243 lines
35 KiB
XML
243 lines
35 KiB
XML
<document id="D9ABA26FB164E09EF41E47FD530188E9" ID-CLB-Dataset="187408" ID-DOI="10.1080/00222933.2022.2117107" ID-GBIF-Dataset="25e45e0a-f320-409b-8923-5ab29eabf122" ID-ISSN="1464-5262" ID-Zenodo-Dep="7318178" ID-ZooBank="A6F9B9FB-D3F9-49F2-8FC1-5E6A5E0AB3B1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1668419502508" checkinUser="plazi" docAuthor="Fachin, Diego Aguilar & Hauser, Martin" docDate="2022" docId="561187CDFFF132619BF84917FDEE1501" docLanguage="en" docName="JNATHIST.56.37-40.1513-1562.pdf" docOrigin="Journal of Natural History 56 (37 - 40)" docSource="http://dx.doi.org/10.1080/00222933.2022.2117107" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Formosargus borneensis Fachin & Hauser 2022, sp. nov." docType="treatment" docUuid="D4B99C35-6994-422E-9EEB-18D3C56E4CB6" docUuidSource="ZooBank" docVersion="7" lastPageNumber="1553" masterDocId="AA28FFB5FFD932489A3F4D31FFA11374" masterDocTitle="Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym" masterLastPageNumber="1562" masterPageNumber="1513" pageNumber="1552" updateTime="1699411179050" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="AD4583D5C3FEA6EC03390FD97DED5314">Revision of Formosargus James, 1939 (Diptera: Stratiomyidae): five new species in an otherwise rare genus and Amsaria Adisoemarto, 1974 as a generic synonym</mods:title>
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<mods:name id="6201F6CFB6CEC97BFF9A85EC90DE8FA2" type="personal">
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<mods:roleTerm id="629C735EFF410CBBAAB18EE92E88652A">Author</mods:roleTerm>
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<mods:namePart id="777B2D1D379829C6D93464D8E051A28D">Fachin, Diego Aguilar</mods:namePart>
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<mods:name id="CC0F83B66D55E260AC7945A63C2D8C87" type="personal">
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<mods:roleTerm id="5ED7F49889CE4FFC0B6E61E881BCA1E2">Author</mods:roleTerm>
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<mods:namePart id="9799EB3DE4253CE7108369D24F0623C4">Hauser, Martin</mods:namePart>
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<mods:typeOfResource id="6702307E0CEBBA26398E0F8EAC54D66F">text</mods:typeOfResource>
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<mods:title id="D3F2EAB19365C9C6E38C9AFE190D2CF5">Journal of Natural History</mods:title>
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</mods:titleInfo>
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<mods:part id="5C199AAD9B68BBB2B0BF923EA3C01A2B">
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<mods:date id="0D8922948A7EA171F2294B0064F6AC3F">2022</mods:date>
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<mods:number id="24A9111F1712A20B01972202CE60E170">2022-10-12</mods:number>
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<mods:detail id="836EF88DC414231A05E3D4C2CE953851" type="volume">
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<mods:number id="E48B572295E49643C95158E6E5A20374">56</mods:number>
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<mods:detail id="4E3AF47DF1CC5F9E3BC93EDB6702FC6C" type="issue">
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<mods:number id="633283AB59301E7D746E27E6C35C952D">37 - 40</mods:number>
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<mods:start id="DFA69D291318E424AF74D6729F663E4A">1513</mods:start>
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<mods:end id="94E453FE76E27719725B606F419D9101">1562</mods:end>
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<mods:url id="6EF686F8B0DA73E79366A0083ED5F6B5">http://dx.doi.org/10.1080/00222933.2022.2117107</mods:url>
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<mods:classification id="940CDFE5975654EB60642F695EE1A7BF">journal article</mods:classification>
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<mods:identifier id="E3E8D3D24B932784B0BE535F5EFB20F4" type="CLB-Dataset">187408</mods:identifier>
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<mods:identifier id="722B384313296F00750B60468F76EDA9" type="DOI">10.1080/00222933.2022.2117107</mods:identifier>
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<treatment id="561187CDFFF132619BF84917FDEE1501" ID-DOI="http://doi.org/10.5281/zenodo.7324778" ID-GBIF-Taxon="205007494" ID-Zenodo-Dep="7324778" LSID="urn:lsid:zoobank.org:act:D4B99C35-6994-422E-9EEB-18D3C56E4CB6" httpUri="http://treatment.plazi.org/id/561187CDFFF132619BF84917FDEE1501" lastPageId="41" lastPageNumber="1553" pageId="40" pageNumber="1552">
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<subSubSection id="96A26550FFF132609BF84917FCFD1734" box="[455,860,1062,1088]" pageId="40" pageNumber="1552" type="nomenclature">
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<paragraph id="DE0736DBFFF132609BF84917FCFD1734" blockId="40.[384,931,1062,1121]" box="[455,860,1062,1088]" pageId="40" pageNumber="1552">
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<heading id="854F81B7FFF132609BF84917FCFD1734" box="[455,860,1062,1088]" centered="true" fontSize="11" level="2" pageId="40" pageNumber="1552" reason="2">
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<emphasis id="ECCCEAC9FFF132609BF84917FCFD1734" bold="true" box="[455,860,1062,1088]" pageId="40" pageNumber="1552">
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<taxonomicName id="19B84D58FFF132609BF84917FD531734" ID-CoL="BCY38" authority="Fachin & Hauser, 2022" authorityName="Fachin & Hauser" authorityYear="2022" box="[455,754,1062,1088]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="40" pageNumber="1552" phylum="Arthropoda" rank="species" species="borneensis" status="sp. nov.">
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<emphasis id="ECCCEAC9FFF132609BF84917FD531734" bold="true" box="[455,754,1062,1088]" italics="true" pageId="40" pageNumber="1552">Formosargus borneensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="F7FF57B2FFF1326098C54917FCFD1734" box="[762,860,1062,1088]" pageId="40" pageNumber="1552" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="96A26550FFF132609BBF4976FC021715" box="[384,931,1095,1121]" pageId="40" pageNumber="1552" type="description">
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<paragraph id="DE0736DBFFF132609BBF4976FC021715" blockId="40.[384,931,1062,1121]" box="[384,931,1095,1121]" pageId="40" pageNumber="1552">
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(
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<figureCitation id="46832A5EFFF132609BB74976FE571715" box="[392,502,1095,1121]" captionStart="Figure 4" captionStartId="8.[160,225,1266,1289]" captionTargetBox="[167,1149,145,1232]" captionTargetId="figure-139@8.[163,1153,142,1234]" captionTargetPageId="8" captionText="Figure 4. Heads of Formosargus species, dorsal view. (a) F.berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.7318188" httpUri="https://zenodo.org/record/7318188/files/figure.png" pageId="40" pageNumber="1552">Figures 4</figureCitation>
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(b), 6(b), 7(f), 15(d), 17(g–h), 19(d–e))
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</paragraph>
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</subSubSection>
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<subSubSection id="96A26550FFF132609A9F49AFFE3416FC" pageId="40" pageNumber="1552" type="diagnosis">
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<paragraph id="DE0736DBFFF132609A9F49AFFEDC17CC" blockId="40.[160,1156,1182,1416]" box="[160,381,1182,1208]" pageId="40" pageNumber="1552">
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<emphasis id="ECCCEAC9FFF132609A9F49AFFEDC17CC" bold="true" box="[160,381,1182,1208]" italics="true" pageId="40" pageNumber="1552">Diagnosis (female)</emphasis>
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</paragraph>
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<paragraph id="DE0736DBFFF132609A9F49F0FE3416FC" blockId="40.[160,1156,1182,1416]" pageId="40" pageNumber="1552">
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Species distinctly reddish in colour. Medial band on scutum slightly wider on posterior half (
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<figureCitation id="46832A5EFFF132609B3649D5FECD1789" box="[265,364,1252,1277]" captionStart="Figure 7" captionStartId="12.[160,225,1575,1598]" captionTargetBox="[167,1149,145,1537]" captionTargetId="figure-3@12.[163,1153,142,1543]" captionTargetPageId="12" captionText="Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.7318195" httpUri="https://zenodo.org/record/7318195/files/figure.png" pageId="40" pageNumber="1552">Figure 7</figureCitation>
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(f)), with a narrowing at level of the transverse suture and yellow one-third of scutum as wide as in
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<taxonomicName id="19B84D58FFF13260980D4836FD721654" authority="Fachin & Hauser, 2022" authorityName="Fachin & Hauser" authorityYear="2022" box="[562,723,1287,1312]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="40" pageNumber="1552" phylum="Arthropoda" rank="species" species="berezovskiyi" status="sp. nov.">
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<emphasis id="ECCCEAC9FFF13260980D4836FD721654" box="[562,723,1287,1312]" italics="true" pageId="40" pageNumber="1552">F. berezovskiyi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="F7FF57B2FFF1326098E34836FC991654" box="[732,824,1287,1312]" pageId="40" pageNumber="1552" rank="species">sp. nov.</taxonomicNameLabel>
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, but differs from it by a pair of anterior lateral dark brown maculae over notopleuron (
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<figureCitation id="46832A5EFFF1326099624818FC6E1636" box="[861,975,1321,1346]" captionStart="Figure 19" captionStartId="31.[160,225,1625,1648]" captionTargetBox="[168,1148,145,1587]" captionTargetId="figure-3@31.[163,1153,142,1594]" captionTargetPageId="31" captionText="Figure 19. Habitus of Formosargus species. (a) F. sagittocerus, male, lateral. (b–c) F. berezovskiyi sp. nov., female, paratype, dorsal and lateral, respectively. (d–e) F. borneensis sp. nov., female, holotype, dorsal and lateral, respectively. (f) F. mangoleensis sp. nov., female, holotype, dorsal." figureDoi="http://doi.org/10.5281/zenodo.7318226" httpUri="https://zenodo.org/record/7318226/files/figure.png" pageId="40" pageNumber="1552">Figure 19</figureCitation>
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(d)). Abdominal tergites with subtle brownish markings, mostly conforming with the background colour (
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<figureCitation id="46832A5EFFF132609AC4485EFECA16FC" box="[251,363,1391,1416]" captionStart="Figure 19" captionStartId="31.[160,225,1625,1648]" captionTargetBox="[168,1148,145,1587]" captionTargetId="figure-3@31.[163,1153,142,1594]" captionTargetPageId="31" captionText="Figure 19. Habitus of Formosargus species. (a) F. sagittocerus, male, lateral. (b–c) F. berezovskiyi sp. nov., female, paratype, dorsal and lateral, respectively. (d–e) F. borneensis sp. nov., female, holotype, dorsal and lateral, respectively. (f) F. mangoleensis sp. nov., female, holotype, dorsal." figureDoi="http://doi.org/10.5281/zenodo.7318226" httpUri="https://zenodo.org/record/7318226/files/figure.png" pageId="40" pageNumber="1552">Figure 19</figureCitation>
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(e)).
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</paragraph>
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</subSubSection>
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<subSubSection id="96A26550FFF132609A9F4885FCA51541" pageId="40" pageNumber="1552" type="materials_examined">
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||
<paragraph id="DE0736DBFFF132609A9F4885FEDE16BA" blockId="40.[160,1156,1460,1589]" box="[160,383,1460,1486]" pageId="40" pageNumber="1552">
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<emphasis id="ECCCEAC9FFF132609A9F4885FEDE16BA" bold="true" box="[160,383,1460,1486]" italics="true" pageId="40" pageNumber="1552">Material examined</emphasis>
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</paragraph>
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<paragraph id="DE0736DBFFF132609A9F48E6FCA51541" blockId="40.[160,1156,1460,1589]" pageId="40" pageNumber="1552">
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<materialsCitation id="6ED03C86FFF132609A9F48E6FCA51541" ID-GBIF-Occurrence="3965743329" collectingDate="1929-04-18" collectionCode="NHMUK" collectorName="H. M. Pendlebury" country="Malaysia" county="Borneo" elevation="2134" location="Kamborangah" municipality="Kinabalu" pageId="40" pageNumber="1552" specimenCount="1" specimenCount-female="1" stateProvince="Sabah" typeStatus="holotype">
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<emphasis id="ECCCEAC9FFF132609A9F48E6FE821684" bold="true" box="[160,291,1495,1520]" pageId="40" pageNumber="1552">
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||
<typeStatus id="01038879FFF132609A9F48E6FE821684" box="[160,291,1495,1520]" pageId="40" pageNumber="1552" type="holotype">HOLOTYPE</typeStatus>
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</emphasis>
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,
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<specimenCount id="C8BEFD52FFF132609B0F48E6FEF8169A" box="[304,345,1495,1520]" pageId="40" pageNumber="1552" type="female">1 ♀</specimenCount>
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(genitalia dissected and stored in microvial with glycerine),
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<emphasis id="ECCCEAC9FFF1326099C048E6FBDF1684" bold="true" box="[1023,1150,1495,1520]" pageId="40" pageNumber="1552">
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<collectingCountry id="A6AF764BFFF1326099C048E6FBDF1684" box="[1023,1150,1495,1520]" name="Malaysia" pageId="40" pageNumber="1552">MALAYSIA</collectingCountry>
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</emphasis>
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, B. [British] N. [Northern]
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<collectingCounty id="37664E57FFF132609B8448CBFDAF1567" box="[443,526,1530,1555]" pageId="40" pageNumber="1552">Borneo</collectingCounty>
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, [
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<collectingRegion id="1C7CF839FFF13260981F48CBFDCD1567" box="[544,620,1530,1555]" country="Malaysia" name="Sabah" pageId="40" pageNumber="1552">Sabah</collectingRegion>
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], Mt. [Mount]
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<collectingMunicipality id="3E63ACA1FFF13260993048CBFCD21567" box="[783,883,1530,1555]" pageId="40" pageNumber="1552">Kinabalu</collectingMunicipality>
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,
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||
<location id="DB676000FFF13260994048CBFB811567" LSID="urn:lsid:plazi:treatment:561187CDFFF132619BF84917FDEE1501:DB676000FFF13260994048CBFB811567" box="[895,1056,1530,1555]" country="Malaysia" county="Borneo" municipality="Kinabalu" name="Kamborangah" pageId="40" pageNumber="1552" stateProvince="Sabah">Kamborangah</location>
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||
,
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<quantity id="19409B3EFFF132609E1248CBFB221567" box="[1069,1155,1530,1555]" metricMagnitude="3" metricUnit="m" metricValue="2.1336" pageId="40" pageNumber="1552" unit="ft" value="7000.0">
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<elevation id="5595D1E8FFF132609E1248CBFB221567" box="[1069,1155,1530,1555]" metricMagnitude="3" metricUnit="m" metricValue="2.1336" pageId="40" pageNumber="1552" unit="ft" value="7000.0">7000 ft.</elevation>
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</quantity>
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||
[feet],
|
||
<date id="AA06101BFFF132609AD84B2DFE211541" box="[231,384,1564,1589]" pageId="40" pageNumber="1552" value="1929-04-18">
|
||
<collectingDate id="BA42E9F3FFF132609AD84B2DFE211541" box="[231,384,1564,1589]" pageId="40" pageNumber="1552" value="1929-04-18">18 April 1929</collectingDate>
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||
</date>
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||
, [coll.
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||
<collectorName id="734D530DFFF132609BF94B2DFD2B1541" box="[454,650,1564,1589]" pageId="40" pageNumber="1552">H. M. Pendlebury</collectorName>
|
||
] (
|
||
<collectionCode id="B8A9AE1EFFF1326098A34B2DFD5A1541" box="[668,763,1564,1589]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="40" pageNumber="1552" type="Museum">NHMUK</collectionCode>
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||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="96A26550FFF132619A9F4B50FEB210FB" lastPageId="41" lastPageNumber="1553" pageId="40" pageNumber="1552" type="description">
|
||
<paragraph id="DE0736DBFFF132609A9F4B50FE87150F" blockId="40.[160,1155,1633,1728]" box="[160,294,1633,1659]" pageId="40" pageNumber="1552">
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||
<emphasis id="ECCCEAC9FFF132609A9F4B50FE87150F" bold="true" box="[160,294,1633,1659]" italics="true" pageId="40" pageNumber="1552">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="DE0736DBFFF132619A9F4BB4FE921063" blockId="40.[160,1155,1633,1728]" lastBlockId="41.[160,1156,142,791]" lastPageId="41" lastPageNumber="1553" pageId="40" pageNumber="1552">
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||
<emphasis id="ECCCEAC9FFF132609A9F4BB4FF5615EA" bold="true" box="[160,247,1669,1694]" pageId="40" pageNumber="1552">Female</emphasis>
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||
. Length (n = 1): body, 10.0 mm; wing, 9.0 mm.
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<emphasis id="ECCCEAC9FFF1326099274BB4FCF715EA" bold="true" box="[792,854,1669,1694]" pageId="40" pageNumber="1552">Head</emphasis>
|
||
(
|
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<figureCitation id="46832A5EFFF1326099594BB4FC7115EA" box="[870,976,1669,1694]" captionStart="Figure 4" captionStartId="8.[160,225,1266,1289]" captionTargetBox="[167,1149,145,1232]" captionTargetId="figure-139@8.[163,1153,142,1234]" captionTargetPageId="8" captionText="Figure 4. Heads of Formosargus species, dorsal view. (a) F.berezovskiyi sp. nov., female, paratype. (b) F. borneensis sp. nov., female, holotype. (c) F. mangoleensis sp. nov., female, holotype. (d–e) F. trivittatus sp. nov. (d. male, holotype; e. female, paratype). (f) F. woodleyi sp. nov., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.7318188" httpUri="https://zenodo.org/record/7318188/files/figure.png" pageId="40" pageNumber="1552">Figures 4</figureCitation>
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(b), 6(b)). Vertex area twice longer than ocellar tubercle. Upper frons as wide as long, twice longer than lower frons; upper frons black lateral area broader than ocellar tubercle width, with short dark brownish pilosity; anterior margin of upper frons with a median triangular projection pointing towards antennal insertion. Fourth flagellomere rounded, as wide as each previous flagellomere alone or slightly wider. Face nearly as long as lower frons.
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||
<emphasis id="ECCCEAC9FFF032619E104DC7FB22127B" bold="true" box="[1071,1155,246,271]" pageId="41" pageNumber="1553">Thorax</emphasis>
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||
(
|
||
<figureCitation id="46832A5EFFF032619A974C28FEA51246" box="[168,260,281,306]" captionStart="Figure 7" captionStartId="12.[160,225,1575,1598]" captionTargetBox="[167,1149,145,1537]" captionTargetId="figure-3@12.[163,1153,142,1543]" captionTargetPageId="12" captionText="Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.7318195" httpUri="https://zenodo.org/record/7318195/files/figure.png" pageId="41" pageNumber="1553">Figure 7</figureCitation>
|
||
(f)). Medial scutal band dark brown, nearly as wide as one-third of scutum, with a subtle narrowing at level of transverse suture, wider posteriorly, weakly discontinued at base of scutellum; a subrectangular dark brown macula over notopleuron area, extending over anepisternum proximally; laterotergite reddish yellow, mediotergite with only a small dark brown mark. Basal half of hind femur dark brown.
|
||
<emphasis id="ECCCEAC9FFF0326199624C95FC3A12C9" bold="true" box="[861,923,420,445]" pageId="41" pageNumber="1553">Wing</emphasis>
|
||
. R2+3 arising slightly before r–m, weakly curved towards R1 on basal third, nearly straight on apical two-thirds. M1 and M3 less faint. Halter mostly reddish yellow, distal half of knob dark brown.
|
||
<emphasis id="ECCCEAC9FFF032619A9F4F3DFEB71151" bold="true" box="[160,278,524,549]" pageId="41" pageNumber="1553">Abdomen</emphasis>
|
||
. Reddish, with a central transverse brownish band, nearly covering entire surface of each tergites 1–5 (
|
||
<figureCitation id="46832A5EFFF032619BCF4F1FFDFF1133" box="[496,606,558,583]" captionStart="Figure 19" captionStartId="31.[160,225,1625,1648]" captionTargetBox="[168,1148,145,1587]" captionTargetId="figure-3@31.[163,1153,142,1594]" captionTargetPageId="31" captionText="Figure 19. Habitus of Formosargus species. (a) F. sagittocerus, male, lateral. (b–c) F. berezovskiyi sp. nov., female, paratype, dorsal and lateral, respectively. (d–e) F. borneensis sp. nov., female, holotype, dorsal and lateral, respectively. (f) F. mangoleensis sp. nov., female, holotype, dorsal." figureDoi="http://doi.org/10.5281/zenodo.7318226" httpUri="https://zenodo.org/record/7318226/files/figure.png" pageId="41" pageNumber="1553">Figure 19</figureCitation>
|
||
(d)), conforming in colour with the background; pilosity reddish yellow.
|
||
<emphasis id="ECCCEAC9FFF032619B994F60FD84111E" bold="true" box="[422,549,593,618]" pageId="41" pageNumber="1553">Terminalia</emphasis>
|
||
(
|
||
<figureCitation id="46832A5EFFF03261980B4F60FD0A111E" box="[564,683,593,618]" captionStart="Figure 15" captionStartId="27.[160,225,1701,1724]" captionTargetBox="[210,1104,144,1666]" captionTargetId="figure-1@27.[208,1107,142,1669]" captionTargetPageId="27" captionText="Figure 15. Female terminalia of Formosargus species, dorsal view. (a) F. kerteszi. (b) F. lineatus. (c) F. berezovskiyi sp. nov., paratype. (d) F. borneensis sp. nov., holotype (cerci lost). Abbreviations: cerc, cercus; st, sternite; tg, tergite." figureDoi="http://doi.org/10.5281/zenodo.7318213" httpUri="https://zenodo.org/record/7318213/files/figure.png" pageId="41" pageNumber="1553">Figures 15</figureCitation>
|
||
(d), 17(g–h)). Genital fork wider medially at level of posterolateral process insertion; anterior half subtriangular, narrowing into a relatively slender and rounded anterior end; posterior bridge weakly bilobed; posterolateral process enlarged and rounded at base, much slenderer on next one-third, enlarging on last third towards apex, distal half nearly parallel; genital opening reduced, roughly oval.
|
||
</paragraph>
|
||
<paragraph id="DE0736DBFFF032619A9F4E62FF7B1019" blockId="41.[160,275,851,911]" box="[160,218,851,877]" pageId="41" pageNumber="1553">
|
||
<emphasis id="ECCCEAC9FFF032619A9F4E62FF7B1019" bold="true" box="[160,218,851,877]" italics="true" pageId="41" pageNumber="1553">Male</emphasis>
|
||
</paragraph>
|
||
<paragraph id="DE0736DBFFF032619A9F4E47FEB210FB" blockId="41.[160,275,851,911]" box="[160,275,886,911]" pageId="41" pageNumber="1553">Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="96A26550FFF032619A9F4EFAFE48175E" pageId="41" pageNumber="1553" type="etymology">
|
||
<paragraph id="DE0736DBFFF032619A9F4EFAFEBC1091" blockId="41.[160,1155,971,1066]" box="[160,285,971,997]" pageId="41" pageNumber="1553">
|
||
<emphasis id="ECCCEAC9FFF032619A9F4EFAFEBC1091" bold="true" box="[160,285,971,997]" italics="true" pageId="41" pageNumber="1553">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="DE0736DBFFF032619A9F4EDFFE48175E" blockId="41.[160,1155,971,1066]" pageId="41" pageNumber="1553">
|
||
The specific epithet,
|
||
<taxonomicName id="19B84D58FFF032619BB14EDFFDA01773" authorityName="Fachin & Hauser" authorityYear="2022" box="[398,513,1006,1031]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="1553" phylum="Arthropoda" rank="species" species="borneensis">
|
||
<emphasis id="ECCCEAC9FFF032619BB14EDFFDA01773" box="[398,513,1006,1031]" italics="true" pageId="41" pageNumber="1553">borneensis</emphasis>
|
||
</taxonomicName>
|
||
, refers to the collection site of the
|
||
<typeStatus id="01038879FFF0326199A64EDFFC5C1773" box="[921,1021,1006,1031]" pageId="41" pageNumber="1553" type="holotype">holotype</typeStatus>
|
||
of the new species, the island of Borneo.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="96A26550FFF032619A9F4954FD0B17D6" pageId="41" pageNumber="1553" type="distribution">
|
||
<paragraph id="DE0736DBFFF032619A9F4954FE18170B" blockId="41.[160,682,1125,1186]" box="[160,441,1125,1151]" pageId="41" pageNumber="1553">
|
||
<emphasis id="ECCCEAC9FFF032619A9F4954FE18170B" bold="true" box="[160,441,1125,1151]" italics="true" pageId="41" pageNumber="1553">Geographic distribution</emphasis>
|
||
</paragraph>
|
||
<paragraph id="DE0736DBFFF032619A9F49B8FD0B17D6" blockId="41.[160,682,1125,1186]" box="[160,682,1161,1186]" pageId="41" pageNumber="1553">
|
||
<emphasis id="ECCCEAC9FFF032619A9F49B8FEFC17D6" bold="true" box="[160,349,1161,1186]" pageId="41" pageNumber="1553">Oriental Region</emphasis>
|
||
.
|
||
<collectingCountry id="A6AF764BFFF032619B5549B8FE6B17D6" box="[362,458,1161,1186]" name="Malaysia" pageId="41" pageNumber="1553">Malaysia</collectingCountry>
|
||
(
|
||
<collectingRegion id="1C7CF839FFF032619BE849B8FDBE17D6" box="[471,543,1161,1186]" country="Malaysia" name="Sabah" pageId="41" pageNumber="1553">Sabah</collectingRegion>
|
||
) (
|
||
<figureCitation id="46832A5EFFF03261980B49B8FD3C17D6" box="[564,669,1161,1186]" captionStart="Figure 26" captionStartId="38.[160,226,870,893]" captionTargetBox="[168,1148,146,834]" captionTargetId="figure-167@38.[163,1153,142,839]" captionTargetPageId="38" captionText="Figure 26. Geographic distribution of Formosargus. Squares represent holotypes records; circles represent paratypes, and additional material. Three of the ‘lines’ of faunal division are shown: 1. Wallace’s Line; 2. Weber’s Line; 3. Lydekker’s Line. ‘Wallacea’ is the area that lies between Wallace’s Line and Lydekker’s Line." figureDoi="http://doi.org/10.5281/zenodo.7318240" httpUri="https://zenodo.org/record/7318240/files/figure.png" pageId="41" pageNumber="1553">Figure 26</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="96A26550FFF032619A9F49ECFDEE1501" pageId="41" pageNumber="1553" type="discussion">
|
||
<paragraph id="DE0736DBFFF032619A9F49ECFEBF1783" blockId="41.[160,1156,1245,1653]" box="[160,286,1245,1271]" pageId="41" pageNumber="1553">
|
||
<emphasis id="ECCCEAC9FFF032619A9F49ECFEBF1783" bold="true" box="[160,286,1245,1271]" italics="true" pageId="41" pageNumber="1553">Comments</emphasis>
|
||
</paragraph>
|
||
<paragraph id="DE0736DBFFF032619A9F4830FDEE1501" blockId="41.[160,1156,1245,1653]" pageId="41" pageNumber="1553">
|
||
This species is part of a group of species, including
|
||
<emphasis id="ECCCEAC9FFF03261991E4830FF541649" italics="true" pageId="41" pageNumber="1553">
|
||
<taxonomicName id="19B84D58FFF03261991E4830FC35166E" authorityName="James" authorityYear="1939" box="[801,916,1281,1306]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="1553" phylum="Arthropoda" rank="species" species="kerteszi">F. kerteszi</taxonomicName>
|
||
,
|
||
<taxonomicName id="19B84D58FFF03261999A4830FBBD166E" box="[933,1052,1281,1306]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="1553" phylum="Arthropoda" rank="species" species="lineatus">F. lineatus</taxonomicName>
|
||
,
|
||
<taxonomicName id="19B84D58FFF032619E124830FF541649" authority="Fachin & Hauser, 2022" authorityName="Fachin & Hauser" authorityYear="2022" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="1553" phylum="Arthropoda" rank="species" species="berezovskiyi" status="sp. nov.">F. berezovskiyi</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel id="F7FF57B2FFF032619B3B4815FEC71649" box="[260,358,1316,1341]" pageId="41" pageNumber="1553" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="19B84D58FFF032619B484815FD8E1649" authority="Fachin & Hauser, 2022" authorityName="Fachin & Hauser" authorityYear="2022" box="[375,559,1316,1341]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="1553" phylum="Arthropoda" rank="species" species="mangoleensis" status="sp. nov.">
|
||
<emphasis id="ECCCEAC9FFF032619B484815FD8E1649" box="[375,559,1316,1341]" italics="true" pageId="41" pageNumber="1553">F. mangoleensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F7FF57B2FFF0326198024815FD3C1649" box="[573,669,1316,1341]" pageId="41" pageNumber="1553" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="19B84D58FFF0326198DB4815FC601649" authorityName="Lessard and Woodley" authorityYear="2020" box="[740,961,1316,1341]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="1553" phylum="Arthropoda" rank="species" species="melanogrammus">
|
||
<emphasis id="ECCCEAC9FFF0326198DB4815FC601649" box="[740,961,1316,1341]" italics="true" pageId="41" pageNumber="1553">F. melanogrammus</emphasis>
|
||
</taxonomicName>
|
||
, that share the presence of a single medial dark brown band on the scutum (see
|
||
<figureCitation id="46832A5EFFF0326199954877FBAA162B" box="[938,1035,1350,1375]" captionStart="Figure 7" captionStartId="12.[160,225,1575,1598]" captionTargetBox="[167,1149,145,1537]" captionTargetId="figure-3@12.[163,1153,142,1543]" captionTargetPageId="12" captionText="Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.7318195" httpUri="https://zenodo.org/record/7318195/files/figure.png" pageId="41" pageNumber="1553">Figure 7</figureCitation>
|
||
(a–c, e, g)) and a rounded fourth flagellomere (see
|
||
<figureCitation id="46832A5EFFF0326198B54858FD4F16F6" box="[650,750,1385,1410]" captionStart="Figure 2" captionStartId="6.[160,225,1404,1427]" captionTargetBox="[165,1150,144,1369]" captionTargetId="figure-53@6.[163,1153,142,1372]" captionTargetPageId="6" captionText="Figure 2. Antennae of Formosargus species. (a) F. lineatus, male. (b) F. lineatus, female. (c) F. trivittatus sp. nov., male, holotype. Abbreviations: ar t flgm, arista-like terminal flagellomere; flg, flagellum; ped, pedicel; scp, scape." figureDoi="http://doi.org/10.5281/zenodo.7318184" httpUri="https://zenodo.org/record/7318184/files/figure.png" pageId="41" pageNumber="1553">Figure 2</figureCitation>
|
||
(a, b)).
|
||
<taxonomicName id="19B84D58FFF0326199794858FBF716F6" authority="Fachin & Hauser, 2022" authorityName="Fachin & Hauser" authorityYear="2022" box="[838,1110,1385,1410]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="1553" phylum="Arthropoda" rank="species" species="borneensis" status="sp. nov.">
|
||
<emphasis id="ECCCEAC9FFF0326199794858FBF716F6" box="[838,1110,1385,1410]" italics="true" pageId="41" pageNumber="1553">Formosargus borneensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F7FF57B2FFF032619E5B4858FF7516D1" pageId="41" pageNumber="1553" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, however, has a pair of additional subrectangular dark markings laterally on the anterior part of the scutum, mostly over the notopleuron (
|
||
<figureCitation id="46832A5EFFF032619969489FFC6716B3" box="[854,966,1454,1479]" captionStart="Figure 19" captionStartId="31.[160,225,1625,1648]" captionTargetBox="[168,1148,145,1587]" captionTargetId="figure-3@31.[163,1153,142,1594]" captionTargetPageId="31" captionText="Figure 19. Habitus of Formosargus species. (a) F. sagittocerus, male, lateral. (b–c) F. berezovskiyi sp. nov., female, paratype, dorsal and lateral, respectively. (d–e) F. borneensis sp. nov., female, holotype, dorsal and lateral, respectively. (f) F. mangoleensis sp. nov., female, holotype, dorsal." figureDoi="http://doi.org/10.5281/zenodo.7318226" httpUri="https://zenodo.org/record/7318226/files/figure.png" pageId="41" pageNumber="1553">Figure 19</figureCitation>
|
||
(e); see couplets 1 and
|
||
<quantity id="19409B3EFFF032619AD148E0FEBA169E" box="[238,283,1489,1514]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="41" pageNumber="1553" unit="in" value="2.0">2 in</quantity>
|
||
the key). This feature indeed has similarities to the condition seen in the three-striped species, for example to that of
|
||
<taxonomicName id="19B84D58FFF03261988948C5FC9F1579" authority="Fachin & Hauser, 2022" authorityName="Fachin & Hauser" authorityYear="2022" box="[694,830,1524,1549]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="1553" phylum="Arthropoda" rank="species" species="trivittatus" status="sp. nov.">
|
||
<emphasis id="ECCCEAC9FFF03261988948C5FC9F1579" box="[694,830,1524,1549]" italics="true" pageId="41" pageNumber="1553">F. trivittatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F7FF57B2FFF03261997648C5FC041579" box="[841,933,1524,1549]" pageId="41" pageNumber="1553" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="46832A5EFFF03261998948C5FB861579" box="[950,1063,1524,1549]" captionStart="Figure 20" captionStartId="32.[160,225,1630,1653]" captionTargetBox="[171,1149,154,1593]" captionTargetId="figure-3@32.[163,1153,142,1598]" captionTargetPageId="32" captionText="Figure 20. Habitus of Formosargus species.(a) F. mangoleensis sp. nov., female, holotype, lateral. (b–d) F. trivittatus sp. nov. (b–c. male, holotype, dorsal and lateral, respectively; d. female, paratype, dorsal). (e–f) F. woodleyi sp. nov., male, holotype, dorsal and lateral, respectively." figureDoi="http://doi.org/10.5281/zenodo.7318228" httpUri="https://zenodo.org/record/7318228/files/figure.png" pageId="41" pageNumber="1553">Figure 20</figureCitation>
|
||
(c)). The difference is that in
|
||
<taxonomicName id="19B84D58FFF032619BAA4B27FDBC155B" authority="Fachin & Hauser, 2022" authorityName="Fachin & Hauser" authorityYear="2022" box="[405,541,1558,1583]" class="Insecta" family="Stratiomyidae" genus="Formosargus" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="1553" phylum="Arthropoda" rank="species" species="trivittatus" status="sp. nov.">
|
||
<emphasis id="ECCCEAC9FFF032619BAA4B27FDBC155B" box="[405,541,1558,1583]" italics="true" pageId="41" pageNumber="1553">F. trivittatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F7FF57B2FFF0326198164B27FD24155B" box="[553,645,1558,1583]" pageId="41" pageNumber="1553" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and in the other species related to it, the lateral macula is not only over the notopleuron but also extends over the anterior part of the scutum (see
|
||
<figureCitation id="46832A5EFFF032619BFD4B6DFD841501" box="[450,549,1628,1653]" captionStart="Figure 7" captionStartId="12.[160,225,1575,1598]" captionTargetBox="[167,1149,145,1537]" captionTargetId="figure-3@12.[163,1153,142,1543]" captionTargetPageId="12" captionText="Figure 7. Thorax of Formosargus species, dorsal view. (a–b) F. kerteszi (a. male, holotype [DEI]; b. female, Philippines). (c) F. lineatus, male; arrow: medial band. (d) F. sagittocerus, male. (e) F. berezovskiyi sp. nov., female, paratype; arrow: mediotergite. (f) F. borneensis sp. nov., female, holotype; arrow: notopleural macula. (g) F. mangoleensis sp. nov., female, holotype; arrow: mediotergite. (h) F. trivittatus sp. nov., male, holotype; arrow: anterior lateral band. (i) F. woodleyi sp. nov., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.7318195" httpUri="https://zenodo.org/record/7318195/files/figure.png" pageId="41" pageNumber="1553">Figure 7</figureCitation>
|
||
(h)).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |