150 lines
18 KiB
XML
150 lines
18 KiB
XML
<document id="860EB1338D0D85B9E8074BB7D61DF1BF" ID-DOI="10.11646/zootaxa.5551.1.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="14389841" ID-ZooBank="BEBB4D36-855E-4AD3-829D-A3B2B17B81B7" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1733924247133" checkinUser="plazi" docAuthor="Brown, John W." docDate="2024" docId="626F651B187EFF92FF381613FBCAFB47" docLanguage="en" docName="zootaxa.5551.1.2.pdf" docOrigin="Zootaxa 5551 (1)" docSource="https://doi.org/10.11646/zootaxa.5551.1.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Transtilliana Brown 2024, new genus" docType="treatment" docVersion="2" lastPageNumber="56" masterDocId="9E561D63187AFF97FFAF1134FFAFFFE6" masterDocTitle="New genera and species of tortricid moths from Chile and Argentina (Lepidoptera: Tortricidae)" masterLastPageNumber="90" masterPageNumber="51" pageNumber="55" updateTime="1733925089986" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="5578600B4A28750367943D4FAB1D16A0">New genera and species of tortricid moths from Chile and Argentina (Lepidoptera: Tortricidae)</mods:title>
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<mods:name id="3CCCE81E43F9D9FF2E34CC0286E524F7" type="personal">
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<mods:namePart id="8F3E35E79E94CA8396C9BFE8F9EF3EF3">Brown, John W.</mods:namePart>
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<mods:title id="915AF0912D738FC445E5B2FAF802300F">Zootaxa</mods:title>
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<mods:part id="1B330804E5A3E61886E91DC1FA70CC22">
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<mods:date id="D7A87914B2CB653CFD1A60C2C55A9A80">2024</mods:date>
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<mods:number id="C3D4E24EF169E502A1A697535A65BBA6">2024-12-11</mods:number>
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<mods:number id="DDB2AB335AD1001A2981B3716A909274">5551</mods:number>
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<mods:url id="1A6BFEAC591D1C0C7ABA8EEEA75C7973">https://doi.org/10.11646/zootaxa.5551.1.2</mods:url>
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<mods:classification id="F15D1DA1616D27DB2C645BE6DE822EB9">journal article</mods:classification>
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<mods:identifier id="6005DF18A1EE7349D978789F4CC96E41" type="DOI">10.11646/zootaxa.5551.1.2</mods:identifier>
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<treatment id="626F651B187EFF92FF381613FBCAFB47" LSID="urn:lsid:plazi:treatment:626F651B187EFF92FF381613FBCAFB47" httpUri="http://treatment.plazi.org/id/626F651B187EFF92FF381613FBCAFB47" lastPageId="5" lastPageNumber="56" pageId="4" pageNumber="55">
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<subSubSection id="A2DC8786187EFF93FF381613FBAFF881" pageId="4" pageNumber="55" type="nomenclature">
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<paragraph id="EA79D40D187EFF93FF381613FD8FF8A4" blockId="4.[151,1024,1831,1895]" box="[151,544,1831,1858]" pageId="4" pageNumber="55">
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<heading id="B1316361187EFF93FF381613FD8FF8A4" bold="true" box="[151,544,1831,1858]" fontSize="11" level="1" pageId="4" pageNumber="55" reason="1">
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<emphasis id="D8B2081F187EFF93FF381613FD8FF8A4" bold="true" box="[151,544,1831,1858]" pageId="4" pageNumber="55">
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<taxonomicName id="2DC6AF8E187EFF93FF381613FE3DF8A4" authority="Brown" authorityName="Brown" authorityYear="2024" box="[151,402,1831,1858]" class="Insecta" family="Tortricidae" genus="Transtilliana" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="D8B2081F187EFF93FF381613FE9BF8A4" bold="true" box="[151,308,1831,1858]" italics="true" pageId="4" pageNumber="55">Transtilliana</emphasis>
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Brown
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</taxonomicName>
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,
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<taxonomicNameLabel id="C381B564187EFF93FE31161CFD8FF8A4" box="[414,544,1832,1858]" pageId="4" pageNumber="55" rank="genus">new genus</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="EA79D40D187EFF93FF381678FBAFF881" blockId="4.[151,1024,1831,1895]" box="[151,1024,1868,1895]" pageId="4" pageNumber="55">
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LSIDurn:lsid:zoobank.org:act:
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<uuid id="9E60EED8187EFF93FE401678FBAFF881" box="[495,1024,1868,1895]" pageId="4" pageNumber="55">94CB458B-92A2-41F2-85B1-CB44E188D222</uuid>
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</paragraph>
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</subSubSection>
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<subSubSection id="A2DC8786187EFF93FF3816A0FCA6F848" box="[151,777,1940,1967]" pageId="4" pageNumber="55" type="type_taxon">
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<paragraph id="EA79D40D187EFF93FF3816A0FCA6F848" blockId="4.[151,1436,1940,2039]" box="[151,777,1940,1967]" pageId="4" pageNumber="55">
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<emphasis id="D8B2081F187EFF93FF3816A0FE84F848" bold="true" box="[151,299,1940,1966]" pageId="4" pageNumber="55">
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<typeStatus id="357D6AAF187EFF93FF3816A0FF7DF848" box="[151,210,1940,1966]" pageId="4" pageNumber="55">Type</typeStatus>
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species
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</emphasis>
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:
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<taxonomicName id="2DC6AF8E187EFF93FE9516A0FDC0F849" authority="Brown" authorityName="Brown" authorityYear="2024" box="[314,623,1940,1967]" class="Insecta" family="Tortricidae" genus="Transtilliana" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="biloba" status="sp. nov.">
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<emphasis id="D8B2081F187EFF93FE9516A0FDB5F848" box="[314,538,1940,1966]" italics="true" pageId="4" pageNumber="55">Transtilliana biloba</emphasis>
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Brown
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</taxonomicName>
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,
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<taxonomicNameLabel id="C381B564187EFF93FDD516A1FCABF848" box="[634,772,1940,1967]" pageId="4" pageNumber="55" rank="species">new species</taxonomicNameLabel>
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.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="A2DC8786187EFF92FF68168CFC47FE6F" lastPageId="5" lastPageNumber="56" pageId="4" pageNumber="55" type="diagnosis">
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<paragraph id="EA79D40D187EFF92FF68168CFC47FE6F" blockId="4.[151,1436,1940,2039]" lastBlockId="5.[151,1437,151,1185]" lastPageId="5" lastPageNumber="56" pageId="4" pageNumber="55">
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<emphasis id="D8B2081F187EFF93FF68168CFE95F834" bold="true" box="[199,314,1976,2002]" pageId="4" pageNumber="55">Diagnosis</emphasis>
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. Superficially,
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<taxonomicName id="2DC6AF8E187EFF93FE5C168CFD2BF834" authorityName="Brown" authorityYear="2024" box="[499,644,1976,2002]" class="Insecta" family="Tortricidae" genus="Transtilliana" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="genus">
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<emphasis id="D8B2081F187EFF93FE5C168CFD2BF834" box="[499,644,1976,2002]" italics="true" pageId="4" pageNumber="55">Transtilliana</emphasis>
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</taxonomicName>
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is similar to
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<taxonomicName id="2DC6AF8E187EFF93FC9D168DFC3AF834" authorityName="Brown" authorityYear="2024" box="[818,917,1977,2002]" class="Insecta" family="Phlaeothripidae" genus="Vargasia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thysanoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="genus">
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<emphasis id="D8B2081F187EFF93FC9D168DFC3AF834" box="[818,917,1977,2002]" italics="true" pageId="4" pageNumber="55">Vargasia</emphasis>
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</taxonomicName>
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, but the male genitalia are quite different, in particular, the shapes of the uncus, transtilla, and phallus (
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<figureCitation id="72FDC888187EFF93FCCC16E8FC5CF810" box="[867,1011,2012,2038]" captionStart="FIGURES 21‒28" captionStartId="23.[151,264,1890,1914]" captionTargetBox="[151,1436,481,1867]" captionTargetId="figure-156@23.[151,1436,478,1867]" captionTargetPageId="23" captionText="FIGURES 21‒28. Male genitalia of tortricid species from Chile and Argentina. 21. Terifrona fronteria, USNM slide 145,517. 22. Transtilliana biloba, USNM slide 154,846. 23. Vargasia karsholti, Horak slide GP 733. 24. Hamiamia trancasiana, USNM slide 92,490. 25. Varifula fulvaria, USNM slide 154,848. 26. Varifula geomops, USNM slide 68,492. 27. Proeulia hasta, furca attached to phallus, USNM slide 154,802. 28. Proeulia hasta, furca in situ, USNM slide 81,233." figureDoi="http://doi.org/10.5281/zenodo.14389851" httpUri="https://zenodo.org/record/14389851/files/figure.png" pageId="4" pageNumber="55">Figs. 22, 23</figureCitation>
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). It is also somewhat reminiscent of
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<taxonomicName id="2DC6AF8E187FFF92FF3811A3FEDFFF57" authority="Brown" authorityName="Brown" authorityYear="1998" box="[151,368,151,177]" class="Insecta" family="Tortricidae" genus="Strophotina" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">
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<emphasis id="D8B2081F187FFF92FF3811A3FEB5FF57" box="[151,282,151,177]" italics="true" pageId="5" pageNumber="56">Strophotina</emphasis>
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Brown
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</taxonomicName>
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owing to its size, forewing shape and coloration, and long labial palpi (
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<figureCitation id="72FDC888187FFF92FB0011A3FAB6FF57" box="[1199,1305,151,177]" captionStart="FIGURES 1‒8" captionStartId="3.[151,264,1592,1616]" captionTargetBox="[151,1436,171,1572]" captionTargetId="figure-191@3.[151,1436,171,1572]" captionTargetPageId="3" captionText="FIGURES 1‒8.Adults of tortricid species from Chile andArgentina.1. Argentiliana basipallidana, paratype female. 2. Terifrona fronteria, holotype male. 3. Transtilliana biloba, paratype male. 4. Transtilliana biloba, paratype male. 5. Vargasia karsholti, holotype male. 6. Hamiamia trancasiana, Chile, Ñuble Province. 7. Varifula fulvaria, Chile, Aconcagua Province. 8. Varifula geomops, holotype male." figureDoi="http://doi.org/10.5281/zenodo.14389843" httpUri="https://zenodo.org/record/14389843/files/figure.png" pageId="5" pageNumber="56">Figs 3, 4</figureCitation>
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). However, whereas the forewing of
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<taxonomicName id="2DC6AF8E187FFF92FE1B118FFD98FF33" authorityName="Brown" authorityYear="1998" box="[436,567,187,213]" class="Insecta" family="Tortricidae" genus="Strophotina" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">
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<emphasis id="D8B2081F187FFF92FE1B118FFD98FF33" box="[436,567,187,213]" italics="true" pageId="5" pageNumber="56">Strophotina</emphasis>
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</taxonomicName>
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has a pronounced and well defined semicircular costal patch, only a hint of such a feature is present in
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<taxonomicName id="2DC6AF8E187FFF92FE6511EBFDF4FF1F" authorityName="Brown" authorityYear="2024" box="[458,603,223,249]" class="Insecta" family="Tortricidae" genus="Transtilliana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">
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<emphasis id="D8B2081F187FFF92FE6511EBFDF4FF1F" box="[458,603,223,249]" italics="true" pageId="5" pageNumber="56">Transtilliana</emphasis>
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</taxonomicName>
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. The genitalia of the two genera show no similarities. The combination of the narrow, parallel-sided valvae with a conspicuous excavation at the distal margin, and the bilobed transtilla serve to distinguish
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<taxonomicName id="2DC6AF8E187FFF92FE981013FE67FEA7" authorityName="Brown" authorityYear="2024" box="[311,456,295,321]" class="Insecta" family="Tortricidae" genus="Transtilliana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">
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<emphasis id="D8B2081F187FFF92FE981013FE67FEA7" box="[311,456,295,321]" italics="true" pageId="5" pageNumber="56">Transtilliana</emphasis>
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</taxonomicName>
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from all other genera. In the female genitalia, the apophyses anteriores are rather short (shorter than the apophyses posteriores), and the bilobed sterigma shows some similarities to those of
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<taxonomicName id="2DC6AF8E187FFF92FABF107FFE27FE6F" authority="Brown and Razowski" authorityName="Brown and Razowski" class="Insecta" family="Tortricidae" genus="Ptychocroca" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">
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<emphasis id="D8B2081F187FFF92FABF107FFA33FE83" box="[1296,1436,331,357]" italics="true" pageId="5" pageNumber="56">Ptychocroca</emphasis>
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Brown and Razowski
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</taxonomicName>
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and
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<taxonomicName id="2DC6AF8E187FFF92FE10105BFDC8FE6F" authority="Urra" authorityName="Urra" authorityYear="2017" box="[447,615,367,393]" genus="Foikeulia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="56" rank="genus">
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<emphasis id="D8B2081F187FFF92FE10105BFD85FE6F" box="[447,554,367,393]" italics="true" pageId="5" pageNumber="56">Foikeulia</emphasis>
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Urra
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</taxonomicName>
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, with which adults are not similar.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="A2DC8786187FFF92FF6810A6FC37FB9B" pageId="5" pageNumber="56" type="description">
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<paragraph id="EA79D40D187FFF92FF6810A6FCBDFD87" blockId="5.[151,1437,151,1185]" pageId="5" pageNumber="56">
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<emphasis id="D8B2081F187FFF92FF6810A6FEFEFE4A" bold="true" box="[199,337,402,428]" pageId="5" pageNumber="56">Description</emphasis>
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. Head: Vertex rough-scaled, with scales directed anteriorly over frons, scales of frons shorter, more appressed to surface; antenna ca. 0.5 length of forewing, non-serrate, with one row of scales per flagellomere, sensory setae 1.0‒1.1 times width of flagellomere in male, shorter and sparser in female; ocellus well developed, slightly larger in female; chaetosemata well developed; labial palpus with scaling greatly expanded distally on second segment, nearly concealing third segment, combined length of all segments 2.8‒3.0 times diameter of compound eye in male, ca. 3.5 times diameter in female.
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</paragraph>
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<paragraph id="EA79D40D187FFF92FF68135FFABCFD17" blockId="5.[151,1437,151,1185]" pageId="5" pageNumber="56">
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Thorax: Dorsum smooth-scaled, metathoracic tuft absent.Legs unmodified,without hairpencil or other secondary scales. Forewing length ca. 2.4 times width; costa evenly arched throughout, without costal fold in male; termen straight, oblique; all veins present and separate, chorda and M-stem absent, R
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<subScript id="7642D648187FFF92FC4113F1FC59FD35" attach="left" box="[1006,1014,709,723]" fontSize="6" pageId="5" pageNumber="56">4</subScript>
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||
to costa, R
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<subScript id="7642D648187FFF92FBDF13F1FBD7FD35" attach="left" box="[1136,1144,709,723]" fontSize="6" pageId="5" pageNumber="56">5</subScript>
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to termen. Hindwing with all veins present and separate; CuP weak at margin; frenulum with one bristle in male, three in female.
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</paragraph>
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<paragraph id="EA79D40D187FFF92FF6813CFFC37FB9B" blockId="5.[151,1437,151,1185]" pageId="5" pageNumber="56">
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Abdomen: Male genitalia (
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<figureCitation id="72FDC888187FFF92FE4213CFFD93FCF3" box="[493,572,763,789]" captionStart="FIGURES 21‒28" captionStartId="23.[151,264,1890,1914]" captionTargetBox="[151,1436,481,1867]" captionTargetId="figure-156@23.[151,1436,478,1867]" captionTargetPageId="23" captionText="FIGURES 21‒28. Male genitalia of tortricid species from Chile and Argentina. 21. Terifrona fronteria, USNM slide 145,517. 22. Transtilliana biloba, USNM slide 154,846. 23. Vargasia karsholti, Horak slide GP 733. 24. Hamiamia trancasiana, USNM slide 92,490. 25. Varifula fulvaria, USNM slide 154,848. 26. Varifula geomops, USNM slide 68,492. 27. Proeulia hasta, furca attached to phallus, USNM slide 154,802. 28. Proeulia hasta, furca in situ, USNM slide 81,233." figureDoi="http://doi.org/10.5281/zenodo.14389851" httpUri="https://zenodo.org/record/14389851/files/figure.png" pageId="5" pageNumber="56">Fig. 22</figureCitation>
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) with tegumen and vinculum rather elongate; uncus short, ovate from rounded-triangular expansion of dorsum of tegumen; socius short-digitate, pendant, densely covered with fine setae; gnathos arms simple, sclerotized, joined distally into V-shaped, pointed process; transtilla with a pair of small rounded, submedian lobes directed ventro-posteriorly, lacking spines; valva broadest in basal 0.33, comparatively narrow, somewhat parallel-sided in distal 0.66, outer margin with conspicuous concavity, sacculus weakly sclerotized, with small triangular process from venter ca. 0.66 from base; juxta a broadly V-shaped plate; phallus long and slender, ca. 0.9 length of valva, cornuti absent. Female genitalia (
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<figureCitation id="72FDC888187FFF92FD5F12E7FCEDFC0B" box="[752,834,979,1005]" captionStart="FIGURES 39‒44" captionStartId="27.[151,264,1948,1972]" captionTargetBox="[151,1436,520,1925]" captionTargetId="figure-170@27.[151,1436,520,1928]" captionTargetPageId="27" captionText="FIGURES 39‒44. Female genitalia of tortricid species from Chile and Argentina. 39. Argentiliana basipallida, USNM slide 154,800. 40. Terifrona fronteria, USNM slide 145,518. 41. Transtilliana biloba, USNM slide 154,847. 42. Vargasia karsholti, JWBrown slide 1216. 43. Hamiamia trancasiana, USNM slide 92491. 44. Varifula fulvaria, USNM slide 88,471." figureDoi="http://doi.org/10.5281/zenodo.14389861" httpUri="https://zenodo.org/record/14389861/files/figure.png" pageId="5" pageNumber="56">Fig. 41</figureCitation>
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) with papilla analis slipper-shaped, somewhat angled postero-laterally, ca. 3 times as long as wide; apophyses slender, posteriores ca. 1.25 length of anteriores; sterigma bilobed anteriorly, each lobe subtended posteriorly by a narrow arched sclerite; ductus bursae long, slender, uniform in width throughout, ca. 0.8 times length of corpus bursae, with antrum well-defined, sclerotized, cup-shaped; corpus bursae membranous, somewhat pear-shaped, lacking signum.
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</paragraph>
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</subSubSection>
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<subSubSection id="A2DC8786187FFF92FF6815B3FBCAFB47" box="[199,1125,1159,1185]" pageId="5" pageNumber="56" type="etymology">
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<paragraph id="EA79D40D187FFF92FF6815B3FBCAFB47" blockId="5.[151,1437,151,1185]" box="[199,1125,1159,1185]" pageId="5" pageNumber="56">
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||
<emphasis id="D8B2081F187FFF92FF6815B3FEE7FB47" bold="true" box="[199,328,1159,1185]" pageId="5" pageNumber="56">Etymology</emphasis>
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. The generic name refers to the unusual transtilla in the male genitalia.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |