
|
|
species.
|
|
</taxonomicName>
|
|
</emphasis>
|
|
In the study area, members of this species have been found synotopic with those of
|
|
<taxonomicName id="2BE42983FF9EFFA9D1E1DCFAFCAB6494" authorityName="Kavanaugh and Cueva-Dabkoski" authorityYear="2023" box="[725,865,1179,1204]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="parviculus">
|
|
<emphasis id="DE908E12FF9EFFA9D1E1DCFAFCAB6494" box="[725,865,1179,1204]" italics="true" pageId="17" pageNumber="468">T. parviculus</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="2BE42983FF9EFFA9D0A1DCFAFBD26493" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[917,1048,1181,1204]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="punctatus">
|
|
<emphasis id="DE908E12FF9EFFA9D0A1DCFAFBD26493" box="[917,1048,1181,1204]" italics="true" pageId="17" pageNumber="468">T. punctatus</emphasis>
|
|
</taxonomicName>
|
|
. The distributional range also overlaps that of
|
|
<taxonomicName id="2BE42983FF9EFFA9D2C9DCDAFDBE64F3" box="[509,628,1213,1236]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="arcuatus">
|
|
<emphasis id="DE908E12FF9EFFA9D2C9DCDAFDBE64F3" box="[509,628,1213,1236]" italics="true" pageId="17" pageNumber="468">T. arcuatus</emphasis>
|
|
</taxonomicName>
|
|
in the study area, but the two species have not yet been found syntopic. As noted above for
|
|
<taxonomicName id="2BE42983FF9EFFA9D100DCBAFD6164D3" box="[564,683,1245,1268]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="arcuatus">
|
|
<emphasis id="DE908E12FF9EFFA9D100DCBAFD6164D3" box="[564,683,1245,1268]" italics="true" pageId="17" pageNumber="468">T. arcuatus</emphasis>
|
|
</taxonomicName>
|
|
, the range of
|
|
<taxonomicName id="2BE42983FF9EFFA9D074DCBAFC6664D3" box="[832,940,1244,1268]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="elegans">
|
|
<emphasis id="DE908E12FF9EFFA9D074DCBAFC6664D3" box="[832,940,1244,1268]" italics="true" pageId="17" pageNumber="468">T. elegans</emphasis>
|
|
</taxonomicName>
|
|
broadly overlaps that of
|
|
<taxonomicName id="2BE42983FF9EFFA9D3ADDC9AFEF26533" authorityName="Jian & Tian" authorityYear="2009" box="[153,312,1276,1300]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="microthorax">
|
|
<emphasis id="DE908E12FF9EFFA9D3ADDC9AFEF26533" box="[153,312,1276,1300]" italics="true" pageId="17" pageNumber="468">T. microthorax</emphasis>
|
|
</taxonomicName>
|
|
, but the latter species has not yet been recorded from the study area.
|
|
<taxonomicName id="2BE42983FF9EFFA9D719DC9AFEE86513" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="elegans">
|
|
<emphasis id="DE908E12FF9EFFA9D719DC9AFEE86513" italics="true" pageId="17" pageNumber="468">Tetragonoderus elegans</emphasis>
|
|
</taxonomicName>
|
|
is also likely sympatric with
|
|
<taxonomicName id="2BE42983FF9EFFA9D1B3DD7AFCCE6513" box="[647,772,1309,1332]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="arcuatus">
|
|
<emphasis id="DE908E12FF9EFFA9D1B3DD7AFCCE6513" box="[647,772,1309,1332]" italics="true" pageId="17" pageNumber="468">T. arcuatus</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="2BE42983FF9EFFA9D02DDD7AFC776513" authorityName="Jian & Tian" authorityYear="2009" box="[793,957,1308,1332]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="microthorax">
|
|
<emphasis id="DE908E12FF9EFFA9D02DDD7AFC776513" box="[793,957,1308,1332]" italics="true" pageId="17" pageNumber="468">T. microthorax</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="2BE42983FF9EFFA9D0E5DD7AFB916513" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[977,1115,1309,1332]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="punctatus">
|
|
<emphasis id="DE908E12FF9EFFA9D0E5DD7AFB916513" box="[977,1115,1309,1332]" italics="true" pageId="17" pageNumber="468">T. punctatus</emphasis>
|
|
</taxonomicName>
|
|
, and
|
|
<taxonomicName id="2BE42983FF9EFFA9D34FDD5BFDA06573" authority="(Wiedemann, 1823)" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[123,618,1340,1364]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="taeniatus">
|
|
<emphasis id="DE908E12FF9EFFA9D34FDD5BFE466573" box="[123,396,1340,1364]" italics="true" pageId="17" pageNumber="468">Tetragonoderus taeniatus</emphasis>
|
|
(
|
|
<bibRefCitation id="88752FF1FF9EFFA9D2AEDD5BFDA86573" author="WIEDEMANN, C. R. W." box="[410,610,1340,1364]" pageId="17" pageNumber="468" pagination="162 - 164" refId="ref18095" refString="WIEDEMANN, C. R. W. 1823. Zweihundert neue Kafer von Java, Bengalen und dem Vorgebirge der guten Hoffnung. Zoologisches Magazin 2: 1 - 135, 162 - 164." type="journal article" year="1823">Wiedemann, 1823</bibRefCitation>
|
|
)
|
|
</taxonomicName>
|
|
, in suitable habitats in at least some localities along the southern slope of the Himalaya between Uttarakhan and western
|
|
<collectingRegion id="2E209CE2FF9EFFA9D06BDD3BFC7E6554" box="[863,948,1372,1395]" country="China" name="Yunnan" pageId="17" pageNumber="468">Yunnan</collectingRegion>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |