175 lines
18 KiB
XML
175 lines
18 KiB
XML
<document id="EADC2EB3B213948A668CB1687B6EF062" ID-CLB-Dataset="6214" ID-DOI="10.11646/zootaxa.4974.3.2" ID-GBIF-Dataset="5b4eaeb3-5722-4021-84ad-125ccd80855f" ID-ISSN="1175-5326" ID-Zenodo-Dep="4777839" ID-ZooBank="1BD6220B-7540-4A12-B3D2-5EDC1A3B7DD8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1621588618725" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2021" docId="4772526A0045FFDFFF182B14116A30E5" docLanguage="en" docName="zootaxa.4974.3.2.pdf" docOrigin="Zootaxa 4974 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hydrochus directus Perkins" docType="treatment" docVersion="7" lastPageNumber="474" masterDocId="BB4B2A12004AFFCFFF8F2D18151D3030" masterDocTitle="Taxonomy of Central American water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)" masterLastPageNumber="503" masterPageNumber="459" pageNumber="474" updateTime="1698940277359" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="3FD58D1FF8C5EEC29997F37DF8980D40">Taxonomy of Central American water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)</mods:title>
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<mods:namePart id="BF64DB19D540358CDFA21DC9C7E97598">Perkins, Philip D.</mods:namePart>
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<mods:title id="CC5602F8D1FD60E150AE0FF84228625D">Zootaxa</mods:title>
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<mods:date id="A0EABC196FBD6D4626864C52B86E13C3">2021</mods:date>
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<treatment id="4772526A0045FFDFFF182B14116A30E5" ID-DOI="http://doi.org/10.5281/zenodo.4777918" ID-GBIF-Taxon="182642567" ID-Zenodo-Dep="4777918" LSID="urn:lsid:plazi:treatment:4772526A0045FFDFFF182B14116A30E5" httpUri="http://treatment.plazi.org/id/4772526A0045FFDFFF182B14116A30E5" lastPageId="16" lastPageNumber="474" pageId="15" pageNumber="474">
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<subSubSection id="87C1B0F70045FFC0FF182B1414F13617" box="[151,492,1548,1575]" pageId="15" pageNumber="474" type="nomenclature">
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<paragraph id="CF64E37C0045FFC0FF182B1414F13617" blockId="15.[151,492,1548,1611]" box="[151,492,1548,1575]" pageId="15" pageNumber="474">
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<heading id="942C54100045FFC0FF182B1414F13617" bold="true" box="[151,492,1548,1575]" fontSize="11" level="1" pageId="15" pageNumber="474" reason="1">
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<taxonomicName id="08DB98FF0045FFC0FF182B1414F13617" ID-CoL="8Y4PX" authority="Perkins" authorityName="Perkins" box="[151,492,1548,1575]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="474" phylum="Arthropoda" rank="species" species="directus">
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<emphasis id="FDAF3F6E0045FFC0FF182B1414F13617" bold="true" box="[151,492,1548,1575]" pageId="15" pageNumber="474">
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<emphasis id="FDAF3F6E0045FFC0FF182B1414993617" bold="true" box="[151,388,1548,1575]" italics="true" pageId="15" pageNumber="474">Hydrochus directus</emphasis>
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Perkins
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="87C1B0F70045FFC0FF182B29141E367B" box="[151,259,1585,1611]" pageId="15" pageNumber="474" type="description">
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<paragraph id="CF64E37C0045FFC0FF182B29141E367B" blockId="15.[151,492,1548,1611]" box="[151,259,1585,1611]" pageId="15" pageNumber="474">
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<figureCitation id="57E0FFF90045FFC0FF182B2915C6367B" box="[151,219,1585,1611]" captionStart="FIGURE 2" captionStartId="5.[151,250,1989,2013]" captionTargetBox="[181,1393,181,1953]" captionTargetId="figure-15@5.[158,1429,181,1965]" captionTargetPageId="5" captionText="FIGURE 2. Dorsal habitus of Central American Hydrochus species." figureDoi="http://doi.org/10.5281/zenodo.4777847" httpUri="https://zenodo.org/record/4777847/files/figure.png" pageId="15" pageNumber="474">Fig. 2</figureCitation>
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,
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<figureCitation id="57E0FFF90045FFC0FF682B29141E367B" box="[231,259,1585,1611]" captionStart="FIGURES 31–33" captionStartId="36.[151,264,1943,1967]" captionTargetBox="[170,1417,181,1920]" captionTargetId="figure-22@36.[170,1417,181,1920]" captionTargetPageId="36" captionText="FIGURES 31–33. Geographical distribution maps." figureDoi="http://doi.org/10.5281/zenodo.4777908" httpUri="https://zenodo.org/record/4777908/files/figure.png" pageId="15" pageNumber="474">32</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="87C1B0F70045FFC0FF182B6D102336CA" pageId="15" pageNumber="474" type="reference_group">
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<paragraph id="CF64E37C0045FFC0FF182B6D173036BD" blockId="15.[151,557,1653,1677]" box="[151,557,1653,1677]" pageId="15" pageNumber="474">
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<treatmentCitationGroup id="EFCBC4520045FFC0FF182B6D173036BD" box="[151,557,1653,1677]" pageId="15" pageNumber="474">
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<treatmentCitation id="4E7AC56D0045FFC0FF182B6D173436BD" ID-DOI="http://doi.org/10.5281/zenodo.3883397" ID-Zenodo-Dep="3883397" author="Perkins, P. D." box="[151,553,1653,1677]" httpUri="http://treatment.plazi.org/id/060AF5509B203708FEA51248FAEBFC5D" page="295" pageId="15" pageNumber="474" year="2020">
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<taxonomicName id="08DB98FF0045FFC0FF182B6D173436BD" ID-CoL="8Y4PX" authority="Perkins 2020 a: 295" authorityName="Perkins" authorityPageNumber="295" authorityYear="2020" box="[151,553,1653,1677]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="474" phylum="Arthropoda" rank="species" species="directus">
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<emphasis id="FDAF3F6E0045FFC0FF182B6D144036BD" box="[151,349,1653,1677]" italics="true" pageId="15" pageNumber="474">Hydrochus directus</emphasis>
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<bibRefCitation id="AB4A9E8D0045FFC0FEEC2B6D173436BD" author="Perkins, P. D." box="[355,553,1653,1677]" pageId="15" pageNumber="474" pagination="291 - 301" refId="ref25459" refString="Perkins, P. D. (2020 a) Taxonomy of Guatemalan water beetles in the genus Hydrochus (Coleoptera: Hydrochidae). Acta Entomologica Musei Nationalis Pragae, 60 (1), 291 - 301. https: // doi. org / 10.37520 / aemnp. 2020.016" type="journal article" year="2020">Perkins 2020a: 295</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="CF64E37C0045FFC0FF182BA3102336CA" blockId="15.[151,1437,1723,2038]" pageId="15" pageNumber="474">
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<materialsCitation id="7FB3E9210045FFC0FF182BA3102736CA" ID-GBIF-Occurrence="3128194540" country="Guatemala" county="Images" location="Originally" municipality="Following" pageId="15" pageNumber="474" specimenCount="1" typeStatus="holotype">
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<location id="CA04B5A70045FFC0FF182BA3141736E5" LSID="urn:lsid:plazi:treatment:4772526A0045FFDFFF182B14116A30E5:CA04B5A70045FFC0FF182BA3141736E5" box="[151,266,1723,1749]" country="Guatemala" county="Images" municipality="Following" name="Originally" pageId="15" pageNumber="474">Originally</location>
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described from
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<collectingCountry id="B7CCA3EC0045FFC0FE482BA3175936E5" box="[455,580,1723,1749]" name="Guatemala" pageId="15" pageNumber="474">Guatemala</collectingCountry>
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.
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<collectingMunicipality id="2F0079060045FFC0FDDF2BA317DE36E5" box="[592,707,1723,1749]" pageId="15" pageNumber="474">Following</collectingMunicipality>
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are new locality records for
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<collectingCountry id="B7CCA3EC0045FFC0FB9F2BA3117436E5" box="[1040,1129,1723,1749]" name="Mexico" pageId="15" pageNumber="474">Mexico</collectingCountry>
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.
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<collectingCounty id="26059BF00045FFC0FBFA2BA411DB36E6" box="[1141,1222,1724,1750]" pageId="15" pageNumber="474">Images</collectingCounty>
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||
of the habitus and male genitalia of the
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||
<typeStatus id="10605DDE0045FFC0FE0B2BC714F436C9" box="[388,489,1759,1785]" pageId="15" pageNumber="474" type="holotype">holotype</typeStatus>
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||
, and detailed description of the male genitalia, are given in
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||
<bibRefCitation id="AB4A9E8D0045FFC0FB0A2BC7102736CA" author="Perkins, P. D." box="[1157,1338,1759,1786]" pageId="15" pageNumber="474" pagination="291 - 301" refId="ref25459" refString="Perkins, P. D. (2020 a) Taxonomy of Guatemalan water beetles in the genus Hydrochus (Coleoptera: Hydrochidae). Acta Entomologica Musei Nationalis Pragae, 60 (1), 291 - 301. https: // doi. org / 10.37520 / aemnp. 2020.016" type="journal article" year="2020">Perkins (2020a)</bibRefCitation>
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</materialsCitation>
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.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="87C1B0F70045FFC0FF482A1B17C23756" pageId="15" pageNumber="474" type="materials_examined">
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||
<paragraph id="CF64E37C0045FFC0FF482A1B17C23756" blockId="15.[151,1437,1723,2038]" pageId="15" pageNumber="474">
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||
<emphasis id="FDAF3F6E0045FFC0FF482A1B14CD372D" bold="true" box="[199,464,1795,1821]" pageId="15" pageNumber="474">Material examined (3)</emphasis>
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||
:
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||
<materialsCitation id="7FB3E9210045FFC0FE512A1B102B372E" ID-GBIF-Occurrence="3128194556" box="[478,1334,1795,1822]" collectingDate="1962-06-28" collectionCode="CNC" collectorName="D. H. Janzen" country="Mexico" location="Cotaxtla" pageId="15" pageNumber="474" specimenCount="1">
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||
<emphasis id="FDAF3F6E0045FFC0FE512A1B1728372D" bold="true" box="[478,565,1795,1821]" pageId="15" pageNumber="474">
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||
<collectingCountry id="B7CCA3EC0045FFC0FE512A1B1728372D" box="[478,565,1795,1821]" name="Mexico" pageId="15" pageNumber="474">Mexico</collectingCountry>
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||
</emphasis>
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||
:
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||
<location id="CA04B5A70045FFC0FDCC2A1B17B8372D" LSID="urn:lsid:plazi:treatment:4772526A0045FFDFFF182B14116A30E5:CA04B5A70045FFC0FDCC2A1B17B8372D" box="[579,677,1795,1821]" country="Mexico" name="Cotaxtla" pageId="15" pageNumber="474">Cotaxtla</location>
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||
, Cotaxtla Exp. Sta.,
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||
<date id="BB65C5BC0045FFC0FC072A1B111F372D" box="[904,1026,1795,1821]" pageId="15" pageNumber="474" value="1962-06-28">
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||
<collectingDate id="AB213C540045FFC0FC072A1B111F372D" box="[904,1026,1795,1821]" pageId="15" pageNumber="474" value="1962-06-28">28.vi.1962</collectingDate>
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</date>
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, leg.
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<collectorName id="622E86AA0045FFC0FBB22A1C11D1372E" box="[1085,1228,1796,1822]" pageId="15" pageNumber="474">D. H. Janzen</collectorName>
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||
(1
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||
<collectionCode id="A9CA7BB90045FFC0FB7F2A1C1033372E" box="[1264,1326,1796,1822]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="15" pageNumber="474">CNC</collectionCode>
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||
)
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</materialsCitation>
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||
;
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<materialsCitation id="7FB3E9210045FFC0FACE2A1C11603772" ID-GBIF-Occurrence="3128194597" collectingDate="1981-05-24" collectionCode="NMNH" collectorName="Ao. Chopan & Paul J. Spangler" country="Mexico" location="Bethania" municipality="Tuxtepec" pageId="15" pageNumber="474" specimenCount="1" stateProvince="Oaxaca">
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||
<collectingRegion id="0D1F2D9E0045FFC0FACE2A1C1085372E" box="[1345,1432,1796,1822]" country="Mexico" name="Oaxaca" pageId="15" pageNumber="474">Oaxaca</collectingRegion>
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,
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<collectingMunicipality id="2F0079060045FFC0FF182A3015E33772" box="[151,254,1832,1858]" pageId="15" pageNumber="474">Tuxtepec</collectingMunicipality>
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||
(
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||
<quantity id="08234E990045FFC0FE802A3F144D3771" box="[271,336,1831,1857]" metricMagnitude="4" metricUnit="m" metricValue="3.1" pageId="15" pageNumber="474" unit="km" value="31.0">31km</quantity>
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||
S),
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||
<location id="CA04B5A70045FFC0FE0E2A3F14FA3771" LSID="urn:lsid:plazi:treatment:4772526A0045FFDFFF182B14116A30E5:CA04B5A70045FFC0FE0E2A3F14FA3771" box="[385,487,1831,1857]" country="Mexico" municipality="Tuxtepec" name="Bethania" pageId="15" pageNumber="474" stateProvince="Oaxaca">Bethania</location>
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||
,
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||
<collectorName id="622E86AA0045FFC0FE7D2A3017633771" box="[498,638,1831,1858]" pageId="15" pageNumber="474">Ao. Chopan</collectorName>
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||
,
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||
<date id="BB65C5BC0045FFC0FD052A3017E63772" box="[650,763,1832,1858]" pageId="15" pageNumber="474" value="1981-05-24">
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||
<collectingDate id="AB213C540045FFC0FD052A3017E63772" box="[650,763,1832,1858]" pageId="15" pageNumber="474" value="1981-05-24">24.v.1981</collectingDate>
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||
</date>
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||
, leg.
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||
<collectorName id="622E86AA0045FFC0FCB52A3F16EF3771" box="[826,1010,1831,1858]" pageId="15" pageNumber="474">Paul J. Spangler</collectorName>
|
||
(1
|
||
<collectionCode id="A9CA7BB90045FFC0FB952A3011683772" box="[1050,1141,1832,1858]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="15" pageNumber="474" type="Museum">NMNH</collectionCode>
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||
)
|
||
</materialsCitation>
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||
;
|
||
<materialsCitation id="7FB3E9210045FFC0FB062A3017C63756" ID-GBIF-Occurrence="3128194457" collectingDate="1965-08-27" collectionCode="NMNH" collectorName="W. Poza Rica & Paul J. Spangler" country="Mexico" location="Veracruz" pageId="15" pageNumber="474" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave">
|
||
<collectingRegion id="0D1F2D9E0045FFC0FB062A3011ED3772" box="[1161,1264,1832,1858]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="15" pageNumber="474">Veracruz</collectingRegion>
|
||
,
|
||
<quantity id="08234E990045FFC0FB732A30102A3771" box="[1276,1335,1831,1858]" metricMagnitude="4" metricUnit="m" metricValue="1.4484096" pageId="15" pageNumber="474" unit="mi" value="9.0">9 mi.</quantity>
|
||
<collectorName id="622E86AA0045FFC0FAB12A3015D03755" pageId="15" pageNumber="474">W. Poza Rica</collectorName>
|
||
,
|
||
<date id="BB65C5BC0045FFC0FF562A53147F3756" box="[217,354,1867,1894]" pageId="15" pageNumber="474" value="1965-08-27">
|
||
<collectingDate id="AB213C540045FFC0FF562A53147F3756" box="[217,354,1867,1894]" pageId="15" pageNumber="474" value="1965-08-27">27.viii.1965</collectingDate>
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||
</date>
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||
, leg.
|
||
<collectorName id="622E86AA0045FFC0FE112A5317493755" box="[414,596,1867,1894]" pageId="15" pageNumber="474">Paul J. Spangler</collectorName>
|
||
(1
|
||
<collectionCode id="A9CA7BB90045FFC0FDF62A5417C93756" box="[633,724,1868,1894]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="15" pageNumber="474" type="Museum">NMNH</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="87C1B0F70045FFDFFF482A77116A30E5" lastPageId="16" lastPageNumber="475" pageId="15" pageNumber="474" type="diagnosis">
|
||
<paragraph id="CF64E37C0045FFDFFF482A77116A30E5" blockId="15.[151,1437,1723,2038]" lastBlockId="16.[151,1436,151,213]" lastPageId="16" lastPageNumber="475" pageId="15" pageNumber="474">
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||
<emphasis id="FDAF3F6E0045FFC0FF482A7714D037B9" bold="true" box="[199,461,1903,1929]" pageId="15" pageNumber="474">Differential diagnosis.</emphasis>
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Among Central American species, recognized by the combination of moderate size (ca. 3.00 mm), the densely punctate pronotum with the reliefs slightly less punctate than the depressions, and the male genitalia. This species has a generalized
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<taxonomicName id="08DB98FF0045FFC0FDD92AAF17CD37E1" authorityName="Leach" authorityYear="1817" box="[598,720,1975,2001]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="474" phylum="Arthropoda" rank="genus">
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||
<emphasis id="FDAF3F6E0045FFC0FDD92AAF17CD37E1" box="[598,720,1975,2001]" italics="true" pageId="15" pageNumber="474">Hydrochus</emphasis>
|
||
</taxonomicName>
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||
habitus; reliable determinations will require dissection of males. The male genitalia of
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<taxonomicName id="08DB98FF0045FFC0FE042AC4171B37C5" box="[395,518,2011,2037]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="474" phylum="Arthropoda" rank="species" species="directus">
|
||
<emphasis id="FDAF3F6E0045FFC0FE042AC4171B37C5" box="[395,518,2011,2037]" italics="true" pageId="15" pageNumber="474">H. directus</emphasis>
|
||
</taxonomicName>
|
||
is very slender, therefore superficially somewhat similar to that of
|
||
<taxonomicName id="08DB98FF0045FFC0FB7E2AC4108137C5" authority="Perkins, 2021" authorityName="Perkins" authorityYear="2021" box="[1265,1436,2012,2037]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="474" phylum="Arthropoda" rank="species" species="aeruginosus" status="sp. nov.">
|
||
<emphasis id="FDAF3F6E0045FFC0FB7E2AC4108137C5" box="[1265,1436,2012,2037]" italics="true" pageId="15" pageNumber="474">H. aeruginosus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="FDAF3F6E005AFFDFFF182D8F15CB3081" bold="true" box="[151,214,151,177]" pageId="16" pageNumber="475">
|
||
<taxonomicNameLabel id="E69C8215005AFFDFFF182D8F15CB3081" box="[151,214,151,177]" pageId="16" pageNumber="475" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; however, details of the two distinctly differ, including shape and size of the parameres, size and shape of the basal piece, and form of the adbl. The habitus of the two species also distinctively differ.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |