362 lines
43 KiB
XML
362 lines
43 KiB
XML
<document id="19724308BCC87B2DA8F59B8CCD476292" ID-DOI="10.5281/zenodo.5351926" ID-ISSN="2345-7600" ID-Zenodo-Dep="5351926" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630207279294" checkinUser="tatiana" docAuthor="Hui, Tan Heok" docDate="2013" docId="615C8787D740FFBC4956ABF787DE4F67" docLanguage="en" docName="61rbz323-330.pdf" docOrigin="Raffles Bulletin of Zoology 61 (1)" docStyle="DocumentStyle:C6443723295B2E4C6F22B9B23F08089C.5:RafflesBZool.2006-2013.journal_article" docStyleId="C6443723295B2E4C6F22B9B23F08089C" docStyleName="RafflesBZool.2006-2013.journal_article" docStyleVersion="5" docTitle="Betta rubra Perugia 1893" docType="treatment" docVersion="3" lastPageNumber="326" masterDocId="9D65FFFFD741FFBF484BAD6B85344D52" masterDocTitle="The Identity Of Betta Rubra (Teleostei: Osphronemidae) Revisited, With Description Of A New Species From Sumatra, Indonesia" masterLastPageNumber="330" masterPageNumber="323" pageNumber="324" updateTime="1699185151093" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="B820828A2E39EA7B0DA626F2FD2F1360">The Identity Of Betta Rubra (Teleostei: Osphronemidae) Revisited, With Description Of A New Species From Sumatra, Indonesia</mods:title>
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<mods:name id="D21068E75A4435C59E691ED20F4C85F1" type="personal">
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<mods:namePart id="EAE7CD79414E75F9882FFD02717180D6">Hui, Tan Heok</mods:namePart>
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<mods:title id="7996A48193F1908A14D51E908A9AF55C">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="3CCEA44BD0316EDFD47C6C82BB9D5824">
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<mods:date id="A02A444B9F80A854329533174CD62AF8">2013</mods:date>
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<mods:number id="8BA1848D3CB374A3CC13F0CD03E765BE">2013-02-28</mods:number>
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<mods:detail id="67334A0EB64D7CE413D62B21E044A451" type="volume">
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<mods:number id="7D6F51B1D7834F05C6692A8F62BE29A1">61</mods:number>
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<mods:start id="269FF9ED5ED161E3630586409E5AC754">323</mods:start>
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<mods:identifier id="368F53A05769A899F07EE4BAD8D055BB" type="DOI">10.5281/zenodo.5351926</mods:identifier>
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<treatment id="615C8787D740FFBC4956ABF787DE4F67" LSID="urn:lsid:plazi:treatment:615C8787D740FFBC4956ABF787DE4F67" httpUri="http://treatment.plazi.org/id/615C8787D740FFBC4956ABF787DE4F67" lastPageId="3" lastPageNumber="326" pageId="1" pageNumber="324">
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<subSubSection id="A1EF651AD740FFBE4956ABF787704BE6" box="[285,580,1691,1716]" pageId="1" pageNumber="324" type="nomenclature">
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<paragraph id="E94A3691D740FFBE4956ABF787704BE6" blockId="1.[271,595,1691,1748]" box="[285,580,1691,1716]" pageId="1" pageNumber="324">
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<heading id="B20281FDD740FFBE4956ABF787704BE6" bold="true" box="[285,580,1691,1716]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="324" reason="2">
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<taxonomicName id="2EF54D12D740FFBE4956ABF787704BE6" ID-CoL="LP2M" authority="Perugia, 1893" authorityName="Perugia" authorityYear="1893" box="[285,580,1691,1716]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="324" phylum="Chordata" rank="species" species="rubra">
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<emphasis id="DB81EA83D740FFBE4956ABF787704BE6" bold="true" box="[285,580,1691,1716]" pageId="1" pageNumber="324">
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<emphasis id="DB81EA83D740FFBE4956ABF784AA4BE6" bold="true" box="[285,414,1691,1716]" italics="true" pageId="1" pageNumber="324">Betta rubra</emphasis>
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<bibRefCitation id="8D644B60D740FFBE49ECABF787704BE6" author="Perugia, A" box="[423,580,1692,1716]" pageId="1" pageNumber="324" pagination="241 - 247" refId="ref5115" refString="Perugia, A., 1893. Di alcuni pesci raccolti in Sumatra dal Dott. Elio Modigliani. Annali del Museo Civico di Storia Naturale Giacoma Doria, Genova, 13: 241 - 247." type="journal article" year="1893">Perugia, 1893</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="A1EF651AD740FFBE4944ABD787674B86" box="[271,595,1724,1748]" pageId="1" pageNumber="324" type="description">
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<paragraph id="E94A3691D740FFBE4944ABD787674B86" blockId="1.[271,595,1691,1748]" box="[271,595,1724,1748]" pageId="1" pageNumber="324">
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(
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||
<figureCitation id="71CE2A14D740FFBE495DABD784AC4B86" box="[278,408,1724,1748]" captionStart="Fig" captionStartId="2.[136,171,2037,2059]" captionTargetBox="[136,1469,632,2007]" captionTargetId="figure-8@2.[136,1470,632,2007]" captionTargetPageId="2" captionText="Fig. 1. Comparison of lateral head and opercle pattern, left side with photograph, right side with schematic drawing. A, B, Betta rubra, MSNG 13019, lectotype, 28.6 mm SL; C, D, Betta rubra, ZRC 53989, 34.0 mm SL male; E, F, Betta dennisyongi, MZB 17207, holotype, 27.4 mm SL male." figureDoi="http://doi.org/10.5281/zenodo.5351928" httpUri="https://zenodo.org/record/5351928/files/figure.png" pageId="1" pageNumber="324">Figs. 1A–D</figureCitation>
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,
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||
