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<document id="45A4CC09AD2471EC665B15A91F7CD85B" ID-DOI="10.5281/zenodo.181374" ID-GBIF-Dataset="321f0d1f-9203-4f2e-8594-1dc16a9a1149" ID-ISSN="1175-5326" ID-Zenodo-Dep="181374" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459833167893" checkinUser="plazi" docAuthor="Grossi, Paschoal Coelho &amp; Abadie, Esteban Ignácio" docDate="2008" docId="460003706D38F215FF73411548F8F864" docLanguage="en" docName="zt01733p060.pdf" docOrigin="Zootaxa 1733" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Eremobothynus cornutus Steinheil 1872" docType="treatment" docVersion="9" lastPageNumber="59" masterDocId="BA397B086D3AF217FFE4470E4A7FFFD4" masterDocTitle="A new species of Eremobothynus Ohaus (Coleoptera: Scarabaeidae: Dynastinae) from Argentina, with notes on Eremobothynus cornutus (Steinheil)" masterLastPageNumber="60" masterPageNumber="57" pageNumber="59" updateTime="1698223136634" updateUser="plazi">
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<mods:title id="BAE8F09371F98855B9CA932EF5B5A49F">A new species of Eremobothynus Ohaus (Coleoptera: Scarabaeidae: Dynastinae) from Argentina, with notes on Eremobothynus cornutus (Steinheil)</mods:title>
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<mods:namePart id="36038D6795018C668B74C50B9848249F">Grossi, Paschoal Coelho</mods:namePart>
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<mods:namePart id="71996FF01D5D89D89E6E930C7DA96B69">Abadie, Esteban Ignácio</mods:namePart>
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<mods:date id="C37310BE10E06BCEEDC22A36B2934A01">2008</mods:date>
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<mods:number id="8DB40F3006235DC963FDE843719D34AD">1733</mods:number>
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<treatment id="460003706D38F215FF73411548F8F864" ID-DOI="http://doi.org/10.5281/zenodo.6231148" ID-GBIF-Taxon="119358280" ID-Zenodo-Dep="6231148" LSID="urn:lsid:plazi:treatment:460003706D38F215FF73411548F8F864" httpUri="http://treatment.plazi.org/id/460003706D38F215FF73411548F8F864" lastPageNumber="59" pageId="2" pageNumber="59">
<subSubSection id="86B3E1ED6D38F215FF7341154972F9E1" box="[151,781,1563,1590]" pageId="2" pageNumber="59" type="nomenclature">
<paragraph id="CE16B2666D38F215FF7341154972F9E1" blockId="2.[151,781,1563,1590]" box="[151,781,1563,1590]" pageId="2" pageNumber="59">
<heading id="955E050A6D38F215FF7341154972F9E1" bold="true" box="[151,781,1563,1590]" fontSize="11" level="1" pageId="2" pageNumber="59" reason="1">
<emphasis id="FCDD6E746D38F215FF7341154972F9E1" bold="true" box="[151,781,1563,1590]" pageId="2" pageNumber="59">
Notes on
<taxonomicName id="09A9C9E56D38F215FEEE41124972F9E1" authority="Steinheil, 1872" authorityName="Steinheil" authorityYear="1872" box="[266,781,1563,1590]" class="Insecta" family="Dynastidae" genus="Eremobothynus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="cornutus">
<emphasis id="FCDD6E746D38F215FEEE41124842F9E2" bold="true" box="[266,573,1564,1590]" italics="true" pageId="2" pageNumber="59">Eremobothynus cornutus</emphasis>
(
<bibRefCitation id="AA38CF976D38F215FDA84115497AF9E1" author="Steinheil" box="[588,773,1563,1589]" pageId="2" pageNumber="60" refString="Steinheil, E. (1872) Symbolae ad historiam Coleopterorum Argentinae meridionalis, ossia enumerazione dei coleotteri raccolti dal Prof. Strobel, durante il suo soggiorno in Buenos Aires e nei viaggi de la intrapresi a Mendoza e nal Chili, indi a Bahia Blanca ed al Carmen de los Patagones; e descrizione delle specie nuove. Atti della Societa Italiana di Scienze Naturali, 15, 554 - 578." type="journal article" year="1872">Steinheil, 1872</bibRefCitation>
)
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="86B3E1ED6D38F215FF73416F48F8F864" pageId="2" pageNumber="59" type="description">
<paragraph id="CE16B2666D38F215FF73416F4887F8D7" blockId="2.[151,1436,1633,1968]" pageId="2" pageNumber="59">
This species was described by
<bibRefCitation id="AA38CF976D38F215FE3B416F48F4F9AC" author="Steinheil" box="[479,651,1633,1656]" pageId="2" pageNumber="60" refString="Steinheil, E. (1872) Symbolae ad historiam Coleopterorum Argentinae meridionalis, ossia enumerazione dei coleotteri raccolti dal Prof. Strobel, durante il suo soggiorno in Buenos Aires e nei viaggi de la intrapresi a Mendoza e nal Chili, indi a Bahia Blanca ed al Carmen de los Patagones; e descrizione delle specie nuove. Atti della Societa Italiana di Scienze Naturali, 15, 554 - 578." type="journal article" year="1872">Steinheil (1872)</bibRefCitation>
based on a single male and was originally placed in the genus
<taxonomicName id="09A9C9E56D38F215FACC416F4B06F94F" authority="Burmeister, 1847" authorityName="Burmeister" authorityYear="1847" class="Insecta" family="Scarabaeidae" genus="Scaptophilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="FCDD6E746D38F215FACC416F4AC9F94F" italics="true" pageId="2" pageNumber="59">Scaptophilus</emphasis>
Burmeister, 1847
</taxonomicName>
, which was later synonymized with
<taxonomicName id="09A9C9E56D38F215FCE6418A4992F94F" authority="Hope, 1837" authorityName="Hope" authorityYear="1837" box="[770,1005,1668,1691]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="FCDD6E746D38F215FCE6418A491AF94F" box="[770,869,1668,1691]" italics="true" pageId="2" pageNumber="59">Bothynus</emphasis>
Hope, 1837
</taxonomicName>
.
<bibRefCitation id="AA38CF976D38F215FC1C418A4EF4F94F" author="Ohaus" box="[1016,1163,1668,1691]" pageId="2" pageNumber="60" refString="Ohaus, F. (1910) Neue Coleoptera Lamellicornia aus Argentinien. Deutsche Entomologische Zeitschrift, 2, 173 - 186." type="journal article" year="1910">Ohaus (1910)</bibRefCitation>
described the genus
<taxonomicName id="09A9C9E56D38F215FA89418A4B67F96A" class="Insecta" family="Dynastidae" genus="Eremobothynus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="FCDD6E746D38F215FA89418A4B67F96A" italics="true" pageId="2" pageNumber="59">Eremobothynus</emphasis>
</taxonomicName>
to accommodate this species.
<bibRefCitation id="AA38CF976D38F215FDBE41A84893F969" author="Ohaus" box="[602,748,1702,1725]" pageId="2" pageNumber="60" refString="Ohaus, F. (1910) Neue Coleoptera Lamellicornia aus Argentinien. Deutsche Entomologische Zeitschrift, 2, 173 - 186." type="journal article" year="1910">Ohaus (1910)</bibRefCitation>
also described the female of
<taxonomicName id="09A9C9E56D38F215FBC641A94EE3F96A" box="[1058,1180,1702,1726]" class="Insecta" family="Dynastidae" genus="Eremobothynus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="cornutus">
<emphasis id="FCDD6E746D38F215FBC641A94E4DF96A" box="[1058,1074,1703,1726]" italics="true" pageId="2" pageNumber="59">E</emphasis>
.
<emphasis id="FCDD6E746D38F215FBDB41A94EE3F96A" box="[1087,1180,1703,1726]" italics="true" pageId="2" pageNumber="59">cornutus</emphasis>
</taxonomicName>
, and mentioned that the three males he studied had a more developed horn than the “female” he examined. It is very probable that he was actually describing a very small male and mistook it for a female.
</paragraph>
<paragraph id="CE16B2666D38F215FF22400048F8F864" blockId="2.[151,1436,1633,1968]" pageId="2" pageNumber="59">
During the course of our studies, we examined a large number of male and female specimens of
<taxonomicName id="09A9C9E56D38F215FB2540014F44F8F2" box="[1217,1339,1806,1830]" class="Insecta" family="Dynastidae" genus="Eremobothynus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="cornutus">
<emphasis id="FCDD6E746D38F215FB2540014EAEF8F2" box="[1217,1233,1807,1830]" italics="true" pageId="2" pageNumber="59">E</emphasis>
.
<emphasis id="FCDD6E746D38F215FB3A40014F44F8F2" box="[1246,1339,1807,1830]" italics="true" pageId="2" pageNumber="59">cornutus</emphasis>
</taxonomicName>
from the second authors collection. All females examined were completely unarmed, with no horns or tubercles on the head (
<figureCitation id="5692AEE36D38F215FA95403F4AD8F8BF" captionStart="FIGURES 1 4" captionStartId="1.[151,269,1954,1978]" captionTargetBox="[302,1290,408,1929]" captionTargetId="figure@1.[292,1296,384,1948]" captionTargetPageId="1" captionText="FIGURES 1 4. Eremobothynus spp.: 1 - 2: E. hirsutus sp. nov., holotype, dorsal and lateral views; 3: E. cornutus, male dorsal view; 4: female head and pronotum anterolateral view." httpUri="https://zenodo.org/record/181375/files/figure.png" pageId="2" pageNumber="59">Fig. 4</figureCitation>
). Therefore, all identification keys (e.g.,
<bibRefCitation id="AA38CF976D38F215FDBE405A4891F8BF" author="Endrodi" box="[602,750,1876,1899]" pageId="2" pageNumber="60" refString="Endrodi, S. (1969) Monographie der Dynastinae, 4. Tribus, amerikanische Pentodontini. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden, 37, 1 - 145." type="journal article" year="1969">Endrödi 1969</bibRefCitation>
,
<bibRefCitation id="AA38CF976D38F215FD1D405A49F2F8BF" author="Endrodi" box="[761,909,1876,1899]" pageId="2" pageNumber="60" refString="Endrodi, S. (1985) The Dynastinae of the World. Dr. W. Junk Publisher, Dordrecht Netherlands. 800 pp., 46 plates." type="book" year="1985">Endrödi 1985</bibRefCitation>
,
<bibRefCitation id="AA38CF976D38F215FC73405A4E4DF8BF" author="Ratcliffe" box="[919,1074,1876,1899]" pageId="2" pageNumber="60" refString="Ratcliffe, B. C. (1981) Barutus hartmanni, a new genus and species from Panama with a key to the genera of New World Pentodontini (Coleoptera: Scarabaeidae: Dynastinae). Coleopterists Bulletin, 35, 463 - 472." type="journal article" year="1981">Ratcliffe 1981</bibRefCitation>
,
<bibRefCitation id="AA38CF976D38F215FBD9405A4F44F8BF" author="Ratcliffe" box="[1085,1339,1876,1899]" pageId="2" pageNumber="60" refString="Ratcliffe, B. C. &amp; Hardy, M. (2005) Collagenus dasysternus, a new genus and species of Dynastinae from eastern Venezuela with a key to the New World genera of Pentodontini (Coleoptera: Scarabaeidae: Dynastinae). Coleopterists Bulletin, 59, 143 - 150." type="journal article" year="2005">Ratcliffe &amp; Hardy 2005</bibRefCitation>
) must be corrected to indicate that only males in
<taxonomicName id="09A9C9E56D38F215FDB5407948B1F85A" box="[593,718,1910,1934]" class="Insecta" family="Dynastidae" genus="Eremobothynus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="cornutus">
<emphasis id="FCDD6E746D38F215FDB54079481EF85A" box="[593,609,1911,1934]" italics="true" pageId="2" pageNumber="59">E</emphasis>
.
<emphasis id="FCDD6E746D38F215FD95407948B1F85A" box="[625,718,1911,1934]" italics="true" pageId="2" pageNumber="59">cornutus</emphasis>
</taxonomicName>
have a frontal horn (
<figureCitation id="5692AEE36D38F215FC5840784980F859" box="[956,1023,1910,1933]" captionStart="FIGURES 1 4" captionStartId="1.[151,269,1954,1978]" captionTargetBox="[302,1290,408,1929]" captionTargetId="figure@1.[292,1296,384,1948]" captionTargetPageId="1" captionText="FIGURES 1 4. Eremobothynus spp.: 1 - 2: E. hirsutus sp. nov., holotype, dorsal and lateral views; 3: E. cornutus, male dorsal view; 4: female head and pronotum anterolateral view." httpUri="https://zenodo.org/record/181375/files/figure.png" pageId="2" pageNumber="59">Fig. 3</figureCitation>
), and that the pronotum is convex in females (
<figureCitation id="5692AEE36D38F215FF1C40974B47F864" box="[248,312,1945,1968]" captionStart="FIGURES 1 4" captionStartId="1.[151,269,1954,1978]" captionTargetBox="[302,1290,408,1929]" captionTargetId="figure@1.[292,1296,384,1948]" captionTargetPageId="1" captionText="FIGURES 1 4. Eremobothynus spp.: 1 - 2: E. hirsutus sp. nov., holotype, dorsal and lateral views; 3: E. cornutus, male dorsal view; 4: female head and pronotum anterolateral view." httpUri="https://zenodo.org/record/181375/files/figure.png" pageId="2" pageNumber="59">Fig. 4</figureCitation>
) and not declivous as in males.
</paragraph>
</subSubSection>
</treatment>
</document>