treatments-xml/data/4C/55/C4/4C55C448E13B900D2CAB428E2E6A9196.xml
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<document id="2D23B72B2F360B66882F28609E7FFFF2" ID-CLB-Dataset="31401" ID-DOI="10.11646/zootaxa.4085.2.3" ID-GBIF-Dataset="cee977ff-1011-4121-882d-775b50e41957" ID-ISSN="1175-5326" ID-Zenodo-Dep="1053424" ID-ZooBank="E153BEED-8C53-482F-B70A-25554AEC2D17" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1510909618376" checkinUser="plazi" docAuthor="Yuan, Zhi-Yong, Zhang, Bao-Lin &amp; Che, Jing" docDate="2016" docId="4C55C448E13B900D2CAB428E2E6A9196" docLanguage="en" docName="zootaxa.4085.2.3.pdf" docOrigin="Zootaxa 4085 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pachytriton wuguanfui Yuan &amp; Zhang &amp; Che 2016, sp. nov." docType="treatment" docVersion="8" lastPageNumber="230" masterDocId="B06CBC30E13390062C3C42192B08941B" masterDocTitle="A new species of the genus Pachytriton (Caudata: Salamandridae) from Hunan and Guangxi, southeastern China" pageNumber="227" updateTime="1698500494417" updateUser="ExternalLinkService">
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<mods:title id="7C4873E426F005ACEC704D929A0B7C35">A new species of the genus Pachytriton (Caudata: Salamandridae) from Hunan and Guangxi, southeastern China</mods:title>
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<mods:namePart id="DE085CF03FDE472173239C7A76426867">Yuan, Zhi-Yong</mods:namePart>
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<mods:namePart id="41560EE8EBCC8F7666BC39843097F973">Zhang, Bao-Lin</mods:namePart>
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<mods:namePart id="7F1DE21A0CA4A72C652416FF543854CC">Che, Jing</mods:namePart>
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<mods:date id="9FB0B2884F6E5E00B3D8EE26D196F9A8">2016</mods:date>
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<treatment id="4C55C448E13B900D2CAB428E2E6A9196" ID-DOI="http://doi.org/10.5281/zenodo.5621648" ID-GBIF-Taxon="137350574" ID-Zenodo-Dep="5621648" LSID="urn:lsid:plazi:treatment:4C55C448E13B900D2CAB428E2E6A9196" httpUri="http://treatment.plazi.org/id/4C55C448E13B900D2CAB428E2E6A9196" lastPageId="11" lastPageNumber="230" pageId="8" pageNumber="227">
<subSubSection id="8CE626D5E13B900E2CAB428E291C94A9" box="[151,532,151,178]" pageId="8" pageNumber="227" type="nomenclature">
<paragraph id="C443755EE13B900E2CAB428E291C94A9" blockId="8.[151,532,151,211]" box="[151,532,151,178]" pageId="8" pageNumber="227">
<heading id="9F0BC232E13B900E2CAB428E291C94A9" bold="true" box="[151,532,151,178]" fontSize="11" level="1" pageId="8" pageNumber="227" reason="1">
<emphasis id="F688A94CE13B900E2CAB428E291C94A9" bold="true" box="[151,532,151,178]" pageId="8" pageNumber="227">
<taxonomicName id="03FC0EDDE13B900E2CAB428E2AB894AA" ID-CoL="75J9H" authority="Yuan &amp; Zhang &amp; Che, 2016" authorityName="Yuan &amp; Zhang &amp; Che" authorityYear="2016" box="[151,432,151,177]" class="Amphibia" family="Salamandridae" genus="Pachytriton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="227" phylum="Chordata" rank="species" species="wuguanfui" status="sp. nov.">
<emphasis id="F688A94CE13B900E2CAB428E2AB894AA" bold="true" box="[151,432,151,177]" italics="true" pageId="8" pageNumber="227">Pachytriton wuguanfui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EDBB1437E13B900E2D8B4281291C94A9" box="[439,532,152,178]" pageId="8" pageNumber="227" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<paragraph id="C443755EE13B900E2CAB42A32A7B94C8" blockId="8.[151,532,151,211]" box="[151,371,186,211]" pageId="8" pageNumber="227">
(
<subSubSection id="8CE626D5E13B900E2CA342A32A7B94C8" box="[159,371,186,211]" pageId="8" pageNumber="227" type="description">
<figureCitation id="5CC769DBE13B900E2CA342A32A0394C8" box="[159,267,186,211]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStart-3="FIGURE 7" captionStartId-0="8.[151,250,1997,2019]" captionStartId-1="9.[151,250,1159,1181]" captionStartId-2="9.[151,250,1900,1922]" captionStartId-3="10.[151,250,1902,1924]" captionTargetBox-0="[151,1436,767,1975]" captionTargetBox-1="[170,1431,211,1138]" captionTargetBox-2="[151,1436,1262,1879]" captionTargetBox-3="[151,1436,1021,1881]" captionTargetId-0="figure@8.[151,1436,767,1975]" captionTargetId-1="figure@9.[151,1435,193,1138]" captionTargetId-2="figure@9.[151,1436,1262,1879]" captionTargetId-3="figure@10.[151,1436,1021,1881]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="9" captionTargetPageId-3="10" captionText-0="FIGURE 4. Dorsal (A) and ventral (B) views of living female Pachytriton wuguanfui sp. nov. (KIZ 08755)." captionText-1="FIGURE 5. Dorsal (A) and ventral (B) views of the holotype of Pachytriton wuguanfui sp. nov. (KIZ 08758) in preservation. The scale shows 20 mm." captionText-2="FIGURE 6. Ventral views of (A) forelimb and (B) hindlimb of the holotype of Pachytriton wuguanfui sp. nov. (KIZ 08758) in preservation." captionText-3="FIGURE 7. Habitat at the type locality of Pachytriton wuguanfui sp. nov., Mt. Gupo, Hezhou, Guangxi, China." httpUri-0="https://zenodo.org/record/1053434/files/figure.png" httpUri-1="https://zenodo.org/record/1053436/files/figure.png" httpUri-2="https://zenodo.org/record/1053438/files/figure.png" httpUri-3="https://zenodo.org/record/1053442/files/figure.png" pageId="8" pageNumber="227">Figs. 47</figureCitation>
,
<tableCitation id="897E40E5E13B900E2D2A42A32A6294C8" box="[278,362,186,211]" captionStart="TABLE 2" captionStartId="6.[151,239,155,178]" captionText="TABLE 2. Linear measurements (in mm) of Pachytriton wuguanfui sp. nov. Abbreviations correspond to the text." pageId="8" pageNumber="227">Table 2</tableCitation>
)
</subSubSection>
</paragraph>
<subSubSection id="8CE626D5E13B900E2CAB431B284595D4" pageId="8" pageNumber="227" type="materials_examined">
<paragraph id="C443755EE13B900E2CAB431B2FB69524" blockId="8.[151,1436,258,715]" pageId="8" pageNumber="227">
<materialsCitation id="74947F03E13B900E2CAB431B2FB69524" ID-GBIF-Occurrence="1701202648" collectingDate="2010-08-17" collectionCode="KIZ" collectorName="Zhiyong Yuan" country="China" elevation="1202" latitude="24.64" location="Mt. Gupo, Hezhou city" longLatPrecision="750" longitude="111.53" pageId="8" pageNumber="227" specimenCode="KIZ08758" specimenCount="2" specimenCount-adult="1" specimenCount-female="1" stateProvince="Guangxi Zhuang Autonomous Region" typeStatus="holotype">
<emphasis id="F688A94CE13B900E2CAB431B2A039500" bold="true" box="[151,267,258,283]" pageId="8" pageNumber="227">
<typeStatus id="1B47CBFCE13B900E2CAB431B2A0E9500" box="[151,262,258,283]" pageId="8" pageNumber="227" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="945ADD25E13B900E2D2D431B2A8E9500" box="[273,390,258,283]" pageId="8" pageNumber="227">KIZ08758</specimenCode>
(
<figureCitation id="5CC769DBE13B900E2DA9431B2AD09500" box="[405,472,258,283]" captionStart="FIGURE 5" captionStartId="9.[151,250,1159,1181]" captionTargetBox="[170,1431,211,1138]" captionTargetId="figure@9.[151,1435,193,1138]" captionTargetPageId="9" captionText="FIGURE 5. Dorsal (A) and ventral (B) views of the holotype of Pachytriton wuguanfui sp. nov. (KIZ 08758) in preservation. The scale shows 20 mm." httpUri="https://zenodo.org/record/1053436/files/figure.png" pageId="8" pageNumber="227">Fig. 5</figureCitation>
), an
<specimenCount id="D2FABED7E13B900E2E32431B294E9500" box="[526,582,258,283]" pageId="8" pageNumber="227" type="adult">adult</specimenCount>
<specimenCount id="D2FABED7E13B900E2E71431B29919500" box="[589,665,258,283]" pageId="8" pageNumber="227" type="female">female</specimenCount>
from
<location id="C1232385E13B900E2EE1431A28D79500" LSID="urn:lsid:plazi:treatment:4C55C448E13B900D2CAB428E2E6A9196:C1232385E13B900E2EE1431A28D79500" box="[733,991,258,283]" country="China" latitude="24.64" longLatPrecision="750" longitude="111.53" name="Mt. Gupo, Hezhou city" pageId="8" pageNumber="227" stateProvince="Guangxi Zhuang Autonomous Region">Mt. Gupo, Hezhou city</location>
,
<collectingRegion id="0638BBBCE13B900E2FD6431B2E909500" box="[1002,1432,258,283]" pageId="8" pageNumber="227">Guangxi Zhuang Autonomous Region</collectingRegion>
,
<collectingCountry id="BCEB35CEE13B900E2CAB433F2BD69524" box="[151,222,294,319]" name="China" pageId="8" pageNumber="227">China</collectingCountry>
;
<geoCoordinate id="A1C81399E13B900E2CD4433F2A469525" box="[232,334,294,319]" direction="north" orientation="latitude" pageId="8" pageNumber="227" precision="555" value="24.64">24.64° N</geoCoordinate>
,
<geoCoordinate id="A1C81399E13B900E2D65433F2AC09525" box="[345,456,294,319]" direction="east" orientation="longitude" pageId="8" pageNumber="227" precision="555" value="111.53">111.53° E</geoCoordinate>
, elevation
<elevation id="4FD1926DE13B900E2E7D433E29909525" box="[577,664,295,319]" metricMagnitude="3" metricUnit="m" metricValue="1.202" pageId="8" pageNumber="227" unit="m" value="1202.0">
<quantity id="0304D8BBE13B900E2E7D433E29909525" box="[577,664,295,319]" metricMagnitude="3" metricUnit="m" metricValue="1.202" pageId="8" pageNumber="227" unit="m" value="1202.0">1202 m</quantity>
</elevation>
, collected by
<collectorName id="69091088E13B900E2F0F433F28DB9525" box="[819,979,294,319]" pageId="8" pageNumber="227">Zhiyong Yuan</collectorName>
on
<collectingDate id="A006AA76E13B900E2FC0433E2FB29524" box="[1020,1210,295,319]" pageId="8" pageNumber="227" value="2010-08-17">
<date id="B042539EE13B900E2FC0433E2FB29524" box="[1020,1210,295,319]" pageId="8" pageNumber="227" value="2010-08-17">
<date id="B042539EE13B900E2FC0433E2F7A9524" box="[1020,1138,295,319]" day="17" month="08" monthName="August" pageId="8" pageNumber="227" year="1917">August 17</date>
,
<date id="B042539EE13B900E2842433E2FB29524" box="[1150,1210,295,319]" pageId="8" pageNumber="227" value="2010" year="2010">2010</date>
</date>
</collectingDate>
.
</materialsCitation>
</paragraph>
<paragraph id="C443755EE13B900E2CFB4353284595D4" blockId="8.[151,1436,258,715]" pageId="8" pageNumber="227">
<emphasis id="F688A94CE13B900E2CFB43532A4E9578" bold="true" box="[199,326,330,355]" pageId="8" pageNumber="227">
<typeStatus id="1B47CBFCE13B900E2CFB43532A499578" box="[199,321,330,355]" pageId="8" pageNumber="227" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="A2EDED9BE13B900E2D6C43532A779578" box="[336,383,330,355]" country="China" httpUri="http://grbio.org/cool/ncht-ribt" name="Kunming Institute of Zoology, Chinese Academy of Sciences" pageId="8" pageNumber="227">KIZ</collectionCode>
0875508757 (three females), 0 8759 (male), collected at the same time and locality as the
<typeStatus id="1B47CBFCE13B900E2CAB43772BF5959C" box="[151,253,366,391]" pageId="8" pageNumber="227" type="holotype">holotype</typeStatus>
.
<collectionCode id="A2EDED9BE13B900E2D3443772A3F959C" box="[264,311,366,391]" country="China" httpUri="http://grbio.org/cool/ncht-ribt" name="Kunming Institute of Zoology, Chinese Academy of Sciences" pageId="8" pageNumber="227">KIZ</collectionCode>
<date id="B042539EE13B900E2D0A43772A82959C" box="[310,394,366,391]" pageId="8" pageNumber="227" value="05-02-17">021705</date>
06 (two females), 0 21708 (female), 0 21711 (female), 0 21712 (male),
<collectionCode id="A2EDED9BE13B900E289043762FD2959C" box="[1196,1242,367,391]" country="China" httpUri="http://grbio.org/cool/ncht-ribt" name="Kunming Institute of Zoology, Chinese Academy of Sciences" pageId="8" pageNumber="227">KIZ</collectionCode>
<date id="B042539EE13B900E28E643762E27959C" box="[1242,1327,367,391]" pageId="8" pageNumber="227" value="32-02-18">021832</date>
(female),
<date id="B042539EE13B900E2CAB438B2BE495B0" box="[151,236,402,427]" pageId="8" pageNumber="227" value="33-02-18">021833</date>
35 (three males) from Chahuaping, Dao country,
<collectingRegion id="0638BBBCE13B900E2F15438A28EA95B0" box="[809,994,402,427]" country="China" name="Hunan" pageId="8" pageNumber="227">Hunan province</collectingRegion>
,
<geoCoordinate id="A1C81399E13B900E2FD2438B2F5D95B1" box="[1006,1109,402,427]" direction="north" orientation="latitude" pageId="8" pageNumber="227" precision="555" value="25.22">25.22° N</geoCoordinate>
,
<geoCoordinate id="A1C81399E13B900E285E438A2FD995B1" box="[1122,1233,403,427]" direction="east" orientation="longitude" pageId="8" pageNumber="227" precision="555" value="111.79">111.79° E</geoCoordinate>
, elevation
<quantity id="0304D8BBE13B900E2971438B2E9F95B1" box="[1357,1431,402,427]" metricMagnitude="2" metricUnit="m" metricValue="6.49" pageId="8" pageNumber="227" unit="m" value="649.0">649 m</quantity>
, collected by Zhiyong Yuan and Limin Ding on
<date id="B042539EE13B900E2E9543AE284295D4" box="[681,842,439,463]" pageId="8" pageNumber="227" value="2012-06-18">June 18, 2012</date>
.
</paragraph>
</subSubSection>
<subSubSection id="8CE626D5E13B900E2CFB43C32E0796D1" pageId="8" pageNumber="227" type="diagnosis">
<paragraph id="C443755EE13B900E2CFB43C32E0796D1" blockId="8.[151,1436,258,715]" pageId="8" pageNumber="227">
<emphasis id="F688A94CE13B900E2CFB43C32A4995E8" bold="true" box="[199,321,474,499]" pageId="8" pageNumber="227">Diagnosis.</emphasis>
<taxonomicName id="03FC0EDDE13B900E2D7443C3294295E8" authority="Yuan &amp; Zhang &amp; Che, 2016" authorityName="Yuan &amp; Zhang &amp; Che" authorityYear="2016" box="[328,586,474,499]" class="Amphibia" family="Salamandridae" genus="Pachytriton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="227" phylum="Chordata" rank="species" species="wuguanfui" status="sp. nov.">
<emphasis id="F688A94CE13B900E2D7443C3294295E8" box="[328,586,474,499]" italics="true" pageId="8" pageNumber="227">Pachytriton wuguanfui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EDBB1437E13B900E2E6D43C329A395E8" box="[593,683,474,499]" pageId="8" pageNumber="227" rank="species">
<emphasis id="F688A94CE13B900E2E6D43C329A395E8" bold="true" box="[593,683,474,499]" pageId="8" pageNumber="227">sp. nov.</emphasis>
</taxonomicNameLabel>
is assigned to the genus
<taxonomicName id="03FC0EDDE13B900E2FFF43C32F4F95E8" box="[963,1095,474,499]" class="Amphibia" family="Salamandridae" genus="Pachytriton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="227" phylum="Chordata" rank="genus">
<emphasis id="F688A94CE13B900E2FFF43C32F4F95E8" box="[963,1095,474,499]" italics="true" pageId="8" pageNumber="227">Pachytriton</emphasis>
</taxonomicName>
by its molecular phylogenetic position and the following morphological characters: head oval and flat; skin very smooth; labial folds obvious on upper jaw; limbs short; tips of fore- and hindlimbs widely separated when limbs adpressed against body flank; tail long, base broad; posterior half of tail gradually laterally compressed; dorsal caudal fin evident. This species can be differentiated from congeners by the following combination of morphological characters: dorsum uniformly dark brown without bright orange dots and black spots over the body; tips of fingers and toes orange on dorsal side; costal grooves between axilla and groin not discernable; Ʌ-shaped vomerine tooth series widely open.
</paragraph>
</subSubSection>
<caption id="908325D6E13B900E2CAB45D42FD593F8" box="[151,1245,1997,2019]" httpUri="https://zenodo.org/record/1053434/files/figure.png" pageId="8" pageNumber="227" targetBox="[151,1436,767,1975]" targetPageId="8">
<paragraph id="C443755EE13B900E2CAB45D42FD593F8" blockId="8.[151,1245,1997,2019]" box="[151,1245,1997,2019]" pageId="8" pageNumber="227">
<emphasis id="F688A94CE13B900E2CAB45D42A1A93F8" bold="true" box="[151,274,1997,2019]" pageId="8" pageNumber="227">FIGURE 4.</emphasis>
Dorsal (A) and ventral (B) views of living female
<taxonomicName id="03FC0EDDE13B900E2F2945D428F693F8" authority="Yuan &amp; Zhang &amp; Che, 2016" authorityName="Yuan &amp; Zhang &amp; Che" authorityYear="2016" box="[789,1022,1997,2019]" class="Amphibia" family="Salamandridae" genus="Pachytriton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="227" phylum="Chordata" rank="species" species="wuguanfui" status="sp. nov.">
<emphasis id="F688A94CE13B900E2F2945D428F693F8" box="[789,1022,1997,2019]" italics="true" pageId="8" pageNumber="227">Pachytriton wuguanfui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EDBB1437E13B900E283845D42F5D93F8" box="[1028,1109,1997,2019]" pageId="8" pageNumber="227" rank="species">
<emphasis id="F688A94CE13B900E283845D42F5D93F8" bold="true" box="[1028,1109,1997,2019]" pageId="8" pageNumber="227">sp. nov.</emphasis>
</taxonomicNameLabel>
(KIZ08755).
</paragraph>
</caption>
<caption id="908325D6E13A900F2CAB469E2A9990A7" httpUri="https://zenodo.org/record/1053436/files/figure.png" pageId="9" pageNumber="228" targetBox="[170,1431,211,1138]" targetPageId="9">
<paragraph id="C443755EE13A900F2CAB469E2A9990A7" blockId="9.[151,1437,1159,1212]" pageId="9" pageNumber="228">
<emphasis id="F688A94CE13A900F2CAB469E2A1B9086" bold="true" box="[151,275,1159,1181]" pageId="9" pageNumber="228">FIGURE 5.</emphasis>
Dorsal (A) and ventral (B) views of the holotype of
<taxonomicName id="03FC0EDDE13A900F2F0E469E2F149086" authority="Yuan &amp; Zhang &amp; Che, 2016" authorityName="Yuan &amp; Zhang &amp; Che" authorityYear="2016" box="[818,1052,1159,1181]" class="Amphibia" family="Salamandridae" genus="Pachytriton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="228" phylum="Chordata" rank="species" species="wuguanfui" status="sp. nov.">
<emphasis id="F688A94CE13A900F2F0E469E2F149086" box="[818,1052,1159,1181]" italics="true" pageId="9" pageNumber="228">Pachytriton wuguanfui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EDBB1437E13A900F281F469E2F7D9086" box="[1059,1141,1159,1181]" pageId="9" pageNumber="228" rank="species">
<emphasis id="F688A94CE13A900F281F469E2F7D9086" bold="true" box="[1059,1141,1159,1181]" pageId="9" pageNumber="228">sp. nov.</emphasis>
</taxonomicNameLabel>
(KIZ08758) in preservation. The scale shows 20 mm.
</paragraph>
</caption>
<caption id="908325D6E13A900F2CAB45752A1393BA" httpUri="https://zenodo.org/record/1053438/files/figure.png" pageId="9" pageNumber="228" targetBox="[151,1436,1262,1879]" targetPageId="9">
<paragraph id="C443755EE13A900F2CAB45752A1393BA" blockId="9.[151,1436,1900,1953]" pageId="9" pageNumber="228">
<emphasis id="F688A94CE13A900F2CAB45752A1B9399" bold="true" box="[151,275,1900,1922]" pageId="9" pageNumber="228">FIGURE 6.</emphasis>
Ventral views of (A) forelimb and (B) hindlimb of the holotype of
<taxonomicName id="03FC0EDDE13A900F2FFC45752FA19399" authority="Yuan &amp; Zhang &amp; Che, 2016" authorityName="Yuan &amp; Zhang &amp; Che" authorityYear="2016" box="[960,1193,1900,1922]" class="Amphibia" family="Salamandridae" genus="Pachytriton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="228" phylum="Chordata" rank="species" species="wuguanfui" status="sp. nov.">
<emphasis id="F688A94CE13A900F2FFC45752FA19399" box="[960,1193,1900,1922]" italics="true" pageId="9" pageNumber="228">Pachytriton wuguanfui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EDBB1437E13A900F289345752E089399" box="[1199,1280,1900,1922]" pageId="9" pageNumber="228" rank="species">
<emphasis id="F688A94CE13A900F289345752E089399" bold="true" box="[1199,1280,1900,1922]" pageId="9" pageNumber="228">sp. nov.</emphasis>
</taxonomicNameLabel>
(KIZ08758) in preservation.
</paragraph>
</caption>
<subSubSection id="8CE626D5E139900C2CFB428E2E799747" pageId="10" pageNumber="229" type="description">
<paragraph id="C443755EE139900C2CFB428E2F399603" blockId="10.[151,1437,151,968]" pageId="10" pageNumber="229">
<emphasis id="F688A94CE139900C2CFB428E291F94AB" bold="true" box="[199,535,151,176]" pageId="10" pageNumber="229">
Description of the
<typeStatus id="1B47CBFCE139900C2D96428E291B94AB" box="[426,531,151,176]" pageId="10" pageNumber="229" type="holotype">holotype</typeStatus>
.
</emphasis>
This specimen is a moderately sized newt (TTL =
<quantity id="0304D8BBE139900C2854428E2FE994AB" box="[1128,1249,151,176]" metricMagnitude="-1" metricUnit="m" metricValue="1.557" pageId="10" pageNumber="229" unit="mm" value="155.7">155.7 mm</quantity>
). Head oval and flat, much longer than wide (HL/HW = 1.24). Snout truncate, projects slightly beyond lower jaw. Nostril small, nearly to the tip of snout. Eyes small and do not protuberate in dorsal view. Labial fold well developed on upper jaw. Gular folds weak. Parotoid gland prominent and protuberant. Tongue pad elliptical, adheres to mouth floor. Vomerine tooth series Ʌ-shaped. Dorsal vertebral groove conspicuous along the dorsal midline. Skin smooth. Costal grooves between axilla and groin lacking. Limbs very short (AL/SVL = 0.18, PL/SVL = 0.21). The forelimb is slightly shorter than the hindlimb (AL/PL = 0.85); tips of forelimbs and hindlimbs are well separated when adpressed to body flank. Fingers and toes moderately webbed on the sides (so digits appear flat) and without palmar tubercles. The relative finger lengths are 1 &lt;4 &lt;2 &lt;3 and relative toe lengths are 1 &lt;5 &lt;2 &lt;4 &lt;3, the fifth toes are very short. Dorsal caudal fin evident extends from tail-base to tail-tip. Ventral caudal fin prominent on posterior half of tail. Tail-tip rounded. Cloaca short slit, small and not protuberant.
</paragraph>
<paragraph id="C443755EE139900C2CFB403D2A7E969E" blockId="10.[151,1437,151,968]" pageId="10" pageNumber="229">
<emphasis id="F688A94CE139900C2CFB403D2A6F9626" bold="true" box="[199,359,548,573]" pageId="10" pageNumber="229">Color in life.</emphasis>
In life, dorsal color uniform dark brown; venter and chin brown and scattered with large, irregular, orange blotches; cloaca, underside of limbs and tail also orange. Tips of digits appear orange red when viewed from above.
</paragraph>
<paragraph id="C443755EE139900C2CFB40962A2D96D6" blockId="10.[151,1437,151,968]" pageId="10" pageNumber="229">
<emphasis id="F688A94CE139900C2CFB40962AC596B3" bold="true" box="[199,461,655,680]" pageId="10" pageNumber="229">Color in preservative.</emphasis>
In preservation, dark brown coloration turns to grey black; orange coloration fades to milky white.
</paragraph>
<paragraph id="C443755EE139900C2CFB40CE2E799747" blockId="10.[151,1437,151,968]" pageId="10" pageNumber="229">
<emphasis id="F688A94CE139900C2CFB40CE2A3496EB" bold="true" box="[199,316,727,752]" pageId="10" pageNumber="229">Variation.</emphasis>
The cloaca in males is larger and more swollen than that in females, with papillae on the cloacal wall. Sexual dimorphism in body size could not be determined because only two well-preserved adult males. Ventral coloration varies greatly from a large amount of orange, irregular blotches (KIZ0875608759, KIZ
<date id="B042539EE139900C290641062E869723" box="[1338,1422,799,824]" pageId="10" pageNumber="229" value="05-02-17">021705</date>
0 6, 0 21708, 0 21712, 0 21832, 0 21833,
<date id="B042539EE139900C2E75415D29979746" box="[585,671,836,861]" pageId="10" pageNumber="229" value="35-02-18">021835</date>
) to only a few small orange spots (KIZ08755, 0 21711,
<date id="B042539EE139900C2937415D2E6C9747" box="[1291,1380,836,860]" pageId="10" pageNumber="229" value="34-02-18">021834</date>
).
</paragraph>
</subSubSection>
<subSubSection id="8CE626D5E139900D2CFB417E2FC69053" lastPageId="11" lastPageNumber="230" pageId="10" pageNumber="229" type="etymology">
<paragraph id="C443755EE139900C2CFB417E2F1997D3" blockId="10.[151,1437,151,968]" pageId="10" pageNumber="229">
<emphasis id="F688A94CE139900C2CFB417E2A46979B" bold="true" box="[199,334,871,896]" pageId="10" pageNumber="229">Etymology.</emphasis>
The specific epithet
<taxonomicName id="03FC0EDDE139900C2E6E417129C0979B" authorityName="Yuan &amp; Zhang &amp; Che" authorityYear="2016" box="[594,712,872,896]" class="Amphibia" family="Salamandridae" genus="Pachytriton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="wuguanfui">
<emphasis id="F688A94CE139900C2E6E417129C0979B" box="[594,712,872,896]" italics="true" pageId="10" pageNumber="229">wuguanfui</emphasis>
</taxonomicName>
is a patronym honouring Guanfu Wu (Chengdu Institute of Biology, Chinese Academy of Sciences), a prominent Chinese herpetologist and educator, for his great contribution to the taxonomy, karyology, and osteology of amphibians and reptiles in
<collectingCountry id="BCEB35CEE139900C2FFA41B62F0597D3" box="[966,1037,943,968]" name="China" pageId="10" pageNumber="229">China</collectingCountry>
.
</paragraph>
<caption id="908325D6E139900C2CAB45772E0E939F" box="[151,1286,1902,1924]" httpUri="https://zenodo.org/record/1053442/files/figure.png" pageId="10" pageNumber="229" targetBox="[151,1436,1021,1881]" targetPageId="10">
<paragraph id="C443755EE139900C2CAB45772E0E939F" blockId="10.[151,1286,1902,1924]" box="[151,1286,1902,1924]" pageId="10" pageNumber="229">
<emphasis id="F688A94CE139900C2CAB45772A1A939F" bold="true" box="[151,274,1902,1924]" pageId="10" pageNumber="229">FIGURE 7.</emphasis>
Habitat at the type locality of
<taxonomicName id="03FC0EDDE139900C2E7B45772838939F" authority="Yuan &amp; Zhang &amp; Che, 2016" authorityName="Yuan &amp; Zhang &amp; Che" authorityYear="2016" box="[583,816,1902,1924]" class="Amphibia" family="Salamandridae" genus="Pachytriton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="wuguanfui" status="sp. nov.">
<emphasis id="F688A94CE139900C2E7B45772838939F" box="[583,816,1902,1924]" italics="true" pageId="10" pageNumber="229">Pachytriton wuguanfui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EDBB1437E139900C2F0A4577288F939F" box="[822,903,1902,1924]" pageId="10" pageNumber="229" rank="species">
<emphasis id="F688A94CE139900C2F0A4577288F939F" bold="true" box="[822,903,1902,1924]" pageId="10" pageNumber="229">sp. nov.</emphasis>
</taxonomicNameLabel>
, Mt. Gupo, Hezhou, Guangxi, China.
</paragraph>
</caption>
<caption id="908325D6E138900D2CAB40E52884972A" httpUri="https://zenodo.org/record/1053444/files/figure.png" pageId="11" pageNumber="230" targetBox="[151,1436,193,743]" targetPageId="11">
<paragraph id="C443755EE138900D2CAB40E52884972A" blockId="11.[151,1436,764,817]" pageId="11" pageNumber="230">
<emphasis id="F688A94CE138900D2CAB40E52A1F9709" bold="true" box="[151,279,764,786]" pageId="11" pageNumber="230">FIGURE 8.</emphasis>
Vomerine tooth morphology in adult (A)
<taxonomicName id="03FC0EDDE138900D2EE240E528C49709" authority="Yuan &amp; Zhang &amp; Che, 2016" authorityName="Yuan &amp; Zhang &amp; Che" authorityYear="2016" box="[734,972,764,786]" class="Amphibia" family="Salamandridae" genus="Pachytriton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="11" pageNumber="230" phylum="Chordata" rank="species" species="wuguanfui" status="sp. nov.">
<emphasis id="F688A94CE138900D2EE240E528C49709" box="[734,972,764,786]" italics="true" pageId="11" pageNumber="230">Pachytriton wuguanfui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EDBB1437E138900D2FEA40E52F239709" box="[982,1067,764,786]" pageId="11" pageNumber="230" rank="species">
<emphasis id="F688A94CE138900D2FEA40E52F239709" bold="true" box="[982,1067,764,786]" pageId="11" pageNumber="230">sp. nov.</emphasis>
</taxonomicNameLabel>
(KIZ021834) and (B)
<taxonomicName id="03FC0EDDE138900D291840E52A1E972A" authorityName="Nishikawa, Jiang, Matsui &amp; Mo" authorityYear="2011" class="Amphibia" family="Salamandridae" genus="Pachytriton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="11" pageNumber="230" phylum="Chordata" rank="species" species="inexpectatus">
<emphasis id="F688A94CE138900D291840E52A1E972A" italics="true" pageId="11" pageNumber="230">Pachytriton inexpectatus</emphasis>
</taxonomicName>
(KIZ014508, Mt. Maoer, Guangxi). The scale shows 1.7 mm.
</paragraph>
</caption>
<paragraph id="C443755EE138900D2CFB41412FC69053" blockId="11.[151,1437,856,1421]" pageId="11" pageNumber="230">
<emphasis id="F688A94CE138900D2CFB41412AFA976A" bold="true" box="[199,498,856,881]" pageId="11" pageNumber="230">Habitat and distribution.</emphasis>
This species inhabits small montane streams in broadleaf forests. Streams inhabited by these newts are ca. 2 meters wide and 0.51 meters in depth; slope is steep, with fast flowing clean and cold water. Many large boulders are scattered in or around the streams. Stream substrates include gravels, scattered small rocks, leaves and sands. Frogs in the genera
<taxonomicName id="03FC0EDDE138900D2EDD41DD285797C6" authorityName="Kuhl &amp; van Hasselt" authorityYear="1822" box="[737,863,964,989]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="230" phylum="Chordata" rank="genus">
<emphasis id="F688A94CE138900D2EDD41DD285797C6" box="[737,863,964,989]" italics="true" pageId="11" pageNumber="230">Megophrys</emphasis>
</taxonomicName>
,
<taxonomicName id="03FC0EDDE138900D2F5341DD28D997C6" box="[879,977,964,989]" class="Amphibia" family="Ranidae" genus="Amolops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="230" phylum="Chordata" rank="genus">
<emphasis id="F688A94CE138900D2F5341DD28D997C6" box="[879,977,964,989]" italics="true" pageId="11" pageNumber="230">Amolops</emphasis>
</taxonomicName>
and
<taxonomicName id="03FC0EDDE138900D283041DC2F7197C7" authorityName="Dubois" authorityYear="1992" box="[1036,1145,965,988]" class="Amphibia" family="Dicroglossidae" genus="Quasipaa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="230" phylum="Chordata" rank="genus">
<emphasis id="F688A94CE138900D283041DC2F7197C7" box="[1036,1145,965,988]" italics="true" pageId="11" pageNumber="230">Quasipaa</emphasis>
</taxonomicName>
were also found in these habitats. Newts were found along the stream at night. We did not encounter the species during the day.
<taxonomicName id="03FC0EDDE138900D292441F12A05903E" authority="Yuan &amp; Zhang &amp; Che, 2016" authorityName="Yuan &amp; Zhang &amp; Che" authorityYear="2016" class="Amphibia" family="Salamandridae" genus="Pachytriton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="11" pageNumber="230" phylum="Chordata" rank="species" species="wuguanfui" status="sp. nov.">
<emphasis id="F688A94CE138900D292441F12A05903E" italics="true" pageId="11" pageNumber="230">Pachytriton wuguanfui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EDBB1437E138900D2D2A46152A7A903E" box="[278,370,1036,1061]" pageId="11" pageNumber="230" rank="species">
<emphasis id="F688A94CE138900D2D2A46152A7A903E" bold="true" box="[278,370,1036,1061]" pageId="11" pageNumber="230">sp. nov.</emphasis>
</taxonomicNameLabel>
were found at Mt. Jiuwei in
<collectingRegion id="0638BBBCE138900D2EF946142818903F" box="[709,784,1037,1060]" country="China" name="Hunan" pageId="11" pageNumber="230">Hunan</collectingRegion>
and Mt. Gupo in
<collectingRegion id="0638BBBCE138900D2FDE46152F4A903E" box="[994,1090,1036,1061]" country="China" name="Guangxi" pageId="11" pageNumber="230">Guangxi</collectingRegion>
and it likely occurs in nearby mountains between the distribution of
<taxonomicName id="03FC0EDDE138900D2E7A462829E69053" authorityName="Nishikawa, Jiang, Matsui &amp; Mo" authorityYear="2011" box="[582,750,1073,1096]" class="Amphibia" family="Salamandridae" genus="Pachytriton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="11" pageNumber="230" phylum="Chordata" rank="species" species="inexpectatus">
<emphasis id="F688A94CE138900D2E7A462829E69053" box="[582,750,1073,1096]" italics="true" pageId="11" pageNumber="230">P. inexpectatus</emphasis>
</taxonomicName>
in the west and
<taxonomicName id="03FC0EDDE138900D2F9846282F459052" box="[932,1101,1072,1097]" class="Amphibia" family="Salamandridae" genus="Pachytriton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="11" pageNumber="230" phylum="Chordata" rank="species" species="xanthospilos">
<emphasis id="F688A94CE138900D2F9846282F459052" box="[932,1101,1072,1097]" italics="true" pageId="11" pageNumber="230">P. xanthospilos</emphasis>
</taxonomicName>
in the east.
</paragraph>
</subSubSection>
<subSubSection id="8CE626D5E138900D2CFB464D2E6A9196" pageId="11" pageNumber="230" type="description">
<paragraph id="C443755EE138900D2CFB464D2E6A9196" blockId="11.[151,1437,856,1421]" pageId="11" pageNumber="230">
<emphasis id="F688A94CE138900D2CFB464D2A649076" bold="true" box="[199,364,1108,1133]" pageId="11" pageNumber="230">Comparisons.</emphasis>
<taxonomicName id="03FC0EDDE138900D2D47464D298B9076" authority="Yuan &amp; Zhang &amp; Che, 2016" authorityName="Yuan &amp; Zhang &amp; Che" authorityYear="2016" box="[379,643,1108,1133]" class="Amphibia" family="Salamandridae" genus="Pachytriton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="11" pageNumber="230" phylum="Chordata" rank="species" species="wuguanfui" status="sp. nov.">
<emphasis id="F688A94CE138900D2D47464D298B9076" box="[379,643,1108,1133]" italics="true" pageId="11" pageNumber="230">Pachytriton wuguanfui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EDBB1437E138900D2EAD464D29F99076" box="[657,753,1108,1133]" pageId="11" pageNumber="230" rank="species">
<emphasis id="F688A94CE138900D2EAD464D29F99076" bold="true" box="[657,753,1108,1133]" pageId="11" pageNumber="230">sp. nov.</emphasis>
</taxonomicNameLabel>
described here distinctly differs from
<taxonomicName id="03FC0EDDE138900D28F4464C2E7B9076" authorityName="Shen, Shen &amp; Mo" authorityYear="2008" box="[1224,1395,1108,1133]" class="Amphibia" family="Salamandridae" genus="Pachytriton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="11" pageNumber="230" phylum="Chordata" rank="species" species="archospotus">
<emphasis id="F688A94CE138900D28F4464C2E7B9076" box="[1224,1395,1108,1133]" italics="true" pageId="11" pageNumber="230">P. archospotus</emphasis>
</taxonomicName>
,
<taxonomicName id="03FC0EDDE138900D29BB464C2BFD908A" class="Amphibia" family="Salamandridae" genus="Pachytriton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="11" pageNumber="230" phylum="Chordata" rank="species" species="brevipes">
<emphasis id="F688A94CE138900D29BB464C2BFD908A" italics="true" pageId="11" pageNumber="230">P. brevipes</emphasis>
</taxonomicName>
, and a few populations of
<taxonomicName id="03FC0EDDE138900D2E12466029C1908A" box="[558,713,1144,1169]" class="Amphibia" family="Salamandridae" genus="Pachytriton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="11" pageNumber="230" phylum="Chordata" rank="species" species="granulosus">
<emphasis id="F688A94CE138900D2E12466029C1908A" box="[558,713,1144,1169]" italics="true" pageId="11" pageNumber="230">P. granulosus</emphasis>
</taxonomicName>
by the lack of black spots on the whole body (vs. presence of black spots in these species). It differs from
<taxonomicName id="03FC0EDDE138900D2E814684282490AE" box="[701,812,1180,1205]" class="Amphibia" family="Salamandridae" genus="Pachytriton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="11" pageNumber="230" phylum="Chordata" rank="species" species="changi">
<emphasis id="F688A94CE138900D2E814684282490AE" box="[701,812,1180,1205]" italics="true" pageId="11" pageNumber="230">P. changi</emphasis>
</taxonomicName>
and
<taxonomicName id="03FC0EDDE138900D2F4C46842F2990AE" box="[880,1057,1180,1205]" class="Amphibia" family="Salamandridae" genus="Pachytriton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="11" pageNumber="230" phylum="Chordata" rank="species" species="xanthospilos">
<emphasis id="F688A94CE138900D2F4C46842F2990AE" box="[880,1057,1180,1205]" italics="true" pageId="11" pageNumber="230">P. xanthospilos</emphasis>
</taxonomicName>
by lacking orange dots on the dorsolateral side (vs. presence of orange spots or blotches that extend ribbon-like along the dorsolateral sides of the body in both species) and from
<emphasis id="F688A94CE138900D2DCB46FC293490E7" box="[503,572,1253,1276]" italics="true" pageId="11" pageNumber="230">P. moi</emphasis>
by having orange blotches on venter (vs. lack of orange blotches on venter in
<emphasis id="F688A94CE138900D2CAB47102BD4913B" box="[151,220,1289,1312]" italics="true" pageId="11" pageNumber="230">P. moi</emphasis>
). It differs from
<taxonomicName id="03FC0EDDE138900D2DA547102948913B" authorityName="Nishikawa, Jiang, Matsui &amp; Mo" authorityYear="2011" box="[409,576,1289,1312]" class="Amphibia" family="Salamandridae" genus="Pachytriton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="11" pageNumber="230" phylum="Chordata" rank="species" species="inexpectatus">
<emphasis id="F688A94CE138900D2DA547102948913B" box="[409,576,1289,1312]" italics="true" pageId="11" pageNumber="230">P. inexpectatus</emphasis>
</taxonomicName>
,
<taxonomicName id="03FC0EDDE138900D2E72471029EF913A" box="[590,743,1288,1313]" class="Amphibia" family="Salamandridae" genus="Pachytriton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="11" pageNumber="230" phylum="Chordata" rank="species" species="granulosus">
<emphasis id="F688A94CE138900D2E72471029EF913A" box="[590,743,1288,1313]" italics="true" pageId="11" pageNumber="230">P. granulosus</emphasis>
</taxonomicName>
and
<taxonomicName id="03FC0EDDE138900D2F2147102854913B" authorityName="Nishikawa, Jiang &amp; Matsui" authorityYear="2011" box="[797,860,1288,1312]" class="Amphibia" family="Salamandridae" genus="Pachytriton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="11" pageNumber="230" phylum="Chordata" rank="species" species="feii">
<emphasis id="F688A94CE138900D2F2147102854913B" box="[797,860,1288,1312]" italics="true" pageId="11" pageNumber="230">P. feii</emphasis>
</taxonomicName>
by lacking costal grooves between axilla and groin (vs. all these three species present costal grooves). The anterior end of the vomerine tooth series is also more widely open in the new species than
<taxonomicName id="03FC0EDDE138900D2E0F474829D39173" authorityName="Nishikawa, Jiang, Matsui &amp; Mo" authorityYear="2011" box="[563,731,1361,1384]" class="Amphibia" family="Salamandridae" genus="Pachytriton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="11" pageNumber="230" phylum="Chordata" rank="species" species="inexpectatus">
<emphasis id="F688A94CE138900D2E0F474829D39173" box="[563,731,1361,1384]" italics="true" pageId="11" pageNumber="230">P. inexpectatus</emphasis>
</taxonomicName>
(
<figureCitation id="5CC769DBE138900D2ED6474928269173" box="[746,814,1360,1385]" captionStart="FIGURE 8" captionStartId="11.[151,250,764,786]" captionTargetBox="[151,1436,193,743]" captionTargetId="figure@11.[151,1436,193,743]" captionTargetPageId="11" captionText="FIGURE 8. Vomerine tooth morphology in adult (A) Pachytriton wuguanfui sp. nov. (KIZ 021834) and (B) Pachytriton inexpectatus (KIZ 014508, Mt. Maoer, Guangxi). The scale shows 1.7 mm." httpUri="https://zenodo.org/record/1053444/files/figure.png" pageId="11" pageNumber="230">Fig. 8</figureCitation>
). Finger and toe tips in life appear orange on the dorsal side in
<taxonomicName id="03FC0EDDE138900D2CD4476C2A719196" authority="Yuan &amp; Zhang &amp; Che, 2016" authorityName="Yuan &amp; Zhang &amp; Che" authorityYear="2016" box="[232,377,1397,1421]" class="Amphibia" family="Salamandridae" genus="Pachytriton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="11" pageNumber="230" phylum="Chordata" rank="species" species="wuguanfui" status="sp. nov.">
<emphasis id="F688A94CE138900D2CD4476C2A719196" box="[232,377,1397,1421]" italics="true" pageId="11" pageNumber="230">P. wuguanfui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EDBB1437E138900D2DBC476D2AD29196" box="[384,474,1396,1421]" pageId="11" pageNumber="230" rank="species">
<emphasis id="F688A94CE138900D2DBC476D2AD29196" bold="true" box="[384,474,1396,1421]" pageId="11" pageNumber="230">sp. nov.</emphasis>
</taxonomicNameLabel>
, which distinguishes the new species from
<taxonomicName id="03FC0EDDE138900D2FFD476C2F089196" authorityName="Nishikawa, Jiang &amp; Matsui" authorityYear="2011" box="[961,1024,1397,1421]" class="Amphibia" family="Salamandridae" genus="Pachytriton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="11" pageNumber="230" phylum="Chordata" rank="species" species="feii">
<emphasis id="F688A94CE138900D2FFD476C2F089196" box="[961,1024,1397,1421]" italics="true" pageId="11" pageNumber="230">P. feii</emphasis>
</taxonomicName>
(digit tips appear black in life).
</paragraph>
</subSubSection>
</treatment>
</document>