366 lines
42 KiB
XML
366 lines
42 KiB
XML
<document ID-CLB-Dataset="35706" ID-DOI="http://dx.doi.org/10.3897/zookeys.147.2117" ID-GBIF-Dataset="737d9e49-ca5a-4db8-8ad8-5b26fd663709" ID-PMC="PMC3286262" ID-Pensoft-Pub="1313-2970-147-399" ID-Pensoft-UUID="FFC09D1206663066632323645E417D20" ID-PubMed="22371669" ID-Zenodo-Dep="577410" ID-ZooBank="2296366206F44A6D82D31EFD1AD3D055" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-147-399" ModsDocOrigin="ZooKeys 147" ModsDocTitle="Badister Clairville 1806: A new species and new continental record for the nominate subgenus in Amazonian Perú (Coleoptera, Carabidae, Licinini)" checkinTime="1451249658171" checkinUser="pensoft" docAuthor="Erwin, Terry L. & Ball, George E." docDate="2011" docId="58603F7E4C238B927F604A2986FDED1F" docLanguage="en" docName="ZooKeys 147: 399-417" docOrigin="ZooKeys 147" docSource="http://dx.doi.org/10.3897/zookeys.147.2117" docTitle="Badister amazonus Erwin & Ball 2011, sp. n." docType="treatment" docUuid="D953E83C-2809-4DE7-91F2-40451DEBFDEA" docUuidSource="ZooBank" docVersion="7" id="FFC09D1206663066632323645E417D20" lastPageNumber="410" masterDocId="FFC09D1206663066632323645E417D20" masterDocTitle="Badister Clairville 1806: A new species and new continental record for the nominate subgenus in Amazonian Peru (Coleoptera, Carabidae, Licinini)" masterLastPageNumber="417" masterPageNumber="399" pageNumber="406" updateTime="1732852210654" updateUser="ExternalLinkService">
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<mods:titleInfo id="1A6702A07DF0FE83D357A2C0B3DCE9C3">
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<mods:title id="0551D66F911BD005FEF441897C98D35A">Badister Clairville 1806: A new species and new continental record for the nominate subgenus in Amazonian Peru (Coleoptera, Carabidae, Licinini)</mods:title>
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<mods:name id="052DF812C6FA341C9440AEC79702AEDF" type="personal">
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<mods:roleTerm id="4409F95E5101ACB088AF49ACFD68123B">Author</mods:roleTerm>
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<mods:namePart id="0EF3B41A46E8AB98D55D89FDBB447829">Erwin, Terry L.</mods:namePart>
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<mods:affiliation id="1238C1D9BD2CC521308035448CC30231">Hyper-diversity Group, Department of Entomology, MRC- 187, National Museum of Natural History, Smithsonian Institution, Washington, P. O. Box 37012, DC 20013 - 7012, USA</mods:affiliation>
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<mods:nameIdentifier id="48CC6651A51E61C9359AB5BBB6D531D8" type="email">erwint@si.edu</mods:nameIdentifier>
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<mods:roleTerm id="2E6704EA7FA375C1FA193F62F39D86B8">Author</mods:roleTerm>
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<mods:namePart id="819506F73EFAB442A67BEA6DB138CC0B">Ball, George E.</mods:namePart>
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<mods:affiliation id="3EDFB900175B9F7ABDE35FEDDECD5171">Department of Biological Sciences, University of Alberta, Edmonton, Canada</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="A4B12632FAED7C7DBB1941FF34701007">text</mods:typeOfResource>
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<mods:title id="6166915165FEF9E3D25E5D4DB2F777B0">ZooKeys</mods:title>
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<mods:part id="2063A52189D6F64670DE18A6FCC95813">
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<mods:date id="73C7FB71D7A3007FA77ED5E25364AEB2">2011</mods:date>
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<mods:detail id="651D2FF010A1D07AAA925CADFED1D90B" type="volume">
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<mods:number id="04FE33E7B1A66D3FB86A072873EF5B59">147</mods:number>
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<mods:start id="F5B1F6BE4608B13844D698727ABD3722">399</mods:start>
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<mods:url id="0BCEEDC8BAF8BAEAD9DB81AB7A6A98B6">http://dx.doi.org/10.3897/zookeys.147.2117</mods:url>
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<mods:classification id="3916341677F05FF32DC4815409CA4B39">journal article</mods:classification>
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<mods:identifier id="A8127CDD0FEBAC4AA8A11BD80388ECD2" type="DOI">http://dx.doi.org/10.3897/zookeys.147.2117</mods:identifier>
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<mods:identifier id="FBF04CFFA58D18F5D67D9674DF981E1E" type="Zenodo-Dep">577410</mods:identifier>
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<treatment id="58603F7E4C238B927F604A2986FDED1F" ID-GBIF-Taxon="152032058" LSID="urn:lsid:zoobank.org:act:D953E83C-2809-4DE7-91F2-40451DEBFDEA" httpUri="http://treatment.plazi.org/id/58603F7E4C238B927F604A2986FDED1F" lastPageId="11" lastPageNumber="410" pageId="7" pageNumber="406" scope_family="Carabidae" scope_order="Coleoptera" scope_subGenus="Badister" scope_tribe="Licinini">
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<subSubSection id="886BAF0C86F7846FA11F8C523ABAE82C" pageId="7" pageNumber="406" type="nomenclature">
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<paragraph id="5BD0210BCAD81567672DB4B54522FD0F" pageId="7" pageNumber="406">
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<taxonomicName id="E82B22581ED6CB32F278084C811AAA30" ID-CoL="KBR5" LSID="58603F7E-4C23-8B92-7F60-4A2986FDED1F" authority="Erwin & Ball" authorityName="Erwin & Ball" authorityYear="2011" class="Insecta" family="Carabidae" genus="Badister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Badister amazonus" order="Coleoptera" pageId="7" pageNumber="406" phylum="Arthropoda" rank="species" species="amazonus" status="sp. n.">Badister amazonus Erwin & Ball</taxonomicName>
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<taxonomicNameLabel id="56EB9A427690A8863A1BA60E19FC3FC2" pageId="7" pageNumber="406">sp. n.</taxonomicNameLabel>
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<figureCitation id="2FC0014F9E990E41474D8BA97AC9BDE3" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Badister amazonus sp. n., female, ADP 051544; type locality. Habitus, dorsal aspect. ABL = 7.13 mm." httpUri="https://binary.pensoft.net/fig/19730" pageId="7" pageNumber="406">Figures 1</figureCitation>
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<figureCitation id="C21F66330396F1E3157A029455BC601D" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Distribution dot map for known localities of Badister amazonus sp. n .. along the rivers Samiria and Ucayali (★)." httpUri="https://binary.pensoft.net/fig/19736" pageId="7" pageNumber="406">-7</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="E0BF96F777839A3FF4154B34017F2730" pageId="7" pageNumber="406" type="holotype">
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<paragraph id="01EB0DB705D27687E3C51F6FCCD0B135" pageId="7" pageNumber="406">Holotype.</paragraph>
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<paragraph id="CEF32772E89B1327E883781D149EFAAC" pageId="7" pageNumber="406">
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<emphasis id="BDC95FA5B6E696A1CFAE474443572369" bold="true" pageId="7" pageNumber="406">
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<normalizedToken id="A03D24EF248B9447B27B5BEC1BA4233E" originalValue="Perú">Peru</normalizedToken>
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</emphasis>
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, Loreto, 1.0 km SW boca del Rio Samiria, Vigilante Post 1, 130m, "
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<geoCoordinate id="97536DDA3F49E0DB90C5E4A646F9586E" degrees="04" direction="south" minutes="40.5" orientation="latitude" precision="92" value="-4.675">04°40.5'S</geoCoordinate>
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,
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<geoCoordinate id="2211F6CA043925FED2EC94A7AA4743DA" degrees="074" direction="west" minutes="18.9" orientation="longitude" precision="92" value="-74.315">074°18.9'W</geoCoordinate>
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" 31 August 1991 (T.L. Erwin & M.G. Pogue)(NMNH: ADP051824, male).
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</paragraph>
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</subSubSection>
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<subSubSection id="3564C75632AA61442CC2AE4581E69250" pageId="8" pageNumber="407" type="derivation of specific epithet">
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<paragraph id="EBCD5DBFDE1933C7348306CD4A6E338B" pageId="8" pageNumber="407">
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<pageBreakToken id="702377DFBCA9E946011346772260B883" pageId="8" pageNumber="407" start="start">Derivation</pageBreakToken>
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of specific epithet.
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</paragraph>
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<paragraph id="0D0DD7C4786EA8CBA323B2AE64814BF4" pageId="8" pageNumber="407">
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The epithet
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<normalizedToken id="359F9398EED4C417E8EAC80C5A200965" originalValue="“amazonus”">
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"
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<emphasis id="A9AB100A982BD91A608AB2AFA703F114" italics="true" pageId="8" pageNumber="407">amazonus</emphasis>
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"
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</normalizedToken>
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is a singular Latinized masculine noun in apposition, based on the name of the area in which these beetles were found.
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</paragraph>
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</subSubSection>
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<subSubSection id="64239B0EF83573DF3F59730D24294A5D" pageId="8" pageNumber="407" type="proposed english vernacular name">
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<paragraph id="68D0C4D8E8E756FE6983243582A3DFF5" pageId="8" pageNumber="407">Proposed English vernacular name.</paragraph>
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<paragraph id="0D1E4CA9A41EDC5DFC45FE2A54385F82" pageId="8" pageNumber="407">Amazon Fast-walking Beetle.</paragraph>
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</subSubSection>
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<subSubSection id="057F1F8F28F1FE0572389B4F69F62EF8" lastPageId="9" lastPageNumber="408" pageId="8" pageNumber="407" type="diagnosis">
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<paragraph id="4883C0321BEF1443057DACF3152866F8" pageId="8" pageNumber="407">Diagnosis.</paragraph>
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<paragraph id="2F89731703567D691AC32A6BB9848219" lastPageId="9" lastPageNumber="408" pageId="8" pageNumber="407">
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With the attributes of the subgenus
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<taxonomicName id="1704FA604D63FA19AB406AE5FCAA0B61" authorityName="Clairville" authorityYear="1806" class="Insecta" family="Carabidae" genus="Badister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="8" pageNumber="407" phylum="Arthropoda" rank="genus">
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<emphasis id="8396B08F32B60BA64B80FDF706A981D0" italics="true" pageId="8" pageNumber="407">Badister</emphasis>
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</taxonomicName>
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, as described by
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<bibRefCitation id="CFD170B3274C07B02D3BC131948DE87E" author="Ball, GE" pageId="12" pageNumber="411" refId="B1" refString="Ball, GE, 1959. A taxonomic study of the North American Licinini with notes on the Old World species of the Genus Diplocheila Brulle (Coleoptera). Memoirs of the American Entomological Society, 16, iv + 1-258." title="A taxonomic study of the North American Licinini with notes on the Old World species of the Genus Diplocheila Brulle (Coleoptera). Memoirs of the American Entomological Society, 16, iv + 1 - 258." year="1959">Ball (1959)</bibRefCitation>
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and as noted above, and large-sized for the genus (mean SBL more than 6.1 mm for both males and females of
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<taxonomicName id="C151A5586F2B7315835DBBFDA33D7C18" authorityName="Erwin & Ball" authorityYear="2011" class="Insecta" family="Carabidae" genus="Badister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="8" pageNumber="407" phylum="Arthropoda" rank="species" species="amazonus">
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<emphasis id="72B951E9DD2F20062A9A4A6ADB30DCE3" italics="true" pageId="8" pageNumber="407">Badister amazonus</emphasis>
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</taxonomicName>
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; mean SBL less than 5.9 mm for males and females of all other Western Hemisphere species samples). Adults with black head and elytra, the elytron with flavotestaceous base, sutural interneur, lateral margin, and epipleuron, and shiny throughout; prothorax and venter flavotestaceous; appendages testaceous. Microsculpture mesh pattern of head and pronotum isodiametric, sur
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<pageBreakToken id="3F18CA33887DEA76F6F41C3C39CF28B4" pageId="9" pageNumber="408" start="start">face</pageBreakToken>
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luster somewhat dull; of elytron, very finely etched transverse lines, surface shiny iridescent. Labial palpomere 3 (
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<figureCitation id="AE847B81DC31BEA195D2018C65DEC82E" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Badister amazonus sp. n., dorsal aspect, female, ADP 051544; type locality. Head, including eyes, antennomeres 1 - 3, mouthparts, and anterior part of pronotum. Legend: a 3, antennomere 3; cl, clypeus; lb, labrum; lp 3, labial palpomere 3; mb, mandibular boss; mn, mandibular notch; mp 4, maxillary palpomere 4." httpUri="https://binary.pensoft.net/fig/19731" pageId="9" pageNumber="408">Fig. 2</figureCitation>
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,
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<emphasis id="F99BCB461CA2E58214EEC8588ECC7E2A" bold="true" pageId="9" pageNumber="408">lp3</emphasis>
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) swollen and triangulate (distal margin obliquely truncate) with a very small pointed knob at apex. Pronotum with lateral margin moderately explanate in anterior half, broadly so posteriorly. Male tarsomeres 1-3 (
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<figureCitation id="020E04B82995B78A9171906FD59D019C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Badister amazonus sp. n., dorsal aspect, female, ADP 051544; type locality. A. Scutellar region of elytra: (pss) parascutellar stria; B-C. Foreleg tarsomeres of male (ADP 051713); B. Dorsal aspect, C. ventral aspect; ats, articulo-seta; ta 1, ta 2, ta 3, tarsomeres 1 - 3." httpUri="https://binary.pensoft.net/fig/19732" pageId="9" pageNumber="408">Fig. 3B, 3C</figureCitation>
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) slightly widened, with rows of adhesive vestiture on ventral surface narrow, confined to anterior half).
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</paragraph>
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</subSubSection>
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<subSubSection id="9A364F05FF79EC5E4F4627CF86A8CA5A" lastPageId="11" lastPageNumber="410" pageId="9" pageNumber="408" type="description">
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<paragraph id="1663D16E8C106B8AFC2E19D707D16709" pageId="9" pageNumber="408">Description.</paragraph>
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<paragraph id="7D9C994730C7A3559D245DFBABCA4F0D" pageId="9" pageNumber="408">
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(
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<figureCitation id="971E9CEA19B382E6132287FE683826AB" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Badister amazonus sp. n., female, ADP 051544; type locality. Habitus, dorsal aspect. ABL = 7.13 mm." httpUri="https://binary.pensoft.net/fig/19730" pageId="9" pageNumber="408">Fig. 1</figureCitation>
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,
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<figureCitation id="3B2D3FCD4BDB31C85B7E5CFEA9292B38" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Badister amazonus sp. n., dorsal aspect, female, ADP 051544; type locality. Head, including eyes, antennomeres 1 - 3, mouthparts, and anterior part of pronotum. Legend: a 3, antennomere 3; cl, clypeus; lb, labrum; lp 3, labial palpomere 3; mb, mandibular boss; mn, mandibular notch; mp 4, maxillary palpomere 4." httpUri="https://binary.pensoft.net/fig/19731" pageId="9" pageNumber="408">2</figureCitation>
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,
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<figureCitation id="9CAF0FA980AC892B60BDDF26E879E307" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Badister amazonus sp. n., dorsal aspect, female, ADP 051544; type locality. A. Scutellar region of elytra: (pss) parascutellar stria; B-C. Foreleg tarsomeres of male (ADP 051713); B. Dorsal aspect, C. ventral aspect; ats, articulo-seta; ta 1, ta 2, ta 3, tarsomeres 1 - 3." httpUri="https://binary.pensoft.net/fig/19732" pageId="9" pageNumber="408">
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3
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<normalizedToken id="7084049D1B6A51E479746B151FA9453F" originalValue="A–C">A-C</normalizedToken>
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</figureCitation>
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,
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<figureCitation id="E5AF2355079DC97CCA7106B195484C82" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Badister amazonus sp. n., dorsal aspect, ADP 051543; type locality. Pygidial (defense) gland system, Legend: cc, collecting canal; ed, efferent duct; edbl, efferent duct, basal lobe; gldr, pygidial gland reservoir. Ovipositor sclerites, Legend: gc 1, gonocoxite 1; gc 2, gonocoxite 2; lt, laterotergite" httpUri="https://binary.pensoft.net/fig/19733" pageId="9" pageNumber="408">4</figureCitation>
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,
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<figureCitation id="E43C01553D11B430DE905E1210AA64E8" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Badister amazonus sp. n., ADP 127149; type locality. Male genitalia, median lobe, and parameres, A - left lateral, B - dorsal, and C- right lateral aspects. Legend: a, apical area; bl, basal lobe; bo, basal orifice; lp, left paramere; ml, median lobe; om, ostial membrane; oo, ostial opening; par, preapical ridge; rp, right paramere; sh, shaft; ss, dorsal sclerotized strip." httpUri="https://binary.pensoft.net/fig/19734" pageId="9" pageNumber="408">
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5
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<normalizedToken id="989B13D828D54156F82195E37EAB5E25" originalValue="A–C">A-C</normalizedToken>
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</figureCitation>
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,
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<figureCitation id="5509DA527EAF7A351863D1609389C18B" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Badister amazonus sp. n. ADP 127151; type locality. Female genital tract. A. Ventral aspect, B. Dorsal aspect: Legend, b, base of gonocoxite 2; bl, blade of gonocoxite 2; des, dorsal ensiform seta; gc 1, gonocoxite 1; gc 2, gonocoxite 2; lt, laterotergite; ves, ventral ensiform setae. Ovipositor. C. Ventral aspect, D. Dorsal aspect: Legend, bc, bursa copulatrix; co, common oviduct; sg, spermathecal gland; sgd, spermathecal gland duct; sp, spermatheca. dorsal aspect; vc, villous canal." httpUri="https://binary.pensoft.net/fig/19735" pageId="9" pageNumber="408">
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6
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<normalizedToken id="A3C93960DD04306279F0933443D82BBC" originalValue="A–D">A-D</normalizedToken>
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</figureCitation>
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,
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<figureCitation id="B5BA8AC71DB957BEEC6B3B93879DF52E" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Distribution dot map for known localities of Badister amazonus sp. n .. along the rivers Samiria and Ucayali (★)." httpUri="https://binary.pensoft.net/fig/19736" pageId="9" pageNumber="408">7</figureCitation>
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).
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<emphasis id="C25081A046D1A5FE4B74A96DEAA94132" italics="true" pageId="9" pageNumber="408">Size</emphasis>
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: Moderately large for the subgenus; ABL = 6.2-7.9mm, SBL = 5.36-6.73mm, EW (maximum width) 2.55-3.43mm, LP = 0.97-1.22mm, WP = 1.71-1.91 mm, LE = 3.86-4.84mm.
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<emphasis id="E89ADFD37025EBA2D239A42F40739683" italics="true" pageId="9" pageNumber="408">Color</emphasis>
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: See Diagnosis above.
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<emphasis id="CB9DE32F6008FD2216483884ED3CB681" italics="true" pageId="9" pageNumber="408">Luster</emphasis>
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: See Diagnosis above.
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<emphasis id="684BF4EB47668E372C0DD83B8FA95843" bold="true" pageId="9" pageNumber="408">Head</emphasis>
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(
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<figureCitation id="8C0C82A5BC93AE01552FA7BE165C92FD" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Badister amazonus sp. n., dorsal aspect, female, ADP 051544; type locality. Head, including eyes, antennomeres 1 - 3, mouthparts, and anterior part of pronotum. Legend: a 3, antennomere 3; cl, clypeus; lb, labrum; lp 3, labial palpomere 3; mb, mandibular boss; mn, mandibular notch; mp 4, maxillary palpomere 4." httpUri="https://binary.pensoft.net/fig/19731" pageId="9" pageNumber="408">Fig. 2</figureCitation>
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): clypeus (
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<emphasis id="14EA1EF09C6887BEC1AA4257DAB67267" bold="true" pageId="9" pageNumber="408">cl</emphasis>
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) medially depressed, unisetose laterally, no clear demarcation from frons. Frons markedly depressed, apically with depression extended to eye carina, posteriorly somewhat concave with slight swellings medially raised to occiput; occiput shallowly domed; neck broad. Eye moderately convex; gena short and flat. Antennae and mouthparts typical for genus (see above for details).
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</paragraph>
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<paragraph id="7C695CBE82B43B05C8BAF3B76E2B00E1" pageId="9" pageNumber="408">
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<emphasis id="75297E21448587728E836751AF447372" bold="true" pageId="9" pageNumber="408">Prothorax. Pronotum</emphasis>
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(
|
||
<figureCitation id="FB6FA16BDF6F197A0EA6BC7C9C2F0525" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Badister amazonus sp. n., female, ADP 051544; type locality. Habitus, dorsal aspect. ABL = 7.13 mm." httpUri="https://binary.pensoft.net/fig/19730" pageId="9" pageNumber="408">Fig. 1</figureCitation>
|
||
,
|
||
<figureCitation id="EE68452EFB709362FF23D0694EF0096A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Badister amazonus sp. n., dorsal aspect, female, ADP 051544; type locality. Head, including eyes, antennomeres 1 - 3, mouthparts, and anterior part of pronotum. Legend: a 3, antennomere 3; cl, clypeus; lb, labrum; lp 3, labial palpomere 3; mb, mandibular boss; mn, mandibular notch; mp 4, maxillary palpomere 4." httpUri="https://binary.pensoft.net/fig/19731" pageId="9" pageNumber="408">2</figureCitation>
|
||
) markedly broad, about twice as long as head (mean LP/LH: 1.683 for males, 1.846 for females), cordiform, margin narrowly explanate with seta at anterior third; base deeply and bilaterally depressed; hind angle slightly obtusely produced and setose; moderately broader than long (mean W/L: 1.71 for males, 1.91 for females); surface medially smooth, slightly rugose laterally medial to explanation.
|
||
</paragraph>
|
||
<paragraph id="EC1A423F9481EA83BFB7F5B31F2CFFEB" pageId="10" pageNumber="409">
|
||
<emphasis id="78062D70FD76E40FA91850FAFCAD2ECC" bold="true" pageId="10" pageNumber="409">
|
||
<pageBreakToken id="6FFC7077EFAAFA655DA7DDAF4BD31D16" pageId="10" pageNumber="409" start="start">Pterothorax</pageBreakToken>
|
||
.
|
||
</emphasis>
|
||
Normal for genus.
|
||
</paragraph>
|
||
<paragraph id="0A87D95F90E71C508BDCD1F4DBDF4E38" pageId="10" pageNumber="409">
|
||
<emphasis id="1F514D2BD8AAA65DB85BB1C09C6F74AC" bold="true" pageId="10" pageNumber="409">Elytra</emphasis>
|
||
. Elytron about same width as head across eyes (mean WH/WE: 0.998 for both sexes), moderately convex, intervals moderately convex and slightly more so laterally, interval 3 bisetose with each setigerous puncture adherent to interneur 2.
|
||
</paragraph>
|
||
<paragraph id="8F732D661CD058A7BBEA89713A64714F" pageId="10" pageNumber="409">
|
||
<emphasis id="BAD8EAE470A94CC6B338655BDF4F05F2" bold="true" pageId="10" pageNumber="409">Hind wings</emphasis>
|
||
. Macropterous.
|
||
</paragraph>
|
||
<paragraph id="17D323A9C1B60E932E8E93531E1C9894" pageId="10" pageNumber="409">
|
||
<emphasis id="D03D68A64FD319A4589236595F8FF058" bold="true" pageId="10" pageNumber="409">Legs.</emphasis>
|
||
(
|
||
<figureCitation id="497B72C030928526D7DAD494723783F7" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Badister amazonus sp. n., female, ADP 051544; type locality. Habitus, dorsal aspect. ABL = 7.13 mm." httpUri="https://binary.pensoft.net/fig/19730" pageId="10" pageNumber="409">Fig. 1</figureCitation>
|
||
). Overall, normal for subgenus. Male front tarsus (
|
||
<figureCitation id="DFDB0C374EA6DF1810F4D44A1185234B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Badister amazonus sp. n., dorsal aspect, female, ADP 051544; type locality. A. Scutellar region of elytra: (pss) parascutellar stria; B-C. Foreleg tarsomeres of male (ADP 051713); B. Dorsal aspect, C. ventral aspect; ats, articulo-seta; ta 1, ta 2, ta 3, tarsomeres 1 - 3." httpUri="https://binary.pensoft.net/fig/19732" pageId="10" pageNumber="409">Fig. 3B, 3C</figureCitation>
|
||
) with tarsomeres 1-3 slightly dilated and each ventrally (
|
||
<figureCitation id="BE2E377D07DCBCC5A5D167FF98C87FB3" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Badister amazonus sp. n., dorsal aspect, female, ADP 051544; type locality. A. Scutellar region of elytra: (pss) parascutellar stria; B-C. Foreleg tarsomeres of male (ADP 051713); B. Dorsal aspect, C. ventral aspect; ats, articulo-seta; ta 1, ta 2, ta 3, tarsomeres 1 - 3." httpUri="https://binary.pensoft.net/fig/19732" pageId="10" pageNumber="409">Fig. 3C</figureCitation>
|
||
) with 2-4 rows of white articulo-setae.
|
||
</paragraph>
|
||
<paragraph id="DD1384457BA943610C0D0F46466BBC5B" pageId="10" pageNumber="409">
|
||
<emphasis id="92006169739860CD76294F72F9D59BB0" bold="true" pageId="10" pageNumber="409">Abdominal sterna</emphasis>
|
||
with normal setation for genus.
|
||
</paragraph>
|
||
<paragraph id="1FF3A1B0AFB9BE2A930B2339C215EFD3" pageId="10" pageNumber="409">
|
||
<emphasis id="3DDCD07278AAAE906254125386310814" bold="true" pageId="10" pageNumber="409">Male genitalia</emphasis>
|
||
(
|
||
<figureCitation id="1BEADD4274F6F4A7A267304D86BAB8CC" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Badister amazonus sp. n., ADP 127149; type locality. Male genitalia, median lobe, and parameres, A - left lateral, B - dorsal, and C- right lateral aspects. Legend: a, apical area; bl, basal lobe; bo, basal orifice; lp, left paramere; ml, median lobe; om, ostial membrane; oo, ostial opening; par, preapical ridge; rp, right paramere; sh, shaft; ss, dorsal sclerotized strip." httpUri="https://binary.pensoft.net/fig/19734" pageId="10" pageNumber="409">
|
||
Fig. 5
|
||
<normalizedToken id="37B8F7F76626527FEFB7DDF210A442E7" originalValue="A–">A-</normalizedToken>
|
||
5C
|
||
</figureCitation>
|
||
). Median lobe (
|
||
<emphasis id="12745DFFD25452567C30817E6BB071C8" bold="true" pageId="10" pageNumber="409">ml</emphasis>
|
||
) with basal lobe (
|
||
<emphasis id="04A3118E70F90F7B2601A5D146A0981D" bold="true" pageId="10" pageNumber="409">bl</emphasis>
|
||
) about half length of shaft (
|
||
<emphasis id="9156646ED86DD1E334FC806DF9B74894" bold="true" pageId="10" pageNumber="409">sh</emphasis>
|
||
), basal opening (
|
||
<emphasis id="6579116A9B416A42BD515E73F793F58D" bold="true" pageId="10" pageNumber="409">bo</emphasis>
|
||
) large. Shaft slender, curved ventrally, dorsally membranous (
|
||
<emphasis id="934D5DB297E0DCBBB0C3B3CC09600340" bold="true" pageId="10" pageNumber="409">om</emphasis>
|
||
) except for two long sclerotized strips (
|
||
<emphasis id="7A105632B624F59FF5932DE4DA82EA07" bold="true" pageId="10" pageNumber="409">ss</emphasis>
|
||
) extended from basal lobe to ostial opening (
|
||
<emphasis id="F0E4AA195A8C7B2D0B3C3670F635EAB7" bold="true" pageId="10" pageNumber="409">oo</emphasis>
|
||
); in ventral aspect tapered toward rather broadly rounded apex, preapically with prominent preapical ridge (
|
||
<emphasis id="36F77CDC4924955F95B862AD38EF2D1C" bold="true" pageId="10" pageNumber="409">par</emphasis>
|
||
), in lateral aspect, ridge a prominently projected point. Parameres (
|
||
<emphasis id="A22BD0331B765524D31DB5AC7475352D" bold="true" pageId="10" pageNumber="409">lp, rp</emphasis>
|
||
)broad, apices subtruncate left paramere
|
||
<emphasis id="24E2F4D2BE2F5C89D2D80E982ADDF78A" bold="true" pageId="10" pageNumber="409">(lp</emphasis>
|
||
) longer than right paramere (
|
||
<emphasis id="8AA1C6DFA44289B540E19611D1F97A4D" bold="true" pageId="10" pageNumber="409">rp</emphasis>
|
||
) about three quarters length of shaft (measured in left lateral aspect). Internal sac with apical ring of rather densely distributed microtrichia, without preapical spines.
|
||
</paragraph>
|
||
<paragraph id="00792E385368ACDDE2D0BBC694AE758A" pageId="10" pageNumber="409">
|
||
<emphasis id="66631A785114D07AEE06018C6EA6C967" bold="true" pageId="10" pageNumber="409">Female genitalia.</emphasis>
|
||
(
|
||
<figureCitation id="4A8A671FAD459F9E3854ED04E243FA39" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Badister amazonus sp. n. ADP 127151; type locality. Female genital tract. A. Ventral aspect, B. Dorsal aspect: Legend, b, base of gonocoxite 2; bl, blade of gonocoxite 2; des, dorsal ensiform seta; gc 1, gonocoxite 1; gc 2, gonocoxite 2; lt, laterotergite; ves, ventral ensiform setae. Ovipositor. C. Ventral aspect, D. Dorsal aspect: Legend, bc, bursa copulatrix; co, common oviduct; sg, spermathecal gland; sgd, spermathecal gland duct; sp, spermatheca. dorsal aspect; vc, villous canal." httpUri="https://binary.pensoft.net/fig/19735" pageId="10" pageNumber="409">
|
||
Fig. 6
|
||
<normalizedToken id="6A34AB4FDF6BEAE31C8D2817BB6444EF" originalValue="A–D">A-D</normalizedToken>
|
||
</figureCitation>
|
||
). Ovipositor with broad laterotergite (
|
||
<emphasis id="8D7FFE2FB7B4DDAA11BBFFEF7978EAA5" bold="true" pageId="10" pageNumber="409">lt</emphasis>
|
||
) and two gonocoxites (
|
||
<emphasis id="D934BD3059CE2A0B5AC3919DE32243F6" bold="true" pageId="10" pageNumber="409">gc 1, gc 2</emphasis>
|
||
); gonocoxite 1 asetose; gonocoxite 2 falcate, base (
|
||
<emphasis id="82899A7DAF1141F98804FAD83F189440" bold="true" pageId="10" pageNumber="409">b</emphasis>
|
||
) large, broad, blade (
|
||
<emphasis id="711DDD2D4B9A16E3BDD7C19D2EC8A4B4" bold="true" pageId="10" pageNumber="409">bl</emphasis>
|
||
) rather short, with two dorsal ensiform setae (
|
||
<emphasis id="815629C274103F39284E4DFE97109CBE" bold="true" pageId="10" pageNumber="409">des</emphasis>
|
||
), and one ventral ensiform seta (
|
||
<emphasis id="C99618F5BFF3AD993F21AF89C3FE4D3D" bold="true" pageId="10" pageNumber="409">ves</emphasis>
|
||
), all ensiform setae short; without ventral preapical nematiform setae. Reproductive tract (
|
||
<figureCitation id="82FAB405E8B84AC1307FC4E13C8C7330" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Badister amazonus sp. n. ADP 127151; type locality. Female genital tract. A. Ventral aspect, B. Dorsal aspect: Legend, b, base of gonocoxite 2; bl, blade of gonocoxite 2; des, dorsal ensiform seta; gc 1, gonocoxite 1; gc 2, gonocoxite 2; lt, laterotergite; ves, ventral ensiform setae. Ovipositor. C. Ventral aspect, D. Dorsal aspect: Legend, bc, bursa copulatrix; co, common oviduct; sg, spermathecal gland; sgd, spermathecal gland duct; sp, spermatheca. dorsal aspect; vc, villous canal." httpUri="https://binary.pensoft.net/fig/19735" pageId="10" pageNumber="409">Fig. 6A, 6B</figureCitation>
|
||
) proximally with short, broad bursa copulatrix (
|
||
<emphasis id="5527606CB94486CF60E7E95F99777D97" bold="true" pageId="10" pageNumber="409">bc</emphasis>
|
||
), continuous at its distal end with common oviduct (
|
||
<emphasis id="6BEB7AD82AA6369AC97226E31FB86677" bold="true" pageId="10" pageNumber="409">co</emphasis>
|
||
) and long spermatheca (
|
||
<emphasis id="0CBB1FCFBC1F1A4A0142CC3BB4597B2C" bold="true" pageId="10" pageNumber="409">sp</emphasis>
|
||
), latter coiled distally; villous canal (
|
||
<emphasis id="5966665EFDE23A6AE9EE32BBFCA2848A" bold="true" pageId="10" pageNumber="409">vc</emphasis>
|
||
) extended from near base of spermatheca well up common oviduct; spermathecal gland (
|
||
<emphasis id="8C719FB01716E11B4A2A6FE722CC9B70" bold="true" pageId="10" pageNumber="409">sg</emphasis>
|
||
) bulbous; spermathecal gland duct (
|
||
<emphasis id="42FC047BC8FF55271D07AA59C4C31AC4" bold="true" pageId="10" pageNumber="409">sgd</emphasis>
|
||
) long, slender, attached to spermatheca at base of its coiled portion.
|
||
</paragraph>
|
||
<paragraph id="6BCA9A89154FCF902604641DCFB850D7" pageId="10" pageNumber="409">
|
||
Markedly similar to homologous structures in
|
||
<taxonomicName id="85066A493BFA673022DD429374793108" authorityName="LeConte" authorityYear="1880" class="Insecta" family="Carabidae" genus="Badister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="409" phylum="Arthropoda" rank="species" species="reflexus" subGenus="Baudia">
|
||
<emphasis id="655AEB211F91C092F0FF5020A80D202F" italics="true" pageId="10" pageNumber="409">Badister (Baudia) reflexus</emphasis>
|
||
</taxonomicName>
|
||
LeConte (
|
||
<bibRefCitation id="2E0F2BCE2FF6647D75C386BBB37FD36D" author="Will, KW" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="13" pageNumber="412" pagination="95 - 103" refId="B28" refString="Will, KW, 1998. A new species of Diplocheila Brulle from North America, with notes on female reproductive tract characters in selected Licinini and implications for evolution of the subgenus Isorembus Jeannel (Coleoptera: Carabidae: Licinini). Proceedings of the Entomological Society of Washington 100: 95 - 103" title="A new species of Diplocheila Brulle from North America, with notes on female reproductive tract characters in selected Licinini and implications for evolution of the subgenus Isorembus Jeannel (Coleoptera: Carabidae: Licinini)." volume="100" year="1998">Will 1998</bibRefCitation>
|
||
: 99, Fig. 8), which differ as follows: spermatheca with basal uncoiled portion more extensive; distal coiled portion short; spermathecal gland elongate digitiform.
|
||
</paragraph>
|
||
<paragraph id="9547FFDFE610945AFB69D6155EE43936" pageId="10" pageNumber="409">
|
||
<emphasis id="89CADDF4E987CF7360C50353F3D0113E" bold="true" pageId="10" pageNumber="409">Classification.</emphasis>
|
||
<bibRefCitation id="ED40DE919ED3C67B2F578453A4374E22" author="Ball, GE" pageId="12" pageNumber="411" refId="B1" refString="Ball, GE, 1959. A taxonomic study of the North American Licinini with notes on the Old World species of the Genus Diplocheila Brulle (Coleoptera). Memoirs of the American Entomological Society, 16, iv + 1-258." title="A taxonomic study of the North American Licinini with notes on the Old World species of the Genus Diplocheila Brulle (Coleoptera). Memoirs of the American Entomological Society, 16, iv + 1 - 258." year="1959">Ball (1959</bibRefCitation>
|
||
: 194-195) arranged the species of subgenus
|
||
<taxonomicName id="B3610636B02A2B5883E6A4306323DD05" authorityName="Clairville" authorityYear="1806" class="Insecta" family="Carabidae" genus="Badister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="409" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="049733198FE5483069010699C45F3876" italics="true" pageId="10" pageNumber="409">Badister</emphasis>
|
||
</taxonomicName>
|
||
in two
|
||
<normalizedToken id="75636B8E7FD25B988CC51993FFA233F7" originalValue="“complexes”">"complexes"</normalizedToken>
|
||
characterized principally by features of the male genitalia and ovipositor gonocoxite 2. The combination of these features in
|
||
<taxonomicName id="2A82415B4E426683F8BB3DDEB5C9EF20" authorityName="Erwin & Ball" authorityYear="2011" class="Insecta" family="Carabidae" genus="Badister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="409" phylum="Arthropoda" rank="species" species="amazonus">
|
||
<emphasis id="CF9B6D5070E52F7333DD47FF3F7790C6" italics="true" pageId="10" pageNumber="409">Badister amazonus</emphasis>
|
||
</taxonomicName>
|
||
fits neither complex. Accordingly, we propose here the monobasic
|
||
<emphasis id="C2CE417241982C9D442C69B5B4D710E9" italics="true" pageId="10" pageNumber="409">amazonus</emphasis>
|
||
complex, characterized as follows: median lobe of male genitalia dorsally with two long sclerotized strips, internal sac without sclerotized plates or spines; gonocoxite 2 of ovipositor relatively broad basally, ensiform setae relatively short; nematiform setae absent. Additional diagnostic features are: male fore tarsomeres 1-3 (
|
||
<figureCitation id="82EF0EE0947FE85E466D31612DA15B2B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Badister amazonus sp. n., dorsal aspect, female, ADP 051544; type locality. A. Scutellar region of elytra: (pss) parascutellar stria; B-C. Foreleg tarsomeres of male (ADP 051713); B. Dorsal aspect, C. ventral aspect; ats, articulo-seta; ta 1, ta 2, ta 3, tarsomeres 1 - 3." httpUri="https://binary.pensoft.net/fig/19732" pageId="10" pageNumber="409">Fig. 3B</figureCitation>
|
||
) relatively slender, not much wider than tarsomeres 1-3 of middle and hind tarsi, fore tarsomere 1 longer than wide, ventral adhesive setae (
|
||
<figureCitation id="152C143CA7BF43B72299831D5853817B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Badister amazonus sp. n., dorsal aspect, female, ADP 051544; type locality. A. Scutellar region of elytra: (pss) parascutellar stria; B-C. Foreleg tarsomeres of male (ADP 051713); B. Dorsal aspect, C. ventral aspect; ats, articulo-seta; ta 1, ta 2, ta 3, tarsomeres 1 - 3." httpUri="https://binary.pensoft.net/fig/19732" pageId="10" pageNumber="409">Fig. 3C</figureCitation>
|
||
) concentrated in anterior half of each tarsomere.
|
||
</paragraph>
|
||
<paragraph id="EE78F0E0889D124474A638037FCAF34F" pageId="10" pageNumber="409">
|
||
<emphasis id="13ECEE9A548C013CF3A653C813F5496B" bold="true" pageId="10" pageNumber="409">Dispersal potential</emphasis>
|
||
. These beetles are fully macropterous and are probably capable of flight; they are fast walkers. They also are climbers on grasses (
|
||
<taxonomicName id="C5940722F058B196A804C04E19074A09" class="Liliopsida" family="Poaceae" genus="Paspalum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="10" pageNumber="409" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="BAE5F5B82DF53C85F33103146A909035" italics="true" pageId="10" pageNumber="409">Paspalum</emphasis>
|
||
</taxonomicName>
|
||
spp.).
|
||
</paragraph>
|
||
<paragraph id="151B61399CA44D6D7C8A409D69181333" lastPageId="11" lastPageNumber="410" pageId="10" pageNumber="409">
|
||
<emphasis id="D31D4B7099F94010475286B639AB4442" bold="true" pageId="10" pageNumber="409">Way of life</emphasis>
|
||
. Adults are associated with water: in open grassy marshes, on beaches of black and white water rivers, and in forest swamps at the edges of standing black
|
||
<pageBreakToken id="9DB02B4782743A57578A2DBF08745ACE" pageId="11" pageNumber="410" start="start">water</pageBreakToken>
|
||
ponds in leaf litter on grey clay soil (Kaolin). They are active at night and take cover in the day among grass stems and under flood debris. They are active in May and
|
||
<normalizedToken id="C131F204E81C8B1C2E8B1743A84AA20B" originalValue="August–September">August-September</normalizedToken>
|
||
; no teneral adults were encountered in these months.
|
||
</paragraph>
|
||
<paragraph id="64AE3CE15462C370B3AA3CBC3C47016D" pageId="11" pageNumber="410">
|
||
<emphasis id="D13BFC8EE4BF5CF08CECB48151B6CD34" bold="true" pageId="11" pageNumber="410">Other specimens examined</emphasis>
|
||
.
|
||
<emphasis id="C8C0052CB4ED67AA071294C2AD9D26F3" bold="true" pageId="11" pageNumber="410">
|
||
<normalizedToken id="8852B484D1479FFE33B8860C3B7A76C4" originalValue="Perú">Peru</normalizedToken>
|
||
,
|
||
</emphasis>
|
||
Loreto, SW bank of Rio Ucayali, 90m,
|
||
<geoCoordinate id="5358F7007A8DACC2F3E970EEFE6AC63D" degrees="4.2089" direction="south" orientation="latitude" precision="5" value="-4.2089">4.2089°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="32BF2DD3D1C41A45E93362421EEC1CD9" degrees="73.3657" direction="west" orientation="longitude" precision="5" value="-73.3657">73.3657°W</geoCoordinate>
|
||
, 2 September 1991, (T.L. Erwin)(NMNH: ADP051862, female); Pacaya-Samiria National Reserve, Rio Samiria, Cocha Shinguito, 90m,
|
||
<geoCoordinate id="794A9C11B3A9F34D55716B1506605486" degrees="5.1775" direction="south" orientation="latitude" precision="5" value="-5.1775">5.1775°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="600E914799BBA25BEBF44636EC424DD6" degrees="74.6556" direction="west" orientation="longitude" precision="5" value="-74.6556">74.6556°W</geoCoordinate>
|
||
, 21 May 1990 (T.L. Erwin)(NMNH: ADP093676, 128618, females); 1.0 km SW boca del Rio Samiria, Vigilante Post 1, 130m, "
|
||
<geoCoordinate id="99C0DF2288C57118E9A75A19DB8E4B05" degrees="04" direction="south" minutes="40.5" orientation="latitude" precision="92" value="-4.675">04°40.5'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="1C46DFE227B0F212E8734DB2570324C5" degrees="074" direction="west" minutes="18.9" orientation="longitude" precision="92" value="-74.315">074°18.9'W</geoCoordinate>
|
||
" 31 August 1991-1 November 1991 (G.E. Ball & D. Shpeley)(NMNH: ADP127151, female, 127149, male); 1.0 km SW boca del Rio Samiria, Vigilante Post 1, 130m, "
|
||
<geoCoordinate id="9F2D3CD9668097D19215CBE8E02582DA" degrees="04" direction="south" minutes="40.5" orientation="latitude" precision="92" value="-4.675">04°40.5'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="F421A20D2A8F763FF76A429675EAB58B" degrees="074" direction="west" minutes="18.9" orientation="longitude" precision="92" value="-74.315">074°18.9'W</geoCoordinate>
|
||
" 31 August 1991 (T.L. Erwin & M.G. Pogue)(NMNH: ADP051697, 051603, 051336, 051721, 051672, 051743, 051693, 051698, 051722, 051694, 051724, 051692, 051670, 051717, 051718, 051723, 050590, 050589, 050605, 051696, 051720 females and ADP051824, 051630, 051719, 051713, 051826, 051691, 051695 males); 1.0 km SW boca del Rio Samiria, Vigilante Post 1, 130m, "
|
||
<geoCoordinate id="9E4CCBE0B4F46B5D912EAA2260200CE6" degrees="04" direction="south" minutes="40.5" orientation="latitude" precision="92" value="-4.675">04°40.5'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="4DB49BD5B8252C572F2FA2ADDC83CADA" degrees="074" direction="west" minutes="18.9" orientation="longitude" precision="92" value="-74.315">074°18.9'W</geoCoordinate>
|
||
" 31 August 1991 (T.L. Erwin)(NMNH: ADP051592, 051563, 051544, 051583, 051589, 051559, 051590, 051564, 051567, 051569, 051561, 051581, 051585, 051570, 051584, 051565, 051543 females and ADP051562, 051541, 051591, 051542, 051566, 051588, 051560, 051568 males).
|
||
</paragraph>
|
||
<paragraph id="3D1BB027F0012D87238D26E21CC0C20B" pageId="11" pageNumber="410">
|
||
Two males and 12 females at UASM with the following label data: /PERU Dpto. Loreto/ 1 km SW Boca del Rio/Samiria
|
||
<geoCoordinate id="2387216551C5F3E75AF773CF828C0D22" degrees="04" direction="south" minutes="40" orientation="latitude" precision="15" seconds="29" value="-4.674722">04°40'29"S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="E45AC3E75DAECF07F2E2BBEA72F04C0A" degrees="74" direction="west" minutes="18" orientation="longitude" precision="15" seconds="55" value="-74.31528">74°18'55"W</geoCoordinate>
|
||
130m/marsh, treading/31.VIII.& 1.IX.91 30-91//T. L. ERWIN EXP./Res. Pacaya-Samiria/G.E. Ball & D. Shpeley/collectors 1991//.
|
||
</paragraph>
|
||
<paragraph id="9274B1852FDCA6DB4619B04E49F8B48C" pageId="11" pageNumber="410">
|
||
<emphasis id="1500093A01C8ADB8AADE20A2C1B88873" bold="true" pageId="11" pageNumber="410">Geographic distribution</emphasis>
|
||
. (
|
||
<figureCitation id="43EC49F44650090520DF7F726090EB25" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Distribution dot map for known localities of Badister amazonus sp. n .. along the rivers Samiria and Ucayali (★)." httpUri="https://binary.pensoft.net/fig/19736" pageId="11" pageNumber="410">Fig. 7</figureCitation>
|
||
). This species is currently known from only three localities: two along the
|
||
<normalizedToken id="85E91F3C78F8920F12F9475853C7C0D6" originalValue="Río">Rio</normalizedToken>
|
||
Samiria, and one along the
|
||
<normalizedToken id="E52B7B5E65B9C1663FCF9389AE9E84C3" originalValue="Río">Rio</normalizedToken>
|
||
Ucayali, Loreto Department,
|
||
<normalizedToken id="C781C527AD4E893AB8B652BB0BFDBABD" originalValue="Perú">Peru</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="093E61360A6C0DB9435CAFD10829AFEA" pageId="11" pageNumber="410">
|
||
<emphasis id="D6E30E5B474301600D94FBF98D316C1B" bold="true" pageId="11" pageNumber="410">Notes</emphasis>
|
||
. We are not presenting an analysis of variation in body proportions because this species can be separated from all other members of the genus without resorting to such attributes. In regard to the location (latitude/longitude) above for "1.0 km SW del boca Rio Samiria," Google Earth allows us now to refine what is given on the labels. The actual site is
|
||
<geoCoordinate id="819F3F883D5B57CD1E936D9BFAB397F4" degrees="4.686779" direction="south" orientation="latitude" precision="1" value="-4.686779">4.686779°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="83EE2B052FEF19BEC58C4DFEB503F6B0" degrees="74.336711" direction="west" orientation="longitude" precision="1" value="-74.33671">74.336711°W</geoCoordinate>
|
||
at an elevation of 101m. Also, the female specimen from the
|
||
<normalizedToken id="1569617B5083464B59997E727DF49315" originalValue="Río">Rio</normalizedToken>
|
||
Ucayali was mistakenly labeled
|
||
<normalizedToken id="E7C0D955DDD77B03B81528FE7FF504F9" originalValue="“Río">"Rio</normalizedToken>
|
||
Amazonas." The river changes names in the vicinity of Iquitos,
|
||
<normalizedToken id="69631CE8BCDCA98DC8B4D1C12671933B" originalValue="Perú">Peru</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |