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<document id="C74B130B073EAEE8A3545264C4D3F187" ID-CLB-Dataset="31229" ID-DOI="10.11646/zootaxa.4362.3.2" ID-GBIF-Dataset="375c7e7b-e311-47c3-9599-f0e16d227889" ID-ISSN="1175-5326" ID-Zenodo-Dep="1094896" ID-ZooBank="852AFFC6-42E7-4C52-A5B9-5FFA4E6759CC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1512548824660" checkinUser="plazi" docAuthor="Bianchi, Filipe Michels, Barão, Kim Ribeiro &amp; Grazia, Jocelia" docDate="2017" docId="6033878DF941CC430BD9FA912C54C712" docLanguage="en" docName="zootaxa.4362.3.2.pdf" docOrigin="Zootaxa 4362 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Euschistus (Euschistus) tacitus Bianchi &amp; Barão &amp; Grazia 2017, sp. nov." docType="treatment" docVersion="10" lastPageNumber="357" masterDocId="9C0AFFF5F949CC4A0B4EFF9B2868C222" masterDocTitle="Review of the sulcacitus group of Euschistus (Pentatomidae: Pentatominae: Carpocorini) with description of the internal female genitalia and a new species" masterLastPageNumber="358" masterPageNumber="348" pageNumber="356" updateTime="1698507761827" updateUser="ExternalLinkService">
<mods:mods id="8B3350F5C9A7CFB0B26B8BD6DC2FD550" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B46A4D73EFD7A3CCA499925F6A079ACF">
<mods:title id="040F8A50741168720BF2AA33755F03D9">Review of the sulcacitus group of Euschistus (Pentatomidae: Pentatominae: Carpocorini) with description of the internal female genitalia and a new species</mods:title>
</mods:titleInfo>
<mods:name id="55348DE788EED89CA76E35D92F79E154" type="personal">
<mods:role id="248AE13FF9E7E90C949126F4D6B27620">
<mods:roleTerm id="F0086793CE9C11DE8EB6111BA223CEC3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="67B0E21E0719782C5DD1BDBE75CE99B7">Bianchi, Filipe Michels</mods:namePart>
<mods:affiliation id="A3E52D4C14992EDD45A03E5F807DF45B">Programa de Pós-Graduação em Biologia Animal, Universidade Federal do Rio Grande do Sul, Porto Alegre, RS, Brazil.</mods:affiliation>
<mods:nameIdentifier id="F8F286B3E773F1B49DA0209B2DA650E1" type="email">fmichels2@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="8AB5F581083D3227B95A95C706EF11D5" type="personal">
<mods:role id="768FC12B374B099DEFB516243A32B7F7">
<mods:roleTerm id="450DBC937B18A915DF7845EF7A128793">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="57EC0E035B422A5C062EA8C4D8815A9B">Barão, Kim Ribeiro</mods:namePart>
<mods:affiliation id="3464068D431F3B4E5E05FE270BCEED11">Unidade Educacional Penedo, Campus Arapiraca, Universidade Federal de Alagoas, Penedo, AL, Brazil</mods:affiliation>
<mods:nameIdentifier id="DCE48DD5AC1CF7CAFE66F825481951A8" type="email">kim.barao@penedo.ufal.br</mods:nameIdentifier>
</mods:name>
<mods:name id="E8B1A13EBD6A6B54AA595B5678426494" type="personal">
<mods:role id="AB03680483A99F91CEFA7DC32AB002EC">
<mods:roleTerm id="869267BFC08CD114B38F8873F789852A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7DC2775297C060238400D59B7F377B9E">Grazia, Jocelia</mods:namePart>
<mods:affiliation id="2AD128540B74577B6DC243B6614DF3E1">Programa de Pós-Graduação em Biologia Animal, Universidade Federal do Rio Grande do Sul, Porto Alegre, RS, Brazil.</mods:affiliation>
<mods:nameIdentifier id="8EAD81555056EF2A3E7B5C0AB5349E9F" type="email">jocelia@ufrgs.br</mods:nameIdentifier>
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<mods:typeOfResource id="906E0DB67DEF460BD2F0B9599F2C054A">text</mods:typeOfResource>
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<mods:title id="A83E92FF534CAEBAEB6B76EBFD39833B">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="C003178C7B0A9E44C90C6AFE95DCDAE2">
<mods:date id="F7291B9FEDD36611142ECE70A83A9FD4">2017</mods:date>
<mods:detail id="B37396C93000586D477219A5D50B8AFE" type="pubDate">
<mods:number id="B8CB6C13A6D7F301D44C7A5B87B15394">2017-12-06</mods:number>
</mods:detail>
<mods:detail id="A862EAB6A535D306557E850551B24281" type="volume">
<mods:number id="7BD8633E8D9CA544DAEBA268622B6D23">4362</mods:number>
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<mods:number id="6D173454588E786303F795B2B9EDFD5F">3</mods:number>
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<mods:classification id="6A7E27D3D946EC5243230F6D0F3BE7C5">journal article</mods:classification>
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<mods:identifier id="85C1D30438A885736E9E182031800BF7" type="DOI">10.11646/zootaxa.4362.3.2</mods:identifier>
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<mods:identifier id="18DCCB91F20FD7C756F1E1F22379D2F1" type="ISSN">1175-5326</mods:identifier>
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<treatment id="6033878DF941CC430BD9FA912C54C712" ID-DOI="http://doi.org/10.5281/zenodo.5669925" ID-GBIF-Taxon="137627673" ID-Zenodo-Dep="5669925" LSID="urn:lsid:plazi:treatment:6033878DF941CC430BD9FA912C54C712" httpUri="http://treatment.plazi.org/id/6033878DF941CC430BD9FA912C54C712" lastPageId="9" lastPageNumber="357" pageId="8" pageNumber="356">
<subSubSection id="A0806510F941CC420BD9FA912B32C707" box="[151,858,1290,1317]" pageId="8" pageNumber="356" type="nomenclature">
<paragraph id="E825369BF941CC420BD9FA912B32C707" blockId="8.[151,858,1290,1350]" box="[151,858,1290,1317]" pageId="8" pageNumber="356">
<heading id="B36D81F7F941CC420BD9FA912B32C707" bold="true" box="[151,858,1290,1317]" fontSize="11" level="1" pageId="8" pageNumber="356" reason="1">
<emphasis id="DAEEEA89F941CC420BD9FA912B32C707" bold="true" box="[151,858,1290,1317]" pageId="8" pageNumber="356">
<taxonomicName id="2F9A4D18F941CC420BD9FA912A9DC707" authority="Bianchi &amp; Grazia" authorityName="Bianchi &amp; Barão &amp; Grazia" authorityYear="2017" box="[151,757,1290,1317]" class="Insecta" family="Pentatomidae" genus="Euschistus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="356" phylum="Arthropoda" rank="species" species="tacitus" status="sp. nov." subGenus="Euschistus">
<emphasis id="DAEEEA89F941CC420BD9FA912973C706" bold="true" box="[151,283,1290,1316]" italics="true" pageId="8" pageNumber="356">Euschistus</emphasis>
(
<emphasis id="DAEEEA89F941CC420A62FA9129D8C706" bold="true" box="[300,432,1290,1316]" italics="true" pageId="8" pageNumber="356">Euschistus</emphasis>
)
<emphasis id="DAEEEA89F941CC420A8FFA912A78C706" bold="true" box="[449,528,1290,1316]" italics="true" pageId="8" pageNumber="356">tacitus</emphasis>
Bianchi &amp; Grazia
</taxonomicName>
<taxonomicNameLabel id="C1DD57F2F941CC4209B2FA902B32C707" box="[764,858,1291,1317]" pageId="8" pageNumber="356" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A0806510F941CC420BD9FAB62A57C764" box="[151,575,1325,1350]" pageId="8" pageNumber="356" type="description">
<paragraph id="E825369BF941CC420BD9FAB62A57C764" blockId="8.[151,858,1290,1350]" box="[151,575,1325,1350]" pageId="8" pageNumber="356">
(
<figureCitation id="70A12A1EF941CC420BD1FAB6288FC764" box="[159,231,1325,1350]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1938,1961]" captionTargetBox="[160,1421,1606,1917]" captionTargetId="figure@2.[151,1436,1599,1917]" captionTargetPageId="2" captionText="FIGURES 1–5. Dorsal view of representative specimens of sulcacitus group species. 1, Euschistus (Euschistus) corcoVacitus; 2, Euschistus (Euschistus) olacitus; 3, Euschistus (Euschistus) palacitus; 4, Euschistus (Euschistus) sulcacitus; 5, Euschistus (Euschistus) tacitus sp. nov., holotype. Scale bars: 2 mm." httpUri="https://zenodo.org/record/1094898/files/figure.png" pageId="8" pageNumber="356">Figs 5</figureCitation>
,
<figureCitation id="70A12A1EF941CC420BBDFAB52954C764" box="[243,316,1326,1350]" captionStart="FIGURES 6–20" captionStartId="3.[151,264,1870,1893]" captionTargetBox="[188,1393,193,1826]" captionTargetId="figure@3.[188,1393,193,1840]" captionTargetPageId="3" captionText="FIGURES 6–20. Dissected and macerated pygophore of species of sulcacitus group. Pygophore in dorsal (6, 9, 12, 15, 18), posterior (7, 10, 13, 16, 19), and ventral (8, 11, 14, 17, 20) views. 6–8, Euschistus (Euschistus) corcoVacitus; 9–11, Euschistus (Euschistus) olacitus; 12–14, Euschistus (Euschistus) palacitus; 15–17, Euschistus (Euschistus) sulcacitus; 18–20, Euschistus (Euschistus) tacitus sp. nov.. dr, dorsal rim; mp, median projection; par, paramere; slvr, superior layer of ventral rim; spdr, superior process of dorsal rim; vr, ventral rim. Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1094900/files/figure.png" pageId="8" pageNumber="356">18–20</figureCitation>
,
<figureCitation id="70A12A1EF941CC420A09FAB6290DC764" box="[327,357,1325,1350]" captionStart="FIGURES 21–26" captionStartId="4.[151,264,504,527]" captionTargetBox="[152,1421,194,484]" captionTargetId="figure@4.[151,1436,193,484]" captionTargetPageId="4" captionText="FIGURES 21–26. Right paramere of species of sulcacitus group. 21, schematic drawing of paramere indicating its regionalization; 22, Euschistus (Euschistus) corcoVacitus; 23, Euschistus (Euschistus) olacitus; 24, Euschistus (Euschistus) palacitus; 25, Euschistus (Euschistus) sulcacitus; 26, Euschistus (Euschistus) tacitus sp. nov. app, apical process of paramere; bpp, basal process of paramere; cr, crown; st, stem. Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1094902/files/figure.png" pageId="8" pageNumber="356">26</figureCitation>
,
<figureCitation id="70A12A1EF941CC420A3FFAB529D2C764" box="[369,442,1326,1350]" captionStart="FIGURES 27–41" captionStartId="4.[151,264,1748,1771]" captionTargetBox="[151,1421,668,1727]" captionTargetId="figure@4.[151,1436,668,1727]" captionTargetPageId="4" captionText="FIGURES 27–41. Phallus of species of sulcacitus group. Phallus in dorsal (27, 30, 33, 36, 39), ventral (28, 31, 34, 37, 40) and lateral (29, 32, 35, 38, 41) views. 27–29, Euschistus (Euschistus) corcoVacitus; 30–32, Euschistus (Euschistus) olacitus; 33–35, Euschistus (Euschistus) palacitus; 36–38, Euschistus (Euschistus) sulcacitus; 39–41, Euschistus (Euschistus) tacitus sp. nov. Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1094904/files/figure.png" pageId="8" pageNumber="356">39–41</figureCitation>
,
<figureCitation id="70A12A1EF941CC420A8BFAB62A66C764" box="[453,526,1325,1350]" captionStart="FIGURES 45–59" captionStartId="6.[151,264,1803,1826]" captionTargetBox="[151,1436,193,1782]" captionTargetId="figure@6.[151,1436,193,1782]" captionTargetPageId="6" captionText="FIGURES 45–59. Female genital plates of species of sulcacitus group. Genital plates in posterior (45, 48, 51, 54, 57) and postero-ventral (45, 46, 49, 50, 52, 53, 55, 56, 58, 59) views. 45–47, Euschistus (Euschistus) corcoVacitus; 48–50, Euschistus (Euschistus) olacitus; 51–53, Euschistus (Euschistus) palacitus; 54–56, Euschistus (Euschistus) sulcacitus; 57–59, Euschistus (Euschistus) tacitus sp. nov.. gc, gonocoxites; la, laterotergites. Scale bars: 1 mm." httpUri="https://zenodo.org/record/1094908/files/figure.png" pageId="8" pageNumber="356">57–59</figureCitation>
,
<figureCitation id="70A12A1EF941CC420957FAB62A5EC764" box="[537,566,1325,1350]" captionStart="FIGURES 60–64" captionStartId="7.[151,264,1794,1817]" captionTargetBox="[151,1436,193,1773]" captionTargetId="figure@7.[151,1436,193,1773]" captionTargetPageId="7" captionText="FIGURES 60–64. Female internal genitalia of species of sulcacitus group. 60, Euschistus (Euschistus) corcoVacitus; 61, Euschistus (Euschistus) olacitus; 62, Euschistus (Euschistus) palacitus; 63, Euschistus (Euschistus) sulcacitus; 64, Euschistus (Euschistus) tacitus sp. nov.. Insets are the capsula seminalis. aaf, anterior anular flange; cs, capsula seminalis; dr, ductus receptaculi; g, gonapophysis; gc, gonocoxites; la, laterotergites; paf, posterior anular flange; pi, pars intermedialis; rs, ring sclerites; tvi, thickening of vaginal intima; va, vesicular area. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/1094910/files/figure.png" pageId="8" pageNumber="356">64</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="A0806510F941CC420BD9FAEE298AC7F4" pageId="8" pageNumber="356" type="etymology">
<paragraph id="E825369BF941CC420BD9FAEE298AC7F4" blockId="8.[151,1437,1397,2034]" pageId="8" pageNumber="356">
<emphasis id="DAEEEA89F941CC420BD9FAEE2949C7AC" bold="true" box="[151,289,1397,1422]" pageId="8" pageNumber="356">Etymology:</emphasis>
Epithet alludes to the external morphological similarities among the species of the
<taxonomicName id="2F9A4D18F941CC420FACFAEE2D3AC7AF" authorityName="Rolston. Rolston" authorityYear="1971" box="[1250,1362,1397,1421]" class="Insecta" family="Pentatomidae" genus="Euschistus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="356" phylum="Arthropoda" rank="species" species="sulcacitus">
<emphasis id="DAEEEA89F941CC420FACFAEE2D3AC7AF" box="[1250,1362,1397,1421]" italics="true" pageId="8" pageNumber="356">sulcacitus</emphasis>
</taxonomicName>
group and the new species. From
<emphasis id="DAEEEA89F941CC420A92FA012A71C793" box="[476,537,1434,1457]" italics="true" pageId="8" pageNumber="356">taceo</emphasis>
(Latin), to be silent, say nothing;
<taxonomicName id="2F9A4D18F941CC4208FCFA012B95C793" authorityName="Bianchi &amp; Barão &amp; Grazia" authorityYear="2017" box="[946,1021,1434,1457]" class="Insecta" family="Pentatomidae" genus="Euschistus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="356" phylum="Arthropoda" rank="species" species="tacitus">
<emphasis id="DAEEEA89F941CC4208FCFA012B95C793" box="[946,1021,1434,1457]" italics="true" pageId="8" pageNumber="356">tacitus</emphasis>
</taxonomicName>
, meaning implied, unmentioned, in reference to a hidden species.
</paragraph>
</subSubSection>
<subSubSection id="A0806510F941CC420B89FA7A2C30C5D0" pageId="8" pageNumber="356" type="distribution">
<paragraph id="E825369BF941CC420B89FA7A2BC5C7D8" blockId="8.[151,1437,1397,2034]" box="[199,941,1505,1530]" pageId="8" pageNumber="356">
<emphasis id="DAEEEA89F941CC420B89FA7A290AC7D8" bold="true" box="[199,354,1505,1530]" pageId="8" pageNumber="356">Distribution:</emphasis>
Mexico (Chiapas, Veracruz); Costa Rica (Alajuela).
</paragraph>
<paragraph id="E825369BF941CC420B89F99E2AF4C43C" blockId="8.[151,1437,1397,2034]" box="[199,668,1541,1566]" pageId="8" pageNumber="356">
<emphasis id="DAEEEA89F941CC420B89F99E2901C43C" bold="true" box="[199,361,1541,1566]" pageId="8" pageNumber="356">
<typeStatus id="37218839F941CC420B89F99E296AC43C" box="[199,258,1541,1566]" pageId="8" pageNumber="356">Type</typeStatus>
locality:
</emphasis>
<location id="ED456040F941CC420A21F99E29A3C43C" LSID="urn:lsid:plazi:treatment:6033878DF941CC430BD9FA912C54C712:ED456040F941CC420A21F99E29A3C43C" box="[367,459,1541,1566]" country="Mexico" name="Lacanja" pageId="8" pageNumber="356" stateProvince="Chiapas">Lacanja</location>
, Chiapas, Mexico.
</paragraph>
<paragraph id="E825369BF941CC420B89F9B22C30C5D0" blockId="8.[151,1437,1397,2034]" pageId="8" pageNumber="356">
<materialsCitation id="58F23CC6F941CC420B89F9B22B19C444" ID-GBIF-Occurrence="2636405302" collectingDate="2009-07-08" collectionCode="UNAM" collectorName="D. Forero &amp; G. Zhang" country="Mexico" elevation="310" latitude="16.77" location="Near Lacanjah" longLatPrecision="768" longitude="-91.13" municipality="Near Lacanjah" pageId="8" pageNumber="356" specimenCode="PTM00283" specimenCount="1" specimenCount-male="1" stateProvince="Chiapas" typeStatus="holotype">
<emphasis id="DAEEEA89F941CC420B89F9B22911C460" bold="true" box="[199,377,1577,1602]" pageId="8" pageNumber="356">
<typeStatus id="37218839F941CC420B89F9B2296AC460" box="[199,258,1577,1602]" pageId="8" pageNumber="356">Type</typeStatus>
material:
</emphasis>
<typeStatus id="37218839F941CC420ACCF9B22987C460" box="[386,495,1577,1602]" pageId="8" pageNumber="356" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="908D760BF941CC420AB5F9B12A06C460" box="[507,622,1578,1602]" name="Mexico" pageId="8" pageNumber="356" value="Mexico">MEXICO</collectingCountry>
:
<specimenCount id="FE9CFD12F941CC420934F9B12AF9C463" box="[634,657,1578,1601]" pageId="8" pageNumber="356" type="male">♂</specimenCount>
,
<collectingRegion id="2A5EF879F941CC4209D1F9B22A96C460" box="[671,766,1577,1602]" country="Mexico" name="Chiapas" pageId="8" pageNumber="356">Chiapas</collectingRegion>
:
<collectingMunicipality id="0841ACE1F941CC420844F9B12BDBC460" box="[778,947,1577,1602]" country="Mexico" latitude="16.77" longLatPrecision="768" longitude="-91.13" name="Near Lacanjah" pageId="8" pageNumber="356" stateProvince="Chiapas">Near Lacanjah</collectingMunicipality>
,
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<elevation id="63B7D1A8F941CC42088EF9B12C6BC460" box="[960,1027,1578,1602]" metricMagnitude="2" metricUnit="m" metricValue="3.1" pageId="8" pageNumber="356" unit="m" value="310.0">310m</elevation>
</quantity>
,
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<geoCoordinate id="8DAE505CF941CC420F36F9B12CB5C460" box="[1144,1245,1578,1602]" degrees="91.13" direction="west" orientation="longitude" pageId="8" pageNumber="356" precision="555" value="-91.13">91.13°W</geoCoordinate>
, 0
<collectingDate id="8C60E9B3F941CC420FB7F9B12D1CC460" box="[1273,1396,1577,1602]" pageId="8" pageNumber="356" value="2009-07-08">
<date id="9C24105BF941CC420FB7F9B12D1CC460" box="[1273,1396,1577,1602]" pageId="8" pageNumber="356" value="2009-07-08">8 Jul 2009</date>
</collectingDate>
,
<collectorName id="456F534DF941CC420ECFF9B1288AC447" pageId="8" pageNumber="356">D. Forero</collectorName>
&amp;
<collectorName id="456F534DF941CC420A45F9D5291CC444" box="[267,372,1613,1638]" pageId="8" pageNumber="356">G. Zhang</collectorName>
[MEX09_SM_L15] /
<specimenCode id="B83C9EE0F941CC420939F9D52A9FC444" box="[631,759,1614,1638]" pageId="8" pageNumber="356">PTM00283</specimenCode>
(
<collectionCode id="8E8BAE5EF941CC420846F9D52B0CC444" box="[776,868,1614,1638]" collectionName="Mexico, Mexico D.F., Universidad Nacional Autonoma de Mexico" country="Mexico" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="8" pageNumber="356">UNAM</collectionCode>
).
</materialsCitation>
<materialsCitation id="58F23CC6F941CC420834F9D52AF2C48C" ID-GBIF-Occurrence="1703280306" collectingDate="2008-12-09" collectionCode="UFRG" collectorName="Weirauch &amp; Forero" country="Costa Rica" elevation="139" latitude="10.5" location="Alajuela" longLatPrecision="779" longitude="-84.59" municipality="On" pageId="8" pageNumber="356" specimenCode="PTM0285" specimenCount="4" specimenCount-female="1" specimenCount-male="1" stateProvince="Alajuela" typeStatus="paratype">
<typeStatus id="37218839F941CC420834F9D52B84C444" box="[890,1004,1614,1638]" pageId="8" pageNumber="356" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="908D760BF941CC4208B6F9D52CCBC444" box="[1016,1187,1614,1638]" name="Costa Rica" pageId="8" pageNumber="356" value="Costa Rica">COSTA RICA</collectingCountry>
:
<specimenCount id="FE9CFD12F941CC420FFEF9D52C97C444" box="[1200,1279,1614,1638]" pageId="8" pageNumber="356">1♂ 1♀</specimenCount>
<collectingRegion id="2A5EF879F941CC420E47F9D62D04C444" box="[1289,1388,1613,1638]" country="Costa Rica" name="Alajuela" pageId="8" pageNumber="356">Alajuela</collectingRegion>
:
<locationDeviation id="F9972FB9F941CC420E34F9D52915C4A8" pageId="8" pageNumber="356">On hwy 4S of Mirador</locationDeviation>
,
<quantity id="2F629B7EF941CC420AC3F9E929A7C4A8" box="[397,463,1650,1674]" metricMagnitude="2" metricUnit="m" metricValue="1.39" pageId="8" pageNumber="356" unit="m" value="139.0">
<elevation id="63B7D1A8F941CC420AC3F9E929A7C4A8" box="[397,463,1650,1674]" metricMagnitude="2" metricUnit="m" metricValue="1.39" pageId="8" pageNumber="356" unit="m" value="139.0">139m</elevation>
</quantity>
,
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<geoCoordinate id="8DAE505CF941CC420907F9EA2AC6C4A8" box="[585,686,1649,1674]" degrees="84.59" direction="west" orientation="longitude" pageId="8" pageNumber="356" precision="555" value="-84.59">84.59°W</geoCoordinate>
,
<collectingDate id="8C60E9B3F941CC4209F0F9E92B32C4A8" box="[702,858,1650,1674]" pageId="8" pageNumber="356">
0
<date id="9C24105BF941CC420983F9E92B32C4A8" box="[717,858,1650,1674]" pageId="8" pageNumber="356" value="2008-12-09">9 Dec 2008</date>
</collectingDate>
,
<collectorName id="456F534DF941CC420824F9EA2C24C4AB" box="[874,1100,1649,1674]" pageId="8" pageNumber="356">Weirauch &amp; Forero</collectorName>
[CR 0 8 L12] (
<specimenCount id="FE9CFD12F941CC420E48F9E92D41C4A8" box="[1286,1321,1650,1674]" pageId="8" pageNumber="356" type="male">1♂</specimenCount>
<collectionCode id="8E8BAE5EF941CC420E7BF9E92DE9C4A8" box="[1333,1409,1650,1674]" collectionName="Brazil, Rio Grande do Sul, Porto Alegre, Universidade Federal do Rio Grande do Sul, Instituto de Biologia" country="Brazil" httpUri="http://grbio.org/cool/nen5-04em" name="Instituto de Biologia" pageId="8" pageNumber="356">UFRG</collectionCode>
) /
<specimenCode id="B83C9EE0F941CC420BD9F90E2960C48C" box="[151,264,1685,1710]" collectionCode="UFRG" country="Brazil" httpUri="http://grbio.org/cool/nen5-04em" name="Instituto de Biologia" pageId="8" pageNumber="356">PTM0285</specimenCode>
(
<specimenCount id="FE9CFD12F941CC420A6FF90D292AC48C" box="[289,322,1686,1710]" pageId="8" pageNumber="356" type="female">1♀</specimenCount>
<collectionCode id="8E8BAE5EF941CC420A00F90D29F2C48C" box="[334,410,1686,1710]" collectionName="Brazil, Rio Grande do Sul, Porto Alegre, Universidade Federal do Rio Grande do Sul, Instituto de Biologia" country="Brazil" httpUri="http://grbio.org/cool/nen5-04em" name="Instituto de Biologia" pageId="8" pageNumber="356">UFRG</collectionCode>
) (abdomen removed)
</materialsCitation>
;
<materialsCitation id="58F23CC6F941CC4209E4F90D2D59C4F0" ID-GBIF-Occurrence="1703280307" collectionCode="NMPC" country="Czech Republic" location="Hacienda Los Inocentes" pageId="8" pageNumber="356" specimenCount="2" specimenCount-male="2" stateProvince="Guanacaste">
<specimenCount id="FE9CFD12F941CC4209E4F90D2AA5C48C" box="[682,717,1686,1710]" pageId="8" pageNumber="356" type="male">1♂</specimenCount>
<collectingRegion id="2A5EF879F941CC420994F90D2B37C48C" box="[730,863,1686,1710]" pageId="8" pageNumber="356">Guanacaste</collectingRegion>
:
<locationDeviation id="F9972FB9F941CC420821F90E2C04C48C" box="[879,1132,1685,1710]" pageId="8" pageNumber="356">
<quantity id="2F629B7EF941CC420821F90E2BD4C48C" box="[879,956,1685,1710]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="8" pageNumber="356" unit="km" value="16.0">16 km</quantity>
E of La Cruz
</locationDeviation>
,
<location id="ED456040F941CC420F33F90E2DF0C48F" LSID="urn:lsid:plazi:treatment:6033878DF941CC430BD9FA912C54C712:ED456040F941CC420F33F90E2DF0C48F" box="[1149,1432,1685,1710]" country="Czech Republic" municipality="Guanacaste" name="Hacienda Los Inocentes" pageId="8" pageNumber="356" stateProvince="Guanacaste">Hacienda Los Inocentes</location>
, 6.23.0 4, MV+UV light / COLLECTIO NACIONAL MUSEUM Praha, Czech Republic (
<specimenCount id="FE9CFD12F941CC420FE9F9212CA2C4F0" box="[1191,1226,1722,1746]" pageId="8" pageNumber="356" type="male">1♂</specimenCount>
<collectionCode id="8E8BAE5EF941CC420F9BF9212D41C4F0" box="[1237,1321,1722,1746]" collectionName="Czech Republic, Prague, National Museum (Natural History)" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="8" pageNumber="356">NMPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="58F23CC6F941CC420E0FF9212CBFC4D4" ID-GBIF-Occurrence="1703280312" collectionCode="NMPC" country="Czech Republic" location="Dos Rios," pageId="8" pageNumber="356" specimenCount="2" specimenCount-male="2" stateProvince="Praha">
<specimenCount id="FE9CFD12F941CC420E0FF9212D0CC4F0" box="[1345,1380,1722,1746]" pageId="8" pageNumber="356" type="male">1♂</specimenCount>
<location id="ED456040F941CC420E21F92128B9C4D4" LSID="urn:lsid:plazi:treatment:6033878DF941CC430BD9FA912C54C712:ED456040F941CC420E21F92128B9C4D4" country="Czech Republic" name="Dos Rios," pageId="8" pageNumber="356" stateProvince="Praha">Dos Rios,</location>
8.16.0 2 / COLLECTIO NACIONAL MUSEUM Praha, Czech Republic (
<specimenCount id="FE9CFD12F941CC420F05F9452C06C4D4" box="[1099,1134,1758,1782]" pageId="8" pageNumber="356" type="male">1♂</specimenCount>
<collectionCode id="8E8BAE5EF941CC420F35F9452CA7C4D4" box="[1147,1231,1758,1782]" collectionName="Czech Republic, Prague, National Museum (Natural History)" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="8" pageNumber="356">NMPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="58F23CC6F941CC420FA7F94529E7C51C" ID-GBIF-Occurrence="1703280308" collectionCode="MNRJ" country="Costa Rica" location="Puntarenas" pageId="8" pageNumber="356" specimenCount="2" specimenCount-female="2" stateProvince="Puntarenas">
<specimenCount id="FE9CFD12F941CC420FA7F9452D64C4D4" box="[1257,1292,1758,1782]" pageId="8" pageNumber="356" type="female">1♀</specimenCount>
<collectingRegion id="2A5EF879F941CC420E57F9452DF0C4D7" box="[1305,1432,1758,1781]" pageId="8" pageNumber="356">Puntarenas</collectingRegion>
:
<locationDeviation id="F9972FB9F941CC420BD9F89A2A7EC538" box="[151,534,1793,1818]" pageId="8" pageNumber="356">Hacienda Baru near to Dominical</locationDeviation>
, 2.16.0 4 MV + UV light / COLLECTIO NACIONAL MUSEUM Praha, Czech Republic (
<specimenCount id="FE9CFD12F941CC420A41F8BD2958C51C" box="[271,304,1830,1854]" pageId="8" pageNumber="356" type="female">1♀</specimenCount>
<collectionCode id="8E8BAE5EF941CC420A79F8BD29EFC51C" box="[311,391,1830,1854]" collectionName="Brazil, Rio de Janeiro, Sao Cristovao, Universidade do Rio Janeiro, Museu Nacional" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="8" pageNumber="356">MNRJ</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="58F23CC6F941CC420AD5F8BD2BC7C5A4" ID-GBIF-Occurrence="2636405301" collectedFrom="1 ♂, 2" collectingDate="2009-07-05" collectionCode="UFRG, UCR" collectorName="D. Forero &amp; G. Zhang" country="Mexico" county="Rd" elevation="361" latitude="18.37" location="Veracruz" longLatPrecision="765" longitude="-95.01" municipality="Cuetzalapan" pageId="8" pageNumber="356" specimenCode="MEX09_SM_L5, PTM00287, PTM00288, PTM00289" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Veracruz">
<collectingCountry id="908D760BF941CC420AD5F8BD2A66C51C" box="[411,526,1830,1854]" name="Mexico" pageId="8" pageNumber="356">MEXICO</collectingCountry>
:
<specimenCount id="FE9CFD12F941CC420956F8BD2A54C51C" box="[536,572,1830,1854]" pageId="8" pageNumber="356" type="male">1♂</specimenCount>
,
<specimenCount id="FE9CFD12F941CC42090CF8BD2A0DC51C" box="[578,613,1830,1854]" pageId="8" pageNumber="356" type="female">2♀</specimenCount>
<collectingRegion id="2A5EF879F941CC420923F8BD2ABDC51F" box="[621,725,1830,1853]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="8" pageNumber="356">Veracruz</collectingRegion>
:
<locationDeviation id="F9972FB9F941CC4209AEF8BE2C8CC51C" box="[736,1252,1829,1854]" pageId="8" pageNumber="356">Rd between Cuetzalapan and Miguel Hidalgo</locationDeviation>
,
<quantity id="2F629B7EF941CC420FBEF8BE2D5AC51C" box="[1264,1330,1829,1854]" metricMagnitude="2" metricUnit="m" metricValue="3.61" pageId="8" pageNumber="356" unit="m" value="361.0">
<elevation id="63B7D1A8F941CC420FBEF8BE2D5AC51C" box="[1264,1330,1829,1854]" metricMagnitude="2" metricUnit="m" metricValue="3.61" pageId="8" pageNumber="356" unit="m" value="361.0">361m</elevation>
</quantity>
,
<geoCoordinate id="8DAE505CF941CC420E70F8BD2DF4C51C" box="[1342,1436,1830,1854]" degrees="18.37" direction="north" orientation="latitude" pageId="8" pageNumber="356" precision="555" value="18.37">18.37°N</geoCoordinate>
<geoCoordinate id="8DAE505CF941CC420BD9F8D22893C540" box="[151,251,1865,1890]" degrees="95.01" direction="west" orientation="longitude" pageId="8" pageNumber="356" precision="555" value="-95.01">95.01°W</geoCoordinate>
, 0
<collectingDate id="8C60E9B3F941CC420A56F8D229FBC540" box="[280,403,1865,1890]" pageId="8" pageNumber="356" value="2009-07-05">
<date id="9C24105BF941CC420A56F8D229FBC540" box="[280,403,1865,1890]" pageId="8" pageNumber="356" value="2009-07-05">5 Jul 2009</date>
</collectingDate>
,
<collectorName id="456F534DF941CC420AEFF8D12A78C543" box="[417,528,1866,1889]" pageId="8" pageNumber="356">D. Forero</collectorName>
&amp;
<collectorName id="456F534DF941CC420976F8D12AC8C540" box="[568,672,1865,1890]" pageId="8" pageNumber="356">G. Zhang</collectorName>
[
<specimenCode id="B83C9EE0F941CC4209FEF8D22B1CC540" box="[688,884,1865,1890]" pageId="8" pageNumber="356">MEX09_SM_L5</specimenCode>
] /
<specimenCode id="B83C9EE0F941CC4208DBF8D12C7CC540" box="[917,1044,1866,1890]" pageId="8" pageNumber="356">PTM00287</specimenCode>
(
<specimenCount id="FE9CFD12F941CC420F67F8D12C22C540" box="[1065,1098,1866,1890]" pageId="8" pageNumber="356" type="female">1♀</specimenCount>
<collectionCode id="8E8BAE5EF941CC420F1DF8D12CF7C540" box="[1107,1183,1866,1890]" collectionName="Brazil, Rio Grande do Sul, Porto Alegre, Universidade Federal do Rio Grande do Sul, Instituto de Biologia" country="Brazil" httpUri="http://grbio.org/cool/nen5-04em" name="Instituto de Biologia" pageId="8" pageNumber="356">UFRG</collectionCode>
) (abdomen removed),
<specimenCode id="B83C9EE0F941CC420BD9F8F5297EC5A4" box="[151,278,1902,1926]" pageId="8" pageNumber="356">PTM00288</specimenCode>
(
<specimenCount id="FE9CFD12F941CC420A62F8F52925C5A4" box="[300,333,1902,1926]" pageId="8" pageNumber="356" type="female">1♀</specimenCount>
<collectionCode id="8E8BAE5EF941CC420A18F8F529CAC5A4" box="[342,418,1902,1926]" collectionName="Brazil, Rio Grande do Sul, Porto Alegre, Universidade Federal do Rio Grande do Sul, Instituto de Biologia" pageId="8" pageNumber="356">UFRG</collectionCode>
) (abdomen removed),
<specimenCode id="B83C9EE0F941CC4209E7F8F52B41C5A4" box="[681,809,1902,1926]" pageId="8" pageNumber="356">PTM00289</specimenCode>
(
<specimenCount id="FE9CFD12F941CC420872F8F52B37C5A4" box="[828,863,1902,1926]" pageId="8" pageNumber="356" type="male">1♂</specimenCount>
<collectionCode id="8E8BAE5EF941CC420827F8F52BC0C5A4" box="[873,936,1902,1926]" collectionName="USA, California, Riverside, University of California" pageId="8" pageNumber="356">UCR</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="58F23CC6F941CC4208F3F8F529FBC5EC" ID-GBIF-Occurrence="1703280304" collectingDate="2009-07-06" collectionCode="UFRG" collectorName="D. Forero &amp; G. Zhang" country="Mexico" latitude="17.74" location="Agua Fria" longLatPrecision="767" longitude="-92.08" municipality="Agua Fria" pageId="8" pageNumber="356" specimenCode="MEX09_SM_L7, PTM00286" specimenCount="25" specimenCount-male="25" stateProvince="Chiapas">
<specimenCount id="FE9CFD12F941CC4208F3F8F52B88C5A4" box="[957,992,1902,1926]" pageId="8" pageNumber="356" type="male">1♂</specimenCount>
<collectingRegion id="2A5EF879F941CC4208A4F8F62C21C5A4" box="[1002,1097,1901,1926]" country="Mexico" name="Chiapas" pageId="8" pageNumber="356">Chiapas</collectingRegion>
:
<collectingMunicipality id="0841ACE1F941CC420F18F8F52CA4C5A4" box="[1110,1228,1901,1926]" pageId="8" pageNumber="356">Agua Fria</collectingMunicipality>
,
<locationDeviation id="F9972FB9F941CC420F94F8F62968C588" pageId="8" pageNumber="356">
about
<quantity id="2F629B7EF941CC420E6CF8F62D1BC5A4" box="[1314,1395,1901,1926]" metricMagnitude="3" metricUnit="m" metricValue="8.5" pageId="8" pageNumber="356" unit="km" value="8.5">8.5 km</quantity>
off Hwy 186
</locationDeviation>
,
<specimenCount id="FE9CFD12F941CC420A45F8092928C588" box="[267,320,1938,1962]" pageId="8" pageNumber="356" type="male">23m</specimenCount>
,
<geoCoordinate id="8DAE505CF941CC420A05F80929C1C588" box="[331,425,1938,1962]" degrees="17.74" direction="north" orientation="latitude" pageId="8" pageNumber="356" precision="555" value="17.74">17.74°N</geoCoordinate>
<geoCoordinate id="8DAE505CF941CC420AE1F8092A7DC588" box="[431,533,1938,1962]" degrees="92.08" direction="west" orientation="longitude" pageId="8" pageNumber="356" precision="555" value="-92.08">92.08°W</geoCoordinate>
, 0
<collectingDate id="8C60E9B3F941CC420963F80A2ACCC588" box="[557,676,1937,1962]" pageId="8" pageNumber="356" value="2009-07-06">
<date id="9C24105BF941CC420963F80A2ACCC588" box="[557,676,1937,1962]" pageId="8" pageNumber="356" value="2009-07-06">6 Jul 2009</date>
</collectingDate>
,
<collectorName id="456F534DF941CC4209E0F8092B73C58B" box="[686,795,1938,1961]" pageId="8" pageNumber="356">D. Forero</collectorName>
&amp;
<collectorName id="456F534DF941CC420870F8092BCBC588" box="[830,931,1937,1962]" pageId="8" pageNumber="356">G. Zhang</collectorName>
[
<specimenCode id="B83C9EE0F941CC4208FEF8092C1CC588" box="[944,1140,1938,1962]" pageId="8" pageNumber="356">MEX09_SM_L7</specimenCode>
] /
<specimenCode id="B83C9EE0F941CC420FC1F80A2D67C588" box="[1167,1295,1937,1962]" pageId="8" pageNumber="356">PTM00286</specimenCode>
(
<specimenCount id="FE9CFD12F941CC420E51F8092D2AC588" box="[1311,1346,1938,1962]" pageId="8" pageNumber="356" type="male">1♂</specimenCount>
<collectionCode id="8E8BAE5EF941CC420E06F8092DFCC588" box="[1352,1428,1938,1962]" collectionName="Brazil, Rio Grande do Sul, Porto Alegre, Universidade Federal do Rio Grande do Sul, Instituto de Biologia" country="Brazil" httpUri="http://grbio.org/cool/nen5-04em" name="Instituto de Biologia" pageId="8" pageNumber="356">UFRG</collectionCode>
) (pygophore removed)
</materialsCitation>
;
<materialsCitation id="58F23CC6F941CC420AEDF82D2C3CC5D0" ID-GBIF-Occurrence="1703280305" collectingDate="2009-07-08" collectionCode="UFRG, AMNH, UNAM" collectorName="D. Forero &amp; G. Zhang" country="Mexico" elevation="310" latitude="16.77" location="Near Lacanjah" longLatPrecision="768" longitude="-91.13" municipality="Near Lacanjah" pageId="8" pageNumber="356" specimenCode="MEX09_SM_L15, PTM0284, MNRJ" specimenCount="10" specimenCount-female="4" specimenCount-male="6" stateProvince="Jul 2009">
<specimenCount id="FE9CFD12F941CC420AEDF82D29AFC5EC" box="[419,455,1974,1998]" pageId="8" pageNumber="356" type="male">3♂</specimenCount>
<specimenCount id="FE9CFD12F941CC420A9CF82D299DC5EC" box="[466,501,1974,1998]" pageId="8" pageNumber="356" type="female">2♀</specimenCount>
<collectingMunicipality id="0841ACE1F941CC42094FF82D2AC6C5EC" box="[513,686,1973,1998]" pageId="8" pageNumber="356">Near Lacanjah</collectingMunicipality>
,
<quantity id="2F629B7EF941CC4209F0F82D2B68C5EC" box="[702,768,1974,1998]" metricMagnitude="2" metricUnit="m" metricValue="3.1" pageId="8" pageNumber="356" unit="m" value="310.0">
<elevation id="63B7D1A8F941CC4209F0F82D2B68C5EC" box="[702,768,1974,1998]" metricMagnitude="2" metricUnit="m" metricValue="3.1" pageId="8" pageNumber="356" unit="m" value="310.0">310m</elevation>
</quantity>
,
<geoCoordinate id="8DAE505CF941CC42085FF82E2B07C5EC" box="[785,879,1973,1998]" degrees="16.77" direction="north" orientation="latitude" pageId="8" pageNumber="356" precision="555" value="16.77">16.77°N</geoCoordinate>
<geoCoordinate id="8DAE505CF941CC420835F82D2B88C5EC" box="[891,992,1974,1998]" degrees="91.13" direction="west" orientation="longitude" pageId="8" pageNumber="356" precision="555" value="-91.13">91.13°W</geoCoordinate>
, 0
<collectingDate id="8C60E9B3F941CC420F4EF82D2CE8C5EC" box="[1024,1152,1973,1998]" pageId="8" pageNumber="356" value="2009-07-08">
<date id="9C24105BF941CC420F4EF82D2CE8C5EC" box="[1024,1152,1973,1998]" pageId="8" pageNumber="356" value="2009-07-08">8 Jul 2009</date>
</collectingDate>
,
<collectorName id="456F534DF941CC420FDFF82D2D6BC5EF" box="[1169,1283,1974,1997]" pageId="8" pageNumber="356">D. Forero</collectorName>
&amp;
<collectorName id="456F534DF941CC420E7FF82D2DF4C5EC" box="[1329,1436,1973,1998]" pageId="8" pageNumber="356">G. Zhang</collectorName>
[
<specimenCode id="B83C9EE0F941CC420BD3F8422907C5D0" box="[157,367,2009,2034]" pageId="8" pageNumber="356">MEX09_SM_L15</specimenCode>
] /
<specimenCode id="B83C9EE0F941CC420AC5F8412995C5D0" box="[395,509,2010,2034]" pageId="8" pageNumber="356">PTM0284</specimenCode>
(
<specimenCount id="FE9CFD12F941CC420943F8412A58C5D0" box="[525,560,2010,2034]" pageId="8" pageNumber="356" type="male">1♂</specimenCount>
<collectionCode id="8E8BAE5EF941CC420979F8412AEEC5D0" box="[567,646,2010,2034]" collectionName="Brazil, Rio Grande do Sul, Porto Alegre, Universidade Federal do Rio Grande do Sul, Instituto de Biologia" country="Brazil" httpUri="http://grbio.org/cool/nen5-04em" name="Instituto de Biologia" pageId="8" pageNumber="356">UFRG</collectionCode>
), (
<specimenCount id="FE9CFD12F941CC4209EDF8412AADC5D0" box="[675,709,2010,2034]" pageId="8" pageNumber="356" type="male">1♂</specimenCount>
<specimenCount id="FE9CFD12F941CC420982F8412A87C5D0" box="[716,751,2010,2034]" pageId="8" pageNumber="356" type="female">1♀</specimenCount>
<collectionCode id="8E8BAE5EF941CC4209B8F8412B26C5D0" box="[758,846,2010,2034]" collectionName="USA, New York, New York, American Museum of Natural History" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="8" pageNumber="356">AMNH</collectionCode>
,
<specimenCount id="FE9CFD12F941CC420814F8412B1EC5D0" box="[858,886,2010,2034]" pageId="8" pageNumber="356" type="male">1♂</specimenCount>
<specimenCode id="B83C9EE0F941CC420838F8412BA0C5D0" box="[886,968,2010,2034]" pageId="8" pageNumber="356">MNRJ</specimenCode>
<specimenCount id="FE9CFD12F941CC420881F8412B8EC5D0" box="[975,998,2010,2034]" pageId="8" pageNumber="356" type="female">1♀</specimenCount>
<collectionCode id="8E8BAE5EF941CC4208A8F8412C23C5D0" box="[998,1099,2010,2034]" collectionName="Mexico, Mexico D.F., Universidad Nacional Autonoma de Mexico" country="Mexico" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="8" pageNumber="356">UNAM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A0806510F940CC430B89FF0C2BCBC632" pageId="9" pageNumber="357" type="description">
<paragraph id="E825369BF940CC430B89FF0C292DC33F" blockId="9.[151,1437,151,1328]" pageId="9" pageNumber="357">
<emphasis id="DAEEEA89F940CC430B89FF0C2933C292" bold="true" box="[199,347,151,176]" pageId="9" pageNumber="357">Description:</emphasis>
Dorsal surface light brown; punctures black; punctures smaller on hemelytra than on remaining body parts. Humeri acutely projected laterally, but not developed as a spine, varying from light brown to reddish. Ventral surface greenish to light-yellow; punctures concolorous with the surface. Antennae uniformly yellow to orange (
<figureCitation id="70A12A1EF940CC430BBCFE9F295FC33F" box="[242,311,260,285]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1938,1961]" captionTargetBox="[160,1421,1606,1917]" captionTargetId="figure@2.[151,1436,1599,1917]" captionTargetPageId="2" captionText="FIGURES 1–5. Dorsal view of representative specimens of sulcacitus group species. 1, Euschistus (Euschistus) corcoVacitus; 2, Euschistus (Euschistus) olacitus; 3, Euschistus (Euschistus) palacitus; 4, Euschistus (Euschistus) sulcacitus; 5, Euschistus (Euschistus) tacitus sp. nov., holotype. Scale bars: 2 mm." httpUri="https://zenodo.org/record/1094898/files/figure.png" pageId="9" pageNumber="357">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="E825369BF940CC430B89FEBC290AC042" blockId="9.[151,1437,151,1328]" pageId="9" pageNumber="357">Head longer than wide; head punctate, punctures less dense on disc. Clypeus rounded apically; clypeus slightly longer than mandibular plates, rarely equal in length. Mandibular plates rounded apically. Rostrum reaching metacoxae. Anterolateral margins of pronotum toothed anteriorly and smooth posteriorly; teeth-like projections small, dark brown. Scutellum reaching abdominal segment V; anterolateral angles foveate. Punctures on scutellum and hemelytra smaller apically than basally. Membrane of hemelytra light brown; exocorium reddish in some specimens. Evaporatorium of metasternal glands not surpassing median line between ostiole and lateral margin of mesopleuron; spout peritreme. Presence of 1+1 dark spots at base of leg between epimeron and episternum of each pleurite, plus 1+1 at metepimeron medially. Legs concolorous with ventral surface of thorax; small dark dots on femora and tibiae.</paragraph>
<paragraph id="E825369BF940CC430B89FDF72B97C08A" blockId="9.[151,1437,151,1328]" pageId="9" pageNumber="357">Abdomen: dorsally, disc of connexivum yellow, black stripes on anterior and posterior margins; ventrally, anterior angle with dark spot. Spiracles concolorous with abdominal sternites.</paragraph>
<paragraph id="E825369BF940CC430B89FD2F2D34C17F" blockId="9.[151,1437,151,1328]" pageId="9" pageNumber="357">
<emphasis id="DAEEEA89F940CC430B89FD2F29E9C0EF" bold="true" box="[199,385,692,717]" pageId="9" pageNumber="357">Male genitalia:</emphasis>
Pygophore trapezoidal. Dorsal rim uniformly concave in dorsal view (
<figureCitation id="70A12A1EF940CC430F95FD2F2D5BC0EE" box="[1243,1331,692,717]" captionStart="FIGURES 6–20" captionStartId="3.[151,264,1870,1893]" captionTargetBox="[188,1393,193,1826]" captionTargetId="figure@3.[188,1393,193,1840]" captionTargetPageId="3" captionText="FIGURES 6–20. Dissected and macerated pygophore of species of sulcacitus group. Pygophore in dorsal (6, 9, 12, 15, 18), posterior (7, 10, 13, 16, 19), and ventral (8, 11, 14, 17, 20) views. 6–8, Euschistus (Euschistus) corcoVacitus; 9–11, Euschistus (Euschistus) olacitus; 12–14, Euschistus (Euschistus) palacitus; 15–17, Euschistus (Euschistus) sulcacitus; 18–20, Euschistus (Euschistus) tacitus sp. nov.. dr, dorsal rim; mp, median projection; par, paramere; slvr, superior layer of ventral rim; spdr, superior process of dorsal rim; vr, ventral rim. Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1094900/files/figure.png" pageId="9" pageNumber="357">Fig. 18</figureCitation>
) median projection of dorsal rim long, sinuosity shallow; superior process of dorsal rim tooth-like. Ventral rim slightly sinuous, convex medially in ventral view (
<figureCitation id="70A12A1EF940CC430931FD672ABEC136" box="[639,726,764,789]" captionStart="FIGURES 6–20" captionStartId="3.[151,264,1870,1893]" captionTargetBox="[188,1393,193,1826]" captionTargetId="figure@3.[188,1393,193,1840]" captionTargetPageId="3" captionText="FIGURES 6–20. Dissected and macerated pygophore of species of sulcacitus group. Pygophore in dorsal (6, 9, 12, 15, 18), posterior (7, 10, 13, 16, 19), and ventral (8, 11, 14, 17, 20) views. 6–8, Euschistus (Euschistus) corcoVacitus; 9–11, Euschistus (Euschistus) olacitus; 12–14, Euschistus (Euschistus) palacitus; 15–17, Euschistus (Euschistus) sulcacitus; 18–20, Euschistus (Euschistus) tacitus sp. nov.. dr, dorsal rim; mp, median projection; par, paramere; slvr, superior layer of ventral rim; spdr, superior process of dorsal rim; vr, ventral rim. Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1094900/files/figure.png" pageId="9" pageNumber="357">Fig. 20</figureCitation>
). In posterior view, ventral rim protuberant medially; superior layer of ventral rim excavated in V-shape in posterior view (
<figureCitation id="70A12A1EF940CC43087BFC842BEFC11A" box="[821,903,799,824]" captionStart="FIGURES 6–20" captionStartId="3.[151,264,1870,1893]" captionTargetBox="[188,1393,193,1826]" captionTargetId="figure@3.[188,1393,193,1840]" captionTargetPageId="3" captionText="FIGURES 6–20. Dissected and macerated pygophore of species of sulcacitus group. Pygophore in dorsal (6, 9, 12, 15, 18), posterior (7, 10, 13, 16, 19), and ventral (8, 11, 14, 17, 20) views. 6–8, Euschistus (Euschistus) corcoVacitus; 9–11, Euschistus (Euschistus) olacitus; 12–14, Euschistus (Euschistus) palacitus; 15–17, Euschistus (Euschistus) sulcacitus; 18–20, Euschistus (Euschistus) tacitus sp. nov.. dr, dorsal rim; mp, median projection; par, paramere; slvr, superior layer of ventral rim; spdr, superior process of dorsal rim; vr, ventral rim. Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1094900/files/figure.png" pageId="9" pageNumber="357">Fig. 19</figureCitation>
). Paramere: margin connecting basal and apical process slightly concave; apical process somewhat spatulate at apex; middle process not developed (
<figureCitation id="70A12A1EF940CC430FB5FCDF2D38C17F" box="[1275,1360,836,861]" captionStart="FIGURES 21–26" captionStartId="4.[151,264,504,527]" captionTargetBox="[152,1421,194,484]" captionTargetId="figure@4.[151,1436,193,484]" captionTargetPageId="4" captionText="FIGURES 21–26. Right paramere of species of sulcacitus group. 21, schematic drawing of paramere indicating its regionalization; 22, Euschistus (Euschistus) corcoVacitus; 23, Euschistus (Euschistus) olacitus; 24, Euschistus (Euschistus) palacitus; 25, Euschistus (Euschistus) sulcacitus; 26, Euschistus (Euschistus) tacitus sp. nov. app, apical process of paramere; bpp, basal process of paramere; cr, crown; st, stem. Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1094902/files/figure.png" pageId="9" pageNumber="357">Fig. 26</figureCitation>
).
</paragraph>
<paragraph id="E825369BF940CC430B89FCFC2BCBC632" blockId="9.[151,1437,151,1328]" pageId="9" pageNumber="357">
<emphasis id="DAEEEA89F940CC430B89FCFC29FFC1A2" bold="true" box="[199,407,871,896]" pageId="9" pageNumber="357">Female genitalia:</emphasis>
Posterior margin of gonocoxites VIII concave, posterolateral margin of gonocoxites VIII projected posteriorly, reaching middle of gonocoxites IX. Gonocoxites
<date id="9C24105BF940CC4308F6FC162B83C186" box="[952,1003,908,932]" pageId="9" pageNumber="357">IX 2</date>
times wider than long, lateral margins strongly rounded; anterior margins of gonocoxites IX straight, posterior margin convex (
<figureCitation id="70A12A1EF940CC430F31FC342CBCC1EA" box="[1151,1236,943,968]" captionStart="FIGURES 45–59" captionStartId="6.[151,264,1803,1826]" captionTargetBox="[151,1436,193,1782]" captionTargetId="figure@6.[151,1436,193,1782]" captionTargetPageId="6" captionText="FIGURES 45–59. Female genital plates of species of sulcacitus group. Genital plates in posterior (45, 48, 51, 54, 57) and postero-ventral (45, 46, 49, 50, 52, 53, 55, 56, 58, 59) views. 45–47, Euschistus (Euschistus) corcoVacitus; 48–50, Euschistus (Euschistus) olacitus; 51–53, Euschistus (Euschistus) palacitus; 54–56, Euschistus (Euschistus) sulcacitus; 57–59, Euschistus (Euschistus) tacitus sp. nov.. gc, gonocoxites; la, laterotergites. Scale bars: 1 mm." httpUri="https://zenodo.org/record/1094908/files/figure.png" pageId="9" pageNumber="357">Fig. 57</figureCitation>
). Posterior region of ductus receptaculi proximalis straight, not dilated. Ductus receptaculi distalis dilated adjacent to anterior annular flanges. Pars intermedialis sclerotized at basal three-fourths (
<figureCitation id="70A12A1EF940CC43080CFC6C2BFEC632" box="[834,918,1015,1040]" captionStart="FIGURES 60–64" captionStartId="7.[151,264,1794,1817]" captionTargetBox="[151,1436,193,1773]" captionTargetId="figure@7.[151,1436,193,1773]" captionTargetPageId="7" captionText="FIGURES 60–64. Female internal genitalia of species of sulcacitus group. 60, Euschistus (Euschistus) corcoVacitus; 61, Euschistus (Euschistus) olacitus; 62, Euschistus (Euschistus) palacitus; 63, Euschistus (Euschistus) sulcacitus; 64, Euschistus (Euschistus) tacitus sp. nov.. Insets are the capsula seminalis. aaf, anterior anular flange; cs, capsula seminalis; dr, ductus receptaculi; g, gonapophysis; gc, gonocoxites; la, laterotergites; paf, posterior anular flange; pi, pars intermedialis; rs, ring sclerites; tvi, thickening of vaginal intima; va, vesicular area. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/1094910/files/figure.png" pageId="9" pageNumber="357">Fig. 64</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A0806510F940CC430B89FB872C54C712" pageId="9" pageNumber="357" type="discussion">
<paragraph id="E825369BF940CC430B89FB872C54C712" blockId="9.[151,1437,151,1328]" pageId="9" pageNumber="357">
<emphasis id="DAEEEA89F940CC430B89FB872927C617" bold="true" box="[199,335,1052,1077]" inLineHeading="true" pageId="9" pageNumber="357" reason="1">Comments.</emphasis>
The external morphology of the
<taxonomicName id="2F9A4D18F940CC4309AAFB872B3CC616" authorityName="Rolston. Rolston" authorityYear="1971" box="[740,852,1052,1076]" class="Insecta" family="Pentatomidae" genus="Euschistus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="357" phylum="Arthropoda" rank="species" species="sulcacitus">
<emphasis id="DAEEEA89F940CC4309AAFB872B3CC616" box="[740,852,1052,1076]" italics="true" pageId="9" pageNumber="357">sulcacitus</emphasis>
</taxonomicName>
group is homogeneous among its species. Some characteristics (e.g. coloration of dorsal and ventral surfaces of body, shape and length of humeral angles, and total body length) vary intraspecifically, making them useless to species diagnosis. Otherwise, the genitalic morphology provides valuable characters to species definition.
<taxonomicName id="2F9A4D18F940CC43099FFB132BF3C6BD" authority="Bianchi &amp; Barão &amp; Grazia, 2017" authorityName="Bianchi &amp; Barão &amp; Grazia" authorityYear="2017" box="[721,923,1160,1184]" class="Insecta" family="Pentatomidae" genus="Euschistus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="357" phylum="Arthropoda" rank="species" species="tacitus" status="sp. nov.">
<emphasis id="DAEEEA89F940CC43099FFB132BF3C6BD" box="[721,923,1160,1184]" italics="true" pageId="9" pageNumber="357">Euschistus tacitus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C1DD57F2F940CC4308EDFB1C2B95C682" box="[931,1021,1159,1184]" pageId="9" pageNumber="357" rank="species">
<emphasis id="DAEEEA89F940CC4308EDFB1C2B95C682" bold="true" box="[931,1021,1159,1184]" pageId="9" pageNumber="357">sp. nov.</emphasis>
</taxonomicNameLabel>
is more similar to
<taxonomicName id="2F9A4D18F940CC430F9AFB122D0CC682" authorityName="Rolston. Rolston" authorityYear="1971" box="[1236,1380,1160,1184]" class="Insecta" family="Pentatomidae" genus="Euschistus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="357" phylum="Arthropoda" rank="species" species="sulcacitus">
<emphasis id="DAEEEA89F940CC430F9AFB122D0CC682" box="[1236,1380,1160,1184]" italics="true" pageId="9" pageNumber="357">E. sulcacitus</emphasis>
</taxonomicName>
than to the others species of the group by the posterior margin of pygophore slightly sinuate, convex mesially; and by the laterotergites IX not surpassing dorsal band that unites laterotergites VIII in posteroventral view.
<taxonomicName id="2F9A4D18F940CC430E6AFB4B288AC72E" authority="Bianchi &amp; Barão &amp; Grazia, 2017" authorityName="Bianchi &amp; Barão &amp; Grazia" authorityYear="2017" class="Insecta" family="Pentatomidae" genus="Euschistus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="357" phylum="Arthropoda" rank="species" species="tacitus" status="sp. nov.">
<emphasis id="DAEEEA89F940CC430E6AFB4B288AC72E" italics="true" pageId="9" pageNumber="357">Euschistus tacitus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C1DD57F2F940CC430BA4FB6F292DC72F" box="[234,325,1268,1293]" pageId="9" pageNumber="357" rank="species">
<emphasis id="DAEEEA89F940CC430BA4FB6F292DC72F" bold="true" box="[234,325,1268,1293]" pageId="9" pageNumber="357">sp. nov.</emphasis>
</taxonomicNameLabel>
can be distinguished from
<taxonomicName id="2F9A4D18F940CC430932FB6E2B64C72E" authorityName="Rolston. Rolston" authorityYear="1971" box="[636,780,1268,1292]" class="Insecta" family="Pentatomidae" genus="Euschistus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="357" phylum="Arthropoda" rank="species" species="sulcacitus">
<emphasis id="DAEEEA89F940CC430932FB6E2B64C72E" box="[636,780,1268,1292]" italics="true" pageId="9" pageNumber="357">E. sulcacitus</emphasis>
</taxonomicName>
by the inferior layer of ventral rim mesially tumescent in posterior view, and posterior margin of gonocoxites VIII mesially slightly concave.
</paragraph>
</subSubSection>
</treatment>
</document>
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