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<document ID-CLB-Dataset="7323" ID-DOI="http://dx.doi.org/10.3897/zookeys.1031.63134" ID-GBIF-Dataset="c2a41bd8-2af6-47bb-b117-5c409e3c5ee8" ID-PMC="PMC8062379" ID-Pensoft-Pub="1313-2970-1031-133" ID-Pensoft-UUID="2B9483DEEF9C5CF486971E358333AB4E" ID-PubMed="33958910" ID-ZooBank="15B4DD8014BD42EABB6D11E8FE9392BA" ModsDocID="1313-2970-1031-133" checkinTime="1618544337298" checkinUser="pensoft" docAuthor="Ng, Peter K. L." docDate="2021" docId="62B0437F02305D64A5CD428E177663A1" docLanguage="en" docName="ZooKeys 1031: 133-141" docOrigin="ZooKeys 1031" docPubDate="2021-04-15" docSource="http://dx.doi.org/10.3897/zookeys.1031.63134" docTitle="Geosesarma sodalis Ng 2021, sp. nov." docType="treatment" docUuid="69A4BE4D-8B0B-4243-9B2D-BA1D559A2C28" docUuidSource="ZooBank" docVersion="6" id="2B9483DEEF9C5CF486971E358333AB4E" lastPageNumber="133" masterDocId="2B9483DEEF9C5CF486971E358333AB4E" masterDocTitle="Geosesarma sodalis, a new species of vampire crab (Crustacea, Brachyura, Sesarmidae) from a limestone cave in central Sarawak, Malaysia" masterLastPageNumber="141" masterPageNumber="133" pageNumber="133" updateTime="1732840798367" updateUser="ExternalLinkService">
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<mods:title id="B206099A5433A9E29605A45672AFEED6">Geosesarma sodalis, a new species of vampire crab (Crustacea, Brachyura, Sesarmidae) from a limestone cave in central Sarawak, Malaysia</mods:title>
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<mods:name id="F6E1B37BE55915AD4B303C94774749EE" type="personal">
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</mods:role>
<mods:namePart id="64BE102A3F9CFEEC4DEAFD507BFA2308">Ng, Peter K. L.</mods:namePart>
<mods:affiliation id="4404E0A47F7E69C5ACCB1EF90A6E0BA2">Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, Kent Ridge, Singapore 119260, Republic of Singapore</mods:affiliation>
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<mods:typeOfResource id="35EDC6A93F7D6F34A44BE7B7B6CB1BA0">text</mods:typeOfResource>
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<mods:date id="FE353E5517A4A00E5305D04635364690">2021</mods:date>
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<mods:number id="CD7D7600683ADA382324268C2F47FE4B">2021-04-15</mods:number>
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<mods:number id="E2ACF878D85312D27BD943B4192FECFE">1031</mods:number>
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<mods:classification id="93073E06A156BA66BF087A0022AD1CDA">journal article</mods:classification>
<mods:identifier id="669256AD3FB2D666E03A039E708B699A" type="DOI">http://dx.doi.org/10.3897/zookeys.1031.63134</mods:identifier>
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<subSection id="C6CF7E3DB99850A534569D418D6E6CAC" pageId="0" pageNumber="133" type="systematic accounts">
<treatment id="62B0437F02305D64A5CD428E177663A1" ID-GBIF-Taxon="181019293" LSID="urn:lsid:zoobank.org:act:69A4BE4D-8B0B-4243-9B2D-BA1D559A2C28" httpUri="http://treatment.plazi.org/id/62B0437F02305D64A5CD428E177663A1" lastPageNumber="133" pageId="0" pageNumber="133" scope_family="Sesarmidae" scope_infraOrder="Brachyura">
<subSubSection id="5245052109D93A42AC75316B09BF4095" pageId="0" pageNumber="133" type="nomenclature">
<paragraph id="ACAFCFC65772402D4B5797AA0815C909" pageId="0" pageNumber="133">
<taxonomicName id="1892D67B0B63CADE4E097F58AD730578" ID-CoL="8753M" LSID="http://zoobank.org/69A4BE4D-8B0B-4243-9B2D-BA1D559A2C28" authority="Ng, 2021" authorityName="Ng" authorityYear="2021" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geosesarma sodalis" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="sodalis" status="sp. nov.">Geosesarma sodalis</taxonomicName>
<taxonomicNameLabel id="90473FA08249D0F81E22D1DFBD6A2A7C" pageId="0" pageNumber="133">sp. nov.</taxonomicNameLabel>
<figureCitation id="96AC15F3D2C4CA4804BC29E904972AEA" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Geosesarma sodalis sp. nov., holotype male (10.1 x 9.8 mm) (ZRC 2020.0413), Sarawak A overall dorsal view B dorsal view of carapace C frontal view of cephalothorax D anterior thoracic sternites and sternopleonal cavity." figureDoi="10.3897/zookeys.1031.63134.figure1" httpUri="https://binary.pensoft.net/fig/531984" pageId="0" pageNumber="133">Figures 1</figureCitation>
<figureCitation id="9D51DE65B88FFCD1750C92FFDECC34AA" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Geosesarma sodalis sp. nov., holotype male (10.1 x 9.8 mm) (ZRC 2020.0413), Sarawak A dorsal view of right cheliped B outer view of right chela C subdorsal view of left chela D inner view of right chela E right third ambulatory leg F right fourth ambulatory leg G pleonal somites 2 - 6 and telson H left G 1 (ventral view) I left G 1 (ventral view) J distal part of left G 1 (ventral view) K distal part of left G 1 (ventral view)." figureDoi="10.3897/zookeys.1031.63134.figure2" httpUri="https://binary.pensoft.net/fig/531985" pageId="0" pageNumber="133">, 2</figureCitation>
<figureCitation id="AB04921EB6C30219A2486A6EF4ACC193" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Geosesarma sodalis sp. nov., holotype male (10.1 x 9.8 mm) (ZRC 2020.0413), Sarawak A left third maxilliped (setae denuded) B left G 1 (ventral view) C left G 1 (ventral view) D distal part of left G 1 (ventral view) E distal part of left G 1 (distomesial view) F distal part of left G 1 (ventral view) G distal part of left G 1 (ventromesial view) H left G 2. Scales bars: 0.5 mm (A-C, H); 0.25 mm (D-G)." figureDoi="10.3897/zookeys.1031.63134.figure3" httpUri="https://binary.pensoft.net/fig/531986" pageId="0" pageNumber="133">, 3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="F25D600B3BAD3B4EA9A58B3732977038" pageId="0" pageNumber="133" type="materials_examined">
<paragraph id="2DBA03DF2FC9249EB2E190CD9C374989" pageId="0" pageNumber="133">Material examined.</paragraph>
<paragraph id="CFD78D450A9F50F9E3F40052662A89CB" pageId="0" pageNumber="133">
<materialsCitation id="A8B9CB4BA43F0145AE364DA68DC8C452" ID-GBIF-Occurrence="3075950301" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Stuebing RB" country="Sarawak" location="Sarawak" specimenCount="1" typeStatus="Holotype">
<emphasis id="C5A874331FFD74EACCAECAF17D52F216" bold="true" pageId="0" pageNumber="133">
<emphasis id="B85BBC9AF42D6E3105479527439962B3" italics="true" pageId="0" pageNumber="133">
<typeStatus id="1204CC1F94D4B21605900744DD92FECE">Holotype</typeStatus>
</emphasis>
:
</emphasis>
male (10.1
<normalizedToken id="089EA22BDB5785DBA0B1BC97553D3E51" originalValue="×">x</normalizedToken>
<quantity id="8160D3A03CBE46FCC32597E774095794" metricMagnitude="-3" metricUnit="m" metricValue="9.8" unit="mm" value="9.8">9.8 mm</quantity>
) (ZRC 2020.0413), limestone cave, Bukit Sarang, Bintulu,
<collectingCountry id="58B2932B9F9228977FF6F6FBFE47BE58" name="Malaysia">Sarawak</collectingCountry>
,
<collectingCountry id="27AC15C8880902138841A22524BA626E" name="Malaysia">Malaysia</collectingCountry>
, coll.
<collectorName id="87D61E0E2FD7F9B89B51FE7E6FA16C03">Stuebing RB</collectorName>
, early 2005.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="F8A638F4CA31653C5919101DB2476221" pageId="0" pageNumber="133" type="diagnosis">
<paragraph id="7EB3B05E4B24FCC3FD147B6642303F8D" pageId="0" pageNumber="133">Diagnosis.</paragraph>
<paragraph id="B463CA411A3C030FFFA016BEF255B2F5" pageId="0" pageNumber="133">
Carapace quadrate, slightly wider than long, width to length ratio 1.03, lateral margins gently concave, subparallel (Fig.
<figureCitation id="11B926C2651C3FB655FCE4AC85C13868" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Geosesarma sodalis sp. nov., holotype male (10.1 x 9.8 mm) (ZRC 2020.0413), Sarawak A overall dorsal view B dorsal view of carapace C frontal view of cephalothorax D anterior thoracic sternites and sternopleonal cavity." figureDoi="10.3897/zookeys.1031.63134.figure1" httpUri="https://binary.pensoft.net/fig/531984" pageId="0" pageNumber="133">1A, B</figureCitation>
); dorsal surfaces with well-defined regions, anterior half with low granules, posterior half almost smooth (Fig.
<figureCitation id="ED94F8700465C7DFA94EA00C99BFE6F6" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Geosesarma sodalis sp. nov., holotype male (10.1 x 9.8 mm) (ZRC 2020.0413), Sarawak A overall dorsal view B dorsal view of carapace C frontal view of cephalothorax D anterior thoracic sternites and sternopleonal cavity." figureDoi="10.3897/zookeys.1031.63134.figure1" httpUri="https://binary.pensoft.net/fig/531984" pageId="0" pageNumber="133">1A, B</figureCitation>
); frontal margin distinctly deflexed, frontal lobes broad, with truncated margins in dorsal view, separated by wide shallow median concavity; postfrontal and postorbital cristae sharp, distinct (Fig.
