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<mods:title id="63483F53867980204F02764A08D4B6B5">Revision of the Australopapuan and West Pacific species of plain pumpkinbeetles, the Aulacophora indica species-complex (Coleoptera: Chrysomelidae: Galerucinae)</mods:title>
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<bibRefCitation id="AD6BE8B13F56E5497125C43CFDCA4786" author="Chevrolat, L. A. A." box="[317,515,1914,1940]" pageId="5" pageNumber="6" pagination="361 - 443" refId="ref39698" refString="Chevrolat, L. A. A. (1836) [new taxa]. In: Dejean, P. F. M. A. (Ed.), Catalogue des Coleopteres de la collection de M. le Comte Dejean. Livraison 5. Mequignon-Marvis, Paris, pp. 361 - 443." type="book chapter" year="1836">Chevrolat, 1836</bibRefCitation>
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<emphasis id="FB8E49523F56E549708FC4F9FED247C5" box="[151,283,1983,2007]" italics="true" pageId="5" pageNumber="6">Aulacophora</emphasis>
<bibRefCitation id="AD6BE8B13F56E549713BC4F9FE3647C5" author="Chevrolat, L. A. A." box="[291,511,1983,2007]" pageId="5" pageNumber="6