treatments-xml/data/60/AD/56/60AD56B2B2AC50299E2114539DBDEE9B.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1147.93824" ID-Pensoft-Pub="1313-2970-1147-1" ID-Pensoft-UUID="FF2B6B317B505F9EA0E1000BDCD16CE7" ID-ZooBank="9BB9881B0076473D8E53155D37CA1F50" ModsDocID="1313-2970-1147-1" checkinTime="1676378625502" checkinUser="pensoft" docAuthor="Pall-Gergely, Barna, Hunyadi, Andras, Vermeulen, Jaap J., Grego, Jozef, Sutcharit, Chirasak, Reischuetz, Alexander, Dumrongrojwattana, Pongrat, Botta-Dukat, Zoltan, Oerstan, Aydin, Fekete, Judit &amp; Jochum, Adrienne" docDate="2023" docId="60AD56B2B2AC50299E2114539DBDEE9B" docLanguage="en" docName="ZooKeys 1147: 1-177" docOrigin="ZooKeys 1147" docPubDate="2023-02-13" docSource="http://dx.doi.org/10.3897/zookeys.1147.93824" docTitle="Angustopila parallela Pall-Gergely &amp; Hunyadi 2023, sp. nov." docType="treatment" docUuid="8A123D0E-0E19-4834-A72C-EF96DD247846" docUuidSource="ZooBank" docVersion="1" id="FF2B6B317B505F9EA0E1000BDCD16CE7" lastPageNumber="1" masterDocId="FF2B6B317B505F9EA0E1000BDCD16CE7" masterDocTitle="Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae)" masterLastPageNumber="177" masterPageNumber="1" pageNumber="1" updateTime="1676378625502" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pall-Gergely, Barna</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6167-7221</mods:nameIdentifier>
<mods:affiliation>Centre for Agricultural Research, Plant Protection Institute, Eoetvoes Lorand Research Network, Herman Otto ut 15, H- 1022 Budapest, Hungary</mods:affiliation>
<mods:nameIdentifier type="email">pallgergely2@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hunyadi, Andras</mods:namePart>
<mods:affiliation>Adria setany 10 G 2 / 5., H- 1148 Budapest, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vermeulen, Jaap J.</mods:namePart>
<mods:affiliation>JK Art and Science, Lauwerbes 8, 2318 AT Leiden, Netherlands</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Grego, Jozef</mods:namePart>
<mods:affiliation>Horna Micina 219, SK- 97401 Banska Bystrica, Slovakia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sutcharit, Chirasak</mods:namePart>
<mods:affiliation>Animal Systematic Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Reischuetz, Alexander</mods:namePart>
<mods:affiliation>Puechhaimgasse 52, A- 3580 Horn, Austria</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dumrongrojwattana, Pongrat</mods:namePart>
<mods:affiliation>Department of Biology, Faculty of Science, Burapha University, 169 Longhardbangsaen Road, Muang District, Chonburi, 20131, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Botta-Dukat, Zoltan</mods:namePart>
<mods:affiliation>Centre for Ecological Research, Institute of Ecology and Botany, Alkotmany 2 - 4, H- 2600, Vacratot, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Oerstan, Aydin</mods:namePart>
<mods:affiliation>12501 Milestone Manor Lane, Germantown, Maryland, 20876, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fekete, Judit</mods:namePart>
<mods:affiliation>University of Pannonia, Centre of Natural Science, Research Group of Limnology, Egyetem u. 10, H- 8200 Veszprem, Hungary &amp; Centre for Ecological Research, Institute of Aquatic Ecology, Department of Tisza Research, 18 / c Bem square, H- 4026 Debrecen, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jochum, Adrienne</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6624-6412</mods:nameIdentifier>
<mods:affiliation>Naturhistorisches Museum der Burgergemeinde Bern, CH- 3005 Bern, Switzerland &amp; Institute of Ecology and Evolution, University of Bern, CH- 3012 Bern, Switzerland &amp; Senckenberg Forschungsinstitut und Naturmuseum, 60325 Frankfurt am Main, Germany</mods:affiliation>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-02-13</mods:number>
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<mods:detail type="volume">
<mods:number>1147</mods:number>
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<mods:start>1</mods:start>
<mods:end>177</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1147.93824</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1147.93824</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1147-1</mods:identifier>
<mods:identifier type="ZooBank">9BB9881B0076473D8E53155D37CA1F50</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:8A123D0E-0E19-4834-A72C-EF96DD247846" httpUri="http://treatment.plazi.org/id/60AD56B2B2AC50299E2114539DBDEE9B" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="https://zoobank.org/8A123D0E-0E19-4834-A72C-EF96DD247846" authority="Páll-Gergely &amp; Hunyadi" authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Angustopila parallela" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="parallela" status="sp. nov.">
Angustopila parallela 
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; Hunyadi
</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 74" captionStartId="F74" captionText="Figure 74. Angustopila parallela Pall-Gergely &amp; Hunyadi, sp. nov. A, C-H holotype (HNHM 105298) B paratype. (2020 / 57). Apertural (A, B), lateral (C), apical (D) and ventral (E) sides of the shell; sculpture on the protoconch (F), ventral (G) and frontal (H) surface of the body whorl." figureDoi="10.3897/zookeys.1147.93824.figure74" httpUri="https://binary.pensoft.net/fig/808883" pageId="0" pageNumber="1">Figs 74</figureCitation>
<figureCitation captionStart="Figure 75" captionStartId="F75" captionText="Figure 75. Variability of Angustopila parallela Pall-Gergely &amp; Hunyadi, sp. nov. from the type locality A, B holotype C paratype." figureDoi="10.3897/zookeys.1147.93824.figure75" httpUri="https://binary.pensoft.net/fig/808884" pageId="0" pageNumber="1">, 75</figureCitation>
<figureCitation captionStart="Figure 76" captionStartId="F76" captionText="Figure 76. Angustopila parallela Pall-Gergely &amp; Hunyadi, sp. nov. (2020 / 41). Apertural (A), lateral (B), apical (C) and ventral (D) sides of the shell; sculpture on the protoconch with protoconch-teleoconch boundary (E); aperture (F); ventral (G) and frontal (H) surface of the body whorl." figureDoi="10.3897/zookeys.1147.93824.figure76" httpUri="https://binary.pensoft.net/fig/808885" pageId="0" pageNumber="1">, 76</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
: Vietnam • 1 empty shell (H: 0.97 mm, D: 1.08 mm); Lạng 
<normalizedToken originalValue="Sơn">Sơn</normalizedToken>
Province, Tam Thanh, 
<normalizedToken originalValue="Núi">Nui</normalizedToken>
Vọng Phu, 
<normalizedToken originalValue="Đ">Đ</normalizedToken>
ền 
<normalizedToken originalValue="Chùa">Chua</normalizedToken>
<normalizedToken originalValue="Thư">Thư</normalizedToken>
ợng 
<normalizedToken originalValue="Ngàn">Ngan</normalizedToken>
(locality code: 2020/57); 
<geoCoordinate degrees="21" direction="north" minutes="51.21" orientation="latitude" precision="9" value="21.8535">21°51.21'N</geoCoordinate>
, 
<geoCoordinate degrees="106" direction="east" minutes="44.91" orientation="longitude" precision="9" value="106.7485">106°44.91'E</geoCoordinate>
; 250 m a.s.l.; 21 Feb. 2020; A. Hunyadi leg.; HNHM 105298.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
: Vietnam • 58 shells; same data as for holotype; coll. HA • 569 shells; Lạng 
<normalizedToken originalValue="Sơn">Sơn</normalizedToken>
Province, vicinity of 
<normalizedToken originalValue="Chùa">Chua</normalizedToken>
Tam Thanh (locality code: 2020/58); 
<geoCoordinate degrees="21" direction="north" minutes="51.35" orientation="latitude" precision="9" value="21.855833">21°51.35'N</geoCoordinate>
, 
<geoCoordinate degrees="106" direction="east" minutes="44.81" orientation="longitude" precision="9" value="106.74683">106°44.81'E</geoCoordinate>
; 265 m a.s.l.; 21 Feb. 2020; A. Hunyadi leg.; coll. HA • 10 shells; same data as for preceding; coll. JJV.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Additional material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="2020-02-14" collectingDateMax="2020-02-21" collectingDateMin="2020-02-14" collectorName="A. Hunyadi, Thanh District, H" country="Vietnam" county="Province" elevation="265" latitude="19.618" location="Thanh District" longLatPrecision="12" longitude="105.57716" municipality="Chua Tam Thanh" specimenCount="1" stateProvince="Thanh Hoa Province">
<collectingCountry name="Vietnam">Vietnam</collectingCountry>
• 7 j/b shells; same data as for holotype; coll. HA • 115 j/b shells; Lạng 
<normalizedToken originalValue="Sơn">Sơn</normalizedToken>
<collectingCounty>Province</collectingCounty>
, vicinity of 
<collectingMunicipality>
<normalizedToken originalValue="Chùa">Chua</normalizedToken>
Tam Thanh
</collectingMunicipality>
(locality code: 2020/58); 
<geoCoordinate degrees="21" direction="north" minutes="51.35" orientation="latitude" precision="9" value="21.855833">21°51.35'N</geoCoordinate>
, 
<geoCoordinate degrees="106" direction="east" minutes="44.81" orientation="longitude" precision="9" value="106.74683">106°44.81'E</geoCoordinate>
; 
<elevation metricMagnitude="2" metricUnit="m" metricValue="2.65" unit="m" value="265.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="2.65" unit="m" value="265.0">265 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate value="2020-02-21">21 Feb. 2020</collectingDate>
; 
<collectorName>A. Hunyadi</collectorName>
leg.; coll. HA • 1 figured shell (Fig. 
<figureCitation captionStart="Figure 76" captionStartId="F76" captionText="Figure 76. Angustopila parallela Pall-Gergely &amp; Hunyadi, sp. nov. (2020 / 41). Apertural (A), lateral (B), apical (C) and ventral (D) sides of the shell; sculpture on the protoconch with protoconch-teleoconch boundary (E); aperture (F); ventral (G) and frontal (H) surface of the body whorl." figureDoi="10.3897/zookeys.1147.93824.figure76" httpUri="https://binary.pensoft.net/fig/808885" pageId="0" pageNumber="1">76</figureCitation>
); 
<collectingRegion country="Vietnam" name="Thanh Hoa">
Thanh 
<normalizedToken originalValue="Hóa">Hoa</normalizedToken>
Province
</collectingRegion>
, 
<normalizedToken originalValue="Như">Như</normalizedToken>
<collectorName>
<location LSID="urn:lsid:plazi:treatment:60AD56B2B2AC50299E2114539DBDEE9B:18113168C446132E895F99CEEF97F152" country="Vietnam" county="Province" latitude="19.618" longLatPrecision="12" longitude="105.57716" municipality="Chua Tam Thanh" name="Thanh District" stateProvince="Thanh Hoa Province">Thanh District</location>
, H
</collectorName>
ải 
<normalizedToken originalValue="Vân">Van</normalizedToken>
, Hang 
<normalizedToken originalValue="Lò">Lo</normalizedToken>
Cao 
<normalizedToken originalValue="Kháng">Khang</normalizedToken>
Chiến, vicinity of the cave (locality code: 2020/41); 
<geoCoordinate degrees="19" direction="north" minutes="37.08" orientation="latitude" precision="9" value="19.618">19°37.08'N</geoCoordinate>
, 
<geoCoordinate degrees="105" direction="east" minutes="34.63" orientation="longitude" precision="9" value="105.57716">105°34.63'E</geoCoordinate>
; 
<elevation metricMagnitude="1" metricUnit="m" metricValue="2.0" unit="m" value="20.0">
<quantity metricMagnitude="1" metricUnit="m" metricValue="2.0" unit="m" value="20.0">20 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate value="2020-02-14">14 Feb. 2020</collectingDate>
; 
<collectorName>A. Hunyadi</collectorName>
leg.; HNHM 105299 • 7 shells; same data as for preceding; coll. HA
</materialsCitation>
.
</paragraph>
<caption doi="10.3897/zookeys.1147.93824.figure74" httpUri="https://binary.pensoft.net/fig/808883" pageId="0" pageNumber="1" start="Figure 74" startId="F74">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 74.</emphasis>
<taxonomicName authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila parallela" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="parallela">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila parallela</emphasis>
</taxonomicName>
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; Hunyadi, sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="1">A, C-H</emphasis>
<typeStatus>holotype</typeStatus>
(HNHM 105298) 
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
<typeStatus>paratype</typeStatus>
. (2020/57). Apertural (
<emphasis bold="true" pageId="0" pageNumber="1">A, B</emphasis>
), lateral (
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
), apical (
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
) and ventral (
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
) sides of the shell; sculpture on the protoconch (
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
), ventral (
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
) and frontal (
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
) surface of the body whorl.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1147.93824.figure75" httpUri="https://binary.pensoft.net/fig/808884" pageId="0" pageNumber="1" start="Figure 75" startId="F75">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 75.</emphasis>
Variability of 
<taxonomicName authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila parallela" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="parallela">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila parallela</emphasis>
</taxonomicName>
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; Hunyadi, sp. nov. from the type locality 
<emphasis bold="true" pageId="0" pageNumber="1">A, B</emphasis>
<typeStatus>holotype</typeStatus>
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
<typeStatus>paratype</typeStatus>
.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1147.93824.figure76" httpUri="https://binary.pensoft.net/fig/808885" pageId="0" pageNumber="1" start="Figure 76" startId="F76">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 76.</emphasis>
<taxonomicName authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila parallela" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="parallela">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila parallela</emphasis>
</taxonomicName>
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; Hunyadi, sp. nov. (2020/41). Apertural (
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
), lateral (
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
), apical (
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
) and ventral (
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
) sides of the shell; sculpture on the protoconch with protoconch-teleoconch boundary (
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
); aperture (
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
); ventral (
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
) and frontal (
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
) surface of the body whorl.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
A large, low or high concave-conical 
<taxonomicName authorityName="Jochum, Slapnik &amp; Pall-Gergely" authorityYear="2014" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila</emphasis>
</taxonomicName>
species with a wide umbilicus, dense spiral striation, and two strong apertural barriers (elevated parietal, elongated or pointed lower palatal).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">Shell large for the genus, wider than high or rarely slightly higher than wide; off-white, concave-conical, shell height highly variable from strongly depressed to triangular, with regularly growing whorls except for the last one; last whorl widest from apertural view; protoconch consists of 1.25-1.5 whorls, spirally striated with the exception of the first whorl; teleoconch with strong, rather dense, irregular radial ribs and equidistantly-spaced, dense spiral striae (ca. 19-21 on body whorl from apertural view); whorls 4-4.25, rounded with tendency towards shouldered contour (in apertural view, the body whorl is slightly pushed from baso-lateral direction); aperture oblique to shell axis in lateral view; umbilicus wide; aperture suboval; peristome expanded, not reflected; parietal callus not detached from penultimate whorl; parietal tooth elevated, straight, starts in some distance from peristome; lower palatal tooth runs parallel with the peristome on the lower 1/2 of the palatal region, in some specimens pointed, in others ridge-like, situated inside from peristome.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Measurements (in mm).</paragraph>
<paragraph pageId="0" pageNumber="1">
H = 0.85-1.12, D: 1.04-1.14, H/D*100 = 77.3-109.6 (
<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
= 18), RUD = 29.0-33.3 (
<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
= 5).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila majuscula" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="majuscula">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila majuscula</emphasis>
</taxonomicName>
sp. nov. is similar to this species by the large shell size, wide umbilicus and the possession of two apertural denticles. However, it has a convex conical shell shape, denser spiral striation and the palatal tooth is situated further up in the aperture. See also under 
<taxonomicName lsidName="A. pustulata" pageId="0" pageNumber="1" rank="species" species="pustulata">
<emphasis italics="true" pageId="0" pageNumber="1">A. pustulata</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">Named after the palatal tooth that runs parallel with the peristome.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila parallela" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="parallela">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila parallela</emphasis>
</taxonomicName>
sp. nov. is known from two localities in northern Vietnam (Lạng 
<normalizedToken originalValue="Sơn">Sơn</normalizedToken>
and Thanh 
<normalizedToken originalValue="Hóa">Hoa</normalizedToken>
Provinces) (Fig. 
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Distribution of Angustopila species." figureDoi="10.3897/zookeys.1147.93824.figure19" httpUri="https://binary.pensoft.net/fig/808828" pageId="0" pageNumber="1">19</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">Shells from population 2020/57 are generally more depressed than those of 2020/58, although the two groups overlap. The distance between these populations is ca. 300 m (!) in a straight line. Most shells of population 2020/41 possess an elongated palatal tooth, whereas that tooth is pointed in most shells of the other two populations. However, this difference is not exclusive, because there are individuals with a pointed palatal tooth in population 2020/41 and a more elongated tooth in the other two populations. Thus, even if the geographic distance is relatively large overall (ca. 280 km in a straight line), the morphological differences are insufficient to distinguish the population 2020/41 from the other two.</paragraph>
</subSubSection>
</treatment>
</document>