treatments-xml/data/4B/C3/13/4BC313BC48A451EEABF4F3A8EE529EDA.xml

<document id="72988171C8E7B98F6688A758578443E8" ID-CLB-Dataset="305072" ID-DOI="10.3897/zse.100.136428" ID-GBIF-Dataset="97f910b5-0214-4288-b6a3-1cb400c58d8e" ID-ZooBank="DC99B66D-5862-44E4-B699-0A9CEFE8738F" ID-publisher-id="136428" URI-arpha="3801D9B0-ECD1-554C-9853-18E2A252BEAD" URI-zoobank="http://zoobank.org/DC99B66D-5862-44E4-B699-0A9CEFE8738F" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1731076163562" checkinUser="pensoft" docAuthor="Salvador, Rodrigo Brincalepe &amp; Bichuette, Maria Elina" docDate="2024" docId="4BC313BC48A451EEABF4F3A8EE529EDA" docLanguage="en" docName="Zoosystematics and Evolution 100(4): 1543-1556" docOrigin="Zoosystematics and Evolution 100 (4)" docSource="https://zse.pensoft.net/article/136428/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Idiopyrgus eowynae Salvador &amp; Bichuette, 2024, sp. nov." docType="treatment" docUuid="733E29CA-FF4B-4138-A3A9-05396C501716" docUuidSource="ZooBank" docVersion="4" dtd-version="3.0" lastPageNumber="1556" masterDocId="3801D9B0ECD1554C985318E2A252BEAD" masterDocTitle="Idiopyrgus Pilsbry, 1911 (Gastropoda, Tomichiidae): a relict genus radiating into subterranean environments" masterLastPageNumber="1556" masterPageNumber="1543" pageNumber="1543" updateTime="1731089396751" updateUser="ExternalLinkService">
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<mods:title id="A44D5719D7518924A7DA6EFE70EE7A9F">Idiopyrgus Pilsbry, 1911 (Gastropoda, Tomichiidae): a relict genus radiating into subterranean environments</mods:title>
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<mods:namePart id="6FFA07AF75186A3FDD0ADC187F51EF82">Salvador, Rodrigo Brincalepe</mods:namePart>
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<mods:affiliation id="F3385C31D0F0A265718A5B16E181DFF4">Zoology Unit, Finnish Museum of Natural History, University of Helsinki, Helsinki, Finland &amp; The Arctic University Museum of Norway, UiT – The Arctic University of Norway, Lars Thørings veg 10, 9006, Tromsø, Norway</mods:affiliation>
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<mods:namePart id="80D2839BC4EE0536A2B08253E2D0BD55">Bichuette, Maria Elina</mods:namePart>
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<mods:affiliation id="5BB3FAFD06282D70406FBD5481D8D3CF">Laboratório de Estudos Subterrâneos, Universidade Federal de São Carlos, Rodovia Washington Luís km 235, Caixa Postal 676, 13565 - 905, São Carlos, SP, Brazil</mods:affiliation>
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<subSubSection id="7A311D07ED9A902D70A7FBE48E4DF497" type="nomenclature">
<paragraph id="F7BF6B32769EDB371E28B8238E9185DD">
<taxonomicName id="BDA3E4CD8556497C21193372B92ABC1F" authorityName="Salvador &amp; Bichuette" authorityYear="2024" class="Gastropoda" family="Tomichiidae" genus="Idiopyrgus" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="eowynae" status="sp. nov.">
<emphasis id="528FDB39D53D8047D6C58AC6FE1CE88B" italics="true">Idiopyrgus eowynae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C9DB3590E21CB02A177B211E8D560124" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="FFBD869FE806D7666F60C02D964E0053" type="description">
<paragraph id="61F6C3F75712CA487091E18FCE6BB151">
<figureCitation id="50B7998D67B823CEAADA86A535FA585C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Idiopyrgus eowynae sp. nov. A, B. Holotype LES 29795; . C. Paratype (typical morph) # 1 LES 29796; D. Paratype (typical morph) # 2 LES 29796; E, F. Paratype (dark morph) LES 29797; G. Paratype (typical morph) # 3 LES 29796, juvenile, showing the protoconch in greater detail as well as the sculpture of the first teleoconch whorls; H. Detail of the body whorl of holotype in dorsal view, showing the teleoconch sculpture, in particular the multitude of spiral lines; I. Paratype # 3 in apical view, showing the protoconch." figureDoi="10.3897/zse.100.136428.figure4" httpUri="https://binary.pensoft.net/fig/1170325">Fig. 4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EEHBG" type="type material">
<paragraph id="A52FCE2DA36B0772FAEDA0D178597138">
<heading id="192E2DE5A334AE8F44EBB23D5403A3EB" reason="title">Type material.</heading>
</paragraph>
<paragraph id="8BA28D2AA569D96BC086802076610305">
<materialsCitation id="4D9F31FB6DB8AEAFDFF501EA3EF79AC5" collectingDate="2021-09-10" collectionCode="LES, MNRJ, MZSP" collectorName="M. E. Bichuette &amp; J. E. Gallao" country="Brazil" latitude="-13.796667" location="Gruna do Pedro Cassiano" longLatPrecision="1" longitude="-43.91389" municipality="Carinhanha municipality" specimenCode="LES 29795, LES 29796, LES 29797, MNRJ 37168, MZSP 168419" specimenCount="13" stateProvince="Bahia state" typeStatus="holotype">
<specimenCode id="F21DB0FD14D37CA5876A80D6BA5DFDA1">
<abbrev id="ABBRID0EKHBG" xlink_title="Laboratório de Estudos Subterrâneos, Universidade Federal de São Carlos">LES</abbrev>
29795
</specimenCode>
(
<typeStatus id="1BC7CA03BAF964D6BC30BDC232C21DBD" type="holotype">holotype</typeStatus>
), 
<specimenCode id="14224408905EC7D6752F3257B6AD135C">
<abbrev id="ABBRID0EOHBG" xlink_title="Laboratório de Estudos Subterrâneos, Universidade Federal de São Carlos">LES</abbrev>
29796
</specimenCode>
(4 
<typeStatus id="37B5A8C6F400092369D38C6A5FB2089D">paratypes</typeStatus>
), 
<specimenCode id="72F302FAB1926D98A515E477F9705E87">
<abbrev id="ABBRID0ESHBG" xlink_title="Laboratório de Estudos Subterrâneos, Universidade Federal de São Carlos">LES</abbrev>
29797
</specimenCode>
(2 dark morphs 
<typeStatus id="2657F7F26C797BA0A91E5E66686E1F48" type="paratype">paratype</typeStatus>
), 
<specimenCode id="6933CF57027A29E12D0D259F7C0B3326">
<named-content id="688729706D66528243859E394EDF1EE4" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museu-nacionaluniversidade-federal-de-rio-de-janeiro" xlink_title="Museu Nacional">MNRJ</named-content>
37168
</specimenCode>
(3 
<typeStatus id="A864F456EF96B9BEED18D0829D1D87D7">paratypes</typeStatus>
), 
<specimenCode id="1CCA1823A7AA7C0AB4C5D64E4ED3ABEC">
<abbrev id="ABBRID0E1HBG" xlink_title="Museu de Zoologia da Universidade de São Paulo">MZSP</abbrev>
168419
</specimenCode>
(3 
<typeStatus id="8B894134793F8520FB8B6D6A21C1E733">paratypes</typeStatus>
). All specimens collected on 
<date id="D0E577FFB2D5262E491CB970F76E1431" value="2021-09-10">
<collectingDate id="AF2C6B1A22049E19C81891DE2220CB7A" value="2021-09-10">10. ix. 2021</collectingDate>
</date>
(
<collectorName id="0C6A3B602771C09FF140D5D1FF9D32B0">M. E. Bichuette</collectorName>
&amp; 
<collectorName id="E5AAE274A767FB4DE9CA766D03B86D1F">J. E. Gallão</collectorName>
leg.); all paratypes from type locality (except dark morphs, which are from the same cave but from the entrance zone)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5HBG" type="type locality">
<paragraph id="68017D6D15FC564C42B04A460159036B">
<heading id="EE063E6E890206B21350063D56D2714A" reason="title">Type locality.</heading>
</paragraph>
<paragraph id="69EB249D8FFBFCB188DF0C828D2C3778">
<collectingCountry id="C3CBA709FB209D0EB4B63086CA395C15" name="Brazil">Brazil</collectingCountry>
, 
<collectingRegion id="153898AABDC759889F58B9EFE41B35BB" country="Brazil" name="Bahia">Bahia state</collectingRegion>
, 
<collectingMunicipality id="BFF1CFC21D57A185909C3372AFF0406B">Carinhanha municipality</collectingMunicipality>
, 
<location id="2097E531979DDD594FBA779E6B1BB452" LSID="urn:lsid:plazi:treatment:4BC313BC48A451EEABF4F3A8EE529EDA:2097E531979DDD594FBA779E6B1BB452" country="Brazil" latitude="-13.796667" longLatPrecision="1" longitude="-43.91389" municipality="Carinhanha municipality" name="Gruna do Pedro Cassiano" stateProvince="Bahia state">Gruna do Pedro Cassiano</location>
(“ 
<location id="37DA09B16B60709B179D987189A627C7" LSID="urn:lsid:plazi:treatment:4BC313BC48A451EEABF4F3A8EE529EDA:37DA09B16B60709B179D987189A627C7" country="Brazil" latitude="-13.796667" longLatPrecision="1" longitude="-43.91389" municipality="Carinhanha municipality" name="Pedro Cassiano Cave" stateProvince="Bahia state">Pedro Cassiano Cave</location>
”), 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EHIBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-43.913889,-13.796667]}">
<geoCoordinate id="FC12BC435C1EDDA2FC293F33D278EB0E" degrees="13" direction="south" minutes="47" orientation="latitude" precision="1" seconds="48.0" value="-13.796667">13°47'48.0&quot;S</geoCoordinate>
, 
<geoCoordinate id="F1E4A87CF5F36698FF427B9B7BDE3F78" degrees="43" direction="west" minutes="54" orientation="longitude" precision="1" seconds="50.0" value="-43.91389">43°54'50.0&quot;W</geoCoordinate>
</named-content>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EMIBG" type="etymology">
<paragraph id="C33C43B3DFBF65B78648827C9AE7F360">
<heading id="170D6D223B8B48F82CB805AF37F5FA3B" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="7960D98CE8800127F7EEFC34B46D2059">The species is named in honour of Éowyn, from J. R. R. Tolkien’s “ The Lord of the Rings. ” Éowyn exemplifies courage, resilience, and resistance against darkness, both internal and external, standing against Gríma Wormtongue and the Witch-king of Angmar.</paragraph>
</subSubSection>
<subSubSection id="SECID0ERIBG" type="diagnosis">
<paragraph id="1F08BE938461BC853210D12D90168516">
<heading id="CE381D4A13D41401EDC8CE6EE43AEBEE" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="D4CE68A8D80DDEFCA7E7518FF0558AF4">Shell conical to turriform and relatively tall in comparison to congeners. Body whorl is translucent yellow; previous whorls are slightly darker and brownish. Teleoconch sculpture consisting of few (4–7) radial rows of thorn-like hairs on the apical portion of whorl and a multitude of fine irregular spiral lines below them, reaching all the way to the umbilicus. Aperture large, rounded.</paragraph>
</subSubSection>
<subSubSection id="SECID0EWIBG" type="description">
<paragraph id="E4399EC176F9D99819DAF01FD994D5D2">
<heading id="58C80BBB52A7A45D1E40A73170AADDCA" reason="title">Description.</heading>
</paragraph>
<paragraph id="C6C13F3DCFE5084DBCEC9AE4F56967D3">
Shell conical to turriform, 5–5 ¼ whorls, 
<quantity id="CF8C31B60A69FD633B1B7ABD2A406AC9" metricMagnitude="-3" metricUnit="m" metricValue="4.55" metricValueMax="4.6" metricValueMin="4.5" unit="mm" value="4.55" valueMax="4.6" valueMin="4.5">4.5–4.6 mm</quantity>
high, ~ 
<quantity id="5692FA2AB9B49E3635B486C996FFE54A" metricMagnitude="-3" metricUnit="m" metricValue="2.7" unit="mm" value="2.7">2.7 mm</quantity>
wide. Colour pale beige or yellowish on body whorl, but slightly darker and brownish on earlier whorls; body whorl translucent; early whorls typically with flaked off periostracum. Protoconch of 1 ¼ whorl, rounded, smooth (Fig. 
<figureCitation id="A73B82CA2B27F0A8FF9F67DB87DE0CBE" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Idiopyrgus eowynae sp. nov. A, B. Holotype LES 29795; . C. Paratype (typical morph) # 1 LES 29796; D. Paratype (typical morph) # 2 LES 29796; E, F. Paratype (dark morph) LES 29797; G. Paratype (typical morph) # 3 LES 29796, juvenile, showing the protoconch in greater detail as well as the sculpture of the first teleoconch whorls; H. Detail of the body whorl of holotype in dorsal view, showing the teleoconch sculpture, in particular the multitude of spiral lines; I. Paratype # 3 in apical view, showing the protoconch." figureDoi="10.3897/zse.100.136428.figure4" httpUri="https://binary.pensoft.net/fig/1170325">4 G, I</figureCitation>
). Suture deep. Whorls increasing uniformly in width but more rapidly in height. Teleoconch sculptured by minute triangular thorn-like ‘ periostracal hairs’ arranged in equidistantly spaced spiral rows (4 to 7 rows) and restricted to apical portion of body whorl (Fig. 
<figureCitation id="71BBAAB6C5C2F35A64D1BF0358EFB7AB" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Idiopyrgus eowynae sp. nov. A, B. Holotype LES 29795; . C. Paratype (typical morph) # 1 LES 29796; D. Paratype (typical morph) # 2 LES 29796; E, F. Paratype (dark morph) LES 29797; G. Paratype (typical morph) # 3 LES 29796, juvenile, showing the protoconch in greater detail as well as the sculpture of the first teleoconch whorls; H. Detail of the body whorl of holotype in dorsal view, showing the teleoconch sculpture, in particular the multitude of spiral lines; I. Paratype # 3 in apical view, showing the protoconch." figureDoi="10.3897/zse.100.136428.figure4" httpUri="https://binary.pensoft.net/fig/1170325">4 A – D, H</figureCitation>
). Below the rows of periostracal hairs, there is a multitude of fine, irregular spiral lines, reaching all the way to the umbilicus (Fig. 
<figureCitation id="3CD44CD8E075D65CAEA084DE56E0A071" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Idiopyrgus eowynae sp. nov. A, B. Holotype LES 29795; . C. Paratype (typical morph) # 1 LES 29796; D. Paratype (typical morph) # 2 LES 29796; E, F. Paratype (dark morph) LES 29797; G. Paratype (typical morph) # 3 LES 29796, juvenile, showing the protoconch in greater detail as well as the sculpture of the first teleoconch whorls; H. Detail of the body whorl of holotype in dorsal view, showing the teleoconch sculpture, in particular the multitude of spiral lines; I. Paratype # 3 in apical view, showing the protoconch." figureDoi="10.3897/zse.100.136428.figure4" httpUri="https://binary.pensoft.net/fig/1170325">4 H</figureCitation>
). Periostracal hairs can be worn out in older specimens (Fig. 
<figureCitation id="5B33BAB95F298BCA936F801AAB05121A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Idiopyrgus eowynae sp. nov. A, B. Holotype LES 29795; . C. Paratype (typical morph) # 1 LES 29796; D. Paratype (typical morph) # 2 LES 29796; E, F. Paratype (dark morph) LES 29797; G. Paratype (typical morph) # 3 LES 29796, juvenile, showing the protoconch in greater detail as well as the sculpture of the first teleoconch whorls; H. Detail of the body whorl of holotype in dorsal view, showing the teleoconch sculpture, in particular the multitude of spiral lines; I. Paratype # 3 in apical view, showing the protoconch." figureDoi="10.3897/zse.100.136428.figure4" httpUri="https://binary.pensoft.net/fig/1170325">4 C, D</figureCitation>
), particularly in the early whorls. Peristome complete, not covering penultimate whorl, expanded, wider in abapical region, narrower in columellar region, of same colour as body whorl or slightly whiter. Aperture rounded, with light apical angulation, but with smooth, rounded contour. Umbilicus rimate, nearly closed. Operculum oval, paucispiral; corneous, thin, and translucent, of same colour as body whorl. Soft body completely white in colour.
</paragraph>
<caption id="905F1FD132CB911F82FE63C9BFE37817" ID-DOI="10.3897/zse.100.136428.figure4" ID-arpha="3BC4BB02-D002-5B9B-BBB9-C73BABD6AA68" httpUri="https://binary.pensoft.net/fig/1170325" startId="F4">
<paragraph id="3066A1E4F8C6A8A07483E9460EB4D543">
<label id="E039CEA1492341075CDB8763ACB693D0">Figure 4.</label>
</paragraph>
<paragraph id="000ADB62B00D09D91A3F2F835A20CEC7">
<taxonomicName id="C99AD9119EF862AE628A82D02BC9919D" authorityName="Salvador &amp; Bichuette" authorityYear="2024" class="Gastropoda" family="Pomatiopsidae" genus="Idiopyrgus" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="eowynae" status="sp. nov.">
<emphasis id="45AD4E5765BE7C25A10AB0FCA348CB39" italics="true">Idiopyrgus eowynae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AE062390BA53D37E4B2DA4EE03648670" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="5EA6D05D303D516A308BD7D4DEC760D7" bold="true">A, B.</emphasis>
Holotype 
<abbrev id="ABBRID0EBKBG" xlink_title="Laboratório de Estudos Subterrâneos, Universidade Federal de São Carlos">LES</abbrev>
29795;. 
<emphasis id="55EB71ADB47F99192494128889CD46FB" bold="true">C.</emphasis>
Paratype (typical morph) # 1 
<abbrev id="ABBRID0EHKBG" xlink_title="Laboratório de Estudos Subterrâneos, Universidade Federal de São Carlos">LES</abbrev>
29796; 
<emphasis id="845C8A010170CFC8F32CAE26F1A5AC71" bold="true">D.</emphasis>
Paratype (typical morph) # 2 
<abbrev id="ABBRID0ENKBG" xlink_title="Laboratório de Estudos Subterrâneos, Universidade Federal de São Carlos">LES</abbrev>
29796; 
<emphasis id="EA42DA48A8BA0C047259294735ED6576" bold="true">E, F.</emphasis>
Paratype (dark morph) 
<abbrev id="ABBRID0ETKBG" xlink_title="Laboratório de Estudos Subterrâneos, Universidade Federal de São Carlos">LES</abbrev>
29797; 
<emphasis id="BD74F4957ABD7525538AD3EF1B895E5C" bold="true">G.</emphasis>
Paratype (typical morph) # 3 
<abbrev id="ABBRID0EZKBG" xlink_title="Laboratório de Estudos Subterrâneos, Universidade Federal de São Carlos">LES</abbrev>
29796, juvenile, showing the protoconch in greater detail as well as the sculpture of the first teleoconch whorls; 
<emphasis id="6E332C6982821B6C6AC983D091C983CE" bold="true">H.</emphasis>
Detail of the body whorl of holotype in dorsal view, showing the teleoconch sculpture, in particular the multitude of spiral lines; 
<emphasis id="4382AFA8A905EE69A90FD62652A56C08" bold="true">I.</emphasis>
Paratype # 3 in apical view, showing the protoconch.
</paragraph>
</caption>
<paragraph id="3A2222171532A6F3ADD2912D983FF1D7">
Dark morph (Fig. 
<figureCitation id="8ACBA007FB00A6B4F82BB78AADF1BF6C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Idiopyrgus eowynae sp. nov. A, B. Holotype LES 29795; . C. Paratype (typical morph) # 1 LES 29796; D. Paratype (typical morph) # 2 LES 29796; E, F. Paratype (dark morph) LES 29797; G. Paratype (typical morph) # 3 LES 29796, juvenile, showing the protoconch in greater detail as well as the sculpture of the first teleoconch whorls; H. Detail of the body whorl of holotype in dorsal view, showing the teleoconch sculpture, in particular the multitude of spiral lines; I. Paratype # 3 in apical view, showing the protoconch." figureDoi="10.3897/zse.100.136428.figure4" httpUri="https://binary.pensoft.net/fig/1170325">4 E, F</figureCitation>
) differs from the above-described typical morph by having a black shell with thicker walls, in which the teleoconch sculpture is nearly invisible: a few remnants of the spiral rows of hairs are visible in the adapical portion of the whorls; traces of the sculpture can also be seen through the shell wall of the aperture (Fig. 
<figureCitation id="38B5523DC50F752CF2043A8FDBA45AB4" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Idiopyrgus eowynae sp. nov. A, B. Holotype LES 29795; . C. Paratype (typical morph) # 1 LES 29796; D. Paratype (typical morph) # 2 LES 29796; E, F. Paratype (dark morph) LES 29797; G. Paratype (typical morph) # 3 LES 29796, juvenile, showing the protoconch in greater detail as well as the sculpture of the first teleoconch whorls; H. Detail of the body whorl of holotype in dorsal view, showing the teleoconch sculpture, in particular the multitude of spiral lines; I. Paratype # 3 in apical view, showing the protoconch." figureDoi="10.3897/zse.100.136428.figure4" httpUri="https://binary.pensoft.net/fig/1170325">4 E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQLBG" type="distribution">
<paragraph id="41041CC7009F3BBAB318170A50642AAB">
<heading id="6E3B8606BA31CDCF47538453600B663A" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="1E967CE9921CC37DCA44F2E3B5D372F9">
Known only from type locality (Fig. 
<figureCitation id="9371770DB95B94520E85DEB940024A30" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution map of Idiopyrgus spp. (and Spiripockia punctata) in caves of the state of Minas Gerais and Bahia, Brazil." figureDoi="10.3897/zse.100.136428.figure1" httpUri="https://binary.pensoft.net/fig/1170322">1</figureCitation>
), the twilight and dark (aphotic) zones of the cave (typical morph) and at the cave entrance (dark morph).
</paragraph>
</subSubSection>
<subSubSection id="SECID0E1LBG" type="remarks">
<paragraph id="0193F8E02B45DCE8B589214B5204E79E">
<heading id="BA4AC425F7B935582E50454A2BFB3394" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="A72EF471CD71E6F93927195E6F98A094">
<taxonomicName id="21FC5EAEE6C3E7E3DE6A032453A8886D" authorityName="Salvador &amp; Bichuette" authorityYear="2024" class="Gastropoda" family="Pomatiopsidae" genus="Idiopyrgus" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="eowynae" status="sp. nov.">
<emphasis id="3581EECA2469E8C0659D92DAAA79CBAE" italics="true">Idiopyrgus eowynae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D5C54A21C35DBCE13531C285650F02B6" rank="species">sp. nov.</taxonomicNameLabel>
can be easily distinguished from most of its congeners (including the type species 
<taxonomicName id="D5F5D32EA31375CFB15E31971853C831" authorityName="Pilsbry" authorityYear="1911" class="Gastropoda" family="Pomatiopsidae" genus="Idiopyrgus" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="souleyetianus">
<emphasis id="178AA6344D4A8FD175B9FE36F44FDD86" italics="true">I. souleyetianus</emphasis>
</taxonomicName>
) by its much wider shell and the presence of teleoconch sculpture. It closely resembles 
<taxonomicName id="C0D69A271DD912AD14B40285147455A3" authorityName="Salvador &amp; Bichuette" authorityYear="2024" class="Gastropoda" family="Pomatiopsidae" genus="Idiopyrgus" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="meriadoci" status="sp. nov.">
<emphasis id="DA8892F229E2768770AD5D8086864D4D" italics="true">I. meriadoci</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E07E47A54F82A842E5A503FF23FBA199" rank="species">sp. nov.</taxonomicNameLabel>
(see below) and 
<taxonomicName id="3A7F2F1F32F555849F28EC0890DD57A7" class="Gastropoda" family="Cochliopidae" genus="Spiripockia" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="umbraticola">
<emphasis id="66196E94D182844A7C351487D8B4E925" italics="true">I. umbraticola</emphasis>
</taxonomicName>
. It differs from 
<taxonomicName id="887AD10695607DFA9107072C0D4B3F41" authorityName="Salvador &amp; Bichuette" authorityYear="2024" class="Gastropoda" family="Pomatiopsidae" genus="Idiopyrgus" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="meriadoci" status="sp. nov.">
<emphasis id="82BBD2390B9A622451EF2DC3EC0672A5" italics="true">I. meriadoci</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C4B55747C2B277B21FBCC18C82470792" rank="species">sp. nov.</taxonomicNameLabel>
by a much larger size and larger number of whorls, as well as by having a different teleoconch sculpture, which includes periostracal hairs. Conchologically, 
<taxonomicName id="5DFD4D668EC3742130338EF115BE2827" authorityName="Salvador &amp; Bichuette" authorityYear="2024" class="Gastropoda" family="Pomatiopsidae" genus="Idiopyrgus" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="eowynae" status="sp. nov.">
<emphasis id="DC9B37CC5F466A47847E3118C2BD6965" italics="true">I. eowynae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7FE9DF19AA1AF8781E5B8FCAEA35DFBB" rank="species">sp. nov.</taxonomicNameLabel>
is most similar to 
<taxonomicName id="5BEE8D3F7A071E93A73E934C788581EF" authorityName="Salvador &amp; Bichuette" authorityYear="2024" baseAuthorityName="Salvador &amp; Bichuette" baseAuthorityYear="2024" class="Gastropoda" family="Cochliopidae" genus="Spiripockia" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="umbraticola" status="comb. nov.">
<emphasis id="1AD5437D8B6AFBE7D5939F6862DD9C36" italics="true">I. umbraticola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="47FEE960D0CE9E83733BB908591441BE" rank="species">comb. nov.</taxonomicNameLabel>
, from which it can be distinguished by its smaller size (~ 
<quantity id="1FC33852A6F674E9B9D5041D532C0A50" metricMagnitude="-3" metricUnit="m" metricValue="4.5" unit="mm" value="4.5">4.5 mm</quantity>
vs. ~ 5.0 mm in 
<taxonomicName id="E507B95E832B43B13958AB5DFD1BD0D3" authorityName="Salvador &amp; Bichuette" authorityYear="2024" baseAuthorityName="Salvador &amp; Bichuette" baseAuthorityYear="2024" class="Gastropoda" family="Cochliopidae" genus="Spiripockia" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="umbraticola">
<emphasis id="83BE24972028424B8F7860D328015378" italics="true">I. umbraticola</emphasis>
</taxonomicName>
), higher whorls with a more strongly convex outline, a larger and rounder aperture, and by having a different teleoconch sculpture. In 
<taxonomicName id="D44F5CC4C352CF4891D170908250E518" authorityName="Salvador &amp; Bichuette" authorityYear="2024" class="Gastropoda" family="Pomatiopsidae" genus="Idiopyrgus" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="eowynae" status="sp. nov.">
<emphasis id="1998B8C950F87656DA0C78A7DABE749D" italics="true">I. eowynae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6A8C0E6A9DE5A0CF380E1044408BF169" rank="species">sp. nov.</taxonomicNameLabel>
, there are few spiral rows of the triangular thorn-like hairs, restricted to the apical portion of the whorl, and which are then replaced by irregular spiral lines that continue towards the umbilicus (also visible in juveniles). In 
<taxonomicName id="91838E9B40E4228AD07457E472C30353" authorityName="Salvador &amp; Bichuette" authorityYear="2024" baseAuthorityName="Salvador &amp; Bichuette" baseAuthorityYear="2024" class="Gastropoda" family="Cochliopidae" genus="Spiripockia" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="umbraticola">
<emphasis id="5A2F0FB7DE9744A7BAA87157A111F476" italics="true">I. umbraticola</emphasis>
</taxonomicName>
, there is a larger number of rows of hairs, stretching the median portion of the whorl; the spiral lines are absent. Notably, a dark morph has also been observed in 
<taxonomicName id="02259FF8A113080C65BC8EA1C966DE42" authorityName="Salvador, F. S. Silva &amp; Bichuette" authorityYear="2022" class="Gastropoda" family="Pomatiopsidae" genus="Idiopyrgus" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="adamanteus">
<emphasis id="48CD6C75C6226643EE1B0F8B4A6503A3" italics="true">I. adamanteus</emphasis>
</taxonomicName>
<bibRefCitation id="E4FEC35FA610820775C73D3C6B55334B" DOI="10.3897/zse.98.90797" author="Salvador RB &amp; Silva FS &amp; Bichuette ME" issue="2" journalOrPublisher="Zoosystematics and Evolution" pagination="365-375" refId="B31" refString="Salvador RB, Silva FS, Bichuette ME (2022 a) Phylogenetic position of the relict South American genus Idiopyrgus Pilsbry, 1911 (Gastropoda, Truncatelloidea), with the description of two new cave species. Zoosystematics and Evolution 98 (2): 365 - 375. https: // doi. org / 10.3897 / zse. 98.90797" title="Phylogenetic position of the relict South American genus Idiopyrgus Pilsbry, 1911 (Gastropoda, Truncatelloidea), with the description of two new cave species." volume="98" year="2022">Salvador, Silva &amp; Bichuette, 2022</bibRefCitation>
, and 
<taxonomicName id="5F12986BF24CC67C40C2E92649D0B042" authority="Pilsbry, 1924" authorityName="Pilsbry" authorityYear="1924" class="Gastropoda" family="Pomatiopsidae" genus="Idiopyrgus" isUncertain="true" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="walkeri">
<emphasis id="CDE9C63FD4181DEB2ECE239B09C2F1CE" italics="true">I.</emphasis>
cf. 
<emphasis id="61BB7D552F5EE6B778078048F1002833" italics="true">walkeri</emphasis>
Pilsbry, 1924
</taxonomicName>
(
<bibRefCitation id="BD118BF2F8792795F11E089977CC3D64" DOI="10.3897/zse.98.90797" author="Salvador RB &amp; Silva FS &amp; Bichuette ME" issue="2" journalOrPublisher="Zoosystematics and Evolution" pagination="365-375" refId="B31" refString="Salvador RB, Silva FS, Bichuette ME (2022 a) Phylogenetic position of the relict South American genus Idiopyrgus Pilsbry, 1911 (Gastropoda, Truncatelloidea), with the description of two new cave species. Zoosystematics and Evolution 98 (2): 365–375. https://doi.org/10.3897/zse.98.90797" title="Phylogenetic position of the relict South American genus Idiopyrgus Pilsbry, 1911 (Gastropoda, Truncatelloidea), with the description of two new cave species." volume="98">Salvador et al. 2022 a</bibRefCitation>
).
</paragraph>
<paragraph id="4A10C92DFE69BFE6240961F000F5E77C">
Except for the existence of two morphs (typical and black), there is little variation in shell shape, form, and proportions among the presently available specimens of 
<taxonomicName id="F484CC7B821B69DAF4BC882087C68348" authorityName="Salvador &amp; Bichuette" authorityYear="2024" class="Gastropoda" family="Pomatiopsidae" genus="Idiopyrgus" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="eowynae" status="sp. nov.">
<emphasis id="011860B1E9E3B6025B3AF47D2EDFD54A" italics="true">I. eowynae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="826FA5413BC53B8B4D0DE524F2A3B1F7" rank="species">sp. nov.</taxonomicNameLabel>
(Fig. 
<figureCitation id="F4E542996B7AB9B9521FDED265680A9C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Idiopyrgus eowynae sp. nov. A, B. Holotype LES 29795; . C. Paratype (typical morph) # 1 LES 29796; D. Paratype (typical morph) # 2 LES 29796; E, F. Paratype (dark morph) LES 29797; G. Paratype (typical morph) # 3 LES 29796, juvenile, showing the protoconch in greater detail as well as the sculpture of the first teleoconch whorls; H. Detail of the body whorl of holotype in dorsal view, showing the teleoconch sculpture, in particular the multitude of spiral lines; I. Paratype # 3 in apical view, showing the protoconch." figureDoi="10.3897/zse.100.136428.figure4" httpUri="https://binary.pensoft.net/fig/1170325">4 A – F</figureCitation>
). Notably, the sequences of the COI barcoding gene fragment of the two morphs were 100 % identical, while the sequences of both morphs had a pairwise identity of around 90 % in relation to 
<taxonomicName id="1FECF37A14C25F3793307C6B2F3CD2A1" authorityName="Salvador, F. S. Silva &amp; Bichuette" authorityYear="2022" class="Gastropoda" family="Pomatiopsidae" genus="Idiopyrgus" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="adamanteus">
<emphasis id="A1845197455077F08F3C2EC862550F08" italics="true">I. adamanteus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4362FCB15D7E3CB388A1A64E6F205968" authorityName="Salvador, F. S. Silva &amp; Bichuette" authorityYear="2022" class="Gastropoda" family="Pomatiopsidae" genus="Idiopyrgus" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="minor">
<emphasis id="4DBCCD02E3F2F9DF1310A5C3070AC303" italics="true">I. minor</emphasis>
</taxonomicName>
. According to the results of the phylogenetic analysis (Fig. 
<figureCitation id="BE2DE6DC2AB75BA7083B2441D09EFE44" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Bayesian inference phylogenetic tree based on concatenated but unlinked markers, highlighting the family Tomichiidae and showing the position of the new species in the genus Idiopyrgus and its relationship to its congeners. Posterior probabilities are shown on the nodes. Scale bar is substitutions per site." figureDoi="10.3897/zse.100.136428.figure3" httpUri="https://binary.pensoft.net/fig/1170324">3</figureCitation>
), 
<taxonomicName id="98BAB9785C839D90C2D9899AA61897FA" authorityName="Salvador &amp; Bichuette" authorityYear="2024" class="Gastropoda" family="Pomatiopsidae" genus="Idiopyrgus" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="eowynae" status="sp. nov.">
<emphasis id="781015338B157FA0858DD33EECF32DB8" italics="true">I. eowynae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5432925177D76BA7E92356DD4DB9D892" rank="species">sp. nov.</taxonomicNameLabel>
is sister to 
<taxonomicName id="9C2B9CD2746C251BC4D9AD40A586E3AD" authorityName="Salvador, F. S. Silva &amp; Bichuette" authorityYear="2022" class="Gastropoda" family="Pomatiopsidae" genus="Idiopyrgus" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="adamanteus">
<emphasis id="03BCF579921919E44A7760C66E87DB0E" italics="true">I. adamanteus</emphasis>
</taxonomicName>
; both species together are sister to 
<taxonomicName id="8414CC0862672904B70A7A8E3F7DB14E" authorityName="Salvador, F. S. Silva &amp; Bichuette" authorityYear="2022" class="Gastropoda" family="Pomatiopsidae" genus="Idiopyrgus" kingdom="Animalia" order="Littorinimorpha" phylum="Mollusca" rank="species" species="minor">
<emphasis id="C90168440048F852B65505B49DE087F0" italics="true">I. minor</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>