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<document id="A993E7F23F7D9497971B56E4730525F4" ID-DOI="10.1206/0003-0082(2003)390&lt;0001:TFCRIA&gt;2.0.CO;2" ID-ISSN="0003-0082" ID-Zenodo-Dep="5372693" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630272521932" checkinUser="carolina" docAuthor="KRISHNA, KUMAR &amp; GRIMALDI, DAVID A." docDate="2003" docId="4C52B8352B144259FF22C20710AFD6B1" docLanguage="en" docName="N3390.pdf" docOrigin="American Museum Novitates 3390" docSource="http://www.bioone.org/doi/abs/10.1206/0003-0082%282003%29390%3C0001%3ATFCRIA%3E2.0.CO%3B2" docStyle="DocumentStyle:70DDF1BDF6846024993F05262BAE3001.3:AmMusNovit.2000-2010.journal_article.type1" docStyleId="70DDF1BDF6846024993F05262BAE3001" docStyleName="AmMusNovit.2000-2010.journal_article.type1" docStyleVersion="3" docTitle="Archeorhinotermes rossi KRISHNA &amp; GRIMALDI 2003, new species" docType="treatment" docVersion="3" lastPageNumber="5" masterDocId="B06BC04D2B17425DFFBCC70A1115D562" masterDocTitle="The First Cretaceous Rhinotermitidae (Isoptera): A New Species, Genus, and Subfamily in Burmese Amber" masterLastPageNumber="11" masterPageNumber="1" pageNumber="4" updateTime="1699207917690" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="1A42737A52CF861F2FA40ACFDC458507">The First Cretaceous Rhinotermitidae (Isoptera): A New Species, Genus, and Subfamily in Burmese Amber</mods:title>
</mods:titleInfo>
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<mods:namePart id="05D6EA026FE8BF2AC9402594BCE5A05B">KRISHNA, KUMAR</mods:namePart>
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<mods:namePart id="57E4264C83618639C13C9902A981417F">GRIMALDI, DAVID A.</mods:namePart>
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<mods:title id="7C7875AE98492FF555D8D74D7CF4426B">American Museum Novitates</mods:title>
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<mods:date id="93511464D1745CFD06C524DCC0CF8AF9">2003</mods:date>
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<mods:number id="2453D9BE72EEFCD2B07FA4A57C4C4BFC">2003-02-19</mods:number>
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<mods:classification id="39BC47AF6F98A2E7506389F09C5401AD">journal article</mods:classification>
<mods:identifier id="529698ACF1E5E3CC6C4E12604A0CCA24" type="DOI">10.1206/0003-0082(2003)390&lt;0001:TFCRIA&gt;2.0.CO;2</mods:identifier>
<mods:identifier id="87CB41E40EB12933B49CE55D644BF039" type="ISSN">0003-0082</mods:identifier>
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<treatment id="4C52B8352B144259FF22C20710AFD6B1" LSID="urn:lsid:plazi:treatment:4C52B8352B144259FF22C20710AFD6B1" httpUri="http://treatment.plazi.org/id/4C52B8352B144259FF22C20710AFD6B1" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
<subSubSection id="8CE15AA82B14425EFF22C207139ED3AF" pageId="3" pageNumber="4" type="nomenclature">
<paragraph id="C44409232B14425EFF22C207137DD046" blockId="3.[158,616,1293,1317]" box="[158,616,1293,1317]" pageId="3" pageNumber="4">
<heading id="9F0CBE4F2B14425EFF22C207137DD046" box="[158,616,1293,1317]" centered="true" fontSize="36" level="2" pageId="3" pageNumber="4" reason="8">
<taxonomicName id="03FB72A02B14425EFF22C20710DCD047" ID-CoL="G8BM" authority="KRISHNA &amp; GRIMALDI, 2003" authorityName="KRISHNA &amp; GRIMALDI" authorityYear="2003" box="[158,457,1293,1317]" class="Insecta" family="Archeorhinotermitidae" genus="Archeorhinotermes" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="rossi" status="sp. nov.">
<emphasis id="F68FD5312B14425EFF22C20710DCD047" bold="true" box="[158,457,1293,1317]" italics="true" pageId="3" pageNumber="4">Archeorhinotermes rossi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="EDBC684A2B14425EFE65C207137DD046" box="[473,616,1293,1316]" pageId="3" pageNumber="4" rank="species">new species</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="C44409232B14425EFF24C233139ED3AF" blockId="3.[123,652,1337,1741]" pageId="3" pageNumber="4">
IMAGO (
<figureCitation id="5CC015A62B14425EFF4FC2331029D032" box="[243,316,1337,1360]" captionStart="Fig" captionStartId="5.[149,187,1560,1581]" captionTargetBox="[163,1187,190,1539]" captionTargetId="figure-10@5.[163,1187,187,1539]" captionTargetPageId="5" captionText="Fig. 1. Holotype imago of Archeorhinotermes rossi, n. sp. a, dorsal view of head; b, dorsal view of mandibles; c, ventral view of middle tibia and tarsus; d, dorsal view of forewing." figureDoi="http://doi.org/10.5281/zenodo.5372695" httpUri="https://zenodo.org/record/5372695/files/figure.png" pageId="3" pageNumber="4">figs. 1</figureCitation>
,
<figureCitation id="5CC015A62B14425EFEF7C2331049D032" box="[331,348,1337,1360]" captionStart="Fig" captionStartId="6.[150,188,1629,1650]" captionTargetBox="[210,1145,187,1608]" captionTargetId="figure-11@6.[210,1145,187,1608]" captionTargetPageId="6" captionText="Fig. 2. Photomicrograph of the holotype imago of Archeorhinotermes rossi, n. sp." figureDoi="http://doi.org/10.5281/zenodo.5372697" httpUri="https://zenodo.org/record/5372697/files/figure.png" pageId="3" pageNumber="4">2</figureCitation>
; table 1): Head, antennae, and pronotum light brown, wing scale brownish, lighter than head, legs light yellow. Head moderately covered with long bristles, longest about
<quantity id="0303A4C62B14425EFE37C2A410EFD0A7" box="[395,506,1454,1477]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="3" pageNumber="4" unit="mm" value="0.18">0.18 mm</quantity>
; upper margin of forewing scale with several short bristles; costal margin and radial sector with a few barely visible short bristles; tergites and sternites densely covered with long bristles; fore, middle, and hind tibia with several long, thin bristles; inner margin of middle and hind tibia with a row of additional short, thicker bristles; tarsal segments with several long bristles. Head small, roundish, slightly
</paragraph>
</subSubSection>
<subSubSection id="8CE15AA82B144259FC2CC7C810C6D622" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="description">
<paragraph id="C44409232B14425EFC2CC7C81587D46F" blockId="3.[720,1206,194,269]" pageId="3" pageNumber="4">
<materialsCitation id="7493037E2B14425EFC2CC7C81587D46F" collectionCode="TABLE" pageId="3" pageNumber="4" specimenCode="TABLE 1" specimenCount="1" typeStatus="holotype">
<specimenCode id="945DA1582B14425EFC2CC7C812FBD5B7" box="[912,1006,194,213]" collectionCode="TABLE" pageId="3" pageNumber="4">TABLE 1</specimenCode>
<emphasis id="F68FD5312B14425EFD6CC7D71587D46F" bold="true" pageId="3" pageNumber="4">
Measurements (mm) of
<typeStatus id="1B40B7812B14425EFC5DC7D71552D590" box="[993,1095,221,242]" pageId="3" pageNumber="4" type="holotype">Holotype</typeStatus>
Imago of
<taxonomicName id="03FB72A02B14425EFD48C7F21515D46F" authority="KRISHNA &amp; GRIMALDI, 2003" authorityName="KRISHNA &amp; GRIMALDI" authorityYear="2003" box="[756,1024,248,269]" class="Insecta" family="Archeorhinotermitidae" genus="Archeorhinotermes" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="rossi" status="sp. nov.">
<emphasis id="F68FD5312B14425EFD48C7F21515D46F" bold="true" box="[756,1024,248,269]" italics="true" pageId="3" pageNumber="4">Archeorhinotermes rossi</emphasis>
</taxonomicName>
,
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</emphasis>
</materialsCitation>
</paragraph>
<paragraph id="C44409232B14425EFD07C6201584D717" blockId="3.[699,1169,298,629]" pageId="3" pageNumber="4">Length of head to tip of labrum 0.77 Width of head 0.69 Maximum diameter of eye 0.24 Minimum diameter of eye 0.13 Maximum diameter of ocellus 0.09 Ocellus from eye 0.04 Length of labrum 0.23 Width of labrum 0.23 Length of postclypeus 0.10 Length of forewing scale 0.64 Length of forewing with scale 4.39 Width of forewing 0.97 Length of middle tibia 0.39 Length of hind tibia 0.56</paragraph>
<paragraph id="C44409232B144259FD07C5BE1336D471" blockId="3.[699,1228,692,1741]" lastBlockId="4.[124,653,193,979]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
longer than wide. Eyes medium­sized, oval, not bulging much beyond head. Ocelli large, separated from eyes. Fontanelle present, appearing closer to hind margin than to postclypeus. Labrum oblong, with lateral margins faintly rounded. Postclypeus narrow, not prominently arched. Pronotum tucked under head, not clearly visible, but appearing narrower than head. Left mandible with a prominent rounded projection near the apical tooth; apical tooth long and prominent, longer than first, second, or third marginal tooth, with outer margin sinuate; first, second, and third marginal teeth almost equal in length; molar prominence pointed, appearing as a fourth marginal tooth. Right mandible with a subsidiary tooth at base of upper margin of first marginal tooth (characteristic of
<taxonomicName id="03FB72A02B14425EFB3DC3AC1221D1B9" authorityName="Desneux" authorityYear="1904" class="Insecta" family="Hodotermitidae" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="4" phylum="Arthropoda" rank="family">Hodotermitidae</taxonomicName>
,
<taxonomicName id="03FB72A02B14425EFCFBC3CE12CAD1B9" box="[839,991,1220,1243]" class="Insecta" family="Termopsidae" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="4" phylum="Arthropoda" rank="family">Termopsidae</taxonomicName>
, and
<taxonomicName id="03FB72A02B14425EFB91C3CE13FDD19A" class="Insecta" family="Rhinotermitidae" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="4" phylum="Arthropoda" rank="family">Rhinotermitidae</taxonomicName>
); second marginal tooth prominent, equal in length to first marginal tooth, posterior margin with a medium­sized cutting edge (longer in all other rhinotermitids). Antennae broken, with 13+ articles; second subequal to third; third and fourth subequal. Tibial spurs very hard to distinguish from thick bristles at end of tibia, appear to be 3:2:2. Tarsus with four articles. Arolium absent. Cerci with two articles. Forewing membrane reticulate, particularly between branches of cubitus; R
<subScript id="587F0B662B14425EFC85C13A1257D35C" attach="left" box="[825,834,1584,1598]" fontSize="6" pageId="3" pageNumber="4">1</subScript>
and R
<subScript id="587F0B662B14425EFC20C13A12B0D35C" attach="left" box="[924,933,1584,1598]" fontSize="6" pageId="3" pageNumber="4">2</subScript>
absent; Rs simple, unbranched, running close to the fused subcosta and costal margin; median arising from a common stem with Rs, inside wing scale, unbranched, running almost midway between Rs and cubitus, appearing joined to cubitus, close to apical margin; cubitus arising independently from inside wing scale, with 11 branches, fourth with a sub­branch.
</paragraph>
<paragraph id="C44409232B134259FF25C61310C6D622" blockId="4.[124,653,193,979]" pageId="4" pageNumber="5">
SPECIMEN AND TYPE LOCALITY: Imago (
<typeStatus id="1B40B7812B134259FDD8C61311D3D42F" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
) in amber, NHM, Department of Palaeontology number In. 20160, received from R.C.J. Swinhoe,
<date id="B0452FE32B134259FEF5C67B10D6D4EA" box="[329,451,369,392]" pageId="4" pageNumber="5" value="1920-07">July 1920</date>
.
<collectingCountry id="BCEC49B32B134259FE6EC67B1356D4EA" box="[466,579,369,392]" name="Myanmar" pageId="4" pageNumber="5">Myanmar</collectingCountry>
(
<collectingCountry id="BCEC49B32B134259FDEBC67B11B0D4C7" name="Myanmar" pageId="4" pageNumber="5">Burma</collectingCountry>
):?Hukawng Valley. Spherical piece of amber,
<quantity id="0303A4C62B134259FF6DC6A61037D4A1" box="[209,290,428,451]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="5" unit="mm" value="10.0">10 mm</quantity>
in diameter, with a 2­mm wide hole bored through center (originally a bead), containing one termite (as above), one
<taxonomicName id="03FB72A02B134259FDDEC6EC1012D779" authorityName="Newman" authorityYear="1835" class="Insecta" family="Cecidomyidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Cecidomyidae</taxonomicName>
, one Acarina, one Auchenorrhycha nymph, four
<taxonomicName id="03FB72A02B134259FEEAC52B10C3D75A" authorityName="Linnaeus" authorityYear="1758" box="[342,470,545,568]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
(?family), one
<taxonomicName id="03FB72A02B134259FFC0C53410A7D737" authorityName="Linnaeus" authorityYear="1758" box="[124,434,574,597]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="order">Coleoptera (Mordellidae)</taxonomicName>
, one Heteroptera (
<taxonomicName id="03FB72A02B134259FF38C55611FBD711" authorityName="Amyot &amp; Serville" authorityYear="1843" box="[132,238,604,627]" class="Insecta" family="Saldidae" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Saldidae</taxonomicName>
), two
<taxonomicName id="03FB72A02B134259FEFAC556139CD711" box="[326,649,604,627]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="order">Diptera (Ceratopogonidae)</taxonomicName>
, and
<specimenCount id="D2FDC2AA2B134259FF12C573100CD7F2" box="[174,281,633,656]" pageId="4" pageNumber="5" type="male">one male</specimenCount>
Coccoidea. The
<typeStatus id="1B40B7812B134259FE63C5731350D7F2" box="[479,581,633,656]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
is deposited in the Department of Palaeontology (Invertebrates), Natural History Museum, London. Burmese amber is dated as Upper to Middle Cretaceous, probably Turonian– Cenomanian (90–100 mya) by
<bibRefCitation id="A06A74D22B134259FE57C40610DAD622" author="Grimaldi, D. A. &amp; M. S. Engel &amp; P. C. Nascimbene" pageId="4" pageNumber="5" pagination="1 - 72" refId="ref3273" refString="Grimaldi, D. A., M. S. Engel, and P. C. Nascimbene. 2002. Fossiliferous Cretaceous amber from Myanmar (Burma): its rediscovery, biotic diversity, and paleontological significance. American Museum Novitates 3361: 1 - 72." type="journal article" year="2002">Grimaldi, Engel, and Nascimbene (2002)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8CE15AA82B134259FF25C44C10AFD6B1" pageId="4" pageNumber="5" type="etymology">
<paragraph id="C44409232B134259FF25C44C10AFD6B1" blockId="4.[124,653,193,979]" pageId="4" pageNumber="5">ETYMOLOGY: This species is named in hon­ or of Andrew Ross, Curator of Fossil Arthropods, Department of Palaeontology, Natural History Museum, London, and an authority on Burmese amber.</paragraph>
</subSubSection>
</treatment>
</document>
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