<figureCitation id="471FC8F28CBDBA464BF42FF785B02F57" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Geosesarma sodalis sp. nov., holotype male (10.1 x 9.8 mm) (ZRC 2020.0413), Sarawak A overall dorsal view B dorsal view of carapace C frontal view of cephalothorax D anterior thoracic sternites and sternopleonal cavity." figureDoi="10.3897/zookeys.1031.63134.figure1" httpUri="https://binary.pensoft.net/fig/531984" pageId="0" pageNumber="133">1A-C</figureCitation>
); external orbital angle triangular, directed obliquely anteriorly, extending just beyond lateral carapace margins, outer lateral margin convex; separated from first epibranchial tooth by deep V-shaped cleft; first epibranchial tooth distinct, second epibranchial tooth visible only as low lobe, barely separated from first tooth by shallow concavity (Fig.
<figureCitation id="B68ECF9FC4C263A08DCF36860E1BC154" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Geosesarma sodalis sp. nov., holotype male (10.1 x 9.8 mm) (ZRC 2020.0413), Sarawak A overall dorsal view B dorsal view of carapace C frontal view of cephalothorax D anterior thoracic sternites and sternopleonal cavity." figureDoi="10.3897/zookeys.1031.63134.figure1" httpUri="https://binary.pensoft.net/fig/531984" pageId="0" pageNumber="133">1A, B</figureCitation>
); merus of third maxilliped subovate; exopod slender, flagellum absent (Fig.
<figureCitation id="2619665050879316FC8E9066D4C7D27A" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Geosesarma sodalis sp. nov., holotype male (10.1 x 9.8 mm) (ZRC 2020.0413), Sarawak A left third maxilliped (setae denuded) B left G 1 (ventral view) C left G 1 (ventral view) D distal part of left G 1 (ventral view) E distal part of left G 1 (distomesial view) F distal part of left G 1 (ventral view) G distal part of left G 1 (ventromesial view) H left G 2. Scales bars: 0.5 mm (A-C, H); 0.25 mm (D-G)." figureDoi="10.3897/zookeys.1031.63134.figure3" httpUri="https://binary.pensoft.net/fig/531986" pageId="0" pageNumber="133">3A</figureCitation>
); outer surfaces of palm of chela covered with small rounded granules, inner surface without transverse ridge; fingers longer than palm, dorsal margin of dactylus with 10 or 11 sharp, anteriorly directed sharp tubercles (Fig.
<figureCitation id="6E45B5D107AA55ECFB35BC3D17EA23A8" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Geosesarma sodalis sp. nov., holotype male (10.1 x 9.8 mm) (ZRC 2020.0413), Sarawak A dorsal view of right cheliped B outer view of right chela C subdorsal view of left chela D inner view of right chela E right third ambulatory leg F right fourth ambulatory leg G pleonal somites 2 - 6 and telson H left G 1 (ventral view) I left G 1 (ventral view) J distal part of left G 1 (ventral view) K distal part of left G 1 (ventral view)." figureDoi="10.3897/zookeys.1031.63134.figure2" httpUri="https://binary.pensoft.net/fig/531985" pageId="0" pageNumber="133">2A-D</figureCitation>
); ambulatory merus with sharp subdistal spine on dorsal margin, surface weakly rugose, propodus slender, relatively long (Figs
<figureCitation id="72EFA8B9571E689B826E2A50C2CD6E75" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Geosesarma sodalis sp. nov., holotype male (10.1 x 9.8 mm) (ZRC 2020.0413), Sarawak A overall dorsal view B dorsal view of carapace C frontal view of cephalothorax D anterior thoracic sternites and sternopleonal cavity." figureDoi="10.3897/zookeys.1031.63134.figure1" httpUri="https://binary.pensoft.net/fig/531984" pageId="0" pageNumber="133">1A</figureCitation>
,
<figureCitation id="E8AC90A044236AE4C95665F60A6A568A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Geosesarma sodalis sp. nov., holotype male (10.1 x 9.8 mm) (ZRC 2020.0413), Sarawak A dorsal view of right cheliped B outer view of right chela C subdorsal view of left chela D inner view of right chela E right third ambulatory leg F right fourth ambulatory leg G pleonal somites 2 - 6 and telson H left G 1 (ventral view) I left G 1 (ventral view) J distal part of left G 1 (ventral view) K distal part of left G 1 (ventral view)." figureDoi="10.3897/zookeys.1031.63134.figure2" httpUri="https://binary.pensoft.net/fig/531985" pageId="0" pageNumber="133">2E, F</figureCitation>
); pleon triangular; somite 3 widest, somite 6 with lateral margins gently convex; telson triangular, longer than broad, lateral margins gently convex (Fig.
<figureCitation id="7C6CAEE622204BB10CDAE3B2D0DE9663" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Geosesarma sodalis sp. nov., holotype male (10.1 x 9.8 mm) (ZRC 2020.0413), Sarawak A dorsal view of right cheliped B outer view of right chela C subdorsal view of left chela D inner view of right chela E right third ambulatory leg F right fourth ambulatory leg G pleonal somites 2 - 6 and telson H left G 1 (ventral view) I left G 1 (ventral view) J distal part of left G 1 (ventral view) K distal part of left G 1 (ventral view)." figureDoi="10.3897/zookeys.1031.63134.figure2" httpUri="https://binary.pensoft.net/fig/531985" pageId="0" pageNumber="133">2G</figureCitation>
); G1 relatively slender, proximal, distal part bent at angle of ca. 45° along longitudinal axis, subdistal part of outer margin gently angular with shelf-like feature (Figs
<figureCitation id="B17AFF8CD15370C0A9D2001ED2BAE21D" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Geosesarma sodalis sp. nov., holotype male (10.1 x 9.8 mm) (ZRC 2020.0413), Sarawak A dorsal view of right cheliped B outer view of right chela C subdorsal view of left chela D inner view of right chela E right third ambulatory leg F right fourth ambulatory leg G pleonal somites 2 - 6 and telson H left G 1 (ventral view) I left G 1 (ventral view) J distal part of left G 1 (ventral view) K distal part of left G 1 (ventral view)." figureDoi="10.3897/zookeys.1031.63134.figure2" httpUri="https://binary.pensoft.net/fig/531985" pageId="0" pageNumber="133">2H-K</figureCitation>
,
<figureCitation id="6C3939DC6879D56C447364E8318DE63A" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Geosesarma sodalis sp. nov., holotype male (10.1 x 9.8 mm) (ZRC 2020.0413), Sarawak A left third maxilliped (setae denuded) B left G 1 (ventral view) C left G 1 (ventral view) D distal part of left G 1 (ventral view) E distal part of left G 1 (distomesial view) F distal part of left G 1 (ventral view) G distal part of left G 1 (ventromesial view) H left G 2. Scales bars: 0.5 mm (A-C, H); 0.25 mm (D-G)." figureDoi="10.3897/zookeys.1031.63134.figure3" httpUri="https://binary.pensoft.net/fig/531986" pageId="0" pageNumber="133">3B-D, F</figureCitation>
), distal part elongate, tapering in lateral view, spatuliform in marginal view, with small submedian cleft at tip when viewed mesially (Fig.
<figureCitation id="CA52F7CF7FFBBB1CC5C09A934734DB89" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Geosesarma sodalis sp. nov., holotype male (10.1 x 9.8 mm) (ZRC 2020.0413), Sarawak A left third maxilliped (setae denuded) B left G 1 (ventral view) C left G 1 (ventral view) D distal part of left G 1 (ventral view) E distal part of left G 1 (distomesial view) F distal part of left G 1 (ventral view) G distal part of left G 1 (ventromesial view) H left G 2. Scales bars: 0.5 mm (A-C, H); 0.25 mm (D-G)." figureDoi="10.3897/zookeys.1031.63134.figure3" httpUri="https://binary.pensoft.net/fig/531986" pageId="0" pageNumber="133">3E, G</figureCitation>
).
</paragraph>
<caption id="85544ECFC0F148F359F8F8435018CC6F" doi="10.3897/zookeys.1031.63134.figure1" httpUri="https://binary.pensoft.net/fig/531984" pageId="0" pageNumber="133" start="Figure 1" startId="F1">
<paragraph id="9B165A73F1398D1B8E1406C5CDC4649D" pageId="0" pageNumber="133">
<emphasis id="A9F6CBEF17F7137DFCCCAAF8876B492E" bold="true" pageId="0" pageNumber="133">Figure 1.</emphasis>
<taxonomicName id="610968AE63AEC8290409BBC4B2D32C49" authorityName="Ng" authorityYear="2021" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geosesarma sodalis" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="sodalis">
<emphasis id="5D080026A1A0A24D43085F98C26F994C" italics="true" pageId="0" pageNumber="133">Geosesarma sodalis</emphasis>
</taxonomicName>
sp. nov., holotype male (10.1
<normalizedToken id="7CF52ED729B2C8D4D83188E81284CAEB" originalValue="×">x</normalizedToken>
9.8 mm) (ZRC 2020.0413), Sarawak
<emphasis id="5C278C4DD3D1A508419AA6C8E0E0E63D" bold="true" pageId="0" pageNumber="133">A</emphasis>
overall dorsal view
<emphasis id="97F24391F1DA14F4C163B0E04C189BDC" bold="true" pageId="0" pageNumber="133">B</emphasis>
dorsal view of carapace
<emphasis id="09B0997C57C446D1C93E4883FCEBA484" bold="true" pageId="0" pageNumber="133">C</emphasis>
frontal view of cephalothorax
<emphasis id="E9D69D28606B8A1E1960BCD5161BD2C6" bold="true" pageId="0" pageNumber="133">D</emphasis>
anterior thoracic sternites and sternopleonal cavity.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="0DC362D1C36B3638BF5A0E5107F550A0" pageId="0" pageNumber="133" type="colour">
<paragraph id="3F4E48D00B8C59EA82E6E76F8C5E6EC9" pageId="0" pageNumber="133">Colour.</paragraph>
<paragraph id="FECDF04DB71E01817BA2B1AD7FEC0EAC" pageId="0" pageNumber="133">Not known.</paragraph>
</subSubSection>
<subSubSection id="0A027156E879028C2E35D0546F4A00FA" pageId="0" pageNumber="133" type="females">
<paragraph id="2C83CF5D61191BE61D9E1B9D88904601" pageId="0" pageNumber="133">Females.</paragraph>
<paragraph id="109798569DF4BFD3D3B6B0054045D82A" pageId="0" pageNumber="133">Not known.</paragraph>
</subSubSection>
<subSubSection id="018AE5E8501C0D2F021A06EC62398C4E" pageId="0" pageNumber="133" type="etymology">
<paragraph id="D2F5AD7818BD5404831716E376769ED0" pageId="0" pageNumber="133">Etymology.</paragraph>
<paragraph id="6A8D68F664060109B038EB365ADEB798" pageId="0" pageNumber="133">The name is derived from the Latin noun for comradeship; alluding to the deep friendship the author has had over the last 30 years with the collector, Rob Stuebing, who has collected many interesting species for him.</paragraph>
</subSubSection>
<subSubSection id="BC61A7BC7A98F2DEBA97BB8FB2538A69" pageId="0" pageNumber="133" type="remarks">
<paragraph id="63AC03E5657D0B036F240DD65263BC34" pageId="0" pageNumber="133">Remarks.</paragraph>
<paragraph id="648D4DDF0CD143DD1765D59528EC614C" pageId="0" pageNumber="133">
The island of Borneo has 13 known species of
<taxonomicName id="607C1AC91F46B0AA3032C170C004DBED" authorityName="De Man" authorityYear="1892" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geosesarma" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="0D5E7D350D92B44DF572D5962B273BB6" italics="true" pageId="0" pageNumber="133">Geosesarma</emphasis>
</taxonomicName>
, all of which are endemic to the island. Five species occur in the state of Sarawak (
<bibRefCitation id="64556B21FCF274C16AE0EADF93DD26A0" DOI="https://doi.org/10.11646/zootaxa.4508.4.5" author="Ng, PKL" journalOrPublisher="Zootaxa" pageId="0" pageNumber="133" pagination="569 - 575" refId="B16" refString="Ng, PKL, Grinang, J, 2018. A new species of highland vampire crab (Crustacea: Brachyura: Sesarmidae: Geosesarma) from Serian, Sarawak. Zootaxa 4508 (4): 569 - 575, DOI: https://doi.org/10.11646/zootaxa.4508.4.5" title="A new species of highland vampire crab (Crustacea: Brachyura: Sesarmidae: Geosesarma) from Serian, Sarawak." url="https://doi.org/10.11646/zootaxa.4508.4.5" volume="4508" year="2018">Ng and Grinang 2018</bibRefCitation>
;
<bibRefCitation id="22B6A080AE0093DDF7138907379F2C2F" DOI="https://doi.org/10.11646/zootaxa.4614.3.6" author="Ng, PYC" journalOrPublisher="Zootaxa" pageId="0" pageNumber="133" pagination="529 - 540" refId="B20" refString="Ng, PYC, Ng, PKL, 2019. Geosesarma spectrum, a new species of semiterrestrial vampire crab (Crustacea: Decapoda: Brachyura: Sesarmidae) from Brunei Darussalam, Borneo. Zootaxa 4614 (3): 529 - 540, DOI: https://doi.org/10.11646/zootaxa.4614.3.6" title="Geosesarma spectrum, a new species of semiterrestrial vampire crab (Crustacea: Decapoda: Brachyura: Sesarmidae) from Brunei Darussalam, Borneo." url="https://doi.org/10.11646/zootaxa.4614.3.6" volume="4614" year="2019">Ng and Ng 2019</bibRefCitation>
). One group of
<taxonomicName id="FBF8CC6DA3B58FE931AC86A5D1509A34" authorityName="De Man" authorityYear="1892" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geosesarma" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="B71660E35F04234EDE543A94DFDE88C1" italics="true" pageId="0" pageNumber="133">Geosesarma</emphasis>
</taxonomicName>
species is characterised by their relatively quadrate carapace, presence of a row to sharp tubercles on the dorsal margin of the cheliped dactylus, absence of a flagellum on the third maxilliped exopod, and a relatively stout G1 with a tapering corneous distal part (in lateral view). In Borneo, the species in this group are
<taxonomicName id="7EDE97916C038968A82FB1919885913F" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. gracillimum" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="gracillimum">
<emphasis id="CE72E3436A2BA62D5462187D7F3941C3" italics="true" pageId="0" pageNumber="133">G. gracillimum</emphasis>
</taxonomicName>
(De Man, 1902),
<taxonomicName id="9C2731F0C293AB227BA127BDE8C7B5BB" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. sabanus" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="sabanus">
<emphasis id="31722C2EC234DFC575237F6CC1792C8B" italics="true" pageId="0" pageNumber="133">G. sabanus</emphasis>
</taxonomicName>
Ng, 1992,
<taxonomicName id="7306E53175129C3FEB700F4920B40F2D" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. aurantium" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="aurantium">
<emphasis id="697A3769593B160C65A01AA7CF48A95F" italics="true" pageId="0" pageNumber="133">G. aurantium</emphasis>
</taxonomicName>
Ng, 1995,
<taxonomicName id="FA7B5A0E58E04B2312915A5BABAB44D7" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. katibas" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="katibas">
<emphasis id="2A16BAD37E0A1F0EDBA69636ACC19441" italics="true" pageId="0" pageNumber="133">G. katibas</emphasis>
</taxonomicName>
Ng, 1995,
<taxonomicName id="08892348944F5CDE40671F294B71DEF5" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. danumense" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="danumense">
<emphasis id="6E772F3CF4CA9E49E487D033040777C9" italics="true" pageId="0" pageNumber="133">G. danumense</emphasis>
</taxonomicName>
Ng, 2002,
<taxonomicName id="576279220D98D637EE065111443F3664" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. bau" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="bau">
<emphasis id="C45ABE9B81B5FE77BC1F30A06918198D" italics="true" pageId="0" pageNumber="133">G. bau</emphasis>
</taxonomicName>
Ng &amp; Grinang, 2004,
<taxonomicName id="C38339D63AC911B37C740EB4F9D7814C" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. ambawang" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="ambawang">
<emphasis id="AFE9D74BB4C818584CD4F647EE9B838C" italics="true" pageId="0" pageNumber="133">G. ambawang</emphasis>
</taxonomicName>
Ng, 2015,
<taxonomicName id="42D8AF12D2BD82A43BEF05127D937CC9" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. pontianak" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="pontianak">
<emphasis id="A899E4447675067BD898F88E8BB40A97" italics="true" pageId="0" pageNumber="133">G. pontianak</emphasis>
</taxonomicName>
Ng, 2015,
<taxonomicName id="14DD9873DF06C52D3C27679F8D793398" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. larsi" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="larsi">
<emphasis id="2F12AA56D554A6074AED5E55D2D7D1BE" italics="true" pageId="0" pageNumber="133">G. larsi</emphasis>
</taxonomicName>
Ng &amp; Grinang, 2018, and
<taxonomicName id="ABC2FAFEAEE57B50658AFD63844C61BD" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. spectrum" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="spectrum">
<emphasis id="7C0ECA6294D64488B3E3C9CE79994AB6" italics="true" pageId="0" pageNumber="133">G. spectrum</emphasis>
</taxonomicName>
Ng &amp; Ng, 2019.
</paragraph>
<paragraph id="4839E894FCF7FAB86C1F795631803343" pageId="0" pageNumber="133">
Five of the species in this group are present in Sarawak and Brunei:
<taxonomicName id="8518CA608FD74EE8D10D2F3DA732E11C" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. gracillimum" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="gracillimum">
<emphasis id="CE8A35992B3C76F85B7CC30FBAA0F532" italics="true" pageId="0" pageNumber="133">G. gracillimum</emphasis>
</taxonomicName>
,
<taxonomicName id="282982E6BE442E0BF79D2763240E4C25" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. katibas" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="katibas">
<emphasis id="742EDA860B7BB100A7A3A21AD8EA2B8C" italics="true" pageId="0" pageNumber="133">G. katibas</emphasis>
</taxonomicName>
,
<taxonomicName id="78B9EE359E2BFBED214E449D03729472" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. bau" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="bau">
<emphasis id="D7C4CDDD6B988D40AE1FBCDB9A91CCA8" italics="true" pageId="0" pageNumber="133">G. bau</emphasis>
</taxonomicName>
,
<taxonomicName id="C94E6E1A58781C0E83436ACAA89DA155" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. larsi" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="larsi">
<emphasis id="4597290542FDEB97EAD5CFA082030254" italics="true" pageId="0" pageNumber="133">G. larsi</emphasis>
</taxonomicName>
, and
<taxonomicName id="AD3F7A122DA23A87F09DCDAD94D56D6E" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. sodalis" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="sodalis">
<emphasis id="215363313C7C85F3F7A926CBFDC54CFA" italics="true" pageId="0" pageNumber="133">G. sodalis</emphasis>
</taxonomicName>
sp. nov. Compared to
<taxonomicName id="BA0E02069ADD0E570FF885406DB26397" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. gracillimum" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="gracillimum">
<emphasis id="A5213ED56F3F1B5ADBC7A9EB7E871547" italics="true" pageId="0" pageNumber="133">G. gracillimum</emphasis>
</taxonomicName>
, the carapace of
<taxonomicName id="A1D1B5FF004F08A979EF5718947678C9" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. sodalis" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="sodalis">
<emphasis id="D9C9FC2578BCE7F923E17BD7FE315ADB" italics="true" pageId="0" pageNumber="133">G. sodalis</emphasis>
</taxonomicName>
sp. nov. is more quadrate with the lateral margins subparallel (Fig.
<figureCitation id="47D8508B77A7EA2A4372AFC777FDFFEB" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Geosesarma sodalis sp. nov., holotype male (10.1 x 9.8 mm) (ZRC 2020.0413), Sarawak A overall dorsal view B dorsal view of carapace C frontal view of cephalothorax D anterior thoracic sternites and sternopleonal cavity." figureDoi="10.3897/zookeys.1031.63134.figure1" httpUri="https://binary.pensoft.net/fig/531984" pageId="0" pageNumber="133">1A, b</figureCitation>
) (versus gently diverging in
<taxonomicName id="E8A4788EF2278A394E2F959A42C6F402" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. gracillimum" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="gracillimum">
<emphasis id="2690B2E9483D7122409F34B4CE5E830B" italics="true" pageId="0" pageNumber="133">G. gracillimum</emphasis>
</taxonomicName>
; see
<bibRefCitation id="66A814460F426F29602AEEE07DF24D57" DOI="https://doi.org/10.11646/zootaxa.4048.1.2" author="Ng, PKL" journalOrPublisher="Zootaxa" pageId="0" pageNumber="133" pagination="37 - 56" refId="B12" refString="Ng, PKL, 2015. Semiterrestrial crabs of the genus Geosesarma De Man, 1892 (Crustacea, Brachyura, Sesarmidae) from western Borneo, Indonesia, with descriptions of three new species. Zootaxa 4048 (1): 37 - 56, DOI: https://doi.org/10.11646/zootaxa.4048.1.2" title="Semiterrestrial crabs of the genus Geosesarma De Man, 1892 (Crustacea, Brachyura, Sesarmidae) from western Borneo, Indonesia, with descriptions of three new species." url="https://doi.org/10.11646/zootaxa.4048.1.2" volume="4048" year="2015">Ng 2015</bibRefCitation>
: fig. 14A, B; Ng and
<bibRefCitation id="126692596F81FC299BA4A6D167E6B838" DOI="https://doi.org/10.11646/zootaxa.4048.1.2" author="Ng, PKL" journalOrPublisher="Zootaxa" pageId="0" pageNumber="133" pagination="37 - 56" refId="B12" refString="Ng, PKL, 2015. Semiterrestrial crabs of the genus Geosesarma De Man, 1892 (Crustacea, Brachyura, Sesarmidae) from western Borneo, Indonesia, with descriptions of three new species. Zootaxa 4048 (1): 37 - 56, DOI: https://doi.org/10.11646/zootaxa.4048.1.2" title="Semiterrestrial crabs of the genus Geosesarma De Man, 1892 (Crustacea, Brachyura, Sesarmidae) from western Borneo, Indonesia, with descriptions of three new species." url="https://doi.org/10.11646/zootaxa.4048.1.2" volume="4048" year="2015">Ng 2015</bibRefCitation>
: fig. 5F). The G1 of
<taxonomicName id="A6F60E3C5F708CC89F590E0EB4C56F56" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. sodalis" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="sodalis">
<emphasis id="E6220FEB1AAFA4D242D86822F64A3144" italics="true" pageId="0" pageNumber="133">G. sodalis</emphasis>
</taxonomicName>
sp. nov. (Figs
<figureCitation id="936F9EDF5F8F2ED8304B97CD87296A93" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Geosesarma sodalis sp. nov., holotype male (10.1 x 9.8 mm) (ZRC 2020.0413), Sarawak A dorsal view of right cheliped B outer view of right chela C subdorsal view of left chela D inner view of right chela E right third ambulatory leg F right fourth ambulatory leg G pleonal somites 2 - 6 and telson H left G 1 (ventral view) I left G 1 (ventral view) J distal part of left G 1 (ventral view) K distal part of left G 1 (ventral view)." figureDoi="10.3897/zookeys.1031.63134.figure2" httpUri="https://binary.pensoft.net/fig/531985" pageId="0" pageNumber="133">2H, I</figureCitation>
,
<figureCitation id="801CA5BE2C6A982AADE494A3DB57AA18" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Geosesarma sodalis sp. nov., holotype male (10.1 x 9.8 mm) (ZRC 2020.0413), Sarawak A left third maxilliped (setae denuded) B left G 1 (ventral view) C left G 1 (ventral view) D distal part of left G 1 (ventral view) E distal part of left G 1 (distomesial view) F distal part of left G 1 (ventral view) G distal part of left G 1 (ventromesial view) H left G 2. Scales bars: 0.5 mm (A-C, H); 0.25 mm (D-G)." figureDoi="10.3897/zookeys.1031.63134.figure3" httpUri="https://binary.pensoft.net/fig/531986" pageId="0" pageNumber="133">3B, C</figureCitation>
) is distinct in that it is proportionately more slender than those of
<taxonomicName id="2B75699EED7C8071BF8DA3B4B9DE60D9" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. gracillimum" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="gracillimum">
<emphasis id="994F18B780C9DB0A2E6D3FA329A3B945" italics="true" pageId="0" pageNumber="133">G. gracillimum</emphasis>
</taxonomicName>
,
<taxonomicName id="5A205353DE52D537E5F69465EA3E3CED" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. katibas" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="katibas">
<emphasis id="23B4915457E440171A0A086AEF2EC2E5" italics="true" pageId="0" pageNumber="133">G. katibas</emphasis>
</taxonomicName>
, and
<taxonomicName id="FED36C1BB06C5A1F7B1BB5392B75EDB9" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. larsi" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="larsi">
<emphasis id="A0E47FE9DEACB6B0184EEBD626E5AF5E" italics="true" pageId="0" pageNumber="133">G. larsi</emphasis>
</taxonomicName>
(cf.
<bibRefCitation id="A1A2D1D72B4CB75F2A07D86874981430" author="Ng, PKL" journalOrPublisher="Malayan Nature Journal" pageId="0" pageNumber="133" pagination="65 - 70" refId="B10" refString="Ng, PKL, 1995. Geosesarma aurantium, a new sesarmine land crab (Crustacea: Decapoda: Brachyura: Grapsidae) from Sabah. Malayan Nature Journal 49: 65 - 70" title="Geosesarma aurantium, a new sesarmine land crab (Crustacea: Decapoda: Brachyura: Grapsidae) from Sabah." volume="49" year="1995">Ng 1995</bibRefCitation>
: fig. 12A-E;
<bibRefCitation id="69B6FFDB4D653B5005C8B3885EF99478" DOI="https://doi.org/10.11646/zootaxa.4508.4.5" author="Ng, PKL" journalOrPublisher="Zootaxa" pageId="0" pageNumber="133" pagination="569 - 575" refId="B16" refString="Ng, PKL, Grinang, J, 2018. A new species of highland vampire crab (Crustacea: Brachyura: Sesarmidae: Geosesarma) from Serian, Sarawak. Zootaxa 4508 (4): 569 - 575, DOI: https://doi.org/10.11646/zootaxa.4508.4.5" title="A new species of highland vampire crab (Crustacea: Brachyura: Sesarmidae: Geosesarma) from Serian, Sarawak." url="https://doi.org/10.11646/zootaxa.4508.4.5" volume="4508" year="2018">Ng and Grinang 2018</bibRefCitation>
: fig. 5B-F,
<bibRefCitation id="9A1FD797D2B58C2C184D070C68EA7B20" DOI="https://doi.org/10.11646/zootaxa.4614.3.6" author="Ng, PYC" journalOrPublisher="Zootaxa" pageId="0" pageNumber="133" pagination="529 - 540" refId="B20" refString="Ng, PYC, Ng, PKL, 2019. Geosesarma spectrum, a new species of semiterrestrial vampire crab (Crustacea: Decapoda: Brachyura: Sesarmidae) from Brunei Darussalam, Borneo. Zootaxa 4614 (3): 529 - 540, DOI: https://doi.org/10.11646/zootaxa.4614.3.6" title="Geosesarma spectrum, a new species of semiterrestrial vampire crab (Crustacea: Decapoda: Brachyura: Sesarmidae) from Brunei Darussalam, Borneo." url="https://doi.org/10.11646/zootaxa.4614.3.6" volume="4614" year="2019">Ng and Ng 2019</bibRefCitation>
: fig. 9B-E, G, H, I-M). In addition, the distal corneous part of the G1 is almost straight in
<taxonomicName id="2A56A6768C6F432BFA16290756537448" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. sodalis" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="sodalis">
<emphasis id="94937EF05BE0189BE783F2623483497C" italics="true" pageId="0" pageNumber="133">G. sodalis</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation id="92A47C8A801AC86665460338CEB63320" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Geosesarma sodalis sp. nov., holotype male (10.1 x 9.8 mm) (ZRC 2020.0413), Sarawak A left third maxilliped (setae denuded) B left G 1 (ventral view) C left G 1 (ventral view) D distal part of left G 1 (ventral view) E distal part of left G 1 (distomesial view) F distal part of left G 1 (ventral view) G distal part of left G 1 (ventromesial view) H left G 2. Scales bars: 0.5 mm (A-C, H); 0.25 mm (D-G)." figureDoi="10.3897/zookeys.1031.63134.figure3" httpUri="https://binary.pensoft.net/fig/531986" pageId="0" pageNumber="133">3B-D, F, H-K</figureCitation>
) but gently upcurved in
<taxonomicName id="CE04611E05DE4677402C19A5C9DEF327" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. gracillimum" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="gracillimum">
<emphasis id="6E6823C9387267E5ACEACB5954AA7EF4" italics="true" pageId="0" pageNumber="133">G. gracillimum</emphasis>
</taxonomicName>
(see
<bibRefCitation id="E4A8303F7D95B17099089B1FC40C0BC4" DOI="https://doi.org/10.11646/zootaxa.4614.3.6" author="Ng, PYC" journalOrPublisher="Zootaxa" pageId="0" pageNumber="133" pagination="529 - 540" refId="B20" refString="Ng, PYC, Ng, PKL, 2019. Geosesarma spectrum, a new species of semiterrestrial vampire crab (Crustacea: Decapoda: Brachyura: Sesarmidae) from Brunei Darussalam, Borneo. Zootaxa 4614 (3): 529 - 540, DOI: https://doi.org/10.11646/zootaxa.4614.3.6" title="Geosesarma spectrum, a new species of semiterrestrial vampire crab (Crustacea: Decapoda: Brachyura: Sesarmidae) from Brunei Darussalam, Borneo." url="https://doi.org/10.11646/zootaxa.4614.3.6" volume="4614" year="2019">Ng and Ng 2019</bibRefCitation>
: fig. 9I-M). Compared to
<taxonomicName id="C6CE89B22A1556DBF5D46454295B52CB" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. bau" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="bau">
<emphasis id="ECF1BAA7F764E3A2A8FC53A095F84B79" italics="true" pageId="0" pageNumber="133">G. bau</emphasis>
</taxonomicName>
, which also has a more slender G1,
<taxonomicName id="E3F784A34B1313EE82CEFB4F6536028C" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. sodalis" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="sodalis">
<emphasis id="135D9A2609B9F195100CBDA361FB55D6" italics="true" pageId="0" pageNumber="133">G. sodalis</emphasis>
</taxonomicName>
sp. nov. has the distal part bent at an angle of about 45° along the longitudinal axis and the subdistal part of the outer margin is more angular and shelf-like (Figs
<figureCitation id="F96E189926F241996B36871E8D970362" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Geosesarma sodalis sp. nov., holotype male (10.1 x 9.8 mm) (ZRC 2020.0413), Sarawak A dorsal view of right cheliped B outer view of right chela C subdorsal view of left chela D inner view of right chela E right third ambulatory leg F right fourth ambulatory leg G pleonal somites 2 - 6 and telson H left G 1 (ventral view) I left G 1 (ventral view) J distal part of left G 1 (ventral view) K distal part of left G 1 (ventral view)." figureDoi="10.3897/zookeys.1031.63134.figure2" httpUri="https://binary.pensoft.net/fig/531985" pageId="0" pageNumber="133">2H-K</figureCitation>
,
<figureCitation id="7B852942A9240E9E6D3A3C0953A783AA" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Geosesarma sodalis sp. nov., holotype male (10.1 x 9.8 mm) (ZRC 2020.0413), Sarawak A left third maxilliped (setae denuded) B left G 1 (ventral view) C left G 1 (ventral view) D distal part of left G 1 (ventral view) E distal part of left G 1 (distomesial view) F distal part of left G 1 (ventral view) G distal part of left G 1 (ventromesial view) H left G 2. Scales bars: 0.5 mm (A-C, H); 0.25 mm (D-G)." figureDoi="10.3897/zookeys.1031.63134.figure3" httpUri="https://binary.pensoft.net/fig/531986" pageId="0" pageNumber="133">3B-D, F</figureCitation>
) (versus G1 bent at about 30° along longitudinal axis and subdistal part of outer margin is gradually sloping in
<taxonomicName id="737B3F0A68B47BF9A40AEE84AE73DCFD" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. bau" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="bau">
<emphasis id="3A4F8ED00DC4D00EB20947E6143BB245" italics="true" pageId="0" pageNumber="133">G. bau</emphasis>
</taxonomicName>
; see Ng and Grinang 2004: fig. 9D, F).
</paragraph>
<caption id="DBCC0CDDCD7BFE32F1BBB8E341C9CC78" doi="10.3897/zookeys.1031.63134.figure2" httpUri="https://binary.pensoft.net/fig/531985" pageId="0" pageNumber="133" start="Figure 2" startId="F2">
<paragraph id="491DA5FEDD57234FFB7FA80EEB3F4115" pageId="0" pageNumber="133">
<emphasis id="6C31240C2F0413B83564E2F926B8D81A" bold="true" pageId="0" pageNumber="133">Figure 2.</emphasis>
<taxonomicName id="D6312AE30FA71BC281E673F1359B9F18" authorityName="Ng" authorityYear="2021" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geosesarma sodalis" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="sodalis">
<emphasis id="5ADFAC20ED032D795D047E83A7986AE1" italics="true" pageId="0" pageNumber="133">Geosesarma sodalis</emphasis>
</taxonomicName>
sp. nov., holotype male (10.1
<normalizedToken id="86F412BECF96378B48A50BEC843F69AC" originalValue="×">x</normalizedToken>
9.8 mm) (ZRC 2020.0413), Sarawak
<emphasis id="144F8908E59EE0A2A30743A5720036BB" bold="true" pageId="0" pageNumber="133">A</emphasis>
dorsal view of right cheliped
<emphasis id="5672FA619F6DFD24BD7080A672817531" bold="true" pageId="0" pageNumber="133">B</emphasis>
outer view of right chela
<emphasis id="9F3E6D0ECB6B515F850F095D6FC90FFB" bold="true" pageId="0" pageNumber="133">C</emphasis>
subdorsal view of left chela
<emphasis id="D6BA1883CFE619D5CC9471026548B82F" bold="true" pageId="0" pageNumber="133">D</emphasis>
inner view of right chela
<emphasis id="A032DC61CD4F8DF544CD0BFE2775134B" bold="true" pageId="0" pageNumber="133">E</emphasis>
right third ambulatory leg
<emphasis id="18A3862E3FC105FDE9340D47AB927179" bold="true" pageId="0" pageNumber="133">F</emphasis>
right fourth ambulatory leg
<emphasis id="E6E15A2CFD37EC32BF96D67F5A31B279" bold="true" pageId="0" pageNumber="133">G</emphasis>
pleonal somites 2-6 and telson
<emphasis id="ADBCE5071C8516D15F56E25249B1410E" bold="true" pageId="0" pageNumber="133">H</emphasis>
left G1 (ventral view)
<emphasis id="DAEF595FE5ABC726030A8194D65D601E" bold="true" pageId="0" pageNumber="133">I</emphasis>
left G1 (ventral view)
<emphasis id="BF926156C80BD1ADBD765D076DED7FED" bold="true" pageId="0" pageNumber="133">J</emphasis>
distal part of left G1 (ventral view)
<emphasis id="D94D59B0779D6235C92F4EDC8C85F521" bold="true" pageId="0" pageNumber="133">K</emphasis>
distal part of left G1 (ventral view).
</paragraph>
</caption>
<caption id="E2F7842C4235E7D17837A8E00A95B92E" doi="10.3897/zookeys.1031.63134.figure3" httpUri="https://binary.pensoft.net/fig/531986" pageId="0" pageNumber="133" start="Figure 3" startId="F3">
<paragraph id="35A3612EBE2C6657D054B04BD97E3670" pageId="0" pageNumber="133">
<emphasis id="752F91AE80F2877972E8664FB443A019" bold="true" pageId="0" pageNumber="133">Figure 3.</emphasis>
<taxonomicName id="FBF1FF50344F9650837C1A2B044281D7" authorityName="Ng" authorityYear="2021" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geosesarma sodalis" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="sodalis">
<emphasis id="BE1AD4637DA9D8F3D6E564EC765BB31B" italics="true" pageId="0" pageNumber="133">Geosesarma sodalis</emphasis>
</taxonomicName>
sp. nov., holotype male (10.1
<normalizedToken id="F2B0886B27DF19ACCFC1E01D6D6A3FA7" originalValue="×">x</normalizedToken>
9.8 mm) (ZRC 2020.0413), Sarawak
<emphasis id="0B1867BC033F4BED93ECEDC4075B61F5" bold="true" pageId="0" pageNumber="133">A</emphasis>
left third maxilliped (setae denuded)
<emphasis id="34E1282B20EC9F7302EFD86AC2189F30" bold="true" pageId="0" pageNumber="133">B</emphasis>
left G1 (ventral view)
<emphasis id="A30BA4FF6AACF9402F2B68F8E7A1477C" bold="true" pageId="0" pageNumber="133">C</emphasis>
left G1 (ventral view)
<emphasis id="99CA931DD22BBF13F8C8ACBEEEE8C27F" bold="true" pageId="0" pageNumber="133">D</emphasis>
distal part of left G1 (ventral view)
<emphasis id="4BB17698684D2CE5DD214373AB8B2C94" bold="true" pageId="0" pageNumber="133">E</emphasis>
distal part of left G1 (distomesial view)
<emphasis id="6D0A68D7780384D5E8BFF057883C6AFE" bold="true" pageId="0" pageNumber="133">F</emphasis>
distal part of left G1 (ventral view)
<emphasis id="C006A42DF382C6DC71CB31B4D9D9FE7C" bold="true" pageId="0" pageNumber="133">G</emphasis>
distal part of left G1 (ventromesial view)
<emphasis id="EA9E51263495282AD80AA4AA65123E8A" bold="true" pageId="0" pageNumber="133">H</emphasis>
left G2. Scales bars: 0.5 mm (
<emphasis id="614D9EF4B4FDF128658D1C57D8A8CB19" bold="true" pageId="0" pageNumber="133">A-C, H</emphasis>
); 0.25 mm (
<emphasis id="B40025C45001806ED1B15CB26E9A4D85" bold="true" pageId="0" pageNumber="133">D-G</emphasis>
).
</paragraph>
</caption>
<paragraph id="14ECEEEDD2B9F031D6165ABD1ECF1511" pageId="0" pageNumber="133">
The relatively longer fingers (distinctly longer than the palm) and the outer surface of the chela with fewer small granules in
<taxonomicName id="9CB494A7D6E60BD7C3E5177AAC7597CE" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. sodalis" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="sodalis">
<emphasis id="831244351CB24303BBA7BCDCCE12B643" italics="true" pageId="0" pageNumber="133">G. sodalis</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation id="7AD765414E9DBAE28A4E8D4CF28086ED" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Geosesarma sodalis sp. nov., holotype male (10.1 x 9.8 mm) (ZRC 2020.0413), Sarawak A dorsal view of right cheliped B outer view of right chela C subdorsal view of left chela D inner view of right chela E right third ambulatory leg F right fourth ambulatory leg G pleonal somites 2 - 6 and telson H left G 1 (ventral view) I left G 1 (ventral view) J distal part of left G 1 (ventral view) K distal part of left G 1 (ventral view)." figureDoi="10.3897/zookeys.1031.63134.figure2" httpUri="https://binary.pensoft.net/fig/531985" pageId="0" pageNumber="133">2A-D</figureCitation>
), differ from the condition in
<taxonomicName id="8A6BAFB3D24A9ADA1D20AF7CE1964088" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. katibas" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="katibas">
<emphasis id="306EE11C5F3ED52F690F7DED1C0C610E" italics="true" pageId="0" pageNumber="133">G. katibas</emphasis>
</taxonomicName>
and
<taxonomicName id="E5AE0B1077A486F1CD7FEA03CB9F76C6" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. larsi" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="larsi">
<emphasis id="7C27080D420FDEDAFFA1B91E3430782F" italics="true" pageId="0" pageNumber="133">G. larsi</emphasis>
</taxonomicName>
, with the shorter fingers and the outer surface densely covered with small rounded granules (see
<bibRefCitation id="B10F913D16643532B466FE38EAA37D6E" DOI="https://doi.org/10.11646/zootaxa.4508.4.5" author="Ng, PKL" journalOrPublisher="Zootaxa" pageId="0" pageNumber="133" pagination="569 - 575" refId="B16" refString="Ng, PKL, Grinang, J, 2018. A new species of highland vampire crab (Crustacea: Brachyura: Sesarmidae: Geosesarma) from Serian, Sarawak. Zootaxa 4508 (4): 569 - 575, DOI: https://doi.org/10.11646/zootaxa.4508.4.5" title="A new species of highland vampire crab (Crustacea: Brachyura: Sesarmidae: Geosesarma) from Serian, Sarawak." url="https://doi.org/10.11646/zootaxa.4508.4.5" volume="4508" year="2018">Ng and Grinang 2018</bibRefCitation>
: figs 2D, 3A;
<bibRefCitation id="8BA3085B9BEA57DB4144455EB2DFF095" DOI="https://doi.org/10.11646/zootaxa.4614.3.6" author="Ng, PYC" journalOrPublisher="Zootaxa" pageId="0" pageNumber="133" pagination="529 - 540" refId="B20" refString="Ng, PYC, Ng, PKL, 2019. Geosesarma spectrum, a new species of semiterrestrial vampire crab (Crustacea: Decapoda: Brachyura: Sesarmidae) from Brunei Darussalam, Borneo. Zootaxa 4614 (3): 529 - 540, DOI: https://doi.org/10.11646/zootaxa.4614.3.6" title="Geosesarma spectrum, a new species of semiterrestrial vampire crab (Crustacea: Decapoda: Brachyura: Sesarmidae) from Brunei Darussalam, Borneo." url="https://doi.org/10.11646/zootaxa.4614.3.6" volume="4614" year="2019">Ng and Ng 2019</bibRefCitation>
: fig. 1C). The longer fingers of the chela most closely resemble those of
<taxonomicName id="58FA5BAE6BE58530507AC79CD27C862B" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. gracillimum" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="gracillimum">
<emphasis id="F0B7B2D1185EAC5F4DABB5FD49C4149F" italics="true" pageId="0" pageNumber="133">G. gracillimum</emphasis>
</taxonomicName>
and
<taxonomicName id="7BDFFC9A0DDE94A97971CE901C5AB9DD" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. bau" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="bau">
<emphasis id="43CBD2778D5CBF65E213304E1C219D65" italics="true" pageId="0" pageNumber="133">G. bau</emphasis>
</taxonomicName>
(see
<bibRefCitation id="F908A718277C2F495012A6F62A0B372A" author="Ng, PKL" journalOrPublisher="Malayan Nature Journal" pageId="0" pageNumber="133" pagination="65 - 70" refId="B10" refString="Ng, PKL, 1995. Geosesarma aurantium, a new sesarmine land crab (Crustacea: Decapoda: Brachyura: Grapsidae) from Sabah. Malayan Nature Journal 49: 65 - 70" title="Geosesarma aurantium, a new sesarmine land crab (Crustacea: Decapoda: Brachyura: Grapsidae) from Sabah." volume="49" year="1995">Ng 1995</bibRefCitation>
: fig. 13A; Ng and Grinang 2004: fig. 8A;
<bibRefCitation id="FE6A12B397021FFC019CD0E281D94B1F" DOI="https://doi.org/10.11646/zootaxa.4048.1.2" author="Ng, PKL" journalOrPublisher="Zootaxa" pageId="0" pageNumber="133" pagination="37 - 56" refId="B12" refString="Ng, PKL, 2015. Semiterrestrial crabs of the genus Geosesarma De Man, 1892 (Crustacea, Brachyura, Sesarmidae) from western Borneo, Indonesia, with descriptions of three new species. Zootaxa 4048 (1): 37 - 56, DOI: https://doi.org/10.11646/zootaxa.4048.1.2" title="Semiterrestrial crabs of the genus Geosesarma De Man, 1892 (Crustacea, Brachyura, Sesarmidae) from western Borneo, Indonesia, with descriptions of three new species." url="https://doi.org/10.11646/zootaxa.4048.1.2" volume="4048" year="2015">Ng 2015</bibRefCitation>
: fig. 14E, F). The male pleon of
<taxonomicName id="80808C9D374429FDC6893DD4394F670A" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. sodalis" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="sodalis">
<emphasis id="31A3DC9C1055F74E5770E6D10BD269DF" italics="true" pageId="0" pageNumber="133">G. sodalis</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation id="46FDF9724C5E56EB52AB56F45EE7CCE1" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Geosesarma sodalis sp. nov., holotype male (10.1 x 9.8 mm) (ZRC 2020.0413), Sarawak A dorsal view of right cheliped B outer view of right chela C subdorsal view of left chela D inner view of right chela E right third ambulatory leg F right fourth ambulatory leg G pleonal somites 2 - 6 and telson H left G 1 (ventral view) I left G 1 (ventral view) J distal part of left G 1 (ventral view) K distal part of left G 1 (ventral view)." figureDoi="10.3897/zookeys.1031.63134.figure2" httpUri="https://binary.pensoft.net/fig/531985" pageId="0" pageNumber="133">2G</figureCitation>
) is similar to that of
<taxonomicName id="4874679A25B9A5910667EF1AD1F7B4A1" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. katibas" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="katibas">
<emphasis id="228A6A522D925B74DEBBEB91C7A48712" italics="true" pageId="0" pageNumber="133">G. katibas</emphasis>
</taxonomicName>
(see
<bibRefCitation id="579C2C9FA4DE2B0F54ABE96CDCA11E0F" DOI="https://doi.org/10.11646/zootaxa.4614.3.6" author="Ng, PYC" journalOrPublisher="Zootaxa" pageId="0" pageNumber="133" pagination="529 - 540" refId="B20" refString="Ng, PYC, Ng, PKL, 2019. Geosesarma spectrum, a new species of semiterrestrial vampire crab (Crustacea: Decapoda: Brachyura: Sesarmidae) from Brunei Darussalam, Borneo. Zootaxa 4614 (3): 529 - 540, DOI: https://doi.org/10.11646/zootaxa.4614.3.6" title="Geosesarma spectrum, a new species of semiterrestrial vampire crab (Crustacea: Decapoda: Brachyura: Sesarmidae) from Brunei Darussalam, Borneo." url="https://doi.org/10.11646/zootaxa.4614.3.6" volume="4614" year="2019">Ng and Ng 2019</bibRefCitation>
: fig. 8D), but this character is not reliable to differentiate taxa as it varies some degree in relative widths of the somites and convexity of the lateral margins of somite 6 (
<bibRefCitation id="9A4D2D0D817D2E20E13013095AC165AB" DOI="https://doi.org/10.11646/zootaxa.4614.3.6" author="Ng, PYC" journalOrPublisher="Zootaxa" pageId="0" pageNumber="133" pagination="529 - 540" refId="B20" refString="Ng, PYC, Ng, PKL, 2019. Geosesarma spectrum, a new species of semiterrestrial vampire crab (Crustacea: Decapoda: Brachyura: Sesarmidae) from Brunei Darussalam, Borneo. Zootaxa 4614 (3): 529 - 540, DOI: https://doi.org/10.11646/zootaxa.4614.3.6" title="Geosesarma spectrum, a new species of semiterrestrial vampire crab (Crustacea: Decapoda: Brachyura: Sesarmidae) from Brunei Darussalam, Borneo." url="https://doi.org/10.11646/zootaxa.4614.3.6" volume="4614" year="2019">Ng and Ng 2019</bibRefCitation>
).
</paragraph>
<paragraph id="A779D39C68EAF664E103303BE7115AE4" pageId="0" pageNumber="133">
The male chela and G1 differences between
<taxonomicName id="F202E10EB076355E43D38D999976BBCB" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. sodalis" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="sodalis">
<emphasis id="600D5F34262597C42A8FA5C75692D7F1" italics="true" pageId="0" pageNumber="133">G. sodalis</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName id="4DFA673EFD0191C48AB74C5D014750A1" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. spectrum" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="spectrum">
<emphasis id="8CA904AA0EBFE7EF4A4B0F71182625ED" italics="true" pageId="0" pageNumber="133">G. spectrum</emphasis>
</taxonomicName>
(from Brunei) are the same as for the Sarawakian
<taxonomicName id="28595FDE2CEA9B0029A75CA46AC3B4A1" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. katibas" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="katibas">
<emphasis id="6AA39CCDEF39456094B37695F1ECB3E8" italics="true" pageId="0" pageNumber="133">G. katibas</emphasis>
</taxonomicName>
.
<taxonomicName id="6E2B20A455DAC4BB2A82B19D22B40BC1" authorityName="Ng" authorityYear="2021" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geosesarma sodalis" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="sodalis">
<emphasis id="4B14A25C078EC7B8363A01B4CAE91151" italics="true" pageId="0" pageNumber="133">Geosesarma sodalis</emphasis>
</taxonomicName>
sp. nov. differs markedly from the two species in this group from Indonesian Kalimantan,
<taxonomicName id="85552D491F26FE0C9176C952BF8E8701" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. ambawang" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="ambawang">
<emphasis id="DB6F286273AC7931C199B78EC755AEA7" italics="true" pageId="0" pageNumber="133">G. ambawang</emphasis>
</taxonomicName>
and
<taxonomicName id="8E39B586013E2104D6C3D044A6040297" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. pontianak" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="pontianak">
<emphasis id="9CE0BAF1F6AEC6E37B55717854E7C856" italics="true" pageId="0" pageNumber="133">G. pontianak</emphasis>
</taxonomicName>
, in possessing a G1 that is proportionately stouter and the subdistal part of the outer margin has a prominent right angled hump-like arch (see
<bibRefCitation id="B4B224DC470ADE138E3A02BAC82C0EAC" DOI="https://doi.org/10.11646/zootaxa.4048.1.2" author="Ng, PKL" journalOrPublisher="Zootaxa" pageId="0" pageNumber="133" pagination="37 - 56" refId="B12" refString="Ng, PKL, 2015. Semiterrestrial crabs of the genus Geosesarma De Man, 1892 (Crustacea, Brachyura, Sesarmidae) from western Borneo, Indonesia, with descriptions of three new species. Zootaxa 4048 (1): 37 - 56, DOI: https://doi.org/10.11646/zootaxa.4048.1.2" title="Semiterrestrial crabs of the genus Geosesarma De Man, 1892 (Crustacea, Brachyura, Sesarmidae) from western Borneo, Indonesia, with descriptions of three new species." url="https://doi.org/10.11646/zootaxa.4048.1.2" volume="4048" year="2015">Ng 2015</bibRefCitation>
: figs 9D-G, 13D-H, J-M). The three species in this group from the eastern Malaysia state of Sabah,
<taxonomicName id="FEF389808DB415269B0299B02938BAA2" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. sabanus" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="sabanus">
<emphasis id="1E5698052626EF3D697C9A023E9C011A" italics="true" pageId="0" pageNumber="133">G. sabanus</emphasis>
</taxonomicName>
,
<taxonomicName id="A35FAA6494A3A0D22E053710228EBEB3" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. aurantium" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="aurantium">
<emphasis id="F3EB6A676AE21B5DBB4CEEC85C3D83AC" italics="true" pageId="0" pageNumber="133">G. aurantium</emphasis>
</taxonomicName>
, and
<taxonomicName id="078013ADAD537421874C457A912643F3" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. danumense" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="danumense">
<emphasis id="2297063401EBDFA45534AE1453C03F21" italics="true" pageId="0" pageNumber="133">G. danumense</emphasis>
</taxonomicName>
differ markedly from
<taxonomicName id="27F80A17A3722042141E0519F8C8BA39" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. sodalis" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="sodalis">
<emphasis id="38FF1ADA36A2A173754F46434489AE2D" italics="true" pageId="0" pageNumber="133">G. sodalis</emphasis>
</taxonomicName>
sp. nov. in that the corneous G1 distal part is longer and distinctly spatuliform in lateral view (
<bibRefCitation id="DF6D44A5D1BC21DBB595012A1212808C" DOI="https://doi.org/10.1163/156854092X00596" author="Ng, PKL" journalOrPublisher="Crustaceana" pageId="0" pageNumber="133" pagination="210 - 213" refId="B9" refString="Ng, PKL, 1992. Geosesarma sabanus, a new sesarmine crab (Decapoda, Brachyura, Grapsidae) from the forests of Sabah, East Malaysia, Borneo. Crustaceana 63 (2): 210 - 213, DOI: https://doi.org/10.1163/156854092X00596" title="Geosesarma sabanus, a new sesarmine crab (Decapoda, Brachyura, Grapsidae) from the forests of Sabah, East Malaysia, Borneo." url="https://doi.org/10.1163/156854092X00596" volume="63" year="1992">Ng 1992</bibRefCitation>
,
<bibRefCitation id="E4A264C31150B73C9FB478FE726103B7" author="Ng, PKL" journalOrPublisher="Malayan Nature Journal" pageId="0" pageNumber="133" pagination="65 - 70" refId="B10" refString="Ng, PKL, 1995. Geosesarma aurantium, a new sesarmine land crab (Crustacea: Decapoda: Brachyura: Grapsidae) from Sabah. Malayan Nature Journal 49: 65 - 70" title="Geosesarma aurantium, a new sesarmine land crab (Crustacea: Decapoda: Brachyura: Grapsidae) from Sabah." volume="49" year="1995">1995</bibRefCitation>
,
<bibRefCitation id="4BA61D4EB788F8BFE265DA6E543CECCB" author="Ng, PKL" journalOrPublisher="Malayan Nature Journal" pageId="0" pageNumber="133" pagination="303 - 310" refId="B11" refString="Ng, PKL, 2002. On a new species of terrestrial crab of the genus Geosesarma (Crustacea: Brachyura: Sesarmidae) from Sabah, Malaysia. Malayan Nature Journal 56 (3): 303 - 310" title="On a new species of terrestrial crab of the genus Geosesarma (Crustacea: Brachyura: Sesarmidae) from Sabah, Malaysia." volume="56" year="2002">2002</bibRefCitation>
; Ng and
<bibRefCitation id="C6DAE3B0B0E23099F3108A718B8E8473" DOI="https://doi.org/10.1163/15685403-00003735" author="Ng, PKL" journalOrPublisher="Crustaceana" pageId="0" pageNumber="133" pagination="185 - 197" refId="B14" refString="Ng, PKL, 2018. On two new species of potamid and sesarmid crabs (Decapoda, Brachyura) from caves in Myanmar. Crustaceana 91 (2): 185 - 197, DOI: https://doi.org/10.1163/15685403-00003735" title="On two new species of potamid and sesarmid crabs (Decapoda, Brachyura) from caves in Myanmar." url="https://doi.org/10.1163/15685403-00003735" volume="91" year="2018">Ng 2018</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="6F7128BB9EBDBB223B9BA5B08E7FE72A" pageId="0" pageNumber="133" type="biology_ecology">
<paragraph id="2A706E73618770B6F941E1FAD6D75C65" pageId="0" pageNumber="133">Biology.</paragraph>
<paragraph id="100F627C96282398B4AA3DEBE91141D9" pageId="0" pageNumber="133">
Noteworthy is that
<taxonomicName id="0486161492622B50F156EE6756323D53" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. sodalis" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="sodalis">
<emphasis id="06A7749D81AFF6F73463B3D6F94856AC" italics="true" pageId="0" pageNumber="133">G. sodalis</emphasis>
</taxonomicName>
sp. nov. was collected inside a cave where a cavernicolous species of gecarcinucid,
<taxonomicName id="85B3AB7612176BB746DE5529DC5C8ED7" authorityName="Grinang &amp; P.K.L.Ng" authorityYear="2021" class="Crustacea" family="Gecarcinucidae" genus="Arachnothelphusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arachnothelphusa sarang" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="sarang">
<emphasis id="F73600B58D7FCE98876E35806CA6F6AE" italics="true" pageId="0" pageNumber="133">Arachnothelphusa sarang</emphasis>
</taxonomicName>
Grinang &amp; Ng, 2021, is present. Bukit Sarang is an isolated limestone outcrop with a complex of small caves, most of which probably have subterranean interconnections, and is part of the Tatau river basin in central Sarawak. The type specimen was obtained in moist areas several hundred meters from the cave entrance (RB Stuebing pers. comm.). Although more surveys in and around the Bukit Sarang were conducted in 2006 and more specimens of
<taxonomicName id="282C026CDC6A157EFD2DB3B0288B7B36" lsidName="A. sarang" pageId="0" pageNumber="133" rank="species" species="sarang">
<emphasis id="E4836EF754AF1C7AE3A412ED885BE89B" italics="true" pageId="0" pageNumber="133">A. sarang</emphasis>
</taxonomicName>
were collected (
<bibRefCitation id="371799A925B7766274527CB55AD3C4AD" author="Grinang, J" journalOrPublisher="Raffles Bulletin of Zoology" pageId="0" pageNumber="133" pagination="1 - 7" refId="B5" refString="Grinang, J, Ng, PKL, 2021. A new species of the genus Arachnothelphusa Ng, 1991 (Crustacea: Decapoda: Gecarcinucidae) from a limestone cave in Sarawak (Malaysian, Borneo). Raffles Bulletin of Zoology 69: 1 - 7" title="A new species of the genus Arachnothelphusa Ng, 1991 (Crustacea: Decapoda: Gecarcinucidae) from a limestone cave in Sarawak (Malaysian, Borneo)." volume="69" year="2021">Grinang and Ng 2021</bibRefCitation>
), no other specimens of
<taxonomicName id="47C179DDFBD0FE02EA09DE36B7D0EF55" authorityName="De Man" authorityYear="1892" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geosesarma" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="EAF5DF441279D32C79E4392DC544F5CA" italics="true" pageId="0" pageNumber="133">Geosesarma</emphasis>
</taxonomicName>
were forthcoming.
</paragraph>
<paragraph id="3E6F8617F361C222672889E4257FC120" pageId="0" pageNumber="133">
<taxonomicName id="A992EA8EE66001228A817BDD3A242A48" authorityName="Ng" authorityYear="2021" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geosesarma sodalis" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="sodalis">
<emphasis id="8B836604E2675F30240D34E95F8D62F3" italics="true" pageId="0" pageNumber="133">Geosesarma sodalis</emphasis>
</taxonomicName>
sp. nov., however, does not have prominently elongated legs or reduced eyes typical of true troglobitic taxa, and must be treated as troglophile. It is probably more widespread outside the cave habitat. The site it was collected from is several hundred metres from the cave entrance and there was no light at all. The sympatric
<taxonomicName id="ACA1C189A65ABD20A5BC1A01B7E05E87" authorityName="Grinang &amp; P.K.L.Ng" authorityYear="2021" class="Crustacea" family="Gecarcinucidae" genus="Arachnothelphusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arachnothelphusa sarang" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="sarang">
<emphasis id="735FD46952CE8F6B49D835D99785923D" italics="true" pageId="0" pageNumber="133">Arachnothelphusa sarang</emphasis>
</taxonomicName>
possesses some cave-dwelling characters-there is hardly any pigmentation on the body and legs and the pereopods are elongated, but the eyes are not reduced with the cornea still distinct, with
<bibRefCitation id="1EC7C0DFAF2FEDBE0F66FF3B81F2158D" author="Grinang, J" journalOrPublisher="Raffles Bulletin of Zoology" pageId="0" pageNumber="133" pagination="1 - 7" refId="B5" refString="Grinang, J, Ng, PKL, 2021. A new species of the genus Arachnothelphusa Ng, 1991 (Crustacea: Decapoda: Gecarcinucidae) from a limestone cave in Sarawak (Malaysian, Borneo). Raffles Bulletin of Zoology 69: 1 - 7" title="A new species of the genus Arachnothelphusa Ng, 1991 (Crustacea: Decapoda: Gecarcinucidae) from a limestone cave in Sarawak (Malaysian, Borneo)." volume="69" year="2021">Grinang and Ng (2021)</bibRefCitation>
treating it only as a troglophilic species.
</paragraph>
<paragraph id="62413941DBF93B26C2DF7D8E8CA49002" pageId="0" pageNumber="133">
No
<taxonomicName id="8A0C2E3B12D2788FE5ADEDB5186DD560" authorityName="De Man" authorityYear="1892" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geosesarma" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="A05F7EB22F1F88E6EDF86873C3DAD9F2" italics="true" pageId="0" pageNumber="133">Geosesarma</emphasis>
</taxonomicName>
species had previously been recorded from caves, although one sesarmid genus
<taxonomicName id="337ECF2DF2DCFA8D24252A5C617EAD1E" class="Crustacea" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karstarma" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="BBEE3D9B2957244E6118F4048E92EBBC" italics="true" pageId="0" pageNumber="133">Karstarma</emphasis>
</taxonomicName>
Davie &amp; Ng, 2007, is known to live in or closely associated with limestone caves.
<taxonomicName id="162D9A23FD1FFB710E9AEB4B9229943B" class="Crustacea" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karstarma" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="C59245F0A41C4E540F6A8C4CE8FDC18D" italics="true" pageId="0" pageNumber="133">Karstarma</emphasis>
</taxonomicName>
species are widely distributed in the Indo-West Pacific, with 18 recognised species (see
<bibRefCitation id="9CB44245232C889C909D3E45992E1EDE" DOI="https://doi.org/10.11646/zootaxa.4482.2.7" author="Wowor, D" journalOrPublisher="Zootaxa" pageId="0" pageNumber="133" pagination="355 - 366" refId="B25" refString="Wowor, D, Ng, PKL, 2018. A new sesarmid crab of the genus Karstarma (Crustacea: Decapoda: Brachyura) associated with limestone formations in East Java, Indonesia. Zootaxa 4482 (2): 355 - 366, DOI: https://doi.org/10.11646/zootaxa.4482.2.7" title="A new sesarmid crab of the genus Karstarma (Crustacea: Decapoda: Brachyura) associated with limestone formations in East Java, Indonesia." url="https://doi.org/10.11646/zootaxa.4482.2.7" volume="4482" year="2018">Wowor and Ng 2018</bibRefCitation>
;
<bibRefCitation id="50AF4396CA37FB20A28B1B491294003A" DOI="https://doi.org/10.11646/zootaxa.4497.3.3" author="Poupin, J" journalOrPublisher="Zootaxa" pageId="0" pageNumber="133" pagination="381 - 397" refId="B21" refString="Poupin, J, Crestey, N, Le Guelte, J-P, 2018. Cave-dwelling crabs of the genus Karstarma from lava tubes of the volcano 'Piton de la Fournaise', in Reunion Island, with description of a new species and redescription of Karstarma jacksoni (Balss, 1934) from Christmas Island. (Decapoda, Brachyura, Sesarmidae). Zootaxa 4497 (3): 381 - 397, DOI: https://doi.org/10.11646/zootaxa.4497.3.3" title="Cave-dwelling crabs of the genus Karstarma from lava tubes of the volcano ' Piton de la Fournaise', in Reunion Island, with description of a new species and redescription of Karstarma jacksoni (Balss, 1934) from Christmas Island. (Decapoda, Brachyura, Sesarmidae)." url="https://doi.org/10.11646/zootaxa.4497.3.3" volume="4497" year="2018">Poupin et al. 2018</bibRefCitation>
;
<bibRefCitation id="DC71CD499E3F03EA28A47E8F3F11EAE8" DOI="https://doi.org/10.18353/crustacea.49.0_73" author="Ng, PKL" journalOrPublisher="Crustacean Research" pageId="0" pageNumber="133" pagination="73 - 88" refId="B15" refString="Ng, PKL, 2020. Karstarma umbra, a new species of cavernicolous crab (Brachyura: Sesarmidae) from Vanuatu, with a key to the genus. Crustacean Research 49: 73 - 88, DOI: https://doi.org/10.18353/crustacea.49.0_73" title="Karstarma umbra, a new species of cavernicolous crab (Brachyura: Sesarmidae) from Vanuatu, with a key to the genus." url="https://doi.org/10.18353/crustacea.49.0_73" volume="49" year="2020">Ng 2020</bibRefCitation>
).
<bibRefCitation id="7C448D13A8F2DACC3D0F2EB8DFD902D1" DOI="https://doi.org/10.11646/zootaxa.4482.2.7" author="Wowor, D" journalOrPublisher="Zootaxa" pageId="0" pageNumber="133" pagination="355 - 366" refId="B25" refString="Wowor, D, Ng, PKL, 2018. A new sesarmid crab of the genus Karstarma (Crustacea: Decapoda: Brachyura) associated with limestone formations in East Java, Indonesia. Zootaxa 4482 (2): 355 - 366, DOI: https://doi.org/10.11646/zootaxa.4482.2.7" title="A new sesarmid crab of the genus Karstarma (Crustacea: Decapoda: Brachyura) associated with limestone formations in East Java, Indonesia." url="https://doi.org/10.11646/zootaxa.4482.2.7" volume="4482" year="2018">Wowor and Ng (2018)</bibRefCitation>
recognised three species-groups in
<taxonomicName id="DB15AA56374E3729A346C58622B7F83C" class="Crustacea" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karstarma" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="22C56AC821388F3A148ED40E939BCE0A" italics="true" pageId="0" pageNumber="133">Karstarma</emphasis>
</taxonomicName>
and discussed the affinities of one of these groups with
<taxonomicName id="36845198D289404C61842F043E28AC32" authorityName="De Man" authorityYear="1892" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geosesarma" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="3BA85C52ACD5FC34EB81EB56826D7337" italics="true" pageId="0" pageNumber="133">Geosesarma</emphasis>
</taxonomicName>
. They commented that the characters of some
<taxonomicName id="318DB2B7850F7DDE12D9F09005C1FEFB" class="Crustacea" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karstarma" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="2BB4A5FBAABA9CA7418F483A87398689" italics="true" pageId="0" pageNumber="133">Karstarma</emphasis>
</taxonomicName>
species (e.g,
<taxonomicName id="F72739E2D47F0959043370FD7B03339A" lsidName="K. microphthalmus" pageId="0" pageNumber="133" rank="species" species="microphthalmus">
<emphasis id="F65C349BF1E848C44166C755943442E7" italics="true" pageId="0" pageNumber="133">K. microphthalmus</emphasis>
</taxonomicName>
(Naruse &amp; Ng, 2007) and
<taxonomicName id="AA47AA9126D8E8D1D9EBE728A9D65CE9" lsidName="K. malang" pageId="0" pageNumber="133" rank="species" species="malang">
<emphasis id="43F7CFB3240BA458A3259A84A49A22DF" italics="true" pageId="0" pageNumber="133">K. malang</emphasis>
</taxonomicName>
Wowor &amp; Ng, 2018) are close to
<taxonomicName id="0060E44EB46E6DDE8648E806668002A4" authorityName="De Man" authorityYear="1892" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geosesarma" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="EE900BFBAE500F3E5FB5E4625BCDF81F" italics="true" pageId="0" pageNumber="133">Geosesarma</emphasis>
</taxonomicName>
. Until the present discovery of
<taxonomicName id="A241E8F79D6C987C659C06C94428ED08" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. sodalis" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="sodalis">
<emphasis id="0E6BDA4988F4D73C6B9F49CA67D5527F" italics="true" pageId="0" pageNumber="133">G. sodalis</emphasis>
</taxonomicName>
sp. nov., no species of
<taxonomicName id="1F30C79C1673437BBBDCE8DD736550F7" authorityName="De Man" authorityYear="1892" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geosesarma" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="D20F156BF19BF938EE5052262D8930D7" italics="true" pageId="0" pageNumber="133">Geosesarma</emphasis>
</taxonomicName>
has previously been found in caves.
<taxonomicName id="DBA6B93EE258B66181FE506080DBE61B" authorityName="Ng" authorityYear="2021" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geosesarma sodalis" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="sodalis">
<emphasis id="B8487F1E2935C52B73812AF2E6B5FF17" italics="true" pageId="0" pageNumber="133">Geosesarma sodalis</emphasis>
</taxonomicName>
sp. nov., however, has none of the morphological features associated with a cavernicolous lifestyle, e.g., reduced eyes and/or cornea and elongated pereopods. In any case,
<taxonomicName id="CE0EF09354D29AACD8DFEBDE5E514D99" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="G. sodalis" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="sodalis">
<emphasis id="0B6A3528ED18C59E01CF40BAF11B7D15" italics="true" pageId="0" pageNumber="133">G. sodalis</emphasis>
</taxonomicName>
sp. nov. differs markedly from the group of
<taxonomicName id="C6D3AD28EBEF6F74F92A66C29B468D56" class="Crustacea" family="Sesarmidae" genus="Karstarma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karstarma" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="E34EF09E13B80170E75EEC07DEFD1A23" italics="true" pageId="0" pageNumber="133">Karstarma</emphasis>
</taxonomicName>
species highlighted by
<bibRefCitation id="35BBEE7E415BAAF4872A7E1A2DBA1E8C" DOI="https://doi.org/10.11646/zootaxa.4482.2.7" author="Wowor, D" journalOrPublisher="Zootaxa" pageId="0" pageNumber="133" pagination="355 - 366" refId="B25" refString="Wowor, D, Ng, PKL, 2018. A new sesarmid crab of the genus Karstarma (Crustacea: Decapoda: Brachyura) associated with limestone formations in East Java, Indonesia. Zootaxa 4482 (2): 355 - 366, DOI: https://doi.org/10.11646/zootaxa.4482.2.7" title="A new sesarmid crab of the genus Karstarma (Crustacea: Decapoda: Brachyura) associated with limestone formations in East Java, Indonesia." url="https://doi.org/10.11646/zootaxa.4482.2.7" volume="4482" year="2018">Wowor and Ng (2018)</bibRefCitation>
in its quadrate carapace, proportionately shorter ambulatory legs, and stouter G1, as well as its well-developed eyes with the large pigmented cornea.
</paragraph>
<paragraph id="C52640675B9E0F0CEAD9B4B42B6576EC" pageId="0" pageNumber="133">
Another species of sesarmid which was originally desrribed from near the entrance of a cave in Myanmar,
<taxonomicName id="86FD935A54AC0066FC8A8BD2514B7DE8" authorityName="P.K.L.Ng" authorityYear="2018" class="Malacostraca" family="Sesarmidae" genus="Pseudosesarma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Pseudosesarma brehieri" order="Decapoda" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="brehieri">
<emphasis id="B8942F5DDA052AC39B93A742BD3B6FF8" italics="true" pageId="0" pageNumber="133">Pseudosesarma brehieri</emphasis>
</taxonomicName>
Ng, 2018, is now known to normally live in mangrove habitats (
<bibRefCitation id="9E61F2F1C80B4B9BF2D1F605E22FFBB4" DOI="https://doi.org/10.1163/15685403-00003735" author="Ng, PKL" journalOrPublisher="Crustaceana" pageId="0" pageNumber="133" pagination="185 - 197" refId="B14" refString="Ng, PKL, 2018. On two new species of potamid and sesarmid crabs (Decapoda, Brachyura) from caves in Myanmar. Crustaceana 91 (2): 185 - 197, DOI: https://doi.org/10.1163/15685403-00003735" title="On two new species of potamid and sesarmid crabs (Decapoda, Brachyura) from caves in Myanmar." url="https://doi.org/10.1163/15685403-00003735" volume="91" year="2018">Ng 2018</bibRefCitation>
;
<bibRefCitation id="6BEAD5BE21219823FA4414E6BE68A965" author="Schubart, CD" journalOrPublisher="Raffles Bulletin of Zoology" pageId="0" pageNumber="133" pagination="891 - 994" refId="B22" refString="Schubart, CD, Ng, PKL, 2020. Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serene &amp; Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species. Raffles Bulletin of Zoology 68: 891 - 994" title="Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serene &amp; Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species." volume="68" year="2020">Schubart and Ng 2020</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>
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