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<document id="8744B86844C935900680CC789946DCB5" ID-DOI="10.5281/zenodo.203121" ID-GBIF-Dataset="cabb7848-5df6-4591-90e3-685a13024e8e" ID-ISSN="1175-5326" ID-Zenodo-Dep="203121" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460219715071" checkinUser="plazi" docAuthor="Hoffman, Richard L." docDate="2011" docId="58464001C22ED969FF3AFA40FA94FE8C" docLanguage="en" docName="zt02734p062.pdf" docOrigin="Zootaxa 2734" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Haplogonopus inflatannulus Verhoeff" docType="treatment" docVersion="10" lastPageNumber="55" masterDocId="A47F3879C22CD96DFFADFB32FF9CFFB3" masterDocTitle="A review of the milliped genus Haplogonopus with commentary on the so-called “ Charactopygus-Bildung ” modification of spirostreptid paraprocts (Diplopoda: Spirostreptidae)" masterLastPageNumber="62" masterPageNumber="53" pageNumber="55" updateTime="1698237495865" updateUser="plazi">
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<mods:title id="4BC95B6EEB8358D8EA1741067F6BCFA9">A review of the milliped genus Haplogonopus with commentary on the so-called “ Charactopygus-Bildung ” modification of spirostreptid paraprocts (Diplopoda: Spirostreptidae)</mods:title>
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<mods:namePart id="9A246EA0525FA47E9E2E627523EDE7B5">Hoffman, Richard L.</mods:namePart>
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<treatment id="58464001C22ED969FF3AFA40FA94FE8C" ID-DOI="http://doi.org/10.5281/zenodo.6184239" ID-GBIF-Taxon="119385052" ID-Zenodo-Dep="6184239" LSID="urn:lsid:plazi:treatment:58464001C22ED969FF3AFA40FA94FE8C" httpUri="http://treatment.plazi.org/id/58464001C22ED969FF3AFA40FA94FE8C" lastPageId="4" lastPageNumber="55" pageId="2" pageNumber="55">
<subSubSection id="98F5A29CC22ED96FFF3AFA40FE80FE1E" pageId="2" pageNumber="55" type="nomenclature">
<paragraph id="D050F117C22ED96FFF3AFA40FDF8FE3F" blockId="2.[151,612,370,429]" box="[151,612,370,396]" pageId="2" pageNumber="55">
<heading id="8B18467BC22ED96FFF3AFA40FDF8FE3F" bold="true" box="[151,612,370,396]" fontSize="11" level="1" pageId="2" pageNumber="55" reason="1">
<taxonomicName id="17EF8A94C22ED96FFF3AFA40FDF8FE3F" ID-CoL="3JMQV" authority="Verhoeff" authorityName="Verhoeff" box="[151,612,370,396]" class="Diplopoda" family="Spirostreptidae" genus="Haplogonopus" higherTaxonomySource="CoL" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="inflatannulus">
<emphasis id="E29B2D05C22ED96FFF3AFA40FDF8FE3F" bold="true" box="[151,612,370,396]" pageId="2" pageNumber="55">
<emphasis id="E29B2D05C22ED96FFF3AFA40FE68FE3F" bold="true" box="[151,500,370,396]" italics="true" pageId="2" pageNumber="55">Haplogonopus inflatannulus</emphasis>
Verhoeff
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="D050F117C22ED96FFF3AFAA7FE80FE1E" blockId="2.[151,612,370,429]" box="[151,284,405,429]" pageId="2" pageNumber="55">
<figureCitation id="48D4ED92C22ED96FFF3AFAA7FE80FE1E" box="[151,284,405,429]" captionStart-0="FIGURES 1 5" captionStart-1="FIGURES 6 9" captionStartId-0="3.[151,264,1922,1945]" captionStartId-1="5.[151,264,1899,1922]" captionTargetBox-0="[214,1374,207,1871]" captionTargetBox-1="[238,1352,202,1881]" captionTargetId-0="figure@3.[196,1395,195,1881]" captionTargetId-1="figure@5.[235,1365,193,1885]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionText-0="FIGURES 1 5. Haplogonopus inflatannulus Verhoeff., structural features. 1. Gnathochilarium and adjacent parts of mandibles and collum. 2. Right mandible, posterior aspect, with distal lobe of basomere indicated by the ↔. 3. First pair of legs of male, syncoxa, sternum, and basal podomeres, anterior aspect. 4. The same, posterior aspect, sclerite of uncertain identity indicated by the? 5. Dorsal half of three midbody segments, lateral aspect, showing convex metazona with ovoid depressions." captionText-1="FIGURES 6 9. Haplogonopus, gonopod structure of three species. 6. H. inflatannulus, right gonopod, anterior aspect. 7. H. jeanneli, left gonopod, anterior aspect (adapted from Brolemann, 1920). 8. H. cingulatus, right gonopod, anterior aspect. 9. Midlength section of posttorsal telopodite, enlarged, to show form of cingulum. Abbreviations: a, apicomedial process of proplica; atp, antetorsal process of telopodite, mp, distal end of metaplica." httpUri-0="https://zenodo.org/record/203122/files/figure.png" httpUri-1="https://zenodo.org/record/203123/files/figure.png" pageId="2" pageNumber="55">Figures 16</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="98F5A29CC22ED96FFF3AFAEFFAEEFD5B" pageId="2" pageNumber="55" type="reference_group">
<paragraph id="D050F117C22ED96FFF3AFAEFFCBCFD83" blockId="2.[151,1436,477,744]" pageId="2" pageNumber="55">
<taxonomicName id="17EF8A94C22ED96FFF3AFAEFFDC1FE41" authority="Verhoeff, 1941" authorityName="Verhoeff" authorityYear="1941" box="[151,605,477,499]" class="Diplopoda" family="Spirostreptidae" genus="Haplogonopus" higherTaxonomySource="CoL" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="inflatannulus">
<emphasis id="E29B2D05C22ED96FFF3AFAEFFE26FE40" box="[151,442,477,499]" italics="true" pageId="2" pageNumber="55">Haplogonopus inflatannulus</emphasis>
<bibRefCitation id="B47E8CE6C22ED96FFE6FFAEFFDC1FE41" author="Verhoeff" box="[450,605,477,499]" pageId="2" pageNumber="62" refString="Verhoeff, K. W. (1941) Studien an athiopische Diplopoden. Jenaische Zeitschrift fur Naturwissenschaften, 73, 231 - 274." type="journal article" year="1941">Verhoeff, 1941</bibRefCitation>
</taxonomicName>
, Jenaische Zeitschr. Naturw., 73: 265, figs. 4547. Male
<typeStatus id="0F544FB5C22ED96FFB01FAEFFA99FE40" box="[1196,1285,477,499]" pageId="2" pageNumber="55" type="holotype">holotype</typeStatus>
(Zoologisches Staatssammlung München) from “...dem Gebirge bei Morogoro am Tanganyikasee...” [=North Uluguru Mountains, Morogoro Region and District,
<collectingCountry id="A8F8B187C22ED96FFDAAF928FDFBFD83" box="[519,615,538,560]" name="Tanzania" pageId="2" pageNumber="55">Tanzania</collectingCountry>
], W. V. Harris leg.
</paragraph>
<paragraph id="D050F117C22ED96FFF3AF908FDEBFD3F" blockId="2.[151,1436,477,744]" pageId="2" pageNumber="55">
<taxonomicName id="17EF8A94C22ED96FFF3AF908FD87FDFC" ID-CoL="6ZCH7" authority="Kraus, 1958" authorityName="Kraus" authorityYear="1958" box="[151,539,569,591]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="nebularius">
<emphasis id="E29B2D05C22ED96FFF3AF908FE0DFDFC" box="[151,401,569,591]" italics="true" pageId="2" pageNumber="55">Spirostreptus nebularius</emphasis>
<bibRefCitation id="B47E8CE6C22ED96FFE36F908FD87FDFC" author="Kraus" box="[411,539,569,591]" pageId="2" pageNumber="62" refString="Kraus, O. (1958) Myriapoden aus Ostafrika (Tanganyika Territory). Ve r o ffentlichen der U berseemuseum in Bremen, Ser. A., 3, 1 - 16." type="journal article" year="1958">Kraus, 1958</bibRefCitation>
</taxonomicName>
, Veröff. Überseemus. Bremen, A, 3: 10, text fig. 2, pl. 3, figs. 2729. Male
<typeStatus id="0F544FB5C22ED96FFAEEF90BFA00FDFC" box="[1347,1436,569,591]" pageId="2" pageNumber="55" type="holotype">holotype</typeStatus>
(Überseemuseum Bremen) from Lukwangule Peak, Uluguru Mountains, Morogoro Region and District,
<collectingCountry id="A8F8B187C22ED96FFAB5F96AFAEAFDDD" box="[1304,1398,600,622]" name="Tanzania" pageId="2" pageNumber="55">Tanzania</collectingCountry>
, H. Knipper leg.
<date id="A451D7D7C22ED96FFEE1F945FE59FD3F" box="[332,453,630,652]" pageId="2" pageNumber="55" value="1952-07-02">2 July 1952</date>
.
<emphasis id="E29B2D05C22ED96FFE63F945FDEBFD3F" bold="true" box="[462,631,631,652]" pageId="2" pageNumber="55">New synonymy.</emphasis>
</paragraph>
<paragraph id="D050F117C22ED96FFF3AF9A4FC6DFD19" blockId="2.[151,1436,477,744]" box="[151,1009,661,683]" pageId="2" pageNumber="55">
<taxonomicName id="17EF8A94C22ED96FFF3AF9A4FDDFFD19" authority="Krabbe, 1982" authorityName="Krabbe" authorityYear="1982" box="[151,579,661,683]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="inflatannulus">
<emphasis id="E29B2D05C22ED96FFF3AF9A4FE2CFD18" box="[151,432,661,683]" italics="true" pageId="2" pageNumber="55">Spirostreptus inflatannulus:</emphasis>
<bibRefCitation id="B47E8CE6C22ED96FFE1AF9A7FDDFFD19" author="Krabbe" box="[439,579,661,683]" pageId="2" pageNumber="62" refString="Krabbe, E. (1982) Systematik der Spirostreptidae (Diplopoda; Spirostreptomorpha). Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg (Neue Folge,) 24, 1 - 476." type="journal article" year="1982">Krabbe, 1982</bibRefCitation>
</taxonomicName>
, Abh. Naturw. Ver. Hamburg, NF 24: 204.
</paragraph>
<paragraph id="D050F117C22ED96FFF3AF986FC4AFD7A" blockId="2.[151,1436,477,744]" box="[151,982,691,714]" pageId="2" pageNumber="55">
<taxonomicName id="17EF8A94C22ED96FFF3AF986FDB4FD7A" ID-CoL="6ZCH7" authority="Krabbe, 1982" authorityName="Krabbe" authorityYear="1982" box="[151,552,691,714]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="nebularius">
<emphasis id="E29B2D05C22ED96FFF3AF986FE12FD79" box="[151,398,692,714]" italics="true" pageId="2" pageNumber="55">Spirostreptus nebularius</emphasis>
:
<bibRefCitation id="B47E8CE6C22ED96FFE31F981FDB4FD7A" author="Krabbe" box="[412,552,691,713]" pageId="2" pageNumber="62" refString="Krabbe, E. (1982) Systematik der Spirostreptidae (Diplopoda; Spirostreptomorpha). Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg (Neue Folge,) 24, 1 - 476." type="journal article" year="1982">Krabbe, 1982</bibRefCitation>
</taxonomicName>
, Abh. Naturw. Ver. Hamburg, NF 24: 207.
</paragraph>
<paragraph id="D050F117C22ED96FFF3AF9E0FAEEFD5B" blockId="2.[151,1436,477,744]" box="[151,1394,722,744]" pageId="2" pageNumber="55">
<taxonomicName id="17EF8A94C22ED96FFF3AF9E0FDFFFD5B" authority="Hoffman, 1993" authorityName="Hoffman" authorityYear="1993" box="[151,611,722,744]" class="Diplopoda" family="Spirostreptidae" genus="Haplogonopus" higherTaxonomySource="CoL" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="inflatannulus">
<emphasis id="E29B2D05C22ED96FFF3AF9E0FE5CFD5B" box="[151,448,722,744]" italics="true" pageId="2" pageNumber="55">Haplogonopus inflatannulus:</emphasis>
<bibRefCitation id="B47E8CE6C22ED96FFE6BF9E0FDFFFD5B" author="Hoffman" box="[454,611,722,744]" pageId="2" pageNumber="62" refString="Hoffman, R. L. (1993) Biogeography of East African montane forest millipedes. Chapt. 6, pp. 103 - 114, in: J. C. Lovett &amp; S. K. Wasser, Biogeography and ecology of the rain forests of eastern Africa. Cambridge University Press. 341 pp." type="book chapter" year="1993">Hoffman, 1993</bibRefCitation>
</taxonomicName>
, in:
<emphasis id="E29B2D05C22ED96FFD23F9E0FABAFD5B" box="[654,1318,722,744]" italics="true" pageId="2" pageNumber="55">Biogeography and Ecology of the Rain Forests of Eastern Africa,</emphasis>
p. 106.
</paragraph>
</subSubSection>
<subSubSection id="98F5A29CC22ED96FFF3AF827FB8AFBB6" pageId="2" pageNumber="55" type="materials_examined">
<paragraph id="D050F117C22ED96FFF3AF827FB8AFBB6" blockId="2.[151,1437,789,2010]" pageId="2" pageNumber="55">
<emphasis id="E29B2D05C22ED96FFF3AF827FE9AFC9D" bold="true" box="[151,262,789,814]" pageId="2" pageNumber="55">Material.</emphasis>
<collectingCountry id="A8F8B187C22ED96FFEA3F824FE3DFC9D" box="[270,417,790,814]" name="Tanzania" pageId="2" pageNumber="55">TANZANIA</collectingCountry>
: Morogoro Region and District:
<emphasis id="E29B2D05C22ED96FFCB8F827FBF8FC9E" box="[789,1124,789,813]" italics="true" pageId="2" pageNumber="55">Northern Uluguru Mountains</emphasis>
: Lupanga Peak West,
<quantity id="17175CF2C22ED96FFAC9F827FF33FCE2" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="2" pageNumber="55" unit="m" value="1800.0">1800 m</quantity>
,
<date id="A451D7D7C22ED96FFF16F80BFEDCFCE2" box="[187,320,825,849]" pageId="2" pageNumber="55" value="1981-07-01">1 July 1981</date>
, Nikolaj Scharff and Michael Stoltze, 6 3, 2 Ƥ (
<collectionCode id="B6FE69D2C22ED96FFCF2F80BFC2AFCE2" box="[863,950,825,849]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="2" pageNumber="55">ZMUC</collectionCode>
); north slope of Lupanga,
<date id="A451D7D7C22ED96FFB71F80BFA04FCE2" box="[1244,1432,825,850]" pageId="2" pageNumber="55" value="1980-01-11">11 January 1980</date>
, S. N. Stuart &amp; T. van der Willigen, 1Ƥ (
<collectionCode id="B6FE69D2C22ED96FFDF5F86FFD28FCC6" box="[600,692,861,885]" httpUri="http://grbio.org/cool/zxpe-c0as" name="Virginia Museum of Natural History" pageId="2" pageNumber="55">VMNH</collectionCode>
); in forest near Morningside,
<date id="A451D7D7C22ED96FFBADF86FFB44FCC6" box="[1024,1240,861,885]" pageId="2" pageNumber="55" value="1973-12-23">23 December 1973</date>
, K. M. Howell et al., 5 3 (
<collectionCode id="B6FE69D2C22ED96FFF55F8B3FEC8FC2A" box="[248,340,897,921]" httpUri="http://grbio.org/cool/zxpe-c0as" name="Virginia Museum of Natural History" pageId="2" pageNumber="55">VMNH</collectionCode>
); Morningside,
<date id="A451D7D7C22ED96FFDAEF8B3FD3FFC2A" box="[515,675,897,921]" pageId="2" pageNumber="55" value="1980-04-27">27 April 1980</date>
, K. M. Howell, 13 (
<collectionCode id="B6FE69D2C22ED96FFC2AF8B3FC7FFC2A" box="[903,995,897,921]" httpUri="http://grbio.org/cool/zxpe-c0as" name="Virginia Museum of Natural History" pageId="2" pageNumber="55">VMNH</collectionCode>
).
<emphasis id="E29B2D05C22ED96FFC58F8B3FADBFC2A" box="[1013,1351,897,921]" italics="true" pageId="2" pageNumber="55">Southern Uluguru Mountains:</emphasis>
Hululu Falls, Mgeta River valley south of Bunduki,
<quantity id="17175CF2C22ED96FFD3BF897FD7BFC0E" box="[662,743,933,957]" metricMagnitude="3" metricUnit="m" metricValue="1.8288" pageId="2" pageNumber="55" unit="ft" value="6000.0">6000 ft</quantity>
asl,
<date id="A451D7D7C22ED96FFCB3F897FC60FC0E" box="[798,1020,933,957]" pageId="2" pageNumber="55" value="1979-11-16">16 November 1979</date>
, W. A. Rodgers &amp; K. Bulstrode, 13 (
<collectionCode id="B6FE69D2C22ED96FFF32F8FBFF67FC52" box="[159,251,969,993]" httpUri="http://grbio.org/cool/zxpe-c0as" name="Virginia Museum of Natural History" pageId="2" pageNumber="55">VMNH</collectionCode>
), also 13, 1Ƥ, date and collector not specified (
<collectionCode id="B6FE69D2C22ED96FFCB8F8FBFCEDFC52" box="[789,881,969,993]" httpUri="http://grbio.org/cool/zxpe-c0as" name="Virginia Museum of Natural History" pageId="2" pageNumber="55">VMNH</collectionCode>
); Bondwa Mountain, Uluguru Forest Reserve,
<date id="A451D7D7C22ED96FFA2DF8FBFE88FBB6" pageId="2" pageNumber="55" value="1980-04-24">24 April 1980</date>
, J. Kielland, 13 (
<collectionCode id="B6FE69D2C22ED96FFE75F8DFFDA8FBB6" box="[472,564,1005,1029]" httpUri="http://grbio.org/cool/zxpe-c0as" name="Virginia Museum of Natural History" pageId="2" pageNumber="55">VMNH</collectionCode>
);
<date id="A451D7D7C22ED96FFDEAF8DFFC9EFBB6" box="[583,770,1005,1029]" pageId="2" pageNumber="55" value="1982-02-02">2 February 1982</date>
, Kielland, 1Ƥ (
<collectionCode id="B6FE69D2C22ED96FFC02F8DFFB97FBB6" box="[943,1035,1005,1029]" httpUri="http://grbio.org/cool/zxpe-c0as" name="Virginia Museum of Natural History" pageId="2" pageNumber="55">VMNH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="98F5A29CC22ED96FFF68FF23FB7AFA24" pageId="2" pageNumber="55" type="diagnosis">
<paragraph id="D050F117C22ED96FFF68FF23FC2DFBFE" blockId="2.[151,1437,789,2010]" pageId="2" pageNumber="55">
<emphasis id="E29B2D05C22ED96FFF68FF23FEA3FB99" bold="true" box="[197,319,1041,1066]" pageId="2" pageNumber="55">Diagnosis.</emphasis>
Distinguished from the two known congeneric species (and other Tanzanian spirostreptids) by its larger size, elevated, orange-red metazona, and unmodified paraprocts.
</paragraph>
<paragraph id="D050F117C22ED96FFF6AFF6BFAFBFAFF" blockId="2.[151,1437,789,2010]" pageId="2" pageNumber="55">
<emphasis id="E29B2D05C22ED96FFF6AFF6BFE07FBC1" bold="true" box="[199,411,1113,1138]" pageId="2" pageNumber="55">Descriptive notes.</emphasis>
Male diameter to 6.5 mm, female to
<quantity id="17175CF2C22ED96FFCEEFF6BFC1BFBC2" box="[835,903,1113,1137]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="55" unit="mm" value="9.0">9 mm</quantity>
, female length to about
<quantity id="17175CF2C22ED96FFB35FF6BFB64FBC2" box="[1176,1272,1113,1137]" metricMagnitude="-1" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="55" unit="mm" value="110.0">110 mm</quantity>
. Front of head smooth and convex, unmodified, ocellaria elongate-triangular, about as long as interocellarial space. Mandibular basomere with distal projection (
<figureCitation id="48D4ED92C22ED96FFDABFF93FDD6FB0A" box="[518,586,1185,1209]" captionStart="FIGURES 1 5" captionStartId="3.[151,264,1922,1945]" captionTargetBox="[214,1374,207,1871]" captionTargetId="figure@3.[196,1395,195,1881]" captionTargetPageId="3" captionText="FIGURES 1 5. Haplogonopus inflatannulus Verhoeff., structural features. 1. Gnathochilarium and adjacent parts of mandibles and collum. 2. Right mandible, posterior aspect, with distal lobe of basomere indicated by the ↔. 3. First pair of legs of male, syncoxa, sternum, and basal podomeres, anterior aspect. 4. The same, posterior aspect, sclerite of uncertain identity indicated by the? 5. Dorsal half of three midbody segments, lateral aspect, showing convex metazona with ovoid depressions." httpUri="https://zenodo.org/record/203122/files/figure.png" pageId="2" pageNumber="55">Fig. 2</figureCitation>
).. Elevated part of metazona orange to light brown, prozona grayish, in preserved specimens, legs yellow. Postfemoral and tibial pads of male small, diminishing posterior to gonopods, scarcely evident posterior to legs of 25th segement. Sigilla small, generally bacilliform but highly variable, dispersed as an irregular belt over anterior two-thirds of pigmented metazonal surface. Paraprocts strongly convex, the caudal edges virtually immarginate, no trace of fossae or diastemma. Hypoproct short, transversely elongated.
</paragraph>
<paragraph id="D050F117C22ED96FFF6AFE69FB7AFA24" blockId="2.[151,1437,789,2010]" pageId="2" pageNumber="55">Coxae of 1st pair of legs medially coalesced with only a trace of suture line on posterior side, this syncoxa only slightly prolonged laterad beyond base of prefemora. Sternum of these legs with distinct stigmata.</paragraph>
</subSubSection>
<subSubSection id="98F5A29CC22ED969FF68FE91FA94FE8C" lastPageId="4" lastPageNumber="57" pageId="2" pageNumber="55" type="discussion">
<paragraph id="D050F117C22ED96FFF68FE91FE8FF9B0" blockId="2.[151,1437,789,2010]" pageId="2" pageNumber="55">
<emphasis id="E29B2D05C22ED96FFF68FE91FEA5FA08" bold="true" box="[197,313,1443,1467]" pageId="2" pageNumber="55">Remarks.</emphasis>
Although I have not examined the
<typeStatus id="0F544FB5C22ED96FFD7DFE96FC9CFA0F" box="[720,768,1444,1468]" pageId="2" pageNumber="55">type</typeStatus>
material of either
<taxonomicName id="17EF8A94C22ED96FFC7BFE91FBF6FA08" box="[982,1130,1443,1467]" class="Diplopoda" family="Spirostreptidae" genus="Haplogonopus" higherTaxonomySource="CoL" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="inflatannulus">
<emphasis id="E29B2D05C22ED96FFC7BFE91FBF6FA08" box="[982,1130,1443,1467]" italics="true" pageId="2" pageNumber="55">inflatannulus</emphasis>
</taxonomicName>
or
<taxonomicName id="17EF8A94C22ED96FFB3FFE91FA96FA08" box="[1170,1290,1443,1467]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="nebularius">
<emphasis id="E29B2D05C22ED96FFB3FFE91FA96FA08" box="[1170,1290,1443,1467]" italics="true" pageId="2" pageNumber="55">nebularius</emphasis>
</taxonomicName>
, the original accounts of these names leave no doubt of their actual identity, as further substantiated by the study of topotypic specimens.
</paragraph>
<paragraph id="D050F117C22ED96FFF6AFD3DFC68F926" blockId="2.[151,1437,789,2010]" pageId="2" pageNumber="55">
The legs of the first pair are of the conventional spirostreptid form, prefemoral lobes relatively small and subtriangular, but of interest by the presence of a transverse sclerite (
<figureCitation id="48D4ED92C22ED96FFCD6FD01FC5CF9F8" box="[891,960,1587,1611]" captionStart="FIGURES 1 5" captionStartId="3.[151,264,1922,1945]" captionTargetBox="[214,1374,207,1871]" captionTargetId="figure@3.[196,1395,195,1881]" captionTargetPageId="3" captionText="FIGURES 1 5. Haplogonopus inflatannulus Verhoeff., structural features. 1. Gnathochilarium and adjacent parts of mandibles and collum. 2. Right mandible, posterior aspect, with distal lobe of basomere indicated by the ↔. 3. First pair of legs of male, syncoxa, sternum, and basal podomeres, anterior aspect. 4. The same, posterior aspect, sclerite of uncertain identity indicated by the? 5. Dorsal half of three midbody segments, lateral aspect, showing convex metazona with ovoid depressions." httpUri="https://zenodo.org/record/203122/files/figure.png" pageId="2" pageNumber="55">Fig. 4</figureCitation>
,?) subtending the syncoxon on the posterior side.. Whether this is merely the detatched continuation of the sternum or an intercalary element between the 1st and 2nd leg pairs could not be established with the material at my disposal.
</paragraph>
<paragraph id="D050F117C22ED96FFF6AFD93FD49F8FA" blockId="2.[151,1437,789,2010]" pageId="2" pageNumber="55">
General uniformly within the several samples available for study suggest that apparently the Uluguru Mountains are not extensive or fragmented enough to have engendered any local diversification (although only a few specimens originated in the southern component of the range). Segment counts (including collum but not the epiproct) range between 47 and 51 (avg. 48.7) in
<specimenCount id="C6E93A9EC22ED96FFD6CFC3FFCB4F896" box="[705,808,1805,1829]" pageId="2" pageNumber="55" type="male">14 males</specimenCount>
, and 4851 (avg. 50.2) in five females. Gonopods show what appears to be only minor individual variation.
</paragraph>
<paragraph id="D050F117C22ED969FF68FC67FA94FE8C" blockId="2.[151,1437,789,2010]" lastBlockId="4.[151,1436,151,320]" lastPageId="4" lastPageNumber="57" pageId="2" pageNumber="55">
The gonopods of
<taxonomicName id="17EF8A94C22ED96FFE21FC64FDBCF8DD" box="[396,544,1878,1902]" class="Diplopoda" family="Spirostreptidae" genus="Haplogonopus" higherTaxonomySource="CoL" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="inflatannulus">
<emphasis id="E29B2D05C22ED96FFE21FC64FDBCF8DD" box="[396,544,1878,1902]" italics="true" pageId="2" pageNumber="55">inflatannulus</emphasis>
</taxonomicName>
are so similar to those of
<taxonomicName id="17EF8A94C22ED96FFCEAFC64FC3DF8DD" box="[839,929,1878,1902]" class="Diplopoda" family="Spirostreptidae" genus="Charactopygus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="jeanneli">
<emphasis id="E29B2D05C22ED96FFCEAFC64FC3DF8DD" box="[839,929,1878,1902]" italics="true" pageId="2" pageNumber="55">jeanneli</emphasis>
</taxonomicName>
that I initially suspected that the latter was a synonym based on specimens transported from the Ulugurus into
<collectingCountry id="A8F8B187C22ED96FFC36FC48FC78F821" box="[923,996,1914,1938]" name="Kenya" pageId="2" pageNumber="55">Kenya</collectingCountry>
in plant material although the virtual absence of commerce between
<collectingCountry id="A8F8B187C22ED96FFE55FCACFDDDF805" box="[504,577,1950,1974]" name="Kenya" pageId="2" pageNumber="55">Kenya</collectingCountry>
and Tanganyika in the early colonial period seemed to militate against such an option. My colleague J.-P. Mauriès examined the
<typeStatus id="0F544FB5C22ED96FFD4AFCF0FC84F869" box="[743,792,1986,2010]" pageId="2" pageNumber="55">type</typeStatus>
specimen of
<taxonomicName id="17EF8A94C22ED96FFC02FCF0FB96F869" box="[943,1034,1986,2010]" class="Diplopoda" family="Spirostreptidae" genus="Charactopygus" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="jeanneli">
<emphasis id="E29B2D05C22ED96FFC02FCF0FB96F869" box="[943,1034,1986,2010]" italics="true" pageId="2" pageNumber="55">jeanneli</emphasis>
</taxonomicName>
and reported that the metazona are absolutely flat and smooth, confirming the status of
<taxonomicName id="17EF8A94C228D969FD42FBAAFCD6FF03" box="[751,842,152,176]" class="Diplopoda" family="Spirostreptidae" genus="Charactopygus" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="57" phylum="Arthropoda" rank="species" species="jeanneli">
<emphasis id="E29B2D05C228D969FD42FBAAFCD6FF03" box="[751,842,152,176]" italics="true" pageId="4" pageNumber="57">jeanneli</emphasis>
</taxonomicName>
as a valid species, as do additional characters mentioned in the following accounts. This, with another “smooth-bodied” species discovered in southern
<collectingCountry id="A8F8B187C228D969FA9DFB8EFA0BFF67" box="[1328,1431,188,212]" name="Tanzania" pageId="4" pageNumber="57">Tanzania</collectingCountry>
, gives the impression of a fragmented generic range resulting from late Tertiary climatic changes, and the evolution of singular peripheral characters in the Uluguru isolate. Such a scenario evokes the possibility, even liklihood, that additional highly localized species may yet be discovered in, e.g., the Usambara or Nguru mountains.
</paragraph>
</subSubSection>
</treatment>
</document>