412 lines
54 KiB
XML
412 lines
54 KiB
XML
<document id="E3999FE91CE181B9AC334AD753F208DF" ID-DOI="10.11646/zootaxa.3980.3.4" ID-GBIF-Dataset="4788bd35-7502-421d-8331-fc980ce1afab" ID-ISSN="1175-5326" ID-Zenodo-Dep="236576" ID-ZooBank="B88E0AA3-260A-43E1-AEC8-9973A283FB11" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461243030218" checkinUser="plazi" docAuthor="Jono, Teppei, Bauer, Aaron M., Brennan, Ian & Mori, Akira" docDate="2015" docId="452EF71F6909FFD9FF5CF56AC0DFFE75" docLanguage="en" docName="zt03980p416.pdf" docOrigin="Zootaxa 3980 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Blaesodactylus microtuberculatus Jono, Bauer, Brennan & Mori, 2015, sp. nov." docType="treatment" docVersion="8" lastPageNumber="413" masterDocId="B9178F67690AFFDEFFCBF32BC36AFFBA" masterDocTitle="New species of Blaesodactylus (Squamata: Gekkonidae) from Tsingy karstic outcrops in Ankarana National Park, northern Madagascar" masterLastPageNumber="416" masterPageNumber="406" pageNumber="409" updateTime="1698605450761" updateUser="plazi">
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<mods:title id="8DC3B2953BBA21514DEAFFB6556D421B">New species of Blaesodactylus (Squamata: Gekkonidae) from Tsingy karstic outcrops in Ankarana National Park, northern Madagascar</mods:title>
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<mods:name id="BBA65D0BA02CB4F8A3B779E6BE5F2477" type="personal">
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<mods:roleTerm id="2361DB4B1B28B20570AC91A14B2B83E1">Author</mods:roleTerm>
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<mods:namePart id="835D2FA92FA215F52A8E567151242EEC">Jono, Teppei</mods:namePart>
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<mods:roleTerm id="8F370191BD947921356BE03974FB9A87">Author</mods:roleTerm>
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<mods:namePart id="4DECBCA38A5B5F4DCDCCF1C2B625EEEC">Bauer, Aaron M.</mods:namePart>
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<mods:roleTerm id="D76AC2A204D268174FA4970CDB8EC68C">Author</mods:roleTerm>
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<mods:namePart id="26E2AA1E477167E3FD19790F1C2BCE8B">Brennan, Ian</mods:namePart>
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<mods:name id="C9FA044F6912746219743E328EEF87F1" type="personal">
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<mods:roleTerm id="D152A9999C8532066A97245AC35584FA">Author</mods:roleTerm>
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<mods:namePart id="D0FF183565F946D3DE25A7384BD5454A">Mori, Akira</mods:namePart>
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<mods:typeOfResource id="361DEB13F1C9F3F0A6A8C3397D2C8CB6">text</mods:typeOfResource>
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<mods:title id="4740AC24CA9093416A61785806434E56">Zootaxa</mods:title>
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<mods:date id="ECDBAC8711208410347AC45D0F6A069E">2015</mods:date>
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<mods:number id="28C8BEF0A64ED8CA6F8C6084A1AA720F">3980</mods:number>
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<treatment id="452EF71F6909FFD9FF5CF56AC0DFFE75" ID-DOI="http://doi.org/10.5281/zenodo.5671558" ID-GBIF-Taxon="119632976" ID-Zenodo-Dep="5671558" LSID="urn:lsid:plazi:treatment:452EF71F6909FFD9FF5CF56AC0DFFE75" httpUri="http://treatment.plazi.org/id/452EF71F6909FFD9FF5CF56AC0DFFE75" lastPageId="7" lastPageNumber="413" pageId="3" pageNumber="409">
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<subSubSection id="859D15826909FFDDFF5CF56AC200F9C7" pageId="3" pageNumber="409" type="nomenclature">
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<paragraph id="CD3846096909FFDDFF5CF56AC1FFF9E1" blockId="3.[151,661,1601,1661]" box="[151,661,1601,1627]" pageId="3" pageNumber="409">
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<heading id="9670F1656909FFDDFF5CF56AC1FFF9E1" bold="true" box="[151,661,1601,1627]" fontSize="11" level="1" pageId="3" pageNumber="409" reason="1">
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<emphasis id="FFF39A1B6909FFDDFF5CF56AC1FFF9E1" bold="true" box="[151,661,1601,1627]" pageId="3" pageNumber="409">
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<taxonomicName id="0A873D8A6909FFDDFF5CF56AC15BF9E1" box="[151,561,1601,1627]" class="Reptilia" family="Gekkonidae" genus="Blaesodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="409" phylum="Chordata" rank="species" species="microtuberculatus" status="sp. nov.">
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<emphasis id="FFF39A1B6909FFDDFF5CF56AC15BF9E1" bold="true" box="[151,561,1601,1627]" italics="true" pageId="3" pageNumber="409">Blaesodactylus microtuberculatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="E4C027606909FFDDFDFCF56AC1FFF9E1" box="[567,661,1601,1627]" pageId="3" pageNumber="409" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="CD3846096909FFDDFF5CF54FC200F9C7" blockId="3.[151,661,1601,1661]" box="[151,362,1636,1661]" pageId="3" pageNumber="409">
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(
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<figureCitation id="55BC5A8C6909FFDDFF54F54FC386F9C6" box="[159,236,1636,1661]" captionStart="FIGURE 2" captionStartId="4.[151,250,1801,1823]" captionTargetBox="[174,1410,777,1766]" captionTargetId="figure@4.[167,1419,749,1779]" captionTargetPageId="4" captionText="FIGURE 2. Holotype of Blaesodactylus microtuberculatus sp. nov. and representatives of three of its congeners. A. B. microtuberculatus sp. nov. (KUZ 069431). B. B. boivini (KUZ 069429). C. B. sakalava (KUZ 069432). D. B. ambonihazo (KUZ 069433)." httpUri="https://zenodo.org/record/236578/files/figure.png" pageId="3" pageNumber="409">Figs. 2</figureCitation>
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A, 4A, 5A)
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</paragraph>
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</subSubSection>
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<subSubSection id="859D15826909FFDDFF5CF587C2FBF8B6" pageId="3" pageNumber="409" type="materials_examined">
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<paragraph id="CD3846096909FFDDFF5CF587C6FCF952" blockId="3.[151,1436,1708,2021]" pageId="3" pageNumber="409">
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<emphasis id="FFF39A1B6909FFDDFF5CF587C261F97E" bold="true" box="[151,267,1708,1732]" pageId="3" pageNumber="409">
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<typeStatus id="123CF8AB6909FFDDFF5CF587C26CF97E" box="[151,262,1708,1732]" pageId="3" pageNumber="409" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectionCode id="AB96DECC6909FFDDFED9F587C226F97F" box="[274,332,1708,1733]" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="3" pageNumber="409">KUZ</collectionCode>
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069431 (field number
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<collectionCode id="AB96DECC6909FFDDFD82F587C115F97E" box="[585,639,1708,1732]" pageId="3" pageNumber="409">AFP</collectionCode>
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2012-002), adult male (
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<figureCitation id="55BC5A8C6909FFDDFC4DF587C0A0F97E" box="[902,970,1708,1733]" captionStart="FIGURE 2" captionStartId="4.[151,250,1801,1823]" captionTargetBox="[174,1410,777,1766]" captionTargetId="figure@4.[167,1419,749,1779]" captionTargetPageId="4" captionText="FIGURE 2. Holotype of Blaesodactylus microtuberculatus sp. nov. and representatives of three of its congeners. A. B. microtuberculatus sp. nov. (KUZ 069431). B. B. boivini (KUZ 069429). C. B. sakalava (KUZ 069432). D. B. ambonihazo (KUZ 069433)." httpUri="https://zenodo.org/record/236578/files/figure.png" pageId="3" pageNumber="409">Fig. 2</figureCitation>
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A); Ankarana National Park, Antsiranana Province, northern
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<collectingCountry id="B59006996909FFDDFEA6F5E4C29EF952" box="[365,500,1743,1768]" name="Madagascar" pageId="3" pageNumber="409">Madagascar</collectingCountry>
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(
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<geoCoordinate id="A8B320CE6909FFDDFDCFF5E4C108F952" box="[516,610,1743,1768]" direction="south" orientation="latitude" pageId="3" pageNumber="409" precision="925" value="-12.95">12°57’S</geoCoordinate>
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,
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<geoCoordinate id="A8B320CE6909FFDDFDA5F5FBC1A5F952" box="[622,719,1744,1768]" direction="east" orientation="longitude" pageId="3" pageNumber="409" precision="925" value="49.116665">49°07’E</geoCoordinate>
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;
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<figureCitation id="55BC5A8C6909FFDDFD11F5E4C076F952" box="[730,796,1743,1768]" captionStart="FIGURE 3" captionStartId="5.[151,249,1948,1970]" captionTargetBox="[172,1415,848,1926]" captionTargetId="figure@5.[152,1435,841,1927]" captionTargetPageId="5" captionText="FIGURE 3. Localities and habitat of two allotopic Blaesodactylus spp. A. Map of Ankarana National Park. Note that the Tsingy karstic outcroppings are inside the forest. Open circle indicates the site where B. microtuberculatus sp. nov. was found and closed circles indicate the sites where B. boivini was found. B. Habitat of B. microtuberculatus sp. nov. C. Habitat of B. boivini." httpUri="https://zenodo.org/record/236579/files/figure.png" pageId="3" pageNumber="409">Fig. 3</figureCitation>
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), collected by T. Jono and A. Mori on
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<date id="B93960C96909FFDDFB02F5FBC6F9F952" box="[1225,1427,1743,1768]" pageId="3" pageNumber="409" value="2012-12-04">4 December 2012</date>
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.
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</paragraph>
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<paragraph id="CD3846096909FFDDFF0CF5DFC2FBF8B6" blockId="3.[151,1436,1708,2021]" box="[199,401,1780,1804]" pageId="3" pageNumber="409">
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<emphasis id="FFF39A1B6909FFDDFF0CF5DFC22CF8B6" bold="true" box="[199,326,1780,1804]" pageId="3" pageNumber="409">
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<typeStatus id="123CF8AB6909FFDDFF0CF5DFC22BF8B6" box="[199,321,1780,1804]" pageId="3" pageNumber="409" type="paratype">Paratypes</typeStatus>
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.
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</emphasis>
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None.
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</paragraph>
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</subSubSection>
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<subSubSection id="859D15826909FFDDFF0CF433C12CF8EF" pageId="3" pageNumber="409" type="etymology">
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<paragraph id="CD3846096909FFDDFF0CF433C12CF8EF" blockId="3.[151,1436,1708,2021]" pageId="3" pageNumber="409">
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<emphasis id="FFF39A1B6909FFDDFF0CF433C224F88A" bold="true" box="[199,334,1816,1840]" pageId="3" pageNumber="409">Etymology.</emphasis>
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The species epithet “
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<taxonomicName id="0A873D8A6909FFDDFD8BF433C061F88A" box="[576,779,1816,1840]" class="Reptilia" family="Gekkonidae" genus="Blaesodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="409" phylum="Chordata" rank="species" species="microtuberculatus">
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<emphasis id="FFF39A1B6909FFDDFD8BF433C061F88A" box="[576,779,1816,1840]" italics="true" pageId="3" pageNumber="409">microtuberculatus</emphasis>
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</taxonomicName>
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” refers to the presence of small tubercles, compared to all other congeners, on their trunk region.
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</paragraph>
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</subSubSection>
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<subSubSection id="859D15826909FFDAFF0CF474C777FD77" lastPageId="4" lastPageNumber="410" pageId="3" pageNumber="409" type="diagnosis">
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<paragraph id="CD3846096909FFDAFF0CF474C077FF6E" blockId="3.[151,1436,1708,2021]" lastBlockId="4.[151,1436,151,717]" lastPageId="4" lastPageNumber="410" pageId="3" pageNumber="409">
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<emphasis id="FFF39A1B6909FFDDFF0CF474C22BF8C2" bold="true" box="[199,321,1887,1912]" pageId="3" pageNumber="409">Diagnosis.</emphasis>
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A large sized (SVL
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<quantity id="0A7FEBEC6909FFDDFDF8F44BC1C0F8C2" box="[563,682,1888,1912]" metricMagnitude="-1" metricUnit="m" metricValue="1.174" pageId="3" pageNumber="409" unit="mm" value="117.4">117.4 mm</quantity>
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), robust-bodied
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<taxonomicName id="0A873D8A6909FFDDFCAFF44BC764F8C2" box="[868,1038,1888,1912]" class="Reptilia" family="Gekkonidae" genus="Blaesodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="409" phylum="Chordata" rank="genus">
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<emphasis id="FFF39A1B6909FFDDFCAFF44BC764F8C2" box="[868,1038,1888,1912]" italics="true" pageId="3" pageNumber="409">Blaesodactylus</emphasis>
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</taxonomicName>
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with tail slightly longer than SVL (
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<figureCitation id="55BC5A8C6909FFDDFF54F4AFC38FF826" box="[159,229,1924,1949]" captionStart="FIGURE 2" captionStartId="4.[151,250,1801,1823]" captionTargetBox="[174,1410,777,1766]" captionTargetId="figure@4.[167,1419,749,1779]" captionTargetPageId="4" captionText="FIGURE 2. Holotype of Blaesodactylus microtuberculatus sp. nov. and representatives of three of its congeners. A. B. microtuberculatus sp. nov. (KUZ 069431). B. B. boivini (KUZ 069429). C. B. sakalava (KUZ 069432). D. B. ambonihazo (KUZ 069433)." httpUri="https://zenodo.org/record/236578/files/figure.png" pageId="3" pageNumber="409">Fig. 2</figureCitation>
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A). Snout elongate, gular granules small and homogenous, internarial region convex (
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<figureCitation id="55BC5A8C6909FFDDFB75F4AFC66EF826" box="[1214,1284,1924,1949]" captionStart="FIGURE 4" captionStartId="6.[151,250,892,914]" captionTargetBox="[151,1436,193,870]" captionTargetId="figure@6.[151,1436,193,871]" captionTargetPageId="6" captionText="FIGURE 4. Lateral and ventral views of heads of Blaesodactylus spp. A. B. microtuberculatus sp. nov. (KUZ 069431). B. B. boivini (KUZ 069429). C. B. sakalava (KUZ 069432). D. B. ambonihazo (KUZ 069433)." httpUri="https://zenodo.org/record/236580/files/figure.png" pageId="3" pageNumber="409">Fig. 4</figureCitation>
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A). 162 scale rows around midbody, 15–17 longitudinal rows of relatively small tubercles on dorsum. Fourth digit of pes with 21 undivided subdigital lamellae. No tubercles on tail dorsum; midventral subcaudal scales transversely enlarged, about half width of tail. Body dorsum grayish, mid-dorsal row of five blotches, original tail weakly banded, venter pale, only gular region with very faint mottling (
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<figureCitation id="55BC5A8C690EFFDAFD7DF397C193FF6E" box="[694,761,188,213]" captionStart="FIGURE 4" captionStartId="6.[151,250,892,914]" captionTargetBox="[151,1436,193,870]" captionTargetId="figure@6.[151,1436,193,871]" captionTargetPageId="6" captionText="FIGURE 4. Lateral and ventral views of heads of Blaesodactylus spp. A. B. microtuberculatus sp. nov. (KUZ 069431). B. B. boivini (KUZ 069429). C. B. sakalava (KUZ 069432). D. B. ambonihazo (KUZ 069433)." httpUri="https://zenodo.org/record/236580/files/figure.png" pageId="4" pageNumber="410">Fig. 4</figureCitation>
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A).
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</paragraph>
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<paragraph id="CD384609690EFFDAFF0CF3CBC777FD77" blockId="4.[151,1436,151,717]" pageId="4" pageNumber="410">
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Among its congeners
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<taxonomicName id="0A873D8A690EFFDAFE0AF3CBC054FF42" box="[449,830,224,248]" class="Reptilia" family="Gekkonidae" genus="Blaesodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="410" phylum="Chordata" rank="species" species="microtuberculatus" status="sp. nov.">
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<emphasis id="FFF39A1B690EFFDAFE0AF3CBC054FF42" box="[449,830,224,248]" italics="true" pageId="4" pageNumber="410">Blaesodactylus microtuberculatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="E4C02760690EFFDAFC8CF3CBC0C8FF42" box="[839,930,224,248]" pageId="4" pageNumber="410" rank="species">
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<emphasis id="FFF39A1B690EFFDAFC8CF3CBC0C8FF42" bold="true" box="[839,930,224,248]" pageId="4" pageNumber="410">sp. nov.</emphasis>
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</taxonomicNameLabel>
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may be differentiated from
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<taxonomicName id="0A873D8A690EFFDAFB2EF3CAC63EFF42" box="[1253,1364,224,248]" class="Reptilia" family="Gekkonidae" genus="Blaesodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="410" phylum="Chordata" rank="species" species="boivini">
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<emphasis id="FFF39A1B690EFFDAFB2EF3CAC63EFF42" box="[1253,1364,224,248]" italics="true" pageId="4" pageNumber="410">B. boivini</emphasis>
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</taxonomicName>
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by its smaller dorsal tubercles (
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<figureCitation id="55BC5A8C690EFFDAFE73F22FC295FEA7" box="[440,511,260,285]" captionStart="FIGURE 5" captionStartId="7.[151,250,1696,1718]" captionTargetBox="[174,1415,519,1673]" captionTargetId="figure@7.[168,1418,519,1674]" captionTargetPageId="7" captionText="FIGURE 5. Dorsal, ventral, tail, and subcaudal scalations of Blaesodactylus spp. A. B. microtubaclata sp. nov. (KUZ 069431). B. B. boivini (KUZ 069429). C. B. sakalava (KUZ 069432). D. B. ambonihazo (KUZ 069433). Arrows indicate tubercles. Magnification of all photos is identical." httpUri="https://zenodo.org/record/236581/files/figure.png" pageId="4" pageNumber="410">Fig. 5</figureCitation>
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; ~3 times size of adjacent scales
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<emphasis id="FFF39A1B690EFFDAFC4FF22EC0A6FEA6" box="[900,972,261,284]" italics="true" pageId="4" pageNumber="410">versus</emphasis>
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~10 times), absence of caudal tubercles (
|
||
<figureCitation id="55BC5A8C690EFFDAFF54F20CC38EFEFA" box="[159,228,295,320]" captionStart="FIGURE 5" captionStartId="7.[151,250,1696,1718]" captionTargetBox="[174,1415,519,1673]" captionTargetId="figure@7.[168,1418,519,1674]" captionTargetPageId="7" captionText="FIGURE 5. Dorsal, ventral, tail, and subcaudal scalations of Blaesodactylus spp. A. B. microtubaclata sp. nov. (KUZ 069431). B. B. boivini (KUZ 069429). C. B. sakalava (KUZ 069432). D. B. ambonihazo (KUZ 069433). Arrows indicate tubercles. Magnification of all photos is identical." httpUri="https://zenodo.org/record/236581/files/figure.png" pageId="4" pageNumber="410">Fig. 5</figureCitation>
|
||
;
|
||
<emphasis id="FFF39A1B690EFFDAFF3AF202C253FEFA" box="[241,313,297,320]" italics="true" pageId="4" pageNumber="410">versus</emphasis>
|
||
caudal tubercles ~4 times size of adjacent scales and present on more than half of the tail), uniform pale venter except for gular region (
|
||
<emphasis id="FFF39A1B690EFFDAFDFAF266C113FEDE" box="[561,633,333,356]" italics="true" pageId="4" pageNumber="410">versus</emphasis>
|
||
mottled with areas of dark pigmentation), and lower number of dorsal tubercle rows (15–17
|
||
<emphasis id="FFF39A1B690EFFDAFE40F25AC2B9FE32" box="[395,467,369,392]" italics="true" pageId="4" pageNumber="410">versus</emphasis>
|
||
18–21). It differs from
|
||
<taxonomicName id="0A873D8A690EFFDAFD11F25AC037FE32" box="[730,861,368,392]" class="Reptilia" family="Gekkonidae" genus="Blaesodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="410" phylum="Chordata" rank="species" species="sakalava">
|
||
<emphasis id="FFF39A1B690EFFDAFD11F25AC037FE32" box="[730,861,368,392]" italics="true" pageId="4" pageNumber="410">B. sakalava</emphasis>
|
||
</taxonomicName>
|
||
in its smaller dorsal tubercles (
|
||
<figureCitation id="55BC5A8C690EFFDAFB73F244C796FE32" box="[1208,1276,367,392]" captionStart="FIGURE 5" captionStartId="7.[151,250,1696,1718]" captionTargetBox="[174,1415,519,1673]" captionTargetId="figure@7.[168,1418,519,1674]" captionTargetPageId="7" captionText="FIGURE 5. Dorsal, ventral, tail, and subcaudal scalations of Blaesodactylus spp. A. B. microtubaclata sp. nov. (KUZ 069431). B. B. boivini (KUZ 069429). C. B. sakalava (KUZ 069432). D. B. ambonihazo (KUZ 069433). Arrows indicate tubercles. Magnification of all photos is identical." httpUri="https://zenodo.org/record/236581/files/figure.png" pageId="4" pageNumber="410">Fig. 5</figureCitation>
|
||
; ~3 times size of adjacent scales
|
||
<emphasis id="FFF39A1B690EFFDAFEA3F2BEC2DAFE16" box="[360,432,405,428]" italics="true" pageId="4" pageNumber="410">versus</emphasis>
|
||
~6 times, with tubercles finely keel), absence of caudal tubercles (
|
||
<figureCitation id="55BC5A8C690EFFDAFB65F2BFC79EFE17" box="[1198,1268,404,429]" captionStart="FIGURE 5" captionStartId="7.[151,250,1696,1718]" captionTargetBox="[174,1415,519,1673]" captionTargetId="figure@7.[168,1418,519,1674]" captionTargetPageId="7" captionText="FIGURE 5. Dorsal, ventral, tail, and subcaudal scalations of Blaesodactylus spp. A. B. microtubaclata sp. nov. (KUZ 069431). B. B. boivini (KUZ 069429). C. B. sakalava (KUZ 069432). D. B. ambonihazo (KUZ 069433). Arrows indicate tubercles. Magnification of all photos is identical." httpUri="https://zenodo.org/record/236581/files/figure.png" pageId="4" pageNumber="410">Fig. 5</figureCitation>
|
||
;
|
||
<emphasis id="FFF39A1B690EFFDAFAC8F2BEC621FE16" box="[1283,1355,405,428]" italics="true" pageId="4" pageNumber="410">versus</emphasis>
|
||
keeled tubercles ~6 times size of adjacent scales and present on more than half of the tail), wider midventral subcaudal scales (
|
||
<figureCitation id="55BC5A8C690EFFDAFF20F2F7C25BFE4F" box="[235,305,476,501]" captionStart="FIGURE 5" captionStartId="7.[151,250,1696,1718]" captionTargetBox="[174,1415,519,1673]" captionTargetId="figure@7.[168,1418,519,1674]" captionTargetPageId="7" captionText="FIGURE 5. Dorsal, ventral, tail, and subcaudal scalations of Blaesodactylus spp. A. B. microtubaclata sp. nov. (KUZ 069431). B. B. boivini (KUZ 069429). C. B. sakalava (KUZ 069432). D. B. ambonihazo (KUZ 069433). Arrows indicate tubercles. Magnification of all photos is identical." httpUri="https://zenodo.org/record/236581/files/figure.png" pageId="4" pageNumber="410">Fig. 5</figureCitation>
|
||
; about half width of tail
|
||
<emphasis id="FFF39A1B690EFFDAFD99F2F6C1F0FE4E" box="[594,666,477,500]" italics="true" pageId="4" pageNumber="410">versus</emphasis>
|
||
less than one-third width of tail), and mottled gular region (
|
||
<figureCitation id="55BC5A8C690EFFDAFA9BF2F7C6FDFE4E" box="[1360,1431,476,501]" captionStart="FIGURE 4" captionStartId="6.[151,250,892,914]" captionTargetBox="[151,1436,193,870]" captionTargetId="figure@6.[151,1436,193,871]" captionTargetPageId="6" captionText="FIGURE 4. Lateral and ventral views of heads of Blaesodactylus spp. A. B. microtuberculatus sp. nov. (KUZ 069431). B. B. boivini (KUZ 069429). C. B. sakalava (KUZ 069432). D. B. ambonihazo (KUZ 069433)." httpUri="https://zenodo.org/record/236580/files/figure.png" pageId="4" pageNumber="410">Fig. 4</figureCitation>
|
||
;
|
||
<emphasis id="FFF39A1B690EFFDAFF5CF12AC3B5FDA2" box="[151,223,513,536]" italics="true" pageId="4" pageNumber="410">versus</emphasis>
|
||
pale). It is distinguished from
|
||
<taxonomicName id="0A873D8A690EFFDAFD98F12AC06AFDA2" box="[595,768,512,536]" class="Reptilia" family="Gekkonidae" genus="Blaesodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="410" phylum="Chordata" rank="species" species="ambonihazo">
|
||
<emphasis id="FFF39A1B690EFFDAFD98F12AC06AFDA2" box="[595,768,512,536]" italics="true" pageId="4" pageNumber="410">B. ambonihazo</emphasis>
|
||
</taxonomicName>
|
||
by its smaller dorsal tubercles (
|
||
<figureCitation id="55BC5A8C690EFFDAFB4AF2D4C7A0FDA2" box="[1153,1226,511,536]" captionStart="FIGURE 5" captionStartId="7.[151,250,1696,1718]" captionTargetBox="[174,1415,519,1673]" captionTargetId="figure@7.[168,1418,519,1674]" captionTargetPageId="7" captionText="FIGURE 5. Dorsal, ventral, tail, and subcaudal scalations of Blaesodactylus spp. A. B. microtubaclata sp. nov. (KUZ 069431). B. B. boivini (KUZ 069429). C. B. sakalava (KUZ 069432). D. B. ambonihazo (KUZ 069433). Arrows indicate tubercles. Magnification of all photos is identical." httpUri="https://zenodo.org/record/236581/files/figure.png" pageId="4" pageNumber="410">Fig. 5</figureCitation>
|
||
; ~3 times size of adjacent scales
|
||
<emphasis id="FFF39A1B690EFFDAFE87F10EC2FEFD86" box="[332,404,549,572]" italics="true" pageId="4" pageNumber="410">versus</emphasis>
|
||
~6 times), absence of caudal tubercles (
|
||
<figureCitation id="55BC5A8C690EFFDAFCBBF10FC0D2FD87" box="[880,952,548,573]" captionStart="FIGURE 5" captionStartId="7.[151,250,1696,1718]" captionTargetBox="[174,1415,519,1673]" captionTargetId="figure@7.[168,1418,519,1674]" captionTargetPageId="7" captionText="FIGURE 5. Dorsal, ventral, tail, and subcaudal scalations of Blaesodactylus spp. A. B. microtubaclata sp. nov. (KUZ 069431). B. B. boivini (KUZ 069429). C. B. sakalava (KUZ 069432). D. B. ambonihazo (KUZ 069433). Arrows indicate tubercles. Magnification of all photos is identical." httpUri="https://zenodo.org/record/236581/files/figure.png" pageId="4" pageNumber="410">Fig. 5</figureCitation>
|
||
;
|
||
<emphasis id="FFF39A1B690EFFDAFC02F10EC77AFD86" box="[969,1040,549,572]" italics="true" pageId="4" pageNumber="410">versus</emphasis>
|
||
keeled tubercles ~3 times size of adjacent scales present on proximal ~40% of the tail), wider midventral subcaudal scales (
|
||
<figureCitation id="55BC5A8C690EFFDAFB5AF16CC7BCFDDA" box="[1169,1238,583,608]" captionStart="FIGURE 5" captionStartId="7.[151,250,1696,1718]" captionTargetBox="[174,1415,519,1673]" captionTargetId="figure@7.[168,1418,519,1674]" captionTargetPageId="7" captionText="FIGURE 5. Dorsal, ventral, tail, and subcaudal scalations of Blaesodactylus spp. A. B. microtubaclata sp. nov. (KUZ 069431). B. B. boivini (KUZ 069429). C. B. sakalava (KUZ 069432). D. B. ambonihazo (KUZ 069433). Arrows indicate tubercles. Magnification of all photos is identical." httpUri="https://zenodo.org/record/236581/files/figure.png" pageId="4" pageNumber="410">Fig. 5</figureCitation>
|
||
; about half width of tail
|
||
<emphasis id="FFF39A1B690EFFDAFF29F146C240FD3E" box="[226,298,621,644]" italics="true" pageId="4" pageNumber="410">versus</emphasis>
|
||
less than one-third width of tail), mottled gular region (
|
||
<figureCitation id="55BC5A8C690EFFDAFC63F147C084FD3E" box="[936,1006,620,645]" captionStart="FIGURE 4" captionStartId="6.[151,250,892,914]" captionTargetBox="[151,1436,193,870]" captionTargetId="figure@6.[151,1436,193,871]" captionTargetPageId="6" captionText="FIGURE 4. Lateral and ventral views of heads of Blaesodactylus spp. A. B. microtuberculatus sp. nov. (KUZ 069431). B. B. boivini (KUZ 069429). C. B. sakalava (KUZ 069432). D. B. ambonihazo (KUZ 069433)." httpUri="https://zenodo.org/record/236580/files/figure.png" pageId="4" pageNumber="410">Fig. 4</figureCitation>
|
||
;
|
||
<emphasis id="FFF39A1B690EFFDAFC30F146C728FD3E" box="[1019,1090,621,644]" italics="true" pageId="4" pageNumber="410">versus</emphasis>
|
||
pale), lower number of dorsal tubercle rows (15–17
|
||
<emphasis id="FFF39A1B690EFFDAFE50F1BAC289FD12" box="[411,483,657,680]" italics="true" pageId="4" pageNumber="410">versus</emphasis>
|
||
17–21). It is distinguished from
|
||
<taxonomicName id="0A873D8A690EFFDAFCB9F1BAC741FD12" box="[882,1067,656,680]" class="Reptilia" family="Gekkonidae" genus="Blaesodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="410" phylum="Chordata" rank="species" species="antongilensis">
|
||
<emphasis id="FFF39A1B690EFFDAFCB9F1BAC741FD12" box="[882,1067,656,680]" italics="true" pageId="4" pageNumber="410">B. antongilensis</emphasis>
|
||
</taxonomicName>
|
||
by its lower number of dorsal tubercles (15–17
|
||
<emphasis id="FFF39A1B690EFFDAFE92F19EC2CAFD76" box="[345,416,693,716]" italics="true" pageId="4" pageNumber="410">versus</emphasis>
|
||
up to 24), and mottled gular region (
|
||
<figureCitation id="55BC5A8C690EFFDAFCF5F19FC0EEFD76" box="[830,900,692,717]" captionStart="FIGURE 4" captionStartId="6.[151,250,892,914]" captionTargetBox="[151,1436,193,870]" captionTargetId="figure@6.[151,1436,193,871]" captionTargetPageId="6" captionText="FIGURE 4. Lateral and ventral views of heads of Blaesodactylus spp. A. B. microtuberculatus sp. nov. (KUZ 069431). B. B. boivini (KUZ 069429). C. B. sakalava (KUZ 069432). D. B. ambonihazo (KUZ 069433)." httpUri="https://zenodo.org/record/236580/files/figure.png" pageId="4" pageNumber="410">Fig. 4</figureCitation>
|
||
;
|
||
<emphasis id="FFF39A1B690EFFDAFC44F19EC0BCFD76" box="[911,982,693,716]" italics="true" pageId="4" pageNumber="410">versus</emphasis>
|
||
pale).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="99F81681690EFFDAFF5CF422C245F8E6" httpUri="https://zenodo.org/record/236578/files/figure.png" pageId="4" pageNumber="410" targetBox="[174,1410,777,1766]" targetPageId="4">
|
||
<paragraph id="CD384609690EFFDAFF5CF422C245F8E6" blockId="4.[151,1436,1801,1884]" pageId="4" pageNumber="410">
|
||
<emphasis id="FFF39A1B690EFFDAFF5CF422C272F8A5" bold="true" box="[151,280,1801,1823]" pageId="4" pageNumber="410">FIGURE 2.</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="0A873D8A690EFFDAFE67F422C063F8A5" box="[428,777,1801,1823]" class="Reptilia" family="Gekkonidae" genus="Blaesodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="410" phylum="Chordata" rank="species" species="microtuberculatus" status="sp. nov.">
|
||
<emphasis id="FFF39A1B690EFFDAFE67F422C063F8A5" box="[428,777,1801,1823]" italics="true" pageId="4" pageNumber="410">Blaesodactylus microtuberculatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E4C02760690EFFDAFCDFF422C003F8A5" box="[788,873,1801,1823]" pageId="4" pageNumber="410" rank="species">
|
||
<emphasis id="FFF39A1B690EFFDAFCDFF422C003F8A5" bold="true" box="[788,873,1801,1823]" pageId="4" pageNumber="410">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and representatives of three of its congeners. A.
|
||
<taxonomicName id="0A873D8A690EFFDAFA4DF421C225F887" class="Reptilia" family="Gekkonidae" genus="Blaesodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="410" phylum="Chordata" rank="species" species="microtuberculatus" status="sp. nov.">
|
||
<emphasis id="FFF39A1B690EFFDAFA4DF421C225F887" italics="true" pageId="4" pageNumber="410">B. microtuberculatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E4C02760690EFFDAFE91F40CC2C5F887" box="[346,431,1831,1853]" pageId="4" pageNumber="410" rank="species">
|
||
<emphasis id="FFF39A1B690EFFDAFE91F40CC2C5F887" bold="true" box="[346,431,1831,1853]" pageId="4" pageNumber="410">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(KUZ069431). B.
|
||
<taxonomicName id="0A873D8A690EFFDAFDB5F402C18FF887" box="[638,741,1831,1854]" class="Reptilia" family="Gekkonidae" genus="Blaesodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="410" phylum="Chordata" rank="species" species="boivini">
|
||
<emphasis id="FFF39A1B690EFFDAFDB5F402C18FF887" box="[638,741,1831,1854]" italics="true" pageId="4" pageNumber="410">B. boivini</emphasis>
|
||
</taxonomicName>
|
||
(KUZ069429). C.
|
||
<taxonomicName id="0A873D8A690EFFDAFC7FF402C745F887" box="[948,1071,1831,1854]" class="Reptilia" family="Gekkonidae" genus="Blaesodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="410" phylum="Chordata" rank="species" species="sakalava">
|
||
<emphasis id="FFF39A1B690EFFDAFC7FF402C745F887" box="[948,1071,1831,1854]" italics="true" pageId="4" pageNumber="410">B. sakalava</emphasis>
|
||
</taxonomicName>
|
||
(KUZ069432). D.
|
||
<taxonomicName id="0A873D8A690EFFDAFB34F402C6F6F887" box="[1279,1436,1831,1854]" class="Reptilia" family="Gekkonidae" genus="Blaesodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="410" phylum="Chordata" rank="species" species="ambonihazo">
|
||
<emphasis id="FFF39A1B690EFFDAFB34F402C6F6F887" box="[1279,1436,1831,1854]" italics="true" pageId="4" pageNumber="410">B. ambonihazo</emphasis>
|
||
</taxonomicName>
|
||
(KUZ069433).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="859D1582690EFFD8FF0CF4A3C101F85B" lastPageId="6" lastPageNumber="412" pageId="4" pageNumber="410" type="description">
|
||
<paragraph id="CD384609690EFFDBFF0CF4A3C104FCAF" blockId="4.[151,1436,1928,2026]" lastBlockId="5.[151,1436,151,789]" lastPageId="5" lastPageNumber="411" pageId="4" pageNumber="410">
|
||
<emphasis id="FFF39A1B690EFFDAFF0CF4A3C160F81B" bold="true" box="[199,522,1928,1953]" pageId="4" pageNumber="410">
|
||
Description of the
|
||
<typeStatus id="123CF8AB690EFFDAFE68F4A2C160F81B" box="[419,522,1929,1953]" pageId="4" pageNumber="410" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
. KUZ069431, adult male. SVL
|
||
<quantity id="0A7FEBEC690EFFDAFCB8F4A2C08CF81B" box="[883,998,1929,1953]" metricMagnitude="-1" metricUnit="m" metricValue="1.174" pageId="4" pageNumber="410" unit="mm" value="117.4">117.4 mm</quantity>
|
||
, TailL
|
||
<quantity id="0A7FEBEC690EFFDAFBFEF4A2C7CDF81B" box="[1077,1191,1929,1953]" metricMagnitude="-1" metricUnit="m" metricValue="1.3419999999999999" pageId="4" pageNumber="410" unit="mm" value="134.2">134.2 mm</quantity>
|
||
(regenerated). Part of liver removed through mid-flank for tissue sample. Head relatively long (HeadL =
|
||
<quantity id="0A7FEBEC690EFFDAFB8EF487C7C7F87F" box="[1093,1197,1964,1989]" metricMagnitude="-2" metricUnit="m" metricValue="3.06" pageId="4" pageNumber="410" unit="mm" value="30.6">30.6 mm</quantity>
|
||
; HeadL/SVL = 0.26) and wide (HeadW =
|
||
<quantity id="0A7FEBEC690EFFDAFE45F4FAC293F853" box="[398,505,2001,2026]" metricMagnitude="-2" metricUnit="m" metricValue="2.36" pageId="4" pageNumber="410" unit="mm" value="23.6">23.6 mm</quantity>
|
||
; HeadW/HeadL = 0.77), depressed (HeadD =
|
||
<quantity id="0A7FEBEC690EFFDAFBD2F4FAC7E9F853" box="[1049,1155,2001,2025]" metricMagnitude="-2" metricUnit="m" metricValue="1.27" pageId="4" pageNumber="410" unit="mm" value="12.7">12.7 mm</quantity>
|
||
; HeadD/HeadL = 0.42); obviously broader than neck. Frontonasal region concave, snout elongate (SnEye =
|
||
<quantity id="0A7FEBEC690FFFDBFBA1F3B3C7BFFF15" box="[1130,1237,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="1.49" pageId="5" pageNumber="411" unit="mm" value="14.9">14.9 mm</quantity>
|
||
; SnEye/HeadL = 0.49), blunt, longer than eye diameter (OrbD =
|
||
<quantity id="0A7FEBEC690FFFDBFD19F397C045FF6E" box="[722,815,188,213]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="5" pageNumber="411" unit="mm" value="6.1">6.1 mm</quantity>
|
||
; OrbD/SnEye = 0.41); internarial distance
|
||
<quantity id="0A7FEBEC690FFFDBFAE6F397C6F2FF6E" box="[1325,1432,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="1.1099999999999999" pageId="5" pageNumber="411" unit="mm" value="11.1">11.1 mm</quantity>
|
||
; interorbital distance
|
||
<quantity id="0A7FEBEC690FFFDBFE4FF3F4C28AFF4D" box="[388,480,223,248]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="5" pageNumber="411" unit="mm" value="4.6">4.6 mm</quantity>
|
||
; scales on snout and forehead small, granular, and heterogeneous; scales on snout larger than those on occipital region except for scattered conical tubercles (~3 times size of adjacent scales); 24 scales across narrowest point of frontals, 52 between superciliary scale rows. Eye large (OrbD/HeadL = 0.20); pupil vertical with crenelated margins; superciliaries forming a short brillar fold with small spines at anterior and posterior margins. Ear opening obliquely oval, large (EarL =
|
||
<quantity id="0A7FEBEC690FFFDBFC95F25BC0D0FE3D" box="[862,954,368,392]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="5" pageNumber="411" unit="mm" value="3.8">3.8 mm</quantity>
|
||
; EarL/HeadL = 0.12); eye to ear distance longer than diameter of eye (EyeEar =
|
||
<quantity id="0A7FEBEC690FFFDBFD80F2BFC1CEFE16" box="[587,676,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="5" pageNumber="411" unit="mm" value="9.3">9.3 mm</quantity>
|
||
; EyeEar/OrbD = 1.54). Rostral quadrangular, much wider (
|
||
<quantity id="0A7FEBEC690FFFDBFAF7F2BFC6FEFE16" box="[1340,1428,404,429]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="5" pageNumber="411" unit="mm" value="5.2">5.2 mm</quantity>
|
||
) than high (
|
||
<quantity id="0A7FEBEC690FFFDBFEDAF29CC207FE6A" box="[273,365,439,464]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="411" unit="mm" value="2.5">2.5 mm</quantity>
|
||
), with no median groove. Enlarged supranasals separated by two internasal scales, one smaller scale (same size of snout granules) anteriorly, bordering rostral, and one larger scale (twice size of snout granules) posteriorly; rostral in contact with first supralabials, supranasals, and one internasal; nostrils round, each surrounded by supranasal, rostral, first supralabial, and a crescentic nasal, itself bordered posteriorly by 4 postnasal scales; 3–4 rows of small scales separate rim of orbit from supralabials. Mental pentagonal, wider (
|
||
<quantity id="0A7FEBEC690FFFDBFACAF163C631FDDA" box="[1281,1371,584,608]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="5" pageNumber="411" unit="mm" value="4.2">4.2 mm</quantity>
|
||
) than deep (
|
||
<quantity id="0A7FEBEC690FFFDBFF15F147C256FD3E" box="[222,316,620,644]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="5" pageNumber="411" unit="mm" value="2.7">2.7 mm</quantity>
|
||
); median pair of postmentals elongated (
|
||
<quantity id="0A7FEBEC690FFFDBFCDDF147C01AFD3E" box="[790,880,620,644]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="5" pageNumber="411" unit="mm" value="3.7">3.7 mm</quantity>
|
||
long), each bordered anteromedially by mental, medially in broad contact with other postmental along most of their entire length, but separated anteriorly by posterior tip of mental, and bordered anterolaterally by first infralabial, laterally by second postmental, posteriorly by 3 slightly enlarged chin scales grading posteriorly and laterally into smaller gular scales; 14 (right) to 13 (left) supralabials, 12 infralabials on both sides.
|
||
</paragraph>
|
||
<caption id="99F81681690FFFDBFF5CF4B7C6FEF84A" httpUri="https://zenodo.org/record/236579/files/figure.png" pageId="5" pageNumber="411" targetBox="[172,1415,848,1926]" targetPageId="5">
|
||
<paragraph id="CD384609690FFFDBFF5CF4B7C6FEF84A" blockId="5.[151,1436,1948,2032]" pageId="5" pageNumber="411">
|
||
<emphasis id="FFF39A1B690FFFDBFF5CF4B7C27BF808" bold="true" box="[151,273,1948,1970]" pageId="5" pageNumber="411">FIGURE 3.</emphasis>
|
||
Localities and habitat of two allotopic
|
||
<taxonomicName id="0A873D8A690FFFDBFD52F4B7C05AF808" box="[665,816,1948,1970]" class="Reptilia" family="Gekkonidae" genus="Blaesodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="411" phylum="Chordata" rank="genus">
|
||
<emphasis id="FFF39A1B690FFFDBFD52F4B7C05AF808" box="[665,816,1948,1970]" italics="true" pageId="5" pageNumber="411">Blaesodactylus</emphasis>
|
||
</taxonomicName>
|
||
spp. A. Map of Ankarana National Park. Note that the Tsingy karstic outcroppings are inside the forest. Open circle indicates the site where
|
||
<taxonomicName id="0A873D8A690FFFDBFC02F497C7F7F86B" box="[969,1181,1979,2001]" class="Reptilia" family="Gekkonidae" genus="Blaesodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="411" phylum="Chordata" rank="species" species="microtuberculatus" status="sp. nov.">
|
||
<emphasis id="FFF39A1B690FFFDBFC02F497C0B2F86B" box="[969,984,1980,2001]" italics="true" pageId="5" pageNumber="411">B</emphasis>
|
||
.
|
||
<emphasis id="FFF39A1B690FFFDBFC23F490C7F7F86B" box="[1000,1181,1979,2001]" italics="true" pageId="5" pageNumber="411">microtuberculatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E4C02760690FFFDBFB6CF490C790F86B" box="[1191,1274,1979,2001]" pageId="5" pageNumber="411" rank="species">
|
||
<emphasis id="FFF39A1B690FFFDBFB6CF490C790F86B" bold="true" box="[1191,1274,1979,2001]" pageId="5" pageNumber="411">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
was found and closed circles indicate the sites where
|
||
<taxonomicName id="0A873D8A690FFFDBFDDFF4F0C11FF84A" box="[532,629,2010,2032]" class="Reptilia" family="Gekkonidae" genus="Blaesodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="411" phylum="Chordata" rank="species" species="boivini">
|
||
<emphasis id="FFF39A1B690FFFDBFDDFF4F0C149F84A" box="[532,547,2011,2032]" italics="true" pageId="5" pageNumber="411">B</emphasis>
|
||
.
|
||
<emphasis id="FFF39A1B690FFFDBFDFBF4F1C11FF84A" box="[560,629,2010,2032]" italics="true" pageId="5" pageNumber="411">boivini</emphasis>
|
||
</taxonomicName>
|
||
was found. B. Habitat of
|
||
<taxonomicName id="0A873D8A690FFFDBFCB3F4F0C723F84A" box="[888,1097,2010,2032]" class="Reptilia" family="Gekkonidae" genus="Blaesodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="411" phylum="Chordata" rank="species" species="microtuberculatus" status="sp. nov.">
|
||
<emphasis id="FFF39A1B690FFFDBFCB3F4F0C723F84A" box="[888,1097,2010,2032]" italics="true" pageId="5" pageNumber="411">B. microtuberculatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E4C02760690FFFDBFB84F4F1C7F5F84A" box="[1103,1183,2010,2032]" pageId="5" pageNumber="411" rank="species">
|
||
<emphasis id="FFF39A1B690FFFDBFB84F4F1C7F5F84A" bold="true" box="[1103,1183,2010,2032]" pageId="5" pageNumber="411">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
C. Habitat of
|
||
<taxonomicName id="0A873D8A690FFFDBFAE6F4F0C6E4F84A" box="[1325,1422,2010,2032]" class="Reptilia" family="Gekkonidae" genus="Blaesodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="411" phylum="Chordata" rank="species" species="boivini">
|
||
<emphasis id="FFF39A1B690FFFDBFAE6F4F0C6E4F84A" box="[1325,1422,2010,2032]" italics="true" pageId="5" pageNumber="411">B. boivini</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="99F81681690CFFD8FF5CF057C767FC0B" httpUri="https://zenodo.org/record/236580/files/figure.png" pageId="6" pageNumber="412" targetBox="[151,1436,193,870]" targetPageId="6">
|
||
<paragraph id="CD384609690CFFD8FF5CF057C767FC0B" blockId="6.[151,1436,892,945]" pageId="6" pageNumber="412">
|
||
<emphasis id="FFF39A1B690CFFD8FF5CF057C27EFC28" bold="true" box="[151,276,892,914]" pageId="6" pageNumber="412">FIGURE 4.</emphasis>
|
||
Lateral and ventral views of heads of
|
||
<taxonomicName id="0A873D8A690CFFD8FD6AF057C050FC28" box="[673,826,892,914]" class="Reptilia" family="Gekkonidae" genus="Blaesodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="412" phylum="Chordata" rank="genus">
|
||
<emphasis id="FFF39A1B690CFFD8FD6AF057C050FC28" box="[673,826,892,914]" italics="true" pageId="6" pageNumber="412">Blaesodactylus</emphasis>
|
||
</taxonomicName>
|
||
spp. A.
|
||
<taxonomicName id="0A873D8A690CFFD8FC58F056C702FC28" box="[915,1128,892,914]" class="Reptilia" family="Gekkonidae" genus="Blaesodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="412" phylum="Chordata" rank="species" species="microtuberculatus" status="sp. nov.">
|
||
<emphasis id="FFF39A1B690CFFD8FC58F056C702FC28" box="[915,1128,892,914]" italics="true" pageId="6" pageNumber="412">B. microtuberculatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E4C02760690CFFD8FBA4F057C7ABFC28" box="[1135,1217,892,914]" pageId="6" pageNumber="412" rank="species">
|
||
<emphasis id="FFF39A1B690CFFD8FBA4F057C7ABFC28" bold="true" box="[1135,1217,892,914]" pageId="6" pageNumber="412">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(KUZ069431). B.
|
||
<taxonomicName id="0A873D8A690CFFD8FA4DF056C3B7FC0B" class="Reptilia" family="Gekkonidae" genus="Blaesodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="412" phylum="Chordata" rank="species" species="boivini">
|
||
<emphasis id="FFF39A1B690CFFD8FA4DF056C3B7FC0B" italics="true" pageId="6" pageNumber="412">B. boivini</emphasis>
|
||
</taxonomicName>
|
||
(KUZ069429). C.
|
||
<taxonomicName id="0A873D8A690CFFD8FE55F0B7C17FFC0B" box="[414,533,923,945]" class="Reptilia" family="Gekkonidae" genus="Blaesodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="412" phylum="Chordata" rank="species" species="sakalava">
|
||
<emphasis id="FFF39A1B690CFFD8FE55F0B7C17FFC0B" box="[414,533,923,945]" italics="true" pageId="6" pageNumber="412">B. sakalava</emphasis>
|
||
</taxonomicName>
|
||
(KUZ069432). D.
|
||
<taxonomicName id="0A873D8A690CFFD8FD1CF0B7C01AFC0B" box="[727,880,923,945]" class="Reptilia" family="Gekkonidae" genus="Blaesodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="412" phylum="Chordata" rank="species" species="ambonihazo">
|
||
<emphasis id="FFF39A1B690CFFD8FD1CF0B7C01AFC0B" box="[727,880,923,945]" italics="true" pageId="6" pageNumber="412">B. ambonihazo</emphasis>
|
||
</taxonomicName>
|
||
(KUZ069433).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="CD384609690CFFD8FF0CF0F3C245FA8F" blockId="6.[151,1437,984,2017]" pageId="6" pageNumber="412">
|
||
Body stout, relatively short (TrunkL =
|
||
<quantity id="0A7FEBEC690CFFD8FDB0F0F2C1B1FC4B" box="[635,731,985,1009]" metricMagnitude="-2" metricUnit="m" metricValue="4.72" pageId="6" pageNumber="412" unit="mm" value="47.2">47.2mm</quantity>
|
||
; TrunkL/SVL = 0.40) with moderately developed ventrolateral folds. Dorsal scales smooth, granular to conical and recumbent, juxtaposed; 162 scale rows around midbody, intermixed with enlarged, smooth rounded tubercles (~3 times size of adjacent scales, largest on flanks, and smallest in occipital region), extending from occipital regions to tail base; tubercles in 15–17 rows at midbody. Ventral scales much larger than dorsals, smooth, imbricate, free margin rounded, largest on abdomen except for rows immediately anterior to vent; 46 scale rows across venter between ventrolateral folds; gular region with relatively homogeneous, smooth, rounded to oval scales, juxtaposed anteriorly to subimbricate posteriorly. No precloacal or femoral pores, no enlarged femoral scales. Preaxial scalation of limbs subimbricate, postaxial granular, dorsal surface of limbs without tubercles. Scales of palms and soles smooth, flattened, round, subimbricate.
|
||
</paragraph>
|
||
<paragraph id="CD384609690CFFD8FF0CF66BC7D5F9B6" blockId="6.[151,1437,984,2017]" pageId="6" pageNumber="412">
|
||
Limbs short and robust (ForeaL =
|
||
<quantity id="0A7FEBEC690CFFD8FD8EF66AC1C0FAE2" box="[581,682,1345,1369]" metricMagnitude="-2" metricUnit="m" metricValue="1.13" pageId="6" pageNumber="412" unit="mm" value="11.3">11.3 mm</quantity>
|
||
; ForeaL/SVL = 0.10; CrusL =
|
||
<quantity id="0A7FEBEC690CFFD8FBCBF66AC70CFAE2" box="[1024,1126,1345,1369]" metricMagnitude="-2" metricUnit="m" metricValue="2.09" pageId="6" pageNumber="412" unit="mm" value="20.9">20.9 mm</quantity>
|
||
; CrusL/SVL = 0.18). Digits broadly dilated, distal portion of digits II–V free of pad, bearing a prominent recurved claw partly sheathed between a pair of scales, distal portion of digit I not free of pad, claw minute and lying in a groove in the adhesive pad; number of broad lamellae beneath each digit (
|
||
<date id="B93960C9690CFFD8FD14F687C010FA7F" box="[735,890,1452,1477]" pageId="6" pageNumber="412" value="1921-10-17" valueMax="1921-10-21" valueMin="1921-10-17">10–17–21–21</date>
|
||
–15 manus; 15–14–17–21–17 pes); all lamellae, except distalmost slightly bowed, undivided except for distalmost lamellae of digit I; interdigital webbing weakly developed. Relative length of digits (manus): IV> III> III> II>V> I; (pes): IV>III> V> II> I.
|
||
</paragraph>
|
||
<paragraph id="CD384609690CFFD8FF0CF533C61EF8B3" blockId="6.[151,1437,984,2017]" pageId="6" pageNumber="412">Tail slender, approximately one-third length of whole tail regenerated, tapering to tip; somewhat longer than snout-vent length (TailL/SVL = 1.14), dorso-ventrally depressed. Tail base with 4 (right) and 3 (left) smooth cloacal spurs on each side, largest anteriormost. Scales of tail dorsum heterogeneous — rectangular to pentagonal or hexagonal, subimbricate; midventral subcaudal scales transversely enlarged and about half of tail width; tail clearly verticillate, each segment 8 dorsal scale rows and 3 transversely enlarged subcaudal scales in length; posterior edge of each segment without a row of enlarged tubercles. Regenerated portion of tail with some segmentation evident but scalation irregular. Hemipenes partly everted, but detailed morphology unobservable.</paragraph>
|
||
<paragraph id="CD384609690CFFD8FF0CF43FC101F85B" blockId="6.[151,1437,984,2017]" pageId="6" pageNumber="412">Coloration (in preservative). Body dorsum grayish without a series of crossbars, mid-dorsal row of five lighter color blotches, indistinctly separated from each other along the dorsal midline. Irregular dark vermiculation on dorsum. Flanks with some faint mottling. Limbs mottled. Tail with faint alternating dark-gray and light-gray bands. Head medium gray with beige labial scales, rictus, and posteroventral orbital margin. Venter cream anteriorly and light gray posteriorly; chin with very faint mottling. Palms and soles grayish. Coloration in life generally similar to that in preservative, but more contrasting.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="859D1582690DFFD9FF0CF3BCC0DFFE75" pageId="7" pageNumber="413" type="distribution">
|
||
<paragraph id="CD384609690DFFD9FF0CF3BCC3A3FF6E" blockId="7.[151,1436,151,464]" pageId="7" pageNumber="413">
|
||
<emphasis id="FFF39A1B690DFFD9FF0CF3BCC20AFF0A" bold="true" box="[199,352,151,176]" pageId="7" pageNumber="413">Distribution.</emphasis>
|
||
This species is currently known only from Ankarana National Park in northern
|
||
<collectingCountry id="B5900699690DFFD9FB13F3BCC635FF0A" box="[1240,1375,151,176]" name="Madagascar" pageId="7" pageNumber="413">Madagascar</collectingCountry>
|
||
(
|
||
<figureCitation id="55BC5A8C690DFFD9FAA5F3BCC3CFFF6E" captionStart="FIGURE 3" captionStartId="5.[151,249,1948,1970]" captionTargetBox="[172,1415,848,1926]" captionTargetId="figure@5.[152,1435,841,1927]" captionTargetPageId="5" captionText="FIGURE 3. Localities and habitat of two allotopic Blaesodactylus spp. A. Map of Ankarana National Park. Note that the Tsingy karstic outcroppings are inside the forest. Open circle indicates the site where B. microtuberculatus sp. nov. was found and closed circles indicate the sites where B. boivini was found. B. Habitat of B. microtuberculatus sp. nov. C. Habitat of B. boivini." httpUri="https://zenodo.org/record/236579/files/figure.png" pageId="7" pageNumber="413">Fig. 3</figureCitation>
|
||
A).
|
||
</paragraph>
|
||
<paragraph id="CD384609690DFFD9FF0CF3CBC0DFFE75" blockId="7.[151,1436,151,464]" pageId="7" pageNumber="413">
|
||
<emphasis id="FFF39A1B690DFFD9FF0CF3CBC2E3FF42" bold="true" box="[199,393,223,248]" pageId="7" pageNumber="413">Natural history.</emphasis>
|
||
The specimen was collected in Tsingy karstic outcropping surrounded by dry deciduous forest in Ankarana National Park at 22:19 on
|
||
<date id="B93960C9690DFFD9FD50F22FC00EFEA6" box="[667,868,260,285]" pageId="7" pageNumber="413" value="2012-12-04">4 December 2012</date>
|
||
(
|
||
<figureCitation id="55BC5A8C690DFFD9FCBFF22FC0DDFEA6" box="[884,951,260,285]" captionStart="FIGURE 3" captionStartId="5.[151,249,1948,1970]" captionTargetBox="[172,1415,848,1926]" captionTargetId="figure@5.[152,1435,841,1927]" captionTargetPageId="5" captionText="FIGURE 3. Localities and habitat of two allotopic Blaesodactylus spp. A. Map of Ankarana National Park. Note that the Tsingy karstic outcroppings are inside the forest. Open circle indicates the site where B. microtuberculatus sp. nov. was found and closed circles indicate the sites where B. boivini was found. B. Habitat of B. microtuberculatus sp. nov. C. Habitat of B. boivini." httpUri="https://zenodo.org/record/236579/files/figure.png" pageId="7" pageNumber="413">Fig. 3</figureCitation>
|
||
A). It perched motionless on the surface of an outcrop (
|
||
<figureCitation id="55BC5A8C690DFFD9FEFBF20CC216FEFA" box="[304,380,295,320]" captionStart="FIGURE 3" captionStartId="5.[151,249,1948,1970]" captionTargetBox="[172,1415,848,1926]" captionTargetId="figure@5.[152,1435,841,1927]" captionTargetPageId="5" captionText="FIGURE 3. Localities and habitat of two allotopic Blaesodactylus spp. A. Map of Ankarana National Park. Note that the Tsingy karstic outcroppings are inside the forest. Open circle indicates the site where B. microtuberculatus sp. nov. was found and closed circles indicate the sites where B. boivini was found. B. Habitat of B. microtuberculatus sp. nov. C. Habitat of B. boivini." httpUri="https://zenodo.org/record/236579/files/figure.png" pageId="7" pageNumber="413">Fig. 3</figureCitation>
|
||
B). We were unable to capture two additional individuals, similar in appearance and morphological characteristics, which were found in a similar posture on limestones in the karstic outcropping. In the deciduous forest of this park (
|
||
<figureCitation id="55BC5A8C690DFFD9FDC5F244C13AFE32" box="[526,592,367,392]" captionStart="FIGURE 3" captionStartId="5.[151,249,1948,1970]" captionTargetBox="[172,1415,848,1926]" captionTargetId="figure@5.[152,1435,841,1927]" captionTargetPageId="5" captionText="FIGURE 3. Localities and habitat of two allotopic Blaesodactylus spp. A. Map of Ankarana National Park. Note that the Tsingy karstic outcroppings are inside the forest. Open circle indicates the site where B. microtuberculatus sp. nov. was found and closed circles indicate the sites where B. boivini was found. B. Habitat of B. microtuberculatus sp. nov. C. Habitat of B. boivini." httpUri="https://zenodo.org/record/236579/files/figure.png" pageId="7" pageNumber="413">Fig. 3</figureCitation>
|
||
C), we found 16 individuals of
|
||
<taxonomicName id="0A873D8A690DFFD9FC65F25AC771FE32" box="[942,1051,368,392]" class="Reptilia" family="Gekkonidae" genus="Blaesodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="413" phylum="Chordata" rank="species" species="boivini">
|
||
<emphasis id="FFF39A1B690DFFD9FC65F25AC771FE32" box="[942,1051,368,392]" italics="true" pageId="7" pageNumber="413">B. boivini</emphasis>
|
||
</taxonomicName>
|
||
on tree trunks both in the daytime and at night, but no geckos similar to
|
||
<taxonomicName id="0A873D8A690DFFD9FD90F2BEC022FE16" box="[603,840,404,428]" class="Reptilia" family="Gekkonidae" genus="Blaesodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="413" phylum="Chordata" rank="species" species="microtuberculatus">
|
||
<emphasis id="FFF39A1B690DFFD9FD90F2BEC022FE16" box="[603,840,404,428]" italics="true" pageId="7" pageNumber="413">B. microtuberculatus</emphasis>
|
||
</taxonomicName>
|
||
were found in this habitat. The minimum distance between the locations of
|
||
<taxonomicName id="0A873D8A690DFFD9FE64F292C1F3FE6A" box="[431,665,440,464]" class="Reptilia" family="Gekkonidae" genus="Blaesodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="413" phylum="Chordata" rank="species" species="microtuberculatus">
|
||
<emphasis id="FFF39A1B690DFFD9FE64F292C2AAFE6A" box="[431,448,441,464]" italics="true" pageId="7" pageNumber="413">B</emphasis>
|
||
.
|
||
<emphasis id="FFF39A1B690DFFD9FE05F293C1F3FE6A" box="[462,665,440,464]" italics="true" pageId="7" pageNumber="413">microtuberculatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="0A873D8A690DFFD9FD04F292C056FE6A" box="[719,828,440,464]" class="Reptilia" family="Gekkonidae" genus="Blaesodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="413" phylum="Chordata" rank="species" species="boivini">
|
||
<emphasis id="FFF39A1B690DFFD9FD04F292C18AFE6A" box="[719,736,441,464]" italics="true" pageId="7" pageNumber="413">B</emphasis>
|
||
.
|
||
<emphasis id="FFF39A1B690DFFD9FD25F293C056FE6A" box="[750,828,440,464]" italics="true" pageId="7" pageNumber="413">boivini</emphasis>
|
||
</taxonomicName>
|
||
was
|
||
<specimenCount id="DB818D80690DFFD9FCBDF293C0DFFE75" box="[886,949,440,464]" pageId="7" pageNumber="413" type="generic">
|
||
<quantity id="0A7FEBEC690DFFD9FCBDF293C0DBFE75" box="[886,945,440,464]" metricMagnitude="1" metricUnit="m" metricValue="1.1" pageId="7" pageNumber="413" unit="m" value="11.0">11 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |