474 lines
72 KiB
XML
474 lines
72 KiB
XML
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<mods:title id="C1FB553C5238A11107B4FA8CA6330E69">Description of two new genera and five new species of clade K 92 from Bahia and Espírito Santo, southeastern Brazil (Arachnida, Opiliones, Gonyleptidae)</mods:title>
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<mods:name id="EDE2F8E24D9BD7CA3373FE3DE6160CFE" type="personal">
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<mods:role id="B91259781711070A636CC9D9C3FA65B0">
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<mods:roleTerm id="CA18C745F939CBF293A12DCB39CDECB4">Author</mods:roleTerm>
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<mods:namePart id="5839A85CA9F4A03DA337F3D42F38C650">Kury, Adriano B.</mods:namePart>
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<mods:nameIdentifier id="481CF0AF7185CCB716DE3DD30AA307FE" type="ORCID">0000-0002-8334-6204</mods:nameIdentifier>
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<mods:affiliation id="EC91BD2A9AA331A2CDD55A82C80774B0">Departamento de Invertebrados, Museu Nacional / Universidade Federal do Rio de Janeiro, Quinta da Boa Vista, São Cristóvão, 20.940 - 040 Rio de Janeiro (RJ), Brazil. adrianok @ gmail. com; https: // orcid. org / 0000 - 0002 - 8334 - 6204</mods:affiliation>
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<mods:nameIdentifier id="CF81D990EACD6DD9FCBA687FA9126EA1" type="email">adrianok@gmail.com</mods:nameIdentifier>
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<mods:namePart id="36BB763DBE54A9C4592433563ED7B526">Bernabé, Tiago</mods:namePart>
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<mods:affiliation id="54FB7E3146137976379FD74657D47EBE">Programa de Pós-Graduação em Biologia Animal, Departamento de Zoologia e Botânica, IBILCE, Universidade Estadual Paulista, UNESP, Rua Cristóvão Colombo 2265, CEP 15054 - 000, São José do Rio Preto, SP, Brazil.</mods:affiliation>
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<mods:nameIdentifier id="BEEFF77FEBC9E689A4FF3F61994EE564" type="email">bernabe.t@gmail.com</mods:nameIdentifier>
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<mods:title id="B49FE31D2720CF1136197F8ABDEC4691">Zootaxa</mods:title>
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<mods:part id="6FC1F3C3F2D28EFDEE397A45AE36FF9F">
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<mods:date id="7825E255402E3D298F5631D8DCA5165A">2023</mods:date>
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<mods:number id="3B51C8A316332E4A9729B7EA51A4F4EB">2023-09-28</mods:number>
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<mods:number id="8B901911673862F0C84B0DCA98EA8FA1">5351</mods:number>
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<treatment id="44270927FFA757546C8EF96AFA749B71" ID-DOI="http://doi.org/10.5281/zenodo.8400097" ID-GBIF-Taxon="213635620" ID-Zenodo-Dep="8400097" LSID="urn:lsid:zoobank.org:act:C662C684-A0C1-459C-A8B9-905D9E91CA23" httpUri="http://treatment.plazi.org/id/44270927FFA757546C8EF96AFA749B71" lastPageId="17" lastPageNumber="418" pageId="14" pageNumber="415">
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<subSubSection id="8494EBBAFFA7574B6C8EF96AFE489CD2" box="[151,416,1676,1703]" pageId="14" pageNumber="415" type="nomenclature">
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<paragraph id="CC31B831FFA7574B6C8EF96AFE489CD2" blockId="14.[151,980,1676,1739]" box="[151,416,1676,1703]" pageId="14" pageNumber="415">
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<heading id="97790F5DFFA7574B6C8EF96AFE489CD2" bold="true" box="[151,416,1676,1703]" fontSize="11" level="1" pageId="14" pageNumber="415" reason="1">
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<emphasis id="FEFA6423FFA7574B6C8EF96AFE489CD2" bold="true" box="[151,416,1676,1703]" pageId="14" pageNumber="415">
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<taxonomicName id="0B8EC3B2FFA7574B6C8EF96AFEC29CD3" ID-CoL="BTLWJ" authority="Kury & Bernabé, 2023" authorityName="Kury & Bernabé" authorityYear="2023" box="[151,298,1676,1703]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="14" pageNumber="415" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="FEFA6423FFA7574B6C8EF96AFEC29CD3" bold="true" box="[151,298,1676,1703]" italics="true" pageId="14" pageNumber="415">Titanoleptes</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="E5C9D958FFA7574B6D2BF96AFE489CD2" box="[306,416,1676,1702]" pageId="14" pageNumber="415" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="8494EBBAFFA7574B6C8EF956FCCF9D79" pageId="14" pageNumber="415" type="description">
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<paragraph id="CC31B831FFA7574B6C8EF956FC3C9CBF" blockId="14.[151,980,1676,1739]" box="[151,980,1712,1739]" pageId="14" pageNumber="415">
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<uri id="B81FB433FFA7574B6C8EF956FC3C9CBF" box="[151,980,1712,1739]" pageId="14" pageNumber="415">
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urn:lsid:zoobank.org:act:
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<uuid id="B82882E4FFA7574B6DAFF956FC3C9CBF" box="[438,980,1712,1739]" pageId="14" pageNumber="415">C662C684-A0C1-459C-A8B9-905D9E91CA23</uuid>
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</uri>
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</paragraph>
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<paragraph id="CC31B831FFA7574B6C87F913FCCF9D79" blockId="14.[158,807,1781,1805]" box="[158,807,1781,1805]" pageId="14" pageNumber="415">
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“Undescribed genera SOD-01, SOD-02”—
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<bibRefCitation id="A81FC5C0FFA7574B6E4BF913FCCB9D79" author="Kury, A. B. & Bernabe, T. N. & de Azara, L. N. & Araujo, D. C. & Benedetti, A. R." box="[594,803,1781,1805]" pageId="14" pageNumber="415" pagination="70 - 122" refId="ref16599" refString="Kury, A. B., Bernabe, T. N., de Azara, L. N., Araujo, D. C. & Benedetti, A. R. (2022) Phylogeny of the clade K 92 (Opiliones, Laniatores, Gonyleptidae) with description of a new subfamily and discussion on the evolution of caelopygine facies and sexual dimorphism. Zoologischer Anzeiger, 298, 70 - 122. https: // doi. org / 10.1016 / j. jcz. 2022.03.004" type="journal article" year="2022">
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Kury
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<emphasis id="FEFA6423FFA7574B6E94F913FD299D79" box="[653,705,1781,1805]" italics="true" pageId="14" pageNumber="415">et al.</emphasis>
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2022: 79
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</bibRefCitation>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="8494EBBAFFA7574B6C8EF8DDFADC9D0D" pageId="14" pageNumber="415" type="etymology">
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||
<paragraph id="CC31B831FFA7574B6C8EF8DDFADC9D0D" blockId="14.[151,1436,1851,2021]" pageId="14" pageNumber="415">
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||
<emphasis id="FEFA6423FFA7574B6C8EF8DDFEF69D21" bold="true" box="[151,286,1851,1877]" pageId="14" pageNumber="415">Etymology.</emphasis>
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From the Greek Τῑτάν (one of the titans, divine beings descending from the primordial deities) +
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<emphasis id="FEFA6423FFA7574B6942F8DDFA749D21" box="[1371,1436,1851,1877]" italics="true" pageId="14" pageNumber="415">leptes</emphasis>
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(truncation of generic name
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<taxonomicName id="0B8EC3B2FFA7574B6DCDF8B9FDBA9D0D" authorityName="Kirby" authorityYear="1819" box="[468,594,1887,1913]" class="Arachnida" family="Gonyleptidae" genus="Gonyleptes" kingdom="Animalia" order="Opiliones" pageId="14" pageNumber="415" phylum="Arthropoda" rank="genus">
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<emphasis id="FEFA6423FFA7574B6DCDF8B9FDBA9D0D" box="[468,594,1887,1913]" italics="true" pageId="14" pageNumber="415">Gonyleptes</emphasis>
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</taxonomicName>
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), referring to the very large size of its species. Gender masculine.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="8494EBBAFFA7574B6CDEF865FD559DE9" box="[199,701,1923,1949]" pageId="14" pageNumber="415" type="type_taxon">
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||
<paragraph id="CC31B831FFA7574B6CDEF865FD559DE9" blockId="14.[151,1436,1851,2021]" box="[199,701,1923,1949]" pageId="14" pageNumber="415">
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||
<emphasis id="FEFA6423FFA7574B6CDEF865FE8A9DE9" bold="true" box="[199,354,1923,1949]" pageId="14" pageNumber="415">
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||
<typeStatus id="13350693FFA7574B6CDEF865FEEA9DE9" box="[199,258,1923,1949]" pageId="14" pageNumber="415">Type</typeStatus>
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species.