<figureCitation id="71CE2A14D740FFBE49EEABD784DC4B86" box="[421,488,1724,1748]" captionStart="Fig" captionStartId="3.[113,148,2031,2053]" captionTargetBox="[113,751,1426,2008]" captionTargetId="figure-11@3.[113,753,1425,2008]" captionTargetPageId="3" captionText="Fig. 2. Betta rubra: A, MSNG 13019a, lectotype, 28.6 mm SL, left side; B, lectotype, right side, Sibolga; C, BMNH 1893.5.29:1, paralectotype, 33.3 mm SL, Lake Toba." figureDoi="http://doi.org/10.5281/zenodo.5351934" httpUri="https://zenodo.org/record/5351934/files/figure.png" pageId="1" pageNumber="324">2A–C</figureCitation>
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,
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<figureCitation id="71CE2A14D740FFBE49BFABD7871B4B86" box="[500,559,1724,1748]" captionStart="Fig" captionStartId="3.[812,847,977,999]" captionTargetBox="[812,1453,543,951]" captionTargetId="figure-41@3.[812,1455,540,954]" captionTargetPageId="3" captionText="Fig. 3. Betta rubra: A, ZRC 53989, 34.0 mm SL male; B, ZRC 53989, 37.9 mm SL female; freshly collected specimens from southern Aceh." figureDoi="http://doi.org/10.5281/zenodo.5351930" httpUri="https://zenodo.org/record/5351930/files/figure.png" pageId="1" pageNumber="324">3A,B</figureCitation>
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||
,
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||
<figureCitation id="71CE2A14D740FFBE4A76ABD7877F4B86" box="[573,587,1724,1748]" captionStart="Fig" captionStartId="3.[812,847,1577,1599]" captionTargetBox="[812,1451,1130,1554]" captionTargetId="figure-43@3.[812,1453,1129,1554]" captionTargetPageId="3" captionText="Fig. 4. Betta rubra live male specimen from southern Aceh (from same series as ZRC 53989, not preserved)." figureDoi="http://doi.org/10.5281/zenodo.5351932" httpUri="https://zenodo.org/record/5351932/files/figure.png" pageId="1" pageNumber="324">4</figureCitation>
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)
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="A1EF651AD740FFBE483AAB9787EB4A35" pageId="1" pageNumber="324" type="reference_group">
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<paragraph id="E94A3691D740FFBE483AAB9787EB4A35" blockId="1.[113,753,1787,1895]" pageId="1" pageNumber="324">
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<treatmentCitationGroup id="C9E511BFD740FFBE483AAB9787EB4A35" pageId="1" pageNumber="324">
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<treatmentCitation id="68541080D740FFBE483AAB9784964A42" author="Perugia, A" box="[113,418,1787,1809]" page="242" pageId="1" pageNumber="324" year="1893">
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||
<taxonomicName id="2EF54D12D740FFBE483AAB9784964A42" ID-CoL="LP2M" authority="Perugia, 1893: 242" authorityName="Perugia" authorityPageNumber="242" authorityYear="1893" box="[113,418,1787,1809]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="324" phylum="Chordata" rank="species" species="rubra">
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||
<emphasis id="DB81EA83D740FFBE483AAB9785D64A42" box="[113,226,1788,1808]" italics="true" pageId="1" pageNumber="324">Betta rubra</emphasis>
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||
<bibRefCitation id="8D644B60D740FFBE48A3AB9084964A42" author="Perugia, A" box="[232,418,1787,1809]" pageId="1" pageNumber="324" pagination="241 - 247" refId="ref5115" refString="Perugia, A., 1893. Di alcuni pesci raccolti in Sumatra dal Dott. Elio Modigliani. Annali del Museo Civico di Storia Naturale Giacoma Doria, Genova, 13: 241 - 247." type="journal article" year="1893">Perugia, 1893: 242</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
;
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||
<treatmentCitation id="68541080D740FFBE49E7AB9087724A42" author="Volz, W" box="[428,582,1787,1809]" page="127" pageId="1" pageNumber="324" year="1905">
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||
<bibRefCitation id="8D644B60D740FFBE49E7AB9087724A42" author="Volz, W" box="[428,582,1787,1809]" pageId="1" pageNumber="324" pagination="35 - 250" refId="ref5377" refString="Volz, W., 1905. Catalogue of the fishes of Sumatra. Natuurkundig Tijdschrift voor Nederlandsch Indie, 9: 35 - 250." type="journal article" year="1905">Volz, 1905: 127</bibRefCitation>
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||
</treatmentCitation>
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||
(part), 1907: 127 (part); Regan, 1910: 781, pl. 77,
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||
<figureCitation id="71CE2A14D740FFBE4980AA7C87374A7F" box="[459,515,1815,1837]" captionStart="Fig" captionStartId="2.[136,171,2037,2059]" captionTargetBox="[136,1469,632,2007]" captionTargetId="figure-8@2.[136,1470,632,2007]" captionTargetPageId="2" captionText="Fig. 1. Comparison of lateral head and opercle pattern, left side with photograph, right side with schematic drawing. A, B, Betta rubra, MSNG 13019, lectotype, 28.6 mm SL; C, D, Betta rubra, ZRC 53989, 34.0 mm SL male; E, F, Betta dennisyongi, MZB 17207, holotype, 27.4 mm SL male." figureDoi="http://doi.org/10.5281/zenodo.5351928" httpUri="https://zenodo.org/record/5351928/files/figure.png" pageId="1" pageNumber="324">Fig. 1</figureCitation>
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||
; Witte & Schmidt, 1992: 325;
|
||
<treatmentCitation id="68541080D740FFBE4881AA5F84F74A18" author="Kottelat, M & Whitten, S" box="[202,451,1844,1866]" page="163" pageId="1" pageNumber="324" year="1993">
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||
<bibRefCitation id="8D644B60D740FFBE4881AA5F84F74A18" author="Kottelat, M & Whitten, S" box="[202,451,1844,1866]" pageId="1" pageNumber="324" refId="ref5034" refString="Kottelat, M., A. J. Whitten, S. N. Kartikasari & S. Wirjoatmodjo, 1993. Freshwater Fishes of Western Indonesia and Sulawesi. Periplus Editions, Hong Kong. 259 pp., 84 pls." type="book" year="1993">Kottelat et al., 1993: 163</bibRefCitation>
|
||
</treatmentCitation>
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||
, pl. 76; Tan & Ng, 2005a: 58; Tan & Ng, 2005b: 119,
|
||
<figureCitation id="71CE2A14D740FFBE49CFAA3A84EE4A35" box="[388,474,1873,1895]" captionStart="Fig" captionStartId="2.[136,171,2037,2059]" captionTargetBox="[136,1469,632,2007]" captionTargetId="figure-8@2.[136,1470,632,2007]" captionTargetPageId="2" captionText="Fig. 1. Comparison of lateral head and opercle pattern, left side with photograph, right side with schematic drawing. A, B, Betta rubra, MSNG 13019, lectotype, 28.6 mm SL; C, D, Betta rubra, ZRC 53989, 34.0 mm SL male; E, F, Betta dennisyongi, MZB 17207, holotype, 27.4 mm SL male." figureDoi="http://doi.org/10.5281/zenodo.5351928" httpUri="https://zenodo.org/record/5351928/files/figure.png" pageId="1" pageNumber="324">Fig. 1a,b</figureCitation>
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||
(part);
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||
<treatmentCitation id="68541080D740FFBE4A6EAA3987954A35" author="Tan, H" box="[549,673,1873,1895]" page="5" pageId="1" pageNumber="324" year="2008">
|
||
<bibRefCitation id="8D644B60D740FFBE4A6EAA3987954A35" author="Tan, H" box="[549,673,1873,1895]" pageId="1" pageNumber="324" pagination="5 - 8" refId="ref5235" refString="Tan, H. H., 2008. The enigma of the red Sumatran fighter (Betta rubra) unveiled [printed as: (Betta rubra) unveiled]. Anabantoid Association of Great Britain newsletter, Labyrinth, 152: 5 - 8." type="journal article" year="2008">Tan, 2008: 5</bibRefCitation>
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||
</treatmentCitation>
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||
(part)
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||
</treatmentCitationGroup>
|
||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="A1EF651AD740FFBE483AAAE186E24C17" pageId="1" pageNumber="324" type="materials_examined">
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||
<paragraph id="E94A3691D740FFBE483AAAE187C54562" blockId="1.[113,753,1930,2097]" pageId="1" pageNumber="324">
|
||
<emphasis id="DB81EA83D740FFBE483AAAE1847A4AF2" bold="true" box="[113,334,1930,1952]" pageId="1" pageNumber="324">
|
||
<emphasis id="DB81EA83D740FFBE483AAAE184184AF2" bold="true" box="[113,300,1930,1952]" italics="true" pageId="1" pageNumber="324">Material examined</emphasis>
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||
. —
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||
</emphasis>
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||
<materialsCitation id="599D3CCCD740FFBE4918AAE084BD4AEF" collectionCode="MSNG" location="West Sumatra Province" pageId="1" pageNumber="324" specimenCode="MSNG 13019" specimenCount="1" stateProvince="Sumatera Barat" typeStatus="lectotype">
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||
<emphasis id="DB81EA83D740FFBE4918AAE087774AF2" bold="true" box="[339,579,1930,1952]" pageId="1" pageNumber="324">
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<collectingRegion id="2B31F873D740FFBE4918AAE087774AF2" box="[339,579,1930,1952]" country="Indonesia" name="Sumatera Barat" pageId="1" pageNumber="324">West Sumatra Province</collectingRegion>
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||
</emphasis>
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||
–
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||
<specimenCode id="B9539EEAD740FFBE4A12AAE087D54AF2" box="[601,737,1931,1952]" collectionCode="MSNG" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="1" pageNumber="324">MSNG 13019</specimenCode>
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a,
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<specimenCount id="FFF3FD18D740FFBE483AAAC385964AEF" box="[113,162,1960,1981]" pageId="1" pageNumber="324" type="generic">1 ex.</specimenCount>
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,
|
||
<typeStatus id="364E8833D740FFBE48E0AACC84314AEF" box="[171,261,1959,1981]" pageId="1" pageNumber="324" type="lectotype">lectotype</typeStatus>
|
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,
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||
<quantity id="2E0D9B74D740FFBE4944AACC84504AEF" box="[271,356,1959,1981]" metricMagnitude="-2" metricUnit="m" metricValue="2.86" pageId="1" pageNumber="324" unit="mm" value="28.6">28.6 mm</quantity>
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||
SL
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="599D3CCCD740FFBE49D8AAC3877C4A88" collectionCode="MSNG" collectorName="Siboga & E. Modigliani" pageId="1" pageNumber="324" specimenCode="MSNG 13019" specimenCount="4" stateProvince="West Sumatra Province" typeStatus="paralectotype">
|
||
<specimenCode id="B9539EEAD740FFBE49D8AAC387284AEF" box="[403,540,1960,1981]" collectionCode="MSNG" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="1" pageNumber="324">MSNG 13019</specimenCode>
|
||
,
|
||
<specimenCount id="FFF3FD18D740FFBE4A6EAAC387624AEF" box="[549,598,1960,1981]" pageId="1" pageNumber="324" type="generic">4 ex.</specimenCount>
|
||
,
|
||
<typeStatus id="364E8833D740FFBE4A2BAACC87D94AEF" box="[608,749,1959,1981]" pageId="1" pageNumber="324" type="paralectotype">paralectotypes</typeStatus>
|
||
,
|
||
<quantity id="2E0D9B74D740FFBE483AAAAF85CA4A88" box="[113,254,1988,2010]" metricMagnitude="-2" metricUnit="m" metricValue="2.915" metricValueMax="3.1399999999999997" metricValueMin="2.69" pageId="1" pageNumber="324" unit="mm" value="29.15" valueMax="31.4" valueMin="26.9">26.9–31.4 mm</quantity>
|
||
SL;
|
||
<collectorName id="44005347D740FFBE497BAAAF84424A88" box="[304,374,1988,2010]" pageId="1" pageNumber="324">Siboga</collectorName>
|
||
;
|
||
<collectorName id="44005347D740FFBE49CBAAAE873E4A88" box="[384,522,1988,2010]" pageId="1" pageNumber="324">E. Modigliani</collectorName>
|
||
, 1886
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="599D3CCCD740FFBE4A19AAAF84454546" collectingDate="1893-05-29" collectionCode="BMNH" collectorName="E. Modigliani" location="Lake Toba" pageId="1" pageNumber="324" specimenCount="1" stateProvince="Sumatera Utara" typeStatus="paralectotype">
|
||
<emphasis id="DB81EA83D740FFBE4A19AAAF85FF4AA5" bold="true" pageId="1" pageNumber="324">
|
||
<collectingRegion id="2B31F873D740FFBE4A19AAAF85FF4AA5" country="Indonesia" name="Sumatera Utara" pageId="1" pageNumber="324">North Sumatra Province</collectingRegion>
|
||
</emphasis>
|
||
–
|
||
<collectionCode id="8FE4AE54D740FFBE48A9AA89841E4AA5" box="[226,298,2018,2039]" pageId="1" pageNumber="324">BMNH</collectionCode>
|
||
<date id="9D4B1051D740FFBE497BAA8A84AE4AA5" box="[304,410,2017,2039]" pageId="1" pageNumber="324" value="1893-05-29">
|
||
<collectingDate id="8D0FE9B9D740FFBE497BAA8A84AE4AA5" box="[304,410,2017,2039]" pageId="1" pageNumber="324" value="1893-05-29">1893.5.29</collectingDate>
|
||
</date>
|
||
:1,
|
||
<specimenCount id="FFF3FD18D740FFBE49E5AA8A84EA4AA5" box="[430,478,2017,2039]" pageId="1" pageNumber="324" type="generic">1 ex.</specimenCount>
|
||
,
|
||
<typeStatus id="364E8833D740FFBE49ACAA8A875F4AA5" box="[487,619,2017,2039]" pageId="1" pageNumber="324" type="paralectotype">paralectotype</typeStatus>
|
||
,
|
||
<quantity id="2E0D9B74D740FFBE4A38AA8A87FC4AA5" box="[627,712,2017,2039]" metricMagnitude="-2" metricUnit="m" metricValue="3.3299999999999996" pageId="1" pageNumber="324" unit="mm" value="33.3">33.3 mm</quantity>
|
||
SL;
|
||
<location id="EC2A604AD740FFBE483AAA9585E84546" LSID="urn:lsid:plazi:treatment:615C8787D740FFBC4956ABF787DE4F67:EC2A604AD740FFBE483AAA9585E84546" box="[113,220,2046,2068]" name="Lake Toba" pageId="1" pageNumber="324" stateProvince="Sumatera Utara">Lake Toba</location>
|
||
;
|
||
<collectorName id="44005347D740FFBE48ACAA9584454546" box="[231,369,2046,2068]" pageId="1" pageNumber="324">E. Modigliani</collectorName>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="599D3CCCD740FFBE4937AA9587A54562" accessionNumber="ZRC 53989, CMK 23280" collectionCode="ZRC" location="Aceh Province" pageId="1" pageNumber="324" specimenCount="44" stateProvince="Aceh" typeStatus="paralectotype">
|
||
<emphasis id="DB81EA83D740FFBE4937AA9587264546" bold="true" box="[380,530,2046,2068]" pageId="1" pageNumber="324">
|
||
<collectingRegion id="2B31F873D740FFBE4937AA9587264546" box="[380,530,2046,2068]" country="Indonesia" name="Aceh" pageId="1" pageNumber="324">Aceh Province</collectingRegion>
|
||
</emphasis>
|
||
–
|
||
<collectionCode id="8FE4AE54D740FFBE4A66AA9587684541" box="[557,604,2046,2067]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="324">ZRC</collectionCode>
|
||
53989,
|
||
<specimenCount id="FFF3FD18D740FFBE4AE5AA9587D94541" box="[686,749,2046,2067]" pageId="1" pageNumber="324" type="generic">40 ex.</specimenCount>
|
||
,
|
||
<quantity id="2E0D9B74D740FFBE483AA57085CF4562" box="[113,251,2075,2097]" metricMagnitude="-2" metricUnit="m" metricValue="3.4299999999999997" metricValueMax="4.21" metricValueMin="2.65" pageId="1" pageNumber="324" unit="mm" value="34.3" valueMax="42.1" valueMin="26.5">26.5–42.1 mm</quantity>
|
||
SL; CMK 23280,
|
||
<specimenCount id="FFF3FD18D740FFBE49E1A57084EE4562" box="[426,474,2075,2096]" pageId="1" pageNumber="324" type="generic">4 ex.</specimenCount>
|
||
,
|
||
<quantity id="2E0D9B74D740FFBE49A8A57087594562" box="[483,621,2075,2097]" metricMagnitude="-2" metricUnit="m" metricValue="3.37" metricValueMax="3.56" metricValueMin="3.18" pageId="1" pageNumber="324" unit="mm" value="33.7" valueMax="35.6" valueMin="31.8">31.8–35.6 mm</quantity>
|
||
SL
|
||
</materialsCitation>
|
||
;
|
||
<collectingRegion id="2B31F873D740FFBE4AD2A57087D94562" box="[665,749,2075,2096]" country="Indonesia" name="Sumatera Utara" pageId="1" pageNumber="324">Sumatra</collectingRegion>
|
||
:
|
||
</paragraph>
|
||
<paragraph id="E94A3691D740FFBE4B66ADD786E24C17" blockId="1.[813,1452,188,325]" pageId="1" pageNumber="324">
|
||
<materialsCitation id="599D3CCCD740FFBE4B66ADD786E34D80" box="[813,983,188,210]" collectorName="Kabupaten" location="Aceh" pageId="1" pageNumber="324" specimenCount="1" stateProvince="Aceh">
|
||
<collectingRegion id="2B31F873D740FFBE4B66ADD786574D80" box="[813,867,188,210]" country="Indonesia" name="Aceh" pageId="1" pageNumber="324">Aceh</collectingRegion>
|
||
:
|
||
<collectorName id="44005347D740FFBE4B24ADD786E34D80" box="[879,983,188,210]" pageId="1" pageNumber="324">Kabupaten</collectorName>
|
||
</materialsCitation>
|
||
<materialsCitation id="599D3CCCD740FFBE4B95ADD780464C59" collectingDate="2009-04-19" collectionCode="T" collectorName="Sim" county="Singkil" elevation="12" location="Singkil-Subulussalam" municipality="Gosong Telaga" pageId="1" pageNumber="324" specimenCount="1" stateProvince="Aceh">
|
||
<collectingRegion id="2B31F873D740FFBE4B95ADD781254D80" box="[990,1041,188,210]" country="Indonesia" name="Aceh" pageId="1" pageNumber="324">Aceh</collectingRegion>
|
||
<collectingCounty id="002B4E1DD740FFBE4C52ADD781564D80" box="[1049,1122,188,210]" pageId="1" pageNumber="324">Singkil</collectingCounty>
|
||
: remnant peat swamp forest near
|
||
<collectingMunicipality id="092EACEBD740FFBE4B66ADB286FC4DBD" box="[813,968,217,239]" pageId="1" pageNumber="324">Gosong Telaga</collectingMunicipality>
|
||
, along road
|
||
<location id="EC2A604AD740FFBE4C07ADB2801E4DBD" LSID="urn:lsid:plazi:treatment:615C8787D740FFBC4956ABF787DE4F67:EC2A604AD740FFBE4C07ADB2801E4DBD" box="[1100,1322,217,239]" county="Singkil" municipality="Gosong Telaga" name="Singkil-Subulussalam" pageId="1" pageNumber="324" stateProvince="Aceh">Singkil-Subulussalam</location>
|
||
, outskirts of
|
||
<location id="EC2A604AD740FFBE4B66AD9D869D4C59" LSID="urn:lsid:plazi:treatment:615C8787D740FFBC4956ABF787DE4F67:EC2A604AD740FFBE4B66AD9D869D4C59" box="[813,937,246,268]" county="Singkil" municipality="Gosong Telaga" name="Singkil town" pageId="1" pageNumber="324" stateProvince="Aceh">Singkil town</location>
|
||
(
|
||
<elevation id="62D8D1A2D740FFBE4BF3AD9D813B4C5E" box="[952,1039,246,268]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="324" unit="m" value="12.0">
|
||
<quantity id="2E0D9B74D740FFBE4BF3AD9D86DF4C59" box="[952,1003,246,267]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="324" unit="m" value="12.0">12 m</quantity>
|
||
asl
|
||
</elevation>
|
||
, pH 5.5);
|
||
<collectionCode id="8FE4AE54D740FFBE4C39AD9D81B64C59" box="[1138,1154,246,267]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="324">T</collectionCode>
|
||
.
|
||
<collectorName id="44005347D740FFBE4CC7AD9D81874C5E" box="[1164,1203,246,268]" pageId="1" pageNumber="324">Sim</collectorName>
|
||
et al.,
|
||
<date id="9D4B1051D740FFBE4CBCAD9D80464C59" box="[1271,1394,246,267]" pageId="1" pageNumber="324" value="2009-04-19">
|
||
<collectingDate id="8D0FE9B9D740FFBE4CBCAD9D80464C59" box="[1271,1394,246,267]" pageId="1" pageNumber="324" value="2009-04-19">19 Apr.2009</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="599D3CCCD740FFBE4D36AD9D81134C7A" accessionNumber="ZRC 54000" collectionCode="ZRC" pageId="1" pageNumber="324" specimenCount="1" stateProvince="Aceh">
|
||
<collectionCode id="8FE4AE54D740FFBE4D36AD9D80984C59" box="[1405,1452,246,267]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="324">ZRC</collectionCode>
|
||
54000,
|
||
<specimenCount id="FFF3FD18D740FFBE4B3FAC7886914C7A" box="[884,933,275,296]" pageId="1" pageNumber="324" type="generic">1 ex.</specimenCount>
|
||
, 34.0 mm SL
|
||
</materialsCitation>
|
||
;
|
||
<collectingRegion id="2B31F873D740FFBE4C7BAC7881B04C7A" box="[1072,1156,275,296]" country="Indonesia" name="Sumatera Utara" pageId="1" pageNumber="324">Sumatra</collectingRegion>
|
||
: southern
|
||
<materialsCitation id="599D3CCCD740FFBE4CA8AC7986E64C17" collectingDate="2008-02" location="Ishizu" pageId="1" pageNumber="324" specimenCount="1" stateProvince="Aceh">
|
||
<collectingRegion id="2B31F873D740FFBE4CA8AC79802C4C7A" box="[1251,1304,274,296]" country="Indonesia" name="Aceh" pageId="1" pageNumber="324">Aceh</collectingRegion>
|
||
, trade material;
|
||
<location id="EC2A604AD740FFBE4B66AC44865E4C17" LSID="urn:lsid:plazi:treatment:615C8787D740FFBC4956ABF787DE4F67:EC2A604AD740FFBE4B66AC44865E4C17" box="[813,874,303,325]" name="Ishizu" pageId="1" pageNumber="324" stateProvince="Aceh">Ishizu</location>
|
||
,
|
||
<date id="9D4B1051D740FFBE4B3EAC4486E64C17" box="[885,978,303,325]" pageId="1" pageNumber="324" value="2008-02">
|
||
<collectingDate id="8D0FE9B9D740FFBE4B3EAC4486E64C17" box="[885,978,303,325]" pageId="1" pageNumber="324" value="2008-02">Feb.2008</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A1EF651AD740FFBE4B66AC0680154E14" pageId="1" pageNumber="324" type="diagnosis">
|
||
<paragraph id="E94A3691D740FFBE4B66AC0680154E14" blockId="1.[813,1452,365,838]" pageId="1" pageNumber="324">
|
||
<emphasis id="DB81EA83D740FFBE4B66AC06869F4CD4" bold="true" box="[813,939,365,390]" pageId="1" pageNumber="324">
|
||
<emphasis id="DB81EA83D740FFBE4B66AC0686974CD7" bold="true" box="[813,931,365,389]" italics="true" pageId="1" pageNumber="324">Diagnosis</emphasis>
|
||
.
|
||
</emphasis>
|
||
—
|
||
<taxonomicName id="2EF54D12D740FFBE4B95AC0481534CD4" authorityName="Perugia" authorityYear="1893" box="[990,1127,367,390]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="324" phylum="Chordata" rank="species" species="rubra">
|
||
<emphasis id="DB81EA83D740FFBE4B95AC0481534CD4" box="[990,1127,367,390]" italics="true" pageId="1" pageNumber="324">Betta rubra</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from
|
||
<taxonomicName id="2EF54D12D740FFBE4B66ACE486E24CF4" authorityName="Hui" authorityYear="2013" box="[813,982,399,422]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="324" phylum="Chordata" rank="species" species="dennisyongi">
|
||
<emphasis id="DB81EA83D740FFBE4B66ACE486E24CF4" box="[813,982,399,422]" italics="true" pageId="1" pageNumber="324">B. dennisyongi</emphasis>
|
||
</taxonomicName>
|
||
by the following suite of characters: an interrupted black postorbital stripe from posterior of eye to mid-opercle, with a large black bar-like marking on posterior half of opercle not extending to opercle edge (vs. a continuous stripe), leaving a clear cream edge over entire opercle margin (vs. not entire opercel margin); less broad suborbital stripe below eye forming a triangular black mark (2–3 scale rows wide, vs. 4–5); more intense body colouration with smaller interspaces between the black body bars; a greater number of anal-fin rays (mode 27, vs. 25); fewer subdorsal scales (5–5½, vs. 6–7); fewer lateral scales (mode 30, vs. 31); fewer vertebrae (mode 29, vs. 30); shorter dorsal-fin base length (11.0–13.4, vs. 13.0–18.4% SL); and larger maximum size (42.1, vs.
|
||
<quantity id="2E0D9B74D740FFBE4CC3AE4681DC4E17" box="[1160,1256,813,837]" metricMagnitude="-2" metricUnit="m" metricValue="3.54" pageId="1" pageNumber="324" unit="mm" value="35.4">35.4 mm</quantity>
|
||
SL).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A1EF651AD740FFBD4B66AE06801D4F67" lastPageId="2" lastPageNumber="325" pageId="1" pageNumber="324" type="description">
|
||
<paragraph id="E94A3691D740FFBE4B66AE06819549B4" blockId="1.[813,1452,877,1254]" pageId="1" pageNumber="324">
|
||
<emphasis id="DB81EA83D740FFBE4B66AE0686874ED4" bold="true" box="[813,947,877,902]" pageId="1" pageNumber="324">
|
||
<emphasis id="DB81EA83D740FFBE4B66AE0686994ED7" bold="true" box="[813,941,877,901]" italics="true" pageId="1" pageNumber="324">Description</emphasis>
|
||
.
|
||
</emphasis>
|
||
— General appearance as illustrated in
|
||
<figureCitation id="71CE2A14D740FFBE4D3CAE06865C4EF4" captionStart="Fig" captionStartId="3.[812,847,977,999]" captionTargetBox="[812,1453,543,951]" captionTargetId="figure-41@3.[812,1455,540,954]" captionTargetPageId="3" captionText="Fig. 3. Betta rubra: A, ZRC 53989, 34.0 mm SL male; B, ZRC 53989, 37.9 mm SL female; freshly collected specimens from southern Aceh." figureDoi="http://doi.org/10.5281/zenodo.5351930" httpUri="https://zenodo.org/record/5351930/files/figure.png" pageId="1" pageNumber="324">Figs. 3A,B</figureCitation>
|
||
,
|
||
<figureCitation id="71CE2A14D740FFBE4B3CAEE586B34EF4" box="[887,903,910,934]" captionStart="Fig" captionStartId="3.[812,847,1577,1599]" captionTargetBox="[812,1451,1130,1554]" captionTargetId="figure-43@3.[812,1453,1129,1554]" captionTargetPageId="3" captionText="Fig. 4. Betta rubra live male specimen from southern Aceh (from same series as ZRC 53989, not preserved)." figureDoi="http://doi.org/10.5281/zenodo.5351932" httpUri="https://zenodo.org/record/5351932/files/figure.png" pageId="1" pageNumber="324">4</figureCitation>
|
||
; meristic and morphometric data of
|
||
<taxonomicName id="2EF54D12D740FFBE4D69AEE480B44EF4" authorityName="Perugia" authorityYear="1893" box="[1314,1408,911,934]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="324" phylum="Chordata" rank="species" species="rubra">
|
||
<emphasis id="DB81EA83D740FFBE4D69AEE480B44EF4" box="[1314,1408,911,934]" italics="true" pageId="1" pageNumber="324">B. rubra</emphasis>
|
||
</taxonomicName>
|
||
are listed in Table 1. Body relatively slender (body depth 25.0– 28.1% SL), head relatively short (head length 30.4–35.7% SL). Dorsal and anal fins pointed, caudal fin rounded with extended median rays and interradial membrane; dorsal fin placed relatively far back (predorsal length 65.4–73.2% SL); anal-fin base length about half of standard length (52.2–58.3% SL); pelvic fin falcate with first ray long and filamentous (30.7–48.9% SL); pectoral fin rounded. Vertebral count: 9–10 + 18–20 = 27–29 (mode = 29, n = 15). Maximum size:
|
||
<quantity id="2E0D9B74D740FFBE4C5AA9A5814549B7" box="[1041,1137,1230,1254]" metricMagnitude="-2" metricUnit="m" metricValue="4.21" pageId="1" pageNumber="324" unit="mm" value="42.1">42.1 mm</quantity>
|
||
SL.
|
||
</paragraph>
|
||
<paragraph id="E94A3691D740FFBE4B66A86781C34AF7" blockId="1.[813,1452,1292,1957]" pageId="1" pageNumber="324">
|
||
<emphasis id="DB81EA83D740FFBE4B66A867810B4874" bold="true" box="[813,1087,1292,1318]" pageId="1" pageNumber="324">
|
||
<emphasis id="DB81EA83D740FFBE4B66A86781034877" bold="true" box="[813,1079,1292,1317]" italics="true" pageId="1" pageNumber="324">Preserved colouration</emphasis>
|
||
.
|
||
</emphasis>
|
||
— See
|
||
<figureCitation id="71CE2A14D740FFBE4CEDA86680064874" box="[1190,1330,1293,1318]" captionStart="Fig" captionStartId="3.[113,148,2031,2053]" captionTargetBox="[113,751,1426,2008]" captionTargetId="figure-11@3.[113,753,1425,2008]" captionTargetPageId="3" captionText="Fig. 2. Betta rubra: A, MSNG 13019a, lectotype, 28.6 mm SL, left side; B, lectotype, right side, Sibolga; C, BMNH 1893.5.29:1, paralectotype, 33.3 mm SL, Lake Toba." figureDoi="http://doi.org/10.5281/zenodo.5351934" httpUri="https://zenodo.org/record/5351934/files/figure.png" pageId="1" pageNumber="324">Figs. 2A–C</figureCitation>
|
||
,
|
||
<figureCitation id="71CE2A14D740FFBE4D09A865804A4874" box="[1346,1406,1294,1318]" captionStart="Fig" captionStartId="3.[812,847,977,999]" captionTargetBox="[812,1453,543,951]" captionTargetId="figure-41@3.[812,1455,540,954]" captionTargetPageId="3" captionText="Fig. 3. Betta rubra: A, ZRC 53989, 34.0 mm SL male; B, ZRC 53989, 37.9 mm SL female; freshly collected specimens from southern Aceh." figureDoi="http://doi.org/10.5281/zenodo.5351930" httpUri="https://zenodo.org/record/5351930/files/figure.png" pageId="1" pageNumber="324">3A,B</figureCitation>
|
||
for general appearance. Dorsum of body dark brown to black, lateral reddish, ventrum lighter brown to cream. Dorsum of head brown with black spots sometimes indistinct. Lower lip black. Distinct pre- and postorbital dark-brown to black stripes on head, an interrupted postorbital black stripe from posterior of eye to mid-opercle, with a large black almost bar-like marking on posterior half of opercle not extending to opercle edge, leaving a clear cream edge over entire opercular margin. Broad dark brown to black chin-bar/ suborbital stripe present, forming a triangular dark brown to black mark below eye, up to eye diameter in width. Opercle region basal colour brownish to reddish (depending on period of preservation). Male body with irregularly spaced blackish vertical 4–7 bars, restricted to lower half of body; bars prominent on anterior two-thirds of body. Female body dark brownish red, with reddish patches on ventral region. Caudal peduncle spot faint. Fins brownish, sometimes reddish. Dorsal, caudal and anal fins without transverse bars on interradial membrane; with hyaline edge. Pectoral fin hyaline; base black. Pelvic fin reddish.
|
||
</paragraph>
|
||
<paragraph id="E94A3691D740FFBE4B66AAA680984577" blockId="1.[813,1452,1996,2085]" pageId="1" pageNumber="324">
|
||
<emphasis id="DB81EA83D740FFBE4B66AAA686C74AB4" bold="true" box="[813,1011,1996,2022]" pageId="1" pageNumber="324">
|
||
<emphasis id="DB81EA83D740FFBE4B66AAA686DF4AB7" bold="true" box="[813,1003,1996,2021]" italics="true" pageId="1" pageNumber="324">Live colouration</emphasis>
|
||
.
|
||
</emphasis>
|
||
— See
|
||
<figureCitation id="71CE2A14D740FFBE4C1EAAA681AC4AB4" box="[1109,1176,1997,2022]" captionStart="Fig" captionStartId="3.[812,847,1577,1599]" captionTargetBox="[812,1451,1130,1554]" captionTargetId="figure-43@3.[812,1453,1129,1554]" captionTargetPageId="3" captionText="Fig. 4. Betta rubra live male specimen from southern Aceh (from same series as ZRC 53989, not preserved)." figureDoi="http://doi.org/10.5281/zenodo.5351932" httpUri="https://zenodo.org/record/5351932/files/figure.png" pageId="1" pageNumber="324">Fig. 4</figureCitation>
|
||
for general appearance. Basal body colour reddish to brownish. Dorsum darker than ventrum. Dorsum of head with dark brown to black spots.
|
||
</paragraph>
|
||
<paragraph id="E94A3691D743FFBD48C3ADD687CE4C87" blockId="2.[136,776,189,470]" pageId="2" pageNumber="325">Mouth with lower lip black. Lower half of iris of eye bright bluish-green. Head and opercle pattern as above; bar-like marking on posterior half of opercle with gold iridescence; opercle edge bright red; rest of opercle region reddish. Body with 4–7 irregularly spaced blackish bars, restricted to lower half of body. Fins reddish. Dorsal, caudal and anal fins with bright bluish-green edge; base of fins with greenish iridescence. Pectoral fin base with an iridescent bluish-green spot. Pelvic fin reddish with greenish flush and green tip.</paragraph>
|
||
<paragraph id="E94A3691D743FFBD48C3AC97801D4F67" blockId="2.[136,776,508,597]" lastBlockId="2.[835,1475,189,566]" pageId="2" pageNumber="325">
|
||
<emphasis id="DB81EA83D743FFBD48C3AC97843F4F47" bold="true" box="[136,267,508,533]" pageId="2" pageNumber="325">
|
||
<emphasis id="DB81EA83D743FFBD48C3AC9784304F47" bold="true" box="[136,260,508,533]" italics="true" pageId="2" pageNumber="325">Field notes</emphasis>
|
||
.
|
||
</emphasis>
|
||
—
|
||
<taxonomicName id="2EF54D12D743FFBD497EAC9484864F44" authorityName="Perugia" authorityYear="1893" box="[309,434,511,534]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="325" phylum="Chordata" rank="species" species="rubra">
|
||
<emphasis id="DB81EA83D743FFBD497EAC9484864F44" box="[309,434,511,534]" italics="true" pageId="2" pageNumber="325">Betta rubra</emphasis>
|
||
</taxonomicName>
|
||
has not been collected since its original description and no authoritative details of its habitat have ever been published in the scientific literature. The present series of fresh material of this species was obtained from a highly disturbed and fragmented environment. The surrounding peat swamp habitat in the Singkil area had already been cleared, apparently for the planting of oil palm. The surrounding peatlands had been drained by ditches and was hot and dry at the time of the visit in 2009 (
|
||
<figureCitation id="71CE2A14D743FFBD4D08AC3680B04C27" box="[1347,1412,349,373]" captionStart="Fig" captionStartId="3.[812,847,2062,2084]" captionTargetBox="[812,1450,1699,2039]" captionTargetId="figure-481@3.[812,1451,1696,2041]" captionTargetPageId="3" captionText="Fig. 5. Fragmented peat swamp locality in Singkil where a remnant population of B. rubra was collected (2009)." figureDoi="http://doi.org/10.5281/zenodo.5351936" httpUri="https://zenodo.org/record/5351936/files/figure.png" pageId="2" pageNumber="325">Fig. 5</figureCitation>
|
||
). The remnant population of
|
||
<taxonomicName id="2EF54D12D743FFBD4C0BAC1481A94CC4" authorityName="Perugia" authorityYear="1893" box="[1088,1181,383,406]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="325" phylum="Chordata" rank="species" species="rubra">
|
||
<emphasis id="DB81EA83D743FFBD4C0BAC1481A94CC4" box="[1088,1181,383,406]" italics="true" pageId="2" pageNumber="325">B. rubra</emphasis>
|
||
</taxonomicName>
|
||
was obtained from a pool of stagnant black water next to an uprooted tree. The pool had no other fish in it. The surface was tepid but the deeper portions cool. Other smaller and shallower water pools were sampled but did not contain any fish. A river nearby was also sampled but did not yield any
|
||
<taxonomicName id="2EF54D12D743FFBD4C8DAF7480164F64" authorityName="Perugia" authorityYear="1893" box="[1222,1314,543,566]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="325" phylum="Chordata" rank="species" species="rubra">
|
||
<emphasis id="DB81EA83D743FFBD4C8DAF7480164F64" box="[1222,1314,543,566]" italics="true" pageId="2" pageNumber="325">B. rubra</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="BD8A6619D743FFBD48C3AA9E84744517" ID-DOI="http://doi.org/10.5281/zenodo.5351928" ID-Zenodo-Dep="5351928" httpUri="https://zenodo.org/record/5351928/files/figure.png" pageId="2" pageNumber="325" startId="2.[136,171,2037,2059]" targetBox="[136,1469,632,2007]" targetPageId="2">
|
||
<paragraph id="E94A3691D743FFBD48C3AA9E84744517" blockId="2.[136,1474,2037,2117]" pageId="2" pageNumber="325">
|
||
Fig. 1. Comparison of lateral head and opercle pattern, left side with photograph, right side with schematic drawing. A, B,
|
||
<taxonomicName id="2EF54D12D743FFBD4D01AA9C85F94575" authority=", MSNG" authorityName="MSNG" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="325" phylum="Chordata" rank="species" species="rubra">
|
||
<emphasis id="DB81EA83D743FFBD4D01AA9C80884559" box="[1354,1468,2039,2059]" italics="true" pageId="2" pageNumber="325">Betta rubra</emphasis>
|
||
, MSNG
|
||
</taxonomicName>
|
||
13019, lectotype, 28.6 mm SL; C, D,
|
||
<taxonomicName id="2EF54D12D743FFBD4A09A57887DA4575" authority=", ZRC" authorityName="ZRC" box="[578,750,2066,2088]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="325" phylum="Chordata" rank="species" species="rubra">
|
||
<emphasis id="DB81EA83D743FFBD4A09A57887874575" box="[578,691,2067,2087]" italics="true" pageId="2" pageNumber="325">Betta rubra</emphasis>
|
||
, ZRC
|
||
</taxonomicName>
|
||
53989, 34.0 mm SL male; E, F,
|
||
<taxonomicName id="2EF54D12D743FFBD4C60A578802D457A" authority=", MZB" authorityName="MZB" box="[1067,1305,2067,2088]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="325" phylum="Chordata" rank="species" species="dennisyongi">
|
||
<emphasis id="DB81EA83D743FFBD4C60A57881EC4575" box="[1067,1240,2067,2087]" italics="true" pageId="2" pageNumber="325">Betta dennisyongi</emphasis>
|
||
, MZB
|
||
</taxonomicName>
|
||
17207, holotype, 27.4 mm SL male.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="A1EF651AD742FFBC483AADD787AE4C07" pageId="3" pageNumber="326" type="distribution">
|
||
<paragraph id="E94A3691D742FFBC483AADD787C44D87" blockId="3.[113,752,191,341]" box="[113,752,188,214]" pageId="3" pageNumber="326">
|
||
<emphasis id="DB81EA83D742FFBC483AADD785CA4D87" bold="true" box="[113,254,188,213]" pageId="3" pageNumber="326">
|
||
<emphasis id="DB81EA83D742FFBC483AADD785C24D87" bold="true" box="[113,246,188,213]" italics="true" pageId="3" pageNumber="326">Distribution</emphasis>
|
||
.
|
||
</emphasis>
|
||
—
|
||
<taxonomicName id="2EF54D12D742FFBC496EADD484954D84" box="[293,417,191,214]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="326" phylum="Chordata" rank="species" species="rubra">
|
||
<emphasis id="DB81EA83D742FFBC496EADD484954D84" box="[293,417,191,214]" italics="true" pageId="3" pageNumber="326">Betta rubra</emphasis>
|
||
</taxonomicName>
|
||
is currently known definitively
|
||
</paragraph>
|
||
<paragraph id="E94A3691D742FFBC483AADB687AE4C07" blockId="3.[113,752,191,341]" pageId="3" pageNumber="326">
|
||
from the peat swamps in the Singkil area in
|
||
<collectingRegion id="2B31F873D742FFBC4AE1ADB687DE4DA7" box="[682,746,221,245]" country="Indonesia" name="Aceh" pageId="3" pageNumber="326">Aceh</collectingRegion>
|
||
; possibly Lake Toba (though possibly erroneous; see earlier Introduction) area near Medan in
|
||
<collectingRegion id="2B31F873D742FFBC4A42AC76875C4C67" box="[521,616,285,309]" country="Indonesia" name="Sumatera Utara" pageId="3" pageNumber="326">Sumatra</collectingRegion>
|
||
Utara; and Sibolga in northern part of
|
||
<collectingRegion id="2B31F873D742FFBC49D6AC56870C4C07" box="[413,568,317,341]" country="Indonesia" name="Sumatera Barat" pageId="3" pageNumber="326">Sumatra Barat</collectingRegion>
|
||
(
|
||
<figureCitation id="71CE2A14D742FFBC4A02AC5687B94C07" box="[585,653,317,341]" captionStart="Fig" captionStartId="4.[137,172,2060,2082]" captionTargetBox="[137,776,1164,2039]" captionTargetId="figure-8@4.[137,776,1164,2039]" captionTargetPageId="4" captionText="Fig. 6. Map of Sumatra showing distribution of Betta rubra (round symbols) and B. dennisyongi (square symbols)." figureDoi="http://doi.org/10.5281/zenodo.5351940" httpUri="https://zenodo.org/record/5351940/files/figure.png" pageId="3" pageNumber="326">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A1EF651AD742FFBC483AAC1787DE4F67" pageId="3" pageNumber="326" type="discussion">
|
||
<paragraph id="E94A3691D742FFBC483AAC1787DE4F67" blockId="3.[113,752,380,565]" pageId="3" pageNumber="326">
|
||
<emphasis id="DB81EA83D742FFBC483AAC1785EE4CC7" bold="true" box="[113,218,380,405]" pageId="3" pageNumber="326">
|
||
<emphasis id="DB81EA83D742FFBC483AAC1785E74CC7" bold="true" box="[113,211,380,405]" italics="true" pageId="3" pageNumber="326">Remarks</emphasis>
|
||
.
|
||
</emphasis>
|
||
— See discussion in Tan & Ng (2005a, 2005b) for details of a 1982 series (incorrectly referred to as
|
||
<taxonomicName id="2EF54D12D742FFBC4AC7ACF487D34CE4" box="[652,743,415,438]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="326" phylum="Chordata" rank="species" species="rubra">
|
||
<emphasis id="DB81EA83D742FFBC4AC7ACF487D34CE4" box="[652,743,415,438]" italics="true" pageId="3" pageNumber="326">B. rubra</emphasis>
|
||
</taxonomicName>
|
||
) from MZB which belongs to the new species described herein; and the apparent reddish tinge of the original
|
||
<typeStatus id="364E8833D742FFBC4A89ACB587C44CA4" box="[706,752,478,502]" pageId="3" pageNumber="326">type</typeStatus>
|
||
series. This species has been recorded to be a paternal oralbrooder by aquarist reports and personal observations.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |