480 lines
76 KiB
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480 lines
76 KiB
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<document ID-CLB-Dataset="20724" ID-DOI="http://dx.doi.org/10.3897/dez.69.79931" ID-GBIF-Dataset="e58ae50b-2b62-4c1c-b1fd-09b0f6aa001d" ID-Pensoft-Pub="1860-1324-1-19" ID-Pensoft-UUID="024B3AF3339C5F8CAA644EA4F78EA082" ID-ZooBank="02E8488BDDA7464CABC239424200939E" ModsDocID="1860-1324-69-1-19" checkinTime="1644448212315" checkinUser="pensoft" docAuthor="Schmidt, Joachim, Scholz, Stephan & Will, Kipling" docDate="2022" docId="5B07C8171ABA53069E983FDE8CF9A50B" docLanguage="en" docName="DeutEntomolZeit 69(1): 19-44" docOrigin="Deutsche Entomologische Zeitschrift 69 (1)" docPubDate="2022-02-09" docSource="http://dx.doi.org/10.3897/dez.69.79931" docTitle="Quasicalathus agonicollis Schmidt & Will 2022, sp. nov." docType="treatment" docUuid="0555D647-EDC0-4A5F-BDDE-05FA1C5BB140" docUuidSource="ZooBank" docVersion="6" id="024B3AF3339C5F8CAA644EA4F78EA082" lastPageNumber="19" masterDocId="024B3AF3339C5F8CAA644EA4F78EA082" masterDocTitle="Character analysis and descriptions of Eocene sphodrine fossils (Coleoptera, Carabidae) using light microscopy, micro-CT scanning, and 3 D imaging" masterLastPageNumber="44" masterPageNumber="19" pageNumber="19" updateTime="1732797172808" updateUser="ExternalLinkService">
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<mods:titleInfo id="2179303A890BCE4985499BE2C02A2F5C">
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<mods:title id="510BB2E3756DE1F4BBF40E3F6CC59CE8">Character analysis and descriptions of Eocene sphodrine fossils (Coleoptera, Carabidae) using light microscopy, micro-CT scanning, and 3 D imaging</mods:title>
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<mods:name id="C97A34793083FB0C4FB06E3E0731B001" type="personal">
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<mods:roleTerm id="8B43ABEE16393A63EF9AE62FF72F3A1D">Author</mods:roleTerm>
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<mods:namePart id="630AFC9A52CCF1E2F0B6E6121FFDEE43">Schmidt, Joachim</mods:namePart>
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<mods:affiliation id="03F687C2F9BA147A121CA880F07823EA">University of Rostock, Institute of Biosciences, General and Systematic Zoology, Universitaetsplatz 2, 18055 Rostock, German</mods:affiliation>
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<mods:nameIdentifier id="9B488163EF849323ABF01890F13C909E" type="email">agonumschmidt@hotmail.com</mods:nameIdentifier>
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<mods:roleTerm id="41C1BB2064D14DFC5DA5C5223E0E48B2">Author</mods:roleTerm>
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<mods:namePart id="B8CDFE48D1BFDC7672E91A20D4E650A0">Scholz, Stephan</mods:namePart>
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<mods:affiliation id="7B5CA5FACB3EC30205BF758C02729589">University of Rostock, Institute of Biosciences, General and Systematic Zoology, Universitaetsplatz 2, 18055 Rostock, German</mods:affiliation>
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<mods:name id="B9CDEEE8BA6061CC20D8C37C809A5F16" type="personal">
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<mods:namePart id="B77F13D27AE5906024C674E1294B57F7">Will, Kipling</mods:namePart>
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<mods:nameIdentifier id="E2A8655B30F7FF6D4E70ED1DDA09B6DA" type="ORCID">https://orcid.org/0000-0002-7056-9011</mods:nameIdentifier>
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<mods:affiliation id="8A7C037C51E5116D3E6BCEEBA6FCBBEE">Essig Museum of Entomology, 1101 Valley Life Sciences Building, # 4780 University of California, Berkeley, CA 94720 - 4780, USA</mods:affiliation>
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<mods:typeOfResource id="E0D158DF46FD2492DB1D7DB4D13BC03E">text</mods:typeOfResource>
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<mods:title id="1B56A02C1E373ADFCD9A70E45A28431D">Deutsche Entomologische Zeitschrift</mods:title>
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<mods:date id="9A0803B8937A2ADA00D30FB1FBF52ECB">2022</mods:date>
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<mods:number id="5236436BEC98029F7AF0533421156B15">2022-02-09</mods:number>
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<mods:number id="05EF0599CF2D2056851DAC4920542B69">69</mods:number>
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<mods:start id="025B27CA706A7D9A043778C5CA8CA082">19</mods:start>
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<mods:url id="6A054218653F7525C419A4A745E269FF">http://dx.doi.org/10.3897/dez.69.79931</mods:url>
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<mods:classification id="E7D61E3A5B55ED47A14FE07FF7EB7855">journal article</mods:classification>
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<mods:identifier id="87E0CFA67CC8F06B766EE73CE3256898" type="DOI">http://dx.doi.org/10.3897/dez.69.79931</mods:identifier>
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<treatment id="5B07C8171ABA53069E983FDE8CF9A50B" ID-GBIF-Taxon="193323627" LSID="urn:lsid:zoobank.org:act:0555D647-EDC0-4A5F-BDDE-05FA1C5BB140" httpUri="http://treatment.plazi.org/id/5B07C8171ABA53069E983FDE8CF9A50B" lastPageNumber="19" pageId="0" pageNumber="19" scope_family="Carabidae" scope_order="Coleoptera">
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<subSubSection id="5EEACF6E3FB87B3047F84BF744E258C3" pageId="0" pageNumber="19" type="nomenclature">
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<paragraph id="D7E9BA214B3123E13ECFE262E76352B8" pageId="0" pageNumber="19">
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<taxonomicName id="B229E1BAD528773868430C1B54EEE18B" LSID="http://zoobank.org0555D647-EDC0-4A5F-BDDE-05FA1C5BB140" authority="Schmidt & Will" authorityName="Schmidt & Will" authorityYear="2022" class="Insecta" family="Carabidae" genus="Quasicalathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quasicalathus agonicollis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="agonicollis" status="sp. nov.">Quasicalathus agonicollis Schmidt & Will</taxonomicName>
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<taxonomicNameLabel id="F93A4D166A300B9BC52570ADE0AFA779" pageId="0" pageNumber="19">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="389B341D7B92C5EE94F766C5567B64C3" pageId="0" pageNumber="19" type="description">
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<paragraph id="0FEDAE4C2692A36CD271D7C144A361D5" pageId="0" pageNumber="19">
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<figureCitation id="BF5E40257D8EA989F6B0094796ED5067" captionStart="Figures 18–24" captionStartId="F4" captionText="Figures 18 - 24. Quasicalathus, light microscopic images of Q. elpis Ortuno & Arillo, 2009 (18 - 20.) and Q. agonicollis sp. nov. (21 - 24.). 18. General view of the amber piece " MAIG 76 " (only that part of the large amber piece bearing the Quasicalathus fossil is shown; the fossil is widely covered by milky coating); 19. General view of the two fragments of specimen " GZG 16185 "; the left one bears only the negative imprint of the left elytra on the inclusion wall; 20. Right anterior part of body of specimen " GZG 16185 " showing part of head, pronotum and humerus; 21, 22. General view of the amber piece " GZG 16188 " (21. With fossil in dorsal view; 22. In ventral view); 23. Anterior part of specimen " GZG 16188 "; 24. Head of specimen " GZG 16188 ". Abbreviations: a 1 - a 6 - antennomeres 1 - 6; as - anterior supraorbital seta; bs - pronotal laterobasal seta; ms - pronotal lateral seta; ps - posterior supraorbital seta." figureDoi="10.3897/dez.69.79931.figures18-24" httpUri="https://binary.pensoft.net/fig/643954" pageId="0" pageNumber="19">Figs 21-24</figureCitation>
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<figureCitation id="4CAB4E76C077A6007150A2C5660F05D6" captionStart="Figures 25–30" captionStartId="F5" captionText="Figures 25 - 30. Quasicalathus agonicollis sp. nov., light microscopic images of specimen " GZG 16188 " (25 - 26.) and the holotype (28 - 30.). 25. Posterior part of pronotum and anterior part of elytra, right side; 26. Anterior part of fifth interval of left elytra showing microsculpture; 27. Posterior part of elytra (the arrows point to the insertions of the discal setae); 28. General view of the amber piece; 29. Right dorsal view of beetle body (the arrow points to the insertion of the discal seta on left elytron); 30. Left ventral view. Abbreviations: bs - insertion of the pronotal laterobasal seta; hm - humerus; m - mite (syninclusion)." figureDoi="10.3897/dez.69.79931.figures25-30" httpUri="https://binary.pensoft.net/fig/643955" pageId="0" pageNumber="19">, 25-30</figureCitation>
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<figureCitation id="79D44D3B9242C05440E7F1288C846A87" captionStart="Figures 31–33" captionStartId="F6" captionText="Figures 31 - 33. Quasicalathus, light microscopic images of the holotypes of Q. agonicollis sp. nov. (31.) and Q. conservans sp. nov. (32, 33.). 31. Ventral side of head showing chaetotaxy of mentum; 32. General view of the amber piece with fossil in dorsal view; 32. Left lateral view. Abbreviations: ce - compound eye; el - elytron; ems - external seta of submentum; gu - gula; ims - internal seta of submentum; msf - mesofemur; mt - mentum; mtf - metafemur; prf - profemur; pt - pronotum." figureDoi="10.3897/dez.69.79931.figures31-33" httpUri="https://binary.pensoft.net/fig/643956" pageId="0" pageNumber="19">, 31</figureCitation>
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<figureCitation id="E8C4F032933EE630889E9F925F483914" captionStart="Figures 65–72" captionStartId="F13" captionText="Figures 65 - 72. Quasicalathus agonicollis sp. nov., volume rendering of the holotype using different grey scales of the Amira software. 65. Dorsal aspect (the negative imprint of the fossil on the inclusion wall is shown); 66. Basal portion of pronotum and anterior part of elytra (positive of the fossilized beetle is shown; the arrows point to the insertions of the pronotal basolateral setae and the parascutellar setae); 67. Metacoxa (the arrows point to the insertions of the three coxal setae each side); 68 - 71. Aedeagus in dorsal aspect (68.), Right lateral aspect (69.), Ventral aspect (70.), Left lateral aspect (71.); The remains of the parameres are coloured (red: left paramere; green: right paramere); 72. left lateral aspect of beetle body; the aedeagus (highlighted by red colour) was separated by the segmentation function of Amira software. Abbreviations: bb - basal bulb of aedeagal median lobe; hm - humerus; os - distal ostium of median lobe; sc - scutellum; tl - terminal lamella of median lobe." figureDoi="10.3897/dez.69.79931.figures65-72" httpUri="https://binary.pensoft.net/fig/643963" pageId="0" pageNumber="19">, 65-72</figureCitation>
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<figureCitation id="8B07A28280156F0521DCB51DC623F8C5" captionStart="Figures 73–77" captionStartId="F14" captionText="Figures 73 - 77. Quasicalathus agonicollis sp. nov., volume rendering of specimen " GZG 16188 ". 73. Dorsal aspect; 74. Basal portion of pronotum and anterior part of elytra (right side of body; the arrows point to the insertions of the pronotal basolateral seta and the parascutellar seta); 75. Prosternum with basal portion of prolegs; 76. Left apical gonocoxite, ventral aspect; 77. Gonocoxites, ventral aspect. Abbreviations: cx - procoxa; des - dorsal ensiform setae; fm - profemur; gx 1 - basal gonocoxite; gx 2 - apical gonocoxite; hm - humerus; pst - prosternum; sc - scutellum; sps - setae of sensory pit; tr - protrochanter; ves - ventral ensiform setae." figureDoi="10.3897/dez.69.79931.figures73-77" httpUri="https://binary.pensoft.net/fig/643964" pageId="0" pageNumber="19">, 73-77</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="227DC402027F0B59CAD66173A87E8D72" pageId="0" pageNumber="19" type="holotype">
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<paragraph id="514AFF82088ED3F0AF1C03503A7450FE" pageId="0" pageNumber="19">Holotype.</paragraph>
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<paragraph id="0A9DEED7986D039244FCE07D3E41D4E0" pageId="0" pageNumber="19">
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Male in Baltic amber, with specimen label data "
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<emphasis id="FA4C50B614F7A7A942C7C9C8F075FF76" bold="true" pageId="0" pageNumber="19">SDEI-Amb-002528</emphasis>
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", deposited in Senckenberg Deutsches Entomologisches Institut,
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<normalizedToken id="B7690D029516AC042F98BE52F41020F6" originalValue="Müncheberg">Muencheberg</normalizedToken>
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, Germany. The original size of the amber piece was 60
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<normalizedToken id="DB872470C2473FA58C0891C4E7D8DD95" originalValue="×">x</normalizedToken>
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23
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<normalizedToken id="DAFE34F3F773C3F08B4C582DD76F3BAE" originalValue="×">x</normalizedToken>
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7 mm and was separated into three pieces (SDEI Amb-002528 a, b, c) in order to get better micro-CT scanning results. The size of the amber piece bearing the calathine fossil measures approx. 15
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<normalizedToken id="273F6BBC8B804C39D464B9EA1E071C83" originalValue="×">x</normalizedToken>
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9
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<normalizedToken id="07F1817BEF2EF6111AA4965B8C4D516C" originalValue="×">x</normalizedToken>
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4 mm (Fig.
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<figureCitation id="8572CBB16D8E3CECFF44D0F9FF883620" captionStart="Figures 25–30" captionStartId="F5" captionText="Figures 25 - 30. Quasicalathus agonicollis sp. nov., light microscopic images of specimen " GZG 16188 " (25 - 26.) and the holotype (28 - 30.). 25. Posterior part of pronotum and anterior part of elytra, right side; 26. Anterior part of fifth interval of left elytra showing microsculpture; 27. Posterior part of elytra (the arrows point to the insertions of the discal setae); 28. General view of the amber piece; 29. Right dorsal view of beetle body (the arrow points to the insertion of the discal seta on left elytron); 30. Left ventral view. Abbreviations: bs - insertion of the pronotal laterobasal seta; hm - humerus; m - mite (syninclusion)." figureDoi="10.3897/dez.69.79931.figures25-30" httpUri="https://binary.pensoft.net/fig/643955" pageId="0" pageNumber="19">28</figureCitation>
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).
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</paragraph>
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<caption id="23A677A399992984DFEE8B34963DAD3E" doi="10.3897/dez.69.79931.figures61-64" httpUri="https://binary.pensoft.net/fig/643962" pageId="0" pageNumber="19" start="Figures 61–64" startId="F12">
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<paragraph id="5A8998D1C206BECFE8F549ECA1B6AA58" pageId="0" pageNumber="19">
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<emphasis id="D1CFFCBEEA2ECFA5265845F3A1722258" bold="true" pageId="0" pageNumber="19">Figures 61-64.</emphasis>
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<taxonomicName id="85B4074E1F8C1885F127AD4909960D58" authorityName="Schmidt & Scholz & Will" authorityYear="2022" baseAuthorityName="Ortuno & Arillo" baseAuthorityYear="2009" class="Insecta" family="Carabidae" genus="Quasicalathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quasicalathus elpis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="elpis">
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<emphasis id="24A1A308D3732DEC4BF4D7D3DD2CC788" italics="true" pageId="0" pageNumber="19">Quasicalathus elpis</emphasis>
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</taxonomicName>
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(
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<bibRefCitation id="6F08CDB03A18736CCFC6ED5AF4BCA27C" DOI="https://doi.org/10.11646/zootaxa.2239.1.5" author="Ortuno, VM" journalOrPublisher="Zootaxa" pageId="0" pageNumber="19" pagination="55 - 61" refId="B25" refString="Ortuno, VM, Arillo, A, 2009. Fossil carabids from Baltic amber - A new species of the genus Calathus Bonelli, 1810 (Coleoptera: Carabidae: Pterostichinae). Zootaxa 2239: 55 - 61, DOI: https://doi.org/10.11646/zootaxa.2239.1.5" title="Fossil carabids from Baltic amber - A new species of the genus Calathus Bonelli, 1810 (Coleoptera: Carabidae: Pterostichinae)." url="https://doi.org/10.11646/zootaxa.2239.1.5" volume="2239" year="2009">
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<normalizedToken id="A28D0CDA51E5C501ECB00E6CCFCC2FA4" originalValue="Ortuño">Ortuno</normalizedToken>
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and Arillo 2009
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</bibRefCitation>
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), volume rendering of specimens "MAIG 76" (
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<emphasis id="57D93D2E41F528B47CB748A49B96DC82" bold="true" pageId="0" pageNumber="19">61-63.</emphasis>
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) and "GZG 16185" (
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<emphasis id="EF7315D23894F02165A64A2B3458F66D" bold="true" pageId="0" pageNumber="19">64.</emphasis>
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);
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<emphasis id="D4FD4CAF62193EB89558E2711657E7A1" bold="true" pageId="0" pageNumber="19">61.</emphasis>
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Dorsal aspect;
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<emphasis id="77FCC5FE024110F804B208F1CBC07D32" bold="true" pageId="0" pageNumber="19">62.</emphasis>
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Right lateral aspect;
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<emphasis id="2BA5A87F8708ECE14CD054956E8D8F94" bold="true" pageId="0" pageNumber="19">63, 64.</emphasis>
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Pronotum (the pronotal outline on left side is highlighted by dotted line in Fig.
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<figureCitation id="383FA542E1DF2649FD01B638ADFE51FC" captionStart="Figures 61–64" captionStartId="F12" captionText="Figures 61 - 64. Quasicalathus elpis (Ortuno and Arillo 2009), volume rendering of specimens " MAIG 76 " (61 - 63.) and " GZG 16185 " (64.); 61. Dorsal aspect; 62. Right lateral aspect; 63, 64. Pronotum (the pronotal outline on left side is highlighted by dotted line in Fig. 64)." figureDoi="10.3897/dez.69.79931.figures61-64" httpUri="https://binary.pensoft.net/fig/643962" pageId="0" pageNumber="19">64</figureCitation>
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).
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</paragraph>
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</caption>
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<paragraph id="046DABE327DE05941BD103CF96F60325" pageId="0" pageNumber="19">
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Preservation status: A clear piece of amber with the embedded carabid fossil well visible, however, extensive flowlines are present in front of the head and on the right side of the beetle body. Several parts of the beetle body are additionally covered by a white coating so that details of the exoskeleton are not visible using light microscopy; this includes the dorsal surface of head and pronotum, lateral parts of the elytra, right side of the ventral surface (Figs
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<figureCitation id="FC9DBA7F97539A2D6A5C6E2A63D8C482" captionStart="Figures 25–30" captionStartId="F5" captionText="Figures 25 - 30. Quasicalathus agonicollis sp. nov., light microscopic images of specimen " GZG 16188 " (25 - 26.) and the holotype (28 - 30.). 25. Posterior part of pronotum and anterior part of elytra, right side; 26. Anterior part of fifth interval of left elytra showing microsculpture; 27. Posterior part of elytra (the arrows point to the insertions of the discal setae); 28. General view of the amber piece; 29. Right dorsal view of beetle body (the arrow points to the insertion of the discal seta on left elytron); 30. Left ventral view. Abbreviations: bs - insertion of the pronotal laterobasal seta; hm - humerus; m - mite (syninclusion)." figureDoi="10.3897/dez.69.79931.figures25-30" httpUri="https://binary.pensoft.net/fig/643955" pageId="0" pageNumber="19">29</figureCitation>
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,
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<figureCitation id="B31157EDAF851355D9968E7ACAC94206" captionStart="Figures 25–30" captionStartId="F5" captionText="Figures 25 - 30. Quasicalathus agonicollis sp. nov., light microscopic images of specimen " GZG 16188 " (25 - 26.) and the holotype (28 - 30.). 25. Posterior part of pronotum and anterior part of elytra, right side; 26. Anterior part of fifth interval of left elytra showing microsculpture; 27. Posterior part of elytra (the arrows point to the insertions of the discal setae); 28. General view of the amber piece; 29. Right dorsal view of beetle body (the arrow points to the insertion of the discal seta on left elytron); 30. Left ventral view. Abbreviations: bs - insertion of the pronotal laterobasal seta; hm - humerus; m - mite (syninclusion)." figureDoi="10.3897/dez.69.79931.figures25-30" httpUri="https://binary.pensoft.net/fig/643955" pageId="0" pageNumber="19">30</figureCitation>
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). The external surface of the fossilized beetle body yields low (head, thorax) or relatively moderate contrast (elytra, abdomen) during micro-CT scan and therefore, the anterior part of the body could only be coarsely imaged (Fig.
|
||
<figureCitation id="CF2ECE091DA2732D413F74007B7F220C" captionStart="Figures 65–72" captionStartId="F13" captionText="Figures 65 - 72. Quasicalathus agonicollis sp. nov., volume rendering of the holotype using different grey scales of the Amira software. 65. Dorsal aspect (the negative imprint of the fossil on the inclusion wall is shown); 66. Basal portion of pronotum and anterior part of elytra (positive of the fossilized beetle is shown; the arrows point to the insertions of the pronotal basolateral setae and the parascutellar setae); 67. Metacoxa (the arrows point to the insertions of the three coxal setae each side); 68 - 71. Aedeagus in dorsal aspect (68.), Right lateral aspect (69.), Ventral aspect (70.), Left lateral aspect (71.); The remains of the parameres are coloured (red: left paramere; green: right paramere); 72. left lateral aspect of beetle body; the aedeagus (highlighted by red colour) was separated by the segmentation function of Amira software. Abbreviations: bb - basal bulb of aedeagal median lobe; hm - humerus; os - distal ostium of median lobe; sc - scutellum; tl - terminal lamella of median lobe." figureDoi="10.3897/dez.69.79931.figures65-72" httpUri="https://binary.pensoft.net/fig/643963" pageId="0" pageNumber="19">65</figureCitation>
|
||
). The aedeagus with median lobe and right paramere is moderately well preserved and could partly be imaged using micro-CT data (Figs
|
||
<figureCitation id="C7056916E0215105D0ABFD98BFF1EA0C" captionStart="Figures 65–72" captionStartId="F13" captionText="Figures 65 - 72. Quasicalathus agonicollis sp. nov., volume rendering of the holotype using different grey scales of the Amira software. 65. Dorsal aspect (the negative imprint of the fossil on the inclusion wall is shown); 66. Basal portion of pronotum and anterior part of elytra (positive of the fossilized beetle is shown; the arrows point to the insertions of the pronotal basolateral setae and the parascutellar setae); 67. Metacoxa (the arrows point to the insertions of the three coxal setae each side); 68 - 71. Aedeagus in dorsal aspect (68.), Right lateral aspect (69.), Ventral aspect (70.), Left lateral aspect (71.); The remains of the parameres are coloured (red: left paramere; green: right paramere); 72. left lateral aspect of beetle body; the aedeagus (highlighted by red colour) was separated by the segmentation function of Amira software. Abbreviations: bb - basal bulb of aedeagal median lobe; hm - humerus; os - distal ostium of median lobe; sc - scutellum; tl - terminal lamella of median lobe." figureDoi="10.3897/dez.69.79931.figures65-72" httpUri="https://binary.pensoft.net/fig/643963" pageId="0" pageNumber="19">68-72</figureCitation>
|
||
); the left paramere provided contrast during CT scan that was too low for reconstruction.
|
||
</paragraph>
|
||
<paragraph id="0A46D88028112952E1A2D5F67D9F221C" pageId="0" pageNumber="19">
|
||
Syninclusions: SDEI-Amb-002528a: seven very tiny insect larva, dust particles; one mite on the carabid fossil near the
|
||
<normalizedToken id="90F77981BCF686A239C46D1C4159964A" originalValue="beetle’s">beetle's</normalizedToken>
|
||
scutellum. SDEI-Amb-002528b: one
|
||
<taxonomicName id="3A184C912E246AF1CDF39CDD2736067C" authorityName="Lepeletier de Saint-Fargeau" authorityYear="1835" lsidName="" pageId="0" pageNumber="19" rank="subFamily" subFamily="Myrmicinae">Myrmicinae</taxonomicName>
|
||
ant, three
|
||
<taxonomicName id="C6134A514404E5BB42F51EA6BB3D8E6F" authorityName="Schmidt & Will" authorityYear="2022" class="Insecta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="subOrder" subOrder="Nematocera">Nematocera</taxonomicName>
|
||
flies, stellate hairs, dust particles. SDEI-Amb-002528c: dust particles.
|
||
</paragraph>
|
||
<caption id="5C0D09521C221FBF776A5AB39C2D3749" doi="10.3897/dez.69.79931.figures65-72" httpUri="https://binary.pensoft.net/fig/643963" pageId="0" pageNumber="19" start="Figures 65–72" startId="F13">
|
||
<paragraph id="2A6778BC1F96480CD46EF0E45F7A7F46" pageId="0" pageNumber="19">
|
||
<emphasis id="6AD34F1A7A086429EE22E8020FFBC032" bold="true" pageId="0" pageNumber="19">Figures 65-72.</emphasis>
|
||
<taxonomicName id="5013EBB96B895E4B34FAA4B5A31C3BD6" authorityName="Schmidt & Will" authorityYear="2022" class="Insecta" family="Carabidae" genus="Quasicalathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quasicalathus agonicollis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="agonicollis">
|
||
<emphasis id="D4FFDA3A5C07FE8E6B1D5DCFB3D21F87" italics="true" pageId="0" pageNumber="19">Quasicalathus agonicollis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., volume rendering of the holotype using different grey scales of the Amira software.
|
||
<emphasis id="E5F6672C2A776BFC0B60C303C4480102" bold="true" pageId="0" pageNumber="19">65.</emphasis>
|
||
Dorsal aspect (the negative imprint of the fossil on the inclusion wall is shown);
|
||
<emphasis id="5F43AAA38EAB8F7DC3C9D6E49DA15B3F" bold="true" pageId="0" pageNumber="19">66.</emphasis>
|
||
Basal portion of pronotum and anterior part of elytra (positive of the fossilized beetle is shown; the arrows point to the insertions of the pronotal basolateral setae and the parascutellar setae);
|
||
<emphasis id="33BD87B565A10A1C7C50CFF431B09E58" bold="true" pageId="0" pageNumber="19">67.</emphasis>
|
||
Metacoxa (the arrows point to the insertions of the three coxal setae each side);
|
||
<emphasis id="8A11FB8C634739DC51CEFD251307051B" bold="true" pageId="0" pageNumber="19">68-71.</emphasis>
|
||
Aedeagus in dorsal aspect (
|
||
<emphasis id="DBA8E025FB87EE406783926CF3458B8C" bold="true" pageId="0" pageNumber="19">68.</emphasis>
|
||
), Right lateral aspect (
|
||
<emphasis id="3AEAB6228DA82F71A11F02DC57B2F3A8" bold="true" pageId="0" pageNumber="19">69.</emphasis>
|
||
), Ventral aspect (
|
||
<emphasis id="7B3174EC018503126F7FBDB73C82E63F" bold="true" pageId="0" pageNumber="19">70.</emphasis>
|
||
), Left lateral aspect (
|
||
<emphasis id="1A60273D24DCCCF95D092B1C572F55B1" bold="true" pageId="0" pageNumber="19">71.</emphasis>
|
||
); The remains of the parameres are coloured (red: left paramere; green: right paramere);
|
||
<emphasis id="184E114FDD36D306D721F03D9284857A" bold="true" pageId="0" pageNumber="19">72.</emphasis>
|
||
left lateral aspect of beetle body; the aedeagus (highlighted by red colour) was separated by the segmentation function of Amira software. Abbreviations:
|
||
<emphasis id="7C75BD4091303CD44A23D648513EBECC" bold="true" pageId="0" pageNumber="19">bb</emphasis>
|
||
- basal bulb of aedeagal median lobe;
|
||
<emphasis id="269B917B021CA0FF05D32A68FE66C7B2" bold="true" pageId="0" pageNumber="19">hm</emphasis>
|
||
- humerus;
|
||
<emphasis id="AB6684222D4BD3308841109474C3A77B" bold="true" pageId="0" pageNumber="19">os</emphasis>
|
||
- distal ostium of median lobe;
|
||
<emphasis id="6A0715CEB2722F41BD7F0E01D1052057" bold="true" pageId="0" pageNumber="19">sc</emphasis>
|
||
- scutellum;
|
||
<emphasis id="BC7DFD2EAAF60F0A7AAF037D54988B72" bold="true" pageId="0" pageNumber="19">tl</emphasis>
|
||
- terminal lamella of median lobe.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="2008D44AD42071C3FB4CEAFE192571FD" pageId="0" pageNumber="19" type="materials_examined">
|
||
<paragraph id="77E3A7796C7B1D036FABEF2F774652E2" pageId="0" pageNumber="19">Additional material.</paragraph>
|
||
<paragraph id="184E7D4D2AD4A8C6BC1CDEC62CF4821B" pageId="0" pageNumber="19">
|
||
Female in Baltic amber, with specimen label data "
|
||
<emphasis id="1C7ACE0B43B4F908602790E6829DF3E6" bold="true" pageId="0" pageNumber="19">
|
||
GZG
|
||
<accessionNumber id="569F2D1A11D744757B1ADDEAD91A5800" httpUri="http://www.ncbi.nlm.nih.gov/protein/BST16188">BST 16188</accessionNumber>
|
||
</emphasis>
|
||
" and
|
||
<normalizedToken id="2237B8A9007C2AAD45F41B878FBA5E9D" originalValue="“K2192”">"K2192"</normalizedToken>
|
||
(ex coll. Klebs), deposited in Geoscience Museum, University of
|
||
<normalizedToken id="741D56EDF1FF870A723ED35339276585" originalValue="Göttingen">Goettingen</normalizedToken>
|
||
,
|
||
<collectingCountry id="FB14297A9104D38D244D21B52825326E" name="Germany">Germany</collectingCountry>
|
||
. Size of the amber piece approx. 12
|
||
<normalizedToken id="DD56968D287B23C17BAF4FC5DCC4E9CB" originalValue="×">x</normalizedToken>
|
||
7
|
||
<normalizedToken id="E7BDD56154F617309DB947EE7E967134" originalValue="×">x</normalizedToken>
|
||
<quantity id="4DBC07F84D0FACCD123297E81646813D" metricMagnitude="-3" metricUnit="m" metricValue="4.0" unit="mm" value="4.0">4 mm</quantity>
|
||
, with seven polished edges (Figs
|
||
<figureCitation id="A5FC686C883CA0CF1DCD8AE7E8001356" captionStart="Figures 18–24" captionStartId="F4" captionText="Figures 18 - 24. Quasicalathus, light microscopic images of Q. elpis Ortuno & Arillo, 2009 (18 - 20.) and Q. agonicollis sp. nov. (21 - 24.). 18. General view of the amber piece " MAIG 76 " (only that part of the large amber piece bearing the Quasicalathus fossil is shown; the fossil is widely covered by milky coating); 19. General view of the two fragments of specimen " GZG 16185 "; the left one bears only the negative imprint of the left elytra on the inclusion wall; 20. Right anterior part of body of specimen " GZG 16185 " showing part of head, pronotum and humerus; 21, 22. General view of the amber piece " GZG 16188 " (21. With fossil in dorsal view; 22. In ventral view); 23. Anterior part of specimen " GZG 16188 "; 24. Head of specimen " GZG 16188 ". Abbreviations: a 1 - a 6 - antennomeres 1 - 6; as - anterior supraorbital seta; bs - pronotal laterobasal seta; ms - pronotal lateral seta; ps - posterior supraorbital seta." figureDoi="10.3897/dez.69.79931.figures18-24" httpUri="https://binary.pensoft.net/fig/643954" pageId="0" pageNumber="19">21</figureCitation>
|
||
,
|
||
<figureCitation id="08B470B947B98BCF60CC1B3BC43678F4" captionStart="Figures 18–24" captionStartId="F4" captionText="Figures 18 - 24. Quasicalathus, light microscopic images of Q. elpis Ortuno & Arillo, 2009 (18 - 20.) and Q. agonicollis sp. nov. (21 - 24.). 18. General view of the amber piece " MAIG 76 " (only that part of the large amber piece bearing the Quasicalathus fossil is shown; the fossil is widely covered by milky coating); 19. General view of the two fragments of specimen " GZG 16185 "; the left one bears only the negative imprint of the left elytra on the inclusion wall; 20. Right anterior part of body of specimen " GZG 16185 " showing part of head, pronotum and humerus; 21, 22. General view of the amber piece " GZG 16188 " (21. With fossil in dorsal view; 22. In ventral view); 23. Anterior part of specimen " GZG 16188 "; 24. Head of specimen " GZG 16188 ". Abbreviations: a 1 - a 6 - antennomeres 1 - 6; as - anterior supraorbital seta; bs - pronotal laterobasal seta; ms - pronotal lateral seta; ps - posterior supraorbital seta." figureDoi="10.3897/dez.69.79931.figures18-24" httpUri="https://binary.pensoft.net/fig/643954" pageId="0" pageNumber="19">22</figureCitation>
|
||
); one edge bears the inscription
|
||
<normalizedToken id="92646BD12D3E46A618761DE4A42A34F9" originalValue="“G644”">"G644"</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="9268FE8BC36814E8964328457E837369" pageId="0" pageNumber="19">
|
||
Preservation status: The amber stone is clear in most parts but its surface shows several corrosion cracks (Figs
|
||
<figureCitation id="91765CB6B22E612B9D45256C17D5D974" captionStart="Figures 18–24" captionStartId="F4" captionText="Figures 18 - 24. Quasicalathus, light microscopic images of Q. elpis Ortuno & Arillo, 2009 (18 - 20.) and Q. agonicollis sp. nov. (21 - 24.). 18. General view of the amber piece " MAIG 76 " (only that part of the large amber piece bearing the Quasicalathus fossil is shown; the fossil is widely covered by milky coating); 19. General view of the two fragments of specimen " GZG 16185 "; the left one bears only the negative imprint of the left elytra on the inclusion wall; 20. Right anterior part of body of specimen " GZG 16185 " showing part of head, pronotum and humerus; 21, 22. General view of the amber piece " GZG 16188 " (21. With fossil in dorsal view; 22. In ventral view); 23. Anterior part of specimen " GZG 16188 "; 24. Head of specimen " GZG 16188 ". Abbreviations: a 1 - a 6 - antennomeres 1 - 6; as - anterior supraorbital seta; bs - pronotal laterobasal seta; ms - pronotal lateral seta; ps - posterior supraorbital seta." figureDoi="10.3897/dez.69.79931.figures18-24" httpUri="https://binary.pensoft.net/fig/643954" pageId="0" pageNumber="19">23</figureCitation>
|
||
,
|
||
<figureCitation id="7A5B162A884B703D392EFBB30717899A" captionStart="Figures 18–24" captionStartId="F4" captionText="Figures 18 - 24. Quasicalathus, light microscopic images of Q. elpis Ortuno & Arillo, 2009 (18 - 20.) and Q. agonicollis sp. nov. (21 - 24.). 18. General view of the amber piece " MAIG 76 " (only that part of the large amber piece bearing the Quasicalathus fossil is shown; the fossil is widely covered by milky coating); 19. General view of the two fragments of specimen " GZG 16185 "; the left one bears only the negative imprint of the left elytra on the inclusion wall; 20. Right anterior part of body of specimen " GZG 16185 " showing part of head, pronotum and humerus; 21, 22. General view of the amber piece " GZG 16188 " (21. With fossil in dorsal view; 22. In ventral view); 23. Anterior part of specimen " GZG 16188 "; 24. Head of specimen " GZG 16188 ". Abbreviations: a 1 - a 6 - antennomeres 1 - 6; as - anterior supraorbital seta; bs - pronotal laterobasal seta; ms - pronotal lateral seta; ps - posterior supraorbital seta." figureDoi="10.3897/dez.69.79931.figures18-24" httpUri="https://binary.pensoft.net/fig/643954" pageId="0" pageNumber="19">24</figureCitation>
|
||
); the ventral surface of the fossil is completely covered by a milky coating (Fig.
|
||
<figureCitation id="8FFBC9D8F53ED42BF6CA36976CAE515A" captionStart="Figures 18–24" captionStartId="F4" captionText="Figures 18 - 24. Quasicalathus, light microscopic images of Q. elpis Ortuno & Arillo, 2009 (18 - 20.) and Q. agonicollis sp. nov. (21 - 24.). 18. General view of the amber piece " MAIG 76 " (only that part of the large amber piece bearing the Quasicalathus fossil is shown; the fossil is widely covered by milky coating); 19. General view of the two fragments of specimen " GZG 16185 "; the left one bears only the negative imprint of the left elytra on the inclusion wall; 20. Right anterior part of body of specimen " GZG 16185 " showing part of head, pronotum and humerus; 21, 22. General view of the amber piece " GZG 16188 " (21. With fossil in dorsal view; 22. In ventral view); 23. Anterior part of specimen " GZG 16188 "; 24. Head of specimen " GZG 16188 ". Abbreviations: a 1 - a 6 - antennomeres 1 - 6; as - anterior supraorbital seta; bs - pronotal laterobasal seta; ms - pronotal lateral seta; ps - posterior supraorbital seta." figureDoi="10.3897/dez.69.79931.figures18-24" httpUri="https://binary.pensoft.net/fig/643954" pageId="0" pageNumber="19">22</figureCitation>
|
||
). The exoskeleton of the fossil is moderately well preserved and could therefore be imaged in most details using micro-CT (Figs
|
||
<figureCitation id="3ED936DF50686C0D459E710E1BD2B2C8" captionStart="Figures 73–77" captionStartId="F14" captionText="Figures 73 - 77. Quasicalathus agonicollis sp. nov., volume rendering of specimen " GZG 16188 ". 73. Dorsal aspect; 74. Basal portion of pronotum and anterior part of elytra (right side of body; the arrows point to the insertions of the pronotal basolateral seta and the parascutellar seta); 75. Prosternum with basal portion of prolegs; 76. Left apical gonocoxite, ventral aspect; 77. Gonocoxites, ventral aspect. Abbreviations: cx - procoxa; des - dorsal ensiform setae; fm - profemur; gx 1 - basal gonocoxite; gx 2 - apical gonocoxite; hm - humerus; pst - prosternum; sc - scutellum; sps - setae of sensory pit; tr - protrochanter; ves - ventral ensiform setae." figureDoi="10.3897/dez.69.79931.figures73-77" httpUri="https://binary.pensoft.net/fig/643964" pageId="0" pageNumber="19">73-75</figureCitation>
|
||
), including the gonocoxites (Figs
|
||
<figureCitation id="6FBC11907B30CE179EF0D1E26A9424E2" captionStart="Figures 73–77" captionStartId="F14" captionText="Figures 73 - 77. Quasicalathus agonicollis sp. nov., volume rendering of specimen " GZG 16188 ". 73. Dorsal aspect; 74. Basal portion of pronotum and anterior part of elytra (right side of body; the arrows point to the insertions of the pronotal basolateral seta and the parascutellar seta); 75. Prosternum with basal portion of prolegs; 76. Left apical gonocoxite, ventral aspect; 77. Gonocoxites, ventral aspect. Abbreviations: cx - procoxa; des - dorsal ensiform setae; fm - profemur; gx 1 - basal gonocoxite; gx 2 - apical gonocoxite; hm - humerus; pst - prosternum; sc - scutellum; sps - setae of sensory pit; tr - protrochanter; ves - ventral ensiform setae." figureDoi="10.3897/dez.69.79931.figures73-77" httpUri="https://binary.pensoft.net/fig/643964" pageId="0" pageNumber="19">76</figureCitation>
|
||
,
|
||
<figureCitation id="7266F90D159B46E9745061BD33C2F807" captionStart="Figures 73–77" captionStartId="F14" captionText="Figures 73 - 77. Quasicalathus agonicollis sp. nov., volume rendering of specimen " GZG 16188 ". 73. Dorsal aspect; 74. Basal portion of pronotum and anterior part of elytra (right side of body; the arrows point to the insertions of the pronotal basolateral seta and the parascutellar seta); 75. Prosternum with basal portion of prolegs; 76. Left apical gonocoxite, ventral aspect; 77. Gonocoxites, ventral aspect. Abbreviations: cx - procoxa; des - dorsal ensiform setae; fm - profemur; gx 1 - basal gonocoxite; gx 2 - apical gonocoxite; hm - humerus; pst - prosternum; sc - scutellum; sps - setae of sensory pit; tr - protrochanter; ves - ventral ensiform setae." figureDoi="10.3897/dez.69.79931.figures73-77" httpUri="https://binary.pensoft.net/fig/643964" pageId="0" pageNumber="19">77</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="C55A5496C0AC753D47C028A2F804FD55" pageId="0" pageNumber="19">Syninclusions: Stellate hairs, dust particles.</paragraph>
|
||
<caption id="B78E244BBA94688FC20CD76CA21D4BE9" doi="10.3897/dez.69.79931.figures73-77" httpUri="https://binary.pensoft.net/fig/643964" pageId="0" pageNumber="19" start="Figures 73–77" startId="F14">
|
||
<paragraph id="EBD8F6C69007924EBFC84BB195CF6CDA" pageId="0" pageNumber="19">
|
||
<emphasis id="E7A402E9102ED38D9F2C7DD405FDC1F5" bold="true" pageId="0" pageNumber="19">Figures 73-77.</emphasis>
|
||
<taxonomicName id="60BA9B6D4628750CECDF7EE98E19599B" authorityName="Schmidt & Will" authorityYear="2022" class="Insecta" family="Carabidae" genus="Quasicalathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quasicalathus agonicollis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="agonicollis">
|
||
<emphasis id="C31E84CAB8936519230F532127705FB6" italics="true" pageId="0" pageNumber="19">Quasicalathus agonicollis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., volume rendering of specimen "
|
||
<accessionNumber id="B3BF6227710E34151A0C563C94277659" httpUri="http://www.ncbi.nlm.nih.gov/protein/GZG16188">GZG 16188</accessionNumber>
|
||
".
|
||
<emphasis id="6473A7A744F4F217E0AF842C32A979D9" bold="true" pageId="0" pageNumber="19">73.</emphasis>
|
||
Dorsal aspect;
|
||
<emphasis id="906BF537612BFD8E4F128AC325DFFE96" bold="true" pageId="0" pageNumber="19">74.</emphasis>
|
||
Basal portion of pronotum and anterior part of elytra (right side of body; the arrows point to the insertions of the pronotal basolateral seta and the parascutellar seta);
|
||
<emphasis id="F3C90076943031DB2A017567CB0C484D" bold="true" pageId="0" pageNumber="19">75.</emphasis>
|
||
Prosternum with basal portion of prolegs;
|
||
<emphasis id="11F12F86913E968FBCE3AEAF876FD990" bold="true" pageId="0" pageNumber="19">76.</emphasis>
|
||
Left apical gonocoxite, ventral aspect;
|
||
<emphasis id="A49B580826BF592F41842517FD8CCDA0" bold="true" pageId="0" pageNumber="19">77.</emphasis>
|
||
Gonocoxites, ventral aspect. Abbreviations:
|
||
<emphasis id="ECDD3B4DA7A65897DB3C0ACD85C69623" bold="true" pageId="0" pageNumber="19">cx</emphasis>
|
||
- procoxa;
|
||
<emphasis id="84BDEDF1486AD3769AF3517AADD34129" bold="true" pageId="0" pageNumber="19">des</emphasis>
|
||
- dorsal ensiform setae;
|
||
<emphasis id="560036B8F09BF39C89D4A0580E2884FB" bold="true" pageId="0" pageNumber="19">fm</emphasis>
|
||
- profemur;
|
||
<emphasis id="7F17DFBD60599152E0AD2659C6BC446A" bold="true" pageId="0" pageNumber="19">gx1</emphasis>
|
||
- basal gonocoxite;
|
||
<emphasis id="5B798C246F2E1304826C753CFEC91B9E" bold="true" pageId="0" pageNumber="19">gx2</emphasis>
|
||
- apical gonocoxite;
|
||
<emphasis id="98B7E3F69810BEDE882F85B7D56ACA94" bold="true" pageId="0" pageNumber="19">hm</emphasis>
|
||
- humerus;
|
||
<emphasis id="3E24CEDCFC802D322DCB20294B36FFD4" bold="true" pageId="0" pageNumber="19">pst</emphasis>
|
||
- prosternum;
|
||
<emphasis id="D0E105D1871BCBC6B5D5287CCF401B11" bold="true" pageId="0" pageNumber="19">sc</emphasis>
|
||
- scutellum;
|
||
<emphasis id="4A937B07DC158B62EC7DB716441AFCA4" bold="true" pageId="0" pageNumber="19">sps</emphasis>
|
||
- setae of sensory pit;
|
||
<emphasis id="6D2DF69B5456E232C8D67C1583BADC31" bold="true" pageId="0" pageNumber="19">tr</emphasis>
|
||
- protrochanter;
|
||
<emphasis id="1A83CDCC565546E14C52A534A36D6DDE" bold="true" pageId="0" pageNumber="19">ves</emphasis>
|
||
- ventral ensiform setae.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="BA9AC9D5FDD8297E288A62027B2C52DF" pageId="0" pageNumber="19" type="remarks">
|
||
<paragraph id="67DC2AA42A2B388DE3A72D5F3E05EF49" pageId="0" pageNumber="19">Remarks.</paragraph>
|
||
<paragraph id="8CB53101B88A71CF2BF4365C43684473" pageId="0" pageNumber="19">
|
||
The specific identity of this second fossil specimen with the holotype of
|
||
<taxonomicName id="03BA8C2B98592574348BC2516B40278E" class="Insecta" kingdom="Animalia" lsidName="Q. agonicollis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="agonicollis">
|
||
<emphasis id="DA35C2E9E6C517B4CB739DF97B8C5C51" italics="true" pageId="0" pageNumber="19">Q. agonicollis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., given the current state is difficult to substantiate. This is due to the poor preservation state of the holotype specimen. In the specimen GZG 16188, the pattern of head microsculpture is quite differently developed from that what we found in
|
||
<taxonomicName id="FECA234F27E634051F1BD19C0F870A3C" class="Insecta" kingdom="Animalia" lsidName="Q. elpis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="elpis">
|
||
<emphasis id="A33A1523A3FE0048B8D42514E09E7F93" italics="true" pageId="0" pageNumber="19">Q. elpis</emphasis>
|
||
</taxonomicName>
|
||
and the below described
|
||
<taxonomicName id="6E8C5E2370C017463D52CDCF32FDA169" class="Insecta" kingdom="Animalia" lsidName="Q. conservans" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="conservans">
|
||
<emphasis id="4B6DEE06A0F0B5E71FC252E7095FD5F8" italics="true" pageId="0" pageNumber="19">Q. conservans</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., however, this character state is unknown for the
|
||
<taxonomicName id="EC34733608AB68674B71847700314DF1" class="Insecta" kingdom="Animalia" lsidName="Q. agonicollis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="agonicollis">
|
||
<emphasis id="327271C1E594A2A25046AC42309AAF76" italics="true" pageId="0" pageNumber="19">Q. agonicollis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. holotype. Therefore, our decision to identify GZG 16188 as
|
||
<taxonomicName id="4E97DD109E6BD602D53369AEFEE3B503" class="Insecta" kingdom="Animalia" lsidName="Q. agonicollis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="agonicollis">
|
||
<emphasis id="FCACC430902D5A6B17382E1148901A63" italics="true" pageId="0" pageNumber="19">Q. agonicollis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. must be considered provisional. It is based on the following four interspecific diagnostic character states that
|
||
<taxonomicName id="9260237A2A49977F2F206C008AAACF1D" class="Insecta" kingdom="Animalia" lsidName="Q. agonicollis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="agonicollis">
|
||
<emphasis id="AF2C5AE8BF87A531674C75FF0753352B" italics="true" pageId="0" pageNumber="19">Q. agonicollis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. holotype and specimen GZG 16188 share: i) body size small, SBL below 7 mm. ii) pronotal laterobasal angles more markedly obtuse; iii) elytral basal margin moderately concave; iv) humerus slightly protruded with humeral angle more markedly obtuse. Therefore, in the description of
|
||
<taxonomicName id="C33C11CC8790956880DC055D4295FC07" class="Insecta" kingdom="Animalia" lsidName="Q. agonicollis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="agonicollis">
|
||
<emphasis id="ACA61C3FC63B9DF6F32C8827CC2FE50F" italics="true" pageId="0" pageNumber="19">Q. agonicollis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. we separate the descriptions of the
|
||
<taxonomicName id="6613DA32F715C6C3607C08FDF1CC9BCC" class="Insecta" kingdom="Animalia" lsidName="Q. agonicollis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="agonicollis">
|
||
<emphasis id="135ECB5826F266EA3881B1BBDE0A71DA" italics="true" pageId="0" pageNumber="19">Q. agonicollis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. holotype and the specimen GZG 16188.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FC22412CD9A5F28A7C0E08AD82C3869F" pageId="0" pageNumber="19" type="description">
|
||
<paragraph id="E435A5FB0AB743717ED7AE4A7133ACB1" pageId="0" pageNumber="19">Description of the holotype.</paragraph>
|
||
<paragraph id="55D7B83B56E233DEBE4B1A509EC69321" pageId="0" pageNumber="19">
|
||
Measurements see Table
|
||
<tableCitation id="888B5408B78489209CADB08C3403D6DD" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Measurements [µm]." httpUri="http://table.plazi.org/id/C19891A363A03B4DD93DC20ECFC36EE4" pageId="0" pageNumber="19" tableUuid="C19891A363A03B4DD93DC20ECFC36EE4">2</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="1A32F29459E061B236BA085B6F961771" pageId="0" pageNumber="19">Standardized body length: 6.7 mm.</paragraph>
|
||
<paragraph id="778E12DE79DD8B80E4D858CF020BEE36" pageId="0" pageNumber="19">Proportions: A3L/HL = 0.45;</paragraph>
|
||
<paragraph id="2787E2ED9490C56254AC87AB30ECCADC" pageId="0" pageNumber="19">EyL/ HW(-) = 0.62;</paragraph>
|
||
<paragraph id="6E9E96F492433D6455B357575B1A468D" pageId="0" pageNumber="19">PW/HW(+) = 1.45;</paragraph>
|
||
<paragraph id="3C3C4351FE0476BD73A7CB391280AC1C" pageId="0" pageNumber="19">PW/PL = 1.26;</paragraph>
|
||
<paragraph id="17D81D485F00838F41D09D1CA1960908" pageId="0" pageNumber="19">PW/PWb = 1.17;</paragraph>
|
||
<paragraph id="D020BDEE4E2327BDCA81898A7D341835" pageId="0" pageNumber="19">PWb/PWa = not available;</paragraph>
|
||
<paragraph id="029161A6F0CD0B83EE5E4C4CD5BBBCB0" pageId="0" pageNumber="19">EW/PW = 1.60;</paragraph>
|
||
<paragraph id="2E46261A17899D9E9F436179349F8FE3" pageId="0" pageNumber="19">EL/EW = 1.55;</paragraph>
|
||
<paragraph id="E8CEF7A45E24A61DCAC802B5D5B63C02" pageId="0" pageNumber="19">EpL/EpW = 1.40;</paragraph>
|
||
<paragraph id="F86036835086D44478D2638F35FFF89E" pageId="0" pageNumber="19">EL/FL = 2.42;</paragraph>
|
||
<paragraph id="8AC31B54C62FC3041AA3E33023A984A3" pageId="0" pageNumber="19">EL/AedL = 4.07.</paragraph>
|
||
<paragraph id="7CA489A35C4008F74758C4B775916634" pageId="0" pageNumber="19">Head: Patterns of microsculpture could not be studied. In all other characters as described for the new genus, above.</paragraph>
|
||
<caption id="872EC154E4FF296EBC71FF7ADB45B9F3" doi="10.3897/dez.69.79931.figures78-80" httpUri="https://binary.pensoft.net/fig/643965" pageId="0" pageNumber="19" start="Figures 78–80" startId="F15">
|
||
<paragraph id="A95A4EA53461E9ED2AD402F772920E21" pageId="0" pageNumber="19">
|
||
<emphasis id="E8B85881D5AC4EB96B071FF67B1894C5" bold="true" pageId="0" pageNumber="19">Figures 78-80.</emphasis>
|
||
<taxonomicName id="A3CB7A913062DD5A0461C507D9368582" authorityName="Schmidt & Will" authorityYear="2022" class="Insecta" family="Carabidae" genus="Quasicalathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quasicalathus conservans" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="conservans">
|
||
<emphasis id="97EB12A9189564364C20E8A11FDDA468" italics="true" pageId="0" pageNumber="19">Quasicalathus conservans</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., volume rendering of the holotype using different grey scales of the Amira software.
|
||
<emphasis id="D9DFA2FAAAD09AAF025868A2D6507079" bold="true" pageId="0" pageNumber="19">78.</emphasis>
|
||
Dorsal aspect;
|
||
<emphasis id="9F081D8B13E776B2838E16B82F35635D" bold="true" pageId="0" pageNumber="19">79.</emphasis>
|
||
Left lateral aspect (
|
||
<emphasis id="E674756D8899A1AF54249A932D4AF747" bold="true" pageId="0" pageNumber="19">aed</emphasis>
|
||
- aedeagus);
|
||
<emphasis id="3910CDA517FDEEA63516C768EB5F7BF4" bold="true" pageId="0" pageNumber="19">80.</emphasis>
|
||
Ventral aspect; the aedeagus (highlighted by red colour) was separated by the segmentation function of Amira software.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="EB692D81C75833B3C64F528F9A5F64BD" pageId="0" pageNumber="19">
|
||
Prothorax: Pronotal lateral margin more markedly narrowed toward base than in
|
||
<taxonomicName id="087AC2CC41DAF5B24FDC855312CD1823" class="Insecta" kingdom="Animalia" lsidName="Q. elpis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="elpis">
|
||
<emphasis id="50BDAEDBDE471990817722780F4DE4F9" italics="true" pageId="0" pageNumber="19">Q. elpis</emphasis>
|
||
</taxonomicName>
|
||
, straight just before laterobasal angles, angles markedly obtuse (Figs
|
||
<figureCitation id="D090733AFFF45139B88B6239330789AC" captionStart="Figures 65–72" captionStartId="F13" captionText="Figures 65 - 72. Quasicalathus agonicollis sp. nov., volume rendering of the holotype using different grey scales of the Amira software. 65. Dorsal aspect (the negative imprint of the fossil on the inclusion wall is shown); 66. Basal portion of pronotum and anterior part of elytra (positive of the fossilized beetle is shown; the arrows point to the insertions of the pronotal basolateral setae and the parascutellar setae); 67. Metacoxa (the arrows point to the insertions of the three coxal setae each side); 68 - 71. Aedeagus in dorsal aspect (68.), Right lateral aspect (69.), Ventral aspect (70.), Left lateral aspect (71.); The remains of the parameres are coloured (red: left paramere; green: right paramere); 72. left lateral aspect of beetle body; the aedeagus (highlighted by red colour) was separated by the segmentation function of Amira software. Abbreviations: bb - basal bulb of aedeagal median lobe; hm - humerus; os - distal ostium of median lobe; sc - scutellum; tl - terminal lamella of median lobe." figureDoi="10.3897/dez.69.79931.figures65-72" httpUri="https://binary.pensoft.net/fig/643963" pageId="0" pageNumber="19">65</figureCitation>
|
||
,
|
||
<figureCitation id="E20B4CA81BF4D661584D786E35175516" captionStart="Figures 65–72" captionStartId="F13" captionText="Figures 65 - 72. Quasicalathus agonicollis sp. nov., volume rendering of the holotype using different grey scales of the Amira software. 65. Dorsal aspect (the negative imprint of the fossil on the inclusion wall is shown); 66. Basal portion of pronotum and anterior part of elytra (positive of the fossilized beetle is shown; the arrows point to the insertions of the pronotal basolateral setae and the parascutellar setae); 67. Metacoxa (the arrows point to the insertions of the three coxal setae each side); 68 - 71. Aedeagus in dorsal aspect (68.), Right lateral aspect (69.), Ventral aspect (70.), Left lateral aspect (71.); The remains of the parameres are coloured (red: left paramere; green: right paramere); 72. left lateral aspect of beetle body; the aedeagus (highlighted by red colour) was separated by the segmentation function of Amira software. Abbreviations: bb - basal bulb of aedeagal median lobe; hm - humerus; os - distal ostium of median lobe; sc - scutellum; tl - terminal lamella of median lobe." figureDoi="10.3897/dez.69.79931.figures65-72" httpUri="https://binary.pensoft.net/fig/643963" pageId="0" pageNumber="19">66</figureCitation>
|
||
). Prosternal process with traces of lateral bead evident. In all other characters as described for the new genus, above.
|
||
</paragraph>
|
||
<paragraph id="D2965C37E504EF380A6BE6CD1161E671" pageId="0" pageNumber="19">
|
||
Pterothorax: Elytra with basal margin moderately concave and humerus moderately protruded anteriorly; basal margin forming a more obtuse angle (ca. 125°) with lateral margin (Figs
|
||
<figureCitation id="B73A659E3816F23F98056279BFC14E66" captionStart="Figures 65–72" captionStartId="F13" captionText="Figures 65 - 72. Quasicalathus agonicollis sp. nov., volume rendering of the holotype using different grey scales of the Amira software. 65. Dorsal aspect (the negative imprint of the fossil on the inclusion wall is shown); 66. Basal portion of pronotum and anterior part of elytra (positive of the fossilized beetle is shown; the arrows point to the insertions of the pronotal basolateral setae and the parascutellar setae); 67. Metacoxa (the arrows point to the insertions of the three coxal setae each side); 68 - 71. Aedeagus in dorsal aspect (68.), Right lateral aspect (69.), Ventral aspect (70.), Left lateral aspect (71.); The remains of the parameres are coloured (red: left paramere; green: right paramere); 72. left lateral aspect of beetle body; the aedeagus (highlighted by red colour) was separated by the segmentation function of Amira software. Abbreviations: bb - basal bulb of aedeagal median lobe; hm - humerus; os - distal ostium of median lobe; sc - scutellum; tl - terminal lamella of median lobe." figureDoi="10.3897/dez.69.79931.figures65-72" httpUri="https://binary.pensoft.net/fig/643963" pageId="0" pageNumber="19">65</figureCitation>
|
||
,
|
||
<figureCitation id="8C54621D76FABFF289DE4F5995595DD2" captionStart="Figures 65–72" captionStartId="F13" captionText="Figures 65 - 72. Quasicalathus agonicollis sp. nov., volume rendering of the holotype using different grey scales of the Amira software. 65. Dorsal aspect (the negative imprint of the fossil on the inclusion wall is shown); 66. Basal portion of pronotum and anterior part of elytra (positive of the fossilized beetle is shown; the arrows point to the insertions of the pronotal basolateral setae and the parascutellar setae); 67. Metacoxa (the arrows point to the insertions of the three coxal setae each side); 68 - 71. Aedeagus in dorsal aspect (68.), Right lateral aspect (69.), Ventral aspect (70.), Left lateral aspect (71.); The remains of the parameres are coloured (red: left paramere; green: right paramere); 72. left lateral aspect of beetle body; the aedeagus (highlighted by red colour) was separated by the segmentation function of Amira software. Abbreviations: bb - basal bulb of aedeagal median lobe; hm - humerus; os - distal ostium of median lobe; sc - scutellum; tl - terminal lamella of median lobe." figureDoi="10.3897/dez.69.79931.figures65-72" httpUri="https://binary.pensoft.net/fig/643963" pageId="0" pageNumber="19">66</figureCitation>
|
||
). Elytral striae moderately deeply engraved, intervals moderately convex. In all other characters as described for the new genus, above.
|
||
</paragraph>
|
||
<paragraph id="E77B68AD274D9CCE30B4C3B2E940D97D" pageId="0" pageNumber="19">
|
||
Aedeagus: Length of median lobe 1.12 mm. Median lobe terminal lamella almost evenly narrowed from base to apex with side margins almost straight; apex slightly bent ventrally (Fig.
|
||
<figureCitation id="73D46966E53E0F61B6DAE3F0FAAE1FA2" captionStart="Figures 65–72" captionStartId="F13" captionText="Figures 65 - 72. Quasicalathus agonicollis sp. nov., volume rendering of the holotype using different grey scales of the Amira software. 65. Dorsal aspect (the negative imprint of the fossil on the inclusion wall is shown); 66. Basal portion of pronotum and anterior part of elytra (positive of the fossilized beetle is shown; the arrows point to the insertions of the pronotal basolateral setae and the parascutellar setae); 67. Metacoxa (the arrows point to the insertions of the three coxal setae each side); 68 - 71. Aedeagus in dorsal aspect (68.), Right lateral aspect (69.), Ventral aspect (70.), Left lateral aspect (71.); The remains of the parameres are coloured (red: left paramere; green: right paramere); 72. left lateral aspect of beetle body; the aedeagus (highlighted by red colour) was separated by the segmentation function of Amira software. Abbreviations: bb - basal bulb of aedeagal median lobe; hm - humerus; os - distal ostium of median lobe; sc - scutellum; tl - terminal lamella of median lobe." figureDoi="10.3897/dez.69.79931.figures65-72" httpUri="https://binary.pensoft.net/fig/643963" pageId="0" pageNumber="19">70</figureCitation>
|
||
). In all other characters as described for the new genus, above.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3284F8EF08089550DDE6961914C025CF" pageId="0" pageNumber="19" type="description">
|
||
<paragraph id="3D3977FA5481C35CD3784F8F8E494ACA" pageId="0" pageNumber="19">Description of specimen GZG 16188.</paragraph>
|
||
<paragraph id="D4EE9A25A0300DC41883AB05EE437579" pageId="0" pageNumber="19">
|
||
Measurements see Table
|
||
<tableCitation id="DB73AEB355A807A7CE9287DB2FBA5F0F" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Measurements [µm]." httpUri="http://table.plazi.org/id/C19891A363A03B4DD93DC20ECFC36EE4" pageId="0" pageNumber="19" tableUuid="C19891A363A03B4DD93DC20ECFC36EE4">2</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="724D14FD61426135DEEFAF41DA75B3EA" pageId="0" pageNumber="19">Standardized body length: 6.9 mm.</paragraph>
|
||
<paragraph id="5DFEF59A2B554DCD57AC5ED36791EBEA" pageId="0" pageNumber="19">Proportions: A3L/HL = 0.44;</paragraph>
|
||
<paragraph id="F8CFB085EB9044640AF1A39333DA582E" pageId="0" pageNumber="19">EyL/ HW(-) = 0.65;</paragraph>
|
||
<paragraph id="F49F6FD8CF1D3DCEF516237113A5E97A" pageId="0" pageNumber="19">PW/HW(+) = 1.46;</paragraph>
|
||
<paragraph id="E75CF5D621A4ADC12E5ECEBF76629FC1" pageId="0" pageNumber="19">PW/PL = 1.26;</paragraph>
|
||
<paragraph id="A2260C5AE75E74AA3563F649EA3A6881" pageId="0" pageNumber="19">PW/PWb = 1.12;</paragraph>
|
||
<paragraph id="9F3DF2F31CCAF1BB2F896DF6BA656EFC" pageId="0" pageNumber="19">PWb/PWa = 1.40;</paragraph>
|
||
<paragraph id="673CC82B1EB82DB5550DB713550F867B" pageId="0" pageNumber="19">EW/PW = 1.63;</paragraph>
|
||
<paragraph id="AA4631DC93B9D0621E73A8C21FFD6416" pageId="0" pageNumber="19">EL/EW = 1.53;</paragraph>
|
||
<paragraph id="EE26777D8E2B33C4F175B02C6547D2C9" pageId="0" pageNumber="19">EpL/EpW = 1.62;</paragraph>
|
||
<paragraph id="1E82D8D28468715FFBC28D5F0BCBE9B7" pageId="0" pageNumber="19">EL/FL = 2.55.</paragraph>
|
||
<paragraph id="009D2A6C2ADAB731A213650DE14291F0" pageId="0" pageNumber="19">
|
||
Head: Microsculpture consists of moderately large isodiametric sculpticells (Fig.
|
||
<figureCitation id="8863A8D531FA8B3C6A7C1A8956155620" captionStart="Figures 18–24" captionStartId="F4" captionText="Figures 18 - 24. Quasicalathus, light microscopic images of Q. elpis Ortuno & Arillo, 2009 (18 - 20.) and Q. agonicollis sp. nov. (21 - 24.). 18. General view of the amber piece " MAIG 76 " (only that part of the large amber piece bearing the Quasicalathus fossil is shown; the fossil is widely covered by milky coating); 19. General view of the two fragments of specimen " GZG 16185 "; the left one bears only the negative imprint of the left elytra on the inclusion wall; 20. Right anterior part of body of specimen " GZG 16185 " showing part of head, pronotum and humerus; 21, 22. General view of the amber piece " GZG 16188 " (21. With fossil in dorsal view; 22. In ventral view); 23. Anterior part of specimen " GZG 16188 "; 24. Head of specimen " GZG 16188 ". Abbreviations: a 1 - a 6 - antennomeres 1 - 6; as - anterior supraorbital seta; bs - pronotal laterobasal seta; ms - pronotal lateral seta; ps - posterior supraorbital seta." figureDoi="10.3897/dez.69.79931.figures18-24" httpUri="https://binary.pensoft.net/fig/643954" pageId="0" pageNumber="19">24</figureCitation>
|
||
). In all other characters as described for the new genus, above.
|
||
</paragraph>
|
||
<paragraph id="509A8270686FE5AD90F828B3AD508BE8" pageId="0" pageNumber="19">
|
||
Prothorax: Pronotal lateral margin almost completely rounded, straight just before laterobasal angles, angles markedly obtuse (Figs
|
||
<figureCitation id="047ECF4EC7E0330885899BC1CD5963E4" captionStart="Figures 18–24" captionStartId="F4" captionText="Figures 18 - 24. Quasicalathus, light microscopic images of Q. elpis Ortuno & Arillo, 2009 (18 - 20.) and Q. agonicollis sp. nov. (21 - 24.). 18. General view of the amber piece " MAIG 76 " (only that part of the large amber piece bearing the Quasicalathus fossil is shown; the fossil is widely covered by milky coating); 19. General view of the two fragments of specimen " GZG 16185 "; the left one bears only the negative imprint of the left elytra on the inclusion wall; 20. Right anterior part of body of specimen " GZG 16185 " showing part of head, pronotum and humerus; 21, 22. General view of the amber piece " GZG 16188 " (21. With fossil in dorsal view; 22. In ventral view); 23. Anterior part of specimen " GZG 16188 "; 24. Head of specimen " GZG 16188 ". Abbreviations: a 1 - a 6 - antennomeres 1 - 6; as - anterior supraorbital seta; bs - pronotal laterobasal seta; ms - pronotal lateral seta; ps - posterior supraorbital seta." figureDoi="10.3897/dez.69.79931.figures18-24" httpUri="https://binary.pensoft.net/fig/643954" pageId="0" pageNumber="19">23</figureCitation>
|
||
,
|
||
<figureCitation id="5D338D89812960EA6B8B6100104F5282" captionStart="Figures 25–30" captionStartId="F5" captionText="Figures 25 - 30. Quasicalathus agonicollis sp. nov., light microscopic images of specimen " GZG 16188 " (25 - 26.) and the holotype (28 - 30.). 25. Posterior part of pronotum and anterior part of elytra, right side; 26. Anterior part of fifth interval of left elytra showing microsculpture; 27. Posterior part of elytra (the arrows point to the insertions of the discal setae); 28. General view of the amber piece; 29. Right dorsal view of beetle body (the arrow points to the insertion of the discal seta on left elytron); 30. Left ventral view. Abbreviations: bs - insertion of the pronotal laterobasal seta; hm - humerus; m - mite (syninclusion)." figureDoi="10.3897/dez.69.79931.figures25-30" httpUri="https://binary.pensoft.net/fig/643955" pageId="0" pageNumber="19">25</figureCitation>
|
||
,
|
||
<figureCitation id="71BDBFF4330C114351EFDD07AD30913E" captionStart="Figures 73–77" captionStartId="F14" captionText="Figures 73 - 77. Quasicalathus agonicollis sp. nov., volume rendering of specimen " GZG 16188 ". 73. Dorsal aspect; 74. Basal portion of pronotum and anterior part of elytra (right side of body; the arrows point to the insertions of the pronotal basolateral seta and the parascutellar seta); 75. Prosternum with basal portion of prolegs; 76. Left apical gonocoxite, ventral aspect; 77. Gonocoxites, ventral aspect. Abbreviations: cx - procoxa; des - dorsal ensiform setae; fm - profemur; gx 1 - basal gonocoxite; gx 2 - apical gonocoxite; hm - humerus; pst - prosternum; sc - scutellum; sps - setae of sensory pit; tr - protrochanter; ves - ventral ensiform setae." figureDoi="10.3897/dez.69.79931.figures73-77" httpUri="https://binary.pensoft.net/fig/643964" pageId="0" pageNumber="19">73</figureCitation>
|
||
,
|
||
<figureCitation id="7C73FC181485F11744909C0495272CE5" captionStart="Figures 73–77" captionStartId="F14" captionText="Figures 73 - 77. Quasicalathus agonicollis sp. nov., volume rendering of specimen " GZG 16188 ". 73. Dorsal aspect; 74. Basal portion of pronotum and anterior part of elytra (right side of body; the arrows point to the insertions of the pronotal basolateral seta and the parascutellar seta); 75. Prosternum with basal portion of prolegs; 76. Left apical gonocoxite, ventral aspect; 77. Gonocoxites, ventral aspect. Abbreviations: cx - procoxa; des - dorsal ensiform setae; fm - profemur; gx 1 - basal gonocoxite; gx 2 - apical gonocoxite; hm - humerus; pst - prosternum; sc - scutellum; sps - setae of sensory pit; tr - protrochanter; ves - ventral ensiform setae." figureDoi="10.3897/dez.69.79931.figures73-77" httpUri="https://binary.pensoft.net/fig/643964" pageId="0" pageNumber="19">74</figureCitation>
|
||
). Prosternal process with traces of an apical bead (Fig.
|
||
<figureCitation id="E5218BE20259F92A86DCFAFFEAA732EF" captionStart="Figures 73–77" captionStartId="F14" captionText="Figures 73 - 77. Quasicalathus agonicollis sp. nov., volume rendering of specimen " GZG 16188 ". 73. Dorsal aspect; 74. Basal portion of pronotum and anterior part of elytra (right side of body; the arrows point to the insertions of the pronotal basolateral seta and the parascutellar seta); 75. Prosternum with basal portion of prolegs; 76. Left apical gonocoxite, ventral aspect; 77. Gonocoxites, ventral aspect. Abbreviations: cx - procoxa; des - dorsal ensiform setae; fm - profemur; gx 1 - basal gonocoxite; gx 2 - apical gonocoxite; hm - humerus; pst - prosternum; sc - scutellum; sps - setae of sensory pit; tr - protrochanter; ves - ventral ensiform setae." figureDoi="10.3897/dez.69.79931.figures73-77" httpUri="https://binary.pensoft.net/fig/643964" pageId="0" pageNumber="19">75</figureCitation>
|
||
). In all other characters as described for the new genus, above.
|
||
</paragraph>
|
||
<paragraph id="533EC6700D9510314070DC4C8A321517" pageId="0" pageNumber="19">
|
||
Pterothorax: Elytra with basal margin moderately concave and humerus moderately protruded anteriorly; basal margin forming a more obtuse angle (ca. 125°) with lateral margin (Figs
|
||
<figureCitation id="C8026AE05F081F52A473D49476F047B8" captionStart="Figures 18–24" captionStartId="F4" captionText="Figures 18 - 24. Quasicalathus, light microscopic images of Q. elpis Ortuno & Arillo, 2009 (18 - 20.) and Q. agonicollis sp. nov. (21 - 24.). 18. General view of the amber piece " MAIG 76 " (only that part of the large amber piece bearing the Quasicalathus fossil is shown; the fossil is widely covered by milky coating); 19. General view of the two fragments of specimen " GZG 16185 "; the left one bears only the negative imprint of the left elytra on the inclusion wall; 20. Right anterior part of body of specimen " GZG 16185 " showing part of head, pronotum and humerus; 21, 22. General view of the amber piece " GZG 16188 " (21. With fossil in dorsal view; 22. In ventral view); 23. Anterior part of specimen " GZG 16188 "; 24. Head of specimen " GZG 16188 ". Abbreviations: a 1 - a 6 - antennomeres 1 - 6; as - anterior supraorbital seta; bs - pronotal laterobasal seta; ms - pronotal lateral seta; ps - posterior supraorbital seta." figureDoi="10.3897/dez.69.79931.figures18-24" httpUri="https://binary.pensoft.net/fig/643954" pageId="0" pageNumber="19">23</figureCitation>
|
||
,
|
||
<figureCitation id="4430971A01F12B2924C5C1145F7B74A3" captionStart="Figures 73–77" captionStartId="F14" captionText="Figures 73 - 77. Quasicalathus agonicollis sp. nov., volume rendering of specimen " GZG 16188 ". 73. Dorsal aspect; 74. Basal portion of pronotum and anterior part of elytra (right side of body; the arrows point to the insertions of the pronotal basolateral seta and the parascutellar seta); 75. Prosternum with basal portion of prolegs; 76. Left apical gonocoxite, ventral aspect; 77. Gonocoxites, ventral aspect. Abbreviations: cx - procoxa; des - dorsal ensiform setae; fm - profemur; gx 1 - basal gonocoxite; gx 2 - apical gonocoxite; hm - humerus; pst - prosternum; sc - scutellum; sps - setae of sensory pit; tr - protrochanter; ves - ventral ensiform setae." figureDoi="10.3897/dez.69.79931.figures73-77" httpUri="https://binary.pensoft.net/fig/643964" pageId="0" pageNumber="19">73</figureCitation>
|
||
,
|
||
<figureCitation id="49D76BE46E6C6C919255630CD97B5968" captionStart="Figures 73–77" captionStartId="F14" captionText="Figures 73 - 77. Quasicalathus agonicollis sp. nov., volume rendering of specimen " GZG 16188 ". 73. Dorsal aspect; 74. Basal portion of pronotum and anterior part of elytra (right side of body; the arrows point to the insertions of the pronotal basolateral seta and the parascutellar seta); 75. Prosternum with basal portion of prolegs; 76. Left apical gonocoxite, ventral aspect; 77. Gonocoxites, ventral aspect. Abbreviations: cx - procoxa; des - dorsal ensiform setae; fm - profemur; gx 1 - basal gonocoxite; gx 2 - apical gonocoxite; hm - humerus; pst - prosternum; sc - scutellum; sps - setae of sensory pit; tr - protrochanter; ves - ventral ensiform setae." figureDoi="10.3897/dez.69.79931.figures73-77" httpUri="https://binary.pensoft.net/fig/643964" pageId="0" pageNumber="19">74</figureCitation>
|
||
). Elytral striae shallowly engraved, intervals rather flat. In all other characters as described for the new genus, above.
|
||
</paragraph>
|
||
<paragraph id="9E858ECF7A56B5DB242A6F7964F38D4E" pageId="0" pageNumber="19">
|
||
Female genitalia: Length of apical gonocoxite about 0.15 mm; shape see Figs
|
||
<figureCitation id="0E7C81C1ADB6AA473EF97D8FC274F877" captionStart="Figures 73–77" captionStartId="F14" captionText="Figures 73 - 77. Quasicalathus agonicollis sp. nov., volume rendering of specimen " GZG 16188 ". 73. Dorsal aspect; 74. Basal portion of pronotum and anterior part of elytra (right side of body; the arrows point to the insertions of the pronotal basolateral seta and the parascutellar seta); 75. Prosternum with basal portion of prolegs; 76. Left apical gonocoxite, ventral aspect; 77. Gonocoxites, ventral aspect. Abbreviations: cx - procoxa; des - dorsal ensiform setae; fm - profemur; gx 1 - basal gonocoxite; gx 2 - apical gonocoxite; hm - humerus; pst - prosternum; sc - scutellum; sps - setae of sensory pit; tr - protrochanter; ves - ventral ensiform setae." figureDoi="10.3897/dez.69.79931.figures73-77" httpUri="https://binary.pensoft.net/fig/643964" pageId="0" pageNumber="19">76</figureCitation>
|
||
,
|
||
<figureCitation id="96073724D5E26EDD7F516198A978BADF" captionStart="Figures 73–77" captionStartId="F14" captionText="Figures 73 - 77. Quasicalathus agonicollis sp. nov., volume rendering of specimen " GZG 16188 ". 73. Dorsal aspect; 74. Basal portion of pronotum and anterior part of elytra (right side of body; the arrows point to the insertions of the pronotal basolateral seta and the parascutellar seta); 75. Prosternum with basal portion of prolegs; 76. Left apical gonocoxite, ventral aspect; 77. Gonocoxites, ventral aspect. Abbreviations: cx - procoxa; des - dorsal ensiform setae; fm - profemur; gx 1 - basal gonocoxite; gx 2 - apical gonocoxite; hm - humerus; pst - prosternum; sc - scutellum; sps - setae of sensory pit; tr - protrochanter; ves - ventral ensiform setae." figureDoi="10.3897/dez.69.79931.figures73-77" httpUri="https://binary.pensoft.net/fig/643964" pageId="0" pageNumber="19">77</figureCitation>
|
||
; bursa copulatrix is not preserved. In all other characters as described for the new genus, above.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="41FF611018E584F5F8BECD2176A59F3E" pageId="0" pageNumber="19" type="differential diagnosis">
|
||
<paragraph id="4CFEB8E8B8F4701DD45D80A4FBCBFAFE" pageId="0" pageNumber="19">Differential diagnosis.</paragraph>
|
||
<paragraph id="7AF8E4482DD8A22079CBF61135BA09BD" pageId="0" pageNumber="19">
|
||
<taxonomicName id="1458EB101C88469BDFA63BDEFBEA4EF7" authorityName="Schmidt & Will" authorityYear="2022" class="Insecta" family="Carabidae" genus="Quasicalathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quasicalathus agonicollis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="agonicollis">
|
||
<emphasis id="7921AB3B38B5E95835772EED4E686824" italics="true" pageId="0" pageNumber="19">Quasicalathus agonicollis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. differs from
|
||
<taxonomicName id="91759E08E6CD54EC08EFE0788EDD17E7" class="Insecta" kingdom="Animalia" lsidName="Q. elpis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="elpis">
|
||
<emphasis id="8BC255018C769B47AB505400C459F2C9" italics="true" pageId="0" pageNumber="19">Q. elpis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="8730B61E05AADA0ECB601CE1E8C89572" class="Insecta" kingdom="Animalia" lsidName="Q. conservans" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="conservans">
|
||
<emphasis id="82823D000C628F32C81E443B71BCE0EA" italics="true" pageId="0" pageNumber="19">Q. conservans</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. by the smaller body (SBL <7 mm), more obtuse laterobasal angles of the pronotum, less concave elytral basal margin, less projected humeri, a more obtuse humeral angle (> 120°) and relatively smaller aedeagus. Based on specimen GZG 16188,
|
||
<taxonomicName id="D51C3805FE4F437FBC3CE2F7978AFB3F" class="Insecta" kingdom="Animalia" lsidName="Q. agonicollis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="agonicollis">
|
||
<emphasis id="287E13F6D56C44DF87A2476D7C064F60" italics="true" pageId="0" pageNumber="19">Q. agonicollis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. differs from
|
||
<taxonomicName id="A1F51E88ACEA53B289CEF0A557796673" class="Insecta" kingdom="Animalia" lsidName="Q. elpis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="elpis">
|
||
<emphasis id="6086A11D57B9D1A59C6742B68F567DC0" italics="true" pageId="0" pageNumber="19">Q. elpis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="32151512AD6808A186FE6916064753F0" class="Insecta" kingdom="Animalia" lsidName="Q. conservans" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="conservans">
|
||
<emphasis id="A7531D73B12735C9A31D3E6E1F13DF4E" italics="true" pageId="0" pageNumber="19">Q. conservans</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. additionally by presence of moderately large isodiametric sculpticells on head disc (small and transverse meshes in
|
||
<taxonomicName id="DEE23F394D3AD77067E34AD9AEE2BB87" class="Insecta" kingdom="Animalia" lsidName="Q. elpis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="elpis">
|
||
<emphasis id="96B9FC0A70CC66D866E129119F9130FF" italics="true" pageId="0" pageNumber="19">Q. elpis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="AF438D488613BD02AF5343102E55503A" class="Insecta" kingdom="Animalia" lsidName="Q. conservans" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="conservans">
|
||
<emphasis id="E9D59F4D437D8F01B698572B38B0146C" italics="true" pageId="0" pageNumber="19">Q. conservans</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.) and (from
|
||
<taxonomicName id="11131DC981ACA3954D795D94F1AC6570" class="Insecta" kingdom="Animalia" lsidName="Q. elpis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="elpis">
|
||
<emphasis id="82457B4B0948016D4B3B403D7182AFCF" italics="true" pageId="0" pageNumber="19">Q. elpis</emphasis>
|
||
</taxonomicName>
|
||
) by the smaller apical gonocoxite (unknown in
|
||
<taxonomicName id="338F2C0F26889D58EF6312244CD5F932" class="Insecta" kingdom="Animalia" lsidName="Q. conservans" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="conservans">
|
||
<emphasis id="EE552502D3FD639D3F2BF8AA54E7D2FE" italics="true" pageId="0" pageNumber="19">Q. conservans</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.). Due to the uncertainties in the identification of the
|
||
<taxonomicName id="82CC0BC12DB2740B7F962E8CEA816366" class="Insecta" kingdom="Animalia" lsidName="Q. agonicollis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="agonicollis">
|
||
<emphasis id="F032CEA84EAD8BBF9684E8A18F362162" italics="true" pageId="0" pageNumber="19">Q. agonicollis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. non-type female specimen (see Remarks above), the latter differential characters need confirmation based on additional material. The eyes of the
|
||
<taxonomicName id="C425B16041800D95579E1DD3D5092A04" class="Insecta" kingdom="Animalia" lsidName="Q. agonicollis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="agonicollis">
|
||
<emphasis id="861EB0F60E2B940C586C400904DAC0F6" italics="true" pageId="0" pageNumber="19">Q. agonicollis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. holotype and the GZG 16188 specimen are found to be proportionally smaller [EyL/ HW(-) = 0.62 resp. 0.65] than in
|
||
<taxonomicName id="53B247D7A55925A0A2A30C5E984E7A56" class="Insecta" kingdom="Animalia" lsidName="Q. elpis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="elpis">
|
||
<emphasis id="F3A0E1714A92A321B96BA76561F70E3B" italics="true" pageId="0" pageNumber="19">Q. elpis</emphasis>
|
||
</taxonomicName>
|
||
(0.72) and the
|
||
<taxonomicName id="02B55301E671DC3170FB6921FC99C72F" class="Insecta" kingdom="Animalia" lsidName="Q. conservans" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="conservans">
|
||
<emphasis id="4BB915B581C2E958D6B4ADC32309D535" italics="true" pageId="0" pageNumber="19">Q. conservans</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. holotype (0.72).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |