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<mods:title id="D181DCF55B1A5F73E8C7DC7455DDDA8B">Taxonomic review of the genus Johnius (Perciformes: Sciaenidae) from East Malaysian waters, with comments on newly recorded Johnius species</mods:title>
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<paragraph id="E96F368FFFDD1F78FF2DC213674640E4" blockId="23.[151,554,1654,1754]" box="[151,552,1654,1681]" pageId="23" pageNumber="108">
<heading id="B22781E3FFDD1F78FF2DC213674640E4" bold="true" box="[151,552,1654,1681]" fontSize="11" level="1" pageId="23" pageNumber="108" reason="1">
<taxonomicName id="2ED04D0CFFDD1F78FF2DC213674640E4" ID-CoL="3QQ7F" authority="Sasaki, 1992" authorityName="Sasaki" authorityYear="1992" box="[151,552,1654,1681]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="23" pageNumber="108" phylum="Chordata" rank="species" species="trewavasae">
<emphasis id="DBA4EA9DFFDD1F78FF2DC213674640E4" bold="true" box="[151,552,1654,1681]" pageId="23" pageNumber="108">
<emphasis id="DBA4EA9DFFDD1F78FF2DC21364E840E4" bold="true" box="[151,390,1654,1681]" italics="true" pageId="23" pageNumber="108">Johnius trewavasae</emphasis>
<bibRefCitation id="8D414B7EFFDD1F78FE37C213674640E4" author="Sasaki, K." box="[397,552,1654,1681]" pageId="23" pageNumber="108" pagination="191 - 199" refId="ref23270" refString="Sasaki, K. (1992) Two new and two resurrected species of the sciaenid genus Johnius (Johnius) from the West Pacific. Japanese Journal of Ichthyology, 39 (3), 191 - 199. https: // doi. org / 10.1007 / BF 02905476" type="journal article" year="1992">Sasaki, 1992</bibRefCitation>
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<subSubSection id="A1CA6504FFDD1F78FF2DC2FE674440C0" box="[151,554,1691,1717]" pageId="23" pageNumber="108" type="vernacular_names">
<paragraph id="E96F368FFFDD1F78FF2DC2FE674440C0" blockId="23.[151,554,1654,1754]" box="[151,554,1691,1717]" pageId="23" pageNumber="108">(Common name: Trewavas croaker)</paragraph>
</subSubSection>
<subSubSection id="A1CA6504FFDD1F78FF2DC2DA64F140AC" box="[151,415,1727,1754]" pageId="23" pageNumber="108" type="description">
<paragraph id="E96F368FFFDD1F78FF2DC2DA64F140AC" blockId="23.[151,554,1654,1754]" box="[151,415,1727,1754]" pageId="23" pageNumber="108">
(
<figureCitation id="71EB2A0AFFDD1F78FF25C2DA647240AF" box="[159,284,1727,1754]" captionStart-0="FIGURE 15" captionStart-1="FIGURE 16" captionStartId-0="24.[151,250,831,855]" captionStartId-1="24.[151,250,1650,1674]" captionTargetBox-0="[151,1436,181,807]" captionTargetBox-1="[337,1259,964,1626]" captionTargetId-0="figure-27@24.[151,1436,181,807]" captionTargetId-1="figure-54@24.[328,1259,951,1626]" captionTargetPageId-0="24" captionTargetPageId-1="24" captionText-0="FIGURE 15. Photographs of the comparative materials. Johnius trewavasae, NMMB-P33709, non-type, 170.9 mm SL, Sarikei fish landing port, Sarawak, Malaysia, 14 March 2017." captionText-1="FIGURE 16. New record of site of collection samples of Johnius trewavasae in Sarikei (SRK), Sarawak, East Malaysia." figureDoi-0="http://doi.org/10.5281/zenodo.14388241" figureDoi-1="http://doi.org/10.5281/zenodo.14388243" httpUri-0="https://zenodo.org/record/14388241/files/figure.png" httpUri-1="https://zenodo.org/record/14388243/files/figure.png" pageId="23" pageNumber="108">Fig. 1516</figureCitation>
,
<tableCitation id="A4520334FFDD1F78FE9DC2DA64F940AC" box="[295,407,1727,1753]" captionStart-0="TABLE 8" captionStart-1="TABLE 9" captionStartId-0="25.[151,239,1052,1076]" captionStartId-1="26.[152,244,1601,1625]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionText-0="TABLE 8. Morphometric and meristic measurements (expressed as %HL, %SL and %ED) of Johnius trewavasae." captionText-1="TABLE 9. Genetic distances for the COI gene among congener species with Johnius trewavasae species and Dendrophysa russelii as an outgroup." httpUri-0="http://table.plazi.org/id/BDAF6607FFD31F76FF2DC07960314241" httpUri-1="http://table.plazi.org/id/BDAF6607FFD01F75FF22C22464FF4008" pageId="23" pageNumber="108" tableUuid-0="BDAF6607FFD31F76FF2DC07960314241" tableUuid-1="BDAF6607FFD01F75FF22C22464FF4008">Table 89</tableCitation>
)
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<treatmentCitationGroup id="C9C011A1FFDD1F78FF2DC36660F2416E" box="[151,1436,1795,1820]" pageId="23" pageNumber="108">
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<taxonomicName id="2ED04D0CFFDD1F78FF2DC366674D416E" ID-CoL="3QQ7F" authority="Sasaki, 1992: 191" authorityName="Sasaki" authorityPageNumber="191" authorityYear="1992" box="[151,547,1795,1819]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="23" pageNumber="108" phylum="Chordata" rank="species" species="trewavasae">
<emphasis id="DBA4EA9DFFDD1F78FF2DC366640F416E" box="[151,353,1795,1819]" italics="true" pageId="23" pageNumber="108">Johnius trewavasae</emphasis>
<bibRefCitation id="8D414B7EFFDD1F78FED0C366674D416E" author="Sasaki, K." box="[362,547,1795,1819]" pageId="23" pageNumber="108" pagination="191 - 199" refId="ref23270" refString="Sasaki, K. (1992) Two new and two resurrected species of the sciaenid genus Johnius (Johnius) from the West Pacific. Japanese Journal of Ichthyology, 39 (3), 191 - 199. https: // doi. org / 10.1007 / BF 02905476" type="journal article" year="1992">Sasaki, 1992: 191</bibRefCitation>
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(type locality:
<collectingCountry id="91C7761FFFDD1F78FD79C3666663416E" box="[707,781,1795,1819]" name="Taiwan" pageId="23" pageNumber="108">Taiwan</collectingCountry>
Strait.
<typeStatus id="366B882DFFDD1F78FCE6C36666D0416E" box="[860,958,1795,1819]" pageId="23" pageNumber="108" type="holotype">Holotype</typeStatus>
:
<specimenCode id="B9769EF4FFDD1F78FC73C3616102416E" box="[969,1132,1795,1819]" collectionCode="HUMZ" country="Japan" httpUri="http://grbio.org/cool/8gr0-wfn9" name="Hokkaido University, Laboratory of Marine Zoology" pageId="23" pageNumber="108">HUMZ 109504</specimenCode>
.
<typeStatus id="366B882DFFDD1F78FBCDC36161B04169" box="[1143,1246,1796,1820]" pageId="23" pageNumber="108" type="paratype">Paratypes</typeStatus>
:
<specimenCode id="B9769EF4FFDD1F78FB52C3666001416E" box="[1256,1391,1795,1819]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="23" pageNumber="108">ANSP 52863</specimenCode>
(1),
</treatmentCitationGroup>
53478 (2); CAS-SU 60859 (1), 60870 (1), 60918 (1), 61045 (3, 2 c&amp;s), 61080 (2), 61484 (2); HUMZ 109501 (1).
<taxonomicName id="2ED04D0CFFDD1F78FF2DC3246484412C" ID-CoL="3QQ7F" authority="Sasaki, 1992" authorityName="Sasaki" authorityYear="1992" box="[151,490,1857,1881]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="23" pageNumber="108" phylum="Chordata" rank="species" species="trewavasae">
<emphasis id="DBA4EA9DFFDD1F78FF2DC3246430412D" box="[151,350,1857,1881]" italics="true" pageId="23" pageNumber="108">Johnius trewavasae</emphasis>
<bibRefCitation id="8D414B7EFFDD1F78FEDFC3246484412C" author="Sasaki, K." box="[357,490,1857,1881]" pageId="23" pageNumber="108" pagination="191 - 199" refId="ref23270" refString="Sasaki, K. (1992) Two new and two resurrected species of the sciaenid genus Johnius (Johnius) from the West Pacific. Japanese Journal of Ichthyology, 39 (3), 191 - 199. https: // doi. org / 10.1007 / BF 02905476" type="journal article" year="1992">Sasaki, 1992</bibRefCitation>
</taxonomicName>
:
<bibRefCitation id="8D414B7EFFDD1F78FE4EC32467C2412D" author="Sasaki, K." box="[500,684,1856,1881]" pageId="23" pageNumber="108" refId="ref23416" refString="Sasaki, K. (2000) A checklist of the fishes of the South China Sea. In: Randall, J. E. &amp; Lim, K. K. P. (Eds.), Raffles Bulletin of Zoology Supplement, 8, pp. 621." type="journal volume" year="2000">Sasaki, 2000: 621</bibRefCitation>
;
<bibRefCitation id="8D414B7EFFDD1F78FD0CC3246615412D" author="Sasaki, K." box="[694,891,1856,1881]" pageId="23" pageNumber="108" pagination="3117 - 3174" refId="ref23468" refString="Sasaki, K. (2001) Sciaenidae. In: Carpenter, K. E. &amp; Niem, V. H. FAO Species Identification Guide for Fishery Purposes. The Living Marine Resources of the Western Central Pacific. Vol. 5. Bony Fishes. Part 3. Menidae to Pomacentridae. FAO, Rome, pp. 3117 - 3174." type="book chapter" year="2001">Sasaki, 2001: 3152</bibRefCitation>
;
<bibRefCitation id="8D414B7EFFDD1F78FC3FC324613D412C" author="Ng, H. H. &amp; Tan, H. H. &amp; Lim, K. K. &amp; Ludt, W. B. &amp; Chakrabarty, P." box="[901,1107,1856,1881]" pageId="23" pageNumber="108" pagination="303 - 337" refId="ref22998" refString="Ng, H. H., Tan, H. H., Lim, K. K., Ludt, W. B. &amp; Chakrabarty, P. (2015) Fishes of the eastern Johor Strait. Raffles Bulletin of Zoology, 31, 303 - 337." type="journal article" year="2015">
Ng
<emphasis id="DBA4EA9DFFDD1F78FC10C32466B0412C" box="[938,990,1857,1881]" italics="true" pageId="23" pageNumber="108">et al.</emphasis>
, 2015: 327
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A1CA6504FFDD1F78FF2DC3E36695419C" pageId="23" pageNumber="108" type="materials_examined">
<paragraph id="E96F368FFFDD1F78FF2DC3E36695419C" blockId="23.[151,1436,1926,2025]" pageId="23" pageNumber="108">
<emphasis id="DBA4EA9DFFDD1F78FF2DC3E364EC41D5" bold="true" box="[151,386,1926,1952]" pageId="23" pageNumber="108">Material examined.</emphasis>
Non-types:
<materialsCitation id="59B83CD2FFDD1F78FDB4C3E260F641D4" box="[526,1432,1927,1954]" collectingDate="2010-06-01" country="China" location="Guangdong" pageId="23" pageNumber="108" specimenCode="NMMB-P15218" specimenCount="1" stateProvince="Guangdong">
<specimenCode id="B9769EF4FFDD1F78FDB4C3E267A841D7" box="[526,710,1927,1954]" collectionCode="NMMB-P" pageId="23" pageNumber="108">NMMB-P15218</specimenCode>
(5, 117.11 to
<quantity id="2E289B6AFFDD1F78FCDFC3E2668841D4" box="[869,998,1927,1953]" metricMagnitude="-1" metricUnit="m" metricValue="1.4172999999999998" pageId="23" pageNumber="108" unit="mm" value="141.73">141.73 mm</quantity>
SL),
<collectingRegion id="2B14F86DFFDD1F78FB9DC3E261C341D4" box="[1063,1197,1927,1953]" country="China" name="Guangdong" pageId="23" pageNumber="108">Guangdong</collectingRegion>
,
<collectingCountry id="91C7761FFFDD1F78FB03C3E2619141D4" box="[1209,1279,1927,1953]" name="China" pageId="23" pageNumber="108">China</collectingCountry>
,
<date id="9D6E104FFFDD1F78FAB1C3E260F641D4" box="[1291,1432,1927,1953]" pageId="23" pageNumber="108" value="2010-06-01">
<collectingDate id="8D2AE9A7FFDD1F78FAB1C3E260F641D4" box="[1291,1432,1927,1953]" pageId="23" pageNumber="108" value="2010-06-01">1 June 2010</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="59B83CD2FFDD1F78FF2DC3CE61DE41B0" box="[151,1200,1963,1989]" collectingDate="2017-03-14" country="Malaysia" location="Sarikei" pageId="23" pageNumber="108" specimenCode="NMMB-P37087" specimenCount="1">
<specimenCode id="B9769EF4FFDD1F78FF2DC3CE642141B0" box="[151,335,1963,1989]" collectionCode="NMMB-P" pageId="23" pageNumber="108">NMMB-P37087</specimenCode>
(4, 110.49 to
<quantity id="2E289B6AFFDD1F78FE49C3CE671B41B0" box="[499,629,1963,1989]" metricMagnitude="-1" metricUnit="m" metricValue="1.3629" pageId="23" pageNumber="108" unit="mm" value="136.29">136.29 mm</quantity>
SL),
<location id="EC0F6054FFDD1F78FD0DC3CE666741B0" LSID="urn:lsid:plazi:treatment:61798799FFDD1F74FF2DC21360EC40A7:EC0F6054FFDD1F78FD0DC3CE666741B0" box="[695,777,1963,1989]" country="Malaysia" name="Sarikei" pageId="23" pageNumber="108">Sarikei</location>
,
<collectingCountry id="91C7761FFFDD1F78FCADC3CE661441B0" box="[791,890,1963,1989]" name="Malaysia" pageId="23" pageNumber="108">Sarawak</collectingCountry>
,
<collectingCountry id="91C7761FFFDD1F78FC3DC3CE669E41B0" box="[903,1008,1963,1989]" name="Malaysia" pageId="23" pageNumber="108">Malaysia</collectingCountry>
,
<date id="9D6E104FFFDD1F78FC44C3CE61DE41B0" box="[1022,1200,1963,1989]" pageId="23" pageNumber="108" value="2017-03-14">
<collectingDate id="8D2AE9A7FFDD1F78FC44C3CE61DE41B0" box="[1022,1200,1963,1989]" pageId="23" pageNumber="108" value="2017-03-14">14 March 2017</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="59B83CD2FFDD1F78FB07C3CE6699419C" collectingDate="2014-06-10" country="Malaysia" location="Tanjung Gemok" pageId="23" pageNumber="108" specimenCode="NMMB-P23653" specimenCount="1" stateProvince="Pahang">
<specimenCode id="B9769EF4FFDD1F78FB07C3CE601B41B3" box="[1213,1397,1963,1990]" collectionCode="NMMB-P" pageId="23" pageNumber="108">NMMB-P23653</specimenCode>
(3, 129.67 to
<quantity id="2E289B6AFFDD1F78FEB2C3AA64E6419C" box="[264,392,1999,2025]" metricMagnitude="-1" metricUnit="m" metricValue="1.3381" pageId="23" pageNumber="108" unit="mm" value="133.81">133.81 mm</quantity>
SL),
<location id="EC0F6054FFDD1F78FE7CC3AA6713419C" LSID="urn:lsid:plazi:treatment:61798799FFDD1F74FF2DC21360EC40A7:EC0F6054FFDD1F78FE7CC3AA6713419C" box="[454,637,1999,2025]" country="Malaysia" name="Tanjung Gemok" pageId="23" pageNumber="108" stateProvince="Pahang">Tanjung Gemok</location>
,
<collectingRegion id="2B14F86DFFDD1F78FD33C3AA67B0419C" box="[649,734,1999,2025]" country="Malaysia" name="Pahang" pageId="23" pageNumber="108">Pahang</collectingRegion>
,
<collectingCountry id="91C7761FFFDD1F78FD53C3AA663C419C" box="[745,850,1999,2025]" name="Malaysia" pageId="23" pageNumber="108">Malaysia</collectingCountry>
,
<date id="9D6E104FFFDD1F78FCE4C3AA6699419C" box="[862,1015,1999,2025]" pageId="23" pageNumber="108" value="2014-06-10">
<collectingDate id="8D2AE9A7FFDD1F78FCE4C3AA6699419C" box="[862,1015,1999,2025]" pageId="23" pageNumber="108" value="2014-06-10">10 June 2014</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="BDAF6607FFD21F77FF2DC75A67D5450E" ID-DOI="http://doi.org/10.5281/zenodo.14388241" ID-Zenodo-Dep="14388241" httpUri="https://zenodo.org/record/14388241/files/figure.png" pageId="24" pageNumber="109" startId="24.[151,250,831,855]" targetBox="[151,1436,181,807]" targetPageId="24" targetType="figure">
<paragraph id="E96F368FFFD21F77FF2DC75A67D5450E" blockId="24.[151,1436,831,891]" pageId="24" pageNumber="109">
<emphasis id="DBA4EA9DFFD21F77FF2DC75A64714522" bold="true" box="[151,287,831,855]" pageId="24" pageNumber="109">FIGURE 15.</emphasis>
Photographs of the comparative materials.
<taxonomicName id="2ED04D0CFFD21F77FD6EC75A66F54522" authorityName="Sasaki" authorityYear="1992" box="[724,923,831,855]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="109" phylum="Chordata" rank="species" species="trewavasae">
<emphasis id="DBA4EA9DFFD21F77FD6EC75A66F54522" box="[724,923,831,855]" italics="true" pageId="24" pageNumber="109">Johnius trewavasae</emphasis>
</taxonomicName>
, NMMB-P33709, non-type, 170.9 mm SL, Sarikei fish landing port, Sarawak, Malaysia, 14 March 2017.
</paragraph>
</caption>
<caption id="BDAF6607FFD21F77FF2DC217600E40FF" ID-DOI="http://doi.org/10.5281/zenodo.14388243" ID-Zenodo-Dep="14388243" box="[151,1376,1650,1674]" httpUri="https://zenodo.org/record/14388243/files/figure.png" pageId="24" pageNumber="109" startId="24.[151,250,1650,1674]" targetBox="[337,1259,964,1626]" targetPageId="24" targetType="figure">
<paragraph id="E96F368FFFD21F77FF2DC217600E40FF" blockId="24.[151,1376,1650,1674]" box="[151,1376,1650,1674]" pageId="24" pageNumber="109">
<emphasis id="DBA4EA9DFFD21F77FF2DC217644E40FF" bold="true" box="[151,288,1650,1674]" pageId="24" pageNumber="109">FIGURE 16.</emphasis>
New record of site of collection samples of
<taxonomicName id="2ED04D0CFFD21F77FD5BC21766C640FF" authorityName="Sasaki" authorityYear="1992" box="[737,936,1650,1674]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="109" phylum="Chordata" rank="species" species="trewavasae">
<emphasis id="DBA4EA9DFFD21F77FD5BC21766C640FF" box="[737,936,1650,1674]" italics="true" pageId="24" pageNumber="109">Johnius trewavasae</emphasis>
</taxonomicName>
in Sarikei (SRK), Sarawak, East Malaysia.
</paragraph>
</caption>
<subSubSection id="A1CA6504FFD21F77FF7DC2B965864117" pageId="24" pageNumber="109" type="diagnosis">
<paragraph id="E96F368FFFD21F77FF7DC2B965864117" blockId="24.[151,1437,1756,2035]" pageId="24" pageNumber="109">
<emphasis id="DBA4EA9DFFD21F77FF7DC2B964544083" bold="true" box="[199,314,1756,1782]" pageId="24" pageNumber="109">Diagnosis</emphasis>
. A species of
<taxonomicName id="2ED04D0CFFD21F77FE60C2B9675F4083" authorityName="Bloch" authorityYear="1793" box="[474,561,1756,1782]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="109" phylum="Chordata" rank="genus">
<emphasis id="DBA4EA9DFFD21F77FE60C2B9675F4083" box="[474,561,1756,1782]" italics="true" pageId="24" pageNumber="109">Johnius</emphasis>
</taxonomicName>
with the following combination of characters: snout steep, obtusely rounded; chin lacking barbel; scales above lateral line 4 to 6, below lateral line 7 to 10; gill rakers very short, obtuse, gill rakers at lower limb 8 to 10; second anal spine length 25 to 37% HL; scales ctenoid on body, large, and easily fallout.
</paragraph>
</subSubSection>
<subSubSection id="A1CA6504FFD21F76FF7DC309602D44F0" lastPageId="25" lastPageNumber="110" pageId="24" pageNumber="109" type="description">
<paragraph id="E96F368FFFD21F77FF7DC309641B41BB" blockId="24.[151,1437,1756,2035]" pageId="24" pageNumber="109">
<emphasis id="DBA4EA9DFFD21F77FF7DC309643741F3" bold="true" box="[199,345,1900,1926]" pageId="24" pageNumber="109">Description.</emphasis>
Counts and measurements of the
<typeStatus id="366B882DFFD21F77FD5CC308667841F2" box="[742,790,1901,1927]" pageId="24" pageNumber="109">type</typeStatus>
specimens are shown in
<tableCitation id="A4520334FFD21F77FB84C30961F841F2" box="[1086,1174,1900,1927]" captionStart="TABLE 8" captionStartId="25.[151,239,1052,1076]" captionTargetPageId="25" captionText="TABLE 8. Morphometric and meristic measurements (expressed as %HL, %SL and %ED) of Johnius trewavasae." httpUri="http://table.plazi.org/id/BDAF6607FFD31F76FF2DC07960314241" pageId="24" pageNumber="109" tableUuid="BDAF6607FFD31F76FF2DC07960314241">Table 8</tableCitation>
. The following data is provided for the
<bibRefCitation id="8D414B7EFFD21F77FEE3C3F5649841DE" author="Sasaki, K." box="[345,502,1936,1963]" pageId="24" pageNumber="109" pagination="191 - 199" refId="ref23270" refString="Sasaki, K. (1992) Two new and two resurrected species of the sciaenid genus Johnius (Johnius) from the West Pacific. Japanese Journal of Ichthyology, 39 (3), 191 - 199. https: // doi. org / 10.1007 / BF 02905476" type="journal article" year="1992">Sasaki (1992)</bibRefCitation>
measurement first as comparison, followed by the size range and mean for the 4 voucher specimens.
</paragraph>
<paragraph id="E96F368FFFD21F76FF7DC3BD602D44F0" blockId="24.[151,1437,1756,2035]" lastBlockId="25.[151,1437,151,1005]" lastPageId="25" lastPageNumber="110" pageId="24" pageNumber="109">Small species with a moderately elongated body; body depth 25 to 29% of standard length. Eye moderately large, 18 to 24% of head length. Interorbital width 23 to 27% of head length. Snout steep, obtusely rounded, projecting slightly in front of upper jaw; mouth small, inferior; upper jaw extending backward below front margin of pupil; no barbel on chin; teeth differentiated into large and small in upper jaw only, the large ones close-set, not canine like, forming outer series; lower jaw with a band of villiform teeth. Gill rakers short, obtuse, about 1/5 length of gill filaments at angle of arch, 6 to 8 on lower limb of first gill arch. Dorsal fin with X to XI (most frequently X) spines, followed by a notch, second part of fin with I spine and 26 to 31 soft rays; anal fin with II spines and 7 or 8 (most frequently 7) soft rays, second spine rather short, slender, 25 to 35% of head length; caudal fin rhomboidal. Scales large, those on flanks much larger than those on the lateral line, 4 or 6 scale rows above lateral line to origin of dorsal fin, 6 to 14 scale rows below lateral line to origin of anal fin; scales cycloid (smooth) on head and throat, ctenoid (rough to the touch) on other parts of head and body; small scales present on soft parts of dorsal and anal fins; lateral line scales reaching tip of caudal fin. Swimbladder hammer-shaped, with about 14 pairs of arborescent appendages along its sides, the first pair entering head beyond transverse septum and sending a palmate branch to front of the pectoral arch. Sagitta with a tadpole-shaped impression, with long axis of its head lying obliquely to that of the sagitta and tail expanded and deepened as hollow cone connected with the head by a narrow groove.</paragraph>
</subSubSection>
<subSubSection id="A1CA6504FFD31F76FF7DC6EB66B644DC" box="[199,984,654,681]" pageId="25" pageNumber="110" type="distribution">
<paragraph id="E96F368FFFD31F76FF7DC6EB66B644DC" blockId="25.[151,1437,151,1005]" box="[199,984,654,681]" pageId="25" pageNumber="110">
<emphasis id="DBA4EA9DFFD31F76FF7DC6EB643744DD" bold="true" box="[199,345,654,680]" pageId="25" pageNumber="110">Distribution</emphasis>
.
<collectingCountry id="91C7761FFFD31F76FEDDC6EA64B544DC" box="[359,475,655,681]" name="Singapore" pageId="25" pageNumber="110">Singapore</collectingCountry>
, north of
<collectingCountry id="91C7761FFFD31F76FDFDC6EA67F244DC" box="[583,668,655,681]" name="Taiwan" pageId="25" pageNumber="110">Taiwan</collectingCountry>
,
<collectingCountry id="91C7761FFFD31F76FD1DC6EA664344DC" box="[679,813,655,681]" name="Hong Kong" pageId="25" pageNumber="110">Hong Kong</collectingCountry>
, and Shanghai.
</paragraph>
</subSubSection>
<subSubSection id="A1CA6504FFD31F74FF7DC6D660EC40A7" lastPageId="27" lastPageNumber="112" pageId="25" pageNumber="110" type="discussion">
<paragraph id="E96F368FFFD31F76FF7DC6D6644A4598" blockId="25.[151,1437,151,1005]" pageId="25" pageNumber="110">
<emphasis id="DBA4EA9DFFD31F76FF7DC6D6645A44B8" bold="true" box="[199,308,691,717]" pageId="25" pageNumber="110">Remarks</emphasis>
. Among the
<taxonomicName id="2ED04D0CFFD31F76FE7FC6D667E344B8" box="[453,653,691,717]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="110" phylum="Chordata" rank="subGenus" subGenus="Johnius">
<emphasis id="DBA4EA9DFFD31F76FE7FC6D6677244B8" box="[453,540,691,717]" italics="true" pageId="25" pageNumber="110">Johnius</emphasis>
(
<emphasis id="DBA4EA9DFFD31F76FD96C6D667ED44B8" box="[556,643,691,717]" italics="true" pageId="25" pageNumber="110">Johnius</emphasis>
)
</taxonomicName>
species with pointed snout and large ctenoid scales (46 scales above lateral line),
<taxonomicName id="2ED04D0CFFD31F76FE9EC6B264D24485" authorityName="Sasaki" authorityYear="1992" box="[292,444,727,752]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="110" phylum="Chordata" rank="species" species="trewavasae">
<emphasis id="DBA4EA9DFFD31F76FE9EC6B264D24485" box="[292,444,727,752]" italics="true" pageId="25" pageNumber="110">J. trewavasae</emphasis>
</taxonomicName>
can be mainly differentiated from
<taxonomicName id="2ED04D0CFFD31F76FC86C6B266894484" baseAuthorityName="Lal Mohan" baseAuthorityYear="1976" box="[828,999,727,753]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="110" phylum="Chordata" rank="species" species="macrorhynus">
<emphasis id="DBA4EA9DFFD31F76FC86C6B266894484" box="[828,999,727,753]" italics="true" pageId="25" pageNumber="110">J. macrorhynus</emphasis>
</taxonomicName>
and
<taxonomicName id="2ED04D0CFFD31F76FBA6C6B261DE4484" authorityName="Bleeker" authorityYear="1873" box="[1052,1200,727,753]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="110" phylum="Chordata" rank="species" species="heterolepis">
<emphasis id="DBA4EA9DFFD31F76FBA6C6B261DE4484" box="[1052,1200,727,753]" italics="true" pageId="25" pageNumber="110">J. heterolepis</emphasis>
</taxonomicName>
by the lower number of gills raker count on the lower limb (obtuse, short,
<quantity id="2E289B6AFFD31F76FD5DC69E66524560" box="[743,828,763,789]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" metricValueMax="2.54" metricValueMin="2.032" pageId="25" pageNumber="110" unit="in" value="9.0" valueMax="10.0" valueMin="8.0">810 in</quantity>
<taxonomicName id="2ED04D0CFFD31F76FCF9C69E66AC4561" authorityName="Sasaki" authorityYear="1992" box="[835,962,763,788]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="110" phylum="Chordata" rank="species" species="trewavasae">
<emphasis id="DBA4EA9DFFD31F76FCF9C69E66AC4561" box="[835,962,763,788]" italics="true" pageId="25" pageNumber="110">trewavasae</emphasis>
</taxonomicName>
vs. stumpy, short,
<quantity id="2E289B6AFFD31F76FB2FC69E61B24560" box="[1173,1244,763,789]" metricMagnitude="-1" metricUnit="m" metricValue="1.905" metricValueMax="2.032" metricValueMin="1.778" pageId="25" pageNumber="110" unit="in" value="7.5" valueMax="8.0" valueMin="7.0">78 in</quantity>
<taxonomicName id="2ED04D0CFFD31F76FB59C69E601B4560" baseAuthorityName="Lal Mohan" baseAuthorityYear="1976" box="[1251,1397,763,789]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="110" phylum="Chordata" rank="species" species="macrorhynus">
<emphasis id="DBA4EA9DFFD31F76FB59C69E601B4560" box="[1251,1397,763,789]" italics="true" pageId="25" pageNumber="110">macrorhynus</emphasis>
</taxonomicName>
vs. slender, short,
<quantity id="2E289B6AFFD31F76FE87C77A64FC454C" box="[317,402,799,825]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" metricValueMax="2.794" metricValueMin="2.286" pageId="25" pageNumber="110" unit="in" value="10.0" valueMax="11.0" valueMin="9.0">911 in</quantity>
<taxonomicName id="2ED04D0CFFD31F76FE20C77A677B454C" authorityName="Bleeker" authorityYear="1873" box="[410,533,799,825]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="110" phylum="Chordata" rank="species" species="heterolepis">
<emphasis id="DBA4EA9DFFD31F76FE20C77A677B454C" box="[410,533,799,825]" italics="true" pageId="25" pageNumber="110">heterolepis</emphasis>
</taxonomicName>
) and a shorter lower jaw (33.838.4% HL in
<taxonomicName id="2ED04D0CFFD31F76FBA6C77A61F5454D" authorityName="Sasaki" authorityYear="1992" box="[1052,1179,799,824]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="110" phylum="Chordata" rank="species" species="trewavasae">
<emphasis id="DBA4EA9DFFD31F76FBA6C77A61F5454D" box="[1052,1179,799,824]" italics="true" pageId="25" pageNumber="110">trewavasae</emphasis>
</taxonomicName>
vs. 33.838.4% HL in
<taxonomicName id="2ED04D0CFFD31F76FF2DC72664474528" baseAuthorityName="Lal Mohan" baseAuthorityYear="1976" box="[151,297,835,861]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="110" phylum="Chordata" rank="species" species="macrorhynus">
<emphasis id="DBA4EA9DFFD31F76FF2DC72664474528" box="[151,297,835,861]" italics="true" pageId="25" pageNumber="110">macrorhynus</emphasis>
</taxonomicName>
vs. 40.344.2% HL in
<taxonomicName id="2ED04D0CFFD31F76FD95C72667C44528" authorityName="Bleeker" authorityYear="1873" box="[559,682,835,861]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="110" phylum="Chordata" rank="species" species="heterolepis">
<emphasis id="DBA4EA9DFFD31F76FD95C72667C44528" box="[559,682,835,861]" italics="true" pageId="25" pageNumber="110">heterolepis</emphasis>
</taxonomicName>
). Meanwhile, main criteria of
<taxonomicName id="2ED04D0CFFD31F76FBB9C72661DE4528" baseAuthorityName="Lal Mohan" baseAuthorityYear="1976" box="[1027,1200,835,861]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="110" phylum="Chordata" rank="species" species="macrorhynus">
<emphasis id="DBA4EA9DFFD31F76FBB9C72661DE4528" box="[1027,1200,835,861]" italics="true" pageId="25" pageNumber="110">J. macrorhynus</emphasis>
</taxonomicName>
can be distinguished by
<taxonomicName id="2ED04D0CFFD31F76FF03C702643C45F5" authorityName="Sasaki" authorityYear="1992" box="[185,338,871,896]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="110" phylum="Chordata" rank="species" species="trewavasae">
<emphasis id="DBA4EA9DFFD31F76FF03C702643C45F5" box="[185,338,871,896]" italics="true" pageId="25" pageNumber="110">J. trewavasae</emphasis>
</taxonomicName>
by the presence of an inner, short row of molariform teeth at posteriorly. In addition,
<taxonomicName id="2ED04D0CFFD31F76FABEC70260F345F5" authorityName="Sasaki" authorityYear="1992" box="[1284,1437,871,896]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="110" phylum="Chordata" rank="species" species="trewavasae">
<emphasis id="DBA4EA9DFFD31F76FABEC70260F345F5" box="[1284,1437,871,896]" italics="true" pageId="25" pageNumber="110">J. trewavasae</emphasis>
</taxonomicName>
can be easily differs from
<taxonomicName id="2ED04D0CFFD31F76FE0FC7EE674445D1" baseAuthorityName="Cuvier" baseAuthorityYear="1830" box="[437,554,907,932]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="110" phylum="Chordata" rank="species" species="carouna">
<emphasis id="DBA4EA9DFFD31F76FE0FC7EE674445D1" box="[437,554,907,932]" italics="true" pageId="25" pageNumber="110">J. carouna</emphasis>
</taxonomicName>
and
<taxonomicName id="2ED04D0CFFD31F76FDE4C7EE67AB45D0" box="[606,709,907,933]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="110" phylum="Chordata" rank="species" species="sasakii">
<emphasis id="DBA4EA9DFFD31F76FDE4C7EE67AB45D0" box="[606,709,907,933]" italics="true" pageId="25" pageNumber="110">J. sasakii</emphasis>
</taxonomicName>
in having the slender and rather short second anal spine fin length (2437% of HL in
<taxonomicName id="2ED04D0CFFD31F76FED5C7CA648045BD" authorityName="Sasaki" authorityYear="1992" box="[367,494,943,968]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="110" phylum="Chordata" rank="species" species="trewavasae">
<emphasis id="DBA4EA9DFFD31F76FED5C7CA648045BD" box="[367,494,943,968]" italics="true" pageId="25" pageNumber="110">trewavasae</emphasis>
</taxonomicName>
vs. stiff, long, 3249% of HL in
<taxonomicName id="2ED04D0CFFD31F76FCD2C7CA66AA45BD" baseAuthorityName="Cuvier" baseAuthorityYear="1830" box="[872,964,943,968]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="110" phylum="Chordata" rank="species" species="carouna">
<emphasis id="DBA4EA9DFFD31F76FCD2C7CA66AA45BD" box="[872,964,943,968]" italics="true" pageId="25" pageNumber="110">carouna</emphasis>
</taxonomicName>
vs. stiff, short, 2535% of HL in
<taxonomicName id="2ED04D0CFFD31F76FAFFC7CA60FD45BC" box="[1349,1427,943,969]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="110" phylum="Chordata" rank="species" species="sasakii">
<emphasis id="DBA4EA9DFFD31F76FAFFC7CA60FD45BC" box="[1349,1427,943,969]" italics="true" pageId="25" pageNumber="110">sasakii</emphasis>
</taxonomicName>
) respectively.
</paragraph>
<caption id="BDAF6607FFD31F76FF2DC07960314241" ID-Table-UUID="BDAF6607FFD31F76FF2DC07960314241" box="[151,1375,1052,1077]" httpUri="http://table.plazi.org/id/BDAF6607FFD31F76FF2DC07960314241" pageId="25" pageNumber="110" startId="25.[151,239,1052,1076]" targetBox="[163,1323,1093,1962]" targetIsTable="true" targetPageId="25" targetType="table">
<paragraph id="E96F368FFFD31F76FF2DC07960314241" blockId="25.[151,1375,1052,1077]" box="[151,1375,1052,1077]" pageId="25" pageNumber="110">
<emphasis id="DBA4EA9DFFD31F76FF2DC07964654241" bold="true" box="[151,267,1052,1076]" pageId="25" pageNumber="110">TABLE 8.</emphasis>
Morphometric and meristic measurements (expressed as %HL, %SL and %ED) of
<taxonomicName id="2ED04D0CFFD31F76FB3CC07960354241" authorityName="Sasaki" authorityYear="1992" box="[1158,1371,1052,1076]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="110" phylum="Chordata" rank="species" species="trewavasae">Johnius trewavasae</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="E96F368FFFD31F76FDFEC020604541DF" pageId="25" pageNumber="110">
<table id="9BD0C42FFFD3E090FF19C020604541DF" box="[163,1323,1093,1962]" gridcols="6" gridrows="22" pageId="25" pageNumber="110">
<tr id="57E034CDFFD3E090FF19C02060454228" box="[163,1323,1093,1117]" gridrow="0" pageId="25" pageNumber="110" rowspan-0="1" rowspan-5="1">
<th id="14315DB1FFD3E090FDFEC02067BE4228" box="[580,720,1093,1117]" gridcol="1" gridrow="0" pageId="25" pageNumber="110">
<bibRefCitation id="8D414B7EFFD31F76FDFEC02067BE4228" author="Sasaki, K." box="[580,720,1093,1117]" pageId="25" pageNumber="110" pagination="191 - 199" refId="ref23270" refString="Sasaki, K. (1992) Two new and two resurrected species of the sciaenid genus Johnius (Johnius) from the West Pacific. Japanese Journal of Ichthyology, 39 (3), 191 - 199. https: // doi. org / 10.1007 / BF 02905476" type="journal article" year="1992">Sasaki (1992)</bibRefCitation>
</th>
<th id="14315DB1FFD3E090FD43C02061C14228" box="[761,1199,1093,1117]" colspan="3" colspanRight="2" gridcol="2" gridrow="0" pageId="25" pageNumber="110">Voucher specimen, NMMB-P 37087 (N=4)</th>
</tr>
<tr id="57E034CDFFD3E090FF19C008604542F0" box="[163,1323,1133,1157]" gridrow="1" pageId="25" pageNumber="110" rowspan-0="1" rowspan-1="1">
<td id="14315DB1FFD3E090FD43C008665C42F0" box="[761,818,1133,1157]" gridcol="2" gridrow="1" pageId="25" pageNumber="110">Min</td>
<td id="14315DB1FFD3E090FC08C008668542F0" box="[946,1003,1133,1157]" gridcol="3" gridrow="1" pageId="25" pageNumber="110">Max</td>
<td id="14315DB1FFD3E090FBDAC00861C142F0" box="[1120,1199,1133,1157]" gridcol="4" gridrow="1" pageId="25" pageNumber="110">Mean</td>
<td id="14315DB1FFD3E090FAB1C008604542F0" box="[1291,1323,1133,1157]" gridcol="5" gridrow="1" pageId="25" pageNumber="110">SD</td>
</tr>
<tr id="57E034CDFFD3E090FF19C0F0604542D8" box="[163,1323,1173,1197]" gridrow="2" pageId="25" pageNumber="110" rowspan-5="1">
<th id="14315DB1FFD3E090FF19C0F0648B42D8" box="[163,485,1173,1197]" gridcol="0" gridrow="2" pageId="25" pageNumber="110">Standard Length (mm)</th>
<td id="14315DB1FFD3E090FDFEC0F067BE42D8" box="[580,720,1173,1197]" gridcol="1" gridrow="2" pageId="25" pageNumber="110">110.9</td>
<td id="14315DB1FFD3E090FD43C0F0665C42D8" box="[761,818,1173,1197]" gridcol="2" gridrow="2" pageId="25" pageNumber="110">100.0</td>
<td id="14315DB1FFD3E090FC08C0F0668542D8" box="[946,1003,1173,1197]" gridcol="3" gridrow="2" pageId="25" pageNumber="110">146.9</td>
<td id="14315DB1FFD3E090FBDAC0F061C142D8" box="[1120,1199,1173,1197]" gridcol="4" gridrow="2" pageId="25" pageNumber="110">117.7</td>
</tr>
<tr id="57E034CDFFD3E090FF19C0D8604542A3" box="[163,1323,1213,1238]" gridrow="3" pageId="25" pageNumber="110" rowspan-5="1">
<th id="14315DB1FFD3E090FF19C0D8648B42A3" box="[163,485,1213,1238]" gridcol="0" gridrow="3" pageId="25" pageNumber="110">Head Length (mm)</th>
<td id="14315DB1FFD3E090FDFEC0D867BE42A3" box="[580,720,1213,1238]" gridcol="1" gridrow="3" pageId="25" pageNumber="110">34.4</td>
<td id="14315DB1FFD3E090FD43C0D8665C42A3" box="[761,818,1213,1238]" gridcol="2" gridrow="3" pageId="25" pageNumber="110">29.2</td>
<td id="14315DB1FFD3E090FC08C0D8668542A3" box="[946,1003,1213,1238]" gridcol="3" gridrow="3" pageId="25" pageNumber="110">44.2</td>
<td id="14315DB1FFD3E090FBDAC0D861C142A3" box="[1120,1199,1213,1238]" gridcol="4" gridrow="3" pageId="25" pageNumber="110">35.6</td>
</tr>
<tr id="57E034CDFFD3E090FF19C0836045428B" box="[163,1323,1254,1278]" gridrow="4" pageId="25" pageNumber="110" rowspan-5="1">
<th id="14315DB1FFD3E090FF19C083648B428B" box="[163,485,1254,1278]" gridcol="0" gridrow="4" pageId="25" pageNumber="110">Eye Diameter (mm)</th>
<td id="14315DB1FFD3E090FDFEC08367BE428B" box="[580,720,1254,1278]" gridcol="1" gridrow="4" pageId="25" pageNumber="110">7.4</td>
<td id="14315DB1FFD3E090FD43C083665C428B" box="[761,818,1254,1278]" gridcol="2" gridrow="4" pageId="25" pageNumber="110">6.6</td>
<td id="14315DB1FFD3E090FC08C0836685428B" box="[946,1003,1254,1278]" gridcol="3" gridrow="4" pageId="25" pageNumber="110">8.8</td>
<td id="14315DB1FFD3E090FBDAC08361C1428B" box="[1120,1199,1254,1278]" gridcol="4" gridrow="4" pageId="25" pageNumber="110">7.6</td>
</tr>
<tr id="57E034CDFFD3E090FF19C16B60454353" box="[163,1323,1294,1318]" gridrow="5" pageId="25" pageNumber="110" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="14315DB1FFD3E090FF19C16B648B4353" box="[163,485,1294,1318]" gridcol="0" gridrow="5" pageId="25" pageNumber="110">Counts</th>
</tr>
<tr id="57E034CDFFD3E090FF19C1506045433B" box="[163,1323,1333,1358]" gridrow="6" pageId="25" pageNumber="110">
<th id="14315DB1FFD3E090FF19C150648B433B" box="[163,485,1333,1358]" gridcol="0" gridrow="6" pageId="25" pageNumber="110">1st dorsal-fin spine</th>
<td id="14315DB1FFD3E090FDFEC15067BE433B" box="[580,720,1333,1358]" gridcol="1" gridrow="6" pageId="25" pageNumber="110">10</td>
<td id="14315DB1FFD3E090FD43C150665C433B" box="[761,818,1333,1358]" gridcol="2" gridrow="6" pageId="25" pageNumber="110">10</td>
<td id="14315DB1FFD3E090FC08C1506685433B" box="[946,1003,1333,1358]" gridcol="3" gridrow="6" pageId="25" pageNumber="110">10</td>
<td id="14315DB1FFD3E090FBDAC15061C1433B" box="[1120,1199,1333,1358]" gridcol="4" gridrow="6" pageId="25" pageNumber="110">10</td>
<td id="14315DB1FFD3E090FAB1C1506045433B" box="[1291,1323,1333,1358]" gridcol="5" gridrow="6" pageId="25" pageNumber="110">0.0</td>
</tr>
<tr id="57E034CDFFD3E090FF19C13B60454303" box="[163,1323,1374,1398]" gridrow="7" pageId="25" pageNumber="110">
<th id="14315DB1FFD3E090FF19C13B648B4303" box="[163,485,1374,1398]" gridcol="0" gridrow="7" pageId="25" pageNumber="110">2nd dorsal-fin spine</th>
<td id="14315DB1FFD3E090FDFEC13B67BE4303" box="[580,720,1374,1398]" gridcol="1" gridrow="7" pageId="25" pageNumber="110">1</td>
<td id="14315DB1FFD3E090FD43C13B665C4303" box="[761,818,1374,1398]" gridcol="2" gridrow="7" pageId="25" pageNumber="110">1</td>
<td id="14315DB1FFD3E090FC08C13B66854303" box="[946,1003,1374,1398]" gridcol="3" gridrow="7" pageId="25" pageNumber="110">1</td>
<td id="14315DB1FFD3E090FBDAC13B61C14303" box="[1120,1199,1374,1398]" gridcol="4" gridrow="7" pageId="25" pageNumber="110">1</td>
<td id="14315DB1FFD3E090FAB1C13B60454303" box="[1291,1323,1374,1398]" gridcol="5" gridrow="7" pageId="25" pageNumber="110">0.0</td>
</tr>
<tr id="57E034CDFFD3E090FF19C1E2604543EA" box="[163,1323,1415,1439]" gridrow="8" pageId="25" pageNumber="110">
<th id="14315DB1FFD3E090FF19C1E2648B43EA" box="[163,485,1415,1439]" gridcol="0" gridrow="8" pageId="25" pageNumber="110">2nd dorsal-fin soft rays</th>
<td id="14315DB1FFD3E090FDFEC1E267BE43EA" box="[580,720,1415,1439]" gridcol="1" gridrow="8" pageId="25" pageNumber="110">27</td>
<td id="14315DB1FFD3E090FD43C1E2665C43EA" box="[761,818,1415,1439]" gridcol="2" gridrow="8" pageId="25" pageNumber="110">26</td>
<td id="14315DB1FFD3E090FC08C1E2668543EA" box="[946,1003,1415,1439]" gridcol="3" gridrow="8" pageId="25" pageNumber="110">31</td>
<td id="14315DB1FFD3E090FBDAC1E261C143EA" box="[1120,1199,1415,1439]" gridcol="4" gridrow="8" pageId="25" pageNumber="110">28</td>
<td id="14315DB1FFD3E090FAB1C1E2604543EA" box="[1291,1323,1415,1439]" gridcol="5" gridrow="8" pageId="25" pageNumber="110">0.0</td>
</tr>
<tr id="57E034CDFFD3E090FF19C1CA604543B2" box="[163,1323,1455,1479]" gridrow="9" pageId="25" pageNumber="110">
<th id="14315DB1FFD3E090FF19C1CA648B43B2" box="[163,485,1455,1479]" gridcol="0" gridrow="9" pageId="25" pageNumber="110">Anal-fin spine</th>
<td id="14315DB1FFD3E090FDFEC1CA67BE43B2" box="[580,720,1455,1479]" gridcol="1" gridrow="9" pageId="25" pageNumber="110">2</td>
<td id="14315DB1FFD3E090FD43C1CA665C43B2" box="[761,818,1455,1479]" gridcol="2" gridrow="9" pageId="25" pageNumber="110">2</td>
<td id="14315DB1FFD3E090FC08C1CA668543B2" box="[946,1003,1455,1479]" gridcol="3" gridrow="9" pageId="25" pageNumber="110">2</td>
<td id="14315DB1FFD3E090FBDAC1CA61C143B2" box="[1120,1199,1455,1479]" gridcol="4" gridrow="9" pageId="25" pageNumber="110">2</td>
<td id="14315DB1FFD3E090FAB1C1CA604543B2" box="[1291,1323,1455,1479]" gridcol="5" gridrow="9" pageId="25" pageNumber="110">0.0</td>
</tr>
<tr id="57E034CDFFD3E090FF19C1B260454385" box="[163,1323,1495,1520]" gridrow="10" pageId="25" pageNumber="110">
<th id="14315DB1FFD3E090FF19C1B2648B4385" box="[163,485,1495,1520]" gridcol="0" gridrow="10" pageId="25" pageNumber="110">Anal-fin soft rays</th>
<td id="14315DB1FFD3E090FDFEC1B267BE4385" box="[580,720,1495,1520]" gridcol="1" gridrow="10" pageId="25" pageNumber="110">7</td>
<td id="14315DB1FFD3E090FD43C1B2665C4385" box="[761,818,1495,1520]" gridcol="2" gridrow="10" pageId="25" pageNumber="110">7</td>
<td id="14315DB1FFD3E090FC08C1B266854385" box="[946,1003,1495,1520]" gridcol="3" gridrow="10" pageId="25" pageNumber="110">7</td>
<td id="14315DB1FFD3E090FBDAC1B261C14385" box="[1120,1199,1495,1520]" gridcol="4" gridrow="10" pageId="25" pageNumber="110">7</td>
<td id="14315DB1FFD3E090FAB1C1B260454385" box="[1291,1323,1495,1520]" gridcol="5" gridrow="10" pageId="25" pageNumber="110">0.0</td>
</tr>
<tr id="57E034CDFFD3E090FF19C19A6045406D" box="[163,1323,1535,1560]" gridrow="11" pageId="25" pageNumber="110">
<th id="14315DB1FFD3E090FF19C19A648B406D" box="[163,485,1535,1560]" gridcol="0" gridrow="11" pageId="25" pageNumber="110">Pectoral-fin rays</th>
<td id="14315DB1FFD3E090FDFEC19A67BE406D" box="[580,720,1535,1560]" gridcol="1" gridrow="11" pageId="25" pageNumber="110">17</td>
<td id="14315DB1FFD3E090FD43C19A665C406D" box="[761,818,1535,1560]" gridcol="2" gridrow="11" pageId="25" pageNumber="110">11</td>
<td id="14315DB1FFD3E090FC08C19A6685406D" box="[946,1003,1535,1560]" gridcol="3" gridrow="11" pageId="25" pageNumber="110">18</td>
<td id="14315DB1FFD3E090FBDAC19A61C1406D" box="[1120,1199,1535,1560]" gridcol="4" gridrow="11" pageId="25" pageNumber="110">16</td>
<td id="14315DB1FFD3E090FAB1C19A6045406D" box="[1291,1323,1535,1560]" gridcol="5" gridrow="11" pageId="25" pageNumber="110">0.0</td>
</tr>
<tr id="57E034CDFFD3E090FF19C24260454035" box="[163,1323,1575,1600]" gridrow="12" pageId="25" pageNumber="110">
<th id="14315DB1FFD3E090FF19C242648B4035" box="[163,485,1575,1600]" gridcol="0" gridrow="12" pageId="25" pageNumber="110">Outer gill rakers of 1st arch</th>
<td id="14315DB1FFD3E090FDFEC24267BE4035" box="[580,720,1575,1600]" gridcol="1" gridrow="12" pageId="25" pageNumber="110">15</td>
<td id="14315DB1FFD3E090FD43C242665C4035" box="[761,818,1575,1600]" gridcol="2" gridrow="12" pageId="25" pageNumber="110">13</td>
<td id="14315DB1FFD3E090FC08C24266854035" box="[946,1003,1575,1600]" gridcol="3" gridrow="12" pageId="25" pageNumber="110">15</td>
<td id="14315DB1FFD3E090FBDAC24261C14035" box="[1120,1199,1575,1600]" gridcol="4" gridrow="12" pageId="25" pageNumber="110">14</td>
<td id="14315DB1FFD3E090FAB1C24260454035" box="[1291,1323,1575,1600]" gridcol="5" gridrow="12" pageId="25" pageNumber="110">0.0</td>
</tr>
<tr id="57E034CDFFD3E090FF19C2356045401D" box="[163,1323,1616,1640]" gridrow="13" pageId="25" pageNumber="110">
<th id="14315DB1FFD3E090FF19C235648B401D" box="[163,485,1616,1640]" gridcol="0" gridrow="13" pageId="25" pageNumber="110">upper limb</th>
<td id="14315DB1FFD3E090FDFEC23567BE401D" box="[580,720,1616,1640]" gridcol="1" gridrow="13" pageId="25" pageNumber="110">6</td>
<td id="14315DB1FFD3E090FD43C235665C401D" box="[761,818,1616,1640]" gridcol="2" gridrow="13" pageId="25" pageNumber="110">4</td>
<td id="14315DB1FFD3E090FC08C2356685401D" box="[946,1003,1616,1640]" gridcol="3" gridrow="13" pageId="25" pageNumber="110">6</td>
<td id="14315DB1FFD3E090FBDAC23561C1401D" box="[1120,1199,1616,1640]" gridcol="4" gridrow="13" pageId="25" pageNumber="110">5</td>
<td id="14315DB1FFD3E090FAB1C2356045401D" box="[1291,1323,1616,1640]" gridcol="5" gridrow="13" pageId="25" pageNumber="110">0.0</td>
</tr>
<tr id="57E034CDFFD3E090FF19C21D604540E5" box="[163,1323,1656,1680]" gridrow="14" pageId="25" pageNumber="110">
<th id="14315DB1FFD3E090FF19C21D648B40E5" box="[163,485,1656,1680]" gridcol="0" gridrow="14" pageId="25" pageNumber="110">lower limb</th>
<td id="14315DB1FFD3E090FDFEC21D67BE40E5" box="[580,720,1656,1680]" gridcol="1" gridrow="14" pageId="25" pageNumber="110">9</td>
<td id="14315DB1FFD3E090FD43C21D665C40E5" box="[761,818,1656,1680]" gridcol="2" gridrow="14" pageId="25" pageNumber="110">8</td>
<td id="14315DB1FFD3E090FC08C21D668540E5" box="[946,1003,1656,1680]" gridcol="3" gridrow="14" pageId="25" pageNumber="110">10</td>
<td id="14315DB1FFD3E090FBDAC21D61C140E5" box="[1120,1199,1656,1680]" gridcol="4" gridrow="14" pageId="25" pageNumber="110">9</td>
<td id="14315DB1FFD3E090FAB1C21D604540E5" box="[1291,1323,1656,1680]" gridcol="5" gridrow="14" pageId="25" pageNumber="110">0.0</td>
</tr>
<tr id="57E034CDFFD3E090FF19C2FA604540CD" box="[163,1323,1695,1720]" gridrow="15" pageId="25" pageNumber="110">
<th id="14315DB1FFD3E090FF19C2FA648B40CD" box="[163,485,1695,1720]" gridcol="0" gridrow="15" pageId="25" pageNumber="110">Inner gill rakers of 1st arch</th>
<td id="14315DB1FFD3E090FDFEC2FA67BE40CD" box="[580,720,1695,1720]" gridcol="1" gridrow="15" pageId="25" pageNumber="110">11</td>
<td id="14315DB1FFD3E090FD43C2FA665C40CD" box="[761,818,1695,1720]" gridcol="2" gridrow="15" pageId="25" pageNumber="110">10</td>
<td id="14315DB1FFD3E090FC08C2FA668540CD" box="[946,1003,1695,1720]" gridcol="3" gridrow="15" pageId="25" pageNumber="110">13</td>
<td id="14315DB1FFD3E090FBDAC2FA61C140CD" box="[1120,1199,1695,1720]" gridcol="4" gridrow="15" pageId="25" pageNumber="110">11</td>
<td id="14315DB1FFD3E090FAB1C2FA604540CD" box="[1291,1323,1695,1720]" gridcol="5" gridrow="15" pageId="25" pageNumber="110">0.0</td>
</tr>
<tr id="57E034CDFFD3E090FF19C2AC60454094" box="[163,1323,1737,1761]" gridrow="16" pageId="25" pageNumber="110">
<th id="14315DB1FFD3E090FF19C2AC648B4094" box="[163,485,1737,1761]" gridcol="0" gridrow="16" pageId="25" pageNumber="110">upper limb</th>
<td id="14315DB1FFD3E090FDFEC2AC67BE4094" box="[580,720,1737,1761]" gridcol="1" gridrow="16" pageId="25" pageNumber="110">3</td>
<td id="14315DB1FFD3E090FD43C2AC665C4094" box="[761,818,1737,1761]" gridcol="2" gridrow="16" pageId="25" pageNumber="110">3</td>
<td id="14315DB1FFD3E090FC08C2AC66854094" box="[946,1003,1737,1761]" gridcol="3" gridrow="16" pageId="25" pageNumber="110">4</td>
<td id="14315DB1FFD3E090FBDAC2AC61C14094" box="[1120,1199,1737,1761]" gridcol="4" gridrow="16" pageId="25" pageNumber="110">3</td>
<td id="14315DB1FFD3E090FAB1C2AC60454094" box="[1291,1323,1737,1761]" gridcol="5" gridrow="16" pageId="25" pageNumber="110">0.0</td>
</tr>
<tr id="57E034CDFFD3E090FF19C2946045417C" box="[163,1323,1777,1801]" gridrow="17" pageId="25" pageNumber="110">
<th id="14315DB1FFD3E090FF19C294648B417C" box="[163,485,1777,1801]" gridcol="0" gridrow="17" pageId="25" pageNumber="110">lower limb</th>
<td id="14315DB1FFD3E090FDFEC29467BE417C" box="[580,720,1777,1801]" gridcol="1" gridrow="17" pageId="25" pageNumber="110">8</td>
<td id="14315DB1FFD3E090FD43C294665C417C" box="[761,818,1777,1801]" gridcol="2" gridrow="17" pageId="25" pageNumber="110">6</td>
<td id="14315DB1FFD3E090FC08C2946685417C" box="[946,1003,1777,1801]" gridcol="3" gridrow="17" pageId="25" pageNumber="110">9</td>
<td id="14315DB1FFD3E090FBDAC29461C1417C" box="[1120,1199,1777,1801]" gridcol="4" gridrow="17" pageId="25" pageNumber="110">8</td>
<td id="14315DB1FFD3E090FAB1C2946045417C" box="[1291,1323,1777,1801]" gridcol="5" gridrow="17" pageId="25" pageNumber="110">0.0</td>
</tr>
<tr id="57E034CDFFD3E090FF19C37C60454144" box="[163,1323,1817,1841]" gridrow="18" pageId="25" pageNumber="110">
<th id="14315DB1FFD3E090FF19C37C648B4144" box="[163,485,1817,1841]" gridcol="0" gridrow="18" pageId="25" pageNumber="110">Pored lateral line scales</th>
<td id="14315DB1FFD3E090FDFEC37C67BE4144" box="[580,720,1817,1841]" gridcol="1" gridrow="18" pageId="25" pageNumber="110">51</td>
<td id="14315DB1FFD3E090FD43C37C665C4144" box="[761,818,1817,1841]" gridcol="2" gridrow="18" pageId="25" pageNumber="110">37</td>
<td id="14315DB1FFD3E090FC08C37C66854144" box="[946,1003,1817,1841]" gridcol="3" gridrow="18" pageId="25" pageNumber="110">49</td>
<td id="14315DB1FFD3E090FBDAC37C61C14144" box="[1120,1199,1817,1841]" gridcol="4" gridrow="18" pageId="25" pageNumber="110">47</td>
<td id="14315DB1FFD3E090FAB1C37C60454144" box="[1291,1323,1817,1841]" gridcol="5" gridrow="18" pageId="25" pageNumber="110">0.0</td>
</tr>
<tr id="57E034CDFFD3E090FF19C3246045412C" box="[163,1323,1857,1881]" gridrow="19" pageId="25" pageNumber="110">
<th id="14315DB1FFD3E090FF19C324648B412C" box="[163,485,1857,1881]" gridcol="0" gridrow="19" pageId="25" pageNumber="110">Scales above lateral line</th>
<td id="14315DB1FFD3E090FDFEC32467BE412C" box="[580,720,1857,1881]" gridcol="1" gridrow="19" pageId="25" pageNumber="110">6</td>
<td id="14315DB1FFD3E090FD43C324665C412C" box="[761,818,1857,1881]" gridcol="2" gridrow="19" pageId="25" pageNumber="110">4</td>
<td id="14315DB1FFD3E090FC08C3246685412C" box="[946,1003,1857,1881]" gridcol="3" gridrow="19" pageId="25" pageNumber="110">6</td>
<td id="14315DB1FFD3E090FBDAC32461C1412C" box="[1120,1199,1857,1881]" gridcol="4" gridrow="19" pageId="25" pageNumber="110">5</td>
<td id="14315DB1FFD3E090FAB1C3246045412C" box="[1291,1323,1857,1881]" gridcol="5" gridrow="19" pageId="25" pageNumber="110">0.0</td>
</tr>
<tr id="57E034CDFFD3E090FF19C30C604541F7" box="[163,1323,1897,1922]" gridrow="20" pageId="25" pageNumber="110">
<th id="14315DB1FFD3E090FF19C30C648B41F7" box="[163,485,1897,1922]" gridcol="0" gridrow="20" pageId="25" pageNumber="110">Scales below lateral line</th>
<td id="14315DB1FFD3E090FDFEC30C67BE41F7" box="[580,720,1897,1922]" gridcol="1" gridrow="20" pageId="25" pageNumber="110">9</td>
<td id="14315DB1FFD3E090FD43C30C665C41F7" box="[761,818,1897,1922]" gridcol="2" gridrow="20" pageId="25" pageNumber="110">6</td>
<td id="14315DB1FFD3E090FC08C30C668541F7" box="[946,1003,1897,1922]" gridcol="3" gridrow="20" pageId="25" pageNumber="110">14</td>
<td id="14315DB1FFD3E090FBDAC30C61C141F7" box="[1120,1199,1897,1922]" gridcol="4" gridrow="20" pageId="25" pageNumber="110">10</td>
<td id="14315DB1FFD3E090FAB1C30C604541F7" box="[1291,1323,1897,1922]" gridcol="5" gridrow="20" pageId="25" pageNumber="110">0.0</td>
</tr>
<tr id="57E034CDFFD3E090FF19C3F7604541DF" box="[163,1323,1938,1962]" gridrow="21" pageId="25" pageNumber="110">
<th id="14315DB1FFD3E090FF19C3F7648B41DF" box="[163,485,1938,1962]" gridcol="0" gridrow="21" pageId="25" pageNumber="110">Circumpeduncular scales</th>
<td id="14315DB1FFD3E090FDFEC3F767BE41DF" box="[580,720,1938,1962]" gridcol="1" gridrow="21" pageId="25" pageNumber="110">14</td>
<td id="14315DB1FFD3E090FD43C3F7665C41DF" box="[761,818,1938,1962]" gridcol="2" gridrow="21" pageId="25" pageNumber="110">11</td>
<td id="14315DB1FFD3E090FC08C3F7668541DF" box="[946,1003,1938,1962]" gridcol="3" gridrow="21" pageId="25" pageNumber="110">17</td>
<td id="14315DB1FFD3E090FBDAC3F761C141DF" box="[1120,1199,1938,1962]" gridcol="4" gridrow="21" pageId="25" pageNumber="110">14</td>
<td id="14315DB1FFD3E090FAB1C3F7604541DF" box="[1291,1323,1938,1962]" gridcol="5" gridrow="21" pageId="25" pageNumber="110">0.0</td>
</tr>
</table>
</paragraph>
<paragraph id="E96F368FFFD31F76FBEEC3DE60FF41A7" blockId="25.[1108,1425,1979,2003]" box="[1108,1425,1979,2003]" pageId="25" pageNumber="110">
<emphasis id="DBA4EA9DFFD31F76FBEEC3DE60FF41A7" box="[1108,1425,1979,2003]" italics="true" pageId="25" pageNumber="110">
<tableNote id="14363701FFD31F76FBEEC3DE60FF41A7" box="[1108,1425,1979,2003]" pageId="25" pageNumber="110">......Continued on the next page</tableNote>
</emphasis>
</paragraph>
<caption id="BDAF6607FFD01F75FF19C4C664F346CE" box="[163,413,163,187]" isContinuationCaption="true" pageId="26" pageNumber="111" startId="26.[163,251,163,187]" targetBox="[163,1323,203,1516]" targetIsTable="true" targetPageId="26" targetType="table">
<paragraph id="E96F368FFFD01F75FF19C4C664F346CE" blockId="26.[163,413,163,187]" box="[163,413,163,187]" pageId="26" pageNumber="111">
<emphasis id="DBA4EA9DFFD01F75FF19C4C6647846CE" bold="true" box="[163,278,163,187]" pageId="26" pageNumber="111">TABLE 8.</emphasis>
(Continued)
</paragraph>
</caption>
<paragraph id="E96F368FFFD01F75FDFEC4AE60454399" pageId="26" pageNumber="111">
<table id="9BD0C42FFFD0E090FF19C4AE60454399" box="[163,1323,203,1516]" gridcols="6" gridrows="33" pageId="26" pageNumber="111">
<tr id="57E034CDFFD0E090FF19C4AE60454696" box="[163,1323,203,227]" gridrow="0" pageId="26" pageNumber="111" rowspan-0="1" rowspan-5="1">
<th id="14315DB1FFD0E090FDFEC4AE67BE4696" box="[580,720,203,227]" gridcol="1" gridrow="0" pageId="26" pageNumber="111">
<bibRefCitation id="8D414B7EFFD01F75FDFEC4AE67BE4696" author="Sasaki, K." box="[580,720,203,227]" pageId="26" pageNumber="111" pagination="191 - 199" refId="ref23270" refString="Sasaki, K. (1992) Two new and two resurrected species of the sciaenid genus Johnius (Johnius) from the West Pacific. Japanese Journal of Ichthyology, 39 (3), 191 - 199. https: // doi. org / 10.1007 / BF 02905476" type="journal article" year="1992">Sasaki (1992)</bibRefCitation>
</th>
<th id="14315DB1FFD0E090FD43C4AE61C14696" box="[761,1199,203,227]" colspan="3" colspanRight="2" gridcol="2" gridrow="0" pageId="26" pageNumber="111">Voucher specimen, NMMB-P 37087 (N=4)</th>
</tr>
<tr id="57E034CDFFD0E090FF19C4966045477E" box="[163,1323,243,267]" gridrow="1" pageId="26" pageNumber="111" rowspan-0="1" rowspan-1="1">
<td id="14315DB1FFD0E090FD43C496664B477E" box="[761,805,243,267]" gridcol="2" gridrow="1" pageId="26" pageNumber="111">Min</td>
<td id="14315DB1FFD0E090FC08C496668E477E" box="[946,992,243,267]" gridcol="3" gridrow="1" pageId="26" pageNumber="111">Max</td>
<td id="14315DB1FFD0E090FBDAC49661C1477E" box="[1120,1199,243,267]" gridcol="4" gridrow="1" pageId="26" pageNumber="111">Mean</td>
<td id="14315DB1FFD0E090FAB1C4966045477E" box="[1291,1323,243,267]" gridcol="5" gridrow="1" pageId="26" pageNumber="111">SD</td>
</tr>
<tr id="57E034CDFFD0E090FF19C57960454741" box="[163,1323,284,308]" gridrow="2" pageId="26" pageNumber="111">
<th id="14315DB1FFD0E090FF19C57960454741" box="[163,1323,284,308]" colspan="6" colspanRight="5" gridcol="0" gridrow="2" pageId="26" pageNumber="111">Measurements as %HL</th>
</tr>
<tr id="57E034CDFFD0E090FF19C52160454728" box="[163,1323,324,349]" gridrow="3" pageId="26" pageNumber="111">
<th id="14315DB1FFD0E090FF19C52167784728" box="[163,534,324,349]" gridcol="0" gridrow="3" pageId="26" pageNumber="111">Snout length</th>
<td id="14315DB1FFD0E090FDFEC52167BE4728" box="[580,720,324,349]" gridcol="1" gridrow="3" pageId="26" pageNumber="111">23.2</td>
<td id="14315DB1FFD0E090FD43C521664B4728" box="[761,805,324,349]" gridcol="2" gridrow="3" pageId="26" pageNumber="111">23.0</td>
<td id="14315DB1FFD0E090FC08C521668E4728" box="[946,992,324,349]" gridcol="3" gridrow="3" pageId="26" pageNumber="111">29.5</td>
<td id="14315DB1FFD0E090FBDAC52161C14728" box="[1120,1199,324,349]" gridcol="4" gridrow="3" pageId="26" pageNumber="111">26.3</td>
<td id="14315DB1FFD0E090FAB1C52160454728" box="[1291,1323,324,349]" gridcol="5" gridrow="3" pageId="26" pageNumber="111">2.0</td>
</tr>
<tr id="57E034CDFFD0E090FF19C508604547F0" box="[163,1323,365,389]" gridrow="4" pageId="26" pageNumber="111">
<th id="14315DB1FFD0E090FF19C508677847F0" box="[163,534,365,389]" gridcol="0" gridrow="4" pageId="26" pageNumber="111">Maxillary length</th>
<td id="14315DB1FFD0E090FDFEC50867BE47F0" box="[580,720,365,389]" gridcol="1" gridrow="4" pageId="26" pageNumber="111">41.3</td>
<td id="14315DB1FFD0E090FD43C508664B47F0" box="[761,805,365,389]" gridcol="2" gridrow="4" pageId="26" pageNumber="111">34.0</td>
<td id="14315DB1FFD0E090FC08C508668E47F0" box="[946,992,365,389]" gridcol="3" gridrow="4" pageId="26" pageNumber="111">42.6</td>
<td id="14315DB1FFD0E090FBDAC50861C147F0" box="[1120,1199,365,389]" gridcol="4" gridrow="4" pageId="26" pageNumber="111">38.2</td>
<td id="14315DB1FFD0E090FAB1C508604547F0" box="[1291,1323,365,389]" gridcol="5" gridrow="4" pageId="26" pageNumber="111">2.8</td>
</tr>
<tr id="57E034CDFFD0E090FF19C5F0604547D8" box="[163,1323,405,429]" gridrow="5" pageId="26" pageNumber="111">
<th id="14315DB1FFD0E090FF19C5F0677847D8" box="[163,534,405,429]" gridcol="0" gridrow="5" pageId="26" pageNumber="111">Eye diameter</th>
<td id="14315DB1FFD0E090FDFEC5F067BE47D8" box="[580,720,405,429]" gridcol="1" gridrow="5" pageId="26" pageNumber="111">21.5</td>
<td id="14315DB1FFD0E090FD43C5F0664B47D8" box="[761,805,405,429]" gridcol="2" gridrow="5" pageId="26" pageNumber="111">18.0</td>
<td id="14315DB1FFD0E090FC08C5F0668E47D8" box="[946,992,405,429]" gridcol="3" gridrow="5" pageId="26" pageNumber="111">24.4</td>
<td id="14315DB1FFD0E090FBDAC5F061C147D8" box="[1120,1199,405,429]" gridcol="4" gridrow="5" pageId="26" pageNumber="111">21.6</td>
<td id="14315DB1FFD0E090FAB1C5F0604547D8" box="[1291,1323,405,429]" gridcol="5" gridrow="5" pageId="26" pageNumber="111">2.0</td>
</tr>
<tr id="57E034CDFFD0E090FF19C5D8604547A0" box="[163,1323,445,469]" gridrow="6" pageId="26" pageNumber="111">
<th id="14315DB1FFD0E090FF19C5D8677847A0" box="[163,534,445,469]" gridcol="0" gridrow="6" pageId="26" pageNumber="111">Interorbital width</th>
<td id="14315DB1FFD0E090FDFEC5D867BE47A0" box="[580,720,445,469]" gridcol="1" gridrow="6" pageId="26" pageNumber="111">23.4</td>
<td id="14315DB1FFD0E090FD43C5D8664B47A0" box="[761,805,445,469]" gridcol="2" gridrow="6" pageId="26" pageNumber="111">23.8</td>
<td id="14315DB1FFD0E090FC08C5D8668E47A0" box="[946,992,445,469]" gridcol="3" gridrow="6" pageId="26" pageNumber="111">27.9</td>
<td id="14315DB1FFD0E090FBDAC5D861C147A0" box="[1120,1199,445,469]" gridcol="4" gridrow="6" pageId="26" pageNumber="111">26.0</td>
<td id="14315DB1FFD0E090FAB1C5D8604547A0" box="[1291,1323,445,469]" gridcol="5" gridrow="6" pageId="26" pageNumber="111">1.3</td>
</tr>
<tr id="57E034CDFFD0E090FF19C5806045478B" box="[163,1323,485,510]" gridrow="7" pageId="26" pageNumber="111">
<th id="14315DB1FFD0E090FF19C5806778478B" box="[163,534,485,510]" gridcol="0" gridrow="7" pageId="26" pageNumber="111">2nd spine length</th>
<td id="14315DB1FFD0E090FDFEC58067BE478B" box="[580,720,485,510]" gridcol="1" gridrow="7" pageId="26" pageNumber="111">31.6</td>
<td id="14315DB1FFD0E090FD43C580664B478B" box="[761,805,485,510]" gridcol="2" gridrow="7" pageId="26" pageNumber="111">24.9</td>
<td id="14315DB1FFD0E090FC08C580668E478B" box="[946,992,485,510]" gridcol="3" gridrow="7" pageId="26" pageNumber="111">35.6</td>
<td id="14315DB1FFD0E090FBDAC58061C1478B" box="[1120,1199,485,510]" gridcol="4" gridrow="7" pageId="26" pageNumber="111">29.2</td>
<td id="14315DB1FFD0E090FAB1C5806045478B" box="[1291,1323,485,510]" gridcol="5" gridrow="7" pageId="26" pageNumber="111">2.6</td>
</tr>
<tr id="57E034CDFFD0E090FF19C66860454453" box="[163,1323,525,550]" gridrow="8" pageId="26" pageNumber="111">
<th id="14315DB1FFD0E090FF19C66867784453" box="[163,534,525,550]" gridcol="0" gridrow="8" pageId="26" pageNumber="111">1st ray length</th>
<td id="14315DB1FFD0E090FDFEC66867BE4453" box="[580,720,525,550]" gridcol="1" gridrow="8" pageId="26" pageNumber="111">39.6</td>
<td id="14315DB1FFD0E090FD43C668664B4453" box="[761,805,525,550]" gridcol="2" gridrow="8" pageId="26" pageNumber="111">30.7</td>
<td id="14315DB1FFD0E090FC08C668668E4453" box="[946,992,525,550]" gridcol="3" gridrow="8" pageId="26" pageNumber="111">46.7</td>
<td id="14315DB1FFD0E090FBDAC66861C14453" box="[1120,1199,525,550]" gridcol="4" gridrow="8" pageId="26" pageNumber="111">36.3</td>
<td id="14315DB1FFD0E090FAB1C66860454453" box="[1291,1323,525,550]" gridcol="5" gridrow="8" pageId="26" pageNumber="111">3.5</td>
</tr>
<tr id="57E034CDFFD0E090FF19C6536045443B" box="[163,1323,566,590]" gridrow="9" pageId="26" pageNumber="111">
<th id="14315DB1FFD0E090FF19C6536045443B" box="[163,1323,566,590]" colspan="6" colspanRight="5" gridcol="0" gridrow="9" pageId="26" pageNumber="111">Measurements as %SL</th>
</tr>
<tr id="57E034CDFFD0E090FF19C63B60454403" box="[163,1323,606,630]" gridrow="10" pageId="26" pageNumber="111">
<th id="14315DB1FFD0E090FF19C63B67784403" box="[163,534,606,630]" gridcol="0" gridrow="10" pageId="26" pageNumber="111">Snout to anal-fin origin</th>
<td id="14315DB1FFD0E090FDFEC63B67BE4403" box="[580,720,606,630]" gridcol="1" gridrow="10" pageId="26" pageNumber="111">69.3</td>
<td id="14315DB1FFD0E090FD43C63B664B4403" box="[761,805,606,630]" gridcol="2" gridrow="10" pageId="26" pageNumber="111">67.1</td>
<td id="14315DB1FFD0E090FC08C63B668E4403" box="[946,992,606,630]" gridcol="3" gridrow="10" pageId="26" pageNumber="111">73.3</td>
<td id="14315DB1FFD0E090FBDAC63B61C14403" box="[1120,1199,606,630]" gridcol="4" gridrow="10" pageId="26" pageNumber="111">70.6</td>
<td id="14315DB1FFD0E090FAB1C63B60454403" box="[1291,1323,606,630]" gridcol="5" gridrow="10" pageId="26" pageNumber="111">1.5</td>
</tr>
<tr id="57E034CDFFD0E090FF19C6E3604544EA" box="[163,1323,646,671]" gridrow="11" pageId="26" pageNumber="111">
<th id="14315DB1FFD0E090FF19C6E3677844EA" box="[163,534,646,671]" gridcol="0" gridrow="11" pageId="26" pageNumber="111">Snout to 2nd dorsal-fin origin</th>
<td id="14315DB1FFD0E090FDFEC6E367BE44EA" box="[580,720,646,671]" gridcol="1" gridrow="11" pageId="26" pageNumber="111">53.2</td>
<td id="14315DB1FFD0E090FD43C6E3664B44EA" box="[761,805,646,671]" gridcol="2" gridrow="11" pageId="26" pageNumber="111">49.3</td>
<td id="14315DB1FFD0E090FC08C6E3668E44EA" box="[946,992,646,671]" gridcol="3" gridrow="11" pageId="26" pageNumber="111">55.0</td>
<td id="14315DB1FFD0E090FBDAC6E361C144EA" box="[1120,1199,646,671]" gridcol="4" gridrow="11" pageId="26" pageNumber="111">51.7</td>
<td id="14315DB1FFD0E090FAB1C6E3604544EA" box="[1291,1323,646,671]" gridcol="5" gridrow="11" pageId="26" pageNumber="111">1.7</td>
</tr>
<tr id="57E034CDFFD0E090FF19C6CB604544B2" box="[163,1323,686,711]" gridrow="12" pageId="26" pageNumber="111">
<th id="14315DB1FFD0E090FF19C6CB677844B2" box="[163,534,686,711]" gridcol="0" gridrow="12" pageId="26" pageNumber="111">Snout to 1st dorsal-fin origin</th>
<td id="14315DB1FFD0E090FDFEC6CB67BE44B2" box="[580,720,686,711]" gridcol="1" gridrow="12" pageId="26" pageNumber="111">33.3</td>
<td id="14315DB1FFD0E090FD43C6CB664B44B2" box="[761,805,686,711]" gridcol="2" gridrow="12" pageId="26" pageNumber="111">31.8</td>
<td id="14315DB1FFD0E090FC08C6CB668E44B2" box="[946,992,686,711]" gridcol="3" gridrow="12" pageId="26" pageNumber="111">36.2</td>
<td id="14315DB1FFD0E090FBDAC6CB61C144B2" box="[1120,1199,686,711]" gridcol="4" gridrow="12" pageId="26" pageNumber="111">33.7</td>
<td id="14315DB1FFD0E090FAB1C6CB604544B2" box="[1291,1323,686,711]" gridcol="5" gridrow="12" pageId="26" pageNumber="111">1.3</td>
</tr>
<tr id="57E034CDFFD0E090FF19C6B26045449A" box="[163,1323,727,751]" gridrow="13" pageId="26" pageNumber="111">
<th id="14315DB1FFD0E090FF19C6B26778449A" box="[163,534,727,751]" gridcol="0" gridrow="13" pageId="26" pageNumber="111">Snout to pectoral-fin insertion</th>
<td id="14315DB1FFD0E090FDFEC6B267BE449A" box="[580,720,727,751]" gridcol="1" gridrow="13" pageId="26" pageNumber="111">31.2</td>
<td id="14315DB1FFD0E090FD43C6B2664B449A" box="[761,805,727,751]" gridcol="2" gridrow="13" pageId="26" pageNumber="111">27.4</td>
<td id="14315DB1FFD0E090FC08C6B2668E449A" box="[946,992,727,751]" gridcol="3" gridrow="13" pageId="26" pageNumber="111">32.8</td>
<td id="14315DB1FFD0E090FBDAC6B261C1449A" box="[1120,1199,727,751]" gridcol="4" gridrow="13" pageId="26" pageNumber="111">30.3</td>
<td id="14315DB1FFD0E090FAB1C6B26045449A" box="[1291,1323,727,751]" gridcol="5" gridrow="13" pageId="26" pageNumber="111">1.5</td>
</tr>
<tr id="57E034CDFFD0E090FF19C69A60454562" box="[163,1323,767,791]" gridrow="14" pageId="26" pageNumber="111">
<th id="14315DB1FFD0E090FF19C69A67784562" box="[163,534,767,791]" gridcol="0" gridrow="14" pageId="26" pageNumber="111">Snout to pelvic-fin insertion</th>
<td id="14315DB1FFD0E090FDFEC69A67BE4562" box="[580,720,767,791]" gridcol="1" gridrow="14" pageId="26" pageNumber="111">35.4</td>
<td id="14315DB1FFD0E090FD43C69A664B4562" box="[761,805,767,791]" gridcol="2" gridrow="14" pageId="26" pageNumber="111">31.0</td>
<td id="14315DB1FFD0E090FC08C69A668E4562" box="[946,992,767,791]" gridcol="3" gridrow="14" pageId="26" pageNumber="111">36.5</td>
<td id="14315DB1FFD0E090FBDAC69A61C14562" box="[1120,1199,767,791]" gridcol="4" gridrow="14" pageId="26" pageNumber="111">34.0</td>
<td id="14315DB1FFD0E090FAB1C69A60454562" box="[1291,1323,767,791]" gridcol="5" gridrow="14" pageId="26" pageNumber="111">1.5</td>
</tr>
<tr id="57E034CDFFD0E090FF19C74260454535" box="[163,1323,807,832]" gridrow="15" pageId="26" pageNumber="111">
<th id="14315DB1FFD0E090FF19C74267784535" box="[163,534,807,832]" gridcol="0" gridrow="15" pageId="26" pageNumber="111">Pectoral-fin length</th>
<td id="14315DB1FFD0E090FDFEC74267BE4535" box="[580,720,807,832]" gridcol="1" gridrow="15" pageId="26" pageNumber="111">21.3</td>
<td id="14315DB1FFD0E090FD43C742664B4535" box="[761,805,807,832]" gridcol="2" gridrow="15" pageId="26" pageNumber="111">16.6</td>
<td id="14315DB1FFD0E090FC08C742668E4535" box="[946,992,807,832]" gridcol="3" gridrow="15" pageId="26" pageNumber="111">24.9</td>
<td id="14315DB1FFD0E090FBDAC74261C14535" box="[1120,1199,807,832]" gridcol="4" gridrow="15" pageId="26" pageNumber="111">19.7</td>
<td id="14315DB1FFD0E090FAB1C74260454535" box="[1291,1323,807,832]" gridcol="5" gridrow="15" pageId="26" pageNumber="111">2.3</td>
</tr>
<tr id="57E034CDFFD0E090FF19C7356045451D" box="[163,1323,848,872]" gridrow="16" pageId="26" pageNumber="111">
<th id="14315DB1FFD0E090FF19C7356778451D" box="[163,534,848,872]" gridcol="0" gridrow="16" pageId="26" pageNumber="111">Head length</th>
<td id="14315DB1FFD0E090FDFEC73567BE451D" box="[580,720,848,872]" gridcol="1" gridrow="16" pageId="26" pageNumber="111">31.0</td>
<td id="14315DB1FFD0E090FD43C735664B451D" box="[761,805,848,872]" gridcol="2" gridrow="16" pageId="26" pageNumber="111">27.7</td>
<td id="14315DB1FFD0E090FC08C735668E451D" box="[946,992,848,872]" gridcol="3" gridrow="16" pageId="26" pageNumber="111">32.3</td>
<td id="14315DB1FFD0E090FBDAC73561C1451D" box="[1120,1199,848,872]" gridcol="4" gridrow="16" pageId="26" pageNumber="111">30.2</td>
<td id="14315DB1FFD0E090FAB1C7356045451D" box="[1291,1323,848,872]" gridcol="5" gridrow="16" pageId="26" pageNumber="111">1.2</td>
</tr>
<tr id="57E034CDFFD0E090FF19C71D604545E5" box="[163,1323,888,912]" gridrow="17" pageId="26" pageNumber="111">
<th id="14315DB1FFD0E090FF19C71D677845E5" box="[163,534,888,912]" gridcol="0" gridrow="17" pageId="26" pageNumber="111">Pelvic-fin length</th>
<td id="14315DB1FFD0E090FDFEC71D67BE45E5" box="[580,720,888,912]" gridcol="1" gridrow="17" pageId="26" pageNumber="111">15.3</td>
<td id="14315DB1FFD0E090FD43C71D664B45E5" box="[761,805,888,912]" gridcol="2" gridrow="17" pageId="26" pageNumber="111">12.7</td>
<td id="14315DB1FFD0E090FC08C71D668E45E5" box="[946,992,888,912]" gridcol="3" gridrow="17" pageId="26" pageNumber="111">20.0</td>
<td id="14315DB1FFD0E090FBDAC71D61C145E5" box="[1120,1199,888,912]" gridcol="4" gridrow="17" pageId="26" pageNumber="111">16.1</td>
<td id="14315DB1FFD0E090FAB1C71D604545E5" box="[1291,1323,888,912]" gridcol="5" gridrow="17" pageId="26" pageNumber="111">2.0</td>
</tr>
<tr id="57E034CDFFD0E090FF19C7C5604545CD" box="[163,1323,928,952]" gridrow="18" pageId="26" pageNumber="111">
<th id="14315DB1FFD0E090FF19C7C5677845CD" box="[163,534,928,952]" gridcol="0" gridrow="18" pageId="26" pageNumber="111">Body depth (D1-P2)</th>
<td id="14315DB1FFD0E090FDFEC7C567BE45CD" box="[580,720,928,952]" gridcol="1" gridrow="18" pageId="26" pageNumber="111">27.0</td>
<td id="14315DB1FFD0E090FD43C7C5664B45CD" box="[761,805,928,952]" gridcol="2" gridrow="18" pageId="26" pageNumber="111">25.2</td>
<td id="14315DB1FFD0E090FC08C7C5668E45CD" box="[946,992,928,952]" gridcol="3" gridrow="18" pageId="26" pageNumber="111">29.0</td>
<td id="14315DB1FFD0E090FBDAC7C561C145CD" box="[1120,1199,928,952]" gridcol="4" gridrow="18" pageId="26" pageNumber="111">27.1</td>
<td id="14315DB1FFD0E090FAB1C7C5604545CD" box="[1291,1323,928,952]" gridcol="5" gridrow="18" pageId="26" pageNumber="111">1.2</td>
</tr>
<tr id="57E034CDFFD0E090FF19C7AD60454595" box="[163,1323,968,992]" gridrow="19" pageId="26" pageNumber="111">
<th id="14315DB1FFD0E090FF19C7AD67784595" box="[163,534,968,992]" gridcol="0" gridrow="19" pageId="26" pageNumber="111">Body width (P1-P1)</th>
<td id="14315DB1FFD0E090FDFEC7AD67BE4595" box="[580,720,968,992]" gridcol="1" gridrow="19" pageId="26" pageNumber="111">15.6</td>
<td id="14315DB1FFD0E090FD43C7AD664B4595" box="[761,805,968,992]" gridcol="2" gridrow="19" pageId="26" pageNumber="111">13.4</td>
<td id="14315DB1FFD0E090FC08C7AD668E4595" box="[946,992,968,992]" gridcol="3" gridrow="19" pageId="26" pageNumber="111">18.5</td>
<td id="14315DB1FFD0E090FBDAC7AD61C14595" box="[1120,1199,968,992]" gridcol="4" gridrow="19" pageId="26" pageNumber="111">15.5</td>
<td id="14315DB1FFD0E090FAB1C7AD60454595" box="[1291,1323,968,992]" gridcol="5" gridrow="19" pageId="26" pageNumber="111">1.8</td>
</tr>
<tr id="57E034CDFFD0E090FF19C7956045427C" box="[163,1323,1008,1033]" gridrow="20" pageId="26" pageNumber="111">
<th id="14315DB1FFD0E090FF19C7956778427C" box="[163,534,1008,1033]" gridcol="0" gridrow="20" pageId="26" pageNumber="111">1st dorsal-fin base length</th>
<td id="14315DB1FFD0E090FDFEC79567BE427C" box="[580,720,1008,1033]" gridcol="1" gridrow="20" pageId="26" pageNumber="111">20.1</td>
<td id="14315DB1FFD0E090FD43C795664B427C" box="[761,805,1008,1033]" gridcol="2" gridrow="20" pageId="26" pageNumber="111">15.9</td>
<td id="14315DB1FFD0E090FC08C795668E427C" box="[946,992,1008,1033]" gridcol="3" gridrow="20" pageId="26" pageNumber="111">23.1</td>
<td id="14315DB1FFD0E090FBDAC79561C1427C" box="[1120,1199,1008,1033]" gridcol="4" gridrow="20" pageId="26" pageNumber="111">19.3</td>
<td id="14315DB1FFD0E090FAB1C7956045427C" box="[1291,1323,1008,1033]" gridcol="5" gridrow="20" pageId="26" pageNumber="111">2.0</td>
</tr>
<tr id="57E034CDFFD0E090FF19C07C60454244" box="[163,1323,1049,1073]" gridrow="21" pageId="26" pageNumber="111">
<th id="14315DB1FFD0E090FF19C07C67784244" box="[163,534,1049,1073]" gridcol="0" gridrow="21" pageId="26" pageNumber="111">longest spine length</th>
<td id="14315DB1FFD0E090FDFEC07C67BE4244" box="[580,720,1049,1073]" gridcol="1" gridrow="21" pageId="26" pageNumber="111">14.9</td>
<td id="14315DB1FFD0E090FD43C07C664B4244" box="[761,805,1049,1073]" gridcol="2" gridrow="21" pageId="26" pageNumber="111">11.9</td>
<td id="14315DB1FFD0E090FC08C07C668E4244" box="[946,992,1049,1073]" gridcol="3" gridrow="21" pageId="26" pageNumber="111">16.7</td>
<td id="14315DB1FFD0E090FBDAC07C61C14244" box="[1120,1199,1049,1073]" gridcol="4" gridrow="21" pageId="26" pageNumber="111">13.8</td>
<td id="14315DB1FFD0E090FAB1C07C60454244" box="[1291,1323,1049,1073]" gridcol="5" gridrow="21" pageId="26" pageNumber="111">1.3</td>
</tr>
<tr id="57E034CDFFD0E090FF19C0246045422C" box="[163,1323,1089,1113]" gridrow="22" pageId="26" pageNumber="111">
<th id="14315DB1FFD0E090FF19C0246778422C" box="[163,534,1089,1113]" gridcol="0" gridrow="22" pageId="26" pageNumber="111">2nd dorsal-fin base length</th>
<td id="14315DB1FFD0E090FDFEC02467BE422C" box="[580,720,1089,1113]" gridcol="1" gridrow="22" pageId="26" pageNumber="111">40.1</td>
<td id="14315DB1FFD0E090FD43C024664B422C" box="[761,805,1089,1113]" gridcol="2" gridrow="22" pageId="26" pageNumber="111">35.8</td>
<td id="14315DB1FFD0E090FC08C024668E422C" box="[946,992,1089,1113]" gridcol="3" gridrow="22" pageId="26" pageNumber="111">44.8</td>
<td id="14315DB1FFD0E090FBDAC02461C1422C" box="[1120,1199,1089,1113]" gridcol="4" gridrow="22" pageId="26" pageNumber="111">39.7</td>
<td id="14315DB1FFD0E090FAB1C0246045422C" box="[1291,1323,1089,1113]" gridcol="5" gridrow="22" pageId="26" pageNumber="111">2.5</td>
</tr>
<tr id="57E034CDFFD0E090FF19C00C604542F7" box="[163,1323,1129,1154]" gridrow="23" pageId="26" pageNumber="111">
<th id="14315DB1FFD0E090FF19C00C677842F7" box="[163,534,1129,1154]" gridcol="0" gridrow="23" pageId="26" pageNumber="111">longest ray length</th>
<td id="14315DB1FFD0E090FDFEC00C67BE42F7" box="[580,720,1129,1154]" gridcol="1" gridrow="23" pageId="26" pageNumber="111">8.9</td>
<td id="14315DB1FFD0E090FD43C00C664B42F7" box="[761,805,1129,1154]" gridcol="2" gridrow="23" pageId="26" pageNumber="111">7.0</td>
<td id="14315DB1FFD0E090FC08C00C668E42F7" box="[946,992,1129,1154]" gridcol="3" gridrow="23" pageId="26" pageNumber="111">14.4</td>
<td id="14315DB1FFD0E090FBDAC00C61C142F7" box="[1120,1199,1129,1154]" gridcol="4" gridrow="23" pageId="26" pageNumber="111">9.8</td>
<td id="14315DB1FFD0E090FAB1C00C604542F7" box="[1291,1323,1129,1154]" gridcol="5" gridrow="23" pageId="26" pageNumber="111">1.7</td>
</tr>
<tr id="57E034CDFFD0E090FF19C0F4604542DF" box="[163,1323,1169,1194]" gridrow="24" pageId="26" pageNumber="111">
<th id="14315DB1FFD0E090FF19C0F4677842DF" box="[163,534,1169,1194]" gridcol="0" gridrow="24" pageId="26" pageNumber="111">Anal-fin base length</th>
<td id="14315DB1FFD0E090FDFEC0F467BE42DF" box="[580,720,1169,1194]" gridcol="1" gridrow="24" pageId="26" pageNumber="111">9.5</td>
<td id="14315DB1FFD0E090FD43C0F4664B42DF" box="[761,805,1169,1194]" gridcol="2" gridrow="24" pageId="26" pageNumber="111">6.8</td>
<td id="14315DB1FFD0E090FC08C0F4668E42DF" box="[946,992,1169,1194]" gridcol="3" gridrow="24" pageId="26" pageNumber="111">10.2</td>
<td id="14315DB1FFD0E090FBDAC0F461C142DF" box="[1120,1199,1169,1194]" gridcol="4" gridrow="24" pageId="26" pageNumber="111">8.8</td>
<td id="14315DB1FFD0E090FAB1C0F4604542DF" box="[1291,1323,1169,1194]" gridcol="5" gridrow="24" pageId="26" pageNumber="111">0.9</td>
</tr>
<tr id="57E034CDFFD0E090FF19C0DF604542A7" box="[163,1323,1210,1234]" gridrow="25" pageId="26" pageNumber="111">
<th id="14315DB1FFD0E090FF19C0DF677842A7" box="[163,534,1210,1234]" gridcol="0" gridrow="25" pageId="26" pageNumber="111">2nd spine length</th>
<td id="14315DB1FFD0E090FDFEC0DF67BE42A7" box="[580,720,1210,1234]" gridcol="1" gridrow="25" pageId="26" pageNumber="111">9.8</td>
<td id="14315DB1FFD0E090FD43C0DF664B42A7" box="[761,805,1210,1234]" gridcol="2" gridrow="25" pageId="26" pageNumber="111">7.4</td>
<td id="14315DB1FFD0E090FC08C0DF668E42A7" box="[946,992,1210,1234]" gridcol="3" gridrow="25" pageId="26" pageNumber="111">10.2</td>
<td id="14315DB1FFD0E090FBDAC0DF61C142A7" box="[1120,1199,1210,1234]" gridcol="4" gridrow="25" pageId="26" pageNumber="111">8.5</td>
<td id="14315DB1FFD0E090FAB1C0DF604542A7" box="[1291,1323,1210,1234]" gridcol="5" gridrow="25" pageId="26" pageNumber="111">0.8</td>
</tr>
<tr id="57E034CDFFD0E090FF19C0846045428E" box="[163,1323,1249,1275]" gridrow="26" pageId="26" pageNumber="111">
<th id="14315DB1FFD0E090FF19C0846778428E" box="[163,534,1249,1275]" gridcol="0" gridrow="26" pageId="26" pageNumber="111">1st ray length</th>
<td id="14315DB1FFD0E090FDFEC08467BE428E" box="[580,720,1249,1275]" gridcol="1" gridrow="26" pageId="26" pageNumber="111">12.3</td>
<td id="14315DB1FFD0E090FD43C084664B428E" box="[761,805,1249,1275]" gridcol="2" gridrow="26" pageId="26" pageNumber="111">9.0</td>
<td id="14315DB1FFD0E090FC08C084668E428E" box="[946,992,1249,1275]" gridcol="3" gridrow="26" pageId="26" pageNumber="111">13.3</td>
<td id="14315DB1FFD0E090FBDAC08461C1428E" box="[1120,1199,1249,1275]" gridcol="4" gridrow="26" pageId="26" pageNumber="111">10.9</td>
<td id="14315DB1FFD0E090FAB1C0846045428E" box="[1291,1323,1249,1275]" gridcol="5" gridrow="26" pageId="26" pageNumber="111">1.0</td>
</tr>
<tr id="57E034CDFFD0E090FF19C16F60454357" box="[163,1323,1290,1314]" gridrow="27" pageId="26" pageNumber="111">
<th id="14315DB1FFD0E090FF19C16F67784357" box="[163,534,1290,1314]" gridcol="0" gridrow="27" pageId="26" pageNumber="111">Caudal-peduncle depth</th>
<td id="14315DB1FFD0E090FDFEC16F67BE4357" box="[580,720,1290,1314]" gridcol="1" gridrow="27" pageId="26" pageNumber="111">10.6</td>
<td id="14315DB1FFD0E090FD43C16F664B4357" box="[761,805,1290,1314]" gridcol="2" gridrow="27" pageId="26" pageNumber="111">8.7</td>
<td id="14315DB1FFD0E090FC08C16F668E4357" box="[946,992,1290,1314]" gridcol="3" gridrow="27" pageId="26" pageNumber="111">10.7</td>
<td id="14315DB1FFD0E090FBDAC16F61C14357" box="[1120,1199,1290,1314]" gridcol="4" gridrow="27" pageId="26" pageNumber="111">9.5</td>
<td id="14315DB1FFD0E090FAB1C16F60454357" box="[1291,1323,1290,1314]" gridcol="5" gridrow="27" pageId="26" pageNumber="111">0.4</td>
</tr>
<tr id="57E034CDFFD0E090FF19C1576045433E" box="[163,1323,1330,1355]" gridrow="28" pageId="26" pageNumber="111">
<th id="14315DB1FFD0E090FF19C1576778433E" box="[163,534,1330,1355]" gridcol="0" gridrow="28" pageId="26" pageNumber="111">Pelvic-fin tip to vent</th>
<td id="14315DB1FFD0E090FDFEC15767BE433E" box="[580,720,1330,1355]" gridcol="1" gridrow="28" pageId="26" pageNumber="111">12.7</td>
<td id="14315DB1FFD0E090FD43C157664B433E" box="[761,805,1330,1355]" gridcol="2" gridrow="28" pageId="26" pageNumber="111">13.8</td>
<td id="14315DB1FFD0E090FC08C157668E433E" box="[946,992,1330,1355]" gridcol="3" gridrow="28" pageId="26" pageNumber="111">22.8</td>
<td id="14315DB1FFD0E090FBDAC15761C1433E" box="[1120,1199,1330,1355]" gridcol="4" gridrow="28" pageId="26" pageNumber="111">18.4</td>
<td id="14315DB1FFD0E090FAB1C1576045433E" box="[1291,1323,1330,1355]" gridcol="5" gridrow="28" pageId="26" pageNumber="111">2.7</td>
</tr>
<tr id="57E034CDFFD0E090FF19C13E60454306" box="[163,1323,1371,1395]" gridrow="29" pageId="26" pageNumber="111">
<th id="14315DB1FFD0E090FF19C13E67784306" box="[163,534,1371,1395]" gridcol="0" gridrow="29" pageId="26" pageNumber="111">Vent to anal-fin origin</th>
<td id="14315DB1FFD0E090FDFEC13E67BE4306" box="[580,720,1371,1395]" gridcol="1" gridrow="29" pageId="26" pageNumber="111">4.4</td>
<td id="14315DB1FFD0E090FD43C13E664B4306" box="[761,805,1371,1395]" gridcol="2" gridrow="29" pageId="26" pageNumber="111">3.2</td>
<td id="14315DB1FFD0E090FC08C13E668E4306" box="[946,992,1371,1395]" gridcol="3" gridrow="29" pageId="26" pageNumber="111">7.9</td>
<td id="14315DB1FFD0E090FBDAC13E61C14306" box="[1120,1199,1371,1395]" gridcol="4" gridrow="29" pageId="26" pageNumber="111">5.7</td>
<td id="14315DB1FFD0E090FAB1C13E60454306" box="[1291,1323,1371,1395]" gridcol="5" gridrow="29" pageId="26" pageNumber="111">1.2</td>
</tr>
<tr id="57E034CDFFD0E090FF19C1E6604543EE" box="[163,1323,1411,1435]" gridrow="30" pageId="26" pageNumber="111" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="14315DB1FFD0E090FF19C1E6677843EE" box="[163,534,1411,1435]" gridcol="0" gridrow="30" pageId="26" pageNumber="111">Measurements as %ED</th>
</tr>
<tr id="57E034CDFFD0E090FF19C1CF604543B1" box="[163,1323,1450,1476]" gridrow="31" pageId="26" pageNumber="111">
<th id="14315DB1FFD0E090FF19C1CF677843B1" box="[163,534,1450,1476]" gridcol="0" gridrow="31" pageId="26" pageNumber="111">Longest gill raker on 1st arch</th>
<td id="14315DB1FFD0E090FDFEC1CF67BE43B1" box="[580,720,1450,1476]" gridcol="1" gridrow="31" pageId="26" pageNumber="111">14.9</td>
<td id="14315DB1FFD0E090FD43C1CF664B43B1" box="[761,805,1450,1476]" gridcol="2" gridrow="31" pageId="26" pageNumber="111">11.3</td>
<td id="14315DB1FFD0E090FC08C1CF668E43B1" box="[946,992,1450,1476]" gridcol="3" gridrow="31" pageId="26" pageNumber="111">19.9</td>
<td id="14315DB1FFD0E090FBDAC1CF61C143B1" box="[1120,1199,1450,1476]" gridcol="4" gridrow="31" pageId="26" pageNumber="111">14.6</td>
<td id="14315DB1FFD0E090FAB1C1CF604543B1" box="[1291,1323,1450,1476]" gridcol="5" gridrow="31" pageId="26" pageNumber="111">2.4</td>
</tr>
<tr id="57E034CDFFD0E090FF19C1B660454399" box="[163,1323,1491,1516]" gridrow="32" pageId="26" pageNumber="111">
<th id="14315DB1FFD0E090FF19C1B667784399" box="[163,534,1491,1516]" gridcol="0" gridrow="32" pageId="26" pageNumber="111">Longest gill filament on 1st arch</th>
<td id="14315DB1FFD0E090FDFEC1B667BE4399" box="[580,720,1491,1516]" gridcol="1" gridrow="32" pageId="26" pageNumber="111">55.7</td>
<td id="14315DB1FFD0E090FD43C1B6664B4399" box="[761,805,1491,1516]" gridcol="2" gridrow="32" pageId="26" pageNumber="111">37.7</td>
<td id="14315DB1FFD0E090FC08C1B6668E4399" box="[946,992,1491,1516]" gridcol="3" gridrow="32" pageId="26" pageNumber="111">59.9</td>
<td id="14315DB1FFD0E090FBDAC1B661C14399" box="[1120,1199,1491,1516]" gridcol="4" gridrow="32" pageId="26" pageNumber="111">51.9</td>
<td id="14315DB1FFD0E090FAB1C1B660454399" box="[1291,1323,1491,1516]" gridcol="5" gridrow="32" pageId="26" pageNumber="111">3.4</td>
</tr>
</table>
</paragraph>
<caption id="BDAF6607FFD01F75FF22C22464FF4008" ID-Table-UUID="BDAF6607FFD01F75FF22C22464FF4008" httpUri="http://table.plazi.org/id/BDAF6607FFD01F75FF22C22464FF4008" pageId="26" pageNumber="111" startId="26.[152,244,1601,1625]" targetBox="[163,1313,1676,1942]" targetIsTable="true" targetPageId="26" targetType="table">
<paragraph id="E96F368FFFD01F75FF22C22464FF4008" blockId="26.[151,1437,1601,1661]" pageId="26" pageNumber="111">
<emphasis id="DBA4EA9DFFD01F75FF22C2246463402C" bold="true" box="[152,269,1601,1625]" pageId="26" pageNumber="111">TABLE 9.</emphasis>
Genetic distances for the COI gene among congener species with
<taxonomicName id="2ED04D0CFFD01F75FC06C22461E5402C" authorityName="Sasaki" authorityYear="1992" box="[956,1163,1601,1625]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="26" pageNumber="111" phylum="Chordata" rank="species" species="trewavasae">
<emphasis id="DBA4EA9DFFD01F75FC06C22461E5402C" box="[956,1163,1601,1625]" italics="true" pageId="26" pageNumber="111">Johnius trewavasae</emphasis>
</taxonomicName>
species and
<taxonomicName id="2ED04D0CFFD01F75FAB4C22465884008" baseAuthorityName="Cuvier" baseAuthorityYear="1829" family="Sciaenidae" genus="Dendrophysa" kingdom="Animalia" order="Perciformes" pageId="26" pageNumber="111" phylum="Chordata" rank="species" species="russelii">
<emphasis id="DBA4EA9DFFD01F75FAB4C22465884008" italics="true" pageId="26" pageNumber="111">Dendrophysa russelii</emphasis>
</taxonomicName>
as an outgroup.
</paragraph>
</caption>
<paragraph id="E96F368FFFD01F75FF19C2E961C841E3" pageId="26" pageNumber="111">
<table id="9BD0C42FFFD0E090FF19C2E9604F41E3" box="[163,1313,1676,1942]" gridcols="7" gridrows="7" pageId="26" pageNumber="111">
<tr id="57E034CDFFD0E090FF19C2E9604F40D0" box="[163,1313,1676,1701]" gridrow="0" pageId="26" pageNumber="111">
<th id="14315DB1FFD0E090FF19C2E964D440D0" box="[163,442,1676,1701]" gridcol="0" gridrow="0" pageId="26" pageNumber="111">
<emphasis id="DBA4EA9DFFD01F75FF19C2E9659C40D1" bold="true" box="[163,242,1676,1700]" pageId="26" pageNumber="111">Species</emphasis>
</th>
<th id="14315DB1FFD0E090FE58C2E9676040D0" box="[482,526,1676,1701]" gridcol="1" gridrow="0" pageId="26" pageNumber="111">
<emphasis id="DBA4EA9DFFD01F75FE58C2E8648140D0" bold="true" box="[482,495,1677,1701]" pageId="26" pageNumber="111">1</emphasis>
</th>
<th id="14315DB1FFD0E090FD31C2E967D940D0" box="[651,695,1676,1701]" gridcol="2" gridrow="0" pageId="26" pageNumber="111">
<emphasis id="DBA4EA9DFFD01F75FD31C2E867F640D0" bold="true" box="[651,664,1677,1701]" pageId="26" pageNumber="111">2</emphasis>
</th>
<th id="14315DB1FFD0E090FC8AC2E9663240D0" box="[816,860,1676,1701]" gridcol="3" gridrow="0" pageId="26" pageNumber="111">
<emphasis id="DBA4EA9DFFD01F75FC8AC2E8665340D0" bold="true" box="[816,829,1677,1701]" pageId="26" pageNumber="111">3</emphasis>
</th>
<th id="14315DB1FFD0E090FC6FC2E9616F40D0" box="[981,1025,1676,1701]" gridcol="4" gridrow="0" pageId="26" pageNumber="111">
<emphasis id="DBA4EA9DFFD01F75FC6FC2E8668C40D0" bold="true" box="[981,994,1677,1701]" pageId="26" pageNumber="111">4</emphasis>
</th>
<th id="14315DB1FFD0E090FBC0C2E961C840D0" box="[1146,1190,1676,1701]" gridcol="5" gridrow="0" pageId="26" pageNumber="111">
<emphasis id="DBA4EA9DFFD01F75FBC0C2E861E940D0" bold="true" box="[1146,1159,1677,1701]" pageId="26" pageNumber="111">5</emphasis>
</th>
<th id="14315DB1FFD0E090FAAEC2E9604F40D0" box="[1300,1313,1676,1701]" gridcol="6" gridrow="0" pageId="26" pageNumber="111">
<emphasis id="DBA4EA9DFFD01F75FAAEC2E8604F40D0" bold="true" box="[1300,1313,1677,1701]" pageId="26" pageNumber="111">6</emphasis>
</th>
</tr>
<tr id="57E034CDFFD0E090FF19C2D1604F40B9" box="[163,1313,1716,1740]" gridrow="1" pageId="26" pageNumber="111">
<th id="14315DB1FFD0E090FF19C2D1604F40B9" box="[163,1313,1716,1740]" colspan="7" colspanRight="6" gridcol="0" gridrow="1" pageId="26" pageNumber="111">
<emphasis id="DBA4EA9DFFD01F75FF19C2D164E140B9" bold="true" box="[163,399,1716,1740]" italics="true" pageId="26" pageNumber="111">
1.
<taxonomicName id="2ED04D0CFFD01F75FF06C2D164E140B9" authorityName="Bleeker" authorityYear="1873" box="[188,399,1716,1740]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="26" pageNumber="111" phylum="Chordata" rank="species" species="heterolepis">Johnius_heterolepis</taxonomicName>
</emphasis>
</th>
</tr>
<tr id="57E034CDFFD0E090FF19C2B9604F4080" box="[163,1313,1756,1781]" gridrow="2" pageId="26" pageNumber="111" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th id="14315DB1FFD0E090FF19C2B964D44080" box="[163,442,1756,1781]" gridcol="0" gridrow="2" pageId="26" pageNumber="111">
<emphasis id="DBA4EA9DFFD01F75FF19C2B8641A4081" bold="true" box="[163,372,1756,1781]" italics="true" pageId="26" pageNumber="111">
2.
<taxonomicName id="2ED04D0CFFD01F75FF06C2B9641A4081" baseAuthorityName="Cuvier" baseAuthorityYear="1830" box="[188,372,1756,1780]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="26" pageNumber="111" phylum="Chordata" rank="species" species="carouna">Johnius_carouna</taxonomicName>
</emphasis>
</th>
<td id="14315DB1FFD0E090FE58C2B967604080" box="[482,526,1756,1781]" gridcol="1" gridrow="2" pageId="26" pageNumber="111">0.13</td>
</tr>
<tr id="57E034CDFFD0E090FF19C360604F4168" box="[163,1313,1797,1821]" gridrow="3" pageId="26" pageNumber="111" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th id="14315DB1FFD0E090FF19C36064D44168" box="[163,442,1797,1821]" gridcol="0" gridrow="3" pageId="26" pageNumber="111">
<emphasis id="DBA4EA9DFFD01F75FF19C36064FE4168" bold="true" box="[163,400,1797,1821]" italics="true" pageId="26" pageNumber="111">
3.
<taxonomicName id="2ED04D0CFFD01F75FF06C36064FE4168" authorityName="Sasaki" authorityYear="1992" box="[188,400,1797,1821]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="26" pageNumber="111" phylum="Chordata" rank="species" species="trewavasae">Johnius_trewavasae</taxonomicName>
</emphasis>
</th>
<td id="14315DB1FFD0E090FE58C36067604168" box="[482,526,1797,1821]" gridcol="1" gridrow="3" pageId="26" pageNumber="111">0.15</td>
<td id="14315DB1FFD0E090FD31C36067D94168" box="[651,695,1797,1821]" gridcol="2" gridrow="3" pageId="26" pageNumber="111">0.16</td>
</tr>
<tr id="57E034CDFFD0E090FF19C348604F4133" box="[163,1313,1837,1862]" gridrow="4" pageId="26" pageNumber="111" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th id="14315DB1FFD0E090FF19C34864D44133" box="[163,442,1837,1862]" gridcol="0" gridrow="4" pageId="26" pageNumber="111">
<emphasis id="DBA4EA9DFFD01F75FF19C348640B4130" bold="true" box="[163,357,1837,1861]" italics="true" pageId="26" pageNumber="111">
4.
<taxonomicName id="2ED04D0CFFD01F75FF06C348640B4130" box="[188,357,1837,1861]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="26" pageNumber="111" phylum="Chordata" rank="species" species="sasakii">Johnius_sasakii</taxonomicName>
</emphasis>
</th>
<td id="14315DB1FFD0E090FE58C34867604133" box="[482,526,1837,1862]" gridcol="1" gridrow="4" pageId="26" pageNumber="111">0.15</td>
<td id="14315DB1FFD0E090FD31C34867D94133" box="[651,695,1837,1862]" gridcol="2" gridrow="4" pageId="26" pageNumber="111">0.17</td>
<td id="14315DB1FFD0E090FC8AC34866324133" box="[816,860,1837,1862]" gridcol="3" gridrow="4" pageId="26" pageNumber="111">0.16</td>
</tr>
<tr id="57E034CDFFD0E090FF19C330604F411B" box="[163,1313,1877,1902]" gridrow="5" pageId="26" pageNumber="111" rowspan-5="1" rowspan-6="1">
<th id="14315DB1FFD0E090FF19C33064D4411B" box="[163,442,1877,1902]" gridcol="0" gridrow="5" pageId="26" pageNumber="111">
<emphasis id="DBA4EA9DFFD01F75FF19C33364D44118" bold="true" box="[163,442,1877,1901]" italics="true" pageId="26" pageNumber="111">
5.
<taxonomicName id="2ED04D0CFFD01F75FF06C33064D44118" baseAuthorityName="Bleeker" baseAuthorityYear="1855" box="[188,442,1877,1901]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="26" pageNumber="111" phylum="Chordata" rank="species" species="amblycephalus">Johnius_amblycephalus</taxonomicName>
</emphasis>
</th>
<td id="14315DB1FFD0E090FE58C3306760411B" box="[482,526,1877,1902]" gridcol="1" gridrow="5" pageId="26" pageNumber="111">0.33</td>
<td id="14315DB1FFD0E090FD31C33067D9411B" box="[651,695,1877,1902]" gridcol="2" gridrow="5" pageId="26" pageNumber="111">0.34</td>
<td id="14315DB1FFD0E090FC8AC3306632411B" box="[816,860,1877,1902]" gridcol="3" gridrow="5" pageId="26" pageNumber="111">0.36</td>
<td id="14315DB1FFD0E090FC6FC330616F411B" box="[981,1025,1877,1902]" gridcol="4" gridrow="5" pageId="26" pageNumber="111">0.37</td>
</tr>
<tr id="57E034CDFFD0E090FF19C318604F41E3" box="[163,1313,1917,1942]" gridrow="6" pageId="26" pageNumber="111" rowspan-6="1">
<th id="14315DB1FFD0E090FF19C31864D441E3" box="[163,442,1917,1942]" gridcol="0" gridrow="6" pageId="26" pageNumber="111">
<emphasis id="DBA4EA9DFFD01F75FF19C31B64CE41E0" bold="true" box="[163,416,1917,1942]" italics="true" pageId="26" pageNumber="111">
6.
<taxonomicName id="2ED04D0CFFD01F75FF06C31864CE41E0" baseAuthorityName="Cuvier" baseAuthorityYear="1829" box="[188,416,1917,1941]" family="Sciaenidae" genus="Dendrophysa" kingdom="Animalia" order="Perciformes" pageId="26" pageNumber="111" phylum="Chordata" rank="species" species="russelii">Dendrophysa_russelii</taxonomicName>
</emphasis>
</th>
<td id="14315DB1FFD0E090FE58C318676041E3" box="[482,526,1917,1942]" gridcol="1" gridrow="6" pageId="26" pageNumber="111">0.34</td>
<td id="14315DB1FFD0E090FD31C31867D941E3" box="[651,695,1917,1942]" gridcol="2" gridrow="6" pageId="26" pageNumber="111">0.37</td>
<td id="14315DB1FFD0E090FC8AC318663241E3" box="[816,860,1917,1942]" gridcol="3" gridrow="6" pageId="26" pageNumber="111">0.36</td>
<td id="14315DB1FFD0E090FC6FC318616F41E3" box="[981,1025,1917,1942]" gridcol="4" gridrow="6" pageId="26" pageNumber="111">0.34</td>
<td id="14315DB1FFD0E090FBC0C31861C841E3" box="[1146,1190,1917,1942]" gridcol="5" gridrow="6" pageId="26" pageNumber="111">0.24</td>
</tr>
</table>
</paragraph>
<caption id="BDAF6607FFD11F74FF2DC7A064A04250" ID-DOI="http://doi.org/10.5281/zenodo.14388245" ID-Zenodo-Dep="14388245" httpUri="https://zenodo.org/record/14388245/files/figure.png" pageId="27" pageNumber="112" startId="27.[151,250,965,989]" targetBox="[325,1261,188,933]" targetPageId="27" targetType="figure">
<paragraph id="E96F368FFFD11F74FF2DC7A064A04250" blockId="27.[151,1437,965,1061]" pageId="27" pageNumber="112">
<emphasis id="DBA4EA9DFFD11F74FF2DC7A0644C45A8" bold="true" box="[151,290,965,989]" pageId="27" pageNumber="112">FIGURE 17.</emphasis>
Neighbor joining tree of COI gene was reconstructed with sequences of
<taxonomicName id="2ED04D0CFFD11F74FBA4C7A0606345A8" baseAuthorityName="Bleeker" baseAuthorityYear="1855" box="[1054,1293,965,989]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="27" pageNumber="112" phylum="Chordata" rank="species" species="amblycephalus">
<emphasis id="DBA4EA9DFFD11F74FBA4C7A0606345A8" box="[1054,1293,965,989]" italics="true" pageId="27" pageNumber="112">Johnius amblycephalus</emphasis>
</taxonomicName>
,
<emphasis id="DBA4EA9DFFD11F74FAA6C7A364EF4274" italics="true" pageId="27" pageNumber="112">
<taxonomicName id="2ED04D0CFFD11F74FAA6C7A3601145A8" box="[1308,1407,965,989]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="27" pageNumber="112" phylum="Chordata" rank="species" species="sasakii">J. sasakii</taxonomicName>
,
<taxonomicName id="2ED04D0CFFD11F74FA31C7A365834274" baseAuthorityName="Cuvier" baseAuthorityYear="1830" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="27" pageNumber="112" phylum="Chordata" rank="species" species="carouna">J. carouna</taxonomicName>
,
<taxonomicName id="2ED04D0CFFD11F74FF42C78F64EF4274" authorityName="Bleeker" authorityYear="1873" box="[248,385,1001,1025]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="27" pageNumber="112" phylum="Chordata" rank="species" species="heterolepis">J. heterolepis</taxonomicName>
</emphasis>
and
<taxonomicName id="2ED04D0CFFD11F74FE0EC78F67514274" authorityName="Sasaki" authorityYear="1992" box="[436,575,1002,1025]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="27" pageNumber="112" phylum="Chordata" rank="species" species="trewavasae">
<emphasis id="DBA4EA9DFFD11F74FE0EC78F67514274" box="[436,575,1002,1025]" italics="true" pageId="27" pageNumber="112">J. trewavasae</emphasis>
</taxonomicName>
. Bootstrap support values were showed at nodes. Name with asterisk represents the sample from the present study.