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||
</emphasis>
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||
<taxonomicName id="0B8EC3B2FFA7574B6D70F865FDB59DE9" ID-CoL="BTLX7" authority="Kury & Bernabé, 2023" authorityName="Kury & Bernabé" authorityYear="2023" box="[361,605,1923,1949]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="14" pageNumber="415" phylum="Arthropoda" rank="species" species="hyperion" status="sp. nov.">
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||
<emphasis id="FEFA6423FFA7574B6D70F865FDB59DE9" box="[361,605,1923,1949]" italics="true" pageId="14" pageNumber="415">Titanoleptes hyperion</emphasis>
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</taxonomicName>
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<emphasis id="FEFA6423FFA7574B6E7DF865FD559DE9" bold="true" box="[612,701,1923,1949]" pageId="14" pageNumber="415">
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<taxonomicNameLabel id="E5C9D958FFA7574B6E7DF865FD559DE9" box="[612,701,1923,1949]" pageId="14" pageNumber="415" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="8494EBBAFFA7574B6CDEF841FCA99D91" pageId="14" pageNumber="415" type="taxon_list">
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<paragraph id="CC31B831FFA7574B6CDEF841FCA99D91" blockId="14.[151,1436,1851,2021]" pageId="14" pageNumber="415">
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<emphasis id="FEFA6423FFA7574B6CDEF841FE709DB5" bold="true" box="[199,408,1959,1985]" pageId="14" pageNumber="415">Included species.</emphasis>
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<taxonomicName id="0B8EC3B2FFA7574B6DBCF841FCAE9DB5" authority="(Roewer, 1913)" baseAuthorityName="Roewer" baseAuthorityYear="1913" box="[421,838,1959,1985]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="14" pageNumber="415" phylum="Arthropoda" rank="species" species="calcar">
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<emphasis id="FEFA6423FFA7574B6DBCF841FD6B9DB5" box="[421,643,1959,1985]" italics="true" pageId="14" pageNumber="415">Titanoleptes calcar</emphasis>
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(
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<bibRefCitation id="A81FC5C0FFA7574B6E8EF841FCD69DB5" author="Roewer, C. - F." box="[663,830,1959,1985]" pageId="14" pageNumber="415" pagination="1 - 256" refId="ref16917" refString="Roewer, C. - F. (1913) Die Familie der Gonyleptiden der Opiliones-Laniatores. Archiv f ¸ r Naturgeschichte, 79 (4), 1 - 256, pl. 1 a." type="journal article" year="1913">Roewer, 1913</bibRefCitation>
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)
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</taxonomicName>
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,
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<taxonomicName id="0B8EC3B2FFA7574B6F41F841FA709DB5" authority="(Soares & Soares, 1946)" authorityName="Kury & Bernabé" authorityYear="2023" baseAuthorityName="Soares & Soares" baseAuthorityYear="1946" box="[856,1432,1959,1985]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="14" pageNumber="415" phylum="Arthropoda" rank="species" species="cupidensis">
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<emphasis id="FEFA6423FFA7574B6F41F841FB8C9DB5" box="[856,1124,1959,1985]" italics="true" pageId="14" pageNumber="415">Titanoleptes cupidensis</emphasis>
|
||
(
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<bibRefCitation id="A81FC5C0FFA7574B6860F841FA789DB5" author="Soares, B. A. M. & Soares, H. E. M." box="[1145,1424,1959,1985]" pageId="14" pageNumber="415" pagination="195 - 212" refId="ref16998" refString="Soares, B. A. M. & Soares, H. E. M. (1946) Novos opilioes do Espirito Santo e um novo opiliao do estado do Para. Papeis Avulsos do Departamento de Zoologia, Secretaria da Agricultura, Sao Paulo, 7, 195 - 212." type="journal article" year="1946">Soares & Soares, 1946</bibRefCitation>
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)
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||
</taxonomicName>
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,
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<taxonomicName id="0B8EC3B2FFA7574B6C8EF82DFEBE9D91" authority="Kury & Bernabé, 2023" authorityName="Kury & Bernabé" authorityYear="2023" box="[151,342,1995,2021]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="14" pageNumber="415" phylum="Arthropoda" rank="species" species="eros" status="sp. nov.">
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<emphasis id="FEFA6423FFA7574B6C8EF82DFEBE9D91" box="[151,342,1995,2021]" italics="true" pageId="14" pageNumber="415">Titanoleptes eros</emphasis>
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||
</taxonomicName>
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||
<emphasis id="FEFA6423FFA7574B6D44F82DFE5F9D91" bold="true" box="[349,439,1995,2021]" pageId="14" pageNumber="415">
|
||
<taxonomicNameLabel id="E5C9D958FFA7574B6D44F82DFE5F9D91" box="[349,439,1995,2021]" pageId="14" pageNumber="415" rank="species">sp. nov.</taxonomicNameLabel>
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||
</emphasis>
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and
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<taxonomicName id="0B8EC3B2FFA7574B6DF4F82DFD099D91" authority="Kury & Bernabé, 2023" authorityName="Kury & Bernabé" authorityYear="2023" box="[493,737,1995,2021]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="14" pageNumber="415" phylum="Arthropoda" rank="species" species="hyperion" status="sp. nov.">
|
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<emphasis id="FEFA6423FFA7574B6DF4F82DFD099D91" box="[493,737,1995,2021]" italics="true" pageId="14" pageNumber="415">Titanoleptes hyperion</emphasis>
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</taxonomicName>
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||
<emphasis id="FEFA6423FFA7574B6EF1F82DFCA99D91" bold="true" box="[744,833,1995,2021]" pageId="14" pageNumber="415">
|
||
<taxonomicNameLabel id="E5C9D958FFA7574B6EF1F82DFCA99D91" box="[744,833,1995,2021]" pageId="14" pageNumber="415" rank="species">sp. nov.</taxonomicNameLabel>
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||
</emphasis>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="8494EBBAFFA6574A6CDEFF70FCA59BFD" pageId="15" pageNumber="416" type="diagnosis">
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||
<paragraph id="CC31B831FFA6574A6CDEFF70FCA59BFD" blockId="15.[151,1437,150,501]" pageId="15" pageNumber="416">
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||
<emphasis id="FEFA6423FFA6574A6CDEFF70FEAA9AC4" bold="true" box="[199,322,150,176]" pageId="15" pageNumber="416">Diagnosis.</emphasis>
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Coxa IV of male without retrolateral apical apophysis (
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<figureCitation id="54B5A4B4FFA6574A6FB6FF71FBE69AC5" box="[943,1038,151,177]" captionStart="FIGURE 16" captionStartId="17.[151,250,1909,1933]" captionTargetBox="[151,1435,625,1885]" captionTargetId="figure-236@17.[151,1435,625,1885]" captionTargetPageId="17" captionText="FIGURE 16. Titanoleptes calcar (Roewer, 1913), (MNRJ 17818) from Sooretama: a. Male, habitus, dorsal view; b. Same, detail of dorsal scutum; c. Male, coxae I–IV, stigmatic area and free sternites, ventral view; d. Male, body, sinistrolateral view; e. Same, frontal view; f. Same, posterior view; g. Female, dorsal scutum, dorsal view; h. Same, sinistrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8391942" httpUri="https://zenodo.org/record/8391942/files/figure.png" pageId="15" pageNumber="416">Fig. 16c</figureCitation>
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), except in
|
||
<taxonomicName id="0B8EC3B2FFA6574A6896FF71FAC89AC5" authorityName="Kury & Bernabé" authorityYear="2023" baseAuthorityName="Soares & Soares" baseAuthorityYear="1946" box="[1167,1312,151,177]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="15" pageNumber="416" phylum="Arthropoda" rank="species" species="cupidensis">
|
||
<emphasis id="FEFA6423FFA6574A6896FF71FAC89AC5" box="[1167,1312,151,177]" italics="true" pageId="15" pageNumber="416">T. cupidensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="54B5A4B4FFA6574A6936FF71FA789AC5" box="[1327,1424,151,177]" captionStart="FIGURE 20" captionStartId="22.[151,250,1276,1300]" captionTargetBox="[151,1435,181,1252]" captionTargetId="figure-20@22.[151,1435,181,1252]" captionTargetPageId="22" captionText="FIGURE 20. Titanoleptes cupidensis (Soares & Soares, 1946), male (MNRJ 17869), from Sooretama:a. Left coxa andtrochanter IV, dorsal view; b. Left patella IV, prolateral view; c. Left coxa to proximal femur IV, prolateral view; d. Left leg IV, ventral view. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.8391950" httpUri="https://zenodo.org/record/8391950/files/figure.png" pageId="15" pageNumber="416">Fig. 20a</figureCitation>
|
||
). Trochanter IV of male armed with a strong, retrolateral, apical apophysis (
|
||
<figureCitation id="54B5A4B4FFA6574A6FD6FF5DFBBB9AA1" box="[975,1107,187,213]" captionStart="FIGURE 21" captionStartId="23.[151,250,1838,1862]" captionTargetBox="[151,1435,417,1815]" captionTargetId="figure-118@23.[151,1435,417,1815]" captionTargetPageId="23" captionText="FIGURE 21. Titanoleptes cupidensis (Soares & Soares, 1946), male (MNRJ 17869), from Sooretama, left Fe IV: a. Dorsal view; b. Retrolateral (external) view; c. Ventral view; d. Prolateral (internal) view. Anterior direction upwards in a to c. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8391953" httpUri="https://zenodo.org/record/8391953/files/figure.png" pageId="15" pageNumber="416">Figs. 21a, c</figureCitation>
|
||
). Patella and tibia IV covered with yellow acuminate tubercles (
|
||
<figureCitation id="54B5A4B4FFA6574A6E16FF39FD939A8D" box="[527,635,223,249]" captionStart="FIGURE 20" captionStartId="22.[151,250,1276,1300]" captionTargetBox="[151,1435,181,1252]" captionTargetId="figure-20@22.[151,1435,181,1252]" captionTargetPageId="22" captionText="FIGURE 20. Titanoleptes cupidensis (Soares & Soares, 1946), male (MNRJ 17869), from Sooretama:a. Left coxa andtrochanter IV, dorsal view; b. Left patella IV, prolateral view; c. Left coxa to proximal femur IV, prolateral view; d. Left leg IV, ventral view. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.8391950" httpUri="https://zenodo.org/record/8391950/files/figure.png" pageId="15" pageNumber="416">Figs. 20d</figureCitation>
|
||
,
|
||
<figureCitation id="54B5A4B4FFA6574A6E9FFF39FD5C9A8D" box="[646,692,223,249]" captionStart="FIGURE 24" captionStartId="26.[151,250,1463,1487]" captionTargetBox="[151,1436,181,1439]" captionTargetId="figure-20@26.[151,1436,181,1439]" captionTargetPageId="26" captionText="FIGURE 24. Titanoleptes eros sp. nov., male paratype (MNRJ 7077) from ES, Santa Teresa, free tergites and proximal left leg IV: a. Dorsal view; b. Prolateral view; c. Ventral view; d. Retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8391959" httpUri="https://zenodo.org/record/8391959/files/figure.png" pageId="15" pageNumber="416">24b</figureCitation>
|
||
). Femur IV of male either substraight (
|
||
<figureCitation id="54B5A4B4FFA6574A687DFF39FB299A8D" box="[1124,1217,223,249]" captionStart="FIGURE 19" captionStartId="21.[151,250,1943,1967]" captionTargetBox="[181,1406,181,1919]" captionTargetId="figure-17@21.[181,1406,181,1919]" captionTargetPageId="21" captionText="FIGURE 19. Titanoleptes cupidensis (Soares & Soares, 1946), (MNRJ 17869), male (A–F), and female (G, H), from Sooretama: a. Male, habitus, dorsal view; b. Same, detail of dorsal scutum; c. Same, ventral view; d. Same, dextrolateral view; e. Same, posterior view; f. Same, frontal view; g. Female, habitus, dextrolateral view; h. Same, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8391948" httpUri="https://zenodo.org/record/8391948/files/figure.png" pageId="15" pageNumber="416">Fig. 19a</figureCitation>
|
||
) or strongly curved to the midline on basal 1/
|
||
<quantity id="0B7615D4FFA6574A6DA0FEE5FE009B69" box="[441,488,259,285]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="15" pageNumber="416" unit="in" value="4.0">4 in</quantity>
|
||
dorsal view (
|
||
<figureCitation id="54B5A4B4FFA6574A6E9DFEE5FCE49B69" box="[644,780,259,285]" captionStart="FIGURE 17" captionStartId="18.[151,250,1209,1233]" captionTargetBox="[151,1436,181,1185]" captionTargetId="figure-20@18.[151,1436,181,1185]" captionTargetPageId="18" captionText="FIGURE 17. Titanoleptes calcar (Roewer, 1913), male (MNRJ 17818) from Sooretama: a. Left femur IV, dorsal view; b. Same, prolateral view; c. Same, ventral view; d. Same, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8391944" httpUri="https://zenodo.org/record/8391944/files/figure.png" pageId="15" pageNumber="416">Figs. 17a, c</figureCitation>
|
||
). Tibia IV armed with rows of acuminate tubercles (
|
||
<figureCitation id="54B5A4B4FFA6574A697DFEE5FF2C9B35" captionStart="FIGURE 20" captionStartId="22.[151,250,1276,1300]" captionTargetBox="[151,1435,181,1252]" captionTargetId="figure-20@22.[151,1435,181,1252]" captionTargetPageId="22" captionText="FIGURE 20. Titanoleptes cupidensis (Soares & Soares, 1946), male (MNRJ 17869), from Sooretama:a. Left coxa andtrochanter IV, dorsal view; b. Left patella IV, prolateral view; c. Left coxa to proximal femur IV, prolateral view; d. Left leg IV, ventral view. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.8391950" httpUri="https://zenodo.org/record/8391950/files/figure.png" pageId="15" pageNumber="416">Figs. 20d</figureCitation>
|
||
,
|
||
<figureCitation id="54B5A4B4FFA6574A6CD6FEC1FF159B35" box="[207,253,295,321]" captionStart="FIGURE 24" captionStartId="26.[151,250,1463,1487]" captionTargetBox="[151,1436,181,1439]" captionTargetId="figure-20@26.[151,1436,181,1439]" captionTargetPageId="26" captionText="FIGURE 24. Titanoleptes eros sp. nov., male paratype (MNRJ 7077) from ES, Santa Teresa, free tergites and proximal left leg IV: a. Dorsal view; b. Prolateral view; c. Ventral view; d. Retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8391959" httpUri="https://zenodo.org/record/8391959/files/figure.png" pageId="15" pageNumber="416">24b</figureCitation>
|
||
). VP of penis with ear-flaps small, triangular, ectal vertex connected but not fused with flange. Flabellum quadrangular, as a stylized shell, proximal sides gently arched, non-serrate, distal half extremely reduced, with distal sides coalesced into one, arched and deeply serrate (
|
||
<figureCitation id="54B5A4B4FFA6574A6EF9FE89FCA89BFD" box="[736,832,367,393]" captionStart="FIGURE 22" captionStartId="24.[151,250,1655,1679]" captionTargetBox="[151,1436,181,1632]" captionTargetId="figure-20@24.[151,1436,181,1632]" captionTargetPageId="24" captionText="FIGURE 22. Titanoleptes cupidensis (Soares & Soares, 1946), (MNRJ 17869), from Sooretama, male genitalia, distal part: a. Dorsal view; b. Ventral view; c. Dextrolateral view; d. Same, panoramic; e. Latero-apical view; f. Dorso-apical view; g. Glans, dextrolateral view. Scale bars = 100 μm (a, b, d, e), 50 μm (c, f, g)." figureDoi="http://doi.org/10.5281/zenodo.8391955" httpUri="https://zenodo.org/record/8391955/files/figure.png" pageId="15" pageNumber="416">Fig. 22e</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8494EBBAFFA6574A6CDEFE74FDC49B81" pageId="15" pageNumber="416" type="distribution">
|
||
<paragraph id="CC31B831FFA6574A6CDEFE74FDC49B81" blockId="15.[151,1437,150,501]" pageId="15" pageNumber="416">
|
||
<emphasis id="FEFA6423FFA6574A6CDEFE74FEB19BD8" bold="true" box="[199,345,402,428]" pageId="15" pageNumber="416">Distribution</emphasis>
|
||
(
|
||
<figureCitation id="54B5A4B4FFA6574A6D75FE75FE1A9BD9" box="[364,498,402,429]" captionStart-0="FIGURE 14" captionStart-1="FIGURE 15" captionStartId-0="15.[151,250,1612,1636]" captionStartId-1="16.[151,250,1468,1492]" captionTargetBox-0="[151,1435,525,1588]" captionTargetBox-1="[151,1435,381,1444]" captionTargetId-0="figure-176@15.[151,1435,525,1588]" captionTargetId-1="figure-141@16.[151,1435,381,1444]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionText-0="FIGURE 14. Eastern Brazil, showing the distribution of the two robust species of Titanoleptes gen. nov., which are allopatrid, distributed in opposite margins of the Doce river." captionText-1="FIGURE 15. Eastern Brazil, showing the distribution of the two gracile species of Titanoleptes gen. nov. which are allopatrid, distributed in opposite margins of the Doce River. These two species are less common in collections. The extension of the map is the same as that of the previous one for comparison." figureDoi-0="http://doi.org/10.5281/zenodo.8391938" figureDoi-1="http://doi.org/10.5281/zenodo.8391940" httpUri-0="https://zenodo.org/record/8391938/files/figure.png" httpUri-1="https://zenodo.org/record/8391940/files/figure.png" pageId="15" pageNumber="416">Figs 14–15</figureCitation>
|
||
). Southern
|
||
<collectingRegion id="0E4A76D3FFA6574A6E61FE75FD149BD9" box="[632,764,403,429]" country="Brazil" name="Bahia" pageId="15" pageNumber="416">Bahia State</collectingRegion>
|
||
and
|
||
<collectingRegion id="0E4A76D3FFA6574A6F20FE75FBCB9BD9" box="[825,1059,403,429]" country="Brazil" name="Espirito Santo" pageId="15" pageNumber="416">Espírito Santo State</collectingRegion>
|
||
. All four species of
|
||
<taxonomicName id="0B8EC3B2FFA6574A690AFE75FA749BD9" authorityName="Kury & Bernabé" authorityYear="2023" box="[1299,1436,403,429]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="15" pageNumber="416" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FEFA6423FFA6574A690AFE75FA749BD9" box="[1299,1436,403,429]" italics="true" pageId="15" pageNumber="416">Titanoleptes</emphasis>
|
||
</taxonomicName>
|
||
occur in the WWF ecoregion NT0103 (
|
||
<collectingRegion id="0E4A76D3FFA6574A6E52FE51FD669BA5" box="[587,654,439,465]" country="Brazil" name="Bahia" pageId="15" pageNumber="416">Bahia</collectingRegion>
|
||
coastal forests) and two subsets of two species each are distributed on opposite margins of the Doce River.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="98F1E8B9FFA6574A6C8EF9AAFD629CFC" ID-DOI="http://doi.org/10.5281/zenodo.8391938" ID-Zenodo-Dep="8391938" httpUri="https://zenodo.org/record/8391938/files/figure.png" pageId="15" pageNumber="416" startId="15.[151,250,1612,1636]" targetBox="[151,1435,525,1588]" targetPageId="15" targetType="figure">
|
||
<paragraph id="CC31B831FFA6574A6C8EF9AAFD629CFC" blockId="15.[151,1437,1612,1672]" pageId="15" pageNumber="416">
|
||
<emphasis id="FEFA6423FFA6574A6C8EF9AAFEC89C10" bold="true" box="[151,288,1612,1636]" pageId="15" pageNumber="416">FIGURE 14.</emphasis>
|
||
Eastern Brazil, showing the distribution of the two robust species of
|
||
<taxonomicName id="0B8EC3B2FFA6574A6FC4F9AAFBB19C10" authority="Kury & Bernabé, 2023" authorityName="Kury & Bernabé" authorityYear="2023" box="[989,1113,1612,1636]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="15" pageNumber="416" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="FEFA6423FFA6574A6FC4F9AAFBB19C10" box="[989,1113,1612,1636]" italics="true" pageId="15" pageNumber="416">Titanoleptes</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="FEFA6423FFA6574A6879F9AAFB569C10" bold="true" box="[1120,1214,1612,1636]" pageId="15" pageNumber="416">
|
||
<taxonomicNameLabel id="E5C9D958FFA6574A6879F9AAFB569C10" box="[1120,1214,1612,1636]" pageId="15" pageNumber="416" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, which are allopatrid, distributed in opposite margins of the Doce river.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="8494EBBAFFA657556CDEF950FAFF9B11" lastPageId="16" lastPageNumber="417" pageId="15" pageNumber="416" type="discussion">
|
||
<paragraph id="CC31B831FFA657556CDEF950FAFF9B11" blockId="15.[151,1437,1717,2033]" lastBlockId="16.[151,1437,151,357]" lastPageId="16" lastPageNumber="417" pageId="15" pageNumber="416">
|
||
<emphasis id="FEFA6423FFA6574A6CDEF950FBF89CA4" bold="true" box="[199,1040,1717,1744]" pageId="15" pageNumber="416">
|
||
Internal and external relationships of the four species of
|
||
<taxonomicName id="0B8EC3B2FFA6574A6F65F953FBE19CA4" authorityName="Kury & Bernabé" authorityYear="2023" box="[892,1033,1717,1744]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="15" pageNumber="416" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FEFA6423FFA6574A6F65F953FBE19CA4" bold="true" box="[892,1033,1717,1744]" italics="true" pageId="15" pageNumber="416">Titanoleptes</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
At a first glance, one could easily identify two forms in
|
||
<taxonomicName id="0B8EC3B2FFA6574A6D88F93CFDF29C80" authority="Kury & Bernabé, 2023" authorityName="Kury & Bernabé" authorityYear="2023" box="[401,538,1754,1780]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="15" pageNumber="416" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="FEFA6423FFA6574A6D88F93CFDF29C80" box="[401,538,1754,1780]" italics="true" pageId="15" pageNumber="416">Titanoleptes</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="FEFA6423FFA6574A6E3BF93CFD649C80" bold="true" box="[546,652,1754,1780]" pageId="15" pageNumber="416">
|
||
<taxonomicNameLabel id="E5C9D958FFA6574A6E3BF93CFD649C80" box="[546,652,1754,1780]" pageId="15" pageNumber="416" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: (1)
|
||
<taxonomicName id="0B8EC3B2FFA6574A6EDCF93DFCF99C80" authority="Kury & Bernabé, 2023" authorityName="Kury & Bernabé" authorityYear="2023" box="[709,785,1755,1780]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="15" pageNumber="416" phylum="Arthropoda" rank="species" species="eros" status="sp. nov.">
|
||
<emphasis id="FEFA6423FFA6574A6EDCF93DFCF99C80" box="[709,785,1755,1780]" italics="true" pageId="15" pageNumber="416">T. eros</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="FEFA6423FFA6574A6F00F93CFC9C9C80" bold="true" box="[793,884,1754,1780]" pageId="15" pageNumber="416">
|
||
<taxonomicNameLabel id="E5C9D958FFA6574A6F00F93CFC9C9C80" box="[793,884,1754,1780]" pageId="15" pageNumber="416" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="0B8EC3B2FFA6574A6FB4F93DFBA89C80" authorityName="Kury & Bernabé" authorityYear="2023" baseAuthorityName="Soares & Soares" baseAuthorityYear="1946" box="[941,1088,1754,1780]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="15" pageNumber="416" phylum="Arthropoda" rank="species" species="cupidensis">
|
||
<emphasis id="FEFA6423FFA6574A6FB4F93DFBA89C80" box="[941,1088,1754,1780]" italics="true" pageId="15" pageNumber="416">T. cupidensis</emphasis>
|
||
</taxonomicName>
|
||
are gracile and have a median apophysis on free tergite III in both sexes, substraight femur IV, and naked glans; (2)
|
||
<taxonomicName id="0B8EC3B2FFA6574A6898F919FAEC9D6C" authority="Kury & Bernabé, 2023" authorityName="Kury & Bernabé" authorityYear="2023" box="[1153,1284,1790,1816]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="15" pageNumber="416" phylum="Arthropoda" rank="species" species="hyperion" status="sp. nov.">
|
||
<emphasis id="FEFA6423FFA6574A6898F919FAEC9D6C" box="[1153,1284,1790,1816]" italics="true" pageId="15" pageNumber="416">T. hyperion</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="FEFA6423FFA6574A6914F918FA829D6C" bold="true" box="[1293,1386,1790,1816]" pageId="15" pageNumber="416">
|
||
<taxonomicNameLabel id="E5C9D958FFA6574A6914F918FA829D6C" box="[1293,1386,1790,1816]" pageId="15" pageNumber="416" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="0B8EC3B2FFA6574A6C8EF8C5FF149D48" baseAuthorityName="Roewer" baseAuthorityYear="1913" box="[151,252,1826,1852]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="15" pageNumber="416" phylum="Arthropoda" rank="species" species="calcar">
|
||
<emphasis id="FEFA6423FFA6574A6C8EF8C5FF149D48" box="[151,252,1826,1852]" italics="true" pageId="15" pageNumber="416">T. calcar</emphasis>
|
||
</taxonomicName>
|
||
are robust and lack an apophysis on free tergite III of males, have a heavily curved femur IV, and have a unique scutum covering the glans. The parapatrid pattern of these two pairs—
|
||
<taxonomicName id="0B8EC3B2FFA6574A6859F8A1FB679D14" authorityName="Kury & Bernabé" authorityYear="2023" box="[1088,1167,1863,1888]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="15" pageNumber="416" phylum="Arthropoda" rank="species" species="eros">
|
||
<emphasis id="FEFA6423FFA6574A6859F8A1FB679D14" box="[1088,1167,1863,1888]" italics="true" pageId="15" pageNumber="416">T. eros</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName id="0B8EC3B2FFA6574A68ADF8A1FAA19D14" authorityName="Kury & Bernabé" authorityYear="2023" baseAuthorityName="Soares & Soares" baseAuthorityYear="1946" box="[1204,1353,1862,1888]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="15" pageNumber="416" phylum="Arthropoda" rank="species" species="cupidensis">
|
||
<emphasis id="FEFA6423FFA6574A68ADF8A1FAA19D14" box="[1204,1353,1862,1888]" italics="true" pageId="15" pageNumber="416">T. cupidensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="0B8EC3B2FFA6574A6991F8A1FF139DF0" authorityName="Kury & Bernabé" authorityYear="2023" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="15" pageNumber="416" phylum="Arthropoda" rank="species" species="hyperion">
|
||
<emphasis id="FEFA6423FFA6574A6991F8A1FF139DF0" italics="true" pageId="15" pageNumber="416">T. hyperion</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName id="0B8EC3B2FFA6574A6D04F88DFE6B9DF0" baseAuthorityName="Roewer" baseAuthorityYear="1913" box="[285,387,1898,1924]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="15" pageNumber="416" phylum="Arthropoda" rank="species" species="calcar">
|
||
<emphasis id="FEFA6423FFA6574A6D04F88DFE6B9DF0" box="[285,387,1898,1924]" italics="true" pageId="15" pageNumber="416">T. calcar</emphasis>
|
||
</taxonomicName>
|
||
are found on opposite margins of the Doce River (
|
||
<figureCitation id="54B5A4B4FFA6574A6FCCF88CFBB39DF0" box="[981,1115,1898,1924]" captionStart-0="FIGURE 13" captionStart-1="FIGURE 14" captionStartId-0="14.[151,250,936,960]" captionStartId-1="15.[151,250,1612,1636]" captionTargetBox-0="[151,1436,151,921]" captionTargetBox-1="[151,1435,525,1588]" captionTargetId-0="figure-22@14.[151,1436,199,913]" captionTargetId-1="figure-176@15.[151,1435,525,1588]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionText-0="FIGURE 13. Cajango vindicator sp. nov., male holotype (MNRJ 9053), from Camacan, genital morphology: a. Penis, distal part, dorsal view; b. Same, sinistrolateral view; c. Same, ventral view." captionText-1="FIGURE 14. Eastern Brazil, showing the distribution of the two robust species of Titanoleptes gen. nov., which are allopatrid, distributed in opposite margins of the Doce river." figureDoi-0="http://doi.org/10.5281/zenodo.8391934" figureDoi-1="http://doi.org/10.5281/zenodo.8391938" httpUri-0="https://zenodo.org/record/8391934/files/figure.png" httpUri-1="https://zenodo.org/record/8391938/files/figure.png" pageId="15" pageNumber="416">Figs. 13–14</figureCitation>
|
||
)—is highly suggestive of a sister relationship between each of the members of these pairs, regardless of whether the pairs themselves form a more inclusive clade or not. This hypothesis, though, is not corroborated by any of the analyses. The cladistic analysis of
|
||
<bibRefCitation id="A81FC5C0FFA6574A6D03F831FE019D84" author="Kury, A. B. & Bernabe, T. N. & de Azara, L. N. & Araujo, D. C. & Benedetti, A. R." box="[282,489,2006,2033]" pageId="15" pageNumber="416" pagination="70 - 122" refId="ref16599" refString="Kury, A. B., Bernabe, T. N., de Azara, L. N., Araujo, D. C. & Benedetti, A. R. (2022) Phylogeny of the clade K 92 (Opiliones, Laniatores, Gonyleptidae) with description of a new subfamily and discussion on the evolution of caelopygine facies and sexual dimorphism. Zoologischer Anzeiger, 298, 70 - 122. https: // doi. org / 10.1016 / j. jcz. 2022.03.004" type="journal article" year="2022">
|
||
Kury
|
||
<emphasis id="FEFA6423FFA6574A6D45F831FE7E9D84" box="[348,406,2006,2032]" italics="true" pageId="15" pageNumber="416">et al.</emphasis>
|
||
(2022)
|
||
</bibRefCitation>
|
||
explored different parameters to assess the arrangement of the four
|
||
<taxonomicName id="0B8EC3B2FFA6574A68ECF830FA969D84" authorityName="Kury & Bernabé" authorityYear="2023" box="[1269,1406,2006,2032]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="15" pageNumber="416" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FEFA6423FFA6574A68ECF830FA969D84" box="[1269,1406,2006,2032]" italics="true" pageId="15" pageNumber="416">Titanoleptes</emphasis>
|
||
</taxonomicName>
|
||
in the phylogeny of Sodreaninae. The favored analysis using the setk script (with implied weights and K value ca. 12) resolved
|
||
<taxonomicName id="0B8EC3B2FFB957556CE5FF5DFE6D9AA1" authorityName="Kury & Bernabé" authorityYear="2023" box="[252,389,187,213]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="417" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FEFA6423FFB957556CE5FF5DFE6D9AA1" box="[252,389,187,213]" italics="true" pageId="16" pageNumber="417">Titanoleptes</emphasis>
|
||
</taxonomicName>
|
||
as a sister clade of
|
||
<taxonomicName id="0B8EC3B2FFB957556E43FF5DFD3D9AA1" box="[602,725,187,213]" class="Arachnida" family="Gonyleptidae" genus="Friburgoia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="417" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FEFA6423FFB957556E43FF5DFD3D9AA1" box="[602,725,187,213]" italics="true" pageId="16" pageNumber="417">Friburgoia</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="0B8EC3B2FFB957556EC7FF5DFC999AA1" box="[734,881,187,213]" class="Arachnida" family="Gonyleptidae" genus="Urodiabunus" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="417" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FEFA6423FFB957556EC7FF5DFC999AA1" box="[734,881,187,213]" italics="true" pageId="16" pageNumber="417">Urodiabunus</emphasis>
|
||
</taxonomicName>
|
||
, with an internal branching sequence of
|
||
<taxonomicName id="0B8EC3B2FFB957556928FF5DFA939AA0" authorityName="Kury & Bernabé" authorityYear="2023" box="[1329,1403,187,212]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="eros">
|
||
<emphasis id="FEFA6423FFB957556928FF5DFA939AA0" box="[1329,1403,187,212]" italics="true" pageId="16" pageNumber="417">T. eros</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="0B8EC3B2FFB957556991FF5DFEE59A8D" authorityName="Kury & Bernabé" authorityYear="2023" baseAuthorityName="Soares & Soares" baseAuthorityYear="1946" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="cupidensis">
|
||
<emphasis id="FEFA6423FFB957556991FF5DFEE59A8D" italics="true" pageId="16" pageNumber="417">T. cupidensis</emphasis>
|
||
</taxonomicName>
|
||
, and then
|
||
<taxonomicName id="0B8EC3B2FFB957556D98FF39FDE89A8D" authorityName="Kury & Bernabé" authorityYear="2023" box="[385,512,223,249]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="hyperion">
|
||
<emphasis id="FEFA6423FFB957556D98FF39FDE89A8D" box="[385,512,223,249]" italics="true" pageId="16" pageNumber="417">T. hyperion</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName id="0B8EC3B2FFB957556E05FF39FD979A8D" baseAuthorityName="Roewer" baseAuthorityYear="1913" box="[540,639,223,249]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="calcar">
|
||
<emphasis id="FEFA6423FFB957556E05FF39FD979A8D" box="[540,639,223,249]" italics="true" pageId="16" pageNumber="417">T. calcar</emphasis>
|
||
</taxonomicName>
|
||
. IE settings of K = 1 and 5, plus equal weights, yielded the same result. However, Bayesian inference did not recover
|
||
<taxonomicName id="0B8EC3B2FFB957556E80FEE5FCCA9B69" authorityName="Kury & Bernabé" authorityYear="2023" box="[665,802,259,285]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="417" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FEFA6423FFB957556E80FEE5FCCA9B69" box="[665,802,259,285]" italics="true" pageId="16" pageNumber="417">Titanoleptes</emphasis>
|
||
</taxonomicName>
|
||
at all, finding only the clade
|
||
<taxonomicName id="0B8EC3B2FFB957556875FEE5FB049B69" authorityName="Kury & Bernabé" authorityYear="2023" box="[1132,1260,259,285]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="hyperion">
|
||
<emphasis id="FEFA6423FFB957556875FEE5FB049B69" box="[1132,1260,259,285]" italics="true" pageId="16" pageNumber="417">T. hyperion</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName id="0B8EC3B2FFB957556910FEE5FA859B69" baseAuthorityName="Roewer" baseAuthorityYear="1913" box="[1289,1389,259,285]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="calcar">
|
||
<emphasis id="FEFA6423FFB957556910FEE5FA859B69" box="[1289,1389,259,285]" italics="true" pageId="16" pageNumber="417">T. calcar</emphasis>
|
||
</taxonomicName>
|
||
and then recovering
|
||
<taxonomicName id="0B8EC3B2FFB957556D57FEC1FE719B34" authorityName="Kury & Bernabé" authorityYear="2023" box="[334,409,295,320]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="eros">
|
||
<emphasis id="FEFA6423FFB957556D57FEC1FE719B34" box="[334,409,295,320]" italics="true" pageId="16" pageNumber="417">T. eros</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="0B8EC3B2FFB957556DC8FEC1FD8B9B35" authorityName="Kury & Bernabé" authorityYear="2023" baseAuthorityName="Soares & Soares" baseAuthorityYear="1946" box="[465,611,295,321]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="cupidensis">
|
||
<emphasis id="FEFA6423FFB957556DC8FEC1FD8B9B35" box="[465,611,295,321]" italics="true" pageId="16" pageNumber="417">T. cupidensis</emphasis>
|
||
</taxonomicName>
|
||
in a trichotomy with the rest of the Sodreaninae. We have decided not to arrange the four species of
|
||
<taxonomicName id="0B8EC3B2FFB957556DDEFEADFDB89B11" authorityName="Kury & Bernabé" authorityYear="2023" box="[455,592,331,357]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="417" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FEFA6423FFB957556DDEFEADFDB89B11" box="[455,592,331,357]" italics="true" pageId="16" pageNumber="417">Titanoleptes</emphasis>
|
||
</taxonomicName>
|
||
into two genera, despite the weak support for their monophyly.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="98F1E8B9FFB957556C8EFA5AFD299C68" ID-DOI="http://doi.org/10.5281/zenodo.8391940" ID-Zenodo-Dep="8391940" httpUri="https://zenodo.org/record/8391940/files/figure.png" pageId="16" pageNumber="417" startId="16.[151,250,1468,1492]" targetBox="[151,1435,381,1444]" targetPageId="16" targetType="figure">
|
||
<paragraph id="CC31B831FFB957556C8EFA5AFD299C68" blockId="16.[151,1437,1468,1564]" pageId="16" pageNumber="417">
|
||
<emphasis id="FEFA6423FFB957556C8EFA5AFEC89FA0" bold="true" box="[151,288,1468,1492]" pageId="16" pageNumber="417">FIGURE 15.</emphasis>
|
||
Eastern Brazil, showing the distribution of the two gracile species of
|
||
<taxonomicName id="0B8EC3B2FFB957556FFAFA5AFBB79FA0" authority="Kury & Bernabé, 2023" authorityName="Kury & Bernabé" authorityYear="2023" box="[995,1119,1468,1492]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="417" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="FEFA6423FFB957556FFAFA5AFBB79FA0" box="[995,1119,1468,1492]" italics="true" pageId="16" pageNumber="417">Titanoleptes</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="FEFA6423FFB95755687FFA5AFB2C9FA0" bold="true" box="[1126,1220,1468,1492]" pageId="16" pageNumber="417">
|
||
<taxonomicNameLabel id="E5C9D958FFB95755687FFA5AFB2C9FA0" box="[1126,1220,1468,1492]" pageId="16" pageNumber="417" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
which are allopatrid, distributed in opposite margins of the Doce River. These two species are less common in collections. The extension of the map is the same as that of the previous one for comparison.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="8494EBBAFFB957546C8EF988FA749B71" lastPageId="17" lastPageNumber="418" pageId="16" pageNumber="417" type="key">
|
||
<paragraph id="CC31B831FFB957556C8EF988FDD89CFC" blockId="16.[151,560,1645,1672]" box="[151,560,1645,1672]" pageId="16" pageNumber="417">
|
||
<emphasis id="FEFA6423FFB957556C8EF988FDD89CFC" bold="true" box="[151,560,1645,1672]" pageId="16" pageNumber="417">
|
||
Key to the species of
|
||
<taxonomicName id="0B8EC3B2FFB957556D84F98BFDD89CFC" ID-CoL="BTLWJ" authorityName="Kury & Bernabé" authorityYear="2023" box="[413,560,1645,1672]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="417" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FEFA6423FFB957556D84F98BFDD89CFC" bold="true" box="[413,560,1645,1672]" italics="true" pageId="16" pageNumber="417">Titanoleptes</emphasis>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
</paragraph>
|
||
<keyStep id="777AA094FFB957556C8EF955FA749D0C" pageId="16" pageNumber="417">
|
||
<paragraph id="CC31B831FFB957556C8EF955FA749D54" blockId="16.[151,1436,1715,2030]" pageId="16" pageNumber="417">
|
||
<keyLead id="777F1B04FFB957556C8EF955FA749D54" pageId="16" pageNumber="417">
|
||
1. Ocularium armed with two small spherical tubercles of lighter color contrasting with background (
|
||
<figureCitation id="54B5A4B4FFB9575568B8F955FB169CBD" box="[1185,1278,1715,1737]" captionStart="FIGURE 16" captionStartId="17.[151,250,1909,1933]" captionTargetBox="[151,1435,625,1885]" captionTargetId="figure-236@17.[151,1435,625,1885]" captionTargetPageId="17" captionText="FIGURE 16. Titanoleptes calcar (Roewer, 1913), (MNRJ 17818) from Sooretama: a. Male, habitus, dorsal view; b. Same, detail of dorsal scutum; c. Male, coxae I–IV, stigmatic area and free sternites, ventral view; d. Male, body, sinistrolateral view; e. Same, frontal view; f. Same, posterior view; g. Female, dorsal scutum, dorsal view; h. Same, sinistrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8391942" httpUri="https://zenodo.org/record/8391942/files/figure.png" pageId="16" pageNumber="417">
|
||
Figs.
|
||
<quantity id="0B7615D4FFB9575568C1F955FB169CBD" box="[1240,1278,1715,1737]" metricMagnitude="-2" metricUnit="kg" metricValue="1.6" pageId="16" pageNumber="417" unit="g" value="16.0">16g</quantity>
|
||
</figureCitation>
|
||
,
|
||
<figureCitation id="54B5A4B4FFB957556910F955FAD99CBD" box="[1289,1329,1715,1737]" captionStart="FIGURE 27" captionStartId="29.[151,250,1378,1402]" captionTargetBox="[151,1435,181,1354]" captionTargetId="figure-17@29.[151,1435,181,1354]" captionTargetPageId="29" captionText="FIGURE 27. Titanoleptes hyperion sp. nov., male paratype (MNRJ 18669) ES, Santa Teresa: a. Habitus, panoramic, dorsal view; b. Habitus, dorsal view; c. Same, ventral view; d. Same, sinistrolateral view; e. Same, frontal view; f. Same, posterior view. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.8391967" httpUri="https://zenodo.org/record/8391967/files/figure.png" pageId="16" pageNumber="417">27b</figureCitation>
|
||
); femur IV of male strongly curved to the median line at proximal one quarter, armed at least with two strong retrolateral spines (
|
||
<figureCitation id="54B5A4B4FFB957556951F936FA709C91" box="[1352,1432,1744,1766]" captionStart="FIGURE 17" captionStartId="18.[151,250,1209,1233]" captionTargetBox="[151,1436,181,1185]" captionTargetId="figure-20@18.[151,1436,181,1185]" captionTargetPageId="18" captionText="FIGURE 17. Titanoleptes calcar (Roewer, 1913), male (MNRJ 17818) from Sooretama: a. Left femur IV, dorsal view; b. Same, prolateral view; c. Same, ventral view; d. Same, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8391944" httpUri="https://zenodo.org/record/8391944/files/figure.png" pageId="16" pageNumber="417">Figs. 17</figureCitation>
|
||
,
|
||
<figureCitation id="54B5A4B4FFB957556CFBF908FF159D77" box="[226,253,1774,1795]" captionStart="FIGURE 28" captionStartId="30.[151,250,1804,1828]" captionTargetBox="[151,1436,211,1667]" captionTargetId="figure-20@30.[151,1436,211,1667]" captionTargetPageId="30" captionText="FIGURE 28. Titanoleptes hyperion sp. nov., male holotype (MNRJ 9076), from Domingos Martins, femur IV (each figure is rotated 90° along the proximal-distal axis relative to each adjacent one). From the top: retrolateral view. ventral view, prolateral view and dorsal view. All scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8391970" httpUri="https://zenodo.org/record/8391970/files/figure.png" pageId="16" pageNumber="417">28</figureCitation>
|
||
); free tergite III unarmed in males, either unarmed (
|
||
<taxonomicName id="0B8EC3B2FFB957556EF4F908FCB19D77" ID-CoL="BTLX7" authorityName="Kury & Bernabé" authorityYear="2023" box="[749,857,1774,1795]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="hyperion">
|
||
<emphasis id="FEFA6423FFB957556EF4F908FCB19D77" box="[749,857,1774,1795]" italics="true" pageId="16" pageNumber="417">T. hyperion</emphasis>
|
||
</taxonomicName>
|
||
) or with median apophysis (
|
||
<taxonomicName id="0B8EC3B2FFB957556872F908FB579D77" ID-CoL="BTLX4" baseAuthorityName="Roewer" baseAuthorityYear="1913" box="[1131,1215,1774,1795]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="calcar">
|
||
<emphasis id="FEFA6423FFB957556872F908FB579D77" box="[1131,1215,1774,1795]" italics="true" pageId="16" pageNumber="417">T. calcar</emphasis>
|
||
</taxonomicName>
|
||
) in females (
|
||
<figureCitation id="54B5A4B4FFB957556925F908FF189D54" captionStart="FIGURE 16" captionStartId="17.[151,250,1909,1933]" captionTargetBox="[151,1435,625,1885]" captionTargetId="figure-236@17.[151,1435,625,1885]" captionTargetPageId="17" captionText="FIGURE 16. Titanoleptes calcar (Roewer, 1913), (MNRJ 17818) from Sooretama: a. Male, habitus, dorsal view; b. Same, detail of dorsal scutum; c. Male, coxae I–IV, stigmatic area and free sternites, ventral view; d. Male, body, sinistrolateral view; e. Same, frontal view; f. Same, posterior view; g. Female, dorsal scutum, dorsal view; h. Same, sinistrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8391942" httpUri="https://zenodo.org/record/8391942/files/figure.png" pageId="16" pageNumber="417">Figs. 16b, g</figureCitation>
|
||
,
|
||
<figureCitation id="54B5A4B4FFB957556CE3F8EDFEF79D55" box="[250,287,1803,1825]" captionStart="FIGURE 27" captionStartId="29.[151,250,1378,1402]" captionTargetBox="[151,1435,181,1354]" captionTargetId="figure-17@29.[151,1435,181,1354]" captionTargetPageId="29" captionText="FIGURE 27. Titanoleptes hyperion sp. nov., male paratype (MNRJ 18669) ES, Santa Teresa: a. Habitus, panoramic, dorsal view; b. Habitus, dorsal view; c. Same, ventral view; d. Same, sinistrolateral view; e. Same, frontal view; f. Same, posterior view. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.8391967" httpUri="https://zenodo.org/record/8391967/files/figure.png" pageId="16" pageNumber="417">27b</figureCitation>
|
||
).............................................................................................. 2
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="CC31B831FFB957556C8EF8CEFA749D0C" blockId="16.[151,1436,1715,2030]" pageId="16" pageNumber="417">
|
||
<keyLead id="777F1B04FFB957556C8EF8CEFA749D0C" pageId="16" pageNumber="417">
|
||
1’. Ocularium with two high erect spines concolor with background (
|
||
<figureCitation id="54B5A4B4FFB957556F63F8CEFC3E9D4A" box="[890,982,1832,1854]" captionStart="FIGURE 19" captionStartId="21.[151,250,1943,1967]" captionTargetBox="[181,1406,181,1919]" captionTargetId="figure-17@21.[181,1406,181,1919]" captionTargetPageId="21" captionText="FIGURE 19. Titanoleptes cupidensis (Soares & Soares, 1946), (MNRJ 17869), male (A–F), and female (G, H), from Sooretama: a. Male, habitus, dorsal view; b. Same, detail of dorsal scutum; c. Same, ventral view; d. Same, dextrolateral view; e. Same, posterior view; f. Same, frontal view; g. Female, habitus, dextrolateral view; h. Same, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8391948" httpUri="https://zenodo.org/record/8391948/files/figure.png" pageId="16" pageNumber="417">Figs. 19f</figureCitation>
|
||
,
|
||
<figureCitation id="54B5A4B4FFB957556FFDF8CEFBE19D49" box="[996,1033,1832,1853]" captionStart="FIGURE 23" captionStartId="25.[151,250,1919,1943]" captionTargetBox="[151,1435,417,1895]" captionTargetId="figure-123@25.[151,1435,417,1895]" captionTargetPageId="25" captionText="FIGURE 23. Titanoleptes eros sp. nov., male/female paratypes (MNRJ 7077) from ES, Santa Teresa: a. Male, habitus, dorsal view; b. Same, ventral view; c. Same, frontal view; d. Same, posterior view; e. Same, dextrolateral view; f. Female, habitus, dorsal view; g. Female, habitus, dextrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8391957" httpUri="https://zenodo.org/record/8391957/files/figure.png" pageId="16" pageNumber="417">23c</figureCitation>
|
||
); femur IV of male substraight, without prominent retrolateral spines (
|
||
<figureCitation id="54B5A4B4FFB957556E1DF8A0FD909D28" box="[516,632,1862,1884]" captionStart="FIGURE 21" captionStartId="23.[151,250,1838,1862]" captionTargetBox="[151,1435,417,1815]" captionTargetId="figure-118@23.[151,1435,417,1815]" captionTargetPageId="23" captionText="FIGURE 21. Titanoleptes cupidensis (Soares & Soares, 1946), male (MNRJ 17869), from Sooretama, left Fe IV: a. Dorsal view; b. Retrolateral (external) view; c. Ventral view; d. Prolateral (internal) view. Anterior direction upwards in a to c. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8391953" httpUri="https://zenodo.org/record/8391953/files/figure.png" pageId="16" pageNumber="417">Figs. 21a–d</figureCitation>
|
||
,
|
||
<figureCitation id="54B5A4B4FFB957556E9BF8A0FD299D28" box="[642,705,1862,1884]" captionStart="FIGURE 24" captionStartId="26.[151,250,1463,1487]" captionTargetBox="[151,1436,181,1439]" captionTargetId="figure-20@26.[151,1436,181,1439]" captionTargetPageId="26" captionText="FIGURE 24. Titanoleptes eros sp. nov., male paratype (MNRJ 7077) from ES, Santa Teresa, free tergites and proximal left leg IV: a. Dorsal view; b. Prolateral view; c. Ventral view; d. Retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8391959" httpUri="https://zenodo.org/record/8391959/files/figure.png" pageId="16" pageNumber="417">24a–d</figureCitation>
|
||
); free tergite III armed with a median apophysis in both sexes (
|
||
<figureCitation id="54B5A4B4FFB95755693AF8A0FA709D28" box="[1315,1432,1862,1884]" captionStart="FIGURE 19" captionStartId="21.[151,250,1943,1967]" captionTargetBox="[181,1406,181,1919]" captionTargetId="figure-17@21.[181,1406,181,1919]" captionTargetPageId="21" captionText="FIGURE 19. Titanoleptes cupidensis (Soares & Soares, 1946), (MNRJ 17869), male (A–F), and female (G, H), from Sooretama: a. Male, habitus, dorsal view; b. Same, detail of dorsal scutum; c. Same, ventral view; d. Same, dextrolateral view; e. Same, posterior view; f. Same, frontal view; g. Female, habitus, dextrolateral view; h. Same, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8391948" httpUri="https://zenodo.org/record/8391948/files/figure.png" pageId="16" pageNumber="417">Figs. 19b, h</figureCitation>
|
||
,
|
||
<figureCitation id="54B5A4B4FFB957556CFBF885FEC89D0D" box="[226,288,1891,1913]" captionStart="FIGURE 23" captionStartId="25.[151,250,1919,1943]" captionTargetBox="[151,1435,417,1895]" captionTargetId="figure-123@25.[151,1435,417,1895]" captionTargetPageId="25" captionText="FIGURE 23. Titanoleptes eros sp. nov., male/female paratypes (MNRJ 7077) from ES, Santa Teresa: a. Male, habitus, dorsal view; b. Same, ventral view; c. Same, frontal view; d. Same, posterior view; e. Same, dextrolateral view; f. Female, habitus, dorsal view; g. Female, habitus, dextrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8391957" httpUri="https://zenodo.org/record/8391957/files/figure.png" pageId="16" pageNumber="417">23 a, f</figureCitation>
|
||
).............................................................................................. 3
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="777AA094FFB957546C8EF866FA749AD9" lastPageId="17" lastPageNumber="418" pageId="16" pageNumber="417">
|
||
<paragraph id="CC31B831FFB957556C8EF866FA749DA4" blockId="16.[151,1436,1715,2030]" pageId="16" pageNumber="417">
|
||
<keyLead id="777F1B04FFB957556C8EF866FA749DA4" pageId="16" pageNumber="417">
|
||
2. Body color mahogany with black reticle (
|
||
<figureCitation id="54B5A4B4FFB957556E72F866FD359DE1" box="[619,733,1920,1942]" captionStart="FIGURE 16" captionStartId="17.[151,250,1909,1933]" captionTargetBox="[151,1435,625,1885]" captionTargetId="figure-236@17.[151,1435,625,1885]" captionTargetPageId="17" captionText="FIGURE 16. Titanoleptes calcar (Roewer, 1913), (MNRJ 17818) from Sooretama: a. Male, habitus, dorsal view; b. Same, detail of dorsal scutum; c. Male, coxae I–IV, stigmatic area and free sternites, ventral view; d. Male, body, sinistrolateral view; e. Same, frontal view; f. Same, posterior view; g. Female, dorsal scutum, dorsal view; h. Same, sinistrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8391942" httpUri="https://zenodo.org/record/8391942/files/figure.png" pageId="16" pageNumber="417">Figs. 16b, g</figureCitation>
|
||
); tubercles of scutal area III strong, acuminate and contrasting dark brown (
|
||
<figureCitation id="54B5A4B4FFB957556CF0F878FEB59DC7" box="[233,349,1950,1972]" captionStart="FIGURE 16" captionStartId="17.[151,250,1909,1933]" captionTargetBox="[151,1435,625,1885]" captionTargetId="figure-236@17.[151,1435,625,1885]" captionTargetPageId="17" captionText="FIGURE 16. Titanoleptes calcar (Roewer, 1913), (MNRJ 17818) from Sooretama: a. Male, habitus, dorsal view; b. Same, detail of dorsal scutum; c. Male, coxae I–IV, stigmatic area and free sternites, ventral view; d. Male, body, sinistrolateral view; e. Same, frontal view; f. Same, posterior view; g. Female, dorsal scutum, dorsal view; h. Same, sinistrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8391942" httpUri="https://zenodo.org/record/8391942/files/figure.png" pageId="16" pageNumber="417">Figs. 16b, g</figureCitation>
|
||
); RD6 of male Fe IV straight (
|
||
<figureCitation id="54B5A4B4FFB957556E9DF878FD3E9DC0" box="[644,726,1950,1972]" captionStart="FIGURE 16" captionStartId="17.[151,250,1909,1933]" captionTargetBox="[151,1435,625,1885]" captionTargetId="figure-236@17.[151,1435,625,1885]" captionTargetPageId="17" captionText="FIGURE 16. Titanoleptes calcar (Roewer, 1913), (MNRJ 17818) from Sooretama: a. Male, habitus, dorsal view; b. Same, detail of dorsal scutum; c. Male, coxae I–IV, stigmatic area and free sternites, ventral view; d. Male, body, sinistrolateral view; e. Same, frontal view; f. Same, posterior view; g. Female, dorsal scutum, dorsal view; h. Same, sinistrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8391942" httpUri="https://zenodo.org/record/8391942/files/figure.png" pageId="16" pageNumber="417">Fig. 16a</figureCitation>
|
||
); FT II-III of female with yellow cones (
|
||
<figureCitation id="54B5A4B4FFB957556845F878FB229DC0" box="[1116,1226,1950,1972]" captionStart="FIGURE 16" captionStartId="17.[151,250,1909,1933]" captionTargetBox="[151,1435,625,1885]" captionTargetId="figure-236@17.[151,1435,625,1885]" captionTargetPageId="17" captionText="FIGURE 16. Titanoleptes calcar (Roewer, 1913), (MNRJ 17818) from Sooretama: a. Male, habitus, dorsal view; b. Same, detail of dorsal scutum; c. Male, coxae I–IV, stigmatic area and free sternites, ventral view; d. Male, body, sinistrolateral view; e. Same, frontal view; f. Same, posterior view; g. Female, dorsal scutum, dorsal view; h. Same, sinistrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8391942" httpUri="https://zenodo.org/record/8391942/files/figure.png" pageId="16" pageNumber="417">
|
||
Figs
|
||
<quantity id="0B7615D4FFB957556894F878FB5B9DC0" box="[1165,1203,1950,1972]" metricMagnitude="-2" metricUnit="kg" metricValue="1.6" pageId="16" pageNumber="417" unit="g" value="16.0">16g</quantity>
|
||
, h
|
||
</figureCitation>
|
||
)................................................................................................
|
||
<taxonomicName id="0B8EC3B2FFB9575568B5F85AFA749DA4" ID-CoL="BTLX4" authority="(Roewer, 1913)" baseAuthorityName="Roewer" baseAuthorityYear="1913" box="[1196,1436,1979,2001]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="calcar">
|
||
<emphasis id="FEFA6423FFB9575568B5F85AFAE89DA4" box="[1196,1280,1979,2001]" italics="true" pageId="16" pageNumber="417">T. calcar</emphasis>
|
||
(
|
||
<bibRefCitation id="A81FC5C0FFB957556914F85DFA7D9DA4" author="Roewer, C. - F." box="[1293,1429,1979,2000]" pageId="16" pageNumber="417" pagination="1 - 256" refId="ref16917" refString="Roewer, C. - F. (1913) Die Familie der Gonyleptiden der Opiliones-Laniatores. Archiv f ¸ r Naturgeschichte, 79 (4), 1 - 256, pl. 1 a." type="journal article" year="1913">Roewer, 1913</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="CC31B831FFB957546C8EF83EFA749AD9" blockId="16.[151,1436,1715,2030]" lastBlockId="17.[151,1436,151,261]" lastPageId="17" lastPageNumber="418" pageId="16" pageNumber="417">
|
||
<keyLead id="777F1B04FFB957546C8EF83EFA749AD9" lastPageId="17" lastPageNumber="418" pageId="16" pageNumber="417">
|
||
2’. Body color dark brown, almost black; tubercles of scutal area III small, rounded and contrasting pale yellow (
|
||
<figureCitation id="54B5A4B4FFB957556911F83EFAB59D9A" box="[1288,1373,2008,2030]" captionStart="FIGURE 27" captionStartId="29.[151,250,1378,1402]" captionTargetBox="[151,1435,181,1354]" captionTargetId="figure-17@29.[151,1435,181,1354]" captionTargetPageId="29" captionText="FIGURE 27. Titanoleptes hyperion sp. nov., male paratype (MNRJ 18669) ES, Santa Teresa: a. Habitus, panoramic, dorsal view; b. Habitus, dorsal view; c. Same, ventral view; d. Same, sinistrolateral view; e. Same, frontal view; f. Same, posterior view. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.8391967" httpUri="https://zenodo.org/record/8391967/files/figure.png" pageId="16" pageNumber="417">Fig. 27b</figureCitation>
|
||
); RD6 of male Fe IV strongly curved (
|
||
<figureCitation id="54B5A4B4FFB857546E09FF71FD8C9AD9" box="[528,612,151,173]" captionStart="FIGURE 27" captionStartId="29.[151,250,1378,1402]" captionTargetBox="[151,1435,181,1354]" captionTargetId="figure-17@29.[151,1435,181,1354]" captionTargetPageId="29" captionText="FIGURE 27. Titanoleptes hyperion sp. nov., male paratype (MNRJ 18669) ES, Santa Teresa: a. Habitus, panoramic, dorsal view; b. Habitus, dorsal view; c. Same, ventral view; d. Same, sinistrolateral view; e. Same, frontal view; f. Same, posterior view. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.8391967" httpUri="https://zenodo.org/record/8391967/files/figure.png" pageId="17" pageNumber="418">Fig. 27a</figureCitation>
|
||
); FT II-III of female entirely unarmed (
|
||
<figureCitation id="54B5A4B4FFB857546FC7FF71FBF79AD9" box="[990,1055,151,173]" captionStart="FIGURE 1" captionStartId="2.[151,250,1831,1855]" captionTargetBox="[151,1436,181,1808]" captionTargetId="figure-20@2.[151,1436,181,1808]" captionTargetPageId="2" captionText="FIGURE 1. Examples in vivo of some of the species treated in this paper: a–c. Cajango camacanensis sp. nov., a. Alpha male, from Una, Bahia State, Brazil; b. Beta male from São José da Vitoria, Bahia State, Brazil; c. Female from Una, Bahia State, Brazil; d. Titanoleptes calcar (Roewer, 1913), male from Prado, Bahia State, Brazil; e–f. Titanoleptes hyperion sp. nov. from Braço do Sul, Espirito Santo State, Brazil, e. Male, f. Female. Photos by A. Anker & P. Martins (a–d) and A.B. Kury (e–f). All used with permission." figureDoi="http://doi.org/10.5281/zenodo.8391906" httpUri="https://zenodo.org/record/8391906/files/figure.png" pageId="17" pageNumber="418">Fig. 1f</figureCitation>
|
||
)...............
|
||
<taxonomicName id="0B8EC3B2FFB8575468C4FF7EFAA19AD8" ID-CoL="BTLX7" authority="Kury & Bernabé, 2023" authorityName="Kury & Bernabé" authorityYear="2023" box="[1245,1353,151,173]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="17" pageNumber="418" phylum="Arthropoda" rank="species" species="hyperion" status="sp. nov.">
|
||
<emphasis id="FEFA6423FFB8575468C4FF7EFAA19AD8" box="[1245,1353,151,173]" italics="true" pageId="17" pageNumber="418">T. hyperion</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="FEFA6423FFB857546949FF7EFA749AD9" bold="true" box="[1360,1436,152,173]" pageId="17" pageNumber="418">
|
||
<taxonomicNameLabel id="E5C9D958FFB857546949FF7EFA749AD9" box="[1360,1436,152,173]" pageId="17" pageNumber="418" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="777AA094FFB857546C8EFF53FA749B71" pageId="17" pageNumber="418">
|
||
<paragraph id="CC31B831FFB857546C8EFF53FA749A9C" blockId="17.[151,1436,151,261]" pageId="17" pageNumber="418">
|
||
<keyLead id="777F1B04FFB857546C8EFF53FA749A9C" pageId="17" pageNumber="418">
|
||
3. Cx IV of male with strong inner apical apophysis (
|
||
<figureCitation id="54B5A4B4FFB857546ED1FF53FCD29ABE" box="[712,826,181,203]" captionStart="FIGURE 19" captionStartId="21.[151,250,1943,1967]" captionTargetBox="[181,1406,181,1919]" captionTargetId="figure-17@21.[181,1406,181,1919]" captionTargetPageId="21" captionText="FIGURE 19. Titanoleptes cupidensis (Soares & Soares, 1946), (MNRJ 17869), male (A–F), and female (G, H), from Sooretama: a. Male, habitus, dorsal view; b. Same, detail of dorsal scutum; c. Same, ventral view; d. Same, dextrolateral view; e. Same, posterior view; f. Same, frontal view; g. Female, habitus, dextrolateral view; h. Same, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8391948" httpUri="https://zenodo.org/record/8391948/files/figure.png" pageId="17" pageNumber="418">Figs. 19b, c</figureCitation>
|
||
,
|
||
<figureCitation id="54B5A4B4FFB857546F5CFF53FC829ABE" box="[837,874,181,202]" captionStart="FIGURE 20" captionStartId="22.[151,250,1276,1300]" captionTargetBox="[151,1435,181,1252]" captionTargetId="figure-20@22.[151,1435,181,1252]" captionTargetPageId="22" captionText="FIGURE 20. Titanoleptes cupidensis (Soares & Soares, 1946), male (MNRJ 17869), from Sooretama:a. Left coxa andtrochanter IV, dorsal view; b. Left patella IV, prolateral view; c. Left coxa to proximal femur IV, prolateral view; d. Left leg IV, ventral view. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.8391950" httpUri="https://zenodo.org/record/8391950/files/figure.png" pageId="17" pageNumber="418">20a</figureCitation>
|
||
); Tr IV with strong apical inner apophysis (
|
||
<figureCitation id="54B5A4B4FFB857546914FF53FAB69ABE" box="[1293,1374,181,203]" captionStart="FIGURE 20" captionStartId="22.[151,250,1276,1300]" captionTargetBox="[151,1435,181,1252]" captionTargetId="figure-20@22.[151,1435,181,1252]" captionTargetPageId="22" captionText="FIGURE 20. Titanoleptes cupidensis (Soares & Soares, 1946), male (MNRJ 17869), from Sooretama:a. Left coxa andtrochanter IV, dorsal view; b. Left patella IV, prolateral view; c. Left coxa to proximal femur IV, prolateral view; d. Left leg IV, ventral view. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.8391950" httpUri="https://zenodo.org/record/8391950/files/figure.png" pageId="17" pageNumber="418">Fig. 20a</figureCitation>
|
||
)..........................................................................
|
||
<taxonomicName id="0B8EC3B2FFB857546836FF35FA749A9C" ID-CoL="BTLX5" authority="(Soares & Soares, 1946)" authorityName="Kury & Bernabé" authorityYear="2023" baseAuthorityName="Soares & Soares" baseAuthorityYear="1946" box="[1071,1436,210,232]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="17" pageNumber="418" phylum="Arthropoda" rank="species" species="cupidensis">
|
||
<emphasis id="FEFA6423FFB857546836FF35FB439A93" box="[1071,1195,210,232]" italics="true" pageId="17" pageNumber="418">T. cupidensis</emphasis>
|
||
(
|
||
<bibRefCitation id="A81FC5C0FFB8575468A1FF34FA7D9A9C" author="Soares, B. A. M. & Soares, H. E. M." box="[1208,1429,210,232]" pageId="17" pageNumber="418" pagination="195 - 212" refId="ref16998" refString="Soares, B. A. M. & Soares, H. E. M. (1946) Novos opilioes do Espirito Santo e um novo opiliao do estado do Para. Papeis Avulsos do Departamento de Zoologia, Secretaria da Agricultura, Sao Paulo, 7, 195 - 212." type="journal article" year="1946">Soares & Soares, 1946</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="CC31B831FFB857546C8EFF16FA749B71" blockId="17.[151,1436,151,261]" box="[151,1436,239,261]" pageId="17" pageNumber="418">
|
||
<keyLead id="777F1B04FFB857546C8EFF16FA749B71" box="[151,1436,239,261]" pageId="17" pageNumber="418">
|
||
3’. Cx IV of male unarmed internally (
|
||
<figureCitation id="54B5A4B4FFB857546E2DFF09FD4F9B71" box="[564,679,239,261]" captionStart="FIGURE 23" captionStartId="25.[151,250,1919,1943]" captionTargetBox="[151,1435,417,1895]" captionTargetId="figure-123@25.[151,1435,417,1895]" captionTargetPageId="25" captionText="FIGURE 23. Titanoleptes eros sp. nov., male/female paratypes (MNRJ 7077) from ES, Santa Teresa: a. Male, habitus, dorsal view; b. Same, ventral view; c. Same, frontal view; d. Same, posterior view; e. Same, dextrolateral view; f. Female, habitus, dorsal view; g. Female, habitus, dextrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8391957" httpUri="https://zenodo.org/record/8391957/files/figure.png" pageId="17" pageNumber="418">Figs. 23a, b</figureCitation>
|
||
); Tr IV with small apical inner apophysis.................
|
||
<taxonomicName id="0B8EC3B2FFB857546913FF16FAA19B71" ID-CoL="BTLX6" authority="Kury & Bernabé, 2023" authorityName="Kury & Bernabé" authorityYear="2023" box="[1290,1353,240,261]" class="Arachnida" family="Gonyleptidae" genus="Titanoleptes" kingdom="Animalia" order="Opiliones" pageId="17" pageNumber="418" phylum="Arthropoda" rank="species" species="eros" status="sp. nov.">
|
||
<emphasis id="FEFA6423FFB857546913FF16FAA19B71" box="[1290,1353,240,261]" italics="true" pageId="17" pageNumber="418">T. eros</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="FEFA6423FFB857546949FF16FA749B71" bold="true" box="[1360,1436,240,261]" pageId="17" pageNumber="418">
|
||
<taxonomicNameLabel id="E5C9D958FFB857546949FF16FA749B71" box="[1360,1436,240,261]" pageId="17" pageNumber="418" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |