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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1184.113255" ID-Pensoft-Pub="1313-2970-1184-291" ID-Pensoft-UUID="A813DA4B651652CC83F81AF764B147C4" ID-ZooBank="1E7795B9D93A41599BB0027625B1B768" ModsDocID="1313-2970-1184-291" checkinTime="1700659040281" checkinUser="pensoft" docAuthor="Tanasevitch, Andrei &amp; Marusik, Yuri" docDate="2023" docId="4622DA4EF9975DA1A2FB563027541B8C" docLanguage="en" docName="ZooKeys 1184: 291-299" docOrigin="ZooKeys 1184" docPubDate="2023-11-21" docSource="http://dx.doi.org/10.3897/zookeys.1184.113255" docTitle="Dactylopisthoides Eskov 1990" docType="treatment" docVersion="1" id="A813DA4B651652CC83F81AF764B147C4" lastPageNumber="291" masterDocId="A813DA4B651652CC83F81AF764B147C4" masterDocTitle="On the synonymy of Dactylopisthoides Eskov, 1990 and Uusitaloia Marusik, Koponen &amp; Danilov, 2001 (Araneae, Linyphiidae)" masterLastPageNumber="299" masterPageNumber="291" pageNumber="291" updateTime="1700659040281" updateUser="pensoft">
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<mods:title id="F225125503A6159216F12F31904EB4B5">On the synonymy of Dactylopisthoides Eskov, 1990 and Uusitaloia Marusik, Koponen &amp; Danilov, 2001 (Araneae, Linyphiidae)</mods:title>
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<mods:namePart id="95DB27BCB73BAAF2EDD4A4A6B38433B0">Tanasevitch, Andrei</mods:namePart>
<mods:nameIdentifier id="7BFF0108134A524EB3233E49018F7EF2" type="ORCID">https://orcid.org/0000-0002-9116-606X</mods:nameIdentifier>
<mods:affiliation id="5C5E1D12CDAC685DB6EA6395D1F5D838">A. N. Severtsov Institute of Ecology and Evolution, Russian Academy of Sciences, Leninsky prospekt 33, Moscow 119071, Russia</mods:affiliation>
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<mods:namePart id="9ED34F2A4C1DD7287F95D91E19BD9E7D">Marusik, Yuri</mods:namePart>
<mods:nameIdentifier id="91B5FE086F989536462478BED0B8E54A" type="ORCID">https://orcid.org/0000-0002-4499-5148</mods:nameIdentifier>
<mods:affiliation id="EF9306952869EB123597949E4B49EB65">Institute for Biological Problems of the North FEB RAS, Portovaya Str. 18, Magadan 685000, Russia &amp; Altai State University, Lenina Pr., 61, Barnaul, RF- 656049, Russia &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa</mods:affiliation>
<mods:nameIdentifier id="448CF8F691A4E886FF51283AEFE55A99" type="email">yurmar@utu.fi</mods:nameIdentifier>
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<mods:typeOfResource id="A1BE3EAC6EBFE8E42C93DC47F215B847">text</mods:typeOfResource>
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<mods:date id="C7772E93DB017110C9FD906CEE9636BC">2023</mods:date>
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<mods:number id="405CC7FD7BADE3C35547543B06061097">2023-11-21</mods:number>
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<mods:detail id="9F54C8F23EF6278E7302B9EEED074D38" type="volume">
<mods:number id="7D29EA2A168AB68BC3109FE7557B2793">1184</mods:number>
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<treatment id="4622DA4EF9975DA1A2FB563027541B8C" LSID="urn:lsid:plazi:treatment:4622DA4EF9975DA1A2FB563027541B8C" httpUri="http://treatment.plazi.org/id/4622DA4EF9975DA1A2FB563027541B8C" lastPageNumber="291" pageId="0" pageNumber="291">
<subSubSection id="BA18D434CC5957A968C47FC56A9EA25C" pageId="0" pageNumber="291" type="nomenclature">
<paragraph id="5C975D19D812809EA3EEECF8214B5153" pageId="0" pageNumber="291">
<taxonomicName id="B9D22C420E1D8DF52EFF262B128CD5D4" ID-CoL="3ZCS" LSID="4622DA4E-F997-5DA1-A2FB-563027541B8C" authority="Eskov, 1990" authorityName="Eskov" authorityYear="1990" class="Arachnida" family="Linyphiidae" genus="Dactylopisthoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dactylopisthoides" order="Araneae" pageId="0" pageNumber="291" phylum="Arthropoda" rank="genus">Dactylopisthoides Eskov, 1990</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="294420441744E2F14104EB1643B3BED3" pageId="0" pageNumber="291" type="reference_group">
<paragraph id="A9E883D1400872D1ED526B2B8BFDB8A9" pageId="0" pageNumber="291">
<taxonomicName id="926998A8F109D916C1CF99BE19132547" authorityName="Eskov" authorityYear="1990" class="Arachnida" family="Linyphiidae" genus="Dactylopisthoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dactylopisthoides" order="Araneae" pageId="0" pageNumber="291" phylum="Arthropoda" rank="genus">Dactylopisthoides</taxonomicName>
Eskov, 1990: 4 (type species:
<taxonomicName id="A31316C6F62D40B5D644029AE343849B" authorityName="Eskov" authorityYear="1990" class="Arachnida" family="Linyphiidae" genus="Dactylopisthoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dactylopisthoides hyperboreus" order="Araneae" pageId="0" pageNumber="291" phylum="Arthropoda" rank="species" species="hyperboreus">Dactylopisthoides hyperboreus</taxonomicName>
Eskov, 1990).
</paragraph>
<paragraph id="15760F8BCD2DEFBA74195A7F3A990E9C" pageId="0" pageNumber="291">
<taxonomicName id="F49A45A158DD782F52AFA020CFAF76F2" authorityName="Marusik, Koponen &amp; Danilov" authorityYear="2001" class="Arachnida" family="Linyphiidae" genus="Uusitaloia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uusitaloia" order="Araneae" pageId="0" pageNumber="291" phylum="Arthropoda" rank="genus">Uusitaloia</taxonomicName>
Marusik, Koponen &amp; Danilov, 2001: 89 (type species:
<taxonomicName id="1C70FF782E1ECE3BA90C59860B380EA1" authorityName="Tanasevitch &amp; Marusik" authorityYear="2023" baseAuthorityName="Tanasevitch &amp; Marusik" baseAuthorityYear="2023" class="Arachnida" family="Linyphiidae" genus="Uusitaloia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uusitaloia transbaicalica" order="Araneae" pageId="0" pageNumber="291" phylum="Arthropoda" rank="species" species="transbaicalica">Uusitaloia transbaicalica</taxonomicName>
Marusik, Koponen &amp; Danilov, 2001), syn. nov.
</paragraph>
</subSubSection>
<subSubSection id="4ACE722F03535C955DE89BF836D066AF" pageId="0" pageNumber="291" type="diagnosis">
<paragraph id="FF6541A47FD08398D993C5F28738CF9E" pageId="0" pageNumber="291">Diagnosis.</paragraph>
<paragraph id="A2779DFD92537A720D9E5EA1391992C5" pageId="0" pageNumber="291">
The genus
<taxonomicName id="D6CBA03A33D15E2E535580948726CBBF" authorityName="Eskov" authorityYear="1990" class="Arachnida" family="Linyphiidae" genus="Dactylopisthoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dactylopisthoides" order="Opiliones" pageId="0" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="5BF1D4977C5306144DBA72C43CB25CA4" italics="true" pageId="0" pageNumber="291">Dactylopisthoides</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="C7D6BD13F14C2AAAA58D934FB8790B8D" authorityName="Simon" authorityYear="1884" class="Arachnida" family="Linyphiidae" genus="Dactylopisthes" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dactylopisthes" order="Araneae" pageId="0" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="097667FFEA23A2D0A13271BE01E2DF3C" italics="true" pageId="0" pageNumber="291">Dactylopisthes</emphasis>
</taxonomicName>
Simon, 1884, with the type species
<taxonomicName id="E07E04BD135CA2A048C1B7E87314A410" lsidName="D. digiticeps" pageId="0" pageNumber="291" rank="species" species="digiticeps">
<emphasis id="2C1F40EEB0A5791A75AF467E8D1589DF" italics="true" pageId="0" pageNumber="291">D. digiticeps</emphasis>
</taxonomicName>
(Simon, 1881). Both genera belong to the
<taxonomicName id="BC95C3F6025CE8E6E0FC2921C91B79C4" class="Arachnida" family="Linyphiidae" genus="Savignia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Savignia" order="Araneae" pageId="0" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="D01C52E8C91CA415274C3F1E8413BCBF" italics="true" pageId="0" pageNumber="291">Savignia</emphasis>
</taxonomicName>
-genus group sensu
<bibRefCitation id="0E947C1A2A2222B3BAFD11393E7F7D8C" author="Millidge, AF" journalOrPublisher="Bulletin - British Arachnological Society" pageId="0" pageNumber="291" pagination="1 - 60" refId="B9" refString="Millidge, AF, 1977. The conformation of the male palpal organs of linyphiid spiders, and its application to the taxonomic and phylogenetic analysis of the family (Araneae: Linyphiidae). Bulletin - British Arachnological Society 4: 1 - 60" title="The conformation of the male palpal organs of linyphiid spiders, and its application to the taxonomic and phylogenetic analysis of the family (Araneae: Linyphiidae)." volume="4" year="1977">Millidge (1977)</bibRefCitation>
. The main diagnostic characters are as follows:
</paragraph>
<paragraph id="0B313AEBC363135BED4955C8022E8658" pageId="0" pageNumber="291">
Formula of leg chaetotaxy:
<taxonomicName id="1AA7F1F98F0D9C252DA790500C46F380" authorityName="Eskov" authorityYear="1990" class="Arachnida" family="Linyphiidae" genus="Dactylopisthoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dactylopisthoides" order="Opiliones" pageId="0" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="D94D6A9DF99CBBA8B6C76BEBC2282B66" italics="true" pageId="0" pageNumber="291">Dactylopisthoides</emphasis>
</taxonomicName>
- 2.2.2.1, vs 2.2.1.1 in
<taxonomicName id="2FE6985C5A9D9AF3AE79D86A4C45003A" authorityName="Simon" authorityYear="1884" class="Arachnida" family="Linyphiidae" genus="Dactylopisthes" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dactylopisthes" order="Araneae" pageId="0" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="641B6F1A78021EEC78A5E53507AA5152" italics="true" pageId="0" pageNumber="291">Dactylopisthes</emphasis>
</taxonomicName>
;
</paragraph>
<paragraph id="79A54650D2B8F31E54C0F737FBF8E2ED" pageId="0" pageNumber="291">
Male carapace in
<taxonomicName id="C770720AD7DBF0EB34231073A46A2261" authorityName="Eskov" authorityYear="1990" class="Arachnida" family="Linyphiidae" genus="Dactylopisthoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dactylopisthoides" order="Opiliones" pageId="0" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="E70326B05BEBBA9C242C32D6CA5B83D2" italics="true" pageId="0" pageNumber="291">Dactylopisthoides</emphasis>
</taxonomicName>
is unmodified, vs highly modified except for three Oriental species of which the generic assignment is doubtful;
</paragraph>
<paragraph id="5D70EE5DC6B97ED5F25DA363F6D48407" pageId="0" pageNumber="291">
Male palpal tibia in
<taxonomicName id="BDEF112687C18ECF74CBACB2668BA289" authorityName="Eskov" authorityYear="1990" class="Arachnida" family="Linyphiidae" genus="Dactylopisthoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dactylopisthoides" order="Opiliones" pageId="0" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="F4697C372F28C3E78F7C3AB15D769FE4" italics="true" pageId="0" pageNumber="291">Dactylopisthoides</emphasis>
</taxonomicName>
is only slightly elongated dorsally, vs highly modified, sickle-shaped in
<taxonomicName id="28FE34C7EC8DFCB4E3067036AA64E0E3" authorityName="Simon" authorityYear="1884" class="Arachnida" family="Linyphiidae" genus="Dactylopisthes" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dactylopisthes" order="Araneae" pageId="0" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="14BEC68BA3019265013419FD6BB2E77D" italics="true" pageId="0" pageNumber="291">Dactylopisthes</emphasis>
</taxonomicName>
, except for
<taxonomicName id="40CD3649E6939BA0D5D007687551BD18" lsidName="D. mirabilis" pageId="0" pageNumber="291" rank="species" species="mirabilis">
<emphasis id="7AAC24FB783AE7482659989E8EDDEA75" italics="true" pageId="0" pageNumber="291">D. mirabilis</emphasis>
</taxonomicName>
(Tanasevitch, 1985), known from Kyrgyzstan (
<bibRefCitation id="DA3A2C85CAFA38F87C92B6068EB20C2B" author="Tanasevitch, AV" journalOrPublisher="Entomologicheskoe Obozrenie" pageId="0" pageNumber="291" pagination="845 - 854" refId="B12" refString="Tanasevitch, AV, 1985. New species of spiders of the family Linyphiidae (Aranei) from Kirghizia. Entomologicheskoe Obozrenie 64: 845 - 854" title="New species of spiders of the family Linyphiidae (Aranei) from Kirghizia." volume="64" year="1985">Tanasevitch 1985</bibRefCitation>
);
</paragraph>
<paragraph id="A206B63FE41CD0ECBC36C37B1C00FB32" pageId="0" pageNumber="291">
Epigyne in
<taxonomicName id="5B5494DCAC3DFD25580DC6A87EB1F8B0" authorityName="Eskov" authorityYear="1990" class="Arachnida" family="Linyphiidae" genus="Dactylopisthoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dactylopisthoides" order="Opiliones" pageId="0" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="C5CD66709C421652C7AEBF3F5240792D" italics="true" pageId="0" pageNumber="291">Dactylopisthoides</emphasis>
</taxonomicName>
with an opened fovea, its lateral walls with a distinct posterior projection on each (Figs
<figureCitation id="98982148D8672BD1A0AF78162127B7E2" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Details of the male palp (A-E) and epigyne (F) of Dactylopisthoides hyperboreus Eskov, 1990, specimens from the type locality A-C right palp, retrolateral (A, B) and dorsal view (C), respectively D, E bulb, lateral view, different aspects F epigyne, ventral view. Abbreviations: dsa - distal suprategular apophysis, ep - embolus proper, mm - median membrane, mt - median tooth, pp - posterior projections of lateral walls, r - radix, rt - radical tooth." figureDoi="10.3897/zookeys.1184.113255.figure1" httpUri="https://binary.pensoft.net/fig/940454" pageId="0" pageNumber="291">1F</figureCitation>
,
<figureCitation id="1962C57AA573382FF7DA426EDC61447D" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Details of the male palp (A-E) and female epigyne (F) of Dactylopisthoides wrangelianus (Marusik &amp; Koponen, 2009), specimens from Mamontovaya River, Wrangel Isl A-D right palp, retrolateral, dorsal, ventral and frontal view, respectively E embolic division, frontal view F epigyne, ventral view. Abbreviations: dsa - distal suprategular apophysis, ep - embolus proper, mm - median membrane, mt - median tooth, pp - posterior projections of lateral walls, r - radix, rt - radical tooth." figureDoi="10.3897/zookeys.1184.113255.figure2" httpUri="https://binary.pensoft.net/fig/940455" pageId="0" pageNumber="291">2F</figureCitation>
), vs fovea absent.
</paragraph>
</subSubSection>
<subSubSection id="7C67BB71A47FE5F926797539568D7283" pageId="0" pageNumber="291" type="description">
<paragraph id="1A26B2BB4A861F64C6D9DAE66C96DC82" pageId="0" pageNumber="291">Description.</paragraph>
<paragraph id="9E210A6D21CA708341839CC2FBDEDF25" pageId="0" pageNumber="291">Small to medium erigonine. Male total length 1.50-1.80, carapace 0.63-0.79 long. Female total length 1.53-1.75, carapace 0.6-0.68 long. Carapace unmodified in both sexes. Formula of leg chaetotaxy 2.2.2.1, tibial spines short and weak. Metatarsi I-III each with trichobothrium. TmI 0.50-0.70. Male palpal tibia slightly modified, somewhat elongated dorsally. Paracymbium small and narrow, L-shaped, uncinate apically. Distal suprategular apophysis greatly enlarged, highly sclerotized, median tooth present. Embolic division relatively large, radix curved, with small outgrowth (= radical tooth); embolus proper short, thick, covered with short median membrane. Epigyne with cavity, lateral walls with distinct posterior projection in each.</paragraph>
</subSubSection>
<subSubSection id="E99E64E26CF1BC3C7560FB51A75AC29B" pageId="0" pageNumber="291" type="comments">
<paragraph id="C060987137755E1DA9A8621C4D7A8DF9" pageId="0" pageNumber="291">Comments.</paragraph>
<paragraph id="D2692E858066C8162CC5662378E93A44" pageId="0" pageNumber="291">
An analysis of both somatic and copulatory organ structures of the type species of
<taxonomicName id="017899DC01F533592BE97E3F72E11DAD" authorityName="Marusik, Koponen &amp; Danilov" authorityYear="2001" class="Arachnida" family="Linyphiidae" genus="Uusitaloia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uusitaloia" order="Araneae" pageId="0" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="384897DB18A0C26A69BE4747867D4609" italics="true" pageId="0" pageNumber="291">Uusitaloia</emphasis>
</taxonomicName>
and
<taxonomicName id="E46962E308B245ACB4F3D146DD2F9AD4" authorityName="Eskov" authorityYear="1990" class="Arachnida" family="Linyphiidae" genus="Dactylopisthoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dactylopisthoides" order="Opiliones" pageId="0" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="CC9ABBCE973F81C9970270A9C11A0901" italics="true" pageId="0" pageNumber="291">Dactylopisthoides</emphasis>
</taxonomicName>
has revealed that they are undoubtedly congeneric: all the species in both genera are of a similar small size, possess the same chaetotaxy (2221) and trichobothriotaxy (I-III; TmI: 0.64-0.65), and are characterized by the same conformation of copulatory organs in both sexes, differing from each other only in minor details: cf. Figs
<figureCitation id="C1D64C73815791D09132A821417F4951" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Details of the male palp (A-E) and epigyne (F) of Dactylopisthoides hyperboreus Eskov, 1990, specimens from the type locality A-C right palp, retrolateral (A, B) and dorsal view (C), respectively D, E bulb, lateral view, different aspects F epigyne, ventral view. Abbreviations: dsa - distal suprategular apophysis, ep - embolus proper, mm - median membrane, mt - median tooth, pp - posterior projections of lateral walls, r - radix, rt - radical tooth." figureDoi="10.3897/zookeys.1184.113255.figure1" httpUri="https://binary.pensoft.net/fig/940454" pageId="0" pageNumber="291">1</figureCitation>
-
<figureCitation id="127426AC4EAEA6F97B99823EB4E9DD69" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of the embolic divisions of Dactylopisthoides hyperboreus Eskov, 1990 (A), D. wrangelianus (Marusik &amp; Koponen, 2009) (B) and D. transbaicalicus (Marusik, Koponen &amp; Danilov, 2001) [reproduced from Marusik and Koponen (2009)] (C). Abbreviations: dsa - distal suprategular apophysis, mm - median membrane, mt - median tooth, rt - radical tooth. Black, thick arrows show additional differences between species. Scale bar: 0.1 mm (S)." figureDoi="10.3897/zookeys.1184.113255.figure3" httpUri="https://binary.pensoft.net/fig/940456" pageId="0" pageNumber="291">3</figureCitation>
. Thus, the genus
<taxonomicName id="6536BE1E6DF2C2E0CB081317CAD971AD" authorityName="Marusik, Koponen &amp; Danilov" authorityYear="2001" class="Arachnida" family="Linyphiidae" genus="Uusitaloia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uusitaloia" order="Araneae" pageId="0" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="7C2EDCEFCBAF4DBA5BFD52D6ED82B232" italics="true" pageId="0" pageNumber="291">Uusitaloia</emphasis>
</taxonomicName>
Marusik, Koponen &amp; Danilov, 2001 is to be considered a junior synonym of
<taxonomicName id="9337B97D849D98D88AC0721A36FEB15D" authorityName="Eskov" authorityYear="1990" class="Arachnida" family="Linyphiidae" genus="Dactylopisthoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dactylopisthoides" order="Opiliones" pageId="0" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="E9E4FF5396B3B1D87A54D9D4F254CA79" italics="true" pageId="0" pageNumber="291">Dactylopisthoides</emphasis>
</taxonomicName>
Eskov, 1990.
</paragraph>
<caption id="416ACBD1D7DFE59DED3FC0E99EE21563" doi="10.3897/zookeys.1184.113255.figure1" httpUri="https://binary.pensoft.net/fig/940454" pageId="0" pageNumber="291" start="Figure 1" startId="F1">
<paragraph id="5DF5B22561B0DE5C6EC5803760EFC694" pageId="0" pageNumber="291">
<emphasis id="4DB7DD1DFC4818A3BEEC07E03CAF9547" bold="true" pageId="0" pageNumber="291">Figure 1.</emphasis>
Details of the male palp (
<emphasis id="D126811319F9AC3C1D0AAAA538236C1E" bold="true" pageId="0" pageNumber="291">A-E</emphasis>
) and epigyne (
<emphasis id="F94A35CAB89CC311CD58202C7E792B5A" bold="true" pageId="0" pageNumber="291">F</emphasis>
) of
<taxonomicName id="301A51651EBAC7A5AFC0193527BF96C5" authorityName="Eskov" authorityYear="1990" class="Arachnida" family="Linyphiidae" genus="Dactylopisthoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dactylopisthoides hyperboreus" order="Opiliones" pageId="0" pageNumber="291" phylum="Arthropoda" rank="species" species="hyperboreus">
<emphasis id="F5809EC43C8C64C9E22F93E0D1FB2679" italics="true" pageId="0" pageNumber="291">Dactylopisthoides hyperboreus</emphasis>
</taxonomicName>
Eskov, 1990, specimens from the type locality
<emphasis id="6EF893197B093475FD3BFF5790353715" bold="true" pageId="0" pageNumber="291">A-C</emphasis>
right palp, retrolateral (
<emphasis id="67A7BDEF19A50BD43EC9A6041B03EBB6" bold="true" pageId="0" pageNumber="291">A, B</emphasis>
) and dorsal view (
<emphasis id="1C3822E18F61055ED00A9803BFE8D14A" bold="true" pageId="0" pageNumber="291">C</emphasis>
), respectively
<emphasis id="AFAA3AF77CE52CF97B0864BF12AFB2D5" bold="true" pageId="0" pageNumber="291">D, E</emphasis>
bulb, lateral view, different aspects
<emphasis id="1A5FACC0A0D1AC9BC448F284D7A93C87" bold="true" pageId="0" pageNumber="291">F</emphasis>
epigyne, ventral view. Abbreviations: dsa - distal suprategular apophysis, ep - embolus proper, mm - median membrane, mt - median tooth, pp - posterior projections of lateral walls, r - radix, rt - radical tooth.
</paragraph>
</caption>
<caption id="7762A32D6BC85F8B123886BE38A78880" doi="10.3897/zookeys.1184.113255.figure2" httpUri="https://binary.pensoft.net/fig/940455" pageId="0" pageNumber="291" start="Figure 2" startId="F2">
<paragraph id="0C1EE6509BDB2623F1A8CCADB78F6531" pageId="0" pageNumber="291">
<emphasis id="A6DF2A6AAE024934D4154A4526E9AC81" bold="true" pageId="0" pageNumber="291">Figure 2.</emphasis>
Details of the male palp (
<emphasis id="38609E6D5EBAD55DD385C6903D428FFC" bold="true" pageId="0" pageNumber="291">A-E</emphasis>
) and female epigyne (
<emphasis id="DA5FAD18B5C02F0974D1C2A052476AAD" bold="true" pageId="0" pageNumber="291">F</emphasis>
) of
<taxonomicName id="F121F3F1853ECC0E71080DE3D5D31BB9" authorityName="Tanasevitch &amp; Marusik" authorityYear="2023" baseAuthorityName="Marusik &amp; Koponen" baseAuthorityYear="2009" class="Arachnida" family="Linyphiidae" genus="Dactylopisthoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dactylopisthoides wrangelianus" order="Opiliones" pageId="0" pageNumber="291" phylum="Arthropoda" rank="species" species="wrangelianus">
<emphasis id="33FB4C47F6929DEC48A882F182E2BB6A" italics="true" pageId="0" pageNumber="291">Dactylopisthoides wrangelianus</emphasis>
</taxonomicName>
(Marusik &amp; Koponen, 2009), specimens from Mamontovaya River, Wrangel Isl
<emphasis id="0540DDB0E1AC4D3EAC1C91677CECC847" bold="true" pageId="0" pageNumber="291">A-D</emphasis>
right palp, retrolateral, dorsal, ventral and frontal view, respectively
<emphasis id="B49B90127D5BB6398C6DAAB0FCAB52E2" bold="true" pageId="0" pageNumber="291">E</emphasis>
embolic division, frontal view
<emphasis id="44B623BDE73974D744CBE6B74FA89373" bold="true" pageId="0" pageNumber="291">F</emphasis>
epigyne, ventral view. Abbreviations: dsa - distal suprategular apophysis, ep - embolus proper, mm - median membrane, mt - median tooth, pp - posterior projections of lateral walls, r - radix, rt - radical tooth.
</paragraph>
</caption>
<caption id="28E1C5D79D3DA883324A531D679A97DF" doi="10.3897/zookeys.1184.113255.figure3" httpUri="https://binary.pensoft.net/fig/940456" pageId="0" pageNumber="291" start="Figure 3" startId="F3">
<paragraph id="D01BEFBA839D6A53AF7C0C16714B1560" pageId="0" pageNumber="291">
<emphasis id="088B8B982C2528247DEC0C39E0CF2D8B" bold="true" pageId="0" pageNumber="291">Figure 3.</emphasis>
Frontal view of the embolic divisions of
<taxonomicName id="D8992C4DF22D9A1798DCA134B29463E0" authorityName="Eskov" authorityYear="1990" class="Arachnida" family="Linyphiidae" genus="Dactylopisthoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dactylopisthoides hyperboreus" order="Opiliones" pageId="0" pageNumber="291" phylum="Arthropoda" rank="species" species="hyperboreus">
<emphasis id="0A6300100009AA44E53D3F048E54265C" italics="true" pageId="0" pageNumber="291">Dactylopisthoides hyperboreus</emphasis>
</taxonomicName>
Eskov, 1990 (
<emphasis id="011F143D6CCD434C12B45248408847ED" bold="true" pageId="0" pageNumber="291">A</emphasis>
),
<taxonomicName id="7A7E6F38BCA1A3FA4F5E5B5904AC6476" lsidName="D. wrangelianus" pageId="0" pageNumber="291" rank="species" species="wrangelianus">
<emphasis id="912E31A69580874CFAC44D07AE271246" italics="true" pageId="0" pageNumber="291">D. wrangelianus</emphasis>
</taxonomicName>
(Marusik &amp; Koponen, 2009) (
<emphasis id="6ACAD8CC8F77931828286857928AE660" bold="true" pageId="0" pageNumber="291">B</emphasis>
) and
<taxonomicName id="3B6C2BBC6A448ABBF65C2660C9CDB87E" lsidName="D. transbaicalicus" pageId="0" pageNumber="291" rank="species" species="transbaicalicus">
<emphasis id="8C078213FA1EA39A4370518109A1315F" italics="true" pageId="0" pageNumber="291">D. transbaicalicus</emphasis>
</taxonomicName>
(Marusik, Koponen &amp; Danilov, 2001) [reproduced from
<bibRefCitation id="A3CA76667FC8849E8996C4E3CB022CA9" DOI="https://doi.org/10.33338/ef.84455" author="Marusik, YM" journalOrPublisher="Entomologica Fennica" pageId="0" pageNumber="291" pagination="18 - 21" refId="B6" refString="Marusik, YM, Koponen, S, 2009. A new species of Uusitaloia Marusik, Koponen &amp; Danilov (Araneae: Linyphiidae: Erigoninae), a genus through to be monotypic, from the Wrangel Island NE Russia. Entomologica Fennica 20 (1): 18 - 21, DOI: https://doi.org/10.33338/ef.84455" title="A new species of Uusitaloia Marusik, Koponen &amp; Danilov (Araneae: Linyphiidae: Erigoninae), a genus through to be monotypic, from the Wrangel Island NE Russia." url="https://doi.org/10.33338/ef.84455" volume="20" year="2009">Marusik and Koponen (2009)</bibRefCitation>
] (
<emphasis id="F6B9001CE9F03B3D6EB6C9A1957EB436" bold="true" pageId="0" pageNumber="291">C</emphasis>
). Abbreviations: dsa - distal suprategular apophysis, mm - median membrane, mt - median tooth, rt - radical tooth. Black, thick arrows show additional differences between species. Scale bar: 0.1 mm (
<emphasis id="17E138A5594439DF6B1DA4F52E16C35E" bold="true" pageId="0" pageNumber="291">
<normalizedToken id="35DD1474AE49B3DFDBDD819C6CE247C0" originalValue="С">S</normalizedToken>
</emphasis>
).
</paragraph>
</caption>
<paragraph id="080E97AD947C055DB8DF0E7E3F9220DF" pageId="0" pageNumber="291">
In the papers dealing with
<taxonomicName id="5C6FAFC2F02D882DA1E33A80DB7884E2" authorityName="Eskov" authorityYear="1990" class="Arachnida" family="Linyphiidae" genus="Dactylopisthoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dactylopisthoides" order="Opiliones" pageId="0" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="2DEFE5BD0450CF2DE854043FCE77A0F2" italics="true" pageId="0" pageNumber="291">Dactylopisthoides</emphasis>
</taxonomicName>
(
<bibRefCitation id="06B1FF78E042BC10980340A8ABB70C54" author="Eskov, KY" journalOrPublisher="Reichenbachia" pageId="0" pageNumber="291" pagination="1 - 5" refId="B1" refString="Eskov, KY, 1990. On the erigonine spider genera Dactylopisthes Simon, 1884 and Dactylopisthoides gen. nov. (Arachnida, Araneae: Linyphiidae). Reichenbachia 28: 1 - 5" title="On the erigonine spider genera Dactylopisthes Simon, 1884 and Dactylopisthoides gen. nov. (Arachnida, Araneae: Linyphiidae)." volume="28" year="1990">Eskov 1990</bibRefCitation>
) and
<taxonomicName id="32050E2BCAF27A42108B624FAB43F1FA" authorityName="Marusik, Koponen &amp; Danilov" authorityYear="2001" class="Arachnida" family="Linyphiidae" genus="Uusitaloia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uusitaloia" order="Araneae" pageId="0" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="E75DCEF449CA1D71422B3AA37016C6DB" italics="true" pageId="0" pageNumber="291">Uusitaloia</emphasis>
</taxonomicName>
(
<bibRefCitation id="9A940427F60307E258CCC726F3D1CAF2" author="Marusik, YM" journalOrPublisher="Bulletin - British Arachnological Society" pageId="0" pageNumber="291" pagination="83 - 92" refId="B7" refString="Marusik, YM, Koponen, S, Danilov, SN, 2001. Taxonomic and faunistic notes on linyphiids of Transbaikalia and south Siberia. Bulletin - British Arachnological Society 12 (2): 83 - 92" title="Taxonomic and faunistic notes on linyphiids of Transbaikalia and south Siberia." volume="12" year="2001">Marusik et al. 2001</bibRefCitation>
;
<bibRefCitation id="6B9C0B6D2361745ADF9ABE485133A072" DOI="https://doi.org/10.33338/ef.84455" author="Marusik, YM" journalOrPublisher="Entomologica Fennica" pageId="0" pageNumber="291" pagination="18 - 21" refId="B6" refString="Marusik, YM, Koponen, S, 2009. A new species of Uusitaloia Marusik, Koponen &amp; Danilov (Araneae: Linyphiidae: Erigoninae), a genus through to be monotypic, from the Wrangel Island NE Russia. Entomologica Fennica 20 (1): 18 - 21, DOI: https://doi.org/10.33338/ef.84455" title="A new species of Uusitaloia Marusik, Koponen &amp; Danilov (Araneae: Linyphiidae: Erigoninae), a genus through to be monotypic, from the Wrangel Island NE Russia." url="https://doi.org/10.33338/ef.84455" volume="20" year="2009">Marusik and Koponen 2009</bibRefCitation>
), the authors provided different tibial chaetotaxy even for different sexes. The reason for this appears to have been the small size of the spines, which could be easily overlooked (for further details, see
<bibRefCitation id="99F54B7173E2C135D5385289B90241C6" DOI="https://doi.org/10.15298/arthsel.31.1.13" author="Tanasevitch, AV" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="291" pagination="111 - 114" refId="B13" refString="Tanasevitch, AV, 2022. A new erigonine genus and species from the Russian Far East (Aranei: Linyphiidae), with notes on chaetotaxy. Arthropoda Selecta 31 (1): 111 - 114, DOI: https://doi.org/10.15298/arthsel.31.1.13" title="A new erigonine genus and species from the Russian Far East (Aranei: Linyphiidae), with notes on chaetotaxy." url="https://doi.org/10.15298/arthsel.31.1.13" volume="31" year="2022">Tanasevitch 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A6A286339DC2797C605BD1CE150660D8" pageId="0" pageNumber="291" type="composition">
<paragraph id="888D2DA0CA425A217FA3B321914505F9" pageId="0" pageNumber="291">Composition.</paragraph>
<paragraph id="5BB4E94A3C09C96EE21ED4699834D19B" pageId="0" pageNumber="291">
The genus
<taxonomicName id="27EB7833273006EE39343D8CBCB63EA4" authorityName="Eskov" authorityYear="1990" class="Arachnida" family="Linyphiidae" genus="Dactylopisthoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dactylopisthoides" order="Opiliones" pageId="0" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="74D8DBC8BB3C68CFB5A2E9E80585F506" italics="true" pageId="0" pageNumber="291">Dactylopisthoides</emphasis>
</taxonomicName>
currently consists of three very similar species:
<taxonomicName id="FB8EA3F4D3D1DAEDFCE70E83F75D9F9F" lsidName="D. hyperboreus" pageId="0" pageNumber="291" rank="species" species="hyperboreus">
<emphasis id="9E126AAA1E313590603A6A1B8160F57C" italics="true" pageId="0" pageNumber="291">D. hyperboreus</emphasis>
</taxonomicName>
Eskov, 1990;
<taxonomicName id="125EEAC6BFF0AE929ED5FCE564EC5E41" lsidName="D. transbaicalicus" pageId="0" pageNumber="291" rank="species" species="transbaicalicus">
<emphasis id="376AA9E3FC7554CAAA74422EC0B6F0E4" italics="true" pageId="0" pageNumber="291">D. transbaicalicus</emphasis>
</taxonomicName>
(Marusik, Koponen &amp; Danilov, 2001) and
<taxonomicName id="D5178F931F5EC3EB24292C280F756914" lsidName="D. wrangelianus" pageId="0" pageNumber="291" rank="species" species="wrangelianus">
<emphasis id="3C98E2B6BB4F5316395717FE34BAC1B8" italics="true" pageId="0" pageNumber="291">D. wrangelianus</emphasis>
</taxonomicName>
(Marusik &amp; Koponen, 2009).
</paragraph>
</subSubSection>
<subSubSection id="5D81A091BBC9262CC47937556A494B5C" pageId="0" pageNumber="291" type="distribution">
<paragraph id="DC3105B3DE432D5BDFCF47100F334362" pageId="0" pageNumber="291">Distribution.</paragraph>
<paragraph id="2EBAAAC5824201649035BD9275FC5AFC" pageId="0" pageNumber="291">
From north-eastern Transbaikalia to Wrangel Island (Fig.
<figureCitation id="3BAFB3A82C13BF3EE2D870F98A26C69F" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Collecting localities of Dactylopisthoides species: D. hyperboreus (dot), D. transbaicalicus (diamond), D. wrangelianus (square), records of D. hyperboreus that could also refer to D. wrangelianus (triangle)." figureDoi="10.3897/zookeys.1184.113255.figure4" httpUri="https://binary.pensoft.net/fig/940457" pageId="0" pageNumber="291">4</figureCitation>
). Its occurrence in Yakutia and Kamchatka is very likely.
</paragraph>
<caption id="BAF494F545216586ED28BA24B06A0840" doi="10.3897/zookeys.1184.113255.figure4" httpUri="https://binary.pensoft.net/fig/940457" pageId="0" pageNumber="291" start="Figure 4" startId="F4">
<paragraph id="88F791C4E73374480278093F716CA01A" pageId="0" pageNumber="291">
<emphasis id="098594A59D46D2923BC94D7F1CEAE403" bold="true" pageId="0" pageNumber="291">Figure 4.</emphasis>
Collecting localities of
<taxonomicName id="299A7FBC7BCEA7F591FF15E983284793" authorityName="Eskov" authorityYear="1990" class="Arachnida" family="Linyphiidae" genus="Dactylopisthoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dactylopisthoides" order="Opiliones" pageId="0" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="18444FCB3F1392719CCE0E48F016EDD6" italics="true" pageId="0" pageNumber="291">Dactylopisthoides</emphasis>
</taxonomicName>
species:
<taxonomicName id="4C5BFC856C07B1FF94B47FE0919BAA6B" lsidName="D. hyperboreus" pageId="0" pageNumber="291" rank="species" species="hyperboreus">
<emphasis id="0F98935D06B72E32D6C6800022CA6441" italics="true" pageId="0" pageNumber="291">D. hyperboreus</emphasis>
</taxonomicName>
(dot),
<taxonomicName id="2F1E8ABDFF9B71E81D7AAD32E4CA5FA6" lsidName="D. transbaicalicus" pageId="0" pageNumber="291" rank="species" species="transbaicalicus">
<emphasis id="711EFA62C544B3D2B57FEE56013887C9" italics="true" pageId="0" pageNumber="291">D. transbaicalicus</emphasis>
</taxonomicName>
(diamond),
<taxonomicName id="64DCD8DDCE8461EC134A1E2C6B5CB14D" lsidName="D. wrangelianus" pageId="0" pageNumber="291" rank="species" species="wrangelianus">
<emphasis id="B7A96EB8E8AE85A91636DD6413B71AE6" italics="true" pageId="0" pageNumber="291">D. wrangelianus</emphasis>
</taxonomicName>
(square), records of
<taxonomicName id="0DA739D0A9ED4E1B124897D3BC46C2D1" lsidName="D. hyperboreus" pageId="0" pageNumber="291" rank="species" species="hyperboreus">
<emphasis id="8A8C90ACE9FA29DF89A28906890A760C" italics="true" pageId="0" pageNumber="291">D. hyperboreus</emphasis>
</taxonomicName>
that could also refer to
<taxonomicName id="ECFCE0F8F4ED4C5A36590DE0BE425371" lsidName="D. wrangelianus" pageId="0" pageNumber="291" rank="species" species="wrangelianus">
<emphasis id="4C450A8E0CA3D8DAC46F246A657E26BE" italics="true" pageId="0" pageNumber="291">D. wrangelianus</emphasis>
</taxonomicName>
(triangle).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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