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<document id="972B8E292F3BD6277DA1091CA42607F3" ID-DOI="10.5281/zenodo.5041512" ID-ISSN="1942-1354" ID-Zenodo-Dep="5041512" ID-ZooBank="37900822-FF60-4386-BF30-9434678DD39B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624977543629" checkinUser="carolina" docAuthor="Eya, Bryan K." docDate="2021" docId="445E87AE2B43FF97FF068DC5E707FDAB" docLanguage="en" docName="InsectaMundi.0851.1-36.pdf" docOrigin="Insecta Mundi 2021 (851)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.3:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="3" docTitle="Stenaspis superba Aurivillius 1908" docType="treatment" docVersion="3" lastPageNumber="21" masterDocId="B867FFD62B55FF81FFC68D13E478F837" masterDocTitle="Recharacterization of Stenaspis Audinet-Serville, 1834 with a new species from Mexico (Coleoptera: Cerambycidae: Cerambycinae: Trachyderini)" masterLastPageNumber="36" masterPageNumber="1" pageNumber="21" updateTime="1698978286134" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="49C7ADB3D2344F73C083733AA51F14CB">Recharacterization of Stenaspis Audinet-Serville, 1834 with a new species from Mexico (Coleoptera: Cerambycidae: Cerambycinae: Trachyderini)</mods:title>
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<mods:namePart id="68CED8892EE2C0F62957A84EE61E5580">Eya, Bryan K.</mods:namePart>
<mods:affiliation id="1B070450AD4587FE4194DA27BBA15317">California Environmental Protection Agency Environmental Health Hazard Assessment, Pesticide and Environmental Toxicology 1001 I Street Sacramento, CA 95812 USA</mods:affiliation>
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<mods:title id="648D37C9C041FD4CDADD638B4F26CD5C">Insecta Mundi</mods:title>
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<mods:date id="1A3DBCB4E853F5C4024FC30A3B05C179">2021</mods:date>
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<subSubSection id="84ED65332B43FF97FF068DC5E603F8C4" box="[192,635,213,243]" pageId="22" pageNumber="21" type="nomenclature">
<paragraph id="CC4836B82B43FF97FF068DC5E603F8C4" blockId="22.[192,635,213,276]" box="[192,635,213,243]" pageId="22" pageNumber="21">
<heading id="970081D42B43FF97FF068DC5E603F8C4" box="[192,635,213,243]" fontSize="12" level="2" pageId="22" pageNumber="21" reason="2">
<taxonomicName id="0BF74D3B2B43FF97FF068DC5E603F8C4" ID-CoL="6ZJHY" authority="Aurivillius, 1908" authorityName="Aurivillius" authorityYear="1908" box="[192,635,213,243]" class="Insecta" family="Cerambycidae" genus="Stenaspis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="21" phylum="Arthropoda" rank="species" species="superba">
<emphasis id="FE83EAAA2B43FF97FF068DC5E603F8C4" bold="true" box="[192,635,213,243]" pageId="22" pageNumber="21">
<emphasis id="FE83EAAA2B43FF97FF068DC5E5E7F8C5" bold="true" box="[192,415,213,242]" italics="true" pageId="22" pageNumber="21">Stenaspis superba</emphasis>
<bibRefCitation id="A8664B492B43FF97FE638DC5E603F8C4" author="Aurivillius C." box="[421,635,214,243]" pageId="22" pageNumber="21" pagination="1 - 13" refId="ref23244" refString="Aurivillius C. 1908. Cerambyciden aus den Grenzgebieten zwischen Peru und Bolivien gesammelt von Dr. Nils Holmgren. Arkiv for Zoologi 5 (1): 1 - 13, 4 fig." type="book chapter" year="1908">Aurivillius, 1908</bibRefCitation>
</emphasis>
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</paragraph>
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<paragraph id="CC4836B82B43FF97FF068DEAE52DF923" blockId="22.[192,635,213,276]" box="[192,341,249,276]" pageId="22" pageNumber="21">
(
<figureCitation id="54CC2A3D2B43FF97FF0E8DEAE536F923" box="[200,334,249,276]" captionStart="Figures 91102" captionStartId="20.[192,279,1342,1368]" captionTargetBox="[232,1396,214,1310]" captionTargetId="figure-292@20.[464,930,645,1053]" captionTargetPageId="20" captionText="Figures 91102. Dorsal and lateral images of Stenaspis solitaria, and images from all sides of S. superba, holotype, female. 9194) S. solitaria, Cochise, AZ, USA. 9192) Male, 29 mm. 9394) Female, 31 mm. 95101) S. superba, holotype, female, Mojos, BOL. 98) Image of head showing apices of mandible acute, genae large, quadrate margins of lower eye lobes well separated from mandibles. 98: 98a) Dorsal anterior margins of genae ridged. 99) Caudal image showing brush of hairs. 100) Lateral image showing coarsely punctured proepisternum and epipleural margin of elytron below humerus. 101) Ventral image showing pronotum narrower than elytra at humeri, lateral tubercles of pronotum slightly behind middle, rounded anterior angle of pronotum, and protuberant prosternal intercoxal process. 102) Label information provided by NHRS along with photos of S. superba." figureDoi="http://doi.org/10.5281/zenodo.5041544" httpUri="https://zenodo.org/record/5041544/files/figure.png" pageId="22" pageNumber="21">Fig. 95101</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="84ED65332B43FF97FF268C31E0F5F96F" pageId="22" pageNumber="21" type="reference_group">
<paragraph id="CC4836B82B43FF97FF268C31E0F5F96F" blockId="22.[224,1440,289,344]" pageId="22" pageNumber="21">
<treatmentCitationGroup id="ECE711962B43FF97FF268C31E0F5F96F" pageId="22" pageNumber="21">
<taxonomicName id="0BF74D3B2B43FF97FF268C31E6C5F90E" ID-CoL="6ZJHY" authority="Aurivillius 1908: 5" authorityName="Aurivillius" authorityPageNumber="5" authorityYear="1908" box="[224,701,289,314]" class="Insecta" family="Cerambycidae" genus="Stenaspis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="21" phylum="Arthropoda" rank="species" species="superba">
<treatmentCitation id="4D5610A92B43FF97FF268C31E629F90E" author="Aurivillius C." box="[224,593,289,314]" page="5" pageId="22" pageNumber="21" year="1908">
<emphasis id="FE83EAAA2B43FF97FF268C31E5F7F90E" box="[224,399,289,313]" italics="true" pageId="22" pageNumber="21">Stenaspis superba</emphasis>
<bibRefCitation id="A8664B492B43FF97FE508C32E629F90E" author="Aurivillius C." box="[406,593,289,314]" pageId="22" pageNumber="21" pagination="1 - 13" refId="ref23244" refString="Aurivillius C. 1908. Cerambyciden aus den Grenzgebieten zwischen Peru und Bolivien gesammelt von Dr. Nils Holmgren. Arkiv for Zoologi 5 (1): 1 - 13, 4 fig." type="book chapter" year="1908">Aurivillius 1908: 5</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="4D5610A92B43FF97FD9A8C31E6C5F90E" author="Aurivillius C." box="[604,701,289,314]" page="458" pageId="22" pageNumber="21" year="1912">
<bibRefCitation id="A8664B492B43FF97FD9A8C31E6C5F90E" author="Aurivillius C." box="[604,701,289,314]" pageId="22" pageNumber="21" pagination="457 - 458" refId="ref23279" refString="Aurivillius C. 1912. Cerambycidae: Cerambycinae. p. 457 - 458. In: Junk W, Schenkling S. (eds.). Coleopterorum Catalogus pars 39 [Vol. 22]. W. Junk; Berlin. 574 p." type="book chapter" year="1912">1912: 458</bibRefCitation>
</treatmentCitation>
</taxonomicName>
(cat.);
<treatmentCitation id="4D5610A92B43FF97FCC38C32E7CEF90D" author="Melzer J." box="[773,950,289,314]" page="422" pageId="22" pageNumber="21" year="1932">
<bibRefCitation id="A8664B492B43FF97FCC38C32E7CEF90D" author="Melzer J." box="[773,950,289,314]" pageId="22" pageNumber="21" pagination="420 - 434" refId="ref26193" refString="Melzer J. 1932. Novos generos e novas especies de Cerambycideos do Brasil (Col. Cerambycidae). Revista de Entomologia, Rio de Janeiro 2 (4): 420 - 434, 12 fig." type="book chapter" year="1932">Melzer 1932: 422</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="4D5610A92B43FF97FC798C32E0DAF90E" author="Blackwelder RE" box="[959,1186,289,313]" page="589" pageId="22" pageNumber="21" year="1946">
<bibRefCitation id="A8664B492B43FF97FC798C32E0DAF90E" author="Blackwelder RE" box="[959,1186,289,313]" pageId="22" pageNumber="21" pagination="551 - 763" refId="ref23548" refString="Blackwelder RE. 1946. Checklist of the coleopterous insects of Mexico, Central America, the West Indies and South America. Part 4. Bulletin of the United States National Museum; Washington D. C. 185 (4): 551 - 763." type="journal article" year="1946">Blackwelder 1946: 589</bibRefCitation>
</treatmentCitation>
(cat.); Linsley 1961: 632;
<treatmentCitation id="4D5610A92B43FF97FEC68C53E64EF96F" author="Monne MA &amp; Giesbert EF" box="[256,566,320,344]" page="152" pageId="22" pageNumber="21" year="1994">
<bibRefCitation id="A8664B492B43FF97FEC68C53E64EF96F" author="Monne MA &amp; Giesbert EF" box="[256,566,320,344]" pageId="22" pageNumber="21" pagination="i - xiv" refId="ref26385" refString="Monne MA, Giesbert EF. 1994. Checklist of the Cerambycidae and Disteniidae (Coleoptera) of the Western Hemisphere. Wolfsgarden Books; Burbank, CA. i - xiv + 410 p." type="book chapter" year="1994">Monné and Giesbert 1994: 152</bibRefCitation>
</treatmentCitation>
(cat.); Monné 1994: 35 (cat.);
<treatmentCitation id="4D5610A92B43FF97FCA38C53E03CF96F" author="Wappes JE &amp; Morris RF &amp; Nearns EH &amp; Thomas MC" box="[869,1092,320,344]" page="23" pageId="22" pageNumber="21" year="2006">
<bibRefCitation id="A8664B492B43FF97FCA38C53E03CF96F" author="Wappes JE &amp; Morris RF &amp; Nearns EH &amp; Thomas MC" box="[869,1092,320,344]" pageId="22" pageNumber="21" pagination="1 - 45" refId="ref27439" refString="Wappes JE, Morris RF, Nearns EH, Thomas MC. 2006. Preliminary checklist of Bolivian Cerambycidae (Coleoptera). Insecta Mundi 20 (1 - 2): 1 - 45." type="journal article" year="2006">Wappes et al. 2006: 23</bibRefCitation>
</treatmentCitation>
(dist.).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="84ED65332B43FF97FF068C74E148F9D1" pageId="22" pageNumber="21" type="materials_examined">
<paragraph id="CC4836B82B43FF97FF068C74E148F9D1" blockId="22.[192,1441,359,487]" pageId="22" pageNumber="21">
<emphasis id="FE83EAAA2B43FF97FF068C74E63DF9B6" bold="true" box="[192,581,359,385]" pageId="22" pageNumber="21">Photographic Material Examined.</emphasis>
<materialsCitation id="7C9F3CE52B43FF97FD8E8C7BE148F9D1" collectionCode="NHRS" collectorName="Mojos, N &amp; J. Bergsten" country="Bolivia" location="Swedish Museum of Natural History" pageId="22" pageNumber="21" specimenCount="1" typeStatus="holotype">
<typeStatus id="134C881A2B43FF97FD8E8C7BE6C9F9B5" box="[584,689,360,386]" pageId="22" pageNumber="21" type="holotype">Holotype</typeStatus>
, female,
<collectingCountry id="B4E076282B43FF97FCCB8C7BE705F9B5" box="[781,893,360,386]" name="Bolivia" pageId="22" pageNumber="21">BOLIVIA</collectingCountry>
:
<collectorName id="6102536E2B43FF97FC438C7BE794F9B5" box="[901,1004,360,386]" pageId="22" pageNumber="21">Mojos, N</collectorName>
. Holmgren, June on the banks of stream, NHRS-JLKB 000071808. Six photographs provided by
<collectorName id="6102536E2B43FF97FCF68C99E7D1F993" box="[816,937,394,420]" pageId="22" pageNumber="21">J. Bergsten</collectorName>
of the
<location id="C92860632B43FF97FC398C9AE1D8F994" LSID="urn:lsid:plazi:treatment:445E87AE2B43FF97FF068DC5E707FDAB:C92860632B43FF97FC398C9AE1D8F994" box="[1023,1440,393,420]" country="Bolivia" name="Swedish Museum of Natural History" pageId="22" pageNumber="21">Swedish Museum of Natural History</location>
(
<collectionCode id="AAE6AE7D2B43FF97FF0E8CB8E56DF9F2" box="[200,277,427,453]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="22" pageNumber="21">NHRS</collectionCode>
, Naturhistoriska Riksmuseet) as follows: NHRS-JLKB 000071808_caud.tif; NHRS-JLKB 000071808_dors. tif; NHRS-JLKB 000071808_fron.tif; NHRS-JLKB 000071808_late.tif; NHRS-JLKB 000071808_vent.tif.
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</paragraph>
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<subSubSection id="84ED65332B43FF97FF068CE5E707FDAB" pageId="22" pageNumber="21" type="discussion">
<paragraph id="CC4836B82B43FF97FF068CE5E78BFBB5" blockId="22.[192,1441,502,1436]" pageId="22" pageNumber="21">
<emphasis id="FE83EAAA2B43FF97FF068CE5E53FFA27" bold="true" box="[192,327,502,528]" pageId="22" pageNumber="21">Discussion.</emphasis>
<bibRefCitation id="A8664B492B43FF97FE8B8CE5E655FA26" author="Aurivillius C." box="[333,557,502,529]" pageId="22" pageNumber="21" pagination="1 - 13" refId="ref23244" refString="Aurivillius C. 1908. Cerambyciden aus den Grenzgebieten zwischen Peru und Bolivien gesammelt von Dr. Nils Holmgren. Arkiv for Zoologi 5 (1): 1 - 13, 4 fig." type="book chapter" year="1908">Aurivillius (1908: 5)</bibRefCitation>
described
<taxonomicName id="0BF74D3B2B43FF97FD608CE5E76EFA27" authorityName="Aurivillius" authorityYear="1908" box="[678,790,502,528]" class="Insecta" family="Cerambycidae" genus="Stenaspis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="21" phylum="Arthropoda" rank="species" species="superba">
<emphasis id="FE83EAAA2B43FF97FD608CE5E76EFA27" box="[678,790,502,528]" italics="true" pageId="22" pageNumber="21">S. superba</emphasis>
</taxonomicName>
(length
<quantity id="0B0F9B5D2B43FF97FCB48CE5E7BBFA26" box="[882,963,502,529]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" pageId="22" pageNumber="21" unit="mm" value="38.0">38 mm</quantity>
) as a species with integument that is black, the pronotum and elytra orange with three nitid black calli on pronotal disc, two in anterior half and one medially in the posterior half. The head is punctate, vertex rugulose and uneven. The antennae barely reach middle of elytra, scape is conical, sub-nitid, sparsely punctate, and third antennomere shorter than the scape. The prothorax is described as being deeply rugose, punctate, convex with prominent tubercles on each side, surface clothed with short, erect golden-brown hairs, and the apical and basal margins narrowly black with dense black pubescence. The scutellum is narrow, elongate, black, canaliculate in the middle and irregularly, sparsely punctate. The elytra are strongly rugose and clothed with a few short hairs at the base, less rugose apically and denudate from the middle, and apices broadly rounded and unarmed. The sternum is all black, the prosternum is very rugulose, meso- and metasternum densely punctate and abdomen sparsely punctate. According to Aurivillius,
<taxonomicName id="0BF74D3B2B43FF97FADC8E36E1F1FB09" authorityName="Aurivillius" authorityYear="1908" box="[1306,1417,804,831]" class="Insecta" family="Cerambycidae" genus="Stenaspis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="21" phylum="Arthropoda" rank="species" species="superba">
<emphasis id="FE83EAAA2B43FF97FADC8E36E1F1FB09" box="[1306,1417,804,831]" italics="true" pageId="22" pageNumber="21">S. superba</emphasis>
</taxonomicName>
is differentiated from all other known species of
<taxonomicName id="0BF74D3B2B43FF97FD018E55E753FB57" authorityName="Audinet-Serville" authorityYear="1834" box="[711,811,838,864]" class="Insecta" family="Cerambycidae" genus="Stenaspis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="FE83EAAA2B43FF97FD018E55E753FB57" box="[711,811,838,864]" italics="true" pageId="22" pageNumber="21">Stenaspis</emphasis>
</taxonomicName>
by the short antennae, and by the coloration and sculpture of the pronotum and elytra, which are reminiscent of an orange skin.
</paragraph>
<paragraph id="CC4836B82B43FF97FEC68E9AE707FDAB" blockId="22.[192,1441,502,1436]" pageId="22" pageNumber="21">
Examination of
<typeStatus id="134C881A2B43FF97FE738E9AE66FFB94" box="[437,535,905,931]" pageId="22" pageNumber="21" type="holotype">holotype</typeStatus>
photographs shows that
<taxonomicName id="0BF74D3B2B43FF97FCF48E99E7DBFB94" authorityName="Aurivillius" authorityYear="1908" box="[818,931,905,932]" class="Insecta" family="Cerambycidae" genus="Stenaspis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="21" phylum="Arthropoda" rank="species" species="superba">
<emphasis id="FE83EAAA2B43FF97FCF48E99E7DBFB94" box="[818,931,905,932]" italics="true" pageId="22" pageNumber="21">S. superba</emphasis>
</taxonomicName>
appears to have the characters representative of
<taxonomicName id="0BF74D3B2B43FF97FF1A8EB8E538FBF2" authorityName="Audinet-Serville" authorityYear="1834" box="[220,320,939,965]" class="Insecta" family="Cerambycidae" genus="Stenaspis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="FE83EAAA2B43FF97FF1A8EB8E538FBF2" box="[220,320,939,965]" italics="true" pageId="22" pageNumber="21">Stenaspis</emphasis>
</taxonomicName>
as follows: (1) head with frons large, square and recessed between the dorsal anterior margin of gena (
<figureCitation id="54CC2A3D2B43FF97FF0E8EDFE537FBD0" box="[200,335,972,999]" captionStart="Figures 91102" captionStartId="20.[192,279,1342,1368]" captionTargetBox="[232,1396,214,1310]" captionTargetId="figure-292@20.[464,930,645,1053]" captionTargetPageId="20" captionText="Figures 91102. Dorsal and lateral images of Stenaspis solitaria, and images from all sides of S. superba, holotype, female. 9194) S. solitaria, Cochise, AZ, USA. 9192) Male, 29 mm. 9394) Female, 31 mm. 95101) S. superba, holotype, female, Mojos, BOL. 98) Image of head showing apices of mandible acute, genae large, quadrate margins of lower eye lobes well separated from mandibles. 98: 98a) Dorsal anterior margins of genae ridged. 99) Caudal image showing brush of hairs. 100) Lateral image showing coarsely punctured proepisternum and epipleural margin of elytron below humerus. 101) Ventral image showing pronotum narrower than elytra at humeri, lateral tubercles of pronotum slightly behind middle, rounded anterior angle of pronotum, and protuberant prosternal intercoxal process. 102) Label information provided by NHRS along with photos of S. superba." figureDoi="http://doi.org/10.5281/zenodo.5041544" httpUri="https://zenodo.org/record/5041544/files/figure.png" pageId="22" pageNumber="21">Fig. 98, 100</figureCitation>
); (2) prosternal intercoxal process protuberant and ridged between coxae and vertical behind (
