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<document id="7666D3B1901450397E4EBD9CC9924284" ID-CLB-Dataset="46393" ID-DOI="10.11646/zootaxa.3765.5.6" ID-GBIF-Dataset="535b33ec-3044-4fec-bafe-b024eb4a64ba" ID-ISSN="1175-5326" ID-Zenodo-Dep="226554" ID-ZooBank="9FCA7DAD-AB4A-4114-8312-CB4EC70AB8B8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460386680217" checkinUser="plazi" docAuthor="Clarkson, Bruno &amp; Ferreira-Jr, Nelson" docDate="2014" docId="5A73E070FFBCFFAC4083F2DBFB8DFD78" docLanguage="en" docName="zt03765p494.pdf" docOrigin="Zootaxa 3765 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chasmogenus lilianae Clarkson &amp; Ferreira-Jr, 2014, sp. nov." docType="treatment" docVersion="8" lastPageNumber="491" masterDocId="A64A9808FFB4FFA64014F715FFD6FF60" masterDocTitle="Four new species and first nominal record of Chasmogenus Sharp, 1882 (Coleoptera: Hydrophilidae) from Brazil" masterLastPageNumber="494" masterPageNumber="481" pageNumber="489" updateTime="1698297696019" updateUser="ExternalLinkService">
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<mods:title id="C981898ED36862C691FA13B7FACFCED4">Four new species and first nominal record of Chasmogenus Sharp, 1882 (Coleoptera: Hydrophilidae) from Brazil</mods:title>
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<mods:namePart id="8E744AC6BD5A6F9FB3E6FB7E41B76674">Clarkson, Bruno</mods:namePart>
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<mods:namePart id="3DA02A6D4D3B2CBE5907D5865CB06D52">Ferreira-Jr, Nelson</mods:namePart>
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<mods:title id="300CCA3894A178E4144C58B1864EDE9E">Zootaxa</mods:title>
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<mods:date id="D2767D310727C9EFBFD49A877E367685">2014</mods:date>
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<treatment id="5A73E070FFBCFFAC4083F2DBFB8DFD78" ID-DOI="http://doi.org/10.5281/zenodo.6131474" ID-GBIF-Taxon="119415650" ID-Zenodo-Dep="6131474" LSID="urn:lsid:plazi:treatment:5A73E070FFBCFFAC4083F2DBFB8DFD78" httpUri="http://treatment.plazi.org/id/5A73E070FFBCFFAC4083F2DBFB8DFD78" lastPageId="10" lastPageNumber="491" pageId="8" pageNumber="489">
<subSubSection id="9AC002EDFFBCFFAE4083F2DBFE6EF96A" pageId="8" pageNumber="489" type="nomenclature">
<paragraph id="D2655166FFBCFFAE4083F2DBFDDFFA88" blockId="8.[151,521,1486,1546]" box="[151,521,1486,1512]" pageId="8" pageNumber="489">
<heading id="892DE60AFFBCFFAE4083F2DBFDDFFA88" bold="true" box="[151,521,1486,1512]" fontSize="11" level="1" pageId="8" pageNumber="489" reason="1">
<emphasis id="E0AE8D74FFBCFFAE4083F2DBFDDFFA88" bold="true" box="[151,521,1486,1512]" pageId="8" pageNumber="489">
<taxonomicName id="15DA2AE5FFBCFFAE4083F2DBFE72FA88" ID-CoL="5XQ2D" box="[151,420,1486,1512]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="489" phylum="Arthropoda" rank="species" species="lilianae" status="sp. nov.">
<emphasis id="E0AE8D74FFBCFFAE4083F2DBFE72FA88" bold="true" box="[151,420,1486,1512]" italics="true" pageId="8" pageNumber="489">Chasmogenus lilianae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FB9D300FFFBCFFAE41B8F2DBFDDFFA88" box="[428,521,1486,1512]" pageId="8" pageNumber="489" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="D2655166FFBCFFAE4083F2E4FE6EF96A" blockId="8.[151,521,1486,1546]" box="[151,440,1521,1546]" pageId="8" pageNumber="489">(Figs. 5, 6, 16, 17 and 24)</paragraph>
</subSubSection>
<subSubSection id="9AC002EDFFBCFFAE4083F12CFCA8F9DE" pageId="8" pageNumber="489" type="materials_examined">
<paragraph id="D2655166FFBCFFAE4083F12CFCA8F9DE" blockId="8.[151,1436,1593,2014]" pageId="8" pageNumber="489">
<emphasis id="E0AE8D74FFBCFFAE4083F12CFDCFF932" bold="true" box="[151,537,1593,1618]" pageId="8" pageNumber="489">
<typeStatus id="0D61EFC4FFBCFFAE4083F12CFF04F932" box="[151,210,1593,1618]" pageId="8" pageNumber="489">Type</typeStatus>
material.
<typeStatus id="0D61EFC4FFBCFFAE415AF12CFE6DF932" box="[334,443,1593,1618]" pageId="8" pageNumber="489" type="holotype">Holotype</typeStatus>
(male):
</emphasis>
<collectingCountry id="AACD11F6FFBCFFAE4238F12FFD40F932" box="[556,662,1594,1618]" name="Brazil" pageId="8" pageNumber="489">BRASIL</collectingCountry>
: Rio de Janeiro,/ Nova Friburgo,/ Macaé de Cima,/ Tributário de 1a Ordem/ do Rio Macaé/
<date id="A66477A6FFBCFFAE418AF14BFDC8F916" box="[414,542,1630,1654]" pageId="8" pageNumber="489" value="2009-09-13">13.IX.2009</date>
,/ Casa amarela, campo/ das hortênsias/
<geoCoordinate id="B7EE37A1FFBCFFAE43C8F148FBA0F916" box="[988,1142,1629,1654]" direction="south" orientation="latitude" pageId="8" pageNumber="489" precision="1" value="-22.426805">22°2536.5”S</geoCoordinate>
<geoCoordinate id="B7EE37A1FFBCFFAE4468F148FACDF916" box="[1148,1307,1629,1654]" direction="east" orientation="longitude" pageId="8" pageNumber="489" precision="1" value="42.532333">
42°3156.4”
<collectingCountry id="AACD11F6FFBCFFAE4512F148FACDF916" box="[1286,1307,1629,1654]" name="Iceland" pageId="8" pageNumber="489">O</collectingCountry>
</geoCoordinate>
,/ Clarkson, B. col.” (
<collectionCode id="B4CBC9A3FFBCFFAE4100F197FE8DF9FA" box="[276,347,1666,1690]" pageId="8" pageNumber="489">DZRJ</collectionCode>
,
<taxonomicName id="15DA2AE5FFBCFFAE4167F194FE38F9FA" box="[371,494,1665,1690]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="489" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
3176);
<emphasis id="E0AE8D74FFBCFFAE4249F194FEECF9DE" bold="true" pageId="8" pageNumber="489">
<typeStatus id="0D61EFC4FFBCFFAE4249F194FD03F9FA" box="[605,725,1665,1690]" pageId="8" pageNumber="489" type="paratype">Paratypes</typeStatus>
(3):
<collectingCountry id="AACD11F6FFBCFFAE4331F194FC42F9FA" box="[805,916,1665,1690]" name="Brazil" pageId="8" pageNumber="489">BRAZIL</collectingCountry>
: Rio de Janeiro State: Nova Friburgo Municipality:
</emphasis>
Same as
<typeStatus id="0D61EFC4FFBCFFAE41B0F1B0FDDBF9DE" box="[420,525,1701,1726]" pageId="8" pageNumber="489" type="holotype">Holotype</typeStatus>
(3 ex.:
<collectionCode id="B4CBC9A3FFBCFFAE4276F1B3FD7EF9DD" box="[610,680,1702,1725]" pageId="8" pageNumber="489">DZRJ</collectionCode>
,
<taxonomicName id="15DA2AE5FFBCFFAE42A7F1B0FCA4F9DE" box="[691,882,1701,1726]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="489" phylum="Arthropoda" rank="order">Coleoptera 2932</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="9AC002EDFFBCFFAF40D3F1DCFE11FD00" lastPageId="9" lastPageNumber="490" pageId="8" pageNumber="489" type="diagnosis">
<paragraph id="D2655166FFBCFFAF40D3F1DCFE1FFFB4" blockId="8.[151,1436,1593,2014]" lastBlockId="9.[151,1436,151,2013]" lastPageId="9" lastPageNumber="490" pageId="8" pageNumber="489">
<emphasis id="E0AE8D74FFBCFFAE40D3F1DCFE97F982" bold="true" box="[199,321,1737,1762]" pageId="8" pageNumber="489">Diagnosis.</emphasis>
Medium to large size species (
<quantity id="1522FC83FFBCFFAE42BDF1DCFC83F982" box="[681,853,1737,1762]" metricMagnitude="-3" metricUnit="m" metricValue="4.175" metricValueMax="4.55" metricValueMin="3.8" pageId="8" pageNumber="489" unit="mm" value="4.175" valueMax="4.55" valueMin="3.8">3.804.55 mm</quantity>
). Head densely punctated. Anterior clypeal margin with narrow emargination, round to angulate shaped, exposing a gap between clypeus and labrum (
<emphasis id="E0AE8D74FFBCFFAE451DF1FBFAE7F865" box="[1289,1329,1774,1797]" italics="true" pageId="8" pageNumber="489">e.g.</emphasis>
Fig. 28). Anterior margin of labrum slightly concave medially. Anterocentral notch developed, wide and semicircular (Figs. 16 and 17). General dorsal punctures of pronotum and elytra fine, much smaller than systematic punctures. Elytra with four weakly distinct longitudinal rows of systematic punctures in dorsal view (Fig. 5). Prosternum not carinate. Mesoventral carina restricted to posterior half, slightly raised (Fig. 6).
<collectingCountry id="AACD11F6FFBCFFAE443EF068FBB4F8F6" box="[1066,1122,1917,1942]" name="India" pageId="8" pageNumber="489">Hind</collectingCountry>
femora pubescent on basal nine-tenths (Fig. 6). Ventrite 5 with shallow apical emargination (Fig. 6). Phallobase short, manubrium distinct, narrow. Parameres as long as median lobe, inner margin slightly emarginated on apical three-tenths, outer margin slightly concave. Median lobe thick, gradually narrowing apicad, apex blunt; basal apophyses slightly convex, within parameres (Fig. 24).
</paragraph>
<paragraph id="D2655166FFBDFFAF40D3F7CAFE11FD00" blockId="9.[151,1436,151,2013]" pageId="9" pageNumber="490">
The new species can be distinguished from other Neotropical species meanly for the male genitalia characteristics (Fig. 24): for the phallobase wider on apical portion, near of the junction with the parameres; parameres as long as median lobe, with outer margin slightly concave and the median lobe apex blunt (by these last two characteristics it resembles
<taxonomicName id="15DA2AE5FFBDFFAF4218F658FD54FE04" box="[524,642,333,356]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="490" phylum="Arthropoda" rank="species" species="itatiaia" status="sp. nov.">
<emphasis id="E0AE8D74FFBDFFAF4218F658FD54FE04" box="[524,642,333,356]" italics="true" pageId="9" pageNumber="490">C. itatiaia</emphasis>
</taxonomicName>
<emphasis id="E0AE8D74FFBDFFAF4298F659FD3FFE05" bold="true" box="[652,745,332,357]" pageId="9" pageNumber="490">
<taxonomicNameLabel id="FB9D300FFFBDFFAF4298F659FD3FFE05" box="[652,745,332,357]" pageId="9" pageNumber="490" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
). Concerning South American species,
<taxonomicName id="15DA2AE5FFBDFFAF44AEF658FAE2FE05" box="[1210,1332,332,357]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="490" phylum="Arthropoda" rank="species" species="lilianae" status="sp. nov.">
<emphasis id="E0AE8D74FFBDFFAF44AEF658FAE2FE05" box="[1210,1332,332,357]" italics="true" pageId="9" pageNumber="490">C. lilianae</emphasis>
</taxonomicName>
<emphasis id="E0AE8D74FFBDFFAF452AF659FA4AFE05" bold="true" box="[1342,1436,332,357]" pageId="9" pageNumber="490">
<taxonomicNameLabel id="FB9D300FFFBDFFAF452AF659FA4AFE05" box="[1342,1436,332,357]" pageId="9" pageNumber="490" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
present anterior clypeal emargination narrow and rounded to angulate (
<emphasis id="E0AE8D74FFBDFFAF43D0F664FC3AFEE8" box="[964,1004,369,392]" italics="true" pageId="9" pageNumber="490">e.g.</emphasis>
Fig. 28), as well as all Brazilian new species,
<taxonomicName id="15DA2AE5FFBDFFAF40E1F680FEA9FECD" box="[245,383,404,429]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="490" phylum="Arthropoda" rank="species" species="bariorum">
<emphasis id="E0AE8D74FFBDFFAF40E1F680FEA9FECD" box="[245,383,404,429]" italics="true" pageId="9" pageNumber="490">C. bariorum</emphasis>
</taxonomicName>
,
<taxonomicName id="15DA2AE5FFBDFFAF4199F680FDE2FECD" box="[397,564,404,429]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="490" phylum="Arthropoda" rank="species" species="occidentalis">
<emphasis id="E0AE8D74FFBDFFAF4199F680FDE2FECD" box="[397,564,404,429]" italics="true" pageId="9" pageNumber="490">C. occidentalis</emphasis>
</taxonomicName>
,
<taxonomicName id="15DA2AE5FFBDFFAF4256F680FD05FECD" box="[578,723,404,429]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="490" phylum="Arthropoda" rank="species" species="yukparum">
<emphasis id="E0AE8D74FFBDFFAF4256F680FD05FECD" box="[578,723,404,429]" italics="true" pageId="9" pageNumber="490">C. yukparum</emphasis>
</taxonomicName>
and
<taxonomicName id="15DA2AE5FFBDFFAF431CF680FC47FECC" box="[776,913,404,428]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="490" phylum="Arthropoda" rank="species" species="rufinasus">
<emphasis id="E0AE8D74FFBDFFAF431CF680FC47FECC" box="[776,913,404,428]" italics="true" pageId="9" pageNumber="490">C. rufinasus</emphasis>
</taxonomicName>
, differing this species from
<taxonomicName id="15DA2AE5FFBDFFAF44DEF680FA9BFECD" box="[1226,1357,404,429]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="490" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="E0AE8D74FFBDFFAF44DEF680FA9BFECD" box="[1226,1357,404,429]" italics="true" pageId="9" pageNumber="490">C. australis</emphasis>
</taxonomicName>
and
<taxonomicName id="15DA2AE5FFBDFFAF4596F680FF27FEB0" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="490" phylum="Arthropoda" rank="species" species="sapucay">
<emphasis id="E0AE8D74FFBDFFAF4596F680FF27FEB0" italics="true" pageId="9" pageNumber="490">C. sapucay</emphasis>
</taxonomicName>
, which present this emargination wide with its mesal portion straight (
<emphasis id="E0AE8D74FFBDFFAF4415F6ACFBFFFEB0" box="[1025,1065,441,464]" italics="true" pageId="9" pageNumber="490">e.g.</emphasis>
Fig. 30). The absence of a strong carina raised into a “tooth” on metaventrite differ this species, as well as all described species here, from Venezuelan species
<taxonomicName id="15DA2AE5FFBDFFAF4163F515FDD4FD78" box="[375,514,511,536]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="490" phylum="Arthropoda" rank="species" species="bariorum">
<emphasis id="E0AE8D74FFBDFFAF4163F515FDD4FD78" box="[375,514,511,536]" italics="true" pageId="9" pageNumber="490">C. bariorum</emphasis>
</taxonomicName>
,
<taxonomicName id="15DA2AE5FFBDFFAF4204F515FD6EFD78" box="[528,696,511,536]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="490" phylum="Arthropoda" rank="species" species="occidentalis">
<emphasis id="E0AE8D74FFBDFFAF4204F515FD6EFD78" box="[528,696,511,536]" italics="true" pageId="9" pageNumber="490">C. occidentalis</emphasis>
</taxonomicName>
and
<taxonomicName id="15DA2AE5FFBDFFAF42FBF515FC56FD78" box="[751,896,511,536]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="490" phylum="Arthropoda" rank="species" species="yukparum">
<emphasis id="E0AE8D74FFBDFFAF42FBF515FC56FD78" box="[751,896,511,536]" italics="true" pageId="9" pageNumber="490">C. yukparum</emphasis>
</taxonomicName>
. hind femora almost totally pubescent (on basal nine-tenths) (Fig. 6), characteristic shared with
<taxonomicName id="15DA2AE5FFBDFFAF42B8F530FCFEFD5D" box="[684,808,548,573]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="490" phylum="Arthropoda" rank="species" species="ubatuba" status="sp. nov.">
<emphasis id="E0AE8D74FFBDFFAF42B8F530FCFEFD5D" box="[684,808,548,573]" italics="true" pageId="9" pageNumber="490">C. ubatuba</emphasis>
</taxonomicName>
<emphasis id="E0AE8D74FFBDFFAF433BF531FC5FFD5D" bold="true" box="[815,905,548,573]" pageId="9" pageNumber="490">
<taxonomicNameLabel id="FB9D300FFFBDFFAF433BF531FC5FFD5D" box="[815,905,548,573]" pageId="9" pageNumber="490" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, is useful to distinguish these two species from the other Brazilian species.
</paragraph>
</subSubSection>
<subSubSection id="9AC002EDFFBDFFAC40D3F579FC47FEE8" lastPageId="10" lastPageNumber="491" pageId="9" pageNumber="490" type="description">
<paragraph id="D2655166FFBDFFAF40D3F579FDDAFDE5" blockId="9.[151,1436,151,2013]" box="[199,524,620,645]" pageId="9" pageNumber="490">
<emphasis id="E0AE8D74FFBDFFAF40D3F579FE8FFDE5" bold="true" box="[199,345,620,645]" pageId="9" pageNumber="490">Description.</emphasis>
<typeStatus id="0D61EFC4FFBDFFAF414BF579FE11FDE5" box="[351,455,620,645]" pageId="9" pageNumber="490" type="holotype">Holotype</typeStatus>
male.
</paragraph>
<paragraph id="D2655166FFBDFFAF40D3F585FAF9FDAD" blockId="9.[151,1436,151,2013]" pageId="9" pageNumber="490">
<emphasis id="E0AE8D74FFBDFFAF40D3F585FEBBFDC8" box="[199,365,655,680]" italics="true" pageId="9" pageNumber="490">Size and form.</emphasis>
Total length
<quantity id="1522FC83FFBDFFAF421CF59AFDA7FDC7" box="[520,625,655,680]" metricMagnitude="-3" metricUnit="m" metricValue="4.05" pageId="9" pageNumber="490" unit="mm" value="4.05">4.05 mm</quantity>
; maximum width
<quantity id="1522FC83FFBDFFAF4357F59AFC7BFDC7" box="[835,941,655,680]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" pageId="9" pageNumber="490" unit="mm" value="2.15">2.15 mm</quantity>
; maximum width of head
<quantity id="1522FC83FFBDFFAF44C9F585FA91FDC7" box="[1245,1351,656,680]" metricMagnitude="-3" metricUnit="m" metricValue="1.03" pageId="9" pageNumber="490" unit="mm" value="1.03">1.03 mm</quantity>
; elytral length
<quantity id="1522FC83FFBDFFAF40F0F5A1FE9DFDAC" box="[228,331,692,717]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" pageId="9" pageNumber="490" unit="mm" value="2.75">2.75 mm</quantity>
; maximum width of pronotum
<quantity id="1522FC83FFBDFFAF42BCF5A1FCD9FDAC" box="[680,783,692,717]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" pageId="9" pageNumber="490" unit="mm" value="1.85">1.85 mm</quantity>
. Body broadly oval, moderately convex (Fig. 5).
</paragraph>
<paragraph id="D2655166FFBDFFAF40D3F5C2FC69FCE0" blockId="9.[151,1436,151,2013]" pageId="9" pageNumber="490">
<emphasis id="E0AE8D74FFBDFFAF40D3F5C2FEDBFD90" box="[199,269,727,752]" italics="true" pageId="9" pageNumber="490">Color.</emphasis>
Dorsum of head and elytra uniformly dark brown, pronotum slightly paler than head with lateral and anterior margins often paler than pronotal disk (Fig. 5). Antennae with antennomeres 1-5 pale brown, club brown (Fig. 6). Maxillary palpi brown, labial palpi pale brown (Fig. 6). Ventral face dark brown, brown in the posteromedial glabrous area and behind metafemur in the metaventrite; lateral part of pronotum and epipleuron brown; legs brown, dark brown on pubescent portion of femora (Fig. 6).
</paragraph>
<paragraph id="D2655166FFBDFFAF40D3F499FE38FA18" blockId="9.[151,1436,151,2013]" pageId="9" pageNumber="490">
<emphasis id="E0AE8D74FFBDFFAF40D3F499FEDDFCC5" box="[199,267,908,933]" italics="true" pageId="9" pageNumber="490">Head.</emphasis>
Ground punctation on clypeus, frons and labrum very fine, distance between punctures 2.03.0× the width of one puncture. Systematic punctures on labrum consisting of several scattered punctures, somewhat larger than surrounding punctures, sometimes bearing short, fine setae. Clypeus with scattered distinct systematic punctures, mainly on anterolateral portion, about twice as large as surrounding punctures, bearing short, fine setae. Frons with irregular row of distinct systematic punctures along frontoclypeal suture, about 2.03.0x larger than surrounding punctures and some less distinct scattered systematic punctures mesally of eyes, both bearing short, fine setae. Anterior clypeal margin with narrow emargination, round to angulate shaped, exposing a gap between clypeus and labrum (
<emphasis id="E0AE8D74FFBDFFAF4193F39CFE79FBC0" box="[391,431,1161,1184]" italics="true" pageId="9" pageNumber="490">e.g.</emphasis>
Fig. 28). Anterior margin of labrum slightly concave medially. Mandibles bifid at apex. Maxillary palpi as long as maximun width of head; apical palpomere subequal in length than penultimate palpomere, palpomere slightly longer than penultimate palpomere (1.3× as long as penultimate one) (Figs. 5 and 6). Labial palpi about half the width of mentum; ultimate palpomere with a preapical long seta (
<emphasis id="E0AE8D74FFBDFFAF4511F3E0FAF8FA6C" box="[1285,1326,1269,1292]" italics="true" pageId="9" pageNumber="490">e.g.</emphasis>
Fig. 12). Mentum bare, with some short setae on lateral margins, rather punctuate, somewhat rugose, with deep notch on anterolateral margin and deeply depressed on anterocentral half, anterocentral noth developed, wide and rounded, semicircular (Figs. 16 and 17).
</paragraph>
<paragraph id="D2655166FFBDFFAF40D3F291FE3DF82D" blockId="9.[151,1436,151,2013]" pageId="9" pageNumber="490">
<emphasis id="E0AE8D74FFBDFFAF40D3F291FEC9FAFD" box="[199,287,1412,1437]" italics="true" pageId="9" pageNumber="490">Thorax.</emphasis>
Ground punctation on pronotum and elytra evenly distributed and very fine. Pronotum with two anterolateral and two posterolateral series of systematic punctures with short fine setae, each systematic puncture at least 2.0x the size of surrounding ground punctures: anterior series forming an irregular row curved backwards, beginning near anterolateral corners toward medial portion of pronutum; posterior series forming an irregular row on posterior half, perpendicular to lateral margins, beginning near of margin, restricted to lateral quarter (Fig. 5). Pronotal margins completely rimmed; lateral margins with sparse row of short setae, on anterior half; posterior margin slightly arcuate (Fig. 5). Elytra with four weakly distinct longitudinal irregular spaced rows of systematic punctures in dorsal view, punctures about 3.04.0x larger than ground punctures, bearing long, fine setae; sutural stria strongly impressed on posterior two-thirds (Fig. 5); Prosternum finely pubescent, weakly convex, not carinate medialy (
<emphasis id="E0AE8D74FFBDFFAF4110F1DCFEFBF980" box="[260,301,1737,1760]" italics="true" pageId="9" pageNumber="490">e.g.</emphasis>
Fig. 27). Mesoventrite with a longitudinal carina on medial part, restricted to posterior half, well defined but only slightly raised (Fig. 6). Metaventrite with posteromedial, enlongate oval glabrous area, twice longer than wide, about two-thirds of total length of metaventrite (Fig. 6).
<collectingCountry id="AACD11F6FFBDFFAF43FBF01AFBF1F848" box="[1007,1063,1807,1832]" name="India" pageId="9" pageNumber="490">Hind</collectingCountry>
femora almost totally pubescent (on basal nine-tenths) (Fig. 6).
</paragraph>
<paragraph id="D2655166FFBDFFAC40D3F042FD51FE20" blockId="9.[151,1436,151,2013]" lastBlockId="10.[151,1436,151,536]" lastPageId="10" lastPageNumber="491" pageId="9" pageNumber="490">
<emphasis id="E0AE8D74FFBDFFAF40D3F042FEEEF810" box="[199,312,1879,1904]" italics="true" pageId="9" pageNumber="490">Abdomen.</emphasis>
Ventrites densely and evenly pubescent. Ventrite 5 with shallow apical emargination, distinctly wider than deep, and lined with coarse yellow setae (Fig. 6). Aedeagus (Fig. 24) about
<quantity id="1522FC83FFBDFFAF44A3F069FAF6F8F4" box="[1207,1312,1916,1941]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" pageId="9" pageNumber="490" unit="mm" value="0.54">0.54 mm</quantity>
in length. Phallobase well developed, as long as wide, symmetrical, shorter than apical portion (ca. 0.30x the aedeagus in length), maximum width on apical portion; manubrium distinct, narrow, rounded at base, about half the length of phallobase. Parameres as long as median lobe, evenly wide from base to apical three-tenths, were gradually narrowing apicad, inner margin slightly emarginated on apical three-tenths in dorsal view, outer margin slightly concave; apex blunt, slightly turned inwards. Median lobe thick, gradually narrowing apicad, apex blunt; corona distinct, semicircular, wider than long, subapical (in apical fifth); basal apophyses slightly convex, within parameres, almost extended into phallobase.
</paragraph>
<paragraph id="D2655166FFBEFFAC40D3F659FC47FEE8" blockId="10.[151,1436,151,536]" pageId="10" pageNumber="491">
<emphasis id="E0AE8D74FFBEFFAC40D3F659FEE9FE05" bold="true" box="[199,319,332,357]" pageId="10" pageNumber="491">Variation.</emphasis>
Body length from
<quantity id="1522FC83FFBEFFAC4200F659FDAEFE04" box="[532,632,332,356]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="10" pageNumber="491" unit="mm" value="3.8">3.80 mm</quantity>
to
<quantity id="1522FC83FFBEFFAC4288F659FCD5FE04" box="[668,771,332,357]" metricMagnitude="-3" metricUnit="m" metricValue="4.55" pageId="10" pageNumber="491" unit="mm" value="4.55">4.55 mm</quantity>
. Significant variation on coloration and sexual dimorphism in the external morphology was not observed in the
<typeStatus id="0D61EFC4FFBEFFAC42C9F665FCDBFEE8" box="[733,781,368,392]" pageId="10" pageNumber="491">type</typeStatus>
specimens.
</paragraph>
</subSubSection>
<subSubSection id="9AC002EDFFBEFFAC40D3F681FCCEFEB0" pageId="10" pageNumber="491" type="etymology">
<paragraph id="D2655166FFBEFFAC40D3F681FCCEFEB0" blockId="10.[151,1436,151,536]" pageId="10" pageNumber="491">
<emphasis id="E0AE8D74FFBEFFAC40D3F681FE98FECD" bold="true" box="[199,334,404,429]" pageId="10" pageNumber="491">Etymology.</emphasis>
This species is dedicated to Dr. Liliana Alicia Fernández in recognition of her significant contributions to South American Hydrophiloidea studies.
</paragraph>
</subSubSection>
<subSubSection id="9AC002EDFFBEFFAC40D3F6C9FB8DFD78" pageId="10" pageNumber="491" type="biology_ecology">
<paragraph id="D2655166FFBEFFAC40D3F6C9FB8DFD78" blockId="10.[151,1436,151,536]" pageId="10" pageNumber="491">
<emphasis id="E0AE8D74FFBEFFAC40D3F6C9FEF1FE95" bold="true" box="[199,295,476,501]" pageId="10" pageNumber="491">Biology.</emphasis>
All
<typeStatus id="0D61EFC4FFBEFFAC414EF6C8FE5CFE95" box="[346,394,477,501]" pageId="10" pageNumber="491">type</typeStatus>
material of
<taxonomicName id="15DA2AE5FFBEFFAC4200F6C8FD5DFE95" box="[532,651,476,501]" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="491" phylum="Arthropoda" rank="species" species="lilianae" status="sp. nov.">
<emphasis id="E0AE8D74FFBEFFAC4200F6C8FD5DFE95" box="[532,651,476,501]" italics="true" pageId="10" pageNumber="491">C. lilianae</emphasis>
</taxonomicName>
<emphasis id="E0AE8D74FFBEFFAC4287F6C9FD3BFE95" bold="true" box="[659,749,476,501]" pageId="10" pageNumber="491">
<taxonomicNameLabel id="FB9D300FFFBEFFAC4287F6C9FD3BFE95" box="[659,749,476,501]" pageId="10" pageNumber="491" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was collected from standing water in a narrow groove about
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of width, among submerged leaf litter, in a mountainous area about
<quantity id="1522FC83FFBEFFAC43CDF515FBF6FD77" box="[985,1056,512,536]" metricMagnitude="2" metricUnit="m" metricValue="9.4" pageId="10" pageNumber="491" unit="m" value="940.0">940 m</quantity>
a.s.l.
</paragraph>
</subSubSection>
</treatment>
</document>