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<document id="74B74DDFC6B366BF9453561F356B434A" ID-DOI="10.5281/zenodo.276270" ID-GBIF-Dataset="91374710-277e-40a4-8cdf-281fe24a7132" ID-ISSN="1175-5326" ID-Zenodo-Dep="276270" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460321189885" checkinUser="plazi" docAuthor="Stüben, Peter E. &amp; Astrin, Jonas J." docDate="2010" docId="45154932A918FD0BFF6129DE2526FCDE" docLanguage="en" docName="zt02662p052.pdf" docOrigin="Zootaxa 2662" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Kyklioacalles atlasicus Stuben &amp; Astrin, sp.n." docType="treatment" docVersion="6" lastPageNumber="44" masterDocId="B92C314AA908FD1BFFF629492602FFFA" masterDocTitle="Molecular phylogeny of the weevil genus Kyklioacalles Stüben, with descriptions of a new subgenus Glaberacalles and two new species (Curculionidae: Cryptorhynchinae)" masterLastPageNumber="52" masterPageNumber="28" pageNumber="44" updateTime="1698256752306" updateUser="plazi">
<mods:mods id="2A0397008A12184D093453014A273A86" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C765A087BCD87D4BD204C2ED079AFE9F">
<mods:title id="D92D771748EFAAB35B9E8D546F9D7734">Molecular phylogeny of the weevil genus Kyklioacalles Stüben, with descriptions of a new subgenus Glaberacalles and two new species (Curculionidae: Cryptorhynchinae)</mods:title>
</mods:titleInfo>
<mods:name id="ED6694874F1E7A10C1456D5B27D902B7" type="personal">
<mods:role id="4DD8A00FC64AE344B07AD0A0B6DE25E2">
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</mods:role>
<mods:namePart id="4C6357DE409085DAE02F6FBC2F304C4C">Stüben, Peter E.</mods:namePart>
</mods:name>
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<mods:roleTerm id="BE19A9B0BA0C46CA4951F53E8B43F714">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5551761EE3327C99FA29B4933F8F44D3">Astrin, Jonas J.</mods:namePart>
</mods:name>
<mods:typeOfResource id="02FDF1E7AD71BB9D80F8D3084310FA4D">text</mods:typeOfResource>
<mods:relatedItem id="80E3E811E43502544855A6484D1E665D" type="host">
<mods:titleInfo id="A48DB31EE092EB448CBB6D36835A1638">
<mods:title id="F4738D5ABC7C40C84F1EDFF32591FD0F">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="2B36593DED05CDB80172C68533A0557E">
<mods:date id="FA0AC5FD8B54AA65F5EC6945E596D057">2010</mods:date>
<mods:detail id="A4A664991F350F7A36EB2291CDA5D278" type="volume">
<mods:number id="4258B9F7717D8AB172C7156522A939BE">2662</mods:number>
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<mods:start id="087526E8ED061FB17398F4D36939D3AD">28</mods:start>
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</mods:extent>
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<mods:classification id="F07C320B1E151B335B804BD1BEF3AA5B">journal article</mods:classification>
<mods:identifier id="C27EC65AA2F65CF3B869E104BC7844F4" type="DOI">10.5281/zenodo.276270</mods:identifier>
<mods:identifier id="FDF8D76119AC4CFEC14336549E4CBFDD" type="GBIF-Dataset">91374710-277e-40a4-8cdf-281fe24a7132</mods:identifier>
<mods:identifier id="15FDCC938EC7561541A8A6BACB735EBE" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="3F23CF32AF5E76E49BC26271203CF45C" type="Zenodo-Dep">276270</mods:identifier>
</mods:mods>
