586 lines
69 KiB
XML
586 lines
69 KiB
XML
<document id="CC0D8F11780120A6CAAFDD4145969FC2" ID-DOI="10.11646/zootaxa.5324.1.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="8213864" ID-ZooBank="5FCE2373-0415-4B1A-8C18-6786956E58A9" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1691128571643" checkinUser="plazi" docAuthor="Ermilov, Sergey G. & Rybalov, Leonid B." docDate="2023" docId="5B6187E7FFF5C97DFF7FF8EAFD090CAB" docLanguage="en" docName="zootaxa.5324.1.5.pdf" docOrigin="Zootaxa 5324 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5324.1.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tyrphonothrus digeluensis Ermilov & Rybalov 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="28" masterDocId="A758FF9FFFF4C979FFE8FFA0FFC00B5F" masterDocTitle="Ontogenetic instars of the oribatid mite Tyrphonothrus digeluensis sp. nov. (Acari Oribatida, Malaconothridae) from Ethiopia" masterLastPageNumber="36" masterPageNumber="24" pageNumber="25" updateTime="1691129144927" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="991955AB3DA97293E3B00E47D109D14B">
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<mods:title id="28FD73B494923E6D753F01B7F5037B2A">Ontogenetic instars of the oribatid mite Tyrphonothrus digeluensis sp. nov. (Acari Oribatida, Malaconothridae) from Ethiopia</mods:title>
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<mods:name id="A8E4A6A50228569DE81F0C21E98B59BB" type="personal">
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<mods:roleTerm id="A18F2B8DA7AC9F3F19E91CFB92018582">Author</mods:roleTerm>
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<mods:namePart id="402A13C4132558A891F6AE78DF661E8A">Ermilov, Sergey G.</mods:namePart>
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<mods:affiliation id="ADA1593CFAC3D0682891E777ED559693">Tyumen State University, Institute of Environmental and Agricultural Biology (X-BIO), Tyumen, Russia.</mods:affiliation>
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<mods:roleTerm id="569F5DC36BD6D3516F049DDCB6F8BE65">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F0D19FFFD560C188E6118E346DDFF69C">Rybalov, Leonid B.</mods:namePart>
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<mods:affiliation id="9F698F0D8D61769C37476357394738F2">Institute of Ecology and Evolution, Russian Academy of Sciences, Laboratory of Soil Zoology and General Entomology, Moscow, Russia.</mods:affiliation>
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<mods:typeOfResource id="B5D3498F49E97BA2924D56847D4F1F87">text</mods:typeOfResource>
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<mods:title id="0E7567A21960D8143D9C5FBF454AA14A">Zootaxa</mods:title>
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<mods:part id="DA0705D6FC7B0867215FC6EA8E97407E">
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<mods:date id="46018767538FF0348DEAD7F55EE6732B">2023</mods:date>
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<mods:number id="359BA6BEEF4F47108E013AD6DADB6A1B">2023-08-04</mods:number>
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<mods:number id="B809ECED0E12A268EA8F47573C851847">5324</mods:number>
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<mods:url id="4FC6556A97779D3C16ED61A11C018F71">http://dx.doi.org/10.11646/zootaxa.5324.1.5</mods:url>
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<mods:classification id="3AC94F8026064CE1A647154642A24688">journal article</mods:classification>
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<mods:identifier id="7B9DA6CA1855E9165F8344F4396B0DFD" type="DOI">10.11646/zootaxa.5324.1.5</mods:identifier>
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<mods:identifier id="059A0EB9716D30AD1E4E5408EE6415C1" type="ISSN">1175-5326</mods:identifier>
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<treatment id="5B6187E7FFF5C97DFF7FF8EAFD090CAB" ID-DOI="http://doi.org/10.5281/zenodo.8213828" ID-Zenodo-Dep="8213828" LSID="urn:lsid:plazi:treatment:5B6187E7FFF5C97DFF7FF8EAFD090CAB" httpUri="http://treatment.plazi.org/id/5B6187E7FFF5C97DFF7FF8EAFD090CAB" lastPageId="4" lastPageNumber="28" pageId="1" pageNumber="25">
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<subSubSection id="9BD2657AFFF5C978FF7FF8EAFD800C3A" box="[151,576,1866,1893]" pageId="1" pageNumber="25" type="nomenclature">
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<paragraph id="D37736F1FFF5C978FF7FF8EAFD800C3A" blockId="1.[151,576,1866,1893]" box="[151,576,1866,1893]" pageId="1" pageNumber="25">
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<heading id="883F819DFFF5C978FF7FF8EAFD800C3A" bold="true" box="[151,576,1866,1893]" fontSize="11" level="1" pageId="1" pageNumber="25" reason="1">
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<emphasis id="E1BCEAE3FFF5C978FF7FF8EAFD800C3A" bold="true" box="[151,576,1866,1893]" pageId="1" pageNumber="25">
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<taxonomicName id="14C84D72FFF5C978FF7FF8EAFE1B0C3A" authority="Ermilov & Rybalov, 2023" authorityName="Ermilov & Rybalov" authorityYear="2023" box="[151,475,1866,1893]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="1" pageNumber="25" phylum="Arthropoda" rank="species" species="digeluensis" status="sp. nov.">
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<emphasis id="E1BCEAE3FFF5C978FF7FF8EAFE1B0C3A" bold="true" box="[151,475,1866,1893]" italics="true" pageId="1" pageNumber="25">Tyrphonothrus digeluensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="FA8F5798FFF5C978FE0BF8EBFD800C3A" box="[483,576,1867,1893]" pageId="1" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="9BD2657AFFF5C97BFF7FF832FD370A3A" lastPageId="2" lastPageNumber="26" pageId="1" pageNumber="25" type="diagnosis">
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<paragraph id="D37736F1FFF5C97BFF7FF832FD370A3A" blockId="1.[151,1437,1938,2043]" lastBlockId="2.[151,1437,151,2013]" lastPageId="2" lastPageNumber="26" pageId="1" pageNumber="25">
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<emphasis id="E1BCEAE3FFF5C978FF7FF832FED20CF3" bold="true" box="[151,274,1938,1964]" pageId="1" pageNumber="25">Diagnosis.</emphasis>
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<emphasis id="E1BCEAE3FFF5C978FEF5F833FE9A0CF2" box="[285,346,1939,1965]" italics="true" pageId="1" pageNumber="25">Adult</emphasis>
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. Body length: 705–735. Rostral, lamellar and interlamellar setae long, subflagellate, smooth (
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<emphasis id="E1BCEAE3FFF5C978FF49F817FF770C8F" box="[161,183,1975,2000]" italics="true" pageId="1" pageNumber="25">in</emphasis>
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>
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<emphasis id="E1BCEAE3FFF5C978FF2EF817FF1A0C8E" box="[198,218,1975,2001]" italics="true" pageId="1" pageNumber="25">le</emphasis>
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>
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<emphasis id="E1BCEAE3FFF5C978FF02F817FEC20C8F" box="[234,258,1975,2000]" italics="true" pageId="1" pageNumber="25">ro</emphasis>
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); exobothridial seta
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<emphasis id="E1BCEAE3FFF5C978FE1EF817FDCF0C8F" box="[502,527,1975,2000]" italics="true" pageId="1" pageNumber="25">ex</emphasis>
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<subScript id="4F4C34B4FFF5C978FDE7F869FDD70C88" attach="left" box="[527,535,1993,2007]" fontSize="6" pageId="1" pageNumber="25">1</subScript>
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short, setiform, barbed. Medioposterior part of notogaster slightly widening. Longitudinal notogastral ridges present. Notogastral setae
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<emphasis id="E1BCEAE3FFF5C978FCC0F87BFCF40CAB" box="[808,820,2011,2036]" italics="true" pageId="1" pageNumber="25">e</emphasis>
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<subScript id="4F4C34B4FFF5C978FCDCF84DFCFC0CA4" attach="left" box="[820,828,2029,2043]" fontSize="6" pageId="1" pageNumber="25">2</subScript>
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,
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<emphasis id="E1BCEAE3FFF5C978FCA0F87BFC960CAA" box="[840,854,2011,2037]" italics="true" pageId="1" pageNumber="25">h</emphasis>
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<subScript id="4F4C34B4FFF5C978FCBEF84DFC9E0CA4" attach="left" box="[854,862,2029,2043]" fontSize="6" pageId="1" pageNumber="25">1</subScript>
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,
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<emphasis id="E1BCEAE3FFF5C978FC85F87BFCBB0CAA" box="[877,891,2011,2037]" italics="true" pageId="1" pageNumber="25">h</emphasis>
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<subScript id="4F4C34B4FFF5C978FC93F84DFC430CA4" attach="left" box="[891,899,2029,2043]" fontSize="6" pageId="1" pageNumber="25">2</subScript>
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,
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<emphasis id="E1BCEAE3FFF5C978FC79F87BFC5F0CAB" box="[913,927,2011,2036]" italics="true" pageId="1" pageNumber="25">p</emphasis>
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<subScript id="4F4C34B4FFF5C978FC77F84DFC670CA4" attach="left" box="[927,935,2029,2043]" fontSize="6" pageId="1" pageNumber="25">2</subScript>
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long, subflagellate, smooth; others medium-sized, setiform, barbed;
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<emphasis id="E1BCEAE3FFF6C97BFE46FF37FE7C0BEF" box="[430,444,151,176]" italics="true" pageId="2" pageNumber="26">p</emphasis>
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<subScript id="4F4C34B4FFF6C97BFE54FF09FE040BE8" attach="left" box="[444,452,169,183]" fontSize="6" pageId="2" pageNumber="26">2</subScript>
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and
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<emphasis id="E1BCEAE3FFF6C97BFDE9FF37FDCF0BEF" box="[513,527,151,176]" italics="true" pageId="2" pageNumber="26">p</emphasis>
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<subScript id="4F4C34B4FFF6C97BFDE7FF09FDD70BE8" attach="left" box="[527,535,169,183]" fontSize="6" pageId="2" pageNumber="26">3</subScript>
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distant from each other. Epimeral setae
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<emphasis id="E1BCEAE3FFF6C97BFC07FF37FBCC0BEE" box="[1007,1036,151,177]" italics="true" pageId="2" pageNumber="26">1a</emphasis>
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,
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<emphasis id="E1BCEAE3FFF6C97BFBF4FF37FBF70BEE" box="[1052,1079,151,177]" italics="true" pageId="2" pageNumber="26">1c</emphasis>
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,
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<emphasis id="E1BCEAE3FFF6C97BFBA0FF37FBA40BEE" box="[1096,1124,151,177]" italics="true" pageId="2" pageNumber="26">2a</emphasis>
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,
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<emphasis id="E1BCEAE3FFF6C97BFB9DFF37FB510BEE" box="[1141,1169,151,177]" italics="true" pageId="2" pageNumber="26">3a</emphasis>
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very short, penicillate; others short, setiform, barbed. Six pairs of genital setae setiform, slightly roughened; anal and adanal setae setiform, barbed. Heterotridactylous.
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<emphasis id="E1BCEAE3FFF6C97BFE38FF7FFD410BA7" box="[464,641,223,249]" italics="true" pageId="2" pageNumber="26">Juvenile instars</emphasis>
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. Gastronotum and anogenital region typically folded. Rostral, lamellar and interlamellar setae setiform, smooth (
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<emphasis id="E1BCEAE3FFF6C97BFD9BFEA3FD490A43" box="[627,649,259,284]" italics="true" pageId="2" pageNumber="26">in</emphasis>
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>
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<emphasis id="E1BCEAE3FFF6C97BFD71FEA3FD6D0A42" box="[665,685,259,285]" italics="true" pageId="2" pageNumber="26">le</emphasis>
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>
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<emphasis id="E1BCEAE3FFF6C97BFD55FEA3FD150A43" box="[701,725,259,284]" italics="true" pageId="2" pageNumber="26">ro</emphasis>
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); exobothridial seta
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<emphasis id="E1BCEAE3FFF6C97BFC29FEA3FC1A0A43" box="[961,986,259,284]" italics="true" pageId="2" pageNumber="26">ex</emphasis>
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<subScript id="4F4C34B4FFF6C97BFC32FEB5FC220A7C" attach="left" box="[986,994,277,291]" fontSize="6" pageId="2" pageNumber="26">1</subScript>
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setiform, barbed. Notogastral setae
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<emphasis id="E1BCEAE3FFF6C97BFA69FEA3FA4D0A43" box="[1409,1421,259,284]" italics="true" pageId="2" pageNumber="26">e</emphasis>
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<subScript id="4F4C34B4FFF6C97BFA65FEB5FA550A7C" attach="left" box="[1421,1429,277,291]" fontSize="6" pageId="2" pageNumber="26">2</subScript>
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,
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<emphasis id="E1BCEAE3FFF6C97BFF7FFE87FF650A1E" box="[151,165,295,321]" italics="true" pageId="2" pageNumber="26">h</emphasis>
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<subScript id="4F4C34B4FFF6C97BFF4DFE99FF6D0A18" attach="left" box="[165,173,313,327]" fontSize="6" pageId="2" pageNumber="26">1</subScript>
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,
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<emphasis id="E1BCEAE3FFF6C97BFF54FE87FF0A0A1E" box="[188,202,295,321]" italics="true" pageId="2" pageNumber="26">h</emphasis>
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<subScript id="4F4C34B4FFF6C97BFF22FE99FF120A18" attach="left" box="[202,210,313,327]" fontSize="6" pageId="2" pageNumber="26">2</subScript>
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,
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<emphasis id="E1BCEAE3FFF6C97BFF0AFE87FF300A1F" box="[226,240,295,320]" italics="true" pageId="2" pageNumber="26">p</emphasis>
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<subScript id="4F4C34B4FFF6C97BFF18FE99FF380A18" attach="left" box="[240,248,313,327]" fontSize="6" pageId="2" pageNumber="26">2</subScript>
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subflagellate, smooth; other setae setiform, barbed. Lateral epimeral setae comparatively short, setiform, barbed. Genital formula for nymphal instars: 1-2-4[5].
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</paragraph>
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</subSubSection>
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<subSubSection id="9BD2657AFFF6C97DFF2FFECEFF310E18" lastPageId="4" lastPageNumber="28" pageId="2" pageNumber="26" type="description">
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<paragraph id="D37736F1FFF6C97BFF2FFECEFC420AF2" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="26">
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<emphasis id="E1BCEAE3FFF6C97BFF2FFECEFE010AD6" bold="true" box="[199,449,366,393]" pageId="2" pageNumber="26">Description of adult.</emphasis>
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<emphasis id="E1BCEAE3FFF6C97BFE23FECFFDAD0AD7" box="[459,621,367,392]" italics="true" pageId="2" pageNumber="26">Measurements</emphasis>
|
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. Body length: 735 (
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||
<typeStatus id="0C738853FFF6C97BFCBEFECFFC7C0AD6" box="[854,956,367,393]" pageId="2" pageNumber="26">holotype</typeStatus>
|
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, female), 705–735 (
|
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<specimenCount id="C5CEFD78FFF6C97BFB4CFECFFAD60AD6" box="[1188,1302,367,393]" pageId="2" pageNumber="26" type="female">37 female</specimenCount>
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<typeStatus id="0C738853FFF6C97BFAC8FECFFA510AD6" box="[1312,1425,367,393]" pageId="2" pageNumber="26">paratypes</typeStatus>
|
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); notogastral width: 405 (
|
||
<typeStatus id="0C738853FFF6C97BFE4AFE33FDCA0AF2" box="[418,522,403,429]" pageId="2" pageNumber="26">holotype</typeStatus>
|
||
), 375–420 (
|
||
<specimenCount id="C5CEFD78FFF6C97BFD66FE33FD3E0AF2" box="[654,766,403,429]" pageId="2" pageNumber="26" type="female">37 female</specimenCount>
|
||
<typeStatus id="0C738853FFF6C97BFCEDFE33FCB60AF2" box="[773,886,403,429]" pageId="2" pageNumber="26">paratypes</typeStatus>
|
||
).
|
||
</paragraph>
|
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<paragraph id="D37736F1FFF6C97BFF2FFE17FE490946" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="26">
|
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<emphasis id="E1BCEAE3FFF6C97BFF2FFE17FE850A8F" box="[199,325,439,464]" italics="true" pageId="2" pageNumber="26">Integument</emphasis>
|
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(
|
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<figureCitation id="4BF32A74FFF6C97BFEBEFE17FE190A8E" box="[342,473,439,465]" captionStart="FIGURE 2" captionStartId="3.[151,250,1782,1806]" captionTargetBox="[163,1428,215,1692]" captionTargetId="figure-25@3.[151,1436,181,1758]" captionTargetPageId="3" captionText="FIGURE 2. Tyrphonothrus digeluensis sp. nov., adult: A—dorsal view; B—ventral view (gnathosoma and legs except trochanters not shown); C— right lateral view (gnathosoma and legs not shown). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8213868" httpUri="https://zenodo.org/record/8213868/files/figure.png" pageId="2" pageNumber="26">Figs 2A, C</figureCitation>
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). Body color light brown. Body surface densely porose and covered by gel-like and sparsely microtuberculate cerotegument; lateral part of notogaster (medial to setae
|
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<emphasis id="E1BCEAE3FFF6C97BFBC5FE7BFB870AAB" box="[1069,1095,475,500]" italics="true" pageId="2" pageNumber="26">cp</emphasis>
|
||
,
|
||
<emphasis id="E1BCEAE3FFF6C97BFBBCFE7BFBA00AAB" box="[1108,1120,475,500]" italics="true" pageId="2" pageNumber="26">e</emphasis>
|
||
<subScript id="4F4C34B4FFF6C97BFB89FE4DFBA90AA4" attach="right" box="[1121,1129,493,507]" fontSize="6" pageId="2" pageNumber="26">2</subScript>
|
||
,
|
||
<emphasis id="E1BCEAE3FFF6C97BFB9CFE7BFBBC0AAA" box="[1140,1148,475,501]" italics="true" pageId="2" pageNumber="26">f</emphasis>
|
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<subScript id="4F4C34B4FFF6C97BFB93FE4DFB430AA4" attach="left" box="[1147,1155,493,507]" fontSize="6" pageId="2" pageNumber="26">2</subScript>
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) with longitudinal striate cerotegumental band.
|
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</paragraph>
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<paragraph id="D37736F1FFF6C97BFF2FFD83FE1E09DA" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="26">
|
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<emphasis id="E1BCEAE3FFF6C97BFF2FFD83FE840962" box="[199,324,547,573]" italics="true" pageId="2" pageNumber="26">Prodorsum</emphasis>
|
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(
|
||
<figureCitation id="4BF32A74FFF6C97BFEBCFD83FE140962" box="[340,468,547,573]" captionStart="FIGURE 2" captionStartId="3.[151,250,1782,1806]" captionTargetBox="[163,1428,215,1692]" captionTargetId="figure-25@3.[151,1436,181,1758]" captionTargetPageId="3" captionText="FIGURE 2. Tyrphonothrus digeluensis sp. nov., adult: A—dorsal view; B—ventral view (gnathosoma and legs except trochanters not shown); C— right lateral view (gnathosoma and legs not shown). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8213868" httpUri="https://zenodo.org/record/8213868/files/figure.png" pageId="2" pageNumber="26">Figs 2A, C</figureCitation>
|
||
). Rostrum broadly rounded. Lateral carina well-developed (distinctly visible in dorsal view). Rostral (64–79), lamellar (64–79) and interlamellar (123–131) setae subflagellate, smooth; exobothridial seta
|
||
<emphasis id="E1BCEAE3FFF6C97BFF7FFDCBFF7009DB" box="[151,176,619,644]" italics="true" pageId="2" pageNumber="26">ex</emphasis>
|
||
<subScript id="4F4C34B4FFF6C97BFF58FDDDFF7809D4" attach="left" box="[176,184,637,651]" fontSize="6" pageId="2" pageNumber="26">1</subScript>
|
||
(15–19) setiform, barbed.
|
||
</paragraph>
|
||
<paragraph id="D37736F1FFF6C97BFF2FFD2FFD910866" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="26">
|
||
<emphasis id="E1BCEAE3FFF6C97BFF2FFD2FFE8309F7" box="[199,323,655,680]" italics="true" pageId="2" pageNumber="26">Notogaster</emphasis>
|
||
(
|
||
<figureCitation id="4BF32A74FFF6C97BFEBAFD2FFE1009F6" box="[338,464,655,681]" captionStart="FIGURE 2" captionStartId="3.[151,250,1782,1806]" captionTargetBox="[163,1428,215,1692]" captionTargetId="figure-25@3.[151,1436,181,1758]" captionTargetPageId="3" captionText="FIGURE 2. Tyrphonothrus digeluensis sp. nov., adult: A—dorsal view; B—ventral view (gnathosoma and legs except trochanters not shown); C— right lateral view (gnathosoma and legs not shown). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8213868" httpUri="https://zenodo.org/record/8213868/files/figure.png" pageId="2" pageNumber="26">Figs 2A, C</figureCitation>
|
||
). Anterior margin slightly convex. Medioposterior part of notogaster slightly widening. Four longitudinal ridges present but often poorly observed in dorsal aspect (versus distinctly visible in dorsolateral aspect). Notogastral setae
|
||
<emphasis id="E1BCEAE3FFF6C97BFE29FD77FE0D09AF" box="[449,461,727,752]" italics="true" pageId="2" pageNumber="26">e</emphasis>
|
||
<subScript id="4F4C34B4FFF6C97BFE25FD49FE1509A8" attach="left" box="[461,469,745,759]" fontSize="6" pageId="2" pageNumber="26">2</subScript>
|
||
,
|
||
<emphasis id="E1BCEAE3FFF6C97BFE09FD77FE2F09AE" box="[481,495,727,753]" italics="true" pageId="2" pageNumber="26">h</emphasis>
|
||
<subScript id="4F4C34B4FFF6C97BFE07FD49FE3709A8" attach="left" box="[495,503,745,759]" fontSize="6" pageId="2" pageNumber="26">1</subScript>
|
||
,
|
||
<emphasis id="E1BCEAE3FFF6C97BFDEFFD77FDD509AE" box="[519,533,727,753]" italics="true" pageId="2" pageNumber="26">h</emphasis>
|
||
<subScript id="4F4C34B4FFF6C97BFDFDFD49FDDD09A8" attach="left" box="[533,541,745,759]" fontSize="6" pageId="2" pageNumber="26">2</subScript>
|
||
,
|
||
<emphasis id="E1BCEAE3FFF6C97BFDC4FD77FDFA09AF" box="[556,570,727,752]" italics="true" pageId="2" pageNumber="26">p</emphasis>
|
||
<subScript id="4F4C34B4FFF6C97BFDD2FD49FD8209A8" attach="left" box="[570,578,745,759]" fontSize="6" pageId="2" pageNumber="26">2</subScript>
|
||
(90–105) subflagellate, smooth; others (41–56) setiform, barbed;
|
||
<emphasis id="E1BCEAE3FFF6C97BFAC5FD77FAFB09AF" box="[1325,1339,727,752]" italics="true" pageId="2" pageNumber="26">p</emphasis>
|
||
<subScript id="4F4C34B4FFF6C97BFAD3FD49FA8309A8" attach="left" box="[1339,1347,745,759]" fontSize="6" pageId="2" pageNumber="26">3</subScript>
|
||
located anteriorly to
|
||
<emphasis id="E1BCEAE3FFF6C97BFEC1FD5BFEF7084B" box="[297,311,763,788]" italics="true" pageId="2" pageNumber="26">p</emphasis>
|
||
<subScript id="4F4C34B4FFF6C97BFEDFFCADFEFF0844" attach="left" box="[311,319,781,795]" fontSize="6" pageId="2" pageNumber="26">2</subScript>
|
||
, both distant from each other; alveolus of
|
||
<emphasis id="E1BCEAE3FFF6C97BFCF5FD5BFCE5084A" box="[797,805,763,789]" italics="true" pageId="2" pageNumber="26">f</emphasis>
|
||
<subScript id="4F4C34B4FFF6C97BFCCDFCADFCED0844" attach="left" box="[805,813,781,795]" fontSize="6" pageId="2" pageNumber="26">2</subScript>
|
||
visible. Opisthonotal gland opening and all notogastral lyrifissures distinct in transmitted light.
|
||
</paragraph>
|
||
<paragraph id="D37736F1FFF6C97BFF2FFCE3FC050896" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="26">
|
||
<emphasis id="E1BCEAE3FFF6C97BFF2FFCE3FE960802" box="[199,342,835,861]" italics="true" pageId="2" pageNumber="26">Gnathosoma</emphasis>
|
||
(
|
||
<figureCitation id="4BF32A74FFF6C97BFE8DFCE3FE250802" box="[357,485,835,861]" captionStart="FIGURE 3" captionStartId="4.[151,250,895,919]" captionTargetBox="[238,1389,207,821]" captionTargetId="figure-22@4.[177,1411,181,871]" captionTargetPageId="4" captionText="FIGURE 3. Tyrphonothrus digeluensis sp. nov., adult: A—subcapitulum, ventral view; B—chelicera, right, antiaxial view; C—palp, right, antiaxial view. Scale bars 20 μm." figureDoi="http://doi.org/10.5281/zenodo.8213870" httpUri="https://zenodo.org/record/8213870/files/figure.png" pageId="2" pageNumber="26">Figs 3A–C</figureCitation>
|
||
). Subcapitulum size: 124–127 × 105–112; subcapitular setae (
|
||
<emphasis id="E1BCEAE3FFF6C97BFB71FCE3FB670803" box="[1177,1191,835,860]" italics="true" pageId="2" pageNumber="26">a</emphasis>
|
||
: 15–19;
|
||
<emphasis id="E1BCEAE3FFF6C97BFAE2FCE3FADE0803" box="[1290,1310,835,860]" italics="true" pageId="2" pageNumber="26">m</emphasis>
|
||
: 11–15;
|
||
<emphasis id="E1BCEAE3FFF6C97BFA6EFCE3FA540802" box="[1414,1428,835,861]" italics="true" pageId="2" pageNumber="26">h</emphasis>
|
||
: 13–15) setiform, barbed; three pairs of adoral setae (9–11) filiform, smooth. Palp (length: 142–150) formula: 0-0- 1-3-9(+ω); solenidion bacilliform; postpalpal seta (7) spiniform, roughened. Chelicera (length: 109–112) with two setae:
|
||
<emphasis id="E1BCEAE3FFF6C97BFF36FC0FFEC60896" box="[222,262,943,969]" italics="true" pageId="2" pageNumber="26">cha</emphasis>
|
||
(4) spiniform, barbed;
|
||
<emphasis id="E1BCEAE3FFF6C97BFDE3FC0FFDF30896" box="[523,563,943,969]" italics="true" pageId="2" pageNumber="26">chb</emphasis>
|
||
(19–22) falciform, dorsally barbed.
|
||
</paragraph>
|
||
<paragraph id="D37736F1FFF6C97BFF2FFC73FC820F4E" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="26">
|
||
<emphasis id="E1BCEAE3FFF6C97BFF2FFC73FEBF08B3" box="[199,383,979,1005]" italics="true" pageId="2" pageNumber="26">Epimeral region</emphasis>
|
||
(
|
||
<figureCitation id="4BF32A74FFF6C97BFE67FC73FDC908B2" box="[399,521,979,1005]" captionStart="FIGURE 2" captionStartId="3.[151,250,1782,1806]" captionTargetBox="[163,1428,215,1692]" captionTargetId="figure-25@3.[151,1436,181,1758]" captionTargetPageId="3" captionText="FIGURE 2. Tyrphonothrus digeluensis sp. nov., adult: A—dorsal view; B—ventral view (gnathosoma and legs except trochanters not shown); C— right lateral view (gnathosoma and legs not shown). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8213868" httpUri="https://zenodo.org/record/8213868/files/figure.png" pageId="2" pageNumber="26">Fig. 2B, C</figureCitation>
|
||
). Epimeral setal formula: 3-1-3-3(rarely 4); setae
|
||
<emphasis id="E1BCEAE3FFF6C97BFBD2FC73FB9708B2" box="[1082,1111,979,1005]" italics="true" pageId="2" pageNumber="26">1a</emphasis>
|
||
,
|
||
<emphasis id="E1BCEAE3FFF6C97BFB8DFC73FBBF08B2" box="[1125,1151,979,1005]" italics="true" pageId="2" pageNumber="26">1c</emphasis>
|
||
,
|
||
<emphasis id="E1BCEAE3FFF6C97BFB66FC73FB6908B2" box="[1166,1193,979,1005]" italics="true" pageId="2" pageNumber="26">2a</emphasis>
|
||
,
|
||
<emphasis id="E1BCEAE3FFF6C97BFB50FC73FB1408B2" box="[1208,1236,979,1005]" italics="true" pageId="2" pageNumber="26">3a</emphasis>
|
||
(5–7) penicillate; others (
|
||
<emphasis id="E1BCEAE3FFF6C97BFF04FC57FEC80F4E" box="[236,264,1015,1041]" italics="true" pageId="2" pageNumber="26">1b</emphasis>
|
||
,
|
||
<emphasis id="E1BCEAE3FFF6C97BFEFEFC57FEF20F4E" box="[278,306,1015,1041]" italics="true" pageId="2" pageNumber="26">3b</emphasis>
|
||
,
|
||
<emphasis id="E1BCEAE3FFF6C97BFEA8FC57FE9A0F4E" box="[320,346,1015,1041]" italics="true" pageId="2" pageNumber="26">3c</emphasis>
|
||
,
|
||
<emphasis id="E1BCEAE3FFF6C97BFE80FC57FE450F4E" box="[360,389,1015,1041]" italics="true" pageId="2" pageNumber="26">4a</emphasis>
|
||
,
|
||
<emphasis id="E1BCEAE3FFF6C97BFE7AFC57FE6D0F4E" box="[402,429,1015,1041]" italics="true" pageId="2" pageNumber="26">4c</emphasis>
|
||
: 15–19;
|
||
<emphasis id="E1BCEAE3FFF6C97BFDF8FC57FDED0F4E" box="[528,557,1015,1041]" italics="true" pageId="2" pageNumber="26">4b</emphasis>
|
||
: 9–11) setiform, barbed.
|
||
</paragraph>
|
||
<paragraph id="D37736F1FFF6C97BFF2FFBBBFB100F06" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="26">
|
||
<emphasis id="E1BCEAE3FFF6C97BFF2FFBBBFE570F6B" box="[199,407,1051,1077]" italics="true" pageId="2" pageNumber="26">Anogenital region</emphasis>
|
||
(
|
||
<figureCitation id="4BF32A74FFF6C97BFE43FBBBFDF30F6A" box="[427,563,1051,1077]" captionStart="FIGURE 2" captionStartId="3.[151,250,1782,1806]" captionTargetBox="[163,1428,215,1692]" captionTargetId="figure-25@3.[151,1436,181,1758]" captionTargetPageId="3" captionText="FIGURE 2. Tyrphonothrus digeluensis sp. nov., adult: A—dorsal view; B—ventral view (gnathosoma and legs except trochanters not shown); C— right lateral view (gnathosoma and legs not shown). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8213868" httpUri="https://zenodo.org/record/8213868/files/figure.png" pageId="2" pageNumber="26">Figs 2B, C</figureCitation>
|
||
). Six pairs of genital setae (52–64) setiform, slightly roughened, directed backwards; anal and adanal setae (26–34) setiform, barbed. Anal and adanal lyrifissures distinct.
|
||
</paragraph>
|
||
<paragraph id="D37736F1FFF6C97BFF2FFBC3FD0A0FB6" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="26">
|
||
<emphasis id="E1BCEAE3FFF6C97BFF2FFBC3FF3C0F23" box="[199,252,1123,1148]" italics="true" pageId="2" pageNumber="26">Legs</emphasis>
|
||
(
|
||
<figureCitation id="4BF32A74FFF6C97BFEE5FBC3FE4B0F22" box="[269,395,1123,1149]" captionStart="FIGURE 4" captionStartId="5.[151,250,1753,1777]" captionTargetBox="[177,1416,191,1669]" captionTargetId="figure-25@5.[151,1436,181,1729]" captionTargetPageId="5" captionText="FIGURE 4. Tyrphonothrus digeluensis sp. nov., adult: A—leg I, right, antiaxial view; B—leg II, right, antiaxial view; C—leg III, left, antiaxial view; D—leg IV, left, antiaxial view. Scale bar 50 μm." figureDoi="http://doi.org/10.5281/zenodo.8213872" httpUri="https://zenodo.org/record/8213872/files/figure.png" pageId="2" pageNumber="26">Figs 4A–D</figureCitation>
|
||
). Heterotridactylous; median claw thicker and shorter than lateral claws; all claws dorsally slightly barbed. Formulas of leg setation and solenidia: I (1-5-4-4-11) [1-1-3], II (1-5-3-4-10) [1-1-1], III (2-3-1-2- 10) [0-1-0], IV (1-2-1-2-10) [0-0-0]; homology of setae and solenidia indicated in
|
||
<tableCitation id="9E4A034AFFF6C97BFBCCFB0BFBB70F9A" box="[1060,1143,1195,1221]" captionStart="TABLE 1" captionStartId="10.[151,239,476,500]" captionText="TABLE 1. Development of leg setation of Tyrphonothrus digeluensis sp. nov." pageId="2" pageNumber="26">Table 1</tableCitation>
|
||
. Famulus of tarsus I short, spiniform; all solenidia rod-like, rounded apically.
|
||
</paragraph>
|
||
<paragraph id="D37736F1FFF6C97BFF2FFB52FC170E0A" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="26">
|
||
<emphasis id="E1BCEAE3FFF6C97BFF2FFB52FDF90E53" bold="true" box="[199,569,1266,1292]" pageId="2" pageNumber="26">Description of juvenile instars.</emphasis>
|
||
<emphasis id="E1BCEAE3FFF6C97BFDA9FB53FD230E53" box="[577,739,1267,1292]" italics="true" pageId="2" pageNumber="26">Measurements</emphasis>
|
||
. Total length of larva: 315–330 (n=4), protonymph: 405–450 (n=5), deutonymph: 480–495 (n=4), tritonymph: 540–675 (n=8). Total width of larva: 150–165 (n=4), protonymph: 195–210 (n=5), deutonymph: 240–285 (n=4), tritonymph: 255–315 (n=8).
|
||
</paragraph>
|
||
<paragraph id="D37736F1FFF6C97BFF2FFAFFFB050E9E" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="26">
|
||
<emphasis id="E1BCEAE3FFF6C97BFF2FFAFFFE850E27" box="[199,325,1375,1400]" italics="true" pageId="2" pageNumber="26">Integument</emphasis>
|
||
(
|
||
<figureCitation id="4BF32A74FFF6C97BFEBFFAFFFE190E26" box="[343,473,1374,1401]" captionStart="FIGURE 5" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[194,1347,206,1691]" captionTargetId="figure-22@6.[151,1436,181,1764]" captionTargetPageId="6" captionText="FIGURE 5. Tyrphonothrus digeluensis sp. nov., larva: A—dorsal view; B—ventral view (gnathosoma and legs except trochanters not shown); C— right lateral view (gnathosoma and legs except trochanters not shown). Scale bar 50 μm." figureDoi="http://doi.org/10.5281/zenodo.8213874" httpUri="https://zenodo.org/record/8213874/files/figure.png" pageId="2" pageNumber="26">Figs 5A–C</figureCitation>
|
||
;
|
||
<figureCitation id="4BF32A74FFF6C97BFE0FFAFFFDC90E26" box="[487,521,1375,1401]" captionStart="FIGURE 6" captionStartId="7.[151,250,1664,1688]" captionTargetBox="[177,1416,203,1556]" captionTargetId="figure-25@7.[151,1436,181,1640]" captionTargetPageId="7" captionText="FIGURE 6. Tyrphonothrus digeluensis sp. nov., larva (A–C) and protonymph (D): A—leg I, right, antiaxial view; B—leg II, right, antiaxial view; C—leg III, left, antiaxial view; D—dorsal view. Scale bars 20 μm (A–C), 100 μm (D)." figureDoi="http://doi.org/10.5281/zenodo.8213876" httpUri="https://zenodo.org/record/8213876/files/figure.png" pageId="2" pageNumber="26">6D</figureCitation>
|
||
;
|
||
<figureCitation id="4BF32A74FFF6C97BFDF3FAFFFDA30E26" box="[539,611,1375,1401]" captionStart="FIGURE 7" captionStartId="8.[151,250,1463,1487]" captionTargetBox="[168,1397,227,1314]" captionTargetId="figure-22@8.[151,1436,181,1439]" captionTargetPageId="8" captionText="FIGURE 7. Tyrphonothrus digeluensis sp. nov., proto- (A) and deutonymph (B): A, B— ventral view (gnathosoma and legs except trochanters not shown). Scale bars 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8213878" httpUri="https://zenodo.org/record/8213878/files/figure.png" pageId="2" pageNumber="26">7A, B</figureCitation>
|
||
;
|
||
<figureCitation id="4BF32A74FFF6C97BFD9BFAFFFD7B0E26" box="[627,699,1375,1401]" captionStart="FIGURE 8" captionStartId="9.[151,250,1491,1515]" captionTargetBox="[157,1429,210,1368]" captionTargetId="figure-25@9.[151,1436,181,1468]" captionTargetPageId="9" captionText="FIGURE 8. Tyrphonothrus digeluensis sp. nov., tritonymph: A—dorsal view; B—ventral view (gnathosoma and legs except trochanters not shown). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8213880" httpUri="https://zenodo.org/record/8213880/files/figure.png" pageId="2" pageNumber="26">8A, B</figureCitation>
|
||
). Body color colorless to light yellowish. Body surface densely porose (especially well visible in epimeral region) and partially covered by gel-like and sparsely microtuberculate cerotegument; additionally, gastronotum, anogenital region and basal part of prodorsum folded.
|
||
</paragraph>
|
||
<paragraph id="D37736F1FFF6C97BFF2FFA6BFB490D2A" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="26">
|
||
<emphasis id="E1BCEAE3FFF6C97BFF2FFA6BFE840EBA" box="[199,324,1483,1509]" italics="true" pageId="2" pageNumber="26">Prodorsum</emphasis>
|
||
(
|
||
<figureCitation id="4BF32A74FFF6C97BFEBEFA6BFE1A0EBA" box="[342,474,1482,1509]" captionStart="FIGURE 5" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[194,1347,206,1691]" captionTargetId="figure-22@6.[151,1436,181,1764]" captionTargetPageId="6" captionText="FIGURE 5. Tyrphonothrus digeluensis sp. nov., larva: A—dorsal view; B—ventral view (gnathosoma and legs except trochanters not shown); C— right lateral view (gnathosoma and legs except trochanters not shown). Scale bar 50 μm." figureDoi="http://doi.org/10.5281/zenodo.8213874" httpUri="https://zenodo.org/record/8213874/files/figure.png" pageId="2" pageNumber="26">Figs 5A, C</figureCitation>
|
||
;
|
||
<figureCitation id="4BF32A74FFF6C97BFE01FA6BFDCB0EBA" box="[489,523,1483,1509]" captionStart="FIGURE 6" captionStartId="7.[151,250,1664,1688]" captionTargetBox="[177,1416,203,1556]" captionTargetId="figure-25@7.[151,1436,181,1640]" captionTargetPageId="7" captionText="FIGURE 6. Tyrphonothrus digeluensis sp. nov., larva (A–C) and protonymph (D): A—leg I, right, antiaxial view; B—leg II, right, antiaxial view; C—leg III, left, antiaxial view; D—dorsal view. Scale bars 20 μm (A–C), 100 μm (D)." figureDoi="http://doi.org/10.5281/zenodo.8213876" httpUri="https://zenodo.org/record/8213876/files/figure.png" pageId="2" pageNumber="26">6D</figureCitation>
|
||
;
|
||
<figureCitation id="4BF32A74FFF6C97BFDF5FA6BFD820EBA" box="[541,578,1483,1509]" captionStart="FIGURE 8" captionStartId="9.[151,250,1491,1515]" captionTargetBox="[157,1429,210,1368]" captionTargetId="figure-25@9.[151,1436,181,1468]" captionTargetPageId="9" captionText="FIGURE 8. Tyrphonothrus digeluensis sp. nov., tritonymph: A—dorsal view; B—ventral view (gnathosoma and legs except trochanters not shown). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8213880" httpUri="https://zenodo.org/record/8213880/files/figure.png" pageId="2" pageNumber="26">8A</figureCitation>
|
||
). Relatively short, about 2/3 (in larva and protonymph) and 1/2 length (in deuto- and tritonymph) of gastronotic region. Rostrum broadly rounded. Lateral carina well-developed (distinctly visible in dorsal view). Rostral (LA: 34–37; PN: 41–45; DN: 49–52; TN: 64–75), lamellar (LA: 34–37; PN: 41–45; DN: 49–52; TN: 64–75) and interlamellar (LA: 67–75; PN: 79–90; DN: 101–109; TN: 112–131) setae setiform, smooth; exobothridial seta
|
||
<emphasis id="E1BCEAE3FFF6C97BFE2FF9FBFE200D2B" box="[455,480,1627,1652]" italics="true" pageId="2" pageNumber="26">ex</emphasis>
|
||
<subScript id="4F4C34B4FFF6C97BFE08F9CDFE280D24" attach="left" box="[480,488,1645,1659]" fontSize="6" pageId="2" pageNumber="26">1</subScript>
|
||
(LA: 11; PN: 15; DN: 15–19; TN: 17–19) setiform, barbed.
|
||
</paragraph>
|
||
<paragraph id="D37736F1FFF6C97BFF2FF9DFFF120C76" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="26">
|
||
<emphasis id="E1BCEAE3FFF6C97BFF2FF9DFFE5C0DC7" box="[199,412,1663,1689]" italics="true" pageId="2" pageNumber="26">Gastronotic region</emphasis>
|
||
(
|
||
<figureCitation id="4BF32A74FFF6C97BFE44F9DFFDEC0DC6" box="[428,556,1662,1689]" captionStart="FIGURE 5" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[194,1347,206,1691]" captionTargetId="figure-22@6.[151,1436,181,1764]" captionTargetPageId="6" captionText="FIGURE 5. Tyrphonothrus digeluensis sp. nov., larva: A—dorsal view; B—ventral view (gnathosoma and legs except trochanters not shown); C— right lateral view (gnathosoma and legs except trochanters not shown). Scale bar 50 μm." figureDoi="http://doi.org/10.5281/zenodo.8213874" httpUri="https://zenodo.org/record/8213874/files/figure.png" pageId="2" pageNumber="26">Figs 5A, C</figureCitation>
|
||
;
|
||
<figureCitation id="4BF32A74FFF6C97BFDD1F9DFFD9B0DC6" box="[569,603,1663,1689]" captionStart="FIGURE 6" captionStartId="7.[151,250,1664,1688]" captionTargetBox="[177,1416,203,1556]" captionTargetId="figure-25@7.[151,1436,181,1640]" captionTargetPageId="7" captionText="FIGURE 6. Tyrphonothrus digeluensis sp. nov., larva (A–C) and protonymph (D): A—leg I, right, antiaxial view; B—leg II, right, antiaxial view; C—leg III, left, antiaxial view; D—dorsal view. Scale bars 20 μm (A–C), 100 μm (D)." figureDoi="http://doi.org/10.5281/zenodo.8213876" httpUri="https://zenodo.org/record/8213876/files/figure.png" pageId="2" pageNumber="26">6D</figureCitation>
|
||
;
|
||
<figureCitation id="4BF32A74FFF6C97BFD82F9DFFD4E0DC6" box="[618,654,1663,1689]" captionStart="FIGURE 8" captionStartId="9.[151,250,1491,1515]" captionTargetBox="[157,1429,210,1368]" captionTargetId="figure-25@9.[151,1436,181,1468]" captionTargetPageId="9" captionText="FIGURE 8. Tyrphonothrus digeluensis sp. nov., tritonymph: A—dorsal view; B—ventral view (gnathosoma and legs except trochanters not shown). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8213880" httpUri="https://zenodo.org/record/8213880/files/figure.png" pageId="2" pageNumber="26">8A</figureCitation>
|
||
). Posteriorly rounded. Length of gastronotic setae: LA:
|
||
<emphasis id="E1BCEAE3FFF6C97BFAE1F9DFFAD50DC7" box="[1289,1301,1663,1688]" italics="true" pageId="2" pageNumber="26">e</emphasis>
|
||
<subScript id="4F4C34B4FFF6C97BFAFEF931FADE0DC0" attach="right" box="[1302,1310,1681,1695]" fontSize="6" pageId="2" pageNumber="26">2</subScript>
|
||
,
|
||
<emphasis id="E1BCEAE3FFF6C97BFAC1F9DFFAF70DC6" box="[1321,1335,1663,1689]" italics="true" pageId="2" pageNumber="26">h</emphasis>
|
||
<subScript id="4F4C34B4FFF6C97BFADFF931FAFF0DC0" attach="left" box="[1335,1343,1681,1695]" fontSize="6" pageId="2" pageNumber="26">1</subScript>
|
||
: 45–49,
|
||
<emphasis id="E1BCEAE3FFF6C97BFF7FF903FF650DE2" box="[151,165,1699,1725]" italics="true" pageId="2" pageNumber="26">h</emphasis>
|
||
<subScript id="4F4C34B4FFF6C97BFF4DF915FF6D0D9C" attach="left" box="[165,173,1717,1731]" fontSize="6" pageId="2" pageNumber="26">2</subScript>
|
||
: 30–34, others: 19–22; PN:
|
||
<emphasis id="E1BCEAE3FFF6C97BFE06F903FE3A0DE3" box="[494,506,1699,1724]" italics="true" pageId="2" pageNumber="26">e</emphasis>
|
||
<subScript id="4F4C34B4FFF6C97BFE12F915FDC20D9C" attach="left" box="[506,514,1717,1731]" fontSize="6" pageId="2" pageNumber="26">2</subScript>
|
||
,
|
||
<emphasis id="E1BCEAE3FFF6C97BFDE6F903FDDC0DE2" box="[526,540,1699,1725]" italics="true" pageId="2" pageNumber="26">h</emphasis>
|
||
<subScript id="4F4C34B4FFF6C97BFDF4F915FDE40D9C" attach="left" box="[540,548,1717,1731]" fontSize="6" pageId="2" pageNumber="26">1</subScript>
|
||
,
|
||
<emphasis id="E1BCEAE3FFF6C97BFDDAF903FD800DE2" box="[562,576,1699,1725]" italics="true" pageId="2" pageNumber="26">h</emphasis>
|
||
<subScript id="4F4C34B4FFF6C97BFDA8F915FD880D9C" attach="left" box="[576,584,1717,1731]" fontSize="6" pageId="2" pageNumber="26">2</subScript>
|
||
: 52–60, others: 22–26; DN:
|
||
<emphasis id="E1BCEAE3FFF6C97BFC65F903FC590DE3" box="[909,921,1699,1724]" italics="true" pageId="2" pageNumber="26">e</emphasis>
|
||
<subScript id="4F4C34B4FFF6C97BFC72F915FC620D9C" attach="right" box="[922,930,1717,1731]" fontSize="6" pageId="2" pageNumber="26">2</subScript>
|
||
,
|
||
<emphasis id="E1BCEAE3FFF6C97BFC45F903FC7B0DE2" box="[941,955,1699,1725]" italics="true" pageId="2" pageNumber="26">h</emphasis>
|
||
<subScript id="4F4C34B4FFF6C97BFC53F915FC030D9C" attach="left" box="[955,963,1717,1731]" fontSize="6" pageId="2" pageNumber="26">1</subScript>
|
||
,
|
||
<emphasis id="E1BCEAE3FFF6C97BFC3AF903FC200DE2" box="[978,992,1699,1725]" italics="true" pageId="2" pageNumber="26">h</emphasis>
|
||
<subScript id="4F4C34B4FFF6C97BFC08F915FC280D9C" attach="left" box="[992,1000,1717,1731]" fontSize="6" pageId="2" pageNumber="26">2</subScript>
|
||
: 79–86,
|
||
<emphasis id="E1BCEAE3FFF6C97BFBA4F903FB9A0DE3" box="[1100,1114,1699,1724]" italics="true" pageId="2" pageNumber="26">p</emphasis>
|
||
<subScript id="4F4C34B4FFF6C97BFBB2F915FBA20D9C" attach="left" box="[1114,1122,1717,1731]" fontSize="6" pageId="2" pageNumber="26">2</subScript>
|
||
: 39–41, others: 26–30; TN:
|
||
<emphasis id="E1BCEAE3FFF6C97BFF7FF967FF630DBF" box="[151,163,1735,1760]" italics="true" pageId="2" pageNumber="26">e</emphasis>
|
||
<subScript id="4F4C34B4FFF6C97BFF4CF979FF6C0DB8" attach="right" box="[164,172,1753,1767]" fontSize="6" pageId="2" pageNumber="26">2</subScript>
|
||
,
|
||
<emphasis id="E1BCEAE3FFF6C97BFF50F967FF060DBE" box="[184,198,1735,1761]" italics="true" pageId="2" pageNumber="26">h</emphasis>
|
||
<subScript id="4F4C34B4FFF6C97BFF2EF979FF0E0DB8" attach="left" box="[198,206,1753,1767]" fontSize="6" pageId="2" pageNumber="26">1</subScript>
|
||
,
|
||
<emphasis id="E1BCEAE3FFF6C97BFF36F967FF2C0DBE" box="[222,236,1735,1761]" italics="true" pageId="2" pageNumber="26">h</emphasis>
|
||
<subScript id="4F4C34B4FFF6C97BFF04F979FF340DB8" attach="left" box="[236,244,1753,1767]" fontSize="6" pageId="2" pageNumber="26">2</subScript>
|
||
: 82–94,
|
||
<emphasis id="E1BCEAE3FFF6C97BFEB3F967FEA90DBF" box="[347,361,1735,1760]" italics="true" pageId="2" pageNumber="26">p</emphasis>
|
||
<subScript id="4F4C34B4FFF6C97BFE81F979FEB10DB8" attach="left" box="[361,369,1753,1767]" fontSize="6" pageId="2" pageNumber="26">2</subScript>
|
||
: 49–64, others: 34–37. Setae
|
||
<emphasis id="E1BCEAE3FFF6C97BFD20F967FD140DBF" box="[712,724,1735,1760]" italics="true" pageId="2" pageNumber="26">e</emphasis>
|
||
<subScript id="4F4C34B4FFF6C97BFD3DF979FD1D0DB8" attach="right" box="[725,733,1753,1767]" fontSize="6" pageId="2" pageNumber="26">2</subScript>
|
||
,
|
||
<emphasis id="E1BCEAE3FFF6C97BFD01F967FD370DBE" box="[745,759,1735,1761]" italics="true" pageId="2" pageNumber="26">h</emphasis>
|
||
<subScript id="4F4C34B4FFF6C97BFD1FF979FD3F0DB8" attach="left" box="[759,767,1753,1767]" fontSize="6" pageId="2" pageNumber="26">1</subScript>
|
||
,
|
||
<emphasis id="E1BCEAE3FFF6C97BFCE7F967FCDD0DBE" box="[783,797,1735,1761]" italics="true" pageId="2" pageNumber="26">h</emphasis>
|
||
<subScript id="4F4C34B4FFF6C97BFCF5F979FCE50DB8" attach="left" box="[797,805,1753,1767]" fontSize="6" pageId="2" pageNumber="26">2</subScript>
|
||
(except LA) and
|
||
<emphasis id="E1BCEAE3FFF6C97BFC1BF967FBC10DBF" box="[1011,1025,1735,1760]" italics="true" pageId="2" pageNumber="26">p</emphasis>
|
||
<subScript id="4F4C34B4FFF6C97BFBE9F979FBC90DB8" attach="left" box="[1025,1033,1753,1767]" fontSize="6" pageId="2" pageNumber="26">2</subScript>
|
||
(except PN) subflagellate, smooth; others setiform, barbed; alveolus of
|
||
<emphasis id="E1BCEAE3FFF6C97BFDC0F94BFDF00C5A" box="[552,560,1771,1797]" italics="true" pageId="2" pageNumber="26">f</emphasis>
|
||
<subScript id="4F4C34B4FFF6C97BFDD8F95DFDF80C54" attach="left" box="[560,568,1789,1803]" fontSize="6" pageId="2" pageNumber="26">2</subScript>
|
||
not visible. Opisthonotal gland opening and all cupules distinct in transmitted light.
|
||
</paragraph>
|
||
<paragraph id="D37736F1FFF6C97BFF2FF893FC0D0C82" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="26">
|
||
<emphasis id="E1BCEAE3FFF6C97BFF2FF893FE960C12" box="[199,342,1843,1869]" italics="true" pageId="2" pageNumber="26">Gnathosoma</emphasis>
|
||
. Generally, similar to adult, except sizes and two pairs of adoral setae in larva. Subcapitulum size: LA: 60 × 49; PN: 75 × 60; DN: 86 × 75; TN: 94 × 82; length of subcapitular setae: LA and PN: 11; DN and TN: 15; length of adoral setae: LA and PN: 5; DN: 7; TN: 7–9. Palp length: LA: 41; PN: 45; DN: 49; TN: 52; length of postpalpal seta: LA, PN and DN: 4; TN: 7. Chelicera length: LA: 60; PN: 75; DN: 86; TN: 94; length of
|
||
<emphasis id="E1BCEAE3FFF6C97BFAA8F83FFAA80CE6" box="[1344,1384,1951,1977]" italics="true" pageId="2" pageNumber="26">cha</emphasis>
|
||
: LA and PN: 2; DN and TN: 4; length of
|
||
<emphasis id="E1BCEAE3FFF6C97BFDD9F863FD990C82" box="[561,601,1987,2013]" italics="true" pageId="2" pageNumber="26">chb</emphasis>
|
||
: LA: 9; PN: 11; DN: 15; TN: 19.
|
||
</paragraph>
|
||
<caption id="87B76679FFF7C97AFF7FF956FB4B0C6D" ID-DOI="http://doi.org/10.5281/zenodo.8213868" ID-Zenodo-Dep="8213868" httpUri="https://zenodo.org/record/8213868/files/figure.png" pageId="3" pageNumber="27" startId="3.[151,250,1782,1806]" targetBox="[163,1428,215,1692]" targetPageId="3" targetType="figure">
|
||
<paragraph id="D37736F1FFF7C97AFF7FF956FB4B0C6D" blockId="3.[151,1436,1782,1842]" pageId="3" pageNumber="27">
|
||
<emphasis id="E1BCEAE3FFF7C97AFF7FF956FED90C51" bold="true" box="[151,281,1782,1806]" pageId="3" pageNumber="27">FIGURE 2.</emphasis>
|
||
<taxonomicName id="14C84D72FFF7C97AFECDF956FDF80C51" authority="Ermilov & Rybalov, 2023" authorityName="Ermilov & Rybalov" authorityYear="2023" box="[293,568,1782,1806]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="27" phylum="Arthropoda" rank="species" species="digeluensis" status="sp. nov.">
|
||
<emphasis id="E1BCEAE3FFF7C97AFECDF956FDF80C51" box="[293,568,1782,1806]" italics="true" pageId="3" pageNumber="27">Tyrphonothrus digeluensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E1BCEAE3FFF7C97AFDACF956FD5A0C51" bold="true" box="[580,666,1782,1806]" pageId="3" pageNumber="27">
|
||
<taxonomicNameLabel id="FA8F5798FFF7C97AFDACF956FD5A0C51" box="[580,666,1782,1806]" pageId="3" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, adult: A—dorsal view; B—ventral view (gnathosoma and legs except trochanters not shown); C— right lateral view (gnathosoma and legs not shown). Scale bar 100 μm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D37736F1FFF7C97AFF2FF8C0FAD80CB9" blockId="3.[151,1436,1888,2023]" pageId="3" pageNumber="27">
|
||
<emphasis id="E1BCEAE3FFF7C97AFF2FF8C0FEBE0C25" box="[199,382,1888,1914]" italics="true" pageId="3" pageNumber="27">Epimeral region</emphasis>
|
||
(
|
||
<figureCitation id="4BF32A74FFF7C97AFE63F8C0FDC70C25" box="[395,519,1888,1915]" captionStart="FIGURE 5" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[194,1347,206,1691]" captionTargetId="figure-22@6.[151,1436,181,1764]" captionTargetPageId="6" captionText="FIGURE 5. Tyrphonothrus digeluensis sp. nov., larva: A—dorsal view; B—ventral view (gnathosoma and legs except trochanters not shown); C— right lateral view (gnathosoma and legs except trochanters not shown). Scale bar 50 μm." figureDoi="http://doi.org/10.5281/zenodo.8213874" httpUri="https://zenodo.org/record/8213874/files/figure.png" pageId="3" pageNumber="27">Figs 5B, C</figureCitation>
|
||
;
|
||
<figureCitation id="4BF32A74FFF7C97AFDFAF8C0FD960C24" box="[530,598,1888,1915]" captionStart="FIGURE 7" captionStartId="8.[151,250,1463,1487]" captionTargetBox="[168,1397,227,1314]" captionTargetId="figure-22@8.[151,1436,181,1439]" captionTargetPageId="8" captionText="FIGURE 7. Tyrphonothrus digeluensis sp. nov., proto- (A) and deutonymph (B): A, B— ventral view (gnathosoma and legs except trochanters not shown). Scale bars 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8213878" httpUri="https://zenodo.org/record/8213878/files/figure.png" pageId="3" pageNumber="27">7A, B</figureCitation>
|
||
;
|
||
<figureCitation id="4BF32A74FFF7C97AFD8AF8C0FD450C25" box="[610,645,1888,1914]" captionStart="FIGURE 8" captionStartId="9.[151,250,1491,1515]" captionTargetBox="[157,1429,210,1368]" captionTargetId="figure-25@9.[151,1436,181,1468]" captionTargetPageId="9" captionText="FIGURE 8. Tyrphonothrus digeluensis sp. nov., tritonymph: A—dorsal view; B—ventral view (gnathosoma and legs except trochanters not shown). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8213880" httpUri="https://zenodo.org/record/8213880/files/figure.png" pageId="3" pageNumber="27">8B</figureCitation>
|
||
). Setal formulas for epimeres: larva: 3-1-2 (
|
||
<emphasis id="E1BCEAE3FFF7C97AFB84F8C0FB470C25" box="[1132,1159,1888,1914]" italics="true" pageId="3" pageNumber="27">1c</emphasis>
|
||
as typical scale covering Claparède’s organ); protonymph 3-1-2-1; deutonymph 3-1-2-2, tritonymph 3-1-2[rarely 3]-3. Length of setae: LA and PN:
|
||
<emphasis id="E1BCEAE3FFF7C97AFF11F808FED50C9D" box="[249,277,1960,1986]" italics="true" pageId="3" pageNumber="27">1b</emphasis>
|
||
,
|
||
<emphasis id="E1BCEAE3FFF7C97AFECAF808FEFE0C9D" box="[290,318,1960,1986]" italics="true" pageId="3" pageNumber="27">3b</emphasis>
|
||
: 11, others: 4; DN:
|
||
<emphasis id="E1BCEAE3FFF7C97AFDF4F808FDF80C9D" box="[540,568,1960,1986]" italics="true" pageId="3" pageNumber="27">1b</emphasis>
|
||
,
|
||
<emphasis id="E1BCEAE3FFF7C97AFDADF808FDA10C9D" box="[581,609,1960,1986]" italics="true" pageId="3" pageNumber="27">3b</emphasis>
|
||
,
|
||
<emphasis id="E1BCEAE3FFF7C97AFD87F808FD4B0C9D" box="[623,651,1960,1986]" italics="true" pageId="3" pageNumber="27">4a</emphasis>
|
||
: 11–15,
|
||
<emphasis id="E1BCEAE3FFF7C97AFD04F808FCC80C9D" box="[748,776,1960,1986]" italics="true" pageId="3" pageNumber="27">4a</emphasis>
|
||
: 7, others: 4; TN:
|
||
<emphasis id="E1BCEAE3FFF7C97AFC3DF808FC310C9D" box="[981,1009,1960,1986]" italics="true" pageId="3" pageNumber="27">1b</emphasis>
|
||
,
|
||
<emphasis id="E1BCEAE3FFF7C97AFC16F808FBDA0C9D" box="[1022,1050,1960,1986]" italics="true" pageId="3" pageNumber="27">3b</emphasis>
|
||
,
|
||
<emphasis id="E1BCEAE3FFF7C97AFBC0F808FB820C9D" box="[1064,1090,1960,1986]" italics="true" pageId="3" pageNumber="27">3c</emphasis>
|
||
(if present),
|
||
<emphasis id="E1BCEAE3FFF7C97AFB3AF808FB2E0C9D" box="[1234,1262,1960,1986]" italics="true" pageId="3" pageNumber="27">4a</emphasis>
|
||
,
|
||
<emphasis id="E1BCEAE3FFF7C97AFB14F808FAD60C9D" box="[1276,1302,1960,1986]" italics="true" pageId="3" pageNumber="27">4c</emphasis>
|
||
: 15–17,
|
||
<emphasis id="E1BCEAE3FFF7C97AFA90F808FA550C9D" box="[1400,1429,1960,1986]" italics="true" pageId="3" pageNumber="27">4a</emphasis>
|
||
: 9–11, others: 4–6; setae
|
||
<emphasis id="E1BCEAE3FFF7C97AFE4FF86CFE030CB9" box="[423,451,1996,2022]" italics="true" pageId="3" pageNumber="27">1a</emphasis>
|
||
,
|
||
<emphasis id="E1BCEAE3FFF7C97AFE39F86CFE2C0CB9" box="[465,492,1996,2022]" italics="true" pageId="3" pageNumber="27">1c</emphasis>
|
||
,
|
||
<emphasis id="E1BCEAE3FFF7C97AFE12F86CFDD50CB9" box="[506,533,1996,2022]" italics="true" pageId="3" pageNumber="27">2a</emphasis>
|
||
,
|
||
<emphasis id="E1BCEAE3FFF7C97AFDCCF86CFDFF0CB9" box="[548,575,1996,2022]" italics="true" pageId="3" pageNumber="27">3a</emphasis>
|
||
, and
|
||
<emphasis id="E1BCEAE3FFF7C97AFD95F86CFD590CB9" box="[637,665,1996,2022]" italics="true" pageId="3" pageNumber="27">4a</emphasis>
|
||
(in larva) spiniform, roughened, others setiform, barbed.
|
||
</paragraph>
|
||
<caption id="87B76679FFF0C97DFF7FFCDFFD4608E4" ID-DOI="http://doi.org/10.5281/zenodo.8213870" ID-Zenodo-Dep="8213870" httpUri="https://zenodo.org/record/8213870/files/figure.png" pageId="4" pageNumber="28" startId="4.[151,250,895,919]" targetBox="[238,1389,207,821]" targetPageId="4" targetType="figure">
|
||
<paragraph id="D37736F1FFF0C97DFF7FFCDFFD4608E4" blockId="4.[151,1436,895,955]" pageId="4" pageNumber="28">
|
||
<emphasis id="E1BCEAE3FFF0C97DFF7FFCDFFED608C8" bold="true" box="[151,278,895,919]" pageId="4" pageNumber="28">FIGURE 3.</emphasis>
|
||
<taxonomicName id="14C84D72FFF0C97DFEF7FCDFFDEE08C8" authority="Ermilov & Rybalov, 2023" authorityName="Ermilov & Rybalov" authorityYear="2023" box="[287,558,895,919]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="28" phylum="Arthropoda" rank="species" species="digeluensis" status="sp. nov.">
|
||
<emphasis id="E1BCEAE3FFF0C97DFEF7FCDFFDEE08C8" box="[287,558,895,919]" italics="true" pageId="4" pageNumber="28">Tyrphonothrus digeluensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E1BCEAE3FFF0C97DFDDFFCDFFD4B08C8" bold="true" box="[567,651,895,919]" pageId="4" pageNumber="28">
|
||
<taxonomicNameLabel id="FA8F5798FFF0C97DFDDFFCDFFD4B08C8" box="[567,651,895,919]" pageId="4" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, adult: A—subcapitulum, ventral view; B—chelicera, right, antiaxial view; C—palp, right, antiaxial view. Scale bars 20 μm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D37736F1FFF0C97DFF2FFC49FE8E0F30" blockId="4.[151,1437,1001,2036]" pageId="4" pageNumber="28">
|
||
<emphasis id="E1BCEAE3FFF0C97DFF2FFC49FE530F5C" box="[199,403,1001,1027]" italics="true" pageId="4" pageNumber="28">Anogenital region</emphasis>
|
||
(
|
||
<figureCitation id="4BF32A74FFF0C97DFE4CFC49FDE60F5C" box="[420,550,1001,1028]" captionStart="FIGURE 5" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[194,1347,206,1691]" captionTargetId="figure-22@6.[151,1436,181,1764]" captionTargetPageId="6" captionText="FIGURE 5. Tyrphonothrus digeluensis sp. nov., larva: A—dorsal view; B—ventral view (gnathosoma and legs except trochanters not shown); C— right lateral view (gnathosoma and legs except trochanters not shown). Scale bar 50 μm." figureDoi="http://doi.org/10.5281/zenodo.8213874" httpUri="https://zenodo.org/record/8213874/files/figure.png" pageId="4" pageNumber="28">Figs 5B, C</figureCitation>
|
||
;
|
||
<figureCitation id="4BF32A74FFF0C97DFDDCFC49FDBC0F5B" box="[564,636,1001,1028]" captionStart="FIGURE 7" captionStartId="8.[151,250,1463,1487]" captionTargetBox="[168,1397,227,1314]" captionTargetId="figure-22@8.[151,1436,181,1439]" captionTargetPageId="8" captionText="FIGURE 7. Tyrphonothrus digeluensis sp. nov., proto- (A) and deutonymph (B): A, B— ventral view (gnathosoma and legs except trochanters not shown). Scale bars 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8213878" httpUri="https://zenodo.org/record/8213878/files/figure.png" pageId="4" pageNumber="28">7A, B</figureCitation>
|
||
;
|
||
<figureCitation id="4BF32A74FFF0C97DFD62FC49FD6E0F5C" box="[650,686,1001,1027]" captionStart="FIGURE 8" captionStartId="9.[151,250,1491,1515]" captionTargetBox="[157,1429,210,1368]" captionTargetId="figure-25@9.[151,1436,181,1468]" captionTargetPageId="9" captionText="FIGURE 8. Tyrphonothrus digeluensis sp. nov., tritonymph: A—dorsal view; B—ventral view (gnathosoma and legs except trochanters not shown). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8213880" httpUri="https://zenodo.org/record/8213880/files/figure.png" pageId="4" pageNumber="28">8B</figureCitation>
|
||
). Ontogeny of genital (PN: 22–26; DN: 26–30; TN: 37–45), anal (TN: 26–30) and adanal (DN: 19; TN: 26–30) setal formulas, proto- to tritonymphs: 1-2-4[rarely 5], 0-0-1, 0-3-3, respectively; all setae setiform, genital setae slightly roughened, anal and adanal setae barbed. Anal and adanal cupules distinct.
|
||
</paragraph>
|
||
<paragraph id="D37736F1FFF0C97DFF2FFBDAFF310E18" blockId="4.[151,1437,1001,2036]" pageId="4" pageNumber="28">
|
||
<emphasis id="E1BCEAE3FFF0C97DFF2FFBDAFF3C0FCC" box="[199,252,1146,1171]" italics="true" pageId="4" pageNumber="28">Legs</emphasis>
|
||
(
|
||
<figureCitation id="4BF32A74FFF0C97DFEE5FBD9FE4F0FCC" box="[269,399,1145,1171]" captionStart="FIGURE 6" captionStartId="7.[151,250,1664,1688]" captionTargetBox="[177,1416,203,1556]" captionTargetId="figure-25@7.[151,1436,181,1640]" captionTargetPageId="7" captionText="FIGURE 6. Tyrphonothrus digeluensis sp. nov., larva (A–C) and protonymph (D): A—leg I, right, antiaxial view; B—leg II, right, antiaxial view; C—leg III, left, antiaxial view; D—dorsal view. Scale bars 20 μm (A–C), 100 μm (D)." figureDoi="http://doi.org/10.5281/zenodo.8213876" httpUri="https://zenodo.org/record/8213876/files/figure.png" pageId="4" pageNumber="28">Figs 6A–C</figureCitation>
|
||
). Claw of each leg dorsally slightly barbed. Formulas of leg setation and solenidia: larva I (0-2-3-4-11) [1-1-1], II (0-2-2-3-10) [1-1-1], III (0-2-1-2-10) [0-1-0]; protonymph I (0-2-3-4-11) [1-1-2], II (0-2-2- 3-10) [1-1-1], III (1-2-1-2-10) [0-1-0], IV (0-0-0-0-7) [0-0-0]; deutonymph I (1-4-3-4-11) [1-1-2], II (1-4-2-4-10) [1-1-1], III (2-2-1-2-10) [0-1-0], IV (0-1-1-2-10) [0-0-0]; tritonymph I (1-5-4-4-11) [1-1-3], II (1-5-3-4-10) [1-1-1], III (2-2[rarely 3]-1-2-10) [0-1-0], IV (1-1[rarely 2]-1-2-10) [0-0-0]; homologies of setae and solenidia indicated in
|
||
<tableCitation id="9E4A034AFFF0C97DFF7FFA8DFF2B0E18" box="[151,235,1325,1351]" captionStart="TABLE 1" captionStartId="10.[151,239,476,500]" captionText="TABLE 1. Development of leg setation of Tyrphonothrus digeluensis sp. nov." pageId="4" pageNumber="28">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9BD2657AFFF0C97DFF2FFAF1FF3B0D38" pageId="4" pageNumber="28" type="materials_examined">
|
||
<paragraph id="D37736F1FFF0C97DFF2FFAF1FD840E88" blockId="4.[151,1437,1001,2036]" pageId="4" pageNumber="28">
|
||
<emphasis id="E1BCEAE3FFF0C97DFF2FFAF1FEBB0E34" bold="true" box="[199,379,1361,1387]" pageId="4" pageNumber="28">Type material.</emphasis>
|
||
<materialsCitation id="63A03CACFFF0C97DFE61FAF1FD840E88" collectingDate="2021-11-28" collectorName="L. B. Rybalov" country="Ethiopia" county="Arsi Zone" elevation="3882" latitude="7.82325" location="Digelu" longLatPrecision="1" longitude="39.392475" municipality="Arsi Mountains National Park" pageId="4" pageNumber="28" specimenCount="59" specimenCount-juvenile="21" specimenCount-larva="1" stateProvince="Oromiya" typeStatus="holotype">
|
||
<typeStatus id="0C738853FFF0C97DFE61FAF1FE310E34" box="[393,497,1361,1387]" pageId="4" pageNumber="28">Holotype</typeStatus>
|
||
(female),
|
||
<specimenCount id="C5CEFD78FFF0C97DFD9AFAF1FCC60E33" box="[626,774,1361,1388]" pageId="4" pageNumber="28" type="generic" typeStatus="paratypes">37 paratypes</specimenCount>
|
||
(all females) and
|
||
<specimenCount id="C5CEFD78FFF0C97DFC03FAF1FBAE0E34" box="[1003,1134,1361,1387]" pageId="4" pageNumber="28" type="juvenile">21 juvenile</specimenCount>
|
||
instars (
|
||
<specimenCount id="C5CEFD78FFF0C97DFB32FAF1FA9F0E34" box="[1242,1375,1361,1387]" pageId="4" pageNumber="28" type="larva">four larvae</specimenCount>
|
||
, five protonymphs, four deutonymphs, eight tritonymphs):
|
||
<collectingCountry id="ABDF7661FFF0C97DFD03FAD5FC8D0ED0" box="[747,845,1397,1423]" name="Ethiopia" pageId="4" pageNumber="28">Ethiopia</collectingCountry>
|
||
,
|
||
<collectingRegion id="110CF813FFF0C97DFCBFFAD5FBC60ED0" box="[855,1030,1397,1423]" country="Ethiopia" name="Oromiya" pageId="4" pageNumber="28">Oromia Region</collectingRegion>
|
||
,
|
||
<collectingCounty id="3A164E7DFFF0C97DFBE6FAD5FB400ECF" box="[1038,1152,1397,1424]" pageId="4" pageNumber="28">Arsi Zone</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="3313AC8BFFF0C97DFB60FAD5FF0D0EEC" pageId="4" pageNumber="28">Arsi Mountains National Park</collectingMunicipality>
|
||
, mountain near
|
||
<location id="D617602AFFF0C97DFE6EFA39FE150EEC" LSID="urn:lsid:plazi:treatment:5B6187E7FFF5C97DFF7FF8EAFD090CAB:D617602AFFF0C97DFE6EFA39FE150EEC" box="[390,469,1433,1459]" country="Ethiopia" county="Arsi Zone" latitude="7.82325" longLatPrecision="1" longitude="39.392475" municipality="Arsi Mountains National Park" name="Digelu" pageId="4" pageNumber="28" stateProvince="Oromiya">Digelu</location>
|
||
,
|
||
<geoCoordinate id="B6FC5036FFF0C97DFE0BFA3BFDBE0EE9" box="[483,638,1435,1462]" degrees="07" direction="north" minutes="49" orientation="latitude" pageId="4" pageNumber="28" precision="1" seconds="23.7" value="7.82325">07º49′23.7′′N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="B6FC5036FFF0C97DFD63FA3BFCF00EE9" box="[651,816,1435,1462]" degrees="039" direction="east" minutes="23" orientation="longitude" pageId="4" pageNumber="28" precision="1" seconds="32.9" value="39.392475">039º23′32.9′′E</geoCoordinate>
|
||
,
|
||
<elevation id="58E5D1C2FFF0C97DFCD6FA39FC140EEC" box="[830,980,1433,1460]" metricMagnitude="3" metricUnit="m" metricValue="3.882" pageId="4" pageNumber="28" unit="m" value="3882.0">
|
||
<quantity id="14309B14FFF0C97DFCD6FA39FC550EEB" box="[830,917,1433,1460]" metricMagnitude="3" metricUnit="m" metricValue="3.882" pageId="4" pageNumber="28" unit="m" value="3882.0">3882 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
, mosses on the swamp at the top of the mountain,
|
||
<date id="A7761031FFF0C97DFEE7FA1DFE4C0E88" box="[271,396,1469,1495]" pageId="4" pageNumber="28" value="2021-11-28">
|
||
<collectingDate id="B732E9D9FFF0C97DFEE7FA1DFE4C0E88" box="[271,396,1469,1495]" pageId="4" pageNumber="28" value="2021-11-28">28.11.2021</collectingDate>
|
||
</date>
|
||
(
|
||
<collectorName id="7E3D5327FFF0C97DFE74FA1EFDF80E88" box="[412,568,1469,1496]" pageId="4" pageNumber="28">L.B. Rybalov</collectorName>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="D37736F1FFF0C97DFF2FFA41FF3B0D38" blockId="4.[151,1437,1001,2036]" pageId="4" pageNumber="28">
|
||
<emphasis id="E1BCEAE3FFF0C97DFF2FFA41FE4C0EA4" bold="true" box="[199,396,1505,1531]" pageId="4" pageNumber="28">Type deposition.</emphasis>
|
||
The
|
||
<typeStatus id="0C738853FFF0C97DFE2FFA41FDE90EA4" box="[455,553,1505,1531]" pageId="4" pageNumber="28">holotype</typeStatus>
|
||
is deposited in the collection of the Senckenberg Museum of Natural History, Görlitz,
|
||
<collectingCountry id="ABDF7661FFF0C97DFF1DF9A5FEA10D40" box="[245,353,1541,1567]" name="Germany" pageId="4" pageNumber="28">Germany</collectingCountry>
|
||
;
|
||
<materialsCitation id="63A03CACFFF0C97DFE84F9A5FD900D1C" country="Russia" location="University Museum of Zoology" pageId="4" pageNumber="28" specimenCount="37" stateProvince="Tyumen'" typeStatus="paratype">
|
||
<specimenCount id="C5CEFD78FFF0C97DFE84F9A5FE3A0D7F" box="[364,506,1541,1568]" pageId="4" pageNumber="28" type="generic" typeStatus="paratypes">37 paratypes</specimenCount>
|
||
and juvenile instars are deposited in the collection of the
|
||
<collectingRegion id="110CF813FFF0C97DFB6CF9A6FADF0D40" box="[1156,1311,1541,1568]" country="Russia" name="Tyumen'" pageId="4" pageNumber="28">Tyumen State</collectingRegion>
|
||
<location id="D617602AFFF0C97DFACEF9A5FE460D1C" LSID="urn:lsid:plazi:treatment:5B6187E7FFF5C97DFF7FF8EAFD090CAB:D617602AFFF0C97DFACEF9A5FE460D1C" country="Russia" name="University Museum of Zoology" pageId="4" pageNumber="28" stateProvince="Tyumen'">University Museum of Zoology</location>
|
||
,
|
||
<collectingRegion id="110CF813FFF0C97DFE7DF98AFE330D1B" box="[405,499,1578,1604]" country="Russia" name="Tyumen'" pageId="4" pageNumber="28">Tyumen</collectingRegion>
|
||
,
|
||
<collectingCountry id="ABDF7661FFF0C97DFDEAF989FD900D1C" box="[514,592,1577,1603]" name="Russia" pageId="4" pageNumber="28">Russia</collectingCountry>
|
||
</materialsCitation>
|
||
. All specimens are preserved in 70% solution of ethanol with a drop of glycerol.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9BD2657AFFF0C97DFF2FF9D1FEC40DF0" pageId="4" pageNumber="28" type="etymology">
|
||
<paragraph id="D37736F1FFF0C97DFF2FF9D1FEC40DF0" blockId="4.[151,1437,1001,2036]" pageId="4" pageNumber="28">
|
||
<emphasis id="E1BCEAE3FFF0C97DFF2FF9D1FE8E0DD4" bold="true" box="[199,334,1649,1675]" pageId="4" pageNumber="28">Etymology.</emphasis>
|
||
The species name
|
||
<taxonomicName id="14C84D72FFF0C97DFDD7F9D1FD7D0DD4" authorityName="Ermilov & Rybalov" authorityYear="2023" box="[575,701,1649,1675]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="28" phylum="Arthropoda" rank="species" species="digeluensis">
|
||
<emphasis id="E1BCEAE3FFF0C97DFDD7F9D1FD7D0DD4" box="[575,701,1649,1675]" italics="true" pageId="4" pageNumber="28">digeluensis</emphasis>
|
||
</taxonomicName>
|
||
refers to the Digelu town, near which the new species was collected.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9BD2657AFFF0C97DFF2FF919FD090CAB" pageId="4" pageNumber="28" type="diagnosis">
|
||
<paragraph id="D37736F1FFF0C97DFF2FF919FD090CAB" blockId="4.[151,1437,1001,2036]" pageId="4" pageNumber="28">
|
||
<emphasis id="E1BCEAE3FFF0C97DFF2FF919FE140D8C" bold="true" box="[199,468,1721,1747]" pageId="4" pageNumber="28">Differential diagnosis.</emphasis>
|
||
Adult of
|
||
<taxonomicName id="14C84D72FFF0C97DFDBCF919FC440D8C" authority="Ermilov & Rybalov, 2023" authorityName="Ermilov & Rybalov" authorityYear="2023" box="[596,900,1721,1747]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="28" phylum="Arthropoda" rank="species" species="digeluensis" status="sp. nov.">
|
||
<emphasis id="E1BCEAE3FFF0C97DFDBCF919FC440D8C" box="[596,900,1721,1747]" italics="true" pageId="4" pageNumber="28">Tyrphonothrus digeluensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E1BCEAE3FFF0C97DFC7AF919FC320D8C" bold="true" box="[914,1010,1721,1747]" pageId="4" pageNumber="28">
|
||
<taxonomicNameLabel id="FA8F5798FFF0C97DFC7AF919FC320D8C" box="[914,1010,1721,1747]" pageId="4" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is morphologically most similar to
|
||
<taxonomicName id="14C84D72FFF0C97DFF7FF97DFD230DA8" authority="(Yamamoto & Coetzee, 2004)" baseAuthorityName="Yamamoto & Coetzee" baseAuthorityYear="2004" box="[151,739,1757,1784]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="28" phylum="Arthropoda" rank="species" species="obesus">
|
||
<emphasis id="E1BCEAE3FFF0C97DFF7FF97DFE4E0DA8" box="[151,398,1757,1783]" italics="true" pageId="4" pageNumber="28">Tyrphonothrus obesus</emphasis>
|
||
(
|
||
<bibRefCitation id="B7594B00FFF0C97DFE76F97EFD1C0DA8" author="Yamamoto, Y. & Coetzee, L." box="[414,732,1757,1784]" pageId="4" pageNumber="28" pagination="13 - 25" refId="ref6909" refString="Yamamoto, Y. & Coetzee, L. (2004) The Trimalaconothrus species from South Africa (Acari, Oribatida, Malaconothridae). Acta Arachnologica, 53 (1), 13 - 25. https: // doi. org / 10.2476 / asjaa. 53.13" type="journal article" year="2004">Yamamoto & Coetzee, 2004</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="14C84D72FFF0C97DFCF0F97EFB0C0DA8" authority="(Yamamoto & Coetzee, 2004)" baseAuthorityName="Yamamoto & Coetzee" baseAuthorityYear="2004" box="[792,1228,1757,1784]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="28" phylum="Arthropoda" rank="species" species="rectus">
|
||
<emphasis id="E1BCEAE3FFF0C97DFCF0F97EFCB60DA8" box="[792,886,1758,1783]" italics="true" pageId="4" pageNumber="28">T. rectus</emphasis>
|
||
(
|
||
<bibRefCitation id="B7594B00FFF0C97DFC6EF97EFB040DA8" author="Yamamoto, Y. & Coetzee, L." box="[902,1220,1757,1784]" pageId="4" pageNumber="28" pagination="13 - 25" refId="ref6909" refString="Yamamoto, Y. & Coetzee, L. (2004) The Trimalaconothrus species from South Africa (Acari, Oribatida, Malaconothridae). Acta Arachnologica, 53 (1), 13 - 25. https: // doi. org / 10.2476 / asjaa. 53.13" type="journal article" year="2004">Yamamoto & Coetzee, 2004</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="ABDF7661FFF0C97DFAE7F97DFA5D0DA8" box="[1295,1437,1757,1783]" name="South Africa" pageId="4" pageNumber="28">South Africa</collectingCountry>
|
||
in having large body size, smooth rostral, lamellar and interlamellar setae, slightly widening medioposterior part of notogaster, different length of notogastral setae, six pairs of genital setae, two pairs of longitudinal notogastral ridges, tridactylous legs, and in the absence of foveolate ornamentation on the notogaster. However, the new species differ from both by the barbed (versus smooth) notogastral setae
|
||
<emphasis id="E1BCEAE3FFF0C97DFC9DF8CEFC410CD8" box="[885,897,1902,1927]" italics="true" pageId="4" pageNumber="28">c</emphasis>
|
||
<subScript id="4F4C34B4FFF0C97DFC6AF820FC4A0CD1" attach="right" box="[898,906,1920,1934]" fontSize="6" pageId="4" pageNumber="28">1</subScript>
|
||
,
|
||
<emphasis id="E1BCEAE3FFF0C97DFC71F8CEFC650CD8" box="[921,933,1902,1927]" italics="true" pageId="4" pageNumber="28">c</emphasis>
|
||
<subScript id="4F4C34B4FFF0C97DFC4DF820FC6D0CD1" attach="left" box="[933,941,1920,1934]" fontSize="6" pageId="4" pageNumber="28">2</subScript>
|
||
,
|
||
<emphasis id="E1BCEAE3FFF0C97DFC54F8CEFC080CD8" box="[956,968,1902,1927]" italics="true" pageId="4" pageNumber="28">c</emphasis>
|
||
<subScript id="4F4C34B4FFF0C97DFC20F820FC100CD1" attach="left" box="[968,976,1920,1934]" fontSize="6" pageId="4" pageNumber="28">3</subScript>
|
||
,
|
||
<emphasis id="E1BCEAE3FFF0C97DFC37F8CEFC390CD8" box="[991,1017,1902,1927]" italics="true" pageId="4" pageNumber="28">cp</emphasis>
|
||
,
|
||
<emphasis id="E1BCEAE3FFF0C97DFBE0F8CDFBD60CD8" box="[1032,1046,1901,1927]" italics="true" pageId="4" pageNumber="28">d</emphasis>
|
||
<subScript id="4F4C34B4FFF0C97DFBFEF820FBDE0CD1" attach="left" box="[1046,1054,1920,1934]" fontSize="6" pageId="4" pageNumber="28">1</subScript>
|
||
,
|
||
<emphasis id="E1BCEAE3FFF0C97DFBC5F8CDFBFB0CD8" box="[1069,1083,1901,1927]" italics="true" pageId="4" pageNumber="28">d</emphasis>
|
||
<subScript id="4F4C34B4FFF0C97DFBD3F820FB830CD1" attach="left" box="[1083,1091,1920,1934]" fontSize="6" pageId="4" pageNumber="28">2</subScript>
|
||
,
|
||
<emphasis id="E1BCEAE3FFF0C97DFBBAF8CEFB9E0CD8" box="[1106,1118,1902,1927]" italics="true" pageId="4" pageNumber="28">e</emphasis>
|
||
<subScript id="4F4C34B4FFF0C97DFBB7F820FBA70CD1" attach="right" box="[1119,1127,1920,1934]" fontSize="6" pageId="4" pageNumber="28">1</subScript>
|
||
,
|
||
<emphasis id="E1BCEAE3FFF0C97DFB9DF8CDFBBD0CD8" box="[1141,1149,1901,1927]" italics="true" pageId="4" pageNumber="28">f</emphasis>
|
||
<subScript id="4F4C34B4FFF0C97DFB95F820FB450CD1" attach="left" box="[1149,1157,1920,1934]" fontSize="6" pageId="4" pageNumber="28">2</subScript>
|
||
,
|
||
<emphasis id="E1BCEAE3FFF0C97DFB7CF8CEFB620CD8" box="[1172,1186,1902,1927]" italics="true" pageId="4" pageNumber="28">p</emphasis>
|
||
<subScript id="4F4C34B4FFF0C97DFB4AF820FB6A0CD1" attach="left" box="[1186,1194,1920,1934]" fontSize="6" pageId="4" pageNumber="28">1</subScript>
|
||
,
|
||
<emphasis id="E1BCEAE3FFF0C97DFB51F8CEFB070CD8" box="[1209,1223,1902,1927]" italics="true" pageId="4" pageNumber="28">p</emphasis>
|
||
<subScript id="4F4C34B4FFF0C97DFB2FF820FB0F0CD1" attach="left" box="[1223,1231,1920,1934]" fontSize="6" pageId="4" pageNumber="28">3</subScript>
|
||
and exobothridial seta
|
||
<emphasis id="E1BCEAE3FFF0C97DFF20F832FF210CF4" box="[200,225,1938,1963]" italics="true" pageId="4" pageNumber="28">ex</emphasis>
|
||
<subScript id="4F4C34B4FFF0C97DFF09F804FF290CED" attach="left" box="[225,233,1956,1970]" fontSize="6" pageId="4" pageNumber="28">1</subScript>
|
||
, and the slightly developed (versus distinctly) notogastral ridges. Also,
|
||
<taxonomicName id="14C84D72FFF0C97DFBE9F832FBA70CF4" baseAuthorityName="Yamamoto & Coetzee" baseAuthorityYear="2004" box="[1025,1127,1937,1963]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="28" phylum="Arthropoda" rank="species" species="obesus">
|
||
<emphasis id="E1BCEAE3FFF0C97DFBE9F832FBA70CF4" box="[1025,1127,1937,1963]" italics="true" pageId="4" pageNumber="28">T. obesus</emphasis>
|
||
</taxonomicName>
|
||
has smooth (versus barbed) epimeral and anoadanal setae;
|
||
<taxonomicName id="14C84D72FFF0C97DFE1AF816FD910C90" baseAuthorityName="Yamamoto & Coetzee" baseAuthorityYear="2004" box="[498,593,1974,1999]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="28" phylum="Arthropoda" rank="species" species="rectus">
|
||
<emphasis id="E1BCEAE3FFF0C97DFE1AF816FD910C90" box="[498,593,1974,1999]" italics="true" pageId="4" pageNumber="28">T. rectus</emphasis>
|
||
</taxonomicName>
|
||
has heavily barbed (versus barbed) epimeral setae and subcapitular seta
|
||
<emphasis id="E1BCEAE3FFF0C97DFA6FF815FA550C90" box="[1415,1429,1973,1999]" italics="true" pageId="4" pageNumber="28">h</emphasis>
|
||
, and smooth (versus barbed) anal and adanal setae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |