906 lines
102 KiB
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906 lines
102 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.866.35394" ID-GBIF-Dataset="4f3c8ff0-c68c-4b0e-a2bd-cab653d0f8fe" ID-PMC="PMC6669213" ID-Pensoft-Pub="1313-2970-866-1" ID-Pensoft-UUID="ECA044C40A7255A0B8F1935457F2E629" ID-PubMed="31388320" ID-ZooBank="EDAEA22F45824B1DB5CB46302E7AA43F" ModsDocID="1313-2970-866-1" checkinTime="1564043551668" checkinUser="pensoft" docAuthor="Tauber, Catherine A." docDate="2019" docId="4CFFA2D7DEBB58A09186E3DA99DD4CC7" docLanguage="en" docName="ZooKeys 866: 1-18" docOrigin="ZooKeys 866" docSource="http://dx.doi.org/10.3897/zookeys.866.35394" docTitle="Nothochrysa ehrenbergi Tauber, sp. nov." docType="treatment" docUuid="528B2ED3-82DF-4A61-8DF2-DD9DD5D77FED" docUuidSource="ZooBank" docVersion="5" lastPageNumber="13" masterDocId="456EE02CFFB9FFED2E1AFF84754DFFF1" masterDocTitle="South American Nothochrysinae (Neuroptera, Chrysopidae): I. Description of Nothochrysa ehrenbergi sp. nov." masterLastPageNumber="18" masterPageNumber="1" pageNumber="4" updateTime="1668167617226" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>South American Nothochrysinae (Neuroptera, Chrysopidae): I. Description of Nothochrysa ehrenbergi sp. nov.</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tauber, Catherine A.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>866</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>18</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.866.35394</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.866.35394</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-866-1</mods:identifier>
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<mods:identifier type="ZooBank">EDAEA22F45824B1DB5CB46302E7AA43F</mods:identifier>
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<mods:identifier type="Pensoft-UUID">ECA044C40A7255A0B8F1935457F2E629</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="159292916" LSID="urn:lsid:zoobank.org:act:528B2ED3-82DF-4A61-8DF2-DD9DD5D77FED" httpUri="http://treatment.plazi.org/id/4CFFA2D7DEBB58A09186E3DA99DD4CC7" lastPageId="12" lastPageNumber="13" pageId="3" pageNumber="4">
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<subSubSection pageId="3" pageNumber="4" type="nomenclature">
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<paragraph pageId="3" pageNumber="4">
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<taxonomicName LSID="4cffa2d7-debb-58a0-9186-e3da99dd4cc7" authority="Tauber" class="Insecta" family="Chrysopidae" genus="Nothochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothochrysa ehrenbergi" order="Neuroptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="ehrenbergi">Nothochrysa ehrenbergi Tauber</taxonomicName>
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<taxonomicNameLabel pageId="3" pageNumber="4">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="4" type="type material">
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<paragraph pageId="3" pageNumber="4">Type material.</paragraph>
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<paragraph pageId="3" pageNumber="4">
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The
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<emphasis bold="true" pageId="3" pageNumber="4">holotype</emphasis>
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(a male) is in the California Academy of Sciences (CAS). Its labels read: [1] "CHILE: Nuble [
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<normalizedToken originalValue="Ñuble">Nuble</normalizedToken>
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] / Las Trancas / 20/25-II-1980 / Luis E. Pena [
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<normalizedToken originalValue="Peña]”">Pena]"</normalizedToken>
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; [2] "
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<taxonomicName class="Insecta" family="Chrysopidae" genus="Suarius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Suarius" order="Neuroptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Suarius</taxonomicName>
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/
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<taxonomicName lsidName="flavescens" pageId="3" pageNumber="4" rank="species" species="flavescens">flavescens</taxonomicName>
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/ (Blanchard) / det. N. Penny, 1988"; [3] "HOLOTYPE /
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<emphasis italics="true" pageId="3" pageNumber="4">
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<taxonomicName class="Insecta" family="Chrysopidae" genus="Nothochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothochrysa" order="Neuroptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Nothochrysa</taxonomicName>
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/
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<taxonomicName lsidName="ehrenbergi" pageId="3" pageNumber="4" rank="species" species="ehrenbergi">ehrenbergi</taxonomicName>
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</emphasis>
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/
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<bibRefCitation pageId="3" pageNumber="4" refId="B22">Tauber 2019</bibRefCitation>
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" (
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Nothochrysa ehrenbergi sp. nov. (Nuble, Chile; Male, CAS): Male genitalia, cleared, and specimen labels (Penny's identification label not included) (a) gonarcal complex, dorsal (b) gonarcal complex, frontal, tilted (c) gonarcal complex, posterior (d) gonarcal complex, lateral (e) hypandrium internum (f) labels. c comes g. a. gonarcal apodeme g. b. gonarcal bridge g. p. gonarcal process gse gonosetae on membranous gonosaccus mu mediuncus." figureDoi="10.3897/zookeys.866.35394.figure7" httpUri="https://binary.pensoft.net/fig/319512" pageId="3" pageNumber="4">Fig. 7f</figureCitation>
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).
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</paragraph>
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<paragraph pageId="3" pageNumber="4">
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This single specimen was found in the CAS collection among the unidentified chrysopids. A subsequent search of the collection did not yield additional examples. Norm
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<normalizedToken originalValue="Penny’s">Penny's</normalizedToken>
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ID label remains on the specimen but was not included in
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Nothochrysa ehrenbergi sp. nov. (Nuble, Chile; Male, CAS): Male genitalia, cleared, and specimen labels (Penny's identification label not included) (a) gonarcal complex, dorsal (b) gonarcal complex, frontal, tilted (c) gonarcal complex, posterior (d) gonarcal complex, lateral (e) hypandrium internum (f) labels. c comes g. a. gonarcal apodeme g. b. gonarcal bridge g. p. gonarcal process gse gonosetae on membranous gonosaccus mu mediuncus." figureDoi="10.3897/zookeys.866.35394.figure7" httpUri="https://binary.pensoft.net/fig/319512" pageId="3" pageNumber="4">Fig. 7f</figureCitation>
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. It refers to
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<taxonomicName class="Insecta" family="Chrysopidae" genus="Suarius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Suarius flavescens" order="Neuroptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="flavescens">
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<emphasis italics="true" pageId="3" pageNumber="4">Suarius flavescens</emphasis>
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</taxonomicName>
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, a species that now is placed in
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<taxonomicName class="Insecta" family="Chrysopidae" genus="Chrysopodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chrysopodes (Neosuarius)" order="Neuroptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="subGenus" subGenus="Neosuarius">Chrysopodes (Neosuarius)</taxonomicName>
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, and with which the new species shares similar coloration and appearance (see
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<bibRefCitation DOI="https://doi.org/10.3897/zookeys.44.387" author="Tauber, CA" journalOrPublisher="Records of the Indian Museum, Calcutta" pageId="14" pageNumber="15" refId="B21" refString="Tauber, CA, 2010. . https://doi.org/10.3897/zookeys.44.387" url="https://doi.org/10.3897/zookeys.44.387" year="2010">Tauber 2010</bibRefCitation>
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).
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</paragraph>
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<paragraph pageId="3" pageNumber="4">When discovered, the specimen was discolored, and its wings were loosely folded around its body. One pair of wings was removed for study and is now attached with water-soluble hide glue to a card mounted on the pin below the specimen. The other pair fell off and was reattached to the specimen with hide glue. The abdomen was cleared and dissected; it is preserved in glycerin within a genitalia vial attached to the pin.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="6" lastPageNumber="7" pageId="3" pageNumber="4" type="diagnosis">
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<paragraph pageId="3" pageNumber="4">Diagnosis.</paragraph>
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<paragraph pageId="3" pageNumber="4">
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Subfamily: This specimen exhibits the following diagnostic features of adult
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<taxonomicName lsidName="" pageId="3" pageNumber="4" rank="subfamily" subfamily="Nothochrysinae">Nothochrysinae</taxonomicName>
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(cf.:
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<bibRefCitation pageId="3" pageNumber="4" refId="B25">Tjeder 1966</bibRefCitation>
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, as
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<taxonomicName lsidName="" pageId="3" pageNumber="4" rank="subfamily" subfamily="Dictyochrysinae">Dictyochrysinae</taxonomicName>
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;
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<bibRefCitation author="Adams, PA" journalOrPublisher="Bulletin of the Museum of Comparative Zoology" pageId="13" pageNumber="14" pagination="215 - 238" refId="B1" refString="Adams, PA, 1967. A review of the Mesochrysinae and Nothochrysinae (Neuroptera: Chrysopidae). . Bulletin of the Museum of Comparative Zoology 135: 215 - 238" title="A review of the Mesochrysinae and Nothochrysinae (Neuroptera: Chrysopidae)." volume="135" year="1967">Adams 1967</bibRefCitation>
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;
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<bibRefCitation author="Brooks, SJ" journalOrPublisher="Bulletin of the British Museum of Natural History, Entomology" pageId="13" pageNumber="14" pagination="117 - 286" refId="B8" refString="Brooks, SJ, Barnard, PC, 1990. The green lacewings of the world: a generic review (Neuroptera: Chrysopidae). . Bulletin of the British Museum of Natural History, Entomology 59: 117 - 286" title="The green lacewings of the world: a generic review (Neuroptera: Chrysopidae)." volume="59" year="1990">Brooks and Barnard 1990</bibRefCitation>
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;
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<bibRefCitation DOI="https://doi.org/10.1666/12-052R.1" author="Makarkin, VN" journalOrPublisher="Journal of Paleontology" pageId="14" pageNumber="15" pagination="123 - 146" refId="B16" refString="Makarkin, VN, Archibald, SB, 2013. A diverse new assemblage of green lacewings (Insecta, Neuroptera, Chrysopidae) from the Early Eocene Okanagan Highlands, western North America. . Journal of Paleontology 87: 123 - 146" title="A diverse new assemblage of green lacewings (Insecta, Neuroptera, Chrysopidae) from the Early Eocene Okanagan Highlands, western North America." url="https://doi.org/10.1666/12-052R.1" volume="87" year="2013">Makarkin and Archibald 2013</bibRefCitation>
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;
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<bibRefCitation author="Breitkreuz, LCV" journalOrPublisher="PhD Thesis, University of Kansas, Lawrence" pageId="13" pageNumber="14" refId="B6" refString="Breitkreuz, LCV, 2018. Systematics and evolution of the family Chrysopidae (Neuroptera), with an emphasis on their morphology. . PhD Thesis, University of Kansas, Lawrence" title="Systematics and evolution of the family Chrysopidae (Neuroptera), with an emphasis on their morphology." year="2018">Breitkreuz 2018</bibRefCitation>
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): (i) wing-coupling mechanism consisting of a large jugal lobe on the forewing (here, folded ventrally;
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Nothochrysa ehrenbergi sp. nov. (Nuble, Chile; Male, CAS): Venation at base of wings (a) left forewing, (b) left hindwing. Note the absence of a tympanal organ at the base of R in the forewing, the independent origin and trajectory of M along the base of R (arrows pointing downward, both wings), and the alignment of RP and MA in the hindwing. A 1, A 2, A 3 first, second, third anal veins Cu cubitus Cu f furcation (division) of cubitus Ju jugal lobe M media MA media anterior m-cu media-cubital crossvein R radius RP radius posterior." figureDoi="10.3897/zookeys.866.35394.figure1" httpUri="https://binary.pensoft.net/fig/319506" pageId="3" pageNumber="4">Fig. 1</figureCitation>
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) and a frenulum on the hindwing (here, broken off); (ii) base of the forewing without tympanal organ (
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Nothochrysa ehrenbergi sp. nov. (Nuble, Chile; Male, CAS): Venation at base of wings (a) left forewing, (b) left hindwing. Note the absence of a tympanal organ at the base of R in the forewing, the independent origin and trajectory of M along the base of R (arrows pointing downward, both wings), and the alignment of RP and MA in the hindwing. A 1, A 2, A 3 first, second, third anal veins Cu cubitus Cu f furcation (division) of cubitus Ju jugal lobe M media MA media anterior m-cu media-cubital crossvein R radius RP radius posterior." figureDoi="10.3897/zookeys.866.35394.figure1" httpUri="https://binary.pensoft.net/fig/319506" pageId="3" pageNumber="4">Fig. 1</figureCitation>
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); (iii) forewing (and hindwing) with stem of the media extending basally, adjacent to the radius and not fused with it (
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Nothochrysa ehrenbergi sp. nov. (Nuble, Chile; Male, CAS): Venation at base of wings (a) left forewing, (b) left hindwing. Note the absence of a tympanal organ at the base of R in the forewing, the independent origin and trajectory of M along the base of R (arrows pointing downward, both wings), and the alignment of RP and MA in the hindwing. A 1, A 2, A 3 first, second, third anal veins Cu cubitus Cu f furcation (division) of cubitus Ju jugal lobe M media MA media anterior m-cu media-cubital crossvein R radius RP radius posterior." figureDoi="10.3897/zookeys.866.35394.figure1" httpUri="https://binary.pensoft.net/fig/319506" pageId="3" pageNumber="4">Fig. 1a, b</figureCitation>
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; cf.
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<bibRefCitation DOI="https://doi.org/10.1206/3890.1" author="Breitkreuz, LCV" journalOrPublisher="American Museum Novitates" pageId="13" pageNumber="14" pagination="1 - 44" refId="B7" refString="Breitkreuz, LCV, Winterton, SL, Engel, MS, 2017. Wing tracheation in Chrysopidae and other Neuropterida (Insecta): a resolution of the confusion about vein fusion. . American Museum Novitates 3890: 1 - 44" title="Wing tracheation in Chrysopidae and other Neuropterida (Insecta): a resolution of the confusion about vein fusion." url="https://doi.org/10.1206/3890.1" volume="3890" year="2017">Breitkreuz et al. 2017</bibRefCitation>
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: 32); (iv) first intramedian cell triangular, with boundaries formed by the MA, the MP, and the crossvein 1ma-mp ("pseudotriangular", sensu
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<bibRefCitation DOI="https://doi.org/10.1206/3890.1" author="Breitkreuz, LCV" journalOrPublisher="American Museum Novitates" pageId="13" pageNumber="14" pagination="1 - 44" refId="B7" refString="Breitkreuz, LCV, Winterton, SL, Engel, MS, 2017. Wing tracheation in Chrysopidae and other Neuropterida (Insecta): a resolution of the confusion about vein fusion. . American Museum Novitates 3890: 1 - 44" title="Wing tracheation in Chrysopidae and other Neuropterida (Insecta): a resolution of the confusion about vein fusion." url="https://doi.org/10.1206/3890.1" volume="3890" year="2017">Breitkreuz et al. 2017</bibRefCitation>
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); (v) pseudomedia ill-defined or appearing to merge with inner (not outer) series of gradates (
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nothochrysa ehrenbergi sp. nov. (Nuble, Chile; Male, CAS): Wings with selected features labeled (a) left forewing, (b) left hindwing. Marginal traces of major veins demarcated; arrow (hindwing) indicates alignment of RP and MA along upper margin of first intramedian cell. A 1, A 2, A 3 first, second, third anal veins CuA, CuP anterior, posterior branches of cubitus icu 1, icu 3 first, third intracubital cells ig inner gradate im 1, im 2 first, second intramedian cells Ju jugal lobe MA media anterior MP media posterior mcua, mpcua second and third medial cells M f furcation of media og outer gradate Psc pseudocubitus Psm pseudomedia R f furcation of radius RP radius posterior RP 1 first branch of radius posterior 1 sc-r first crossvein between subcosta and radius 2 m-cu second crossvein between media and cubitus." figureDoi="10.3897/zookeys.866.35394.figure2" httpUri="https://binary.pensoft.net/fig/319507" pageId="3" pageNumber="4">Fig. 2</figureCitation>
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); (vi) pseudocubitus appearing to merge with outer series of gradates (
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nothochrysa ehrenbergi sp. nov. (Nuble, Chile; Male, CAS): Wings with selected features labeled (a) left forewing, (b) left hindwing. Marginal traces of major veins demarcated; arrow (hindwing) indicates alignment of RP and MA along upper margin of first intramedian cell. A 1, A 2, A 3 first, second, third anal veins CuA, CuP anterior, posterior branches of cubitus icu 1, icu 3 first, third intracubital cells ig inner gradate im 1, im 2 first, second intramedian cells Ju jugal lobe MA media anterior MP media posterior mcua, mpcua second and third medial cells M f furcation of media og outer gradate Psc pseudocubitus Psm pseudomedia R f furcation of radius RP radius posterior RP 1 first branch of radius posterior 1 sc-r first crossvein between subcosta and radius 2 m-cu second crossvein between media and cubitus." figureDoi="10.3897/zookeys.866.35394.figure2" httpUri="https://binary.pensoft.net/fig/319507" pageId="3" pageNumber="4">Fig. 2</figureCitation>
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); (vii) forewing with basal subcostal crossvein present (
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nothochrysa ehrenbergi sp. nov. (Nuble, Chile; Male, CAS): Wings with selected features labeled (a) left forewing, (b) left hindwing. Marginal traces of major veins demarcated; arrow (hindwing) indicates alignment of RP and MA along upper margin of first intramedian cell. A 1, A 2, A 3 first, second, third anal veins CuA, CuP anterior, posterior branches of cubitus icu 1, icu 3 first, third intracubital cells ig inner gradate im 1, im 2 first, second intramedian cells Ju jugal lobe MA media anterior MP media posterior mcua, mpcua second and third medial cells M f furcation of media og outer gradate Psc pseudocubitus Psm pseudomedia R f furcation of radius RP radius posterior RP 1 first branch of radius posterior 1 sc-r first crossvein between subcosta and radius 2 m-cu second crossvein between media and cubitus." figureDoi="10.3897/zookeys.866.35394.figure2" httpUri="https://binary.pensoft.net/fig/319507" pageId="3" pageNumber="4">Fig. 2</figureCitation>
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); (viii) second m-cu crossvein stemming from the proximal half of the first intramedian cell (
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nothochrysa ehrenbergi sp. nov. (Nuble, Chile; Male, CAS): Wings with selected features labeled (a) left forewing, (b) left hindwing. Marginal traces of major veins demarcated; arrow (hindwing) indicates alignment of RP and MA along upper margin of first intramedian cell. A 1, A 2, A 3 first, second, third anal veins CuA, CuP anterior, posterior branches of cubitus icu 1, icu 3 first, third intracubital cells ig inner gradate im 1, im 2 first, second intramedian cells Ju jugal lobe MA media anterior MP media posterior mcua, mpcua second and third medial cells M f furcation of media og outer gradate Psc pseudocubitus Psm pseudomedia R f furcation of radius RP radius posterior RP 1 first branch of radius posterior 1 sc-r first crossvein between subcosta and radius 2 m-cu second crossvein between media and cubitus." figureDoi="10.3897/zookeys.866.35394.figure2" httpUri="https://binary.pensoft.net/fig/319507" pageId="3" pageNumber="4">Fig. 2</figureCitation>
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); (ix) each flagellomere having five or six whorls of setae (
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Nothochrysa ehrenbergi sp. nov. (Nuble, Chile; Male, CAS): Head and prothorax (a) head, frontal (b) head and prothorax, dorsal (c) head and prothorax, lateral (d, e) base of antennae, dorsal, lateral (f) flagellar segments, mid antenna." figureDoi="10.3897/zookeys.866.35394.figure3" httpUri="https://binary.pensoft.net/fig/319508" pageId="3" pageNumber="4">Figs 3e</figureCitation>
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,
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Nothochrysa ehrenbergi sp. nov. (Nuble, Chile; Male, CAS): Head and prothorax (a) head, frontal (b) head and prothorax, dorsal (c) head and prothorax, lateral (d, e) base of antennae, dorsal, lateral (f) flagellar segments, mid antenna." figureDoi="10.3897/zookeys.866.35394.figure3" httpUri="https://binary.pensoft.net/fig/319508" pageId="3" pageNumber="4">3f</figureCitation>
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); and (x) anterodorsal surface of the metascutum displaying small, convex protrusion (
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Nothochrysa ehrenbergi sp. nov. (Nuble, Chile; Male, CAS): Habitus (a) antenna, head, and thorax, lateral (b) mesothorax, metathorax, dorsal (c) metatarsus, dorsal (d) metatarsus, ventral (e) mesotarsus, lateral. p raised metascutal protuberance l. e. mesoscutellar lobate expansion." figureDoi="10.3897/zookeys.866.35394.figure4" httpUri="https://binary.pensoft.net/fig/319509" pageId="3" pageNumber="4">Fig. 4b</figureCitation>
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; cf.
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<bibRefCitation author="Breitkreuz, LCV" journalOrPublisher="PhD Thesis, University of Kansas, Lawrence" pageId="13" pageNumber="14" refId="B6" refString="Breitkreuz, LCV, 2018. Systematics and evolution of the family Chrysopidae (Neuroptera), with an emphasis on their morphology. . PhD Thesis, University of Kansas, Lawrence" title="Systematics and evolution of the family Chrysopidae (Neuroptera), with an emphasis on their morphology." year="2018">Breitkreuz 2018</bibRefCitation>
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,
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<bibRefCitation pageId="3" pageNumber="4" refId="B22">Tauber 2019</bibRefCitation>
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).
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</paragraph>
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<caption doi="10.3897/zookeys.866.35394.figure1" httpUri="https://binary.pensoft.net/fig/319506" pageId="3" pageNumber="4" start="Figure 1" startId="F1">
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<paragraph pageId="3" pageNumber="4">
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Figure 1.
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<taxonomicName class="Insecta" family="Chrysopidae" genus="Nothochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothochrysa ehrenbergi" order="Neuroptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="ehrenbergi">
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<emphasis italics="true" pageId="3" pageNumber="4">Nothochrysa ehrenbergi</emphasis>
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</taxonomicName>
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sp. nov. (
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<normalizedToken originalValue="Ñuble">Nuble</normalizedToken>
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, Chile; Male, CAS): Venation at base of wings (
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<emphasis bold="true" pageId="3" pageNumber="4">a</emphasis>
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) left forewing, (
|
||
<emphasis bold="true" pageId="3" pageNumber="4">b</emphasis>
|
||
) left hindwing. Note the absence of a tympanal organ at the base of R in the forewing, the independent origin and trajectory of M along the base of R (arrows pointing downward, both wings), and the alignment of RP and MA in the hindwing.
|
||
<emphasis bold="true" pageId="3" pageNumber="4">A1, A2, A3</emphasis>
|
||
first, second, third anal veins
|
||
<emphasis bold="true" pageId="3" pageNumber="4">Cu</emphasis>
|
||
cubitus
|
||
<emphasis bold="true" pageId="3" pageNumber="4">
|
||
Cu
|
||
<emphasis italics="true" pageId="3" pageNumber="4">f</emphasis>
|
||
</emphasis>
|
||
furcation (division) of cubitus
|
||
<emphasis bold="true" pageId="3" pageNumber="4">Ju</emphasis>
|
||
jugal lobe
|
||
<emphasis bold="true" pageId="3" pageNumber="4">M</emphasis>
|
||
media
|
||
<emphasis bold="true" pageId="3" pageNumber="4">MA</emphasis>
|
||
media anterior
|
||
<emphasis bold="true" pageId="3" pageNumber="4">m-cu</emphasis>
|
||
media-cubital crossvein
|
||
<emphasis bold="true" pageId="3" pageNumber="4">R</emphasis>
|
||
radius
|
||
<emphasis bold="true" pageId="3" pageNumber="4">RP</emphasis>
|
||
radius posterior.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.866.35394.figure2" httpUri="https://binary.pensoft.net/fig/319507" pageId="3" pageNumber="4" start="Figure 2" startId="F2">
|
||
<paragraph pageId="3" pageNumber="4">
|
||
Figure 2.
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Nothochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothochrysa ehrenbergi" order="Neuroptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="ehrenbergi">
|
||
<emphasis italics="true" pageId="3" pageNumber="4">Nothochrysa ehrenbergi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (
|
||
<normalizedToken originalValue="Ñuble">Nuble</normalizedToken>
|
||
, Chile; Male, CAS): Wings with selected features labeled (
|
||
<emphasis bold="true" pageId="3" pageNumber="4">a</emphasis>
|
||
) left forewing, (
|
||
<emphasis bold="true" pageId="3" pageNumber="4">b</emphasis>
|
||
) left hindwing. Marginal traces of major veins demarcated; arrow (hindwing) indicates alignment of RP and MA along upper margin of first intramedian cell.
|
||
<emphasis bold="true" pageId="3" pageNumber="4">A1, A2, A3</emphasis>
|
||
first, second, third anal veins
|
||
<emphasis bold="true" pageId="3" pageNumber="4">CuA, CuP</emphasis>
|
||
anterior, posterior branches of cubitus
|
||
<emphasis bold="true" pageId="3" pageNumber="4">
|
||
<emphasis italics="true" pageId="3" pageNumber="4">icu1</emphasis>
|
||
,
|
||
<emphasis italics="true" pageId="3" pageNumber="4">icu3</emphasis>
|
||
</emphasis>
|
||
first, third intracubital cells
|
||
<emphasis bold="true" pageId="3" pageNumber="4">ig</emphasis>
|
||
inner gradate
|
||
<emphasis bold="true" italics="true" pageId="3" pageNumber="4">im1, im2</emphasis>
|
||
first, second intramedian cells
|
||
<emphasis bold="true" pageId="3" pageNumber="4">Ju</emphasis>
|
||
jugal lobe
|
||
<emphasis bold="true" pageId="3" pageNumber="4">MA</emphasis>
|
||
media anterior
|
||
<emphasis bold="true" pageId="3" pageNumber="4">MP</emphasis>
|
||
media posterior
|
||
<emphasis bold="true" italics="true" pageId="3" pageNumber="4">mcua, mpcua</emphasis>
|
||
second and third medial cells
|
||
<emphasis bold="true" pageId="3" pageNumber="4">
|
||
M
|
||
<emphasis italics="true" pageId="3" pageNumber="4">f</emphasis>
|
||
</emphasis>
|
||
furcation of media
|
||
<emphasis bold="true" pageId="3" pageNumber="4">og</emphasis>
|
||
outer gradate
|
||
<emphasis bold="true" pageId="3" pageNumber="4">Psc</emphasis>
|
||
pseudocubitus
|
||
<emphasis bold="true" pageId="3" pageNumber="4">Psm</emphasis>
|
||
pseudomedia
|
||
<emphasis bold="true" pageId="3" pageNumber="4">
|
||
R
|
||
<emphasis italics="true" pageId="3" pageNumber="4">f</emphasis>
|
||
</emphasis>
|
||
furcation of radius
|
||
<emphasis bold="true" pageId="3" pageNumber="4">RP</emphasis>
|
||
radius posterior
|
||
<emphasis bold="true" pageId="3" pageNumber="4">RP1</emphasis>
|
||
first branch of radius posterior
|
||
<emphasis bold="true" pageId="3" pageNumber="4">1sc-r</emphasis>
|
||
first crossvein between subcosta and radius
|
||
<emphasis bold="true" pageId="3" pageNumber="4">2m-cu</emphasis>
|
||
second crossvein between media and cubitus.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.866.35394.figure3" httpUri="https://binary.pensoft.net/fig/319508" pageId="3" pageNumber="4" start="Figure 3" startId="F3">
|
||
<paragraph pageId="3" pageNumber="4">
|
||
Figure 3.
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Nothochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothochrysa ehrenbergi" order="Neuroptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="ehrenbergi">
|
||
<emphasis italics="true" pageId="3" pageNumber="4">Nothochrysa ehrenbergi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (
|
||
<normalizedToken originalValue="Ñuble">Nuble</normalizedToken>
|
||
, Chile; Male, CAS): Head and prothorax (
|
||
<emphasis bold="true" pageId="3" pageNumber="4">a</emphasis>
|
||
) head, frontal (
|
||
<emphasis bold="true" pageId="3" pageNumber="4">b</emphasis>
|
||
) head and prothorax, dorsal (
|
||
<emphasis bold="true" pageId="3" pageNumber="4">c</emphasis>
|
||
) head and prothorax, lateral (
|
||
<emphasis bold="true" pageId="3" pageNumber="4">d, e</emphasis>
|
||
) base of antennae, dorsal, lateral
|
||
<emphasis bold="true" pageId="3" pageNumber="4">(f)</emphasis>
|
||
flagellar segments, mid antenna.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.866.35394.figure4" httpUri="https://binary.pensoft.net/fig/319509" pageId="3" pageNumber="4" start="Figure 4" startId="F4">
|
||
<paragraph pageId="3" pageNumber="4">
|
||
Figure 4.
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Nothochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothochrysa ehrenbergi" order="Neuroptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="ehrenbergi">
|
||
<emphasis italics="true" pageId="3" pageNumber="4">Nothochrysa ehrenbergi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (
|
||
<normalizedToken originalValue="Ñuble">Nuble</normalizedToken>
|
||
, Chile; Male, CAS): Habitus (
|
||
<emphasis bold="true" pageId="3" pageNumber="4">a</emphasis>
|
||
) antenna, head, and thorax, lateral (
|
||
<emphasis bold="true" pageId="3" pageNumber="4">b</emphasis>
|
||
) mesothorax, metathorax, dorsal (
|
||
<emphasis bold="true" pageId="3" pageNumber="4">c</emphasis>
|
||
) metatarsus, dorsal
|
||
<emphasis bold="true" pageId="3" pageNumber="4">(d)</emphasis>
|
||
metatarsus, ventral
|
||
<emphasis bold="true" pageId="3" pageNumber="4">(e)</emphasis>
|
||
mesotarsus, lateral.
|
||
<emphasis bold="true" pageId="3" pageNumber="4">p</emphasis>
|
||
raised metascutal protuberance
|
||
<emphasis bold="true" pageId="3" pageNumber="4">l.e.</emphasis>
|
||
mesoscutellar lobate expansion.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="4" pageNumber="5">
|
||
<pageBreakToken pageId="4" pageNumber="5" start="start">Genus</pageBreakToken>
|
||
placement: The Chilean specimen under study here falls into the genus
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Nothochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothochrysa" order="Neuroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="4" pageNumber="5">Nothochrysa</emphasis>
|
||
</taxonomicName>
|
||
on the basis of the following features of its wings (
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Nothochrysa ehrenbergi sp. nov. (Nuble, Chile; Male, CAS): Venation at base of wings (a) left forewing, (b) left hindwing. Note the absence of a tympanal organ at the base of R in the forewing, the independent origin and trajectory of M along the base of R (arrows pointing downward, both wings), and the alignment of RP and MA in the hindwing. A 1, A 2, A 3 first, second, third anal veins Cu cubitus Cu f furcation (division) of cubitus Ju jugal lobe M media MA media anterior m-cu media-cubital crossvein R radius RP radius posterior." figureDoi="10.3897/zookeys.866.35394.figure1" httpUri="https://binary.pensoft.net/fig/319506" pageId="4" pageNumber="5">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nothochrysa ehrenbergi sp. nov. (Nuble, Chile; Male, CAS): Wings with selected features labeled (a) left forewing, (b) left hindwing. Marginal traces of major veins demarcated; arrow (hindwing) indicates alignment of RP and MA along upper margin of first intramedian cell. A 1, A 2, A 3 first, second, third anal veins CuA, CuP anterior, posterior branches of cubitus icu 1, icu 3 first, third intracubital cells ig inner gradate im 1, im 2 first, second intramedian cells Ju jugal lobe MA media anterior MP media posterior mcua, mpcua second and third medial cells M f furcation of media og outer gradate Psc pseudocubitus Psm pseudomedia R f furcation of radius RP radius posterior RP 1 first branch of radius posterior 1 sc-r first crossvein between subcosta and radius 2 m-cu second crossvein between media and cubitus." figureDoi="10.3897/zookeys.866.35394.figure2" httpUri="https://binary.pensoft.net/fig/319507" pageId="4" pageNumber="5">2</figureCitation>
|
||
): (i) forewing and hindwing having well developed pseudomedia and pseudocubitus; (ii) forewing and hindwing with two regular series of gradate veins (inner and outer); (iii) intramedian cell of forewing triangular, elongate, occupying approximately half the width between the pseudomedia and pseudocubitus; (iv) RP of forewing with 10 or more branches (
|
||
<bibRefCitation author="Adams, PA" journalOrPublisher="Bulletin of the Museum of Comparative Zoology" pageId="13" pageNumber="14" pagination="215 - 238" refId="B1" refString="Adams, PA, 1967. A review of the Mesochrysinae and Nothochrysinae (Neuroptera: Chrysopidae). . Bulletin of the Museum of Comparative Zoology 135: 215 - 238" title="A review of the Mesochrysinae and Nothochrysinae (Neuroptera: Chrysopidae)." volume="135" year="1967">Adams 1967</bibRefCitation>
|
||
;
|
||
<bibRefCitation DOI="https://doi.org/10.1666/12-052R.1" author="Makarkin, VN" journalOrPublisher="Journal of Paleontology" pageId="14" pageNumber="15" pagination="123 - 146" refId="B16" refString="Makarkin, VN, Archibald, SB, 2013. A diverse new assemblage of green lacewings (Insecta, Neuroptera, Chrysopidae) from the Early Eocene Okanagan Highlands, western North America. . Journal of Paleontology 87: 123 - 146" title="A diverse new assemblage of green lacewings (Insecta, Neuroptera, Chrysopidae) from the Early Eocene Okanagan Highlands, western North America." url="https://doi.org/10.1666/12-052R.1" volume="87" year="2013">Makarkin and Archibald 2013</bibRefCitation>
|
||
;
|
||
<bibRefCitation DOI="https://doi.org/10.4039/tce.2014.53" author="Archibald, SB" journalOrPublisher="Canadian Entomologist" pageId="13" pageNumber="14" pagination="359 - 369" refId="B3" refString="Archibald, SB, Makarkin, VN, 2015. A new species of Archaeochrysa Adams (Neuroptera: Chrysopidae) from the early Eocene of Driftwood Canyon, British Columbia, Canada. . Canadian Entomologist 147: 359 - 369" title="A new species of Archaeochrysa Adams (Neuroptera: Chrysopidae) from the early Eocene of Driftwood Canyon, British Columbia, Canada." url="https://doi.org/10.4039/tce.2014.53" volume="147" year="2015">Archibald and Makarkin 2015</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Breitkreuz, LCV" journalOrPublisher="PhD Thesis, University of Kansas, Lawrence" pageId="13" pageNumber="14" refId="B6" refString="Breitkreuz, LCV, 2018. Systematics and evolution of the family Chrysopidae (Neuroptera), with an emphasis on their morphology. . PhD Thesis, University of Kansas, Lawrence" title="Systematics and evolution of the family Chrysopidae (Neuroptera), with an emphasis on their morphology." year="2018">Breitkreuz 2018</bibRefCitation>
|
||
: 200). [Note: Some specimens of
|
||
<taxonomicName lsidName="N. californica" pageId="4" pageNumber="5" rank="species" species="californica">
|
||
<emphasis italics="true" pageId="4" pageNumber="5">N. californica</emphasis>
|
||
</taxonomicName>
|
||
are known to have only eight or nine branches from the RP.]
|
||
</paragraph>
|
||
<paragraph lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5">
|
||
Species placement: Apart from being the only known
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Nothochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothochrysa" order="Neuroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="4" pageNumber="5">Nothochrysa</emphasis>
|
||
</taxonomicName>
|
||
species reported from South America,
|
||
<taxonomicName lsidName="N. ehrenbergi" pageId="4" pageNumber="5" rank="species" species="ehrenbergi">
|
||
<emphasis italics="true" pageId="4" pageNumber="5">N. ehrenbergi</emphasis>
|
||
</taxonomicName>
|
||
is distinguishable from other species of
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Nothochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothochrysa" order="Neuroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="4" pageNumber="5">Nothochrysa</emphasis>
|
||
</taxonomicName>
|
||
on the basis of a number of wing characters (
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Nothochrysa ehrenbergi sp. nov. (Nuble, Chile; Male, CAS): Venation at base of wings (a) left forewing, (b) left hindwing. Note the absence of a tympanal organ at the base of R in the forewing, the independent origin and trajectory of M along the base of R (arrows pointing downward, both wings), and the alignment of RP and MA in the hindwing. A 1, A 2, A 3 first, second, third anal veins Cu cubitus Cu f furcation (division) of cubitus Ju jugal lobe M media MA media anterior m-cu media-cubital crossvein R radius RP radius posterior." figureDoi="10.3897/zookeys.866.35394.figure1" httpUri="https://binary.pensoft.net/fig/319506" pageId="4" pageNumber="5">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nothochrysa ehrenbergi sp. nov. (Nuble, Chile; Male, CAS): Wings with selected features labeled (a) left forewing, (b) left hindwing. Marginal traces of major veins demarcated; arrow (hindwing) indicates alignment of RP and MA along upper margin of first intramedian cell. A 1, A 2, A 3 first, second, third anal veins CuA, CuP anterior, posterior branches of cubitus icu 1, icu 3 first, third intracubital cells ig inner gradate im 1, im 2 first, second intramedian cells Ju jugal lobe MA media anterior MP media posterior mcua, mpcua second and third medial cells M f furcation of media og outer gradate Psc pseudocubitus Psm pseudomedia R f furcation of radius RP radius posterior RP 1 first branch of radius posterior 1 sc-r first crossvein between subcosta and radius 2 m-cu second crossvein between media and cubitus." figureDoi="10.3897/zookeys.866.35394.figure2" httpUri="https://binary.pensoft.net/fig/319507" pageId="4" pageNumber="5">2</figureCitation>
|
||
; cf.
|
||
<bibRefCitation author="Adams, PA" journalOrPublisher="Bulletin of the Museum of Comparative Zoology" pageId="13" pageNumber="14" pagination="215 - 238" refId="B1" refString="Adams, PA, 1967. A review of the Mesochrysinae and Nothochrysinae (Neuroptera: Chrysopidae). . Bulletin of the Museum of Comparative Zoology 135: 215 - 238" title="A review of the Mesochrysinae and Nothochrysinae (Neuroptera: Chrysopidae)." volume="135" year="1967">Adams 1967</bibRefCitation>
|
||
;
|
||
<bibRefCitation pageId="4" pageNumber="5" refId="B4">
|
||
<normalizedToken originalValue="Aspöck">Aspoeck</normalizedToken>
|
||
et al. 1980
|
||
</bibRefCitation>
|
||
<pageBreakToken pageId="5" pageNumber="6" start="start">:</pageBreakToken>
|
||
figs 154, 155;
|
||
<bibRefCitation DOI="https://doi.org/10.1080/00379271.2007.10697507" author="Kovanci, B" journalOrPublisher="Annales de la Societe Entomologique de France (NS)" pageId="14" pageNumber="15" pagination="165 - 168" refId="B13" refString="Kovanci, B, Canbulat, S, 2007. A new species of the genus Nothochrysa McLachlan 1868 from northwestern Turkey (Neuroptera: Chrysopidae) with a key to western Palaearctic species. . Annales de la Societe Entomologique de France (NS) 43: 165 - 168" title="A new species of the genus Nothochrysa McLachlan 1868 from northwestern Turkey (Neuroptera: Chrysopidae) with a key to western Palaearctic species." url="https://doi.org/10.1080/00379271.2007.10697507" volume="43" year="2007">Kovanci and Canbulat 2007</bibRefCitation>
|
||
: fig. 2): (i) the first anal vein is not forked; (ii) the basal subcostal crossvein is slightly distal to the furcation of the radius; (iii) as in most
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Nothochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothochrysa" order="Neuroptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="5" pageNumber="6">Nothochrysa</emphasis>
|
||
</taxonomicName>
|
||
species, the first intramedian cell is more wedge shaped
|
||
<pageBreakToken pageId="6" pageNumber="7" start="start">than</pageBreakToken>
|
||
truly quadrangular or triangular (i.e., the MA and MP meet basally at a broadly acute angle); and (iv) the third medial cell (directly below
|
||
<emphasis italics="true" pageId="6" pageNumber="7">im1</emphasis>
|
||
,
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nothochrysa ehrenbergi sp. nov. (Nuble, Chile; Male, CAS): Wings with selected features labeled (a) left forewing, (b) left hindwing. Marginal traces of major veins demarcated; arrow (hindwing) indicates alignment of RP and MA along upper margin of first intramedian cell. A 1, A 2, A 3 first, second, third anal veins CuA, CuP anterior, posterior branches of cubitus icu 1, icu 3 first, third intracubital cells ig inner gradate im 1, im 2 first, second intramedian cells Ju jugal lobe MA media anterior MP media posterior mcua, mpcua second and third medial cells M f furcation of media og outer gradate Psc pseudocubitus Psm pseudomedia R f furcation of radius RP radius posterior RP 1 first branch of radius posterior 1 sc-r first crossvein between subcosta and radius 2 m-cu second crossvein between media and cubitus." figureDoi="10.3897/zookeys.866.35394.figure2" httpUri="https://binary.pensoft.net/fig/319507" pageId="6" pageNumber="7">Fig. 2a</figureCitation>
|
||
) is elongate and extends toward the pseudocubitus well beyond the distal edge of first intramedial cell.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="10" lastPageNumber="11" pageId="6" pageNumber="7" type="description">
|
||
<paragraph pageId="6" pageNumber="7">Morphological characteristics.</paragraph>
|
||
<paragraph lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
|
||
Head (
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Nothochrysa ehrenbergi sp. nov. (Nuble, Chile; Male, CAS): Head and prothorax (a) head, frontal (b) head and prothorax, dorsal (c) head and prothorax, lateral (d, e) base of antennae, dorsal, lateral (f) flagellar segments, mid antenna." figureDoi="10.3897/zookeys.866.35394.figure3" httpUri="https://binary.pensoft.net/fig/319508" pageId="6" pageNumber="7">Fig. 3</figureCitation>
|
||
): Width 1.6 mm (including eyes); ratio of head width to eye width = 3.0: 1. Vertex raised, round; surface pitted anteriorly, with few or no setae, lacking prominent posterior fold. Distance between scapes 0.09 mm; distance between tentorial pits 0.36 mm; length of frons (midway between scapes - midway between tentorial pits) 0.33 mm. Frons relatively wide, with broad longitudinal ridge mesally; surface smooth, shiny, slightly rounded below toruli and at insertion of mouthparts; margin above clypeus straight. Clypeus tapering, with rounded sculpturing basally, indented mesally, slightly expanded distally, with distal margin straight to slightly convex; dorsal surface shiny, smooth, sculptured. Labrum about same width as clypeal margin, with small longitudinal ridge mesally; dorsal surface
|
||
<pageBreakToken pageId="7" pageNumber="8" start="start">sculptured</pageBreakToken>
|
||
, shiny; distal margin bilobed, bearing numerous long setae distally. Antenna 9.7-9.8 mm long (~0.5
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
length of forewing); scape shorter than wide (0.23 mm long, 0.33 mm wide), lateral margin straight, mesal margin strongly convex, surface with short setae throughout; pedicel 0.17 mm long, 0.13 mm wide, with numerous short setae; flagellum with basal flagellomeres distinct, somewhat elongate (0.12-0.14 mm long, 0.07-0.08 mm wide), midantennal flagellomeres twice as long as broad (0.15 mm long, 0.07 mm wide), basal two flagellomeres with 4-5 partially indistinct whorls of thickset brown setae extending distally, third flagellomere and others distally all with five distinct whorls of thickset, brown setae extending distally, 0.3
|
||
<normalizedToken originalValue="–0.5×">-0.5x</normalizedToken>
|
||
width of flagellomere, distal whorl with one or two slender, elongate (~0.75
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
width of flagellomere), pale setae extending laterally.
|
||
</paragraph>
|
||
<paragraph lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
|
||
<emphasis italics="true" pageId="7" pageNumber="8">Head coloration</emphasis>
|
||
: Scape cream, with reddish spot on distolateral tip; pedicel, flagellum cream, unmarked; thickset setae in whorls mostly brown, elongate setae pale. Vertex cream, possibly tinged red laterally; dorsal torulus yellow to cream, apparently
|
||
<pageBreakToken pageId="8" pageNumber="9" start="start">unmarked</pageBreakToken>
|
||
. Frons cream, probably with reddish tinge laterally below torulus; torulus cream, unmarked. Clypeus cream, possibly tinged red laterally; basal, distal margins straight. Genal mark dark red/brown throughout, extending to tentorial pit. Labrum probably cream. Palpomeres probably mostly cream, somewhat darkened distally.
|
||
</paragraph>
|
||
<paragraph pageId="8" pageNumber="9">
|
||
Thorax (
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Nothochrysa ehrenbergi sp. nov. (Nuble, Chile; Male, CAS): Habitus (a) antenna, head, and thorax, lateral (b) mesothorax, metathorax, dorsal (c) metatarsus, dorsal (d) metatarsus, ventral (e) mesotarsus, lateral. p raised metascutal protuberance l. e. mesoscutellar lobate expansion." figureDoi="10.3897/zookeys.866.35394.figure4" httpUri="https://binary.pensoft.net/fig/319509" pageId="8" pageNumber="9">Fig. 4</figureCitation>
|
||
): Cervix not visible. Dorsal thoracic surface with pale longitudinal stripe mesally, probably with broad reddish or brownish stripes or coloration laterally. Prothorax broad, 0.9 mm long, 1.5 mm wide, ratio of length to width = 0.63: 1; pronotum well sclerotized, with textured surface, transverse fold mesally, few or no setae. Legs elongate, slender, probably cream, unmarked, lacking prominent tibial spurs. Tarsus with basal three tarsomeres appearing coalesced, bearing spurs, setae intermixed along undersurface; middle three tarsomeres with expanded lateral lobes bearing spurs, setae in irregular rows; distal tarsomere narrow basally, enlarged distally, bearing numerous elongate, slender, dark setae laterally, distally, terminus bearing pair of claws laterally, large pad mesally; claw amber, with basal enlargement, acute slender hook terminally.
|
||
</paragraph>
|
||
<paragraph pageId="9" pageNumber="10">
|
||
<pageBreakToken pageId="9" pageNumber="10" start="start">Wings</pageBreakToken>
|
||
(
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Nothochrysa ehrenbergi sp. nov. (Nuble, Chile; Male, CAS): Venation at base of wings (a) left forewing, (b) left hindwing. Note the absence of a tympanal organ at the base of R in the forewing, the independent origin and trajectory of M along the base of R (arrows pointing downward, both wings), and the alignment of RP and MA in the hindwing. A 1, A 2, A 3 first, second, third anal veins Cu cubitus Cu f furcation (division) of cubitus Ju jugal lobe M media MA media anterior m-cu media-cubital crossvein R radius RP radius posterior." figureDoi="10.3897/zookeys.866.35394.figure1" httpUri="https://binary.pensoft.net/fig/319506" pageId="9" pageNumber="10">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nothochrysa ehrenbergi sp. nov. (Nuble, Chile; Male, CAS): Wings with selected features labeled (a) left forewing, (b) left hindwing. Marginal traces of major veins demarcated; arrow (hindwing) indicates alignment of RP and MA along upper margin of first intramedian cell. A 1, A 2, A 3 first, second, third anal veins CuA, CuP anterior, posterior branches of cubitus icu 1, icu 3 first, third intracubital cells ig inner gradate im 1, im 2 first, second intramedian cells Ju jugal lobe MA media anterior MP media posterior mcua, mpcua second and third medial cells M f furcation of media og outer gradate Psc pseudocubitus Psm pseudomedia R f furcation of radius RP radius posterior RP 1 first branch of radius posterior 1 sc-r first crossvein between subcosta and radius 2 m-cu second crossvein between media and cubitus." figureDoi="10.3897/zookeys.866.35394.figure2" httpUri="https://binary.pensoft.net/fig/319507" pageId="9" pageNumber="10">2</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Nothochrysa ehrenbergi sp. nov. (Nuble, Chile; Male, CAS): Wings, color slightly enhanced to emphasize pattern of vein markings (a) forewing (b) hindwing." figureDoi="10.3897/zookeys.866.35394.figure5" httpUri="https://binary.pensoft.net/fig/319510" pageId="9" pageNumber="10">5</figureCitation>
|
||
):
|
||
<emphasis italics="true" pageId="9" pageNumber="10">Forewing</emphasis>
|
||
18.5 mm long, 6.5 mm wide (at widest point); ratio of length to maximum width = 2.9: 1. Membrane clear, lacking markings; microtrichia present below base of every major vein, pale. Trichosors (sensu
|
||
<bibRefCitation DOI="https://doi.org/10.1666/12-052R.1" author="Makarkin, VN" journalOrPublisher="Journal of Paleontology" pageId="14" pageNumber="15" pagination="123 - 146" refId="B16" refString="Makarkin, VN, Archibald, SB, 2013. A diverse new assemblage of green lacewings (Insecta, Neuroptera, Chrysopidae) from the Early Eocene Okanagan Highlands, western North America. . Journal of Paleontology 87: 123 - 146" title="A diverse new assemblage of green lacewings (Insecta, Neuroptera, Chrysopidae) from the Early Eocene Okanagan Highlands, western North America." url="https://doi.org/10.1666/12-052R.1" volume="87" year="2013">Makarkin and Archibald 2013</bibRefCitation>
|
||
: 140-142) absent. Costal area relatively enlarged; tallest costal cell (7th from base of wing) 1.8 mm tall, 2.7
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
width of cell, 0.28
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
height of wing; costal crossveins simple, six before 1sc-r, twelve after 1sc-r and before stigma, one (very small) after stigma, none within stigma. Sc extending into stigma, fading but not appearing to merge with C or RA; no crossveins in stigma; first sc-r crossvein slightly distal to R
|
||
<emphasis italics="true" pageId="9" pageNumber="10">f</emphasis>
|
||
, slightly basal to M
|
||
<emphasis italics="true" pageId="9" pageNumber="10">f</emphasis>
|
||
; RA with one very short veinlet extending to wing margin after stigma. Radial area between RA and RP with single row of ten closed cells; tallest cell (3rd from base of wing) 0.6
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as tall as wide. Intramedian cell (
|
||
<emphasis italics="true" pageId="9" pageNumber="10">im1</emphasis>
|
||
=
|
||
<emphasis italics="true" pageId="9" pageNumber="10">mamp1</emphasis>
|
||
) prominent, elongate, triangular, formed by MA, crossvein 1ma-mp, and two abscissae of MP, occupying approximately half the space between MA and CuA, with M
|
||
<emphasis italics="true" pageId="9" pageNumber="10">f</emphasis>
|
||
broadly acute, long sides (MA, MP) roughly parallel for most of span; crossvein 2m-cu proximal to midpoint of
|
||
<emphasis italics="true" pageId="9" pageNumber="10">im1</emphasis>
|
||
. Three medial cells present (
|
||
<emphasis italics="true" pageId="9" pageNumber="10">mcu, mcua, mpcua</emphasis>
|
||
), second, third of these elongate, with roughly parallel sides; MP merging into Psc well beyond
|
||
<emphasis italics="true" pageId="9" pageNumber="10">im1</emphasis>
|
||
. Two series of gradate veins parallel basally, diverging slightly medially, converging distally. Approximately nine inner gradates in regular, sinuous series, continuing from Psm in zigzag pattern across center of wing; approximately ten outer gradates continuing from Psc in regular, upturned series. RP with nine marginal forks beyond Psc. Cu furcated after m-cu crossvein, with two closed, four open
|
||
<emphasis italics="true" pageId="9" pageNumber="10">icu</emphasis>
|
||
cells. CuA with three furcations before meeting MP; CuP furcated below
|
||
<emphasis italics="true" pageId="9" pageNumber="10">icu2</emphasis>
|
||
; thus cubital trace having five terminal veinlets (three from CuA, two from CuP). A1, A2, A3 simple, unforked; a1-a2 and a2-a3 crossveins present; distal part of
|
||
<emphasis italics="true" pageId="9" pageNumber="10">a3</emphasis>
|
||
and jugal lobe with dense patch of microtrichia. Jugal lobe large, quadrate, folded beneath third anal cell, without internal vein; margin bearing long, slender setae basally.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.866.35394.figure5" httpUri="https://binary.pensoft.net/fig/319510" pageId="9" pageNumber="10" start="Figure 5" startId="F5">
|
||
<paragraph pageId="9" pageNumber="10">
|
||
Figure 5.
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Nothochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothochrysa ehrenbergi" order="Neuroptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="ehrenbergi">
|
||
<emphasis italics="true" pageId="9" pageNumber="10">Nothochrysa ehrenbergi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (
|
||
<normalizedToken originalValue="Ñuble">Nuble</normalizedToken>
|
||
, Chile; Male, CAS): Wings, color slightly enhanced to emphasize pattern of vein markings (
|
||
<emphasis bold="true" pageId="9" pageNumber="10">a</emphasis>
|
||
) forewing (
|
||
<emphasis bold="true" pageId="9" pageNumber="10">b</emphasis>
|
||
) hindwing.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="9" pageNumber="10">
|
||
<emphasis italics="true" pageId="9" pageNumber="10">Hindwing</emphasis>
|
||
: 12.4 mm long, 4.2 mm wide. Costal area not enlarged; at least 15 c-sc crossveins before stigma, none within or after stigma. Radial area containing single row of eleven closed cells between RA and RP. Gradate veins in two roughly parallel series, slightly divergent distally; approximately seven inner gradates beyond Psm; approximately 11 outer gradates beyond Psc. Psc with nine marginal forks. MA aligned with RP for approximately one-third length of
|
||
<emphasis italics="true" pageId="9" pageNumber="10">im1</emphasis>
|
||
. CuA with two furcations before meeting MP; CuP undivided; thus, wing margin having three cubital veinlets (two from CuA, one from CuP). A1, A2, A3 simple, unforked; a1-a2 and a2-a3 crossveins present. Jugal lobe without internal vein, basal margin bearing long, slender setae.
|
||
</paragraph>
|
||
<paragraph pageId="9" pageNumber="10">
|
||
<emphasis italics="true" pageId="9" pageNumber="10">Coloration of forewing, hindwing</emphasis>
|
||
(
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Nothochrysa ehrenbergi sp. nov. (Nuble, Chile; Male, CAS): Wings, color slightly enhanced to emphasize pattern of vein markings (a) forewing (b) hindwing." figureDoi="10.3897/zookeys.866.35394.figure5" httpUri="https://binary.pensoft.net/fig/319510" pageId="9" pageNumber="10">Fig. 5</figureCitation>
|
||
): Membrane clear, somewhat glossy. Stigma slightly opaque, without coloration. Costal, subcostal, radial veins brownish; all other longitudinal veins pale with black marks at intersections and (forewing) at bases of setae. Forewing with posterior veinlets extensively marked black; basal inner gradates pale, others becoming increasingly marked black until entirely black distally; outer gradates mostly black. Hindwing with basal inner gradates pale, marked with black at intersections; outer gradates mostly black.
|
||
</paragraph>
|
||
<paragraph pageId="10" pageNumber="11">
|
||
<pageBreakToken pageId="10" pageNumber="11" start="start">Abdomen</pageBreakToken>
|
||
(Male,
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Nothochrysa ehrenbergi sp. nov. (Nuble, Chile; Male, CAS): Abdomen, cleared (a) midsection-terminus, lateral (b) T 8 (distal), T 9, and ectoproct, lateral (c) terminal abdominal segments, lateral (d) terminal abdominal segments, ventral. apo dorsal apodeme extending below T 8 cc callus cerci ect ectoproct k distal knob extending from S 8 + 9 sr spiracle S 4, S 7 fourth, seventh strenites S 8, S 9 partially coalesced eighth and ninth sternites T 7, T 8, T 9 seventh, eighth, ninth tergites." figureDoi="10.3897/zookeys.866.35394.figure6" httpUri="https://binary.pensoft.net/fig/319511" pageId="10" pageNumber="11">Fig. 6</figureCitation>
|
||
; female unknown): Sclerites, integument of pleural region somewhat soft, flexible; tergites, sternites, pleural region covered with setae of uniformly short length; microsetae present, no microtholi. T6: length 0.78 mm, ~1.8
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
height; T7: length 0.80 mm, ~1.6
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
height; S6: length 0.67 mm, 0.72
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
height; S7: length 0.68 mm, ~0.70
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
height. Tergites roughly rectangular, edges acute or slightly rounded, ventral margins straight or slightly concave mesally. Spiracles located approximately in center of lateral membrane, roughly circular externally, not enlarged; atria slightly enlarged, rounded, with bifurcated tracheae. Coloration: body somewhat discolored; setae pale. Tergites probably green, without markings; pleuron mostly tan; sternites with green longitudinal stripe dorsally, tan ventrally; callus cerci white.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.866.35394.figure6" httpUri="https://binary.pensoft.net/fig/319511" pageId="10" pageNumber="11" start="Figure 6" startId="F6">
|
||
<paragraph pageId="10" pageNumber="11">
|
||
Figure 6.
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Nothochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothochrysa ehrenbergi" order="Neuroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ehrenbergi">
|
||
<emphasis italics="true" pageId="10" pageNumber="11">Nothochrysa ehrenbergi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (
|
||
<normalizedToken originalValue="Ñuble">Nuble</normalizedToken>
|
||
, Chile; Male, CAS): Abdomen, cleared (
|
||
<emphasis bold="true" pageId="10" pageNumber="11">a</emphasis>
|
||
) midsection-terminus, lateral (
|
||
<emphasis bold="true" pageId="10" pageNumber="11">b</emphasis>
|
||
) T8 (distal), T9, and ectoproct, lateral (
|
||
<emphasis bold="true" pageId="10" pageNumber="11">c</emphasis>
|
||
) terminal abdominal segments, lateral (
|
||
<emphasis bold="true" pageId="10" pageNumber="11">d</emphasis>
|
||
) terminal abdominal segments, ventral.
|
||
<emphasis bold="true" pageId="10" pageNumber="11">apo</emphasis>
|
||
dorsal apodeme extending below T8
|
||
<emphasis bold="true" pageId="10" pageNumber="11">cc</emphasis>
|
||
callus cerci
|
||
<emphasis bold="true" pageId="10" pageNumber="11">ect</emphasis>
|
||
ectoproct
|
||
<emphasis bold="true" pageId="10" pageNumber="11">k</emphasis>
|
||
distal knob extending from S8+9
|
||
<emphasis bold="true" pageId="10" pageNumber="11">sr</emphasis>
|
||
spiracle
|
||
<emphasis bold="true" pageId="10" pageNumber="11">S4, S7</emphasis>
|
||
fourth, seventh strenites
|
||
<emphasis bold="true" pageId="10" pageNumber="11">S8, S9</emphasis>
|
||
partially coalesced eighth and ninth sternites
|
||
<emphasis bold="true" pageId="10" pageNumber="11">T7, T8, T9</emphasis>
|
||
seventh, eighth, ninth tergites.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="10" pageNumber="11">
|
||
Male terminalia (
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Nothochrysa ehrenbergi sp. nov. (Nuble, Chile; Male, CAS): Male genitalia, cleared, and specimen labels (Penny's identification label not included) (a) gonarcal complex, dorsal (b) gonarcal complex, frontal, tilted (c) gonarcal complex, posterior (d) gonarcal complex, lateral (e) hypandrium internum (f) labels. c comes g. a. gonarcal apodeme g. b. gonarcal bridge g. p. gonarcal process gse gonosetae on membranous gonosaccus mu mediuncus." figureDoi="10.3897/zookeys.866.35394.figure7" httpUri="https://binary.pensoft.net/fig/319512" pageId="10" pageNumber="11">Fig. 7</figureCitation>
|
||
): T8 broadly wedge shaped, with dorsal surface slightly rounded, length 0.83 mm, height 0.49 mm, considerably longer than dorsal surfaces of either T9 or ectoproct; lateral margins tapering inward ventrally, ventral margin roughly straight. T9 and ectoproct separate, not fused; callus cerci ovate, protruding basally from posterior margin of ectoproct, 0.18 mm length, 0.10 mm width, with ~30 trichobothria of various lengths. T9 rectangular, with distoventral margin rounded; elongate, lightly sclerotized ventral apodeme along ventral margin, extending proximally to midsection of A8. Ectoproct dome shaped, rounded distally, slightly convex basally, tightly curved ventrally, sloping dorsally; callus cerci situated on lower proximal margin. S8 and S9 partially fused, without internal ridge; S9 more heavily sclerotized than S8, posterior margin slightly more sclerotized than remainder of sternite. S8+9 (lateral view) with proximal margin straight ventrally, becoming broadly rounded dorsally, distal margin short, straight, ventral margin straight; terminal knob extending well beyond edge of S9, with elongate setae on ventral margin; dorsal surface of knob contiguous with heavy recurrent membrane attached to elongate gonarcal membrane. Subanal plate not found.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.866.35394.figure7" httpUri="https://binary.pensoft.net/fig/319512" pageId="10" pageNumber="11" start="Figure 7" startId="F7">
|
||
<paragraph pageId="10" pageNumber="11">
|
||
Figure 7.
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Nothochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothochrysa ehrenbergi" order="Neuroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ehrenbergi">
|
||
<emphasis italics="true" pageId="10" pageNumber="11">Nothochrysa ehrenbergi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (
|
||
<normalizedToken originalValue="Ñuble">Nuble</normalizedToken>
|
||
, Chile; Male, CAS): Male genitalia, cleared, and specimen labels (
|
||
<normalizedToken originalValue="Penny’s">Penny's</normalizedToken>
|
||
identification label not included) (
|
||
<emphasis bold="true" pageId="10" pageNumber="11">a</emphasis>
|
||
) gonarcal complex, dorsal (
|
||
<emphasis bold="true" pageId="10" pageNumber="11">b</emphasis>
|
||
) gonarcal complex, frontal, tilted (
|
||
<emphasis bold="true" pageId="10" pageNumber="11">c</emphasis>
|
||
) gonarcal complex, posterior (
|
||
<emphasis bold="true" pageId="10" pageNumber="11">d</emphasis>
|
||
) gonarcal complex, lateral (
|
||
<emphasis bold="true" pageId="10" pageNumber="11">e</emphasis>
|
||
) hypandrium internum
|
||
<emphasis bold="true" pageId="10" pageNumber="11">(f)</emphasis>
|
||
labels.
|
||
<emphasis bold="true" pageId="10" pageNumber="11">c</emphasis>
|
||
comes
|
||
<emphasis bold="true" pageId="10" pageNumber="11">g.a.</emphasis>
|
||
gonarcal apodeme
|
||
<emphasis bold="true" pageId="10" pageNumber="11">g.b.</emphasis>
|
||
gonarcal bridge
|
||
<emphasis bold="true" pageId="10" pageNumber="11">g.p.</emphasis>
|
||
gonarcal process
|
||
<emphasis bold="true" pageId="10" pageNumber="11">gse</emphasis>
|
||
gonosetae on membranous gonosaccus
|
||
<emphasis bold="true" pageId="10" pageNumber="11">mu</emphasis>
|
||
mediuncus.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="10" pageNumber="11">
|
||
Gonarcus delicate, slender, broadly arcuate; lateral apodemes slender, quadrate (lateral view), rounded distally, with short, contiguous processes mesally, extending forward. Mediuncus closely attached to dorsal surface of gonarcal arch, flat, recurved into an almost fully circular hood, with two internal sclerotized
|
||
<normalizedToken originalValue="“rods”">"rods"</normalizedToken>
|
||
extending roughly in parallel from mediuncal base to tip, converging slightly at tip; base of mediuncus quadrate (dorsal view), occupying approximately one-fourth span of gonarcal bridge; terminus of mediuncus with expanded lateral wings, rounded mesal protrusion. Gonosaccus transparent, immediately beneath gonarcal arch and mediuncus, with approximately 32 short setae on distinct setal bases uniformly distributed in two equal patches. Hypandrium internum small, located on delicate membrane extending well below gonosaccus, consisting of paired, curved lateral arms meeting mesally at narrow, rounded apex; comes lightly sclerotized, extending forward beyond apex. Gonapsis, gonocristae absent.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="biology_ecology">
|
||
<paragraph pageId="10" pageNumber="11">Biology.</paragraph>
|
||
<paragraph pageId="10" pageNumber="11">
|
||
Nothing is known about the biology or larval morphology of this species. The gut of the
|
||
<taxonomicName lsidName="N. ehrenbergi" pageId="10" pageNumber="11" rank="species" species="ehrenbergi">
|
||
<emphasis italics="true" pageId="10" pageNumber="11">N. ehrenbergi</emphasis>
|
||
</taxonomicName>
|
||
specimen did not contain noteworthy contents.
|
||
</paragraph>
|
||
<paragraph pageId="11" pageNumber="12">
|
||
<pageBreakToken pageId="11" pageNumber="12" start="start">Larval</pageBreakToken>
|
||
descriptions of several
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Nothochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothochrysa" order="Neuroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="11" pageNumber="12">Nothochrysa</emphasis>
|
||
</taxonomicName>
|
||
species are available for comparison if
|
||
<taxonomicName lsidName="N. ehrenbergi" pageId="11" pageNumber="12" rank="species" species="ehrenbergi">
|
||
<emphasis italics="true" pageId="11" pageNumber="12">N. ehrenbergi</emphasis>
|
||
</taxonomicName>
|
||
larval specimens were to become available (see
|
||
<bibRefCitation DOI="https://doi.org/10.1603/AN13163" author="Tauber, CA" journalOrPublisher="Annals of the Entomological Society of America" pageId="14" pageNumber="15" pagination="295 - 314" refId="B23" refString="Tauber, CA, Tauber, MJ, Albuquerque, GS, 2014. Debris-carrying in larval Chrysopidae: unraveling its evolutionary history. . Annals of the Entomological Society of America 107: 295 - 314" title="Debris-carrying in larval Chrysopidae: unraveling its evolutionary history." url="https://doi.org/10.1603/AN13163" volume="107" year="2014">Tauber et al. 2014</bibRefCitation>
|
||
).
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Nothochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothochrysa" order="Neuroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="11" pageNumber="12">Nothochrysa</emphasis>
|
||
</taxonomicName>
|
||
larvae generally are considered debris-carriers, but their packets of debris are small, and their morphology is only moderately modified for debris-carrying. In addition, detailed information on aspects of the developmental and reproductive biology of
|
||
<taxonomicName lsidName="N. californica" pageId="11" pageNumber="12" rank="species" species="californica">
|
||
<emphasis italics="true" pageId="11" pageNumber="12">N. californica</emphasis>
|
||
</taxonomicName>
|
||
is available (
|
||
<bibRefCitation DOI="https://doi.org/10.3733/hilg.v36n11p391" author="Toschi, CA" journalOrPublisher="Hilgardia" pageId="14" pageNumber="15" pagination="391 - 433" refId="B26" refString="Toschi, CA, 1965. The taxonomy, life histories, and mating behavior of the green lacewings of Strawberry Canyon (Neuroptera, Chrysopidae). . Hilgardia 36: 391 - 433" title="The taxonomy, life histories, and mating behavior of the green lacewings of Strawberry Canyon (Neuroptera, Chrysopidae)." url="https://doi.org/10.3733/hilg.v36n11p391" volume="36" year="1965">Toschi 1965</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="11" pageNumber="12">
|
||
For generic-level comparisons, larval descriptions for genera within
|
||
<taxonomicName lsidName="" pageId="11" pageNumber="12" rank="subfamily" subfamily="Nothochrysinae">Nothochrysinae</taxonomicName>
|
||
(
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Kimochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kimochrysa" order="Neuroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="11" pageNumber="12">Kimochrysa</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Pimachrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pimachrysa" order="Neuroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="11" pageNumber="12">Pimachrysa</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Dictyochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dictyochrysa" order="Neuroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="11" pageNumber="12">Dictyochrysa</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Hypochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypochrysa" order="Neuroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="11" pageNumber="12">Hypochrysa</emphasis>
|
||
</taxonomicName>
|
||
) have been published (see
|
||
<bibRefCitation DOI="https://doi.org/10.1603/AN13163" author="Tauber, CA" journalOrPublisher="Annals of the Entomological Society of America" pageId="14" pageNumber="15" pagination="295 - 314" refId="B23" refString="Tauber, CA, Tauber, MJ, Albuquerque, GS, 2014. Debris-carrying in larval Chrysopidae: unraveling its evolutionary history. . Annals of the Entomological Society of America 107: 295 - 314" title="Debris-carrying in larval Chrysopidae: unraveling its evolutionary history." url="https://doi.org/10.1603/AN13163" volume="107" year="2014">Tauber et al. 2014</bibRefCitation>
|
||
). Unfortunately, larvae of
|
||
<emphasis italics="true" pageId="11" pageNumber="12">
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Asthenochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Asthenochrysa" order="Neuroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">Asthenochrysa</taxonomicName>
|
||
,
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Leptochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptochrysa" order="Neuroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">Leptochrysa</taxonomicName>
|
||
, Pamochrysa
|
||
</emphasis>
|
||
, and
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Triplochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Triplochrysa" order="Neuroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="11" pageNumber="12">Triplochrysa</emphasis>
|
||
</taxonomicName>
|
||
are not described.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="11" pageNumber="12" type="distribution">
|
||
<paragraph pageId="11" pageNumber="12">Known distribution.</paragraph>
|
||
<paragraph pageId="11" pageNumber="12">
|
||
Currently, this species has only been reported from the type locality, which presumably is the Valle Las Trancas in the region of
|
||
<normalizedToken originalValue="Ñuble">Nuble</normalizedToken>
|
||
, Chile.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="etymology">
|
||
<paragraph pageId="11" pageNumber="12">Etymology.</paragraph>
|
||
<paragraph pageId="11" pageNumber="12">This species is named in honor of Ronald G. Ehrenberg, Irving M. Ives Professor of Industrial and Labor Relations and Economics at Cornell University, an esteemed and cherished colleague of the author and her late husband (Maurice J. Tauber).</paragraph>
|
||
<subSection lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="characteristics shared with archaeochrysa species">
|
||
<paragraph pageId="11" pageNumber="12">
|
||
Characteristics shared with
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Archaeochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archaeochrysa" order="Neuroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="11" pageNumber="12">Archaeochrysa</emphasis>
|
||
</taxonomicName>
|
||
species
|
||
</paragraph>
|
||
<paragraph pageId="11" pageNumber="12">
|
||
As shown above,
|
||
<taxonomicName lsidName="N. ehrenbergi" pageId="11" pageNumber="12" rank="species" species="ehrenbergi">
|
||
<emphasis italics="true" pageId="11" pageNumber="12">N. ehrenbergi</emphasis>
|
||
</taxonomicName>
|
||
shares many features with other extant
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Nothochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothochrysa" order="Neuroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="11" pageNumber="12">Nothochrysa</emphasis>
|
||
</taxonomicName>
|
||
species, and its inclusion in the genus is well supported. However, the species also expresses many features that differ from
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Nothochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothochrysa" order="Neuroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="11" pageNumber="12">Nothochrysa</emphasis>
|
||
</taxonomicName>
|
||
and that are shared by at least some of the five species in the fossil genus
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Archaeochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archaeochrysa" order="Neuroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="11" pageNumber="12">Archaeochrysa</emphasis>
|
||
</taxonomicName>
|
||
. I discuss four below:
|
||
</paragraph>
|
||
<paragraph pageId="11" pageNumber="12">
|
||
First, in the
|
||
<taxonomicName lsidName="N. ehrenbergi" pageId="11" pageNumber="12" rank="species" species="ehrenbergi">
|
||
<emphasis italics="true" pageId="11" pageNumber="12">N. ehrenbergi</emphasis>
|
||
</taxonomicName>
|
||
forewing, vein A1 is not forked, whereas it is forked in all other
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Nothochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothochrysa" order="Neuroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="11" pageNumber="12">Nothochrysa</emphasis>
|
||
</taxonomicName>
|
||
species (
|
||
<bibRefCitation author="Adams, PA" journalOrPublisher="Bulletin of the Museum of Comparative Zoology" pageId="13" pageNumber="14" pagination="215 - 238" refId="B1" refString="Adams, PA, 1967. A review of the Mesochrysinae and Nothochrysinae (Neuroptera: Chrysopidae). . Bulletin of the Museum of Comparative Zoology 135: 215 - 238" title="A review of the Mesochrysinae and Nothochrysinae (Neuroptera: Chrysopidae)." volume="135" year="1967">Adams 1967</bibRefCitation>
|
||
;
|
||
<bibRefCitation pageId="11" pageNumber="12" refId="B4">
|
||
<normalizedToken originalValue="Aspöck">Aspoeck</normalizedToken>
|
||
et al. 1980
|
||
</bibRefCitation>
|
||
: figs 154, 155;
|
||
<bibRefCitation DOI="https://doi.org/10.1666/12-052R.1" author="Makarkin, VN" journalOrPublisher="Journal of Paleontology" pageId="14" pageNumber="15" pagination="123 - 146" refId="B16" refString="Makarkin, VN, Archibald, SB, 2013. A diverse new assemblage of green lacewings (Insecta, Neuroptera, Chrysopidae) from the Early Eocene Okanagan Highlands, western North America. . Journal of Paleontology 87: 123 - 146" title="A diverse new assemblage of green lacewings (Insecta, Neuroptera, Chrysopidae) from the Early Eocene Okanagan Highlands, western North America." url="https://doi.org/10.1666/12-052R.1" volume="87" year="2013">Makarkin and Archibald 2013</bibRefCitation>
|
||
: 135, 136). The feature is variable in
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Archaeochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archaeochrysa" order="Neuroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="11" pageNumber="12">Archaeochrysa</emphasis>
|
||
</taxonomicName>
|
||
specimens where A1 is visible. It is not forked in two species (
|
||
<bibRefCitation author="Adams, PA" journalOrPublisher="Bulletin of the Museum of Comparative Zoology" pageId="13" pageNumber="14" pagination="215 - 238" refId="B1" refString="Adams, PA, 1967. A review of the Mesochrysinae and Nothochrysinae (Neuroptera: Chrysopidae). . Bulletin of the Museum of Comparative Zoology 135: 215 - 238" title="A review of the Mesochrysinae and Nothochrysinae (Neuroptera: Chrysopidae)." volume="135" year="1967">Adams 1967</bibRefCitation>
|
||
: 237), forked in two species (
|
||
<bibRefCitation author="Adams, PA" journalOrPublisher="Bulletin of the Museum of Comparative Zoology" pageId="13" pageNumber="14" pagination="215 - 238" refId="B1" refString="Adams, PA, 1967. A review of the Mesochrysinae and Nothochrysinae (Neuroptera: Chrysopidae). . Bulletin of the Museum of Comparative Zoology 135: 215 - 238" title="A review of the Mesochrysinae and Nothochrysinae (Neuroptera: Chrysopidae)." volume="135" year="1967">Adams 1967</bibRefCitation>
|
||
: 230,
|
||
<bibRefCitation DOI="https://doi.org/10.1666/12-052R.1" author="Makarkin, VN" journalOrPublisher="Journal of Paleontology" pageId="14" pageNumber="15" pagination="123 - 146" refId="B16" refString="Makarkin, VN, Archibald, SB, 2013. A diverse new assemblage of green lacewings (Insecta, Neuroptera, Chrysopidae) from the Early Eocene Okanagan Highlands, western North America. . Journal of Paleontology 87: 123 - 146" title="A diverse new assemblage of green lacewings (Insecta, Neuroptera, Chrysopidae) from the Early Eocene Okanagan Highlands, western North America." url="https://doi.org/10.1666/12-052R.1" volume="87" year="2013">Makarkin and Archibald 2013</bibRefCitation>
|
||
: 135), and missing from the specimen of the fifth species (
|
||
<bibRefCitation DOI="https://doi.org/10.4039/tce.2014.53" author="Archibald, SB" journalOrPublisher="Canadian Entomologist" pageId="13" pageNumber="14" pagination="359 - 369" refId="B3" refString="Archibald, SB, Makarkin, VN, 2015. A new species of Archaeochrysa Adams (Neuroptera: Chrysopidae) from the early Eocene of Driftwood Canyon, British Columbia, Canada. . Canadian Entomologist 147: 359 - 369" title="A new species of Archaeochrysa Adams (Neuroptera: Chrysopidae) from the early Eocene of Driftwood Canyon, British Columbia, Canada." url="https://doi.org/10.4039/tce.2014.53" volume="147" year="2015">Archibald and Makarkin 2015</bibRefCitation>
|
||
: 363).
|
||
</paragraph>
|
||
<paragraph pageId="11" pageNumber="12">
|
||
Second, in
|
||
<taxonomicName lsidName="N. ehrenbergi" pageId="11" pageNumber="12" rank="species" species="ehrenbergi">
|
||
<emphasis italics="true" pageId="11" pageNumber="12">N. ehrenbergi</emphasis>
|
||
</taxonomicName>
|
||
the basal sc-r crossvein arises distal to the furcation of the radius and almost directly above the furcation of the media. Both of these character states are shared with the fossil genus
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Archaeochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archaeochrysa" order="Neuroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="11" pageNumber="12">Archaeochrysa</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation author="Adams, PA" journalOrPublisher="Bulletin of the Museum of Comparative Zoology" pageId="13" pageNumber="14" pagination="215 - 238" refId="B1" refString="Adams, PA, 1967. A review of the Mesochrysinae and Nothochrysinae (Neuroptera: Chrysopidae). . Bulletin of the Museum of Comparative Zoology 135: 215 - 238" title="A review of the Mesochrysinae and Nothochrysinae (Neuroptera: Chrysopidae)." volume="135" year="1967">Adams 1967</bibRefCitation>
|
||
,
|
||
<bibRefCitation DOI="https://doi.org/10.1666/12-052R.1" author="Makarkin, VN" journalOrPublisher="Journal of Paleontology" pageId="14" pageNumber="15" pagination="123 - 146" refId="B16" refString="Makarkin, VN, Archibald, SB, 2013. A diverse new assemblage of green lacewings (Insecta, Neuroptera, Chrysopidae) from the Early Eocene Okanagan Highlands, western North America. . Journal of Paleontology 87: 123 - 146" title="A diverse new assemblage of green lacewings (Insecta, Neuroptera, Chrysopidae) from the Early Eocene Okanagan Highlands, western North America." url="https://doi.org/10.1666/12-052R.1" volume="87" year="2013">Makarkin and Archibald 2013</bibRefCitation>
|
||
), but not with other known
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Nothochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothochrysa" order="Neuroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="11" pageNumber="12">Nothochrysa</emphasis>
|
||
</taxonomicName>
|
||
species.
|
||
</paragraph>
|
||
<paragraph pageId="11" pageNumber="12">
|
||
Third, in
|
||
<taxonomicName lsidName="N. ehrenbergi" pageId="11" pageNumber="12" rank="species" species="ehrenbergi">
|
||
<emphasis italics="true" pageId="11" pageNumber="12">N. ehrenbergi</emphasis>
|
||
</taxonomicName>
|
||
the distinction between the inner gradate series and the pseudomedia as well as between the outer gradate series and the pseudocubitus is indistinct. Rather, the gradate series and their respective pseudoveins tend to run together more smoothly as a curve, rather than at an angle as in other
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Nothochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothochrysa" order="Neuroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="11" pageNumber="12">Nothochrysa</emphasis>
|
||
</taxonomicName>
|
||
species. Again, this feature of
|
||
<taxonomicName lsidName="N. ehrenbergi" pageId="11" pageNumber="12" rank="species" species="ehrenbergi">
|
||
<emphasis italics="true" pageId="11" pageNumber="12">N. ehrenbergi</emphasis>
|
||
</taxonomicName>
|
||
is shared most closely with
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Archaeochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archaeochrysa" order="Neuroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="11" pageNumber="12">Archaeochrysa</emphasis>
|
||
</taxonomicName>
|
||
species (
|
||
<bibRefCitation author="Adams, PA" journalOrPublisher="Bulletin of the Museum of Comparative Zoology" pageId="13" pageNumber="14" pagination="215 - 238" refId="B1" refString="Adams, PA, 1967. A review of the Mesochrysinae and Nothochrysinae (Neuroptera: Chrysopidae). . Bulletin of the Museum of Comparative Zoology 135: 215 - 238" title="A review of the Mesochrysinae and Nothochrysinae (Neuroptera: Chrysopidae)." volume="135" year="1967">Adams 1967</bibRefCitation>
|
||
,
|
||
<bibRefCitation DOI="https://doi.org/10.1666/12-052R.1" author="Makarkin, VN" journalOrPublisher="Journal of Paleontology" pageId="14" pageNumber="15" pagination="123 - 146" refId="B16" refString="Makarkin, VN, Archibald, SB, 2013. A diverse new assemblage of green lacewings (Insecta, Neuroptera, Chrysopidae) from the Early Eocene Okanagan Highlands, western North America. . Journal of Paleontology 87: 123 - 146" title="A diverse new assemblage of green lacewings (Insecta, Neuroptera, Chrysopidae) from the Early Eocene Okanagan Highlands, western North America." url="https://doi.org/10.1666/12-052R.1" volume="87" year="2013">Makarkin and Archibald 2013</bibRefCitation>
|
||
,
|
||
<bibRefCitation DOI="https://doi.org/10.4039/tce.2014.53" author="Archibald, SB" journalOrPublisher="Canadian Entomologist" pageId="13" pageNumber="14" pagination="359 - 369" refId="B3" refString="Archibald, SB, Makarkin, VN, 2015. A new species of Archaeochrysa Adams (Neuroptera: Chrysopidae) from the early Eocene of Driftwood Canyon, British Columbia, Canada. . Canadian Entomologist 147: 359 - 369" title="A new species of Archaeochrysa Adams (Neuroptera: Chrysopidae) from the early Eocene of Driftwood Canyon, British Columbia, Canada." url="https://doi.org/10.4039/tce.2014.53" volume="147" year="2015">Archibald and Makarkin 2015</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
|
||
Fourth, currently the primary feature used to distinguish between
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Nothochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothochrysa" order="Neuroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="11" pageNumber="12">Nothochrysa</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Archaeochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archaeochrysa" order="Neuroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="11" pageNumber="12">Archaeochrysa</emphasis>
|
||
</taxonomicName>
|
||
is the presence or absence of a crossvein between RP and MA in the basal part of the hindwing. The crossvein is present in all known
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Archaeochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archaeochrysa" order="Neuroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="11" pageNumber="12">Archaeochrysa</emphasis>
|
||
</taxonomicName>
|
||
species and is reported to be absent from
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Nothochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothochrysa" order="Neuroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="11" pageNumber="12">Nothochrysa</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation DOI="https://doi.org/10.1666/12-052R.1" author="Makarkin, VN" journalOrPublisher="Journal of Paleontology" pageId="14" pageNumber="15" pagination="123 - 146" refId="B16" refString="Makarkin, VN, Archibald, SB, 2013. A diverse new assemblage of green lacewings (Insecta, Neuroptera, Chrysopidae) from the Early Eocene Okanagan Highlands, western North America. . Journal of Paleontology 87: 123 - 146" title="A diverse new assemblage of green lacewings (Insecta, Neuroptera, Chrysopidae) from the Early Eocene Okanagan Highlands, western North America." url="https://doi.org/10.1666/12-052R.1" volume="87" year="2013">Makarkin and Archibald 2013</bibRefCitation>
|
||
: 134).
|
||
<pageBreakToken pageId="12" pageNumber="13" start="start">In</pageBreakToken>
|
||
<taxonomicName lsidName="N. ehrenbergi" pageId="12" pageNumber="13" rank="species" species="ehrenbergi">
|
||
<emphasis italics="true" pageId="12" pageNumber="13">N. ehrenbergi</emphasis>
|
||
</taxonomicName>
|
||
, MA aligns with RP for about one-third the length of the upper margin of the
|
||
<emphasis italics="true" pageId="12" pageNumber="13">im1</emphasis>
|
||
cell, and no crossvein is present (
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Nothochrysa ehrenbergi sp. nov. (Nuble, Chile; Male, CAS): Venation at base of wings (a) left forewing, (b) left hindwing. Note the absence of a tympanal organ at the base of R in the forewing, the independent origin and trajectory of M along the base of R (arrows pointing downward, both wings), and the alignment of RP and MA in the hindwing. A 1, A 2, A 3 first, second, third anal veins Cu cubitus Cu f furcation (division) of cubitus Ju jugal lobe M media MA media anterior m-cu media-cubital crossvein R radius RP radius posterior." figureDoi="10.3897/zookeys.866.35394.figure1" httpUri="https://binary.pensoft.net/fig/319506" pageId="12" pageNumber="13">Figs 1b</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nothochrysa ehrenbergi sp. nov. (Nuble, Chile; Male, CAS): Wings with selected features labeled (a) left forewing, (b) left hindwing. Marginal traces of major veins demarcated; arrow (hindwing) indicates alignment of RP and MA along upper margin of first intramedian cell. A 1, A 2, A 3 first, second, third anal veins CuA, CuP anterior, posterior branches of cubitus icu 1, icu 3 first, third intracubital cells ig inner gradate im 1, im 2 first, second intramedian cells Ju jugal lobe MA media anterior MP media posterior mcua, mpcua second and third medial cells M f furcation of media og outer gradate Psc pseudocubitus Psm pseudomedia R f furcation of radius RP radius posterior RP 1 first branch of radius posterior 1 sc-r first crossvein between subcosta and radius 2 m-cu second crossvein between media and cubitus." figureDoi="10.3897/zookeys.866.35394.figure2" httpUri="https://binary.pensoft.net/fig/319507" pageId="12" pageNumber="13">2b</figureCitation>
|
||
). However, even with this character there appears to be a possible exception.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nothochrysa ehrenbergi sp. nov. (Nuble, Chile; Male, CAS): Wings with selected features labeled (a) left forewing, (b) left hindwing. Marginal traces of major veins demarcated; arrow (hindwing) indicates alignment of RP and MA along upper margin of first intramedian cell. A 1, A 2, A 3 first, second, third anal veins CuA, CuP anterior, posterior branches of cubitus icu 1, icu 3 first, third intracubital cells ig inner gradate im 1, im 2 first, second intramedian cells Ju jugal lobe MA media anterior MP media posterior mcua, mpcua second and third medial cells M f furcation of media og outer gradate Psc pseudocubitus Psm pseudomedia R f furcation of radius RP radius posterior RP 1 first branch of radius posterior 1 sc-r first crossvein between subcosta and radius 2 m-cu second crossvein between media and cubitus." figureDoi="10.3897/zookeys.866.35394.figure2" httpUri="https://binary.pensoft.net/fig/319507" pageId="12" pageNumber="13">Figure 2</figureCitation>
|
||
accompanying the original description of
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Nothochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothochrysa turcica" order="Neuroptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="turcica">
|
||
<emphasis italics="true" pageId="12" pageNumber="13">Nothochrysa turcica</emphasis>
|
||
</taxonomicName>
|
||
Kovanci and Canbulat shows a short crossvein between RP and MA; confirmation of the accuracy of this drawing is necessary.
|
||
</paragraph>
|
||
</subSection>
|
||
<subSection pageId="12" pageNumber="13" type="phylogenetic position of nothochrysa ehrenbergi sp. nov">
|
||
<paragraph pageId="12" pageNumber="13">
|
||
Phylogenetic position of
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Nothochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothochrysa ehrenbergi" order="Neuroptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="ehrenbergi">
|
||
<emphasis italics="true" pageId="12" pageNumber="13">Nothochrysa ehrenbergi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
</paragraph>
|
||
<paragraph pageId="12" pageNumber="13">
|
||
Given the above,
|
||
<bibRefCitation DOI="https://doi.org/10.4039/tce.2014.53" author="Archibald, SB" journalOrPublisher="Canadian Entomologist" pageId="13" pageNumber="14" pagination="359 - 369" refId="B3" refString="Archibald, SB, Makarkin, VN, 2015. A new species of Archaeochrysa Adams (Neuroptera: Chrysopidae) from the early Eocene of Driftwood Canyon, British Columbia, Canada. . Canadian Entomologist 147: 359 - 369" title="A new species of Archaeochrysa Adams (Neuroptera: Chrysopidae) from the early Eocene of Driftwood Canyon, British Columbia, Canada." url="https://doi.org/10.4039/tce.2014.53" volume="147" year="2015">
|
||
Archibald and
|
||
<normalizedToken originalValue="Makarkin’s">Makarkin's</normalizedToken>
|
||
(2015)
|
||
</bibRefCitation>
|
||
discussion of the phylogeny of
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Archaeochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archaeochrysa" order="Neuroptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="12" pageNumber="13">Archaeochrysa</emphasis>
|
||
</taxonomicName>
|
||
species is worthy of consideration here. Their paper evaluates how the various
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Archaeochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archaeochrysa" order="Neuroptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="12" pageNumber="13">Archaeochrysa</emphasis>
|
||
</taxonomicName>
|
||
species express three features; each feature has several conditions ranging from presumably plesiomorphic to more derived. Below, the three features are considered, relative to their expression by
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Nothochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothochrysa" order="Neuroptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="12" pageNumber="13">Nothochrysa</emphasis>
|
||
</taxonomicName>
|
||
species, especially
|
||
<taxonomicName lsidName="N. ehrenbergi" pageId="12" pageNumber="13" rank="species" species="ehrenbergi">
|
||
<emphasis italics="true" pageId="12" pageNumber="13">N. ehrenbergi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="12" pageNumber="13">
|
||
(1)
|
||
<emphasis bold="true" pageId="12" pageNumber="13">
|
||
The shape of the
|
||
<emphasis italics="true" pageId="12" pageNumber="13">im1</emphasis>
|
||
cell.
|
||
</emphasis>
|
||
<bibRefCitation DOI="https://doi.org/10.4039/tce.2014.53" author="Archibald, SB" journalOrPublisher="Canadian Entomologist" pageId="13" pageNumber="14" pagination="359 - 369" refId="B3" refString="Archibald, SB, Makarkin, VN, 2015. A new species of Archaeochrysa Adams (Neuroptera: Chrysopidae) from the early Eocene of Driftwood Canyon, British Columbia, Canada. . Canadian Entomologist 147: 359 - 369" title="A new species of Archaeochrysa Adams (Neuroptera: Chrysopidae) from the early Eocene of Driftwood Canyon, British Columbia, Canada." url="https://doi.org/10.4039/tce.2014.53" volume="147" year="2015">Archibald and Makarkin (2015)</bibRefCitation>
|
||
describe two configurations for this character;
|
||
<taxonomicName lsidName="N. ehrenbergi" pageId="12" pageNumber="13" rank="species" species="ehrenbergi">
|
||
<emphasis italics="true" pageId="12" pageNumber="13">N. ehrenbergi</emphasis>
|
||
</taxonomicName>
|
||
expresses the second (more advanced) condition in which the sides of the
|
||
<emphasis italics="true" pageId="12" pageNumber="13">im1</emphasis>
|
||
cell are almost parallel for most of their span and converge basally at a relatively steep angle. The extant species of
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Nothochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothochrysa" order="Neuroptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="12" pageNumber="13">Nothochrysa</emphasis>
|
||
</taxonomicName>
|
||
, including
|
||
<taxonomicName lsidName="N. ehrenbergi" pageId="12" pageNumber="13" rank="species" species="ehrenbergi">
|
||
<emphasis italics="true" pageId="12" pageNumber="13">N. ehrenbergi</emphasis>
|
||
</taxonomicName>
|
||
, share this feature with two species of
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Archaeochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archaeochrysa" order="Neuroptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="12" pageNumber="13">Archaeochrysa</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="12" pageNumber="13">
|
||
(2)
|
||
<emphasis bold="true" pageId="12" pageNumber="13">The position of crossvein 2m-cu.</emphasis>
|
||
<bibRefCitation DOI="https://doi.org/10.4039/tce.2014.53" author="Archibald, SB" journalOrPublisher="Canadian Entomologist" pageId="13" pageNumber="14" pagination="359 - 369" refId="B3" refString="Archibald, SB, Makarkin, VN, 2015. A new species of Archaeochrysa Adams (Neuroptera: Chrysopidae) from the early Eocene of Driftwood Canyon, British Columbia, Canada. . Canadian Entomologist 147: 359 - 369" title="A new species of Archaeochrysa Adams (Neuroptera: Chrysopidae) from the early Eocene of Driftwood Canyon, British Columbia, Canada." url="https://doi.org/10.4039/tce.2014.53" volume="147" year="2015">Archibald and Makarkin (2015)</bibRefCitation>
|
||
list six conditions for this character, each one considered more evolutionarily advanced than the preceding.
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Nothochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothochrysa ehrenbergi" order="Neuroptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="ehrenbergi">
|
||
<emphasis italics="true" pageId="12" pageNumber="13">Nothochrysa ehrenbergi</emphasis>
|
||
</taxonomicName>
|
||
falls into Condition 5, a derived condition in which 2m-cu is located distinctly in the proximal part of
|
||
<emphasis italics="true" pageId="12" pageNumber="13">im1</emphasis>
|
||
(as shown in fig. 2C of
|
||
<bibRefCitation DOI="https://doi.org/10.4039/tce.2014.53" author="Archibald, SB" journalOrPublisher="Canadian Entomologist" pageId="13" pageNumber="14" pagination="359 - 369" refId="B3" refString="Archibald, SB, Makarkin, VN, 2015. A new species of Archaeochrysa Adams (Neuroptera: Chrysopidae) from the early Eocene of Driftwood Canyon, British Columbia, Canada. . Canadian Entomologist 147: 359 - 369" title="A new species of Archaeochrysa Adams (Neuroptera: Chrysopidae) from the early Eocene of Driftwood Canyon, British Columbia, Canada." url="https://doi.org/10.4039/tce.2014.53" volume="147" year="2015">Archibald and Makarkin 2015</bibRefCitation>
|
||
). This character state is typical of at least two
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Archaeochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archaeochrysa" order="Neuroptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="12" pageNumber="13">Archaeochrysa</emphasis>
|
||
</taxonomicName>
|
||
species,
|
||
<taxonomicName lsidName="A. creedi" pageId="12" pageNumber="13" rank="species" species="creedi">
|
||
<emphasis italics="true" pageId="12" pageNumber="13">A. creedi</emphasis>
|
||
</taxonomicName>
|
||
(Adams) and
|
||
<taxonomicName lsidName="A. paranervis" pageId="12" pageNumber="13" rank="species" species="paranervis">
|
||
<emphasis italics="true" pageId="12" pageNumber="13">A. paranervis</emphasis>
|
||
</taxonomicName>
|
||
(Adams), as well as several other extant genera in
|
||
<taxonomicName lsidName="" pageId="12" pageNumber="13" rank="subfamily" subfamily="Nothochrysinae">Nothochrysinae</taxonomicName>
|
||
, including
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Nothochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothochrysa" order="Neuroptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="12" pageNumber="13">Nothochrysa</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="12" pageNumber="13">
|
||
(3)
|
||
<emphasis bold="true" pageId="12" pageNumber="13">The crossveins of Psc.</emphasis>
|
||
<bibRefCitation DOI="https://doi.org/10.4039/tce.2014.53" author="Archibald, SB" journalOrPublisher="Canadian Entomologist" pageId="13" pageNumber="14" pagination="359 - 369" refId="B3" refString="Archibald, SB, Makarkin, VN, 2015. A new species of Archaeochrysa Adams (Neuroptera: Chrysopidae) from the early Eocene of Driftwood Canyon, British Columbia, Canada. . Canadian Entomologist 147: 359 - 369" title="A new species of Archaeochrysa Adams (Neuroptera: Chrysopidae) from the early Eocene of Driftwood Canyon, British Columbia, Canada." url="https://doi.org/10.4039/tce.2014.53" volume="147" year="2015">Archibald and Makarkin (2015</bibRefCitation>
|
||
: 366) describe and illustrate four character states for this feature; interested readers are referred to the original paper. Suffice it to say here,
|
||
<taxonomicName lsidName="N. ehrenbergi" pageId="12" pageNumber="13" rank="species" species="ehrenbergi">
|
||
<emphasis italics="true" pageId="12" pageNumber="13">N. ehrenbergi</emphasis>
|
||
</taxonomicName>
|
||
, as well as three
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Archaeochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archaeochrysa" order="Neuroptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="12" pageNumber="13">Archaeochrysa</emphasis>
|
||
</taxonomicName>
|
||
species but no other
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Nothochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothochrysa" order="Neuroptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="12" pageNumber="13">Nothochrysa</emphasis>
|
||
</taxonomicName>
|
||
species, fall into the second of the four conditions. This position is considered plesiomorphic among
|
||
<taxonomicName lsidName="" pageId="12" pageNumber="13" rank="subfamily" subfamily="Nothochrysinae">Nothochrysinae</taxonomicName>
|
||
, both fossil and extant (
|
||
<bibRefCitation DOI="https://doi.org/10.4039/tce.2014.53" author="Archibald, SB" journalOrPublisher="Canadian Entomologist" pageId="13" pageNumber="14" pagination="359 - 369" refId="B3" refString="Archibald, SB, Makarkin, VN, 2015. A new species of Archaeochrysa Adams (Neuroptera: Chrysopidae) from the early Eocene of Driftwood Canyon, British Columbia, Canada. . Canadian Entomologist 147: 359 - 369" title="A new species of Archaeochrysa Adams (Neuroptera: Chrysopidae) from the early Eocene of Driftwood Canyon, British Columbia, Canada." url="https://doi.org/10.4039/tce.2014.53" volume="147" year="2015">Archibald and Makarkin 2015</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="12" pageNumber="13">
|
||
On the basis of the above information, it appears that
|
||
<taxonomicName lsidName="N. ehrenbergi" pageId="12" pageNumber="13" rank="species" species="ehrenbergi">
|
||
<emphasis italics="true" pageId="12" pageNumber="13">N. ehrenbergi</emphasis>
|
||
</taxonomicName>
|
||
shares a very close phylogenetic relationship with the fossil genus
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Archaeochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archaeochrysa" order="Neuroptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="12" pageNumber="13">Archaeochrysa</emphasis>
|
||
</taxonomicName>
|
||
. At this point, only one character (the absence of a crossvein between the RP and the MA above the first intramedial cell of the hindwing) supports its exclusion from
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Archaeochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archaeochrysa" order="Neuroptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="12" pageNumber="13">Archaeochrysa</emphasis>
|
||
</taxonomicName>
|
||
, and this character may have exceptions within
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Nothochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothochrysa" order="Neuroptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="12" pageNumber="13">Nothochrysa</emphasis>
|
||
</taxonomicName>
|
||
. Indeed, there does not appear to be a synapomorphic character that consistently differentiates
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Nothochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothochrysa" order="Neuroptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="12" pageNumber="13">Nothochrysa</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Archaeochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archaeochrysa" order="Neuroptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="12" pageNumber="13">Archaeochrysa</emphasis>
|
||
</taxonomicName>
|
||
. Thus, given the overall similarity between
|
||
<taxonomicName lsidName="N. ehrenbergi" pageId="12" pageNumber="13" rank="species" species="ehrenbergi">
|
||
<emphasis italics="true" pageId="12" pageNumber="13">N. ehrenbergi</emphasis>
|
||
</taxonomicName>
|
||
and the known
|
||
<taxonomicName class="Insecta" family="Chrysopidae" genus="Archaeochrysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archaeochrysa" order="Neuroptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="12" pageNumber="13">Archaeochrysa</emphasis>
|
||
</taxonomicName>
|
||
species, I recommend that future studies examine the validity of maintaining the generic separation.
|
||
</paragraph>
|
||
</subSection>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |