169 lines
17 KiB
XML
169 lines
17 KiB
XML
<document id="7DF85E37FDA6C4A245DA30727A035D38" ID-DOI="http://dx.doi.org/10.3897/zookeys.692.14706" ID-GBIF-Dataset="81f065e7-a342-495c-9855-21bc26539311" ID-PMC="PMC5672734" ID-Pensoft-Pub="1313-2970-692-1" ID-PubMed="29133987" ID-ZooBank="1B7C3F517CF543338EAB1CB1BD9D8A07" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-692-1" ModsDocOrigin="ZooKeys 692" ModsDocTitle="A review of the land snail genus Alycaeus (Gastropoda, Alycaeidae) in Peninsular Malaysia" checkinTime="1503352913593" checkinUser="pensoft" docAuthor="Foon, Junn Kitt & Liew, Thor-Seng" docDate="2017" docId="4B673B16195781472C33A94D9F4A9C07" docLanguage="en" docName="ZooKeys 692: 1-81" docOrigin="ZooKeys 692" docSource="http://dx.doi.org/10.3897/zookeys.692.14706" docTitle="Alycaeus carinata Maassen 2006" docType="treatment" docVersion="6" lastPageNumber="15" masterDocId="FF8E6C15A17CDA48FFA7FFE36C7CFFA2" masterDocTitle="A review of the land snail genus Alycaeus (Gastropoda, Alycaeidae) in Peninsular Malaysia" masterLastPageNumber="81" masterPageNumber="1" pageNumber="13" updateTime="1701315888619" updateUser="plazi">
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<mods:title id="03F78CBDDA661E61FDD7D449CD7F6AC1">A review of the land snail genus Alycaeus (Gastropoda, Alycaeidae) in Peninsular Malaysia</mods:title>
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<mods:namePart id="D059820AF7854D9ED9D721506CD637AC">Foon, Junn Kitt</mods:namePart>
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<mods:namePart id="1A6D8A9176E70CDB761CDADC287FB60A">Liew, Thor-Seng</mods:namePart>
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<treatment id="4B673B16195781472C33A94D9F4A9C07" ID-GBIF-Taxon="132786257" LSID="urn:lsid:plazi:treatment:4B673B16195781472C33A94D9F4A9C07" httpUri="http://treatment.plazi.org/id/4B673B16195781472C33A94D9F4A9C07" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13">
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<subSubSection id="2134A39A55413EE88CE7FDEA9FDA4D5D" pageId="12" pageNumber="13" type="nomenclature">
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<paragraph id="7E1C79D62206A25DEF9F6B6BAE52F0AA" pageId="12" pageNumber="13">
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<taxonomicName id="E1E2D59F1BB115CFA3D14EAB7FF88D6D" ID-CoL="CBZR" authority="Maassen, 2006" authorityName="Maassen" authorityYear="2006" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus carinata" order="Architaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="carinata">
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<pageBreakToken id="CF16D4A0A8101589A5CA70508C53B552" pageId="12" pageNumber="13" start="start">Alycaeus</pageBreakToken>
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carinata Maassen, 2006
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</taxonomicName>
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Figures 11, 31Q
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</paragraph>
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</subSubSection>
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<subSubSection id="830569A4BCF4D195DE7855FD089CF08A" pageId="13" pageNumber="14" type="reference_group">
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<paragraph id="E19A1E55D1B6E70D222947A0437169B9" pageId="13" pageNumber="14">
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<taxonomicName id="9781B982116989C5760AA5A1E37D5B20" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus carinata" order="Architaenioglossa" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="carinata">
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<pageBreakToken id="6F8F0895917CBE14AD3B3808B85DB8AF" pageId="13" pageNumber="14" start="start">Alycaeus</pageBreakToken>
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carinata
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</taxonomicName>
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Maassen, 2006: 137-138, figures 10-13;
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<bibRefCitation id="54885A3616556D5DEE76248A66F0FD27" author="Tarruella, A" journalOrPublisher="Spira" pageId="53" pageNumber="54" pagination="71 - 76" title="Listado taxonomico ilustrado de la familia Cyclophoridae J. E. Gray, 1847 (Mollusca: Gastropoda): Parte 2. La subfamilia Alycaeinae J. E. Gray, 1850." url="http://www.molluscat.com/assets/spira_4_12_7.pdf" volume="4" year="2011">
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Tarruella and
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<normalizedToken id="A8C972FC8D392058DB40AF9960978B2B" originalValue="Domènech">Domenech</normalizedToken>
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2011
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</bibRefCitation>
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: 72.
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</paragraph>
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</subSubSection>
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<subSubSection id="22EA581F34F79CEB9C8EF0CE1F6E1259" pageId="13" pageNumber="14" type="type locality">
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<paragraph id="84DF8E14A5960B0252F5D554864460E5" pageId="13" pageNumber="14">Type locality.</paragraph>
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<paragraph id="C42E1CB60CD6521BE511B7040CBEEF9D" pageId="13" pageNumber="14">
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PHG 74 Bukit Sagu, Pahang (
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<geoCoordinate id="3799C7E56771FA3BB690DF403146281E" direction="north" orientation="latitude" precision="15" value="3.9816668">3°58'54"N</geoCoordinate>
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,
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<geoCoordinate id="872458C67DD140BC972077E0B4635C6E" direction="east" orientation="longitude" precision="15" value="103.144165">103°08'39"E</geoCoordinate>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="96983939CD284017A22B87948443F21D" pageId="13" pageNumber="14" type="type material">
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<paragraph id="DE8C19042BB33B7AE3FBE226CF2F333F" pageId="13" pageNumber="14">Type material.</paragraph>
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<paragraph id="7051C634323E4E80C9DCBD3D3727DD2F" pageId="13" pageNumber="14">Holotype. PHG 74 Bukit Sagu, Pahang: RMNH 104428 (Not seen). Paratypes. PHG 74 Bukit Sagu, Pahang: RMNH 104427/2 (1 specimen seen), HW /58 (Not seen), ANSP /2 (Not seen), HW /200 (Not seen). PHG 75 Bukit Tenggek, Pahang: RMNH 104426/2 (Not seen), HW /37 (Not seen).</paragraph>
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</subSubSection>
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<subSubSection id="A948DEC3BEE88E0F9163DD8F4BA63A52" pageId="13" pageNumber="14" type="description">
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<paragraph id="CABC82E1B2320E8E3076A04A52A26403" pageId="13" pageNumber="14">Description.</paragraph>
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<paragraph id="9AC2C0CFA30D4277FAE49B3552F19012" pageId="13" pageNumber="14">Protoconch. Smooth.</paragraph>
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<paragraph id="2BBEE886F0D76A041128CB2DC726FDDE" pageId="13" pageNumber="14">
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Shell shape. Almost flat, conical. Shell height: 2.2-3.0 mm. Shell width: 4.0-4.6 mm. Shell measurements derived from
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<bibRefCitation id="DFAA6442985580EB4D618DE8FDDE8FF5" author="Maassen, WJM" journalOrPublisher="Basteria" pageId="51" pageNumber="52" pagination="133 - 139" title="Remarks on Alycaeus species from South-East Asia, with descriptions of four new species with keeled shells (Gastropoda, Caenogastropoda, Cyclophoridae)." url="http://natuurtijdschriften.nl/record/597326" volume="70" year="2006">Maassen (2006)</bibRefCitation>
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and re-examination of one paratype (RMNH 104427).
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</paragraph>
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<paragraph id="FC07F2CB709C5731A386E41C6C865F56" pageId="13" pageNumber="14">Spire. Spire height: 3.0 mm. Spire width: 4.5 mm. Number of whorls: up to 4. Spire shape: oblong conical. Whorl periphery strongly keeled except posterior of the constriction, where it becomes rounded. Umbilicus open.</paragraph>
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<paragraph id="638560A8B4EAC20D29C6B8E97991D53D" pageId="13" pageNumber="14">
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Whorl constriction. At about 3
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<normalizedToken id="A3B64CA651282E81B994057B0FA24812" originalValue="¾">3/4</normalizedToken>
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whorls posterior of protoconch.
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</paragraph>
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<paragraph id="4A601D16007C11E53E1A2BA883221889" pageId="13" pageNumber="14">Breathing tube. Length: 0.3 mm.</paragraph>
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<paragraph id="FA7047D921513A5B7E161900272EDBB3" pageId="13" pageNumber="14">Aperture and peristome. Aperture circular, moderately expanded. Aperture height: 2.2 mm. Aperture width: 2.0 mm. Peristome double, not thickened, winged at suture. Interspace narrow. Peristome orientation is 30-55° oblique with respect to the coiling axis.</paragraph>
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<paragraph id="FBF014E466E5596F85E36059120ED859" pageId="13" pageNumber="14">Spiral lines. Indistinct. Regularly spaced. Approximately 25 lines per 1 mm.</paragraph>
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<paragraph id="37F1A2CF51DB8AB1AB74A29BD8868560" pageId="13" pageNumber="14">Radial ribs running anterior of breathing tube. Radial ribs indistinct to absent, only unevenly spaced radial growth lines present.</paragraph>
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<paragraph id="E98AA124C9A333912AE97746A241D4A5" pageId="13" pageNumber="14">Radial ribs running perpendicular to breathing tube. Radial ribs pronounced, evenly spaced. Approximately 13 ribs per 1 mm.</paragraph>
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<paragraph id="72B963833D0A35BA792E54E1A9BF0548" pageId="13" pageNumber="14">Radial ribs running posterior of breathing tube. Radial ribs indistinct to absent, only unevenly spaced radial growth lines present.</paragraph>
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<paragraph id="69F53E9252C9B482B1C2A9B2D6BBD784" pageId="13" pageNumber="14">
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Operculum. Concave, rounded. Exterior and interior composition and sculpture not examined.
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<bibRefCitation id="75774BC80EFD5A7354C3E1D4F7FFAF22" author="Maassen, WJM" journalOrPublisher="Basteria" pageId="51" pageNumber="52" pagination="133 - 139" title="Remarks on Alycaeus species from South-East Asia, with descriptions of four new species with keeled shells (Gastropoda, Caenogastropoda, Cyclophoridae)." url="http://natuurtijdschriften.nl/record/597326" volume="70" year="2006">Maassen (2006)</bibRefCitation>
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considered the operculum entirely proteinaceous.
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</paragraph>
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<paragraph id="C937F10C929D4BE2CA1C57A161B0717D" pageId="13" pageNumber="14">Shell colour. Whorls yellow to brown. Peristome white.</paragraph>
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<paragraph id="5C9BC923D9DB339E5EF46B66F5112C16" pageId="13" pageNumber="14">Living animal. Unknown.</paragraph>
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</subSubSection>
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||
<subSubSection id="2CDDC9F41E7F16B4B2493F77D6B3D0C3" pageId="13" pageNumber="14" type="habitat">
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<paragraph id="2122C5A4C338DC41C5C6D59158199470" pageId="13" pageNumber="14">Habitat and ecology.</paragraph>
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<paragraph id="6548A382ECD7B65E27C78A90D9DDBD6F" pageId="13" pageNumber="14">
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Lives on limestone walls in a large, collapsed cave (
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<bibRefCitation id="5063EDC52E88D91DEB932D7AAB35EE63" author="Maassen, WJM" journalOrPublisher="Basteria" pageId="51" pageNumber="52" pagination="133 - 139" title="Remarks on Alycaeus species from South-East Asia, with descriptions of four new species with keeled shells (Gastropoda, Caenogastropoda, Cyclophoridae)." url="http://natuurtijdschriften.nl/record/597326" volume="70" year="2006">Maassen 2006</bibRefCitation>
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). Probably prefers very moist environments only (
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<bibRefCitation id="2BF49124EDFA2C9B64CC2EDB2BFC3AB9" author="Maassen, WJM" journalOrPublisher="Basteria" pageId="51" pageNumber="52" pagination="133 - 139" title="Remarks on Alycaeus species from South-East Asia, with descriptions of four new species with keeled shells (Gastropoda, Caenogastropoda, Cyclophoridae)." url="http://natuurtijdschriften.nl/record/597326" volume="70" year="2006">Maassen 2006</bibRefCitation>
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||
).
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="70D3EB3D647FE728DB0FA57FB62991BC" pageId="13" pageNumber="14" type="distribution">
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<paragraph id="4EC684D362193C7BAACE257113B94D32" pageId="13" pageNumber="14">Distribution range.</paragraph>
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<paragraph id="93D514DF37838F088855A65B860EF09D" pageId="13" pageNumber="14">Restricted to Bukit Sagu and Bukit Tenggek, Pahang.</paragraph>
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</subSubSection>
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<subSubSection id="4850A9736C27D05ADA0FF632A3C465FE" pageId="13" pageNumber="14" type="differential diagnosis">
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<paragraph id="948113732510FD8E5EF0753A91F5453B" pageId="13" pageNumber="14">Differential diagnosis.</paragraph>
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<paragraph id="C520F22661C6E4DB9625185E5DFAF30E" pageId="13" pageNumber="14">
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<taxonomicName id="836EFB907427A12F9CE4EED87C96A493" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus carinata" order="Architaenioglossa" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="carinata">Alycaeus carinata</taxonomicName>
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is distinguished from all other Peninsular Malaysian
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<taxonomicName id="8F6F74BB6E1C028CCA0DF176E2587624" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus" order="Architaenioglossa" pageId="13" pageNumber="14" phylum="Mollusca" rank="genus">Alycaeus</taxonomicName>
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species by its strongly keeled periphery, relatively low spire, very expanded ultimate whorls and very oblique peristome orientation with respect to the coiling axis (
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<bibRefCitation id="D5886E5552B5E92F58904F2CAEA95019" author="Maassen, WJM" journalOrPublisher="Basteria" pageId="51" pageNumber="52" pagination="133 - 139" title="Remarks on Alycaeus species from South-East Asia, with descriptions of four new species with keeled shells (Gastropoda, Caenogastropoda, Cyclophoridae)." url="http://natuurtijdschriften.nl/record/597326" volume="70" year="2006">Maassen 2006</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="71AC36F36A016AD167FC81CA6D983C09" pageId="14" pageNumber="15" type="discussion">
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<paragraph id="15B19D2A8975844BC55AE0177ACF89A7" pageId="14" pageNumber="15">
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<pageBreakToken id="8218200143F3F3DD6B684935696D5777" pageId="14" pageNumber="15" start="start">Discussion</pageBreakToken>
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.
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</paragraph>
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<paragraph id="2CB282EBEB181F8DEFC2905D3B0F0328" pageId="14" pageNumber="15">
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Interestingly, no
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<taxonomicName id="00CD316F07961890E1C73ED1687CE5E0" lsidName="A. carinata" pageId="14" pageNumber="15" rank="species" species="carinata">A. carinata</taxonomicName>
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was found during the 1950s University of Malaya, Singapore expedition to the type locality although congeners
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<taxonomicName id="203BEF25ECB584D4E81C8F1C0700CE78" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus charasensis" order="Architaenioglossa" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="charasensis">Alycaeus charasensis</taxonomicName>
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sp. n. and
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<taxonomicName id="560EF0C9AAAE4EEB43592DE0367CF503" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus expansus" order="Architaenioglossa" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="expansus">Alycaeus expansus</taxonomicName>
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sp. n. were obtained (ZRC collection lots). This suggests
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<taxonomicName id="9FE40A11C4E9549EBC73BDF912C7444D" lsidName="A. carinata" pageId="14" pageNumber="15" rank="species" species="carinata">A. carinata</taxonomicName>
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may be restricted to upper sections of limestone hills that became accessible only during quarrying in the 1990s. However, viable habitats at Bukit Sagu and Bukit Tenggek are now very likely lost to quarrying (
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<bibRefCitation id="FF25DCC92F31173FC9A70DD85C328A82" author="Schilthuizen, M" journalOrPublisher="Tentacle" pageId="53" pageNumber="54" pagination="8 - 9" title="Tracking extinction from space." url="https://www.hawaii.edu/cowielab/tentacle/tentacle_16.pdf" volume="16" year="2008">Schilthuizen and Clements 2008</bibRefCitation>
|
||
,
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||
<bibRefCitation id="2BB3FCA125AEB93A5875E74C6C73D8FD" author="Liew, T" journalOrPublisher="ZooKeys" pageId="51" pageNumber="52" pagination="1 - 107" title="A cybertaxonomic revision of the micro-landsnail genus Plectostoma Adam (Mollusca, Caenogastropoda, Diplommatinidae), from Peninsular Malaysia, Sumatra and Indochina." url="https://doi.org/10.3897/zookeys.393.6717" volume="393" year="2014">Liew et al. 2014</bibRefCitation>
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||
).
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||
</paragraph>
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<caption id="C4766E2D33CA2FEE46E7373AC9B4117E" pageId="14" pageNumber="15">
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<paragraph id="D3E3B46C345D051C8EE566560E9E6BC4" pageId="14" pageNumber="15">
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Figure 11.
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<taxonomicName id="D40A059E90C9DEAF5DDDF7283E1DCEA0" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus carinata" order="Architaenioglossa" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="carinata">Alycaeus carinata</taxonomicName>
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Maassen, 2006
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<normalizedToken id="A60E954D198EECDABC69C70F6BD4E6AA" originalValue="A–E">A-E</normalizedToken>
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Shell of paratype, RMNH 104427
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<normalizedToken id="699D74B1E0F63E7D07ACFFEB0758A52F" originalValue="F–H">F-H</normalizedToken>
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Close up of shell of paratype, RMNH 104427. Scale bars:
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<normalizedToken id="483F79759199764B3F9354844AB3E777" originalValue="A–G">A-G</normalizedToken>
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1 mm; H 0.5 mm. All photographs by Thor Seng Liew.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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||
</document> |