163 lines
17 KiB
XML
163 lines
17 KiB
XML
<document id="8A754AFDFD6C98F0894935900544097A" ID-DOI="10.1649/0010-065X(2001)055[0433:NSOHKC]2.0.CO;2" ID-ISSN="1938-4394" ID-Zenodo-Dep="4900383" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1622828115451" checkinUser="carolina" docAuthor="Ratcliffe, Brett C." docDate="2001" docId="4C4A87F74456E03CFE1592E3FE8DFE9E" docLanguage="en" docName="TheColeopBull.55.4.433-443.pdf.imf" docOrigin="The Coleopterists Bulletin 55 (4)" docStyle="DocumentStyle:37D8AFFB4AA6BE9981973FD5310B493E.6:TheColeopBull.2000-2002.journal_article.0cover" docStyleId="37D8AFFB4AA6BE9981973FD5310B493E" docStyleName="TheColeopBull.2000-2002.journal_article.0cover" docStyleVersion="6" docTitle="Hemiphileurus bispinosus Ratcliffe 2001, new species" docType="treatment" docVersion="4" lastPageNumber="437" masterDocId="B073FF8F4455E038FF98931DFFFEFFA1" masterDocTitle="New Species Of Hemiphileurus Kolbe (Coleoptera: Scarabaeidae: Dynastinae) From Mexico, Guatemala, Colombia, And Brazil" masterLastPageNumber="443" masterPageNumber="433" pageNumber="436" updateTime="1698955217201" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:mods id="52B27F0EF405B0195BCD01241C94B0EA" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="47149E19A82CCCAC69F7E159E424261C">
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<mods:title id="4D870BFB9CEA56AE3A023B3746065997">New Species Of Hemiphileurus Kolbe (Coleoptera: Scarabaeidae: Dynastinae) From Mexico, Guatemala, Colombia, And Brazil</mods:title>
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</mods:titleInfo>
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<mods:name id="CEF34C1AA21AD30CA8846C3E03705E1E" type="personal">
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<mods:role id="3E10B92E66D4EE7A5135599DBC495003">
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<mods:roleTerm id="36A67306E56145DFF655BDC5D430609D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="26541FAFE9E300749E3223D32DB808FE">Ratcliffe, Brett C.</mods:namePart>
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<mods:affiliation id="C704C35E3DFDFFDAA9E24A5E51CDDB71">Systematics Research Collections W 436 Nebraska Hall University of Nebraska Lincoln, NE 68588 0514, U. S. A.</mods:affiliation>
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<mods:nameIdentifier id="B2C020FE3528E2C4DB3679939E7E2762" type="email">bratcliffe1@unl.edu</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource id="FF91CB446E7556FE1B7032C385AEA7AB">text</mods:typeOfResource>
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<mods:titleInfo id="482292C038E806C6F04822A760635E23">
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<mods:title id="0BBB179041D102389689A11E14607EE1">The Coleopterists Bulletin</mods:title>
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</mods:titleInfo>
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<mods:part id="86A61A300973431CD95C8FDA75645CFF">
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<mods:date id="E51B95AC44A63C79606FFA22970AE209">2001</mods:date>
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<mods:detail id="0E368BD33EE4AD0F8A633AC666628743" type="pubDate">
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<mods:number id="4792A7A9F345E8F9210176696E14E011">2001-12-01</mods:number>
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</mods:detail>
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<mods:detail id="9C4831834ECB224DEE6A11570B374766" type="volume">
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<mods:number id="5F69CD021C1597B5C1FD9B383A98B646">55</mods:number>
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</mods:detail>
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<mods:detail id="59227F8741CFA062DED3D542A3599DEA" type="issue">
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<mods:number id="0D340BC80E7B64E460F7A9DA6938BC21">4</mods:number>
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</mods:detail>
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<mods:extent id="1077C595456CB4BBE1D226B29DED44F6" unit="page">
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<mods:start id="926D9F168213D0B68767CCDBDE5468BC">433</mods:start>
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<mods:end id="174456ED4FF1326AB553D45BC56D31F6">443</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="DDC05D1F06C4515EC9B253306459F2D8">journal article</mods:classification>
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<mods:identifier id="93D73AA113DF26B3AB5D91ECDBDEFE3F" type="DOI">10.1649/0010-065X(2001)055[0433:NSOHKC]2.0.CO;2</mods:identifier>
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<mods:identifier id="CC1605E9DAA2B86272223A3789223465" type="ISSN">1938-4394</mods:identifier>
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<mods:identifier id="1D0BB4F9BF97455C1CE475ECF49BFFB7" type="Zenodo-Dep">4900383</mods:identifier>
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</mods:mods>
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<treatment id="4C4A87F74456E03CFE1592E3FE8DFE9E" ID-DOI="http://doi.org/10.5281/zenodo.4900463" ID-Zenodo-Dep="4900463" LSID="urn:lsid:plazi:treatment:4C4A87F74456E03CFE1592E3FE8DFE9E" httpUri="http://treatment.plazi.org/id/4C4A87F74456E03CFE1592E3FE8DFE9E" lastPageId="4" lastPageNumber="437" pageId="3" pageNumber="436">
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<subSubSection id="8CF9656A4456E03BFE1592E3FD2FFD8F" pageId="3" pageNumber="436" type="nomenclature">
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<paragraph id="C45C36E14456E03BFE1592E3FC68FDB2" blockId="3.[397,918,510,558]" box="[397,918,510,531]" pageId="3" pageNumber="436">
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<heading id="9F14818D4456E03BFE1592E3FC68FDB2" box="[397,918,510,531]" centered="true" fontSize="9" level="2" pageId="3" pageNumber="436" reason="2">
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<taxonomicName id="03E34D624456E03BFE1592E3FCF8FDB2" authority="Ratcliffe" authorityName="Ratcliffe" authorityYear="2001" box="[397,774,510,531]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="436" phylum="Arthropoda" rank="species" species="bispinosus" status="sp. nov.">
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<emphasis id="F697EAF34456E03BFE1592E3FD61FDB2" box="[397,671,510,531]" italics="true" pageId="3" pageNumber="436">Hemiphileurus bispinosus</emphasis>
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Ratcliffe
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</taxonomicName>
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,
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<emphasis id="F697EAF34456E03BFC8B92E3FC68FDB2" bold="true" box="[787,918,510,531]" pageId="3" pageNumber="436">
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<taxonomicNameLabel id="EDA457884456E03BFC8B92E3FC68FDB2" box="[787,918,510,531]" pageId="3" pageNumber="436" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="C45C36E14456E03BFDC39104FD2FFD8F" blockId="3.[397,918,510,558]" box="[603,721,537,558]" pageId="3" pageNumber="436">(Figs. 4–5)</paragraph>
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</subSubSection>
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||
<subSubSection id="8CF9656A4456E03BFE88915DFE1CFAAB" pageId="3" pageNumber="436" type="materials_examined">
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<paragraph id="C45C36E14456E03BFE88915DFE76FD04" blockId="3.[246,1078,576,677]" pageId="3" pageNumber="436">
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<emphasis id="F697EAF34456E03BFE88915DFE46FDF4" bold="true" box="[272,440,576,597]" pageId="3" pageNumber="436">Type Material.</emphasis>
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<materialsCitation id="748B3CBC4456E03BFE27915DFC32FDD1" collectionCode="UFMG" collectorName="D. Yanega" country="Brazil" location="Estacao Ecologica" municipality="Belo Horizonte" pageId="3" pageNumber="436" specimenCount="1" stateProvince="Minas Gerais" typeStatus="holotype">
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<typeStatus id="1B5888434456E03BFE27915DFDDEFDF4" box="[447,544,576,597]" pageId="3" pageNumber="436" type="holotype">Holotype</typeStatus>
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labeled ‘‘
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<collectingCountry id="BCF476714456E03BFD1B915DFD15FDF4" box="[643,747,576,597]" name="Brazil" pageId="3" pageNumber="436">BRAZIL</collectingCountry>
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:
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<collectingRegion id="0627F8034456E03BFD6E915DFC7BFDF4" box="[758,901,576,597]" country="Brazil" name="Minas Gerais" pageId="3" pageNumber="436">Minas Gerais</collectingRegion>
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,
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<collectingMunicipality id="2438AC9B4456E03BFC17915DFBCCFDF4" box="[911,1074,576,597]" pageId="3" pageNumber="436">Belo Horizonte</collectingMunicipality>
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,
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<collectionCode id="A2F2AE244456E03BFF6E9146FEBAFDD1" box="[246,324,603,624]" country="Brazil" httpUri="http://grbio.org/cool/z0ie-wfm6" name="Universidade Federal de Minas Gerais" pageId="3" pageNumber="436">UFMG</collectionCode>
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,
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<location id="C13C603A4456E03BFEC99146FDE9FDD1" LSID="urn:lsid:plazi:treatment:4C4A87F74456E03CFE1592E3FE8DFE9E:C13C603A4456E03BFEC99146FDE9FDD1" box="[337,535,603,624]" country="Brazil" municipality="Belo Horizonte" name="Estacao Ecologica" pageId="3" pageNumber="436" stateProvince="Minas Gerais">Estação Ecologica</location>
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, IVIV1998,
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<collectorName id="691653374456E03BFD349146FCE0FDD1" box="[684,798,603,624]" pageId="3" pageNumber="436">D. Yanega</collectorName>
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, ex blacklight.’’
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</materialsCitation>
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<materialsCitation id="748B3CBC4456E03BFC4D9146FE7AFD04" collectorName="de Zoologia" country="Brazil" location="Museu de Zoologia" pageId="3" pageNumber="436" specimenCount="1" stateProvince="Sao Paulo, Sao Paulo" typeStatus="holotype">
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<typeStatus id="1B5888434456E03BFC4D9146FBC8FDD1" box="[981,1078,603,624]" pageId="3" pageNumber="436" type="holotype">Holotype</typeStatus>
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deposited at the
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<location id="C13C603A4456E03BFE35916BFD7FFD2A" LSID="urn:lsid:plazi:treatment:4C4A87F74456E03CFE1592E3FE8DFE9E:C13C603A4456E03BFE35916BFD7FFD2A" box="[429,641,630,651]" country="Brazil" name="Museu de Zoologia" pageId="3" pageNumber="436" stateProvince="Sao Paulo, Sao Paulo">
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Museu
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<collectorName id="691653374456E03BFE67916BFD7FFD2A" box="[511,641,630,651]" pageId="3" pageNumber="436">de Zoologia</collectorName>
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</location>
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of the Universidade de
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<collectingRegion id="0627F8034456E03BFC08916BFECAFD04" isEnumeration="true" pageId="3" pageNumber="436">São Paulo, São Paulo</collectingRegion>
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, Brazil
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="C45C36E14456E03BFE9591AAFE1CFAAB" blockId="3.[246,1079,695,1290]" pageId="3" pageNumber="436">
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<emphasis id="F697EAF34456E03BFE9591AAFE99FD6B" bold="true" box="[269,359,695,714]" pageId="3" pageNumber="436">
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<typeStatus id="1B5888434456E03BFE9591AAFE99FD6B" box="[269,359,695,714]" pageId="3" pageNumber="436" type="holotype">Holotype</typeStatus>
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</emphasis>
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. Male. Length 16.0 mm; width
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<quantity id="031B9B044456E03BFD0091AAFD1AFD6B" box="[664,740,695,714]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="3" pageNumber="436" unit="mm" value="7.1">7.1 mm</quantity>
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. Color black.
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<emphasis id="F697EAF34456E03BFCF191AAFC65FD6B" box="[873,923,695,714]" italics="true" pageId="3" pageNumber="436">Head</emphasis>
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: Frons in center with deep fovea, surface smooth; frons on sides with moderatelysized horns, horns erect and curving slightly posteriorly. Clypeus triangular, surface nearly smooth, with a few scattered, small, transverse punctures; apex acute and strongly reflexed, a weak carina extending from apex to base of each horn. Interocular width equals 4.0 transverse eye diameters. Antenna 10segmented, club slightly longer than segments 2–7. Mandibles arcuate on lateral edge, apices acute.
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<emphasis id="F697EAF34456E03BFDC8905AFD53FCFB" box="[592,685,839,858]" italics="true" pageId="3" pageNumber="436">Pronotum</emphasis>
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||
: Surface weakly, finely shagreened, punctate; punctures in longitudinal furrow sparse, large, ocellate, punctures just either side of furrow sparse, small; punctures on disc moderate in density, large, ocellate, punctures becoming dense in anterior angles. Median longitudinal furrow deep, extending from base to anterior fifth of pronotum, furrow widest at middle. Base with complete marginal bead.
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<emphasis id="F697EAF34456E03BFEB790A2FE97FC73" box="[303,361,959,978]" italics="true" pageId="3" pageNumber="436">Elytra</emphasis>
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||
: Surface with 5 punctate striae between suture and humeral umbone; punctures moderate to mostly large, umbilicate, separated from one another in each row by about 1 puncture diameter. Odd intervals slightly convex, becoming transversely wrinkled on sides of disc; even intervals with rows of punctures similar to those of striae.
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<taxonomicName id="03E34D624456E03BFF6E9702FE5CFB93" authority=": Surface" authorityName="Surface" box="[246,418,1055,1074]" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="436" phylum="Chordata" rank="genus">
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<emphasis id="F697EAF34456E03BFF6E9702FEB1FB93" box="[246,335,1055,1074]" italics="true" pageId="3" pageNumber="436">Pygidium</emphasis>
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: Surface
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</taxonomicName>
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moderately densely punctate; punctures moderate to moderately large, most setigerous either side of middle; setae short, tawny in color. In lateral view, surface regularly convex.
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<emphasis id="F697EAF34456E03BFE3E9752FE2CFBC3" box="[422,466,1103,1122]" italics="true" pageId="3" pageNumber="436">Legs</emphasis>
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||
: Foretibia tridentate, teeth subequally separated. Median transverse carina on meso and metatibiae crenulate. Apex of posterior tibia with strong, acute spine on upper angle and with several small serrations and a short, broad spinule between each serration (7 spinules on each side). Apex of first tarsomere of posterior tarsus extended into acute spine dorsally.
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<emphasis id="F697EAF34456E03BFDD497B2FD74FB63" box="[588,650,1199,1218]" italics="true" pageId="3" pageNumber="436">Venter</emphasis>
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: Prosternal process short, subtriangular, flattened from front to back, and with small swelling on posterior face near base. Last sternite transversely rugopunctate to punctate in anterior half, apical half nearly smooth.
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<emphasis id="F697EAF34456E03BFF6E97EAFEA2FAAB" box="[246,348,1271,1290]" italics="true" pageId="3" pageNumber="436">Parameres</emphasis>
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: Figures 4–5.
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</paragraph>
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</subSubSection>
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<subSubSection id="8CF9656A4456E03BFE889603FE18FA25" pageId="3" pageNumber="436" type="etymology">
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<paragraph id="C45C36E14456E03BFE889603FE18FA25" blockId="3.[246,1079,1310,1652]" pageId="3" pageNumber="436">
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<emphasis id="F697EAF34456E03BFE889603FE71FA92" bold="true" box="[272,399,1310,1331]" pageId="3" pageNumber="436">Etymology.</emphasis>
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From the Latin
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<emphasis id="F697EAF34456E03BFDDD9602FDA1FA95" box="[581,607,1311,1332]" italics="true" pageId="3" pageNumber="436">bi,</emphasis>
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meaning two, and
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<emphasis id="F697EAF34456E03BFCAB9602FC8CFA95" box="[819,882,1311,1332]" italics="true" pageId="3" pageNumber="436">spina,</emphasis>
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meaning spine or thorn; this combination of words in reference to the two distinctive, curved spines between the parameres of the male genitalia (Figs. 4–5), hence the twospined
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<emphasis id="F697EAF34456E03BFEDB9672FE18FA25" box="[323,486,1391,1412]" italics="true" pageId="3" pageNumber="436">
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<taxonomicName id="03E34D624456E03BFEDB9672FE1CFA25" authorityName="Kolbe" authorityYear="1910" box="[323,482,1391,1412]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="436" phylum="Arthropoda" rank="genus">Hemiphileurus</taxonomicName>
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.
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="8CF9656A4456E03CFE889694FE8DFE9E" lastPageId="4" lastPageNumber="437" pageId="3" pageNumber="436" type="discussion">
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||
<paragraph id="C45C36E14456E03BFE889694FDB1F9D5" blockId="3.[246,1079,1310,1652]" pageId="3" pageNumber="436">
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<emphasis id="F697EAF34456E03BFE889694FE83FA3F" bold="true" box="[272,381,1417,1438]" pageId="3" pageNumber="436">Remarks.</emphasis>
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<taxonomicName id="03E34D624456E03BFE129697FD5FFA3E" authorityName="Ratcliffe" authorityYear="2001" box="[394,673,1418,1439]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="436" phylum="Arthropoda" rank="species" species="bispinosus">
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<emphasis id="F697EAF34456E03BFE129697FD5FFA3E" box="[394,673,1418,1439]" italics="true" pageId="3" pageNumber="436">Hemiphileurus bispinosus</emphasis>
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</taxonomicName>
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will key to couplet
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<quantity id="031B9B044456E03BFC0F9694FC2DFA3F" box="[919,979,1417,1438]" metricMagnitude="-1" metricUnit="m" metricValue="7.874" pageId="3" pageNumber="436" unit="in" value="31.0">31 in</quantity>
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Endrödi (1985) where neither of the subsequent two choices fit the character states. The parameres of
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<taxonomicName id="03E34D624456E03BFE2D96A2FDBAFA75" authorityName="Ratcliffe" authorityYear="2001" box="[437,580,1471,1492]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="436" phylum="Arthropoda" rank="species" species="bispinosus">
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<emphasis id="F697EAF34456E03BFE2D96A2FDBAFA75" box="[437,580,1471,1492]" italics="true" pageId="3" pageNumber="436">H. bispinosus</emphasis>
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</taxonomicName>
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(Figs. 4–5) are remarkable because of the two, long, slender spinelike processes arising from between the parameres. Among all the other species of
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<emphasis id="F697EAF34456E03BFE7396E9FD73F9A8" box="[491,653,1524,1545]" italics="true" pageId="3" pageNumber="436">
|
||
<taxonomicName id="03E34D624456E03BFE7396E9FD77F9A8" authorityName="Kolbe" authorityYear="1910" box="[491,649,1524,1545]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="436" phylum="Arthropoda" rank="genus">Hemiphileurus</taxonomicName>
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,
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</emphasis>
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only
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<taxonomicName id="03E34D624456E03BFD5296E9FEE9F985" authority="Dupuis and Dechambre" authorityName="Dupuis and Dechambre" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="436" phylum="Arthropoda" rank="species" species="elongatus">
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<emphasis id="F697EAF34456E03BFD5296E9FCAFF9A8" box="[714,849,1524,1545]" italics="true" pageId="3" pageNumber="436">H. elongatus</emphasis>
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Dupuis and Dechambre
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</taxonomicName>
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from
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<collectingCountry id="BCF476714456E03BFEFA9512FE51F985" box="[354,431,1551,1572]" name="Bolivia" pageId="3" pageNumber="436">Bolivia</collectingCountry>
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has parameres with internal processes, but these are very different from those of
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<emphasis id="F697EAF34456E03BFE6F9537FD71F99E" box="[503,655,1578,1599]" italics="true" pageId="3" pageNumber="436">
|
||
<taxonomicName id="03E34D624456E03BFE6F9537FD72F99E" authorityName="Ratcliffe" authorityYear="2001" box="[503,652,1578,1599]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="436" phylum="Arthropoda" rank="species" species="bispinosus">H. bispinosus</taxonomicName>
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.
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</emphasis>
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In addition, the form of the prosternal process is unusually short and ‘‘undeveloped’’ in this new species relative to most other species in the genus.
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</paragraph>
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<paragraph id="C45C36E14451E03CFE8B9212FE8DFE9E" blockId="4.[249,1081,271,319]" pageId="4" pageNumber="437">The biology of this species is unknown. The single specimen was taken at blacklights.</paragraph>
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</subSubSection>
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||
</treatment>
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||
</document> |