<figureCitation id="54CC2A3D2B43FF97FAB28EDFE493FC3F" captionStart="Figures 91102" captionStartId="20.[192,279,1342,1368]" captionTargetBox="[232,1396,214,1310]" captionTargetId="figure-292@20.[464,930,645,1053]" captionTargetPageId="20" captionText="Figures 91102. Dorsal and lateral images of Stenaspis solitaria, and images from all sides of S. superba, holotype, female. 9194) S. solitaria, Cochise, AZ, USA. 9192) Male, 29 mm. 9394) Female, 31 mm. 95101) S. superba, holotype, female, Mojos, BOL. 98) Image of head showing apices of mandible acute, genae large, quadrate margins of lower eye lobes well separated from mandibles. 98: 98a) Dorsal anterior margins of genae ridged. 99) Caudal image showing brush of hairs. 100) Lateral image showing coarsely punctured proepisternum and epipleural margin of elytron below humerus. 101) Ventral image showing pronotum narrower than elytra at humeri, lateral tubercles of pronotum slightly behind middle, rounded anterior angle of pronotum, and protuberant prosternal intercoxal process. 102) Label information provided by NHRS along with photos of S. superba." figureDoi="http://doi.org/10.5281/zenodo.5041544" httpUri="https://zenodo.org/record/5041544/files/figure.png" pageId="22" pageNumber="21">Fig. 101</figureCitation>
); (3) mesosternal intercoxal process not protuberant, level with coxae, and abruptly declivous in front (
<figureCitation id="54CC2A3D2B43FF97FAB28EFDE494FC1D" captionStart="Figures 91102" captionStartId="20.[192,279,1342,1368]" captionTargetBox="[232,1396,214,1310]" captionTargetId="figure-292@20.[464,930,645,1053]" captionTargetPageId="20" captionText="Figures 91102. Dorsal and lateral images of Stenaspis solitaria, and images from all sides of S. superba, holotype, female. 9194) S. solitaria, Cochise, AZ, USA. 9192) Male, 29 mm. 9394) Female, 31 mm. 95101) S. superba, holotype, female, Mojos, BOL. 98) Image of head showing apices of mandible acute, genae large, quadrate margins of lower eye lobes well separated from mandibles. 98: 98a) Dorsal anterior margins of genae ridged. 99) Caudal image showing brush of hairs. 100) Lateral image showing coarsely punctured proepisternum and epipleural margin of elytron below humerus. 101) Ventral image showing pronotum narrower than elytra at humeri, lateral tubercles of pronotum slightly behind middle, rounded anterior angle of pronotum, and protuberant prosternal intercoxal process. 102) Label information provided by NHRS along with photos of S. superba." figureDoi="http://doi.org/10.5281/zenodo.5041544" httpUri="https://zenodo.org/record/5041544/files/figure.png" pageId="22" pageNumber="21">Fig. 100</figureCitation>
); (4) pronotum narrower than base of elytra at humeri, lateral tubercles placed slightly behind middle, and anterior angle broadly callused (
<figureCitation id="54CC2A3D2B43FF97FDE08922E6FBFC7C" box="[550,643,1073,1099]" captionStart="Figures 91102" captionStartId="20.[192,279,1342,1368]" captionTargetBox="[232,1396,214,1310]" captionTargetId="figure-292@20.[464,930,645,1053]" captionTargetPageId="20" captionText="Figures 91102. Dorsal and lateral images of Stenaspis solitaria, and images from all sides of S. superba, holotype, female. 9194) S. solitaria, Cochise, AZ, USA. 9192) Male, 29 mm. 9394) Female, 31 mm. 95101) S. superba, holotype, female, Mojos, BOL. 98) Image of head showing apices of mandible acute, genae large, quadrate margins of lower eye lobes well separated from mandibles. 98: 98a) Dorsal anterior margins of genae ridged. 99) Caudal image showing brush of hairs. 100) Lateral image showing coarsely punctured proepisternum and epipleural margin of elytron below humerus. 101) Ventral image showing pronotum narrower than elytra at humeri, lateral tubercles of pronotum slightly behind middle, rounded anterior angle of pronotum, and protuberant prosternal intercoxal process. 102) Label information provided by NHRS along with photos of S. superba." figureDoi="http://doi.org/10.5281/zenodo.5041544" httpUri="https://zenodo.org/record/5041544/files/figure.png" pageId="22" pageNumber="21">Fig. 101</figureCitation>
); (5) proepisternum of thorax coarsely punctate (
<figureCitation id="54CC2A3D2B43FF97FB608922E08EFC7C" box="[1190,1270,1073,1099]" captionStart="Figures 3341" captionStartId="9.[192,279,1168,1194]" captionTargetBox="[197,1436,213,1140]" captionTargetId="figure-215@9.[661,970,524,833]" captionTargetPageId="9" captionText="Figures 3341. Pronotal and prosternal characteristic of Stenaspis solitaria and Stenaspis superba. 3335) S. solitaria, male, Cochise Co., AZ, USA with disc coarsely punctate with 5 vague calli (yellow arrows). 33: 33a) Pronotum with prominent lateral tubercle. 33: 33b) Rounded anterior angles, which does not extend and merge over lateral tubercles. 34: 34a) Proepisternum finely punctate. 35: 35a) Prosternum with finely densely punctate transverse subrectangular area on each side of middle. 3638) S. solitaria, female, Cochise Co., AZ, USA 36: 36a) Anterior angle of pronotum obtusely callused and obliquely tapering to the apex of pronotum. 37) Coarsely punctate proepisternum of female. 38) Coarsely punctate prosternum of female. 3941) S. superba, holotype, female, Mojos, BOL (photos provided from J. Bergsten, NHRS). 39) Coarsely punctate pronotal disc with two prominent discal calli in anterior half. 40) Convex pronotal disc with coarsely punctate proepisternum. 41) Coarsely punctate prosternum and broadly callused anterior angle of S. superba." figureDoi="http://doi.org/10.5281/zenodo.5041522" httpUri="https://zenodo.org/record/5041522/files/figure.png" pageId="22" pageNumber="21">Fig. 40</figureCitation>
); and (6) elytra distinctly margined laterally (
<figureCitation id="54CC2A3D2B43FF97FDCD8940E612FC5A" box="[523,618,1107,1133]" captionStart="Figures 91102" captionStartId="20.[192,279,1342,1368]" captionTargetBox="[232,1396,214,1310]" captionTargetId="figure-292@20.[464,930,645,1053]" captionTargetPageId="20" captionText="Figures 91102. Dorsal and lateral images of Stenaspis solitaria, and images from all sides of S. superba, holotype, female. 9194) S. solitaria, Cochise, AZ, USA. 9192) Male, 29 mm. 9394) Female, 31 mm. 95101) S. superba, holotype, female, Mojos, BOL. 98) Image of head showing apices of mandible acute, genae large, quadrate margins of lower eye lobes well separated from mandibles. 98: 98a) Dorsal anterior margins of genae ridged. 99) Caudal image showing brush of hairs. 100) Lateral image showing coarsely punctured proepisternum and epipleural margin of elytron below humerus. 101) Ventral image showing pronotum narrower than elytra at humeri, lateral tubercles of pronotum slightly behind middle, rounded anterior angle of pronotum, and protuberant prosternal intercoxal process. 102) Label information provided by NHRS along with photos of S. superba." figureDoi="http://doi.org/10.5281/zenodo.5041544" httpUri="https://zenodo.org/record/5041544/files/figure.png" pageId="22" pageNumber="21">Fig. 100</figureCitation>
), and apices rounded and unarmed (
<figureCitation id="54CC2A3D2B43FF97FBCF8940E0FBFC5A" box="[1033,1155,1107,1134]" captionStart="Figures 91102" captionStartId="20.[192,279,1342,1368]" captionTargetBox="[232,1396,214,1310]" captionTargetId="figure-292@20.[464,930,645,1053]" captionTargetPageId="20" captionText="Figures 91102. Dorsal and lateral images of Stenaspis solitaria, and images from all sides of S. superba, holotype, female. 9194) S. solitaria, Cochise, AZ, USA. 9192) Male, 29 mm. 9394) Female, 31 mm. 95101) S. superba, holotype, female, Mojos, BOL. 98) Image of head showing apices of mandible acute, genae large, quadrate margins of lower eye lobes well separated from mandibles. 98: 98a) Dorsal anterior margins of genae ridged. 99) Caudal image showing brush of hairs. 100) Lateral image showing coarsely punctured proepisternum and epipleural margin of elytron below humerus. 101) Ventral image showing pronotum narrower than elytra at humeri, lateral tubercles of pronotum slightly behind middle, rounded anterior angle of pronotum, and protuberant prosternal intercoxal process. 102) Label information provided by NHRS along with photos of S. superba." figureDoi="http://doi.org/10.5281/zenodo.5041544" httpUri="https://zenodo.org/record/5041544/files/figure.png" pageId="22" pageNumber="21">Fig. 95, 97</figureCitation>
). Compared to other
<taxonomicName id="0BF74D3B2B43FF97FAB08940E525FCB9" class="Insecta" family="Cerambycidae" genus="Stenaspis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="21" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="FE83EAAA2B43FF97FAB08940E57CFCB9" italics="true" pageId="22" pageNumber="21">Stenaspis</emphasis>
species
</taxonomicName>
,
<taxonomicName id="0BF74D3B2B43FF97FEAF8966E5A3FCB9" authorityName="Aurivillius" authorityYear="1908" box="[361,475,1140,1167]" class="Insecta" family="Cerambycidae" genus="Stenaspis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="21" phylum="Arthropoda" rank="species" species="superba">
<emphasis id="FE83EAAA2B43FF97FEAF8966E5A3FCB9" box="[361,475,1140,1167]" italics="true" pageId="22" pageNumber="21">S. superba</emphasis>
</taxonomicName>
has a pronotal disc that is more convex (
<figureCitation id="54CC2A3D2B43FF97FC688967E787FCB8" box="[942,1023,1140,1167]" captionStart="Figures 3341" captionStartId="9.[192,279,1168,1194]" captionTargetBox="[197,1436,213,1140]" captionTargetId="figure-215@9.[661,970,524,833]" captionTargetPageId="9" captionText="Figures 3341. Pronotal and prosternal characteristic of Stenaspis solitaria and Stenaspis superba. 3335) S. solitaria, male, Cochise Co., AZ, USA with disc coarsely punctate with 5 vague calli (yellow arrows). 33: 33a) Pronotum with prominent lateral tubercle. 33: 33b) Rounded anterior angles, which does not extend and merge over lateral tubercles. 34: 34a) Proepisternum finely punctate. 35: 35a) Prosternum with finely densely punctate transverse subrectangular area on each side of middle. 3638) S. solitaria, female, Cochise Co., AZ, USA 36: 36a) Anterior angle of pronotum obtusely callused and obliquely tapering to the apex of pronotum. 37) Coarsely punctate proepisternum of female. 38) Coarsely punctate prosternum of female. 3941) S. superba, holotype, female, Mojos, BOL (photos provided from J. Bergsten, NHRS). 39) Coarsely punctate pronotal disc with two prominent discal calli in anterior half. 40) Convex pronotal disc with coarsely punctate proepisternum. 41) Coarsely punctate prosternum and broadly callused anterior angle of S. superba." figureDoi="http://doi.org/10.5281/zenodo.5041522" httpUri="https://zenodo.org/record/5041522/files/figure.png" pageId="22" pageNumber="21">Fig. 40</figureCitation>
) with two prominent anterior dorsal calli, and a disc surface that is more coarsely punctate. Also, the elytra (E) are more elongate relative to pronotum (P), where the E/P ratio is 4.8 compared to other species where E/P ranges from 33.8. The other four species have the pronotal disc flattened (
<figureCitation id="54CC2A3D2B43FF97FDF589CAE6D4FCC4" box="[563,684,1241,1267]" captionStart="Figures 1523" captionStartId="7.[192,278,1345,1371]" captionTargetBox="[198,1438,213,1315]" captionTargetId="figure-294@7.[611,1024,491,904]" captionTargetPageId="7" captionText="Figures 1523. Pronotal and prosternal characteristic of Stenaspis verticalis: 1517) S. verticalis, male, Jalisco, MEX with sides of pronotum rounded. 15) Disc coarsely punctate with 5 vague calli (red arrows). 16: 16a) Proepisternum inflated and finely punctate (dark reddish area). 16: 16b) Demarcation of proepisternal finely punctate area from coarsely punctate dorsum of disc. 17: 17a) Subrectangular impressed area. 17: 17b) Vague demarcation dividing proepisternum from the subrectangular impressed area. 18, 20) S. verticalis, male, Nuevo Leon, MEX with side of pronotum with distinct lateral tubercles. 19) S. verticalis, male, Puebla, MEX with proepisternum inflated and finely punctate clearly demarcated from dorsum. 2122) S. verticalis, female, Guerrero, MEX with side of pronotum distinctly tuberculate. 21: 21a) Anterior angles obtusely tuberculate. 22) Pronotal disc and proepisternum coarsely punctate. 23) S. verticalis, female, Jalisco, MEX with prosternum coarsely punctate." figureDoi="http://doi.org/10.5281/zenodo.5041518" httpUri="https://zenodo.org/record/5041518/files/figure.png" pageId="22" pageNumber="21">Fig. 16, 22</figureCitation>
,
<figureCitation id="54CC2A3D2B43FF97FD7E89CAE6ADFCC4" box="[696,725,1241,1267]" captionStart="Figures 2432" captionStartId="8.[192,279,1345,1371]" captionTargetBox="[197,1438,214,1314]" captionTargetId="figure-146@8.[610,1023,490,903]" captionTargetPageId="8" captionText="Figures 2432. Pronotal and prosternal characteristic of Stenaspis castaneipennis. 2426) S. castaneipennis, male, Oaxaca, MEX with sides of pronotum rounded, disc coarsely punctate with 5 vague calli. 25) Proepisternum inflated and finely punctate (dark reddish area) clearly demarcated from coarsely punctate dorsum (black area). 2729) S. castaneipennis, male, Oaxaca, MEX with sides of pronotum with distinct lateral tubercles. 28) Proepisternum inflated and finely punctate (reddish area) clearly demarcated from dorsum (black area). 29: 29a) Subrectangular finely punctate area of prosternum. 29: 29b) Demarcation line dividing proepisternum from finely punctate subrectangular area of prosternum. 3032) S. castaneipennis, female, Oaxaca, MEX with side of pronotum distinctly tuberculate. 30: 30a) Anterior angles obtusely tuberculate. 3132) Pronotal disc, proepisternum and prosternum coarsely punctate; proepisternun not demarcated from dorsum and prosternum." figureDoi="http://doi.org/10.5281/zenodo.5041520" httpUri="https://zenodo.org/record/5041520/files/figure.png" pageId="22" pageNumber="21">31</figureCitation>
,
<figureCitation id="54CC2A3D2B43FF97FD2689CAE75EFCC4" box="[736,806,1241,1267]" captionStart="Figures 3341" captionStartId="9.[192,279,1168,1194]" captionTargetBox="[197,1436,213,1140]" captionTargetId="figure-215@9.[661,970,524,833]" captionTargetPageId="9" captionText="Figures 3341. Pronotal and prosternal characteristic of Stenaspis solitaria and Stenaspis superba. 3335) S. solitaria, male, Cochise Co., AZ, USA with disc coarsely punctate with 5 vague calli (yellow arrows). 33: 33a) Pronotum with prominent lateral tubercle. 33: 33b) Rounded anterior angles, which does not extend and merge over lateral tubercles. 34: 34a) Proepisternum finely punctate. 35: 35a) Prosternum with finely densely punctate transverse subrectangular area on each side of middle. 3638) S. solitaria, female, Cochise Co., AZ, USA 36: 36a) Anterior angle of pronotum obtusely callused and obliquely tapering to the apex of pronotum. 37) Coarsely punctate proepisternum of female. 38) Coarsely punctate prosternum of female. 3941) S. superba, holotype, female, Mojos, BOL (photos provided from J. Bergsten, NHRS). 39) Coarsely punctate pronotal disc with two prominent discal calli in anterior half. 40) Convex pronotal disc with coarsely punctate proepisternum. 41) Coarsely punctate prosternum and broadly callused anterior angle of S. superba." figureDoi="http://doi.org/10.5281/zenodo.5041522" httpUri="https://zenodo.org/record/5041522/files/figure.png" pageId="22" pageNumber="21">34, 37</figureCitation>
) and glabrate with vaguely visible dorsal calli. Although
<taxonomicName id="0BF74D3B2B43FF97FF0689E8E549FD22" authorityName="Aurivillius" authorityYear="1908" box="[192,305,1275,1301]" class="Insecta" family="Cerambycidae" genus="Stenaspis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="21" phylum="Arthropoda" rank="species" species="superba">
<emphasis id="FE83EAAA2B43FF97FF0689E8E4B5FD22" box="[192,205,1275,1301]" italics="true" pageId="22" pageNumber="21">S</emphasis>
.
<emphasis id="FE83EAAA2B43FF97FF1D89E8E549FD22" box="[219,305,1275,1301]" italics="true" pageId="22" pageNumber="21">superba</emphasis>
</taxonomicName>
is geographically isolated in
<collectingCountry id="B4E076282B43FF97FDB189E8E6BDFD22" box="[631,709,1275,1301]" name="Bolivia" pageId="22" pageNumber="21">Bolivia</collectingCountry>
compared to all other known species found in
<collectingCountry id="B4E076282B43FF97FB1C89E8E177FD22" box="[1242,1295,1275,1301]" name="United States of America" pageId="22" pageNumber="21">USA</collectingCountry>
and
<collectingCountry id="B4E076282B43FF97FA8E89E8E1E5FD22" box="[1352,1437,1275,1301]" name="Mexico" pageId="22" pageNumber="21">Mexico</collectingCountry>
, it has many characteristics that are commonly found in
<taxonomicName id="0BF74D3B2B43FF97FCF1880EE7E3FD00" authorityName="Audinet-Serville" authorityYear="1834" box="[823,923,1309,1335]" class="Insecta" family="Cerambycidae" genus="Stenaspis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="FE83EAAA2B43FF97FCF1880EE7E3FD00" box="[823,923,1309,1335]" italics="true" pageId="22" pageNumber="21">Stenaspis</emphasis>
</taxonomicName>
; therefore, at this time this species is retained in this genus until a male is captured and the pattern of sexual punctation on the thorax is examined. Males of this group of trachyderines (i.e.,
<taxonomicName id="0BF74D3B2B43FF97FDEA8873E6A9FD4D" authorityName="Eya (Eya" authorityYear="2015" box="[556,721,1376,1402]" class="Insecta" family="Cerambycidae" genus="Callistochroma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="FE83EAAA2B43FF97FDEA8873E6A9FD4D" box="[556,721,1376,1402]" italics="true" pageId="22" pageNumber="21">Callistochroma</emphasis>
</taxonomicName>
,
<taxonomicName id="0BF74D3B2B43FF97FD188873E717FD4D" authorityName="Audinet-Serville" authorityYear="1834" box="[734,879,1376,1402]" class="Insecta" family="Cerambycidae" genus="Crioprosopus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="FE83EAAA2B43FF97FD188873E717FD4D" box="[734,879,1376,1402]" italics="true" pageId="22" pageNumber="21">Crioprosopus</emphasis>
</taxonomicName>
and
<taxonomicName id="0BF74D3B2B43FF97FC608873E072FD4D" authorityName="Audinet-Serville" authorityYear="1834" box="[934,1034,1376,1402]" class="Insecta" family="Cerambycidae" genus="Stenaspis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="FE83EAAA2B43FF97FC608873E072FD4D" box="[934,1034,1376,1402]" italics="true" pageId="22" pageNumber="21">Stenaspis</emphasis>
</taxonomicName>
) have additional characteristics that substantiate placement of species into their appropriate genera.
</paragraph>
</subSubSection>
</treatment>
</document>