<treatment id="45154932A918FD0BFF6129DE2526FCDE" ID-DOI="http://doi.org/10.5281/zenodo.6212166" ID-GBIF-Taxon="119402537" ID-Zenodo-Dep="6212166" LSID="urn:lsid:plazi:treatment:45154932A918FD0BFF6129DE2526FCDE" httpUri="http://treatment.plazi.org/id/45154932A918FD0BFF6129DE2526FCDE" lastPageNumber="44" pageId="16" pageNumber="44">
<subSubSection id="85A6ABAFA918FD0BFF6129DE278EFF2E" pageId="16" pageNumber="44" type="nomenclature">
<paragraph id="CD03F824A918FD0BFF6129DE24C2FF4B" blockId="16.[151,704,151,212]" box="[151,704,151,177]" pageId="16" pageNumber="44">
<heading id="964B4F48A918FD0BFF6129DE24C2FF4B" bold="true" box="[151,704,151,177]" fontSize="11" level="1" pageId="16" pageNumber="44" reason="1">
<emphasis id="FFC82436A918FD0BFF6129DE24C2FF4B" bold="true" box="[151,704,151,177]" pageId="16" pageNumber="44">
<taxonomicName id="0ABC83A7A918FD0BFF6129DE247CFF4B" ID-CoL="3RGXK" authority="Stuben &amp; Astrin" authorityName="Stuben &amp; Astrin" box="[151,638,151,177]" class="Insecta" family="Curculionidae" genus="Kyklioacalles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="44" phylum="Arthropoda" rank="species" species="atlasicus" status="sp. nov.">
<emphasis id="FFC82436A918FD0BFF6129DE27ABFF4B" bold="true" box="[151,425,151,177]" italics="true" pageId="16" pageNumber="44">Kyklioacalles atlasicus</emphasis>
Stüben &amp; Astrin
</taxonomicName>
<taxonomicNameLabel id="E4FB994DA918FD0BFD7329DE24C2FF4B" box="[645,704,151,177]" pageId="16" pageNumber="44" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="CD03F824A918FD0BFF6129F3278EFF2E" blockId="16.[151,704,151,212]" box="[151,396,186,212]" pageId="16" pageNumber="44">
(
<figureCitation id="5587E4A1A918FD0BFF6929F32703FF2E" box="[159,257,186,212]" captionStart="FIGURES 20 – 21" captionStartId="13.[151,277,1842,1866]" captionTargetBox="[151,1436,271,1818]" captionTargetId="figure@13.[151,1436,271,1819]" captionTargetPageId="13" captionText="FIGURES 20 – 21. New species of Kyklioacalles (dorsal / lateral view); 22 – 24. Related species (dorsally): 22. Kyklioacalles bupleuri Stüben, 2004, 23. Kyklioacalles anthyllis Stüben, 2004, 24. Kyklioacalles maroccensis (Stüben, 2001)." httpUri="https://zenodo.org/record/276275/files/figure.png" pageId="16" pageNumber="44">Figs. 21</figureCitation>
,
<figureCitation id="5587E4A1A918FD0BFEF829F3272CFF2E" box="[270,302,186,212]" captionStart="FIGURES 25 – 29" captionStartId="14.[151,269,1845,1869]" captionTargetBox="[153,1433,194,1823]" captionTargetId="figure@14.[153,1434,194,1823]" captionTargetPageId="14" captionText="FIGURES 25 – 29. Male genitalia of the new and related species; 30. Loci typici and host plants of the new species; 31 – 32. Female genitalia." httpUri="https://zenodo.org/record/276276/files/figure.png" pageId="16" pageNumber="44">28</figureCitation>
, 30, 32)
</paragraph>
</subSubSection>
<subSubSection id="85A6ABAFA918FD0BFF6129B622CBFDE5" pageId="16" pageNumber="44" type="materials_examined">
<paragraph id="CD03F824A918FD0BFF6129B624F6FE9E" blockId="16.[151,1437,255,804]" pageId="16" pageNumber="44">
<emphasis id="FFC82436A918FD0BFF6129B627F6FEE3" bold="true" box="[151,500,255,281]" pageId="16" pageNumber="44">
<typeStatus id="12074686A918FD0BFF6129B626DBFEE3" box="[151,217,255,281]" pageId="16" pageNumber="44">Type</typeStatus>
material.
<typeStatus id="12074686A918FD0BFE9A29B627ECFEE3" box="[364,494,255,281]" pageId="16" pageNumber="44" type="holotype">Holotype</typeStatus>
.
</emphasis>
1ɗ: &quot;
<collectingCountry id="B5ABB8B4A918FD0BFDC9284924EBFEE0" box="[575,745,256,282]" name="Morocco" pageId="16" pageNumber="44">MOROCCO</collectingCountry>
: High Atlas,
<quantity id="0A4455C1A918FD0BFC6E284925E9FEE0" box="[920,1003,256,282]" metricMagnitude="4" metricUnit="m" metricValue="5.9" pageId="16" pageNumber="44" unit="km" value="59.0">59 km</quantity>
SE Marrakech, N Taddert, (near Tazouguerte),
<geoCoordinate id="A8889EE3A918FD0BFEB6286C27CDFEC5" box="[320,463,293,319]" direction="north" orientation="latitude" pageId="16" pageNumber="44" precision="15" value="31.471945">N31°28’19”</geoCoordinate>
<geoCoordinate id="A8889EE3A918FD0BFE2E286C2473FEC5" box="[472,625,293,319]" direction="west" orientation="longitude" pageId="16" pageNumber="44" precision="15" value="-7.4061112">W07°24’22”</geoCoordinate>
,
<quantity id="0A4455C1A918FD0BFD8B286C24D8FEC5" box="[637,730,293,319]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="16" pageNumber="44" unit="m" value="1500.0">1500 m</quantity>
,
<date id="B902DEE4A918FD0BFD10286C256FFEC5" box="[742,877,293,319]" pageId="16" pageNumber="44" value="2002-12-28">28.12.2002</date>
,
<emphasis id="FFC82436A918FD0BFC8F286C22CAFEC4" box="[889,1224,293,318]" italics="true" pageId="16" pageNumber="44">
<taxonomicName id="0ABC83A7A918FD0BFC8F286C2213FEC4" box="[889,1041,293,318]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="16" pageNumber="44" phylum="Tracheophyta" rank="species" species="ilex">Quercus ilex</taxonomicName>
,
<taxonomicName id="0ABC83A7A918FD0BFBEB286C22CAFEC4" box="[1053,1224,293,318]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="16" pageNumber="44" phylum="Tracheophyta" rank="species" species="suber">Quercus suber</taxonomicName>
</emphasis>
, leg. Stüben -19 - &quot;, coll. CURCULIO-Institut, D- Mönchengladbach.
</paragraph>
<paragraph id="CD03F824A918FD0BFF33282622CBFDE5" blockId="16.[151,1437,255,804]" pageId="16" pageNumber="44">
<emphasis id="FFC82436A918FD0BFF3328262749FE73" bold="true" box="[197,331,367,393]" pageId="16" pageNumber="44">
<typeStatus id="12074686A918FD0BFF3328262746FE73" box="[197,324,367,393]" pageId="16" pageNumber="44" type="paratype">Paratypes</typeStatus>
.
</emphasis>
58ɗ, 25Ψ: data as for
<typeStatus id="12074686A918FD0BFDAA283924C5FE70" box="[604,711,368,394]" pageId="16" pageNumber="44" type="holotype">holotype</typeStatus>
, coll. Stüben, coll. CURCULIO-Institute, Mönchengladbach (
<emphasis id="FFC82436A918FD0BFF5728DC26B2FE55" bold="true" box="[161,176,405,431]" pageId="16" pageNumber="44">1</emphasis>
Ψ); 2ɗ: &quot;
<collectingCountry id="B5ABB8B4A918FD0BFEEC28DC27BBFE55" box="[282,441,405,431]" name="Morocco" pageId="16" pageNumber="44">MOROCCO</collectingCountry>
: High Atlas, E Marrakech, N Taddert, (near Tazouguerte),
<quantity id="0A4455C1A918FD0BFB7628DC22DCFE55" box="[1152,1246,405,431]" metricMagnitude="3" metricUnit="m" metricValue="1.498" pageId="16" pageNumber="44" unit="m" value="1498.0">1498 m</quantity>
,
<geoCoordinate id="A8889EE3A918FD0BFB1D28DC237BFE55" box="[1259,1401,405,431]" direction="north" orientation="latitude" pageId="16" pageNumber="44" precision="15" value="31.471666">N31°28'18&quot;</geoCoordinate>
<geoCoordinate id="A8889EE3A918FD0BFA7628DC2717FE2E" direction="west" orientation="longitude" pageId="16" pageNumber="44" precision="15" value="-7.4058332">W 07°24'21&quot;</geoCoordinate>
,
<emphasis id="FFC82436A918FD0BFED528F327EEFE29" box="[291,492,442,467]" italics="true" pageId="16" pageNumber="44">
<taxonomicName id="0ABC83A7A918FD0BFED528F32788FE29" box="[291,394,442,467]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="16" pageNumber="44" phylum="Tracheophyta" rank="genus">Quercus</taxonomicName>
(
<taxonomicName id="0ABC83A7A918FD0BFE6A28F327E1FE29" box="[412,483,442,467]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="16" pageNumber="44" phylum="Tracheophyta" rank="species" species="suber">suber</taxonomicName>
)
</emphasis>
,
<date id="B902DEE4A918FD0BFE0B28F3246FFE2E" box="[509,621,442,468]" pageId="16" pageNumber="44" value="2009-05-03">3.5.2009</date>
, leg. Stüben -13-&quot;, coll. Stüben; 3ɗ: &quot;
<collectingCountry id="B5ABB8B4A918FD0BFBB528F322E7FE2E" box="[1091,1253,442,468]" name="Morocco" pageId="16" pageNumber="44">MOROCCO</collectingCountry>
: High Atlas, E Marrakech, N Taddert, (near Tazouguerte),
<quantity id="0A4455C1A918FD0BFD6228A924F2FE00" box="[660,752,480,506]" metricMagnitude="3" metricUnit="m" metricValue="1.7269999999999999" pageId="16" pageNumber="44" unit="m" value="1727.0">1727 m</quantity>
,
<geoCoordinate id="A8889EE3A918FD0BFD0A28A92587FE00" box="[764,901,480,506]" direction="north" orientation="latitude" pageId="16" pageNumber="44" precision="15" value="31.46861">N31°28'07&quot;</geoCoordinate>
<geoCoordinate id="A8889EE3A918FD0BFC7B28A92224FE00" box="[909,1062,480,506]" direction="west" orientation="longitude" pageId="16" pageNumber="44" precision="15" value="-7.416389">W 07°24'59&quot;</geoCoordinate>
,
<taxonomicName id="0ABC83A7A918FD0BFBC528A92297FE03" box="[1075,1173,480,505]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="16" pageNumber="44" phylum="Tracheophyta" rank="genus">
<emphasis id="FFC82436A918FD0BFBC528A92297FE03" box="[1075,1173,480,505]" italics="true" pageId="16" pageNumber="44">Quercus</emphasis>
</taxonomicName>
,
<date id="B902DEE4A918FD0BFB5228A9230FFE00" box="[1188,1293,480,506]" pageId="16" pageNumber="44" value="2009-05-03">3.5.2009</date>
, leg. Stüben -14-&quot;, coll. Stüben, coll.
<collectionCode id="ABAD60E1A918FD0BFE412B4C240FFDE5" LSID="urn:lsid:biocol.org:col:34613" box="[439,525,517,543]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="16" pageNumber="44">ZFMK</collectionCode>
: Zoologisches Forschungsmuseum Alexander Koenig (2ɗ).
</paragraph>
</subSubSection>
<subSubSection id="85A6ABAFA918FD0BFF302B632526FCDE" pageId="16" pageNumber="44" type="discussion">
<paragraph id="CD03F824A918FD0BFF302B632526FCDE" blockId="16.[151,1437,255,804]" pageId="16" pageNumber="44">
<emphasis id="FFC82436A918FD0BFF302B632499FDBE" bold="true" box="[198,667,554,580]" pageId="16" pageNumber="44">Discussion and differential diagnosis.</emphasis>
The new species
<taxonomicName id="0ABC83A7A918FD0BFC872B632280FDB9" box="[881,1154,554,579]" class="Insecta" family="Curculionidae" genus="Kyklioacalles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="44" phylum="Arthropoda" rank="species" species="atlasicus">
<emphasis id="FFC82436A918FD0BFC872B632280FDB9" box="[881,1154,554,579]" italics="true" pageId="16" pageNumber="44">Kyklioacalles atlasicus</emphasis>
</taxonomicName>
from the High Atlas of
<collectingCountry id="B5ABB8B4A918FD0BFF612B192703FD90" box="[151,257,592,618]" name="Morocco" pageId="16" pageNumber="44">Morocco</collectingCountry>
must be compared with the related species from the Anti-Atlas,
<taxonomicName id="0ABC83A7A918FD0BFC0C2B1926E2FD75" authority="Stuben, 2001" authorityName="Stuben" authorityYear="2001" class="Insecta" family="Curculionidae" genus="Kyklioacalles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="44" phylum="Arthropoda" rank="species" species="maroccensis">
<emphasis id="FFC82436A918FD0BFC0C2B192330FD93" box="[1018,1330,592,617]" italics="true" pageId="16" pageNumber="44">Kyklioacalles maroccensis</emphasis>
(
<bibRefCitation id="A92D85D5A918FD0BFAB42B1926D5FD75" author="Stuben" pageId="16" pageNumber="44" refString="Stuben, P. E. (2001) Beschreibung neuer westpalaarktischer Acalles - Arten. (Curculionidae: Cryptorhynchinae). SNUDEBILLER, Studies on taxonomy, biology and ecology of Curculionoidea, 2, 180 - 197." type="journal article" year="2001">Stüben, 2001</bibRefCitation>
)
</taxonomicName>
. The latter was described as an
<taxonomicName id="0ABC83A7A918FD0BFDAA2B3C24B1FD74" box="[604,691,629,654]" class="Insecta" family="Curculionidae" genus="Acalles" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="FFC82436A918FD0BFDAA2B3C24B1FD74" box="[604,691,629,654]" italics="true" pageId="16" pageNumber="44">Acalles</emphasis>
</taxonomicName>
species at first (
<bibRefCitation id="A92D85D5A918FD0BFC812B3C2289FD75" author="Stuben" box="[887,1163,629,655]" pageId="16" pageNumber="44" refString="Stuben, P. E. (2001) Beschreibung neuer westpalaarktischer Acalles - Arten. (Curculionidae: Cryptorhynchinae). SNUDEBILLER, Studies on taxonomy, biology and ecology of Curculionoidea, 2, 180 - 197." type="journal article" year="2001">Stüben 2001: 182–184</bibRefCitation>
); but this was revealed later as a mistake (
<bibRefCitation id="A92D85D5A918FD0BFE832BD3241FFD4E" author="Stuben" box="[373,541,666,692]" pageId="16" pageNumber="44" refString="Stuben, P. E. (2003 a) Revision des Genus Kyklioacalles und Beschreibung der Untergattung Palaeoacalles subg. n. unter Heranziehung phylogenetischer, morphogenetischer und biogeographischer Aspekte (Curculionidae: Cryptorhynchinae). SNUDEBILLER, Studies on taxonomy, biology and ecology of Curculionoidea, 4, 116 - 166." type="journal article" year="2003" yearSuffix="a">Stüben 2003a</bibRefCitation>
:
<specimenCount id="DBBA33ADA918FD0BFDDF2BD32469FD4E" box="[553,619,666,692]" pageId="16" pageNumber="44" type="generic">142f.</specimenCount>
). This species belongs to the Genus
<taxonomicName id="0ABC83A7A918FD0BFBEB2BD322BCFD49" box="[1053,1214,666,691]" class="Insecta" family="Curculionidae" genus="Kyklioacalles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="FFC82436A918FD0BFBEB2BD322BCFD49" box="[1053,1214,666,691]" italics="true" pageId="16" pageNumber="44">Kyklioacalles</emphasis>
</taxonomicName>
, because the basal part of the endophallus consists of a cyclic—even if weakly developed—structure (
<figureCitation id="5587E4A1A918FD0BFB6C2B8922FAFD20" box="[1178,1272,704,730]" captionStart="FIGURES 25 – 29" captionStartId="14.[151,269,1845,1869]" captionTargetBox="[153,1433,194,1823]" captionTargetId="figure@14.[153,1434,194,1823]" captionTargetPageId="14" captionText="FIGURES 25 – 29. Male genitalia of the new and related species; 30. Loci typici and host plants of the new species; 31 – 32. Female genitalia." httpUri="https://zenodo.org/record/276276/files/figure.png" pageId="16" pageNumber="44">Fig. 29</figureCitation>
). The closely related new species has a more highly developed, complete and strongly cyclical structure of the endophallus (
<figureCitation id="5587E4A1A918FD0BFF692A4326F5FCDE" box="[159,247,778,804]" captionStart="FIGURES 25 – 29" captionStartId="14.[151,269,1845,1869]" captionTargetBox="[153,1433,194,1823]" captionTargetId="figure@14.[153,1434,194,1823]" captionTargetPageId="14" captionText="FIGURES 25 – 29. Male genitalia of the new and related species; 30. Loci typici and host plants of the new species; 31 – 32. Female genitalia." httpUri="https://zenodo.org/record/276276/files/figure.png" pageId="16" pageNumber="44">Fig. 28</figureCitation>
). Moreover there are some different characters:
</paragraph>
</subSubSection>
</treatment>
</document>
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