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<document id="614AA25DA060602C3AD89109F46825BA" ID-DOI="10.1206/0003-0090(2004)286&lt;0001:MPTASO&gt;2.0.CO;2" ID-ISSN="0003-0090" ID-Zenodo-Dep="5355565" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630215625017" checkinUser="felipe" docAuthor="CLARKE, JULIA A." docDate="2004" docId="4302B56EFFCAFFB3FFCF73897DBAB654" docLanguage="en" docName="B286.pdf" docOrigin="Bulletin of the American Museum of Natural History 2004 (286)" docSource="http://www.bioone.org/doi/abs/10.1206/0003-0090%282004%29286%3C0001%3AMPTASO%3E2.0.CO%3B2" docStyle="DocumentStyle:2B34016C1EBFE58B07368F9517618743.3:BulAmeMusNatHis.2000-2010.journal_article.0cover.type1" docStyleId="2B34016C1EBFE58B07368F9517618743" docStyleName="BulAmeMusNatHis.2000-2010.journal_article.0cover.type1" docStyleVersion="3" docTitle="Ichthyornis victor : YPM 1775" docType="treatment" docVersion="4" lastPageNumber="42" masterDocId="BF3BCD16FFE2FF9AFFB377697C7DB221" masterDocTitle="Morphology, Phylogenetic Taxonomy, And Systematics Of Ichthyornis And Apatornis (Avialae: Ornithurae)" masterLastPageNumber="179" masterPageNumber="1" pageNumber="41" updateTime="1699189515630" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="CA7BE565A9F655DD18EAB361C655A933">Morphology, Phylogenetic Taxonomy, And Systematics Of Ichthyornis And Apatornis (Avialae: Ornithurae)</mods:title>
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<subSubSection id="83B157F3FFCAFFB2FFCF73897D33B6D6" box="[124,334,1248,1271]" pageId="40" pageNumber="41" type="nomenclature">
<paragraph id="CB140478FFCAFFB2FFCF73897D33B6D6" blockId="40.[124,334,1248,1271]" box="[124,334,1248,1271]" pageId="40" pageNumber="41">
<heading id="905CB314FFCAFFB2FFCF73897D33B6D6" box="[124,334,1248,1271]" fontSize="9" level="2" pageId="40" pageNumber="41" reason="6">
<taxonomicName id="0CAB7FFBFFCAFFB2FFCF73897D33B6D6" authorityName=": YPM" authorityYear="1775" box="[124,334,1248,1271]" class="Aves" family="Ichthyornithidae" genus="Ichthyornis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ichthyornithiformes" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="victor">
<emphasis id="F9DFD86AFFCAFFB2FFCF73897D33B6D6" box="[124,334,1248,1271]" italics="true" pageId="40" pageNumber="41">Ichthyornis victor</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="83B157F3FFCAFFB2FF2A727D7CB5B782" pageId="40" pageNumber="41" type="reference_group">
<paragraph id="CB140478FFCAFFB2FF2A727D7CB5B782" blockId="40.[124,653,1300,1741]" pageId="40" pageNumber="41">
<taxonomicName id="0CAB7FFBFFCAFFB2FF2A727D7D0CB70A" authorityName=": YPM" authorityYear="1775" box="[153,369,1300,1323]" class="Aves" family="Ichthyornithidae" genus="Ichthyornis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ichthyornithiformes" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="victor">
<emphasis id="F9DFD86AFFCAFFB2FF2A727D7D0CB70A" box="[153,369,1300,1323]" italics="true" pageId="40" pageNumber="41">Ichthyornis victor</emphasis>
</taxonomicName>
was named by
<bibRefCitation id="AF3A7989FFCAFFB2FDF1727D7CB4B768" author="Marsh, O. C." pageId="40" pageNumber="41" pagination="509 - 511" refId="ref103742" refString="Marsh, O. C. 1876. Notice of new Odontornithes. American Journal of Science, 3 rd. ser., 11: 509 - 511." type="journal article" year="1876">Marsh (1876)</bibRefCitation>
and differentiated from the
<typeStatus id="1410BADAFFCAFFB2FD94725B7EF0B768" box="[551,653,1330,1353]" pageId="40" pageNumber="41" type="holotype">holotype</typeStatus>
of
<taxonomicName id="0CAB7FFBFFCAFFB2FF1372397E60B746" authority="(YPM 1450)" baseAuthorityName="YPM" baseAuthorityYear="1450" box="[160,541,1360,1383]" class="Aves" family="Ichthyornithidae" genus="Ichthyornis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ichthyornithiformes" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="dispar">
<emphasis id="F9DFD86AFFCAFFB2FF1372397D07B746" box="[160,378,1360,1383]" italics="true" pageId="40" pageNumber="41">Ichthyornis dispar</emphasis>
(YPM 1450)
</taxonomicName>
as being one­third larger than that species (
<bibRefCitation id="AF3A7989FFCAFFB2FD8972077CC6B782" author="Marsh, O. C." pageId="40" pageNumber="41" pagination="509 - 511" refId="ref103742" refString="Marsh, O. C. 1876. Notice of new Odontornithes. American Journal of Science, 3 rd. ser., 11: 509 - 511." type="journal article" year="1876">Marsh, 1876</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="83B157F3FFCAFFB2FF2A72C37F6CB7C2" pageId="40" pageNumber="41" type="materials_examined">
<paragraph id="CB140478FFCAFFB2FF2A72C37F61B6D9" blockId="40.[124,653,1300,1741]" lastBlockId="40.[700,1229,1220,1741]" pageId="40" pageNumber="41">
<typeStatus id="1410BADAFFCAFFB2FF2A72C37D72B7E1" box="[153,271,1450,1473]" pageId="40" pageNumber="41" type="holotype">HOLOTYPE</typeStatus>
SPECIMEN:
<collectionCode id="ADBA9CBDFFCAFFB2FE2F72C37DA6B7E0" box="[412,475,1450,1473]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34880" lsid="urn:lsid:biocol.org:col:34880" name="Peabody Museum of Natural History" pageId="40" pageNumber="41" type="Museum">YPM</collectionCode>
1452 includes a proximal right coracoid, proximal right scapula, and three fragments of a poorly preserved left humerus (fig. 11). As discussed in the Historical Background, some or all of the material incorporated into the
<collectionCode id="ADBA9CBDFFCAFFB2FDA671567E29B477" box="[533,596,1599,1622]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34880" lsid="urn:lsid:biocol.org:col:34880" name="Peabody Museum of Natural History" pageId="40" pageNumber="41" type="Museum">YPM</collectionCode>
<taxonomicName id="0CAB7FFBFFCAFFB2FDED71567D53B455" authorityName=": YPM" authorityYear="1775" class="Aves" family="Ichthyornithidae" genus="Ichthyornis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ichthyornithiformes" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="victor">
<emphasis id="F9DFD86AFFCAFFB2FDED71567D53B455" italics="true" pageId="40" pageNumber="41">Ichthyornis victor</emphasis>
</taxonomicName>
panel mount frequently appears to have been mistaken for
<typeStatus id="1410BADAFFCAFFB2FDB671127E16B4B3" box="[517,619,1659,1682]" pageId="40" pageNumber="41" type="holotype">holotype</typeStatus>
of
<taxonomicName id="0CAB7FFBFFCAFFB2FFCF71F07D2DB491" authorityName=": YPM" authorityYear="1775" box="[124,336,1689,1712]" class="Aves" family="Ichthyornithidae" genus="Ichthyornis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ichthyornithiformes" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="victor">
<emphasis id="F9DFD86AFFCAFFB2FFCF71F07D2DB491" box="[124,336,1689,1712]" italics="true" pageId="40" pageNumber="41">Ichthyornis victor</emphasis>
</taxonomicName>
. The
<collectionCode id="ADBA9CBDFFCAFFB2FE2F71F17DA6B48E" box="[412,475,1688,1711]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34880" lsid="urn:lsid:biocol.org:col:34880" name="Peabody Museum of Natural History" pageId="40" pageNumber="41" type="Museum">YPM</collectionCode>
Catalogue entry for
<collectionCode id="ADBA9CBDFFCAFFB2FF6171DF7D6CB4EC" box="[210,273,1718,1741]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34880" lsid="urn:lsid:biocol.org:col:34880" name="Peabody Museum of Natural History" pageId="40" pageNumber="41" type="Museum">YPM</collectionCode>
1452 included the phrase Ma­ jor portions of skeleton for most of the 20th century.
</paragraph>
<paragraph id="CB140478FFCAFFB2FD6A73977F6CB7C2" blockId="40.[700,1229,1220,1741]" pageId="40" pageNumber="41">
LOCALITY
<collectionCode id="ADBA9CBDFFCAFFB2FCEE726A7FF6B734" box="[861,907,1283,1301]" pageId="40" pageNumber="41">AND</collectionCode>
HORIZON:
<bibRefCitation id="AF3A7989FFCAFFB2FB9173977E8FB712" author="Marsh, O. C." pageId="40" pageNumber="41" refId="ref103841" refString="Marsh, O. C. 1880. Odontornithes: a monograph on the extinct toothed birds of North America. United States Geological Exploration of the 40 th Parallel. Washington, DC: U. S. Government Printing Office, 201 pp." type="book" year="1880">Marsh (1880: 199)</bibRefCitation>
reported that
<collectionCode id="ADBA9CBDFFCAFFB2FC2B72757FAAB712" box="[920,983,1308,1331]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34880" lsid="urn:lsid:biocol.org:col:34880" name="Peabody Museum of Natural History" pageId="40" pageNumber="41" type="Museum">YPM</collectionCode>
1452 was found in
<date id="BF1522B8FFCAFFB2FD0F72507F47B771" box="[700,826,1337,1360]" pageId="40" pageNumber="41" value="1876-05">May 1876</date>
, by Mr. H. A. Brous, in Wallace County,
<collectingRegion id="096FCA9AFFCAFFB2FC99723F7FFAB74C" box="[810,903,1366,1389]" country="United States of America" name="Kansas" pageId="40" pageNumber="41">Kansas</collectingRegion>
. No locality information was given in the original description of the species (
<bibRefCitation id="AF3A7989FFCAFFB2FC9572F87FB8B789" author="Marsh, O. C." box="[806,965,1425,1448]" pageId="40" pageNumber="41" pagination="509 - 511" refId="ref103742" refString="Marsh, O. C. 1876. Notice of new Odontornithes. American Journal of Science, 3 rd. ser., 11: 509 - 511." type="journal article" year="1876">Marsh, 1876</bibRefCitation>
), while the listing in
<bibRefCitation id="AF3A7989FFCAFFB2FD0F72C77F15B7E4" author="Marsh, O. C." box="[700,872,1454,1477]" pageId="40" pageNumber="41" refId="ref103841" refString="Marsh, O. C. 1880. Odontornithes: a monograph on the extinct toothed birds of North America. United States Geological Exploration of the 40 th Parallel. Washington, DC: U. S. Government Printing Office, 201 pp." type="book" year="1880">Marsh (1880)</bibRefCitation>
was repeated in
<bibRefCitation id="AF3A7989FFCAFFB2FBEE72C77F70B7C2" author="Brodkorb, P." pageId="40" pageNumber="41" pagination="99 - 220" refId="ref98286" refString="Brodkorb, P. 1967. Catalogue of fossil birds: part 3 (Ralliformes, Ichthyornithiformes, Charadriiformes). Bulletin of the Florida State Museum (Biological Sciences) 11: 99 - 220." type="journal article" year="1967">Brodkorb (1967)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="83B157F3FFCAFFB3FD6A72807DBAB654" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41" type="discussion">
<paragraph id="CB140478FFCAFFB3FD6A72807D77B63C" blockId="40.[700,1229,1220,1741]" lastBlockId="41.[123,652,854,1141]" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">
DISCUSSION: Several morphologies described for the
<typeStatus id="1410BADAFFCAFFB2FCCF716F7F9FB43C" box="[892,994,1542,1565]" pageId="40" pageNumber="41" type="holotype">holotype</typeStatus>
(
<bibRefCitation id="AF3A7989FFCAFFB2FC44716F78EEB43C" author="Marsh, O. C." box="[1015,1171,1542,1565]" pageId="40" pageNumber="41" pagination="509 - 511" refId="ref103742" refString="Marsh, O. C. 1876. Notice of new Odontornithes. American Journal of Science, 3 rd. ser., 11: 509 - 511." type="journal article" year="1876">Marsh, 1876</bibRefCitation>
) but not used to differentiate it are also present in the
<typeStatus id="1410BADAFFCAFFB2FD5B71287F33B479" box="[744,846,1601,1624]" pageId="40" pageNumber="41" type="holotype">holotype</typeStatus>
of
<taxonomicName id="0CAB7FFBFFCAFFB2FCC571287830B479" baseAuthorityName="YPM" baseAuthorityYear="1450" box="[886,1101,1601,1624]" class="Aves" family="Ichthyornithidae" genus="Ichthyornis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ichthyornithiformes" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="dispar">
<emphasis id="F9DFD86AFFCAFFB2FCC571287830B479" box="[886,1101,1601,1624]" italics="true" pageId="40" pageNumber="41">Ichthyornis dispar</emphasis>
</taxonomicName>
. These are now known to be ancestral for more inclusive avialan clades (e.g., expanded deltopectoral crest and dorsal humerus condyle larger than the ventral; see appendix 1: characters 113, 125). Only humeri can be directly compared between the
<typeStatus id="1410BADAFFCBFFB3FED6741A7DABB1AB" box="[357,470,883,906]" pageId="41" pageNumber="42" type="holotype">holotypes</typeStatus>
of
<taxonomicName id="0CAB7FFBFFCBFFB3FDB4741A7CB8B189" baseAuthorityName="YPM" baseAuthorityYear="1450" class="Aves" family="Ichthyornithidae" genus="Ichthyornis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ichthyornithiformes" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="dispar">
<emphasis id="F9DFD86AFFCBFFB3FDB4741A7CB8B189" italics="true" pageId="41" pageNumber="42">Ichthyornis dispar</emphasis>
</taxonomicName>
and
<taxonomicName id="0CAB7FFBFFCBFFB3FF4C74F87DADB189" authorityName=": YPM" authorityYear="1775" box="[255,464,913,936]" class="Aves" family="Ichthyornithidae" genus="Ichthyornis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ichthyornithiformes" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="victor">
<emphasis id="F9DFD86AFFCBFFB3FF4C74F87DADB189" box="[255,464,913,936]" italics="true" pageId="41" pageNumber="42">Ichthyornis victor</emphasis>
</taxonomicName>
and, while they are different in size, there are no appreciable differences in morphology. The morphology of YPM 1452 is discussed in the Anatomical Description.
</paragraph>
<caption id="9FD454F0FFCBFFB3FF2675897EDBB12E" ID-DOI="http://doi.org/10.5281/zenodo.5355593" ID-Zenodo-Dep="5355593" httpUri="https://zenodo.org/record/5355593/files/figure.png" pageId="41" pageNumber="42" startId="41.[149,187,736,757]" targetBox="[123,1227,188,715]" targetPageId="41">
<paragraph id="CB140478FFCBFFB3FF2675897EDBB12E" blockId="41.[123,1227,735,783]" pageId="41" pageNumber="42">
Fig. 12. The
<taxonomicName id="0CAB7FFBFFCBFFB3FE8475897E7EB0D4" baseAuthorityName="YPM" baseAuthorityYear="1740" box="[311,515,736,757]" class="Aves" family="Ichthyornithidae" genus="Ichthyornis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ichthyornithiformes" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="validus">
<emphasis id="F9DFD86AFFCBFFB3FE8475897E7EB0D4" box="[311,515,736,757]" italics="true" pageId="41" pageNumber="42">Ichthyornis validus</emphasis>
</taxonomicName>
holotype (YPM 1740), a right ulna:
<emphasis id="F9DFD86AFFCBFFB3FC2C75B67FB9B0D5" bold="true" box="[927,964,735,756]" pageId="41" pageNumber="42">(A)</emphasis>
the distal end in dorsal view and
<emphasis id="F9DFD86AFFCBFFB3FF5675937D75B12E" bold="true" box="[229,264,762,783]" pageId="41" pageNumber="42">(B)</emphasis>
the complete element in ventral view.
</paragraph>
</caption>
<paragraph id="CB140478FFCBFFB3FF2B734A7DBAB654" blockId="41.[123,652,854,1141]" pageId="41" pageNumber="42">
REFERRED SPECIMENS: The majority of YPM specimens were referred (
<bibRefCitation id="AF3A7989FFCBFFB3FE5E73297EFEB676" author="Marsh, O. C." box="[493,643,1088,1111]" pageId="41" pageNumber="42" refId="ref103841" refString="Marsh, O. C. 1880. Odontornithes: a monograph on the extinct toothed birds of North America. United States Geological Exploration of the 40 th Parallel. Washington, DC: U. S. Government Printing Office, 201 pp." type="book" year="1880">Marsh, 1880</bibRefCitation>
) to this species (see table 1).
</paragraph>
</subSubSection>
</treatment>
</document>