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<document id="4EC4B3C416BBBBCFFD76B8C2BEE58AFC" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629589581802" checkinUser="felipe" docAuthor="Oyewo, Emmanuel Adeoye &amp; Saether, Ole A." docDate="2008" docId="57405B27FFD9FF81FF13F93F89FDFBF0" docLanguage="en" docName="zt01953p145.pdf" docOrigin="Zootaxa 1953" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Polypedilum (Pentapedilum) toyamadeeum Oyewo &amp; Saether 2008, comb. n." docType="treatment" docVersion="4" lastPageNumber="110" masterDocId="AB79235FFFB5FFECFF84FFCB8B6FFFF3" masterDocTitle="Revision of Polypedilum (Pentapedilum) Kieffer and Ainuyusurika Sasa et Shirasaki (Diptera: Chironomidae)" masterLastPageNumber="145" masterPageNumber="1" pageNumber="109" updateTime="1699081859134" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="9A6FF3B8BF8A1C95C5DCD0AE89E636B8">Revision of Polypedilum (Pentapedilum) Kieffer and Ainuyusurika Sasa et Shirasaki (Diptera: Chironomidae)</mods:title>
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<mods:namePart id="A1CD96577DF844B8DD4A8252D53FB25D">Oyewo, Emmanuel Adeoye</mods:namePart>
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<mods:namePart id="86A55D998709B9A711AAC772BE9E9196">Saether, Ole A.</mods:namePart>
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<mods:date id="989F5EDE2A19FBD318C3D8D581A961D0">2008</mods:date>
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<mods:number id="F648E65234D95928E9DFCF796C44013A">2008-12-05</mods:number>
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<treatment id="57405B27FFD9FF81FF13F93F89FDFBF0" ID-DOI="http://doi.org/10.5281/zenodo.5241899" ID-Zenodo-Dep="5241899" LSID="urn:lsid:plazi:treatment:57405B27FFD9FF81FF13F93F89FDFBF0" httpUri="http://treatment.plazi.org/id/57405B27FFD9FF81FF13F93F89FDFBF0" lastPageId="109" lastPageNumber="110" pageId="108" pageNumber="109">
<subSubSection id="97F3B9BAFFD9FF80FF13F93F880AF8FE" box="[151,869,1779,1806]" pageId="108" pageNumber="109" type="nomenclature">
<paragraph id="DF56EA31FFD9FF80FF13F93F880AF8FE" blockId="108.[151,869,1779,1841]" box="[151,869,1779,1806]" pageId="108" pageNumber="109">
<heading id="841E5D5DFFD9FF80FF13F93F880AF8FE" bold="true" box="[151,869,1779,1806]" fontSize="11" level="1" pageId="108" pageNumber="109" reason="1">
<emphasis id="ED9D3623FFD9FF80FF13F93F880AF8FE" bold="true" box="[151,869,1779,1806]" pageId="108" pageNumber="109">
<taxonomicName id="18E991B2FFD9FF80FF13F93F899DF8FE" ID-CoL="9HTBS" authority="(Sasa)" authorityName="Oyewo &amp; Saether" authorityYear="2008" baseAuthorityName="Sasa" box="[151,754,1779,1806]" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="toyamadeeum" status="comb. nov." subGenus="Pentapedilum">
<emphasis id="ED9D3623FFD9FF80FF13F93F8A41F8FD" bold="true" box="[151,302,1780,1806]" italics="true" pageId="108" pageNumber="109">Polypedilum</emphasis>
(
<emphasis id="ED9D3623FFD9FF80FEBBF93F8A89F8FD" bold="true" box="[319,486,1780,1806]" italics="true" pageId="108" pageNumber="109">Pentapedilum</emphasis>
)
<emphasis id="ED9D3623FFD9FF80FE7CF93F89F0F8FD" bold="true" box="[504,671,1780,1806]" italics="true" pageId="108" pageNumber="109">toyamadeeum</emphasis>
(Sasa)
</taxonomicName>
<taxonomicNameLabel id="F6AE8B58FFD9FF80FD7EF938880AF8FE" box="[762,869,1779,1805]" pageId="108" pageNumber="109" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="97F3B9BAFFD9FF80FF13F8DC8A7CF8C2" box="[151,275,1815,1841]" pageId="108" pageNumber="109" type="description">
<paragraph id="DF56EA31FFD9FF80FF13F8DC8A7CF8C2" blockId="108.[151,869,1779,1841]" box="[151,275,1815,1841]" pageId="108" pageNumber="109">
(
<figureCitation id="47D2F6B4FFD9FF80FF1BF8DC8A66F8C2" box="[159,265,1815,1841]" captionStart="FIGURE 15" captionStartId="53.[151,254,1631,1655]" captionTargetBox="[268,1321,195,1587]" captionTargetId="figure-213@53.[245,1342,181,1693]" captionTargetPageId="53" captionText="FIGURE 15. Polypedilum (Pentapedilum) spp., males, anal point (C) and superior volsella. A—P. (Pe.) elongatus Tokunaga; B, C—P. (Pe.) kamosecundum (Sasa); D—P. (Pe.) mellense (Goetghebuer); E—P. (Pe.) nodosum (Johannsen); F—P. (Pe.) pseudosordens Zhang et Wang; G—P. (Pe.) palauensis Tokunaga; H—P. (Pe.) toyamadeeum (Sasa); I—P. (Pe.) tusimaefeum (Sasa); J—P. (Pe.) tusimafegeum (Sasa)." pageId="108" pageNumber="109">Fig. 15H</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="97F3B9BAFFD9FF80FF13F8AC89F8F88E" box="[151,663,1894,1918]" pageId="108" pageNumber="109" type="reference_group">
<paragraph id="DF56EA31FFD9FF80FF13F8AC89F8F88E" blockId="108.[151,663,1894,1918]" box="[151,663,1894,1918]" pageId="108" pageNumber="109">
<treatmentCitationGroup id="FFF9CD1FFFD9FF80FF13F8AC89F8F88E" box="[151,663,1894,1918]" pageId="108" pageNumber="109">
<treatmentCitation id="5E48CC20FFD9FF80FF13F8AC8904F88E" author="Sasa, M." box="[151,619,1894,1918]" page="23" pageId="108" pageNumber="109" year="1996">
<taxonomicName id="18E991B2FFD9FF80FF13F8AC8904F88E" ID-CoL="9HY4L" authority="Sasa, 1996: 23" authorityName="Sasa" authorityPageNumber="23" authorityYear="1996" box="[151,619,1894,1918]" class="Insecta" family="Chironomidae" genus="Pentapedilum" kingdom="Animalia" order="Diptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="toyamadeeum">
<emphasis id="ED9D3623FFD9FF80FF13F8AC8AA9F88D" box="[151,454,1895,1918]" italics="true" pageId="108" pageNumber="109">Pentapedilum toyamadeeum</emphasis>
<bibRefCitation id="BB7897C0FFD9FF80FE48F8AD8904F88E" author="Sasa, M." box="[460,619,1894,1917]" pageId="108" pageNumber="109" pagination="16 - 112" refId="ref93190" refString="Sasa, M. (1996) Seasonal distribution of the chironomids collected with light traps at the side of two lakes in the Toyama City Family Park. Report from Toyama Prefectural Environmental Pollution Research Centre, 1996 (March), 16 - 112." type="journal article" year="1996">Sasa, 1996: 23</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 48.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="97F3B9BAFFD9FF81FF13F8658990FC28" lastPageId="109" lastPageNumber="110" pageId="108" pageNumber="109" type="materials_examined">
<paragraph id="DF56EA31FFD9FF80FF13F8658953F802" blockId="108.[151,1436,1966,2033]" pageId="108" pageNumber="109">
<emphasis id="ED9D3623FFD9FF80FF13F8658AE3F83B" bold="true" box="[151,396,1966,1992]" pageId="108" pageNumber="109">Material examined.</emphasis>
<materialsCitation id="6F81E06CFFD9FF80FE10F8648956F802" collectingDate="1993-06-23" collectionCode="NMST" collectorName="Lake A &amp; M. Sasa &amp; Type No." country="Japan" location="Family Park" pageId="108" pageNumber="109" specimenCount="1" stateProvince="Toyama" typeStatus="holotype">
<collectingCountry id="A7FEAAA1FFD9FF80FE10F8648A9EF83A" box="[404,497,1967,1993]" name="Japan" pageId="108" pageNumber="109">JAPAN</collectingCountry>
:
<collectingRegion id="1D2D24D3FFD9FF80FE79F86489B1F83A" box="[509,734,1967,1993]" country="Japan" name="Toyama" pageId="108" pageNumber="109">Toyama Prefecture</collectingRegion>
,
<collectingRegion id="1D2D24D3FFD9FF80FD6FF8648823F83A" box="[747,844,1967,1993]" country="Japan" name="Toyama" pageId="108" pageNumber="109">Toyama</collectingRegion>
,
<location id="DA36BCEAFFD9FF80FCDCF8648884F83A" LSID="urn:lsid:plazi:treatment:57405B27FFD9FF81FF13F93F89FDFBF0:DA36BCEAFFD9FF80FCDCF8648884F83A" box="[856,1003,1967,1993]" country="Japan" name="Family Park" pageId="108" pageNumber="109" stateProvince="Toyama">Family Park</location>
,
<collectorName id="721C8FE7FFD9FF80FC7DF8648F3CF83A" box="[1017,1107,1967,1993]" pageId="108" pageNumber="109">Lake A</collectorName>
,
<typeStatus id="00525493FFD9FF80FBE4F8648FA9F83A" box="[1120,1222,1967,1993]" pageId="108" pageNumber="109" type="holotype">holotype</typeStatus>
male,
<date id="AB57CCF1FFD9FF80FA93F8648EF9F83A" box="[1303,1430,1967,1993]" pageId="108" pageNumber="109" value="1993-06-23">
<collectingDate id="BB133519FFD9FF80FA93F8648EF9F83A" box="[1303,1430,1967,1993]" pageId="108" pageNumber="109" value="1993-06-23">23.vi.1993</collectingDate>
</date>
,
<collectorName id="721C8FE7FFD9FF80FF13F81C8B9AF802" box="[151,245,2007,2033]" pageId="108" pageNumber="109">M. Sasa</collectorName>
(
<collectionCode id="B9F872F4FFD9FF80FE80F81C8A38F802" box="[260,343,2007,2033]" pageId="108" pageNumber="109">NMST</collectionCode>
<collectorName id="721C8FE7FFD9FF80FEDBF81C8AA3F802" box="[351,460,2007,2033]" pageId="108" pageNumber="109">Type No.</collectorName>
272: 14)
</materialsCitation>
.
</paragraph>
<paragraph id="DF56EA31FFD8FF81FF43FF5C8F24FEF1" blockId="109.[151,1436,151,1027]" pageId="109" pageNumber="110">
<emphasis id="ED9D3623FFD8FF81FF43FF5C8A8FFF42" bold="true" box="[199,480,151,177]" pageId="109" pageNumber="110">Diagnostic characters.</emphasis>
The species differs from other species of the subgenus, except
<taxonomicName id="18E991B2FFD8FF81FB6DFF538BB3FF2A" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" pageId="109" pageNumber="29" phylum="Arthropoda" rank="species" species="fanjingense" subGenus="Pe.">
<emphasis id="ED9D3623FFD8FF81FB6DFF538F97FF42" box="[1257,1272,152,177]" italics="true" pageId="109" pageNumber="110">P</emphasis>
. (
<emphasis id="ED9D3623FFD8FF81FA97FF538E5DFF42" box="[1299,1330,152,177]" italics="true" pageId="109" pageNumber="110">Pe</emphasis>
.)
<emphasis id="ED9D3623FFD8FF81FACAFF538BB3FF2A" italics="true" pageId="109" pageNumber="110">fanjingense</emphasis>
</taxonomicName>
<emphasis id="ED9D3623FFD8FF81FF58FF0B8B8EFF2A" box="[220,225,192,217]" italics="true" pageId="109" pageNumber="110">,</emphasis>
by the low antennal ratio (0.30.4) combined with a broad anal point, with median furrow reaching apex. It differs from
<taxonomicName id="18E991B2FFD8FF81FE0EFF238900FEF2" box="[394,623,232,258]" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" pageId="109" pageNumber="29" phylum="Arthropoda" rank="species" species="fanjingense" subGenus="Pe.">
<emphasis id="ED9D3623FFD8FF81FE0EFF238AF7FEF2" box="[394,408,232,257]" italics="true" pageId="109" pageNumber="110">P</emphasis>
. (
<emphasis id="ED9D3623FFD8FF81FE35FF238ABFFEF2" box="[433,464,232,257]" italics="true" pageId="109" pageNumber="110">Pe</emphasis>
.)
<emphasis id="ED9D3623FFD8FF81FE6CFF238900FEF2" box="[488,623,232,257]" italics="true" pageId="109" pageNumber="110">fanjingense</emphasis>
</taxonomicName>
by lacking a spur on the foretibial scale.
</paragraph>
<paragraph id="DF56EA31FFD8FF81FF43FEC48A31FED9" blockId="109.[151,1436,151,1027]" box="[199,350,271,298]" pageId="109" pageNumber="110">
<emphasis id="ED9D3623FFD8FF81FF43FEC48A69FEDA" bold="true" box="[199,262,271,297]" pageId="109" pageNumber="110">Male</emphasis>
(n = 1)
</paragraph>
<paragraph id="DF56EA31FFD8FF81FF43FEF38F5FFEA1" blockId="109.[151,1436,151,1027]" box="[199,1072,312,338]" pageId="109" pageNumber="110">
Wing length
<quantity id="181147D4FFD8FF81FEDBFEF38AA5FEA1" box="[351,458,312,338]" metricMagnitude="-3" metricUnit="m" metricValue="1.73" pageId="109" pageNumber="110" unit="mm" value="1.73">1.73 mm</quantity>
. Wing length / length of profemur 2.19. Pale brown.
</paragraph>
<paragraph id="DF56EA31FFD8FF81FF43FEAB8A5EFE39" blockId="109.[151,1436,151,1027]" pageId="109" pageNumber="110">
<emphasis id="ED9D3623FFD8FF81FF43FEAB8A61FE8A" box="[199,270,352,377]" italics="true" pageId="109" pageNumber="110">Head.</emphasis>
AR 0.33. Ultimate flagellomere 191 µm. Temporal setae 15. Clypeus with 26 setae. Tentorium 140 µm long. Stipes 151 µm long. Palpomere lengths (in µm): 44, 40, 93, 127, 175. Fifth palpomere / third palpomere 1.88.
</paragraph>
<paragraph id="DF56EA31FFD8FF81FF43FE138F0FFE01" blockId="109.[151,1436,151,1027]" box="[199,1120,472,498]" pageId="109" pageNumber="110">
<emphasis id="ED9D3623FFD8FF81FF43FE138A4CFE02" box="[199,291,472,497]" italics="true" pageId="109" pageNumber="110">Thorax.</emphasis>
Acrostichals 22, dorsocentrals 30, prealars 7.
<taxonomicName id="18E991B2FFD8FF81FCC4FE1388D8FE01" box="[832,951,472,498]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="109" pageNumber="15" phylum="Arthropoda" rank="genus">Scutellum</taxonomicName>
with 12 setae.
</paragraph>
<paragraph id="DF56EA31FFD8FF81FF43FDCB89A5FD68" blockId="109.[151,1436,151,1027]" pageId="109" pageNumber="110">
<emphasis id="ED9D3623FFD8FF81FF43FDCB8A64FDEA" box="[199,267,512,537]" italics="true" pageId="109" pageNumber="110">Wing.</emphasis>
Anal lobe developed. VR 1.19. Brachiolum with 3 setae, Sc with 21 setae,
<collectionCode id="B9F872F4FFD8FF81FB14FDCB8FCCFDE9" box="[1168,1187,512,538]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="109" pageNumber="110">R</collectionCode>
with 40,
<collectionCode id="B9F872F4FFD8FF81FA92FDCB8E46FDE9" box="[1302,1321,512,538]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="109" pageNumber="110">R</collectionCode>
<subScript id="436DE874FFD8FF81FAAEFDDB8E5DFDED" attach="left" box="[1322,1330,528,542]" fontSize="6" pageId="109" pageNumber="110">1</subScript>
with 34,
<collectionCode id="B9F872F4FFD8FF81FF13FDE18BC5FDB7" box="[151,170,554,580]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="109" pageNumber="110">R</collectionCode>
<subScript id="436DE874FFD8FF81FF2EFDF08BABFDBA" attach="left" box="[170,196,571,585]" fontSize="6" pageId="109" pageNumber="110">4+5</subScript>
with 93, M with 14,
<collectingRegion id="1D2D24D3FFD8FF81FE44FDE18A81FDB7" box="[448,494,554,580]" country="Kenya" name="Lamu" pageId="109" pageNumber="110">RM</collectingRegion>
with 4, M
<subScript id="436DE874FFD8FF81FDE8FDF089EAFDBA" attach="left" box="[620,645,571,585]" fontSize="6" pageId="109" pageNumber="110">1+2</subScript>
with 81, M
<subScript id="436DE874FFD8FF81FC97FDF08842FDBA" attach="left" box="[787,813,571,585]" fontSize="6" pageId="109" pageNumber="110">3+4</subScript>
with 38, Cu with 44, Cu
<subScript id="436DE874FFD8FF81FBD3FDF08F30FDBA" attach="left" box="[1111,1119,571,585]" fontSize="6" pageId="109" pageNumber="110">1</subScript>
with 24, PCu with 55, An with 33 setae. Cells r
<subScript id="436DE874FFD8FF81FE0BFDAE8AC7FD80" attach="left" box="[399,424,613,627]" fontSize="6" pageId="109" pageNumber="110">4+5</subScript>
, m
<subScript id="436DE874FFD8FF81FE4AFDAE8A88FD80" attach="left" box="[462,487,613,627]" fontSize="6" pageId="109" pageNumber="110">1+2</subScript>
, m
<subScript id="436DE874FFD8FF81FD89FDAE8949FD80" attach="left" box="[525,550,613,627]" fontSize="6" pageId="109" pageNumber="110">3+4</subScript>
, cell m basal of
<collectingRegion id="1D2D24D3FFD8FF81FD6EFD9E8875FD9C" box="[746,794,597,623]" country="Kenya" name="Lamu" pageId="109" pageNumber="110">RM</collectingRegion>
, cu plus an each with numerous macrotrichia. Squama lost; with 1016 setae according to
<bibRefCitation id="BB7897C0FFD8FF81FDB3FD4A89A9FD68" author="Sasa, M." box="[567,710,641,667]" pageId="109" pageNumber="110" pagination="16 - 112" refId="ref93190" refString="Sasa, M. (1996) Seasonal distribution of the chironomids collected with light traps at the side of two lakes in the Toyama City Family Park. Report from Toyama Prefectural Environmental Pollution Research Centre, 1996 (March), 16 - 112." type="journal article" year="1996">Sasa (1996)</bibRefCitation>
.
</paragraph>
<paragraph id="DF56EA31FFD8FF81FF43FD628EF3FCE0" blockId="109.[151,1436,151,1027]" pageId="109" pageNumber="110">
<emphasis id="ED9D3623FFD8FF81FF43FD628A69FD31" box="[199,262,681,706]" italics="true" pageId="109" pageNumber="110">Legs.</emphasis>
Scale on foretibia 58 µm long, triangular, without apical spur. Spurs on midtibia 73 µm long, on hind tibia 65 µm long. Comb on midtibia 28 µm long, combs on hind tibia 33 µm and 30 µm long. Width at apex of foretibia 64 µm, of midtibia 63 µm, of hind tibia 70 µm. Length and proportions of legs as in
<tableCitation id="926BDF8AFFD8FF81FAB4FD328EF7FCE0" box="[1328,1432,761,787]" captionStart="TABLE 45" captionStartId="109.[151,239,1080,1104]" captionText="TABLE 45. Length (in µm) and proportions of legs of Polypedilum (Pe.) toyamadeeum (Sasa), male (n = 1)." pageId="109" pageNumber="110">Table 45</tableCitation>
.
</paragraph>
<paragraph id="DF56EA31FFD8FF81FF43FCEA8990FC28" blockId="109.[151,1436,151,1027]" pageId="109" pageNumber="110">
<emphasis id="ED9D3623FFD8FF81FF43FCEA8A0BFCC9" box="[199,356,801,826]" italics="true" pageId="109" pageNumber="110">Hypopygium.</emphasis>
Tergite IX with 20 median setae, laterosternite IX with 3 setae. Anal point parallel-sided, not measurable. Transverse sternapodeme 37 µm long; phallapodeme 73 µm long. Gonocoxite 163 µm long. Superior volsella (
<figureCitation id="47D2F6B4FFD8FF81FEF0FCBA8A8EFC78" box="[372,481,881,907]" captionStart="FIGURE 15" captionStartId="53.[151,254,1631,1655]" captionTargetBox="[268,1321,195,1587]" captionTargetId="figure-213@53.[245,1342,181,1693]" captionTargetPageId="53" captionText="FIGURE 15. Polypedilum (Pentapedilum) spp., males, anal point (C) and superior volsella. A—P. (Pe.) elongatus Tokunaga; B, C—P. (Pe.) kamosecundum (Sasa); D—P. (Pe.) mellense (Goetghebuer); E—P. (Pe.) nodosum (Johannsen); F—P. (Pe.) pseudosordens Zhang et Wang; G—P. (Pe.) palauensis Tokunaga; H—P. (Pe.) toyamadeeum (Sasa); I—P. (Pe.) tusimaefeum (Sasa); J—P. (Pe.) tusimafegeum (Sasa)." pageId="109" pageNumber="110">Fig. 15H</figureCitation>
) slender, 70 µm long; base with few microtrichia and 3 inner setae, projection with pointed apex, with dorsolateral seta. Inferior volsella 84 µm long, parallel-sided and slender, with prominent apical seta. Gonostylus 139 µm long. HR 1.18.
</paragraph>
</subSubSection>
<subSubSection id="97F3B9BAFFD8FF81FF43FC2389FDFBF0" box="[199,658,1000,1027]" pageId="109" pageNumber="110" type="distribution">
<paragraph id="DF56EA31FFD8FF81FF43FC2389FDFBF0" blockId="109.[151,1436,151,1027]" box="[199,658,1000,1027]" pageId="109" pageNumber="110">
<emphasis id="ED9D3623FFD8FF81FF43FC238A08FBF1" bold="true" box="[199,359,1000,1026]" pageId="109" pageNumber="110">Distribution.</emphasis>
Known only from
<collectingCountry id="A7FEAAA1FFD8FF81FDCCFC2289E0FBF0" box="[584,655,1001,1027]" name="Japan" pageId="109" pageNumber="110">Japan</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>