</paragraph>
</caption>
<paragraph id="E96F368FFFD11F74FF7DC01266F1431F" blockId="27.[151,1437,1143,1746]" pageId="27" pageNumber="112">
According to Ng
<emphasis id="DBA4EA9DFFD11F74FE36C01D64AB42E4" box="[396,453,1143,1169]" italics="true" pageId="27" pageNumber="112">et al.</emphasis>
, (2015) and
<bibRefCitation id="8D414B7EFFD11F74FDEFC012679D42E7" author="Sasaki, K." box="[597,755,1143,1170]" pageId="27" pageNumber="112" refId="ref23416" refString="Sasaki, K. (2000) A checklist of the fishes of the South China Sea. In: Randall, J. E. &amp; Lim, K. K. P. (Eds.), Raffles Bulletin of Zoology Supplement, 8, pp. 621." type="journal volume" year="2000">Sasaki (2000)</bibRefCitation>
, both references mentioned
<taxonomicName id="2ED04D0CFFD11F74FB8AC01D61A442E4" authorityName="Sasaki" authorityYear="1992" box="[1072,1226,1144,1169]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="27" pageNumber="112" phylum="Chordata" rank="species" species="trewavasae">
<emphasis id="DBA4EA9DFFD11F74FB8AC01D61A442E4" box="[1072,1226,1144,1169]" italics="true" pageId="27" pageNumber="112">J. trewavasae</emphasis>
</taxonomicName>
has been recorded occurring in
<collectingCountry id="91C7761FFFD11F74FE99C0FE64E442C0" box="[291,394,1179,1205]" name="Malaysia" pageId="27" pageNumber="112">Malaysia</collectingCountry>
waters. Moreover, this species was synonym and frequently misapplied name as
<taxonomicName id="2ED04D0CFFD11F74FAB2C0F960F242C0" authorityName="Bleeker" authorityYear="1873" box="[1288,1436,1179,1205]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="27" pageNumber="112" phylum="Chordata" rank="species" species="heterolepis">
<emphasis id="DBA4EA9DFFD11F74FAB2C0F960F242C0" box="[1288,1436,1179,1205]" italics="true" pageId="27" pageNumber="112">J. heterolepis</emphasis>
</taxonomicName>
and
<taxonomicName id="2ED04D0CFFD11F74FF7DC0A5641D42AC" baseAuthorityName="Lal Mohan" baseAuthorityYear="1976" box="[199,371,1215,1241]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="27" pageNumber="112" phylum="Chordata" rank="species" species="macrorhynus">
<emphasis id="DBA4EA9DFFD11F74FF7DC0A5641D42AC" box="[199,371,1215,1241]" italics="true" pageId="27" pageNumber="112">J. macrorhynus</emphasis>
</taxonomicName>
in Malaysian waters. Indicates from the
<figureCitation id="71EB2A0AFFD11F74FC84C0DA66E042AC" box="[830,910,1215,1241]" captionStart="FIGURE 16" captionStartId="24.[151,250,1650,1674]" captionTargetBox="[337,1259,964,1626]" captionTargetId="figure-54@24.[328,1259,951,1626]" captionTargetPageId="24" captionText="FIGURE 16. New record of site of collection samples of Johnius trewavasae in Sarikei (SRK), Sarawak, East Malaysia." figureDoi="http://doi.org/10.5281/zenodo.14388243" httpUri="https://zenodo.org/record/14388243/files/figure.png" pageId="27" pageNumber="112">Fig. 16</figureCitation>
shown the new extension of record sites in the East
<collectingCountry id="91C7761FFFD11F74FF6AC08664594288" box="[208,311,1251,1277]" name="Malaysia" pageId="27" pageNumber="112">Malaysia</collectingCountry>
waters, concisely at the Sarikei,
<collectingCountry id="91C7761FFFD11F74FD09C086667D4288" box="[691,787,1251,1277]" name="Malaysia" pageId="27" pageNumber="112">Sarawak</collectingCountry>
states. According to Ng
<emphasis id="DBA4EA9DFFD11F74FB8BC08161054288" box="[1073,1131,1251,1277]" italics="true" pageId="27" pageNumber="112">et al.</emphasis>
, (2015) and
<bibRefCitation id="8D414B7EFFD11F74FABAC08660F2428B" author="Sasaki, K." box="[1280,1436,1251,1278]" pageId="27" pageNumber="112" pagination="3117 - 3174" refId="ref23468" refString="Sasaki, K. (2001) Sciaenidae. In: Carpenter, K. E. &amp; Niem, V. H. FAO Species Identification Guide for Fishery Purposes. The Living Marine Resources of the Western Central Pacific. Vol. 5. Bony Fishes. Part 3. Menidae to Pomacentridae. FAO, Rome, pp. 3117 - 3174." type="book chapter" year="2001">Sasaki (2001)</bibRefCitation>
previous collection discovered
<taxonomicName id="2ED04D0CFFD11F74FE40C16D67FB4354" authorityName="Sasaki" authorityYear="1992" box="[506,661,1288,1313]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="27" pageNumber="112" phylum="Chordata" rank="species" species="trewavasae">
<emphasis id="DBA4EA9DFFD11F74FE40C16D67FB4354" box="[506,661,1288,1313]" italics="true" pageId="27" pageNumber="112">J. trewavasae</emphasis>
</taxonomicName>
(LSUMZ 16645) occurring in the Eastern
<collectingRegion id="2B14F86DFFD11F74FB38C16261AE4354" box="[1154,1216,1287,1313]" country="Malaysia" name="Johor" pageId="27" pageNumber="112">Johor</collectingRegion>
Strait,
<collectingCountry id="91C7761FFFD11F74FAAFC16260124354" box="[1301,1404,1287,1313]" name="Malaysia" pageId="27" pageNumber="112">Malaysia</collectingCountry>
or
<collectingCountry id="91C7761FFFD11F74FF2DC14E64674330" box="[151,265,1323,1349]" name="Singapore" pageId="27" pageNumber="112">Singapore</collectingCountry>
waters. Thus, the extension distribution evidence discovered from this study shown several population of
<taxonomicName id="2ED04D0CFFD11F74FF2DC135645F431C" authorityName="Sasaki" authorityYear="1992" box="[151,305,1360,1385]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="27" pageNumber="112" phylum="Chordata" rank="species" species="trewavasae">
<emphasis id="DBA4EA9DFFD11F74FF2DC135645F431C" box="[151,305,1360,1385]" italics="true" pageId="27" pageNumber="112">J. trewavasae</emphasis>
</taxonomicName>
might be has been established in the Malaysian waters.
</paragraph>
<paragraph id="E96F368FFFD11F74FF7DC11160EC40A7" blockId="27.[151,1437,1143,1746]" pageId="27" pageNumber="112">
<emphasis id="DBA4EA9DFFD11F74FF7DC11164B443F8" bold="true" box="[199,474,1395,1422]" pageId="27" pageNumber="112">Molecular verification.</emphasis>
This work produced phylogenetic tree sequences utilising COI genetic markers with congener
<taxonomicName id="2ED04D0CFFD11F74FEB2C1F264D543C4" box="[264,443,1431,1457]" pageId="27" pageNumber="112">
<emphasis id="DBA4EA9DFFD11F74FEB2C1F2643143C4" box="[264,351,1431,1457]" italics="true" pageId="27" pageNumber="112">Johnius</emphasis>
species
</taxonomicName>
in East Malaysian seas and demarcated them with GenBank sequences to clarify the taxonomic status of
<taxonomicName id="2ED04D0CFFD11F74FEC3C1D9677C43A0" authorityName="Sasaki" authorityYear="1992" box="[377,530,1468,1493]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="27" pageNumber="112" phylum="Chordata" rank="species" species="trewavasae">
<emphasis id="DBA4EA9DFFD11F74FEC3C1D9677C43A0" box="[377,530,1468,1493]" italics="true" pageId="27" pageNumber="112">J. trewavasae</emphasis>
</taxonomicName>
with congener species (
<bibRefCitation id="8D414B7EFFD11F74FCA7C1D966A843A3" author="Lo, P. C. &amp; Liu, S. H. &amp; Nor, S. A. M. &amp; Chen, W. J." box="[797,966,1467,1494]" pageId="27" pageNumber="112" refId="ref22425" refString="Lo, P. C., Liu, S. H., Nor, S. A. M. &amp; Chen, W. J. (2017) Molecular exploration of hidden diversity in the Indo-West Pacific sciaenid clade. PLoS ONE, 12 (4), e 0176623. https: // doi. org / 10.1371 / journal. pone. 0176623" type="journal volume" year="2017">
Lo
<emphasis id="DBA4EA9DFFD11F74FCFEC1D9661243A0" box="[836,892,1467,1493]" italics="true" pageId="27" pageNumber="112">et al.</emphasis>
, 2017
</bibRefCitation>
), as shown in
<figureCitation id="71EB2A0AFFD11F74FBD9C1DE61DB43A3" box="[1123,1205,1467,1494]" captionStart="FIGURE 17" captionStartId="27.[151,250,965,989]" captionTargetBox="[325,1261,188,933]" captionTargetId="figure-18@27.[320,1268,181,941]" captionTargetPageId="27" captionText="FIGURE 17. Neighbor joining tree of COI gene was reconstructed with sequences of Johnius amblycephalus, J. sasakii, J. carouna, J. heterolepis and J. trewavasae. Bootstrap support values were showed at nodes. Name with asterisk represents the sample from the present study." figureDoi="http://doi.org/10.5281/zenodo.14388245" httpUri="https://zenodo.org/record/14388245/files/figure.png" pageId="27" pageNumber="112">Fig. 17</figureCitation>
.
<taxonomicName id="2ED04D0CFFD11F74FB7AC1DE60F343A0" authorityName="Sasaki" authorityYear="1992" box="[1216,1437,1467,1493]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="27" pageNumber="112" phylum="Chordata" rank="species" species="trewavasae">
<emphasis id="DBA4EA9DFFD11F74FB7AC1DE60F343A0" box="[1216,1437,1467,1493]" italics="true" pageId="27" pageNumber="112">Johnius trewavasae</emphasis>
</taxonomicName>
was a sister clade to
<taxonomicName id="2ED04D0CFFD11F74FE32C185649C438C" box="[392,498,1503,1529]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="27" pageNumber="112" phylum="Chordata" rank="species" species="sasakii">
<emphasis id="DBA4EA9DFFD11F74FE32C185649C438C" box="[392,498,1503,1529]" italics="true" pageId="27" pageNumber="112">J. sasakii</emphasis>
</taxonomicName>
and a sister group to
<taxonomicName id="2ED04D0CFFD11F74FD55C18566E9438C" authorityName="Bleeker" authorityYear="1873" box="[751,903,1503,1529]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="27" pageNumber="112" phylum="Chordata" rank="species" species="heterolepis">
<emphasis id="DBA4EA9DFFD11F74FD55C18566E9438C" box="[751,903,1503,1529]" italics="true" pageId="27" pageNumber="112">J. heterolepis</emphasis>
</taxonomicName>
and
<taxonomicName id="2ED04D0CFFD11F74FC78C1856154438C" baseAuthorityName="Cuvier" baseAuthorityYear="1830" box="[962,1082,1504,1529]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="27" pageNumber="112" phylum="Chordata" rank="species" species="carouna">
<emphasis id="DBA4EA9DFFD11F74FC78C1856154438C" box="[962,1082,1504,1529]" italics="true" pageId="27" pageNumber="112">J. carouna</emphasis>
</taxonomicName>
, according to the phylogenetic tree connection (
<figureCitation id="71EB2A0AFFD11F74FEEDC26664C3406B" box="[343,429,1539,1566]" captionStart="FIGURE 17" captionStartId="27.[151,250,965,989]" captionTargetBox="[325,1261,188,933]" captionTargetId="figure-18@27.[320,1268,181,941]" captionTargetPageId="27" captionText="FIGURE 17. Neighbor joining tree of COI gene was reconstructed with sequences of Johnius amblycephalus, J. sasakii, J. carouna, J. heterolepis and J. trewavasae. Bootstrap support values were showed at nodes. Name with asterisk represents the sample from the present study." figureDoi="http://doi.org/10.5281/zenodo.14388245" httpUri="https://zenodo.org/record/14388245/files/figure.png" pageId="27" pageNumber="112">Fig. 17</figureCitation>
). Meanwhile,
<taxonomicName id="2ED04D0CFFD11F74FDE8C266662D4068" baseAuthorityName="Cuvier" baseAuthorityYear="1829" box="[594,835,1539,1565]" family="Sciaenidae" genus="Dendrophysa" kingdom="Animalia" order="Perciformes" pageId="27" pageNumber="112" phylum="Chordata" rank="species" species="russelii">
<emphasis id="DBA4EA9DFFD11F74FDE8C266662D4068" box="[594,835,1539,1565]" italics="true" pageId="27" pageNumber="112">Dendrophysa russelii</emphasis>
</taxonomicName>
is the outgroup clade, with a strong bootstrap value confirming the connection and validation species, while
<taxonomicName id="2ED04D0CFFD11F74FCAAC24D66BC4034" baseAuthorityName="Bleeker" baseAuthorityYear="1855" box="[784,978,1575,1601]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="27" pageNumber="112" phylum="Chordata" rank="species" species="amblycephalus">
<emphasis id="DBA4EA9DFFD11F74FCAAC24D66BC4034" box="[784,978,1575,1601]" italics="true" pageId="27" pageNumber="112">J. amblycephalus</emphasis>
</taxonomicName>
is the basal clade. According to pairwise K2P genetic distances (
<tableCitation id="A4520334FFD11F74FE1CC22E64914013" box="[422,511,1611,1638]" captionStart="TABLE 9" captionStartId="26.[152,244,1601,1625]" captionTargetPageId="26" captionText="TABLE 9. Genetic distances for the COI gene among congener species with Johnius trewavasae species and Dendrophysa russelii as an outgroup." httpUri="http://table.plazi.org/id/BDAF6607FFD01F75FF22C22464FF4008" pageId="27" pageNumber="112" tableUuid="BDAF6607FFD01F75FF22C22464FF4008">Table 9</tableCitation>
),
<taxonomicName id="2ED04D0CFFD11F74FDA2C22967DB4010" authorityName="Sasaki" authorityYear="1992" box="[536,693,1612,1637]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="27" pageNumber="112" phylum="Chordata" rank="species" species="trewavasae">
<emphasis id="DBA4EA9DFFD11F74FDA2C22967DB4010" box="[536,693,1612,1637]" italics="true" pageId="27" pageNumber="112">J. trewavasae</emphasis>
</taxonomicName>
is 16% distinct from
<taxonomicName id="2ED04D0CFFD11F74FC0CC229614C4010" box="[950,1058,1611,1637]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="27" pageNumber="112" phylum="Chordata" rank="species" species="sasakii">
<emphasis id="DBA4EA9DFFD11F74FC0CC229614C4010" box="[950,1058,1611,1637]" italics="true" pageId="27" pageNumber="112">J. sasakii</emphasis>
</taxonomicName>
, 16% different from
<taxonomicName id="2ED04D0CFFD11F74FAA1C22960FB4010" baseAuthorityName="Cuvier" baseAuthorityYear="1830" box="[1307,1429,1612,1637]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="27" pageNumber="112" phylum="Chordata" rank="species" species="carouna">
<emphasis id="DBA4EA9DFFD11F74FAA1C22960FB4010" box="[1307,1429,1612,1637]" italics="true" pageId="27" pageNumber="112">J. carouna</emphasis>
</taxonomicName>
, 15% different from
<taxonomicName id="2ED04D0CFFD11F74FEC0C215677E40FC" authorityName="Bleeker" authorityYear="1873" box="[378,528,1647,1673]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="27" pageNumber="112" phylum="Chordata" rank="species" species="heterolepis">
<emphasis id="DBA4EA9DFFD11F74FEC0C215677E40FC" box="[378,528,1647,1673]" italics="true" pageId="27" pageNumber="112">J. heterolepis</emphasis>
</taxonomicName>
, 36% different from
<taxonomicName id="2ED04D0CFFD11F74FCB9C21566A840FC" baseAuthorityName="Bleeker" baseAuthorityYear="1855" box="[771,966,1647,1673]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="27" pageNumber="112" phylum="Chordata" rank="species" species="amblycephalus">
<emphasis id="DBA4EA9DFFD11F74FCB9C21566A840FC" box="[771,966,1647,1673]" italics="true" pageId="27" pageNumber="112">J. amblycephalus</emphasis>
</taxonomicName>
, and 36% different from the outgroup
<taxonomicName id="2ED04D0CFFD11F74FA3BC215658740D8" baseAuthorityName="Cuvier" baseAuthorityYear="1829" family="Sciaenidae" genus="Dendrophysa" kingdom="Animalia" order="Perciformes" pageId="27" pageNumber="112" phylum="Chordata" rank="species" species="russelii">
<emphasis id="DBA4EA9DFFD11F74FA3BC215658740D8" italics="true" pageId="27" pageNumber="112">D. russelii</emphasis>
</taxonomicName>
. In conclusions, the
<taxonomicName id="2ED04D0CFFD11F74FE66C2F667E040D8" box="[476,654,1683,1709]" pageId="27" pageNumber="112">
<emphasis id="DBA4EA9DFFD11F74FE66C2F6675D40D8" box="[476,563,1683,1709]" italics="true" pageId="27" pageNumber="112">Johnius</emphasis>
species
</taxonomicName>
in this section formed a monophyletic clade, which integrated with complementary approach supported by morphological and genetic evidence confirming the
<taxonomicName id="2ED04D0CFFD11F74FB21C2DD605A40A4" authorityName="Sasaki" authorityYear="1992" box="[1179,1332,1720,1745]" family="Sciaenidae" genus="Johnius" kingdom="Animalia" order="Perciformes" pageId="27" pageNumber="112" phylum="Chordata" rank="species" species="trewavasae">
<emphasis id="DBA4EA9DFFD11F74FB21C2DD605A40A4" box="[1179,1332,1720,1745]" italics="true" pageId="27" pageNumber="112">J. trewavasae</emphasis>
</taxonomicName>
status.
</paragraph>
</subSubSection>
</treatment>
</document>