273 lines
40 KiB
XML
273 lines
40 KiB
XML
<document id="0BB1612C718943120C7504974CAF6209" ID-CLB-Dataset="187408" ID-DOI="10.1080/00222933.2022.2117107" ID-GBIF-Dataset="25e45e0a-f320-409b-8923-5ab29eabf122" ID-ISSN="1464-5262" ID-Zenodo-Dep="7318178" ID-ZooBank="A6F9B9FB-D3F9-49F2-8FC1-5E6A5E0AB3B1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1668419502508" checkinUser="plazi" docAuthor="Fachin, Diego Aguilar & Hauser, Martin" docDate="2022" docId="561187CDFFFE32609B824DBFFE2F109D" docLanguage="en" docName="JNATHIST.56.37-40.1513-1562.pdf" docOrigin="Journal of Natural History 56 (37 - 40)" docSource="http://dx.doi.org/10.1080/00222933.2022.2117107" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Formosargus berezovskiyi Fachin & Hauser 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="1552" masterDocId="AA28FFB5FFD932489A3F4D31FFA11374" masterDocTitle="Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym" masterLastPageNumber="1562" masterPageNumber="1513" pageNumber="1551" updateTime="1699411179050" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="61A8B2C8CB79E1BF0C3AAB4A84377631">Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym</mods:title>
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<mods:roleTerm id="EA3D0B47F037732C50C26911132D73A9">Author</mods:roleTerm>
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<mods:namePart id="BE064742A67371FB8A71CA427468C221">Fachin, Diego Aguilar</mods:namePart>
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<mods:name id="DE8F69DD443B7F6B63FE7830BCF3DB8C" type="personal">
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<mods:roleTerm id="8C5B26F7CDC354C21739E3EE4FCDA48D">Author</mods:roleTerm>
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<mods:namePart id="32132C1AF1D8A9AE91D6447E17B2248A">Hauser, Martin</mods:namePart>
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<mods:typeOfResource id="83C21206C5F4CEF1795FA93B11A4DCC0">text</mods:typeOfResource>
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<mods:title id="CAA67CBF6DF6BE40D93B2900F9D1E864">Journal of Natural History</mods:title>
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<mods:part id="5FB4FF865DB67D6145CED1C90278533F">
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<mods:date id="F902A9D9A99E34BE8C21028F102B5A8D">2022</mods:date>
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<mods:number id="261485652BE14790B346021469898043">2022-10-12</mods:number>
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<mods:number id="01331249F5D44BFA73486C86B7B1B670">56</mods:number>
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<mods:detail id="40312C94AD2170083CB14E0BD66A9EA6" type="issue">
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<mods:number id="0DBBEB6897D050DC7E16E95396177BE2">37 - 40</mods:number>
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<mods:url id="30B1200344CB1ED00FE06A04CBABA6F3">http://dx.doi.org/10.1080/00222933.2022.2117107</mods:url>
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<treatment id="561187CDFFFE32609B824DBFFE2F109D" ID-DOI="http://doi.org/10.5281/zenodo.7324776" ID-GBIF-Taxon="205007491" ID-Zenodo-Dep="7324776" LSID="urn:lsid:plazi:treatment:561187CDFFFE32609B824DBFFE2F109D" httpUri="http://treatment.plazi.org/id/561187CDFFFE32609B824DBFFE2F109D" lastPageId="40" lastPageNumber="1552" pageId="39" pageNumber="1551">
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<subSubSection id="96A26550FFFE326F9B824DBFFCC713DC" box="[445,870,142,168]" pageId="39" pageNumber="1551" type="nomenclature">
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<paragraph id="DE0736DBFFFE326F9B824DBFFCC713DC" blockId="39.[445,870,142,168]" box="[445,870,142,168]" pageId="39" pageNumber="1551">
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<heading id="854F81B7FFFE326F9B824DBFFCC713DC" bold="true" box="[445,870,142,168]" centered="true" fontSize="11" level="2" pageId="39" pageNumber="1551" reason="3">
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<emphasis id="ECCCEAC9FFFE326F9B824DBFFCC713DC" bold="true" box="[445,870,142,168]" pageId="39" pageNumber="1551">
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<taxonomicName id="19B84D58FFFE326F9B824DBFFD5D13DC" ID-CoL="BCY37" authority="Fachin & Hauser, 2022" authorityName="Fachin & Hauser" authorityYear="2022" box="[445,764,142,168]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="39" pageNumber="1551" phylum="Arthropoda" rank="species" species="berezovskiyi" status="sp. nov.">
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<emphasis id="ECCCEAC9FFFE326F9B824DBFFD5D13DC" bold="true" box="[445,764,142,168]" italics="true" pageId="39" pageNumber="1551">Formosargus berezovskiyi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="F7FF57B2FFFE326F993B4DBFFCC713DC" box="[772,870,142,168]" pageId="39" pageNumber="1551" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="96A26550FFFE326F9B644DE4FC68139B" box="[347,969,213,239]" pageId="39" pageNumber="1551" type="description">
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<paragraph id="DE0736DBFFFE326F9B644DE4FC68139B" blockId="39.[347,969,213,239]" box="[347,969,213,239]" pageId="39" pageNumber="1551">
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(
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<figureCitation id="46832A5EFFFE326F9B5D4DE4FE6F139B" box="[354,462,213,239]" captionStart="Figure 1" captionStartId="3.[160,225,645,668]" captionTargetBox="[347,969,144,610]" captionTargetId="figure-252@3.[344,972,142,613]" captionTargetPageId="3" captionText="Figure 1. Formosargus berezovskiyi sp. nov., live specimen, male, Taiwan, Orchid Island (© Yi-Xiang Lin (Gaga)). Available from: http://gaga.biodiv.tw/9501bx/536.htm." figureDoi="http://doi.org/10.5281/zenodo.7318182" httpUri="https://zenodo.org/record/7318182/files/figure.png" pageId="39" pageNumber="1551">Figures 1</figureCitation>
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,
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<figureCitation id="46832A5EFFFE326F9BE34DE4FE4A139B" box="[476,491,213,239]" captionStart="Figure 4" captionStartId="8.[160,225,1266,1289]" captionTargetBox="[167,1149,145,1232]" captionTargetId="figure-139@8.[163,1153,142,1234]" captionTargetPageId="8" captionText="Figure 4. Heads of Formosargus species, dorsal view. (a) F.berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.7318188" httpUri="https://zenodo.org/record/7318188/files/figure.png" pageId="39" pageNumber="1551">4</figureCitation>
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(a), 6(a), 7(e), 9(b), 15(c), 17(e–f), 19(b–c))
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</paragraph>
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</subSubSection>
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<subSubSection id="96A26550FFFE326F9A9F4C0DFEF21151" pageId="39" pageNumber="1551" type="diagnosis">
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<paragraph id="DE0736DBFFFE326F9A9F4C0DFEDC1222" blockId="39.[160,1156,316,549]" box="[160,381,316,342]" pageId="39" pageNumber="1551">
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<emphasis id="ECCCEAC9FFFE326F9A9F4C0DFEDC1222" bold="true" box="[160,381,316,342]" italics="true" pageId="39" pageNumber="1551">Diagnosis (female)</emphasis>
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</paragraph>
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<paragraph id="DE0736DBFFFE326F9A9F4C6EFEF21151" blockId="39.[160,1156,316,549]" pageId="39" pageNumber="1551">
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Dark brown pilosity on vertex area and ocellar tubercle (
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<figureCitation id="46832A5EFFFE326F99264C6EFCD4120C" box="[793,885,351,376]" captionStart="Figure 6" captionStartId="10.[160,225,1285,1308]" captionTargetBox="[167,1149,145,1251]" captionTargetId="figure-99@10.[163,1153,142,1253]" captionTargetPageId="10" captionText="Figure 6. Heads of Formosargus species, frontal view. (a) F. berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype. Arrow: anterior margin of the upper frons at the edge with lower frons." figureDoi="http://doi.org/10.5281/zenodo.7318193" httpUri="https://zenodo.org/record/7318193/files/figure.png" pageId="39" pageNumber="1551">Figure 6</figureCitation>
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(a)); upper frons as wide as long, longer than frontal callus and lower frons combined. Scutal medial band broad, roughly as wide as one-third of scutum (
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<figureCitation id="46832A5EFFFE326F985D4C95FD1C12C9" box="[610,701,420,445]" captionStart="Figure 7" captionStartId="12.[160,225,1575,1598]" captionTargetBox="[167,1149,145,1537]" captionTargetId="figure-3@12.[163,1153,142,1543]" captionTargetPageId="12" captionText="Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.7318195" httpUri="https://zenodo.org/record/7318195/files/figure.png" pageId="39" pageNumber="1551">Figure 7</figureCitation>
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(e)), only narrowing at level of transverse suture; mediotergite dark. Abdominal tergites marked with dark brown, a transverse proximal band on tergite 1 and subrectangular spots, narrowed medially, on tergites 2– 5 (
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<figureCitation id="46832A5EFFFE326F9A824F3DFE891151" box="[189,296,524,549]" captionStart="Figure 19" captionStartId="31.[160,225,1625,1648]" captionTargetBox="[168,1148,145,1587]" captionTargetId="figure-3@31.[163,1153,142,1594]" captionTargetPageId="31" captionText="Figure 19. Habitus of Formosargus species. (a) F. sagittocerus, male, lateral. (b–c) F. berezovskiyi sp. nov., female, paratype, dorsal and lateral, respectively. (d–e) F. borneensis sp. nov., female, holotype, dorsal and lateral, respectively. (f) F. mangoleensis sp. nov., female, holotype, dorsal." figureDoi="http://doi.org/10.5281/zenodo.7318226" httpUri="https://zenodo.org/record/7318226/files/figure.png" pageId="39" pageNumber="1551">Figure 19</figureCitation>
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(b)).
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="96A26550FFFE326F9A9F4F50FF6310B7" pageId="39" pageNumber="1551" type="materials_examined">
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||
<paragraph id="DE0736DBFFFE326F9A9F4F50FEDE110F" blockId="39.[160,1156,609,808]" box="[160,383,609,635]" pageId="39" pageNumber="1551">
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||
<emphasis id="ECCCEAC9FFFE326F9A9F4F50FEDE110F" bold="true" box="[160,383,609,635]" italics="true" pageId="39" pageNumber="1551">Material examined</emphasis>
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</paragraph>
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<paragraph id="DE0736DBFFFE326F9A9F4FB5FB25105C" blockId="39.[160,1156,609,808]" pageId="39" pageNumber="1551">
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||
<materialsCitation id="6ED03C86FFFE326F9A9F4FB5FC3A11B4" ID-GBIF-Occurrence="3965743307" collectingDate="1997-04-20" collectingDateMax="1997-05-01" collectingDateMin="1997-04-20" collectionCode="LACM" collectorName="V. Berezovskiy" country="Taiwan" latitude="22.0" location="Chungui Bridge" longLatPrecision="5579" longitude="121.57" municipality="Chungui Bridge" pageId="39" pageNumber="1551" specimenCount="1" specimenCount-female="1" stateProvince="Orchid Island" typeStatus="holotype">
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||
<emphasis id="ECCCEAC9FFFE326F9A9F4FB5FE8211E9" bold="true" box="[160,291,644,669]" pageId="39" pageNumber="1551">
|
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<typeStatus id="01038879FFFE326F9A9F4FB5FE8211E9" box="[160,291,644,669]" pageId="39" pageNumber="1551" type="holotype">HOLOTYPE</typeStatus>
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</emphasis>
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||
,
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||
<specimenCount id="C8BEFD52FFFE326F9B104FB5FEF611E8" box="[303,343,644,669]" pageId="39" pageNumber="1551" type="female">1 ♀</specimenCount>
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(with puparium pinned beneath label),
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||
<emphasis id="ECCCEAC9FFFE326F992A4FB5FCD911E9" bold="true" box="[789,888,644,669]" pageId="39" pageNumber="1551">
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<collectingCountry id="A6AF764BFFFE326F992A4FB5FCD911E9" box="[789,888,644,669]" name="Taiwan" pageId="39" pageNumber="1551">TAIWAN</collectingCountry>
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</emphasis>
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,
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||
<collectingRegion id="1C7CF839FFFE326F99BB4FB5FBBA11E9" box="[900,1051,644,669]" pageId="39" pageNumber="1551">Orchid Island</collectingRegion>
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,
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<location id="DB676000FFFE326F9E1B4FB5FF4A11B4" LSID="urn:lsid:plazi:treatment:561187CDFFFE32609B824DBFFE2F109D:DB676000FFFE326F9E1B4FB5FF4A11B4" country="Taiwan" latitude="22.0" longLatPrecision="5579" longitude="121.57" municipality="Chungui Bridge" name="Chungui Bridge" pageId="39" pageNumber="1551" stateProvince="Orchid Island">Chungui Bridge</location>
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,
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<geoCoordinate id="BB8C501CFFFE326F9ACA4F96FE9F11B4" box="[245,318,679,704]" degrees="22.0" direction="north" orientation="latitude" pageId="39" pageNumber="1551" precision="5555" value="22.0">22.0°N</geoCoordinate>
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<geoCoordinate id="BB8C501CFFFE326F9B7A4F96FE0611B4" box="[325,423,679,704]" degrees="121.57" direction="east" orientation="longitude" pageId="39" pageNumber="1551" precision="555" value="121.57">121.57°E</geoCoordinate>
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,
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<date id="AA06101BFFFE326F9B8E4F96FDFF11B4" box="[433,606,679,704]" pageId="39" pageNumber="1551" value="1997-04-20" valueMax="1997-05-01" valueMin="1997-04-20">
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<collectingDate id="BA42E9F3FFFE326F9B8E4F96FDFF11B4" box="[433,606,679,704]" pageId="39" pageNumber="1551" value="1997-04-20" valueMax="1997-05-01" valueMin="1997-04-20">20.iv–01.v.1997</collectingDate>
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</date>
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||
,
|
||
<collectorName id="734D530DFFFE326F98564F96FCA811B4" box="[617,777,679,704]" pageId="39" pageNumber="1551">V. Berezovskiy</collectorName>
|
||
, coll. (
|
||
<collectionCode id="B8A9AE1EFFFE326F99724F96FC3411B4" box="[845,917,679,704]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="39" pageNumber="1551" type="Museum">LACM</collectionCode>
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)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="6ED03C86FFFE326F999B4F96FB25105C" ID-GBIF-Occurrence="3965743315" collectingDate="1997-04-20" collectingDateMax="1997-05-01" collectingDateMin="1997-04-20" collectionCode="LACM" collectorName="V. Berezovskiy" country="Taiwan" latitude="22.0" location="Orchid Island" longLatPrecision="5579" longitude="121.57" municipality="Chungui Bridge" pageId="39" pageNumber="1551" specimenCount="3" specimenCount-female="3" stateProvince="Orchid Island" typeStatus="paratype">
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<emphasis id="ECCCEAC9FFFE326F999B4F96FB9011B4" bold="true" box="[932,1073,679,704]" pageId="39" pageNumber="1551">
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<typeStatus id="01038879FFFE326F999B4F96FB9011B4" box="[932,1073,679,704]" pageId="39" pageNumber="1551" type="paratype">PARATYPES</typeStatus>
|
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</emphasis>
|
||
,
|
||
<specimenCount id="C8BEFD52FFFE326F9E014F96FBC611CA" box="[1086,1127,679,704]" pageId="39" pageNumber="1551" type="female">3 ♀</specimenCount>
|
||
(1 ♀ with left wing slide-mounted, genitalia dissected and stored in microvial with glycerine) (each puparium pinned beneath label), same data as
|
||
<typeStatus id="01038879FFFE326F98CD4FDDFCF71071" box="[754,854,748,773]" pageId="39" pageNumber="1551" type="holotype">holotype</typeStatus>
|
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(
|
||
<collectionCode id="B8A9AE1EFFFE326F995C4FDDFC0A1071" box="[867,939,748,773]" collectionName="USA, California, Los Angeles, Los Angeles County Museum of Natural History" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="39" pageNumber="1551" type="Museum">LACM</collectionCode>
|
||
). Two females have an additional label mentioning that each adult emerged from the puparium on
|
||
<date id="AA06101BFFFE326F9E2B4E3EFB21105C" box="[1044,1152,783,808]" pageId="39" pageNumber="1551" value="1997-05-21">21.v.1997</date>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="DE0736DBFFFE326F9A9F4E55FF6310B7" blockId="39.[160,1155,868,963]" pageId="39" pageNumber="1551">
|
||
<emphasis id="ECCCEAC9FFFE326F9A9F4E55FE2F100A" bold="true" box="[160,398,868,894]" italics="true" pageId="39" pageNumber="1551">Additional material.</emphasis>
|
||
<materialsCitation id="6ED03C86FFFE326F9BA24E55FF1E10B7" ID-GBIF-Occurrence="3965743312" country="Taiwan" latitude="-22.037306" location="Lanyu Weather Station" longLatPrecision="1" longitude="121.558334" municipality="Lanyu Weather Station" pageId="39" pageNumber="1551" specimenCount="1" specimenCount-male="1" stateProvince="Orchid Island">
|
||
<specimenCount id="C8BEFD52FFFE326F9BA24E55FE671008" box="[413,454,868,893]" pageId="39" pageNumber="1551" type="male">1 ♂</specimenCount>
|
||
,
|
||
<emphasis id="ECCCEAC9FFFE326F9BEC4E55FD971009" bold="true" box="[467,566,868,893]" pageId="39" pageNumber="1551">
|
||
<collectingCountry id="A6AF764BFFFE326F9BEC4E55FD971009" box="[467,566,868,893]" name="Taiwan" pageId="39" pageNumber="1551">TAIWAN</collectingCountry>
|
||
</emphasis>
|
||
,
|
||
<collectingRegion id="1C7CF839FFFE326F987B4E55FD7A1009" box="[580,731,868,893]" pageId="39" pageNumber="1551">Orchid Island</collectingRegion>
|
||
,
|
||
<location id="DB676000FFFE326F98DA4E55FC491009" LSID="urn:lsid:plazi:treatment:561187CDFFFE32609B824DBFFE2F109D:DB676000FFFE326F98DA4E55FC491009" box="[741,1000,868,893]" country="Taiwan" latitude="-22.037306" longLatPrecision="1" longitude="121.558334" municipality="Lanyu Weather Station" name="Lanyu Weather Station" pageId="39" pageNumber="1551" stateProvince="Orchid Island">Lanyu Weather Station</location>
|
||
, -
|
||
<geoCoordinate id="BB8C501CFFFE326F99C74E55FB221009" box="[1016,1155,868,893]" degrees="22.037306" direction="south" orientation="latitude" pageId="39" pageNumber="1551" precision="1" value="-22.037306">22.037306°S</geoCoordinate>
|
||
<geoCoordinate id="BB8C501CFFFE326F9A9F4EB6FE9710D4" box="[160,310,903,928]" degrees="121.558333" direction="east" orientation="longitude" pageId="39" pageNumber="1551" precision="1" value="121.558334">121.558333°E</geoCoordinate>
|
||
[Note: male specimen not collected; only known from photograph (
|
||
<figureCitation id="46832A5EFFFE326F9E034EB6FF0F10B7" captionStart="Figure 1" captionStartId="3.[160,225,645,668]" captionTargetBox="[347,969,144,610]" captionTargetId="figure-252@3.[344,972,142,613]" captionTargetPageId="3" captionText="Figure 1. Formosargus berezovskiyi sp. nov., live specimen, male, Taiwan, Orchid Island (© Yi-Xiang Lin (Gaga)). Available from: http://gaga.biodiv.tw/9501bx/536.htm." figureDoi="http://doi.org/10.5281/zenodo.7318182" httpUri="https://zenodo.org/record/7318182/files/figure.png" pageId="39" pageNumber="1551">Figure 1</figureCitation>
|
||
)]
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="96A26550FFFE32609A9F4ECFFDDE13BF" lastPageId="40" lastPageNumber="1552" pageId="39" pageNumber="1551" type="description">
|
||
<paragraph id="DE0736DBFFFE326F9A9F4ECFFE87176C" blockId="39.[160,1156,1022,1707]" box="[160,294,1022,1048]" pageId="39" pageNumber="1551">
|
||
<emphasis id="ECCCEAC9FFFE326F9A9F4ECFFE87176C" bold="true" box="[160,294,1022,1048]" italics="true" pageId="39" pageNumber="1551">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="DE0736DBFFFE326F9A9F4913FEA515DF" blockId="39.[160,1156,1022,1707]" pageId="39" pageNumber="1551">
|
||
<emphasis id="ECCCEAC9FFFE326F9A9F4913FF56174F" bold="true" box="[160,247,1058,1083]" pageId="39" pageNumber="1551">Female</emphasis>
|
||
. Length (n = 4): body mm, 5.0–6.0; wing mm, 5.0.
|
||
<emphasis id="ECCCEAC9FFFE326F991B4913FCC3174F" bold="true" box="[804,866,1058,1083]" pageId="39" pageNumber="1551">Head</emphasis>
|
||
(
|
||
<figureCitation id="46832A5EFFFE326F994E4913FC79174F" box="[881,984,1058,1083]" captionStart="Figure 4" captionStartId="8.[160,225,1266,1289]" captionTargetBox="[167,1149,145,1232]" captionTargetId="figure-139@8.[163,1153,142,1234]" captionTargetPageId="8" captionText="Figure 4. Heads of Formosargus species, dorsal view. (a) F.berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.7318188" httpUri="https://zenodo.org/record/7318188/files/figure.png" pageId="39" pageNumber="1551">Figures 4</figureCitation>
|
||
(a), 6(a)). Vertex area as long as ocellar tubercle, with medium dark brown pilosity. Upper frons as wide as long, slightly longer than lower frons; black lateral area of upper frons nearly as wide as ocellar tubercle width, with short dark brown pilosity; anterior margin of upper frons with median triangular extension slightly pointing towards antennal insertion. Fourth flagellomere rounded, as wide as each previous flagellomere. Face slightly shorter than lower frons.
|
||
<emphasis id="ECCCEAC9FFFE326F9ADB49C3FE99167F" bold="true" box="[228,312,1266,1291]" pageId="39" pageNumber="1551">Thorax</emphasis>
|
||
(
|
||
<figureCitation id="46832A5EFFFE326F9B7949C3FE00167F" box="[326,417,1266,1291]" captionStart="Figure 7" captionStartId="12.[160,225,1575,1598]" captionTargetBox="[167,1149,145,1537]" captionTargetId="figure-3@12.[163,1153,142,1543]" captionTargetPageId="12" captionText="Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.7318195" httpUri="https://zenodo.org/record/7318195/files/figure.png" pageId="39" pageNumber="1551">Figure 7</figureCitation>
|
||
(e)). Medial dark brown band over scutum broad, nearly as wide as one-third of scutum width, homogeneous in width along its entire extension, except for a narrowing at level of transverse suture; pleuron mostly yellowish, with only a small dark brown macula on anepisternum, mediotergite dark brown.
|
||
<emphasis id="ECCCEAC9FFFE326F9958486BFC071607" bold="true" box="[871,934,1370,1395]" pageId="39" pageNumber="1551">Wing</emphasis>
|
||
(
|
||
<figureCitation id="46832A5EFFFE326F9986486BFBBC1607" box="[953,1053,1370,1395]" captionStart="Figure 9" captionStartId="17.[160,225,1285,1308]" captionTargetBox="[171,1149,145,1249]" captionTargetId="figure-3@17.[163,1153,142,1253]" captionTargetPageId="17" captionText="Figure 9. Wings of Formosargus species. (a) F. lineatus, female. (b) F. berezovskiyi sp. nov., female, paratype. (c) F. woodleyi sp. nov., male, holotype. Abbreviations: A1, first branch of anal vein; al, alula; an lb, anal lobe; bm, basal medial cell; br, basal radial cell; C, costal vein; c, costal cell; Cu, cubital vein or cubitus; CuA, anterior branch of cubital vein; cua, anterior cubital cell; CuA+CuP, anterior branch of cubital vein and posterior branch of cubital vein; CuP, posterior branch of cubital vein; cup, posterior cubital cell; d, discal cell; h, humeral crossvein; M, medial vein or media; M1, first branch of media; m1, first medial cell; M1+2, fused first and second branch of media; M2, second branch of media; m2+3, fused second and third cell of media;M3, third branch of media; M3+4, fused third and fourth branch of media; M4, fourth branch of media; m4, fourth medial cell; m–cu, medial–cubital crossvein; m–m, medial crossvein; R1, anterior branch of radius; r1, first radial cell; R2+3, second branch of radius; r2+3, second and third radial cell; R4, upper branch of third branch of radius; r4, fourth radial cell; R5, lower branch of third branch of radius; r5, fifth radial cell; r–m, radial–medial crossvein; Rs, radial sector; Sc, subcostal vein sc, subcostal cell." figureDoi="http://doi.org/10.5281/zenodo.7318199" httpUri="https://zenodo.org/record/7318199/files/figure.png" pageId="39" pageNumber="1551">Figure 9</figureCitation>
|
||
(b)). R2+3 originating at r–m level, weakly curved on basal third, nearly straight on apical two-thirds.
|
||
<emphasis id="ECCCEAC9FFFE326F9A9F48AEFEB416CC" bold="true" box="[160,277,1439,1464]" pageId="39" pageNumber="1551">Abdomen</emphasis>
|
||
. Tergite 1 with a dark brown transverse band proximally; tergites 2–5 with a pair of dark brown subrectangular spots; only spots of tergite 4 connected medially.
|
||
<emphasis id="ECCCEAC9FFFE326F9A9F48D5FEBE1689" bold="true" box="[160,287,1508,1533]" pageId="39" pageNumber="1551">Terminalia</emphasis>
|
||
(
|
||
<figureCitation id="46832A5EFFFE326F9B1048D5FE061689" box="[303,423,1508,1533]" captionStart="Figure 15" captionStartId="27.[160,225,1701,1724]" captionTargetBox="[210,1104,144,1666]" captionTargetId="figure-1@27.[208,1107,142,1669]" captionTargetPageId="27" captionText="Figure 15. Female terminalia of Formosargus species, dorsal view. (a) F. kerteszi. (b) F. lineatus. (c) F. berezovskiyi sp. nov., paratype. (d) F. borneensis sp. nov., holotype (cerci lost). Abbreviations: cerc, cercus; st, sternite; tg, tergite." figureDoi="http://doi.org/10.5281/zenodo.7318213" httpUri="https://zenodo.org/record/7318213/files/figure.png" pageId="39" pageNumber="1551">Figures 15</figureCitation>
|
||
(c), 17(e–f)). Genital fork equally wide on distal two-thirds, with a subtle widening at level of posterolateral process insertion; posterior, lateral and anterior margins projecting backwards; basal one-third abruptly narrowing into a wide and rounded anterior end; posterior bridge bilobed; posterolateral process wider on basal one-third, narrowing from midway to apex, nearly parallel on distal half; genital opening elliptical.
|
||
</paragraph>
|
||
<paragraph id="DE0736DBFFF132609A9F4DBFFF7B13DC" blockId="40.[160,639,142,203]" box="[160,218,142,168]" pageId="40" pageNumber="1552">
|
||
<emphasis id="ECCCEAC9FFF132609A9F4DBFFF7B13DC" bold="true" box="[160,218,142,168]" italics="true" pageId="40" pageNumber="1552">Male</emphasis>
|
||
</paragraph>
|
||
<paragraph id="DE0736DBFFF132609A9F4D83FDDE13BF" blockId="40.[160,639,142,203]" box="[160,639,178,203]" pageId="40" pageNumber="1552">
|
||
Only known from a photograph (
|
||
<figureCitation id="46832A5EFFF1326098294D83FDD313BF" box="[534,626,178,203]" captionStart="Figure 1" captionStartId="3.[160,225,645,668]" captionTargetBox="[347,969,144,610]" captionTargetId="figure-252@3.[344,972,142,613]" captionTargetPageId="3" captionText="Figure 1. Formosargus berezovskiyi sp. nov., live specimen, male, Taiwan, Orchid Island (© Yi-Xiang Lin (Gaga)). Available from: http://gaga.biodiv.tw/9501bx/536.htm." figureDoi="http://doi.org/10.5281/zenodo.7318182" httpUri="https://zenodo.org/record/7318182/files/figure.png" pageId="40" pageNumber="1552">Figure 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="96A26550FFF132609A9F4DC7FE9212EF" pageId="40" pageNumber="1552" type="etymology">
|
||
<paragraph id="DE0736DBFFF132609A9F4DC7FEBC1264" blockId="40.[160,1155,246,411]" box="[160,285,246,272]" pageId="40" pageNumber="1552">
|
||
<emphasis id="ECCCEAC9FFF132609A9F4DC7FEBC1264" bold="true" box="[160,285,246,272]" italics="true" pageId="40" pageNumber="1552">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="DE0736DBFFF132609A9F4C2BFE9212EF" blockId="40.[160,1155,246,411]" pageId="40" pageNumber="1552">
|
||
The specific epithet is a patronym dedicated to Vladimir V. Berezovskiy, a senior hymenopterist technician at the Entomology Research Museum in Riverside,
|
||
<collectingCountry id="A6AF764BFFF1326099E04C0CFBB01222" box="[991,1041,317,342]" name="United States of America" pageId="40" pageNumber="1552">USA</collectingCountry>
|
||
, who first found the larva and identified the specimens in the LACM collection as belonging to
|
||
<taxonomicName id="19B84D58FFF132609A9F4CB3FE8C12EF" baseAuthorityName="James" baseAuthorityYear="1939" box="[160,301,386,411]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="40" pageNumber="1552" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="ECCCEAC9FFF132609A9F4CB3FE8C12EF" box="[160,301,386,411]" italics="true" pageId="40" pageNumber="1552">Formosargus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="96A26550FFF132609A9F4CF7FD4B1177" pageId="40" pageNumber="1552" type="distribution">
|
||
<paragraph id="DE0736DBFFF132609A9F4CF7FE181294" blockId="40.[160,746,454,515]" box="[160,441,454,480]" pageId="40" pageNumber="1552">
|
||
<emphasis id="ECCCEAC9FFF132609A9F4CF7FE181294" bold="true" box="[160,441,454,480]" italics="true" pageId="40" pageNumber="1552">Geographic distribution</emphasis>
|
||
</paragraph>
|
||
<paragraph id="DE0736DBFFF132609A9F4CDBFD4B1177" blockId="40.[160,746,454,515]" box="[160,746,490,515]" pageId="40" pageNumber="1552">
|
||
<emphasis id="ECCCEAC9FFF132609A9F4CDBFEFC1177" bold="true" box="[160,349,490,515]" pageId="40" pageNumber="1552">Oriental Region</emphasis>
|
||
.
|
||
<collectingCountry id="A6AF764BFFF132609B554CDBFE1B1177" box="[362,442,490,515]" name="Taiwan" pageId="40" pageNumber="1552">Taiwan</collectingCountry>
|
||
(Orchid Island) (
|
||
<figureCitation id="46832A5EFFF13260984B4CDBFD7C1177" box="[628,733,490,515]" captionStart="Figure 26" captionStartId="38.[160,226,870,893]" captionTargetBox="[168,1148,146,834]" captionTargetId="figure-167@38.[163,1153,142,839]" captionTargetPageId="38" captionText="Figure 26. Geographic distribution of Formosargus. Squares represent holotypes records; circles represent paratypes, and additional material. Three of the ‘lines’ of faunal division are shown: 1. Wallace’s Line; 2. Weber’s Line; 3. Lydekker’s Line. ‘Wallacea’ is the area that lies between Wallace’s Line and Lydekker’s Line." figureDoi="http://doi.org/10.5281/zenodo.7318240" httpUri="https://zenodo.org/record/7318240/files/figure.png" pageId="40" pageNumber="1552">Figure 26</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="96A26550FFF132609A9F4F1EFE2F109D" pageId="40" pageNumber="1552" type="discussion">
|
||
<paragraph id="DE0736DBFFF132609A9F4F1EFEBF113D" blockId="40.[160,1156,559,1001]" box="[160,286,559,585]" pageId="40" pageNumber="1552">
|
||
<emphasis id="ECCCEAC9FFF132609A9F4F1EFEBF113D" bold="true" box="[160,286,559,585]" italics="true" pageId="40" pageNumber="1552">Comments</emphasis>
|
||
</paragraph>
|
||
<paragraph id="DE0736DBFFF132609A9F4F63FE2F109D" blockId="40.[160,1156,559,1001]" pageId="40" pageNumber="1552">
|
||
This new species is the second species from
|
||
<collectingCountry id="A6AF764BFFF1326098B04F63FD40111F" box="[655,737,594,619]" name="Taiwan" pageId="40" pageNumber="1552">Taiwan</collectingCountry>
|
||
, but it is only found on Orchid Island, while
|
||
<taxonomicName id="19B84D58FFF132609ADB4F44FE6911FA" authorityName="James" authorityYear="1939" box="[228,456,629,654]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="40" pageNumber="1552" phylum="Arthropoda" rank="species" species="kerteszi">
|
||
<emphasis id="ECCCEAC9FFF132609ADB4F44FE6911FA" box="[228,456,629,654]" italics="true" pageId="40" pageNumber="1552">Formosargus kerteszi</emphasis>
|
||
</taxonomicName>
|
||
is reported from the southern tip of the main island, which is the
|
||
<typeStatus id="01038879FFF132609AF44FA9FF5C11C5" box="[203,253,664,689]" pageId="40" pageNumber="1552">type</typeStatus>
|
||
locality, and now also in the
|
||
<collectingCountry id="A6AF764BFFF1326098044FA9FD1711C5" box="[571,694,664,689]" name="Philippines" pageId="40" pageNumber="1552">Philippines</collectingCountry>
|
||
(
|
||
<figureCitation id="46832A5EFFF1326098FC4FA9FC8A11C5" box="[707,811,664,689]" captionStart="Figure 26" captionStartId="38.[160,226,870,893]" captionTargetBox="[168,1148,146,834]" captionTargetId="figure-167@38.[163,1153,142,839]" captionTargetPageId="38" captionText="Figure 26. Geographic distribution of Formosargus. Squares represent holotypes records; circles represent paratypes, and additional material. Three of the ‘lines’ of faunal division are shown: 1. Wallace’s Line; 2. Weber’s Line; 3. Lydekker’s Line. ‘Wallacea’ is the area that lies between Wallace’s Line and Lydekker’s Line." figureDoi="http://doi.org/10.5281/zenodo.7318240" httpUri="https://zenodo.org/record/7318240/files/figure.png" pageId="40" pageNumber="1552">Figure 26</figureCitation>
|
||
). The male of
|
||
<taxonomicName id="19B84D58FFF1326099FA4FA9FBFF11C5" authority="Fachin & Hauser, 2022" authorityName="Fachin & Hauser" authorityYear="2022" box="[965,1118,664,689]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="40" pageNumber="1552" phylum="Arthropoda" rank="species" species="berezovskiyi" status="sp. nov.">
|
||
<emphasis id="ECCCEAC9FFF1326099FA4FA9FBFF11C5" box="[965,1118,664,689]" italics="true" pageId="40" pageNumber="1552">F. berezovskiyi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F7FF57B2FFF132609E5B4FA9FF7211A7" pageId="40" pageNumber="1552" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, only known from a photograph (
|
||
<figureCitation id="46832A5EFFF1326098774F8BFD0511A7" box="[584,676,698,723]" captionStart="Figure 1" captionStartId="3.[160,225,645,668]" captionTargetBox="[347,969,144,610]" captionTargetId="figure-252@3.[344,972,142,613]" captionTargetPageId="3" captionText="Figure 1. Formosargus berezovskiyi sp. nov., live specimen, male, Taiwan, Orchid Island (© Yi-Xiang Lin (Gaga)). Available from: http://gaga.biodiv.tw/9501bx/536.htm." figureDoi="http://doi.org/10.5281/zenodo.7318182" httpUri="https://zenodo.org/record/7318182/files/figure.png" pageId="40" pageNumber="1552">Figure 1</figureCitation>
|
||
), is very similar in the colour pattern to the male of
|
||
<taxonomicName id="19B84D58FFF132609AC04FECFECB1182" authorityName="James" authorityYear="1939" box="[255,362,733,758]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="40" pageNumber="1552" phylum="Arthropoda" rank="species" species="kerteszi">
|
||
<emphasis id="ECCCEAC9FFF132609AC04FECFECB1182" box="[255,362,733,758]" italics="true" pageId="40" pageNumber="1552">F. kerteszi</emphasis>
|
||
</taxonomicName>
|
||
, with a single medial scutal band, but one important difference, also present in the females, is the shape and width of the band. In
|
||
<taxonomicName id="19B84D58FFF1326099634E31FC59106D" authority="Fachin & Hauser, 2022" authorityName="Fachin & Hauser" authorityYear="2022" box="[860,1016,768,793]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="40" pageNumber="1552" phylum="Arthropoda" rank="species" species="berezovskiyi" status="sp. nov.">
|
||
<emphasis id="ECCCEAC9FFF1326099634E31FC59106D" box="[860,1016,768,793]" italics="true" pageId="40" pageNumber="1552">F. berezovskiyi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F7FF57B2FFF1326099C04E31FBF7106D" box="[1023,1110,768,793]" pageId="40" pageNumber="1552" rank="species">sp. nov.</taxonomicNameLabel>
|
||
it is clearly narrowed at the level of the transverse suture and wider than in other species, as much as one-third the width of the scutum (
|
||
<figureCitation id="46832A5EFFF1326098AE4E74FD56102A" box="[657,759,837,862]" captionStart="Figure 1" captionStartId="3.[160,225,645,668]" captionTargetBox="[347,969,144,610]" captionTargetId="figure-252@3.[344,972,142,613]" captionTargetPageId="3" captionText="Figure 1. Formosargus berezovskiyi sp. nov., live specimen, male, Taiwan, Orchid Island (© Yi-Xiang Lin (Gaga)). Available from: http://gaga.biodiv.tw/9501bx/536.htm." figureDoi="http://doi.org/10.5281/zenodo.7318182" httpUri="https://zenodo.org/record/7318182/files/figure.png" pageId="40" pageNumber="1552">Figures 1</figureCitation>
|
||
,
|
||
<figureCitation id="46832A5EFFF13260993B4E74FCB3102A" box="[772,786,837,862]" captionStart="Figure 7" captionStartId="12.[160,225,1575,1598]" captionTargetBox="[167,1149,145,1537]" captionTargetId="figure-3@12.[163,1153,142,1543]" captionTargetPageId="12" captionText="Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.7318195" httpUri="https://zenodo.org/record/7318195/files/figure.png" pageId="40" pageNumber="1552">7</figureCitation>
|
||
(e)), while in
|
||
<taxonomicName id="19B84D58FFF13260999E4E74FBAB102A" authorityName="James" authorityYear="1939" box="[929,1034,837,862]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="40" pageNumber="1552" phylum="Arthropoda" rank="species" species="kerteszi">
|
||
<emphasis id="ECCCEAC9FFF13260999E4E74FBAB102A" box="[929,1034,837,862]" italics="true" pageId="40" pageNumber="1552">F. kerteszi</emphasis>
|
||
</taxonomicName>
|
||
, the scutal band is more linear and thinner (
|
||
<figureCitation id="46832A5EFFF1326098154E59FD3410F5" box="[554,661,872,897]" captionStart="Figure 7" captionStartId="12.[160,225,1575,1598]" captionTargetBox="[167,1149,145,1537]" captionTargetId="figure-3@12.[163,1153,142,1543]" captionTargetPageId="12" captionText="Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.7318195" httpUri="https://zenodo.org/record/7318195/files/figure.png" pageId="40" pageNumber="1552">Figures 7</figureCitation>
|
||
(a,b)). Besides these features, females of
|
||
<taxonomicName id="19B84D58FFF132609E4E4E59FE8310D7" authority="Fachin & Hauser, 2022" authorityName="Fachin & Hauser" authorityYear="2022" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="40" pageNumber="1552" phylum="Arthropoda" rank="species" species="berezovskiyi" status="sp. nov.">
|
||
<emphasis id="ECCCEAC9FFF132609E4E4E59FE8310D7" italics="true" pageId="40" pageNumber="1552">F. berezovskiyi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F7FF57B2FFF132609B114EBBFE2810D7" box="[302,393,906,931]" pageId="40" pageNumber="1552" rank="species">sp. nov.</taxonomicNameLabel>
|
||
can be separated from all the other one-striped females by the presence of a black mediotergite and abdominal tergites strongly marked with dark brown (
|
||
<figureCitation id="46832A5EFFF132609AC74EE1FEC2109D" box="[248,355,976,1001]" captionStart="Figure 19" captionStartId="31.[160,225,1625,1648]" captionTargetBox="[168,1148,145,1587]" captionTargetId="figure-3@31.[163,1153,142,1594]" captionTargetPageId="31" captionText="Figure 19. Habitus of Formosargus species. (a) F. sagittocerus, male, lateral. (b–c) F. berezovskiyi sp. nov., female, paratype, dorsal and lateral, respectively. (d–e) F. borneensis sp. nov., female, holotype, dorsal and lateral, respectively. (f) F. mangoleensis sp. nov., female, holotype, dorsal." figureDoi="http://doi.org/10.5281/zenodo.7318226" httpUri="https://zenodo.org/record/7318226/files/figure.png" pageId="40" pageNumber="1552">Figure 19</figureCitation>
|
||
(b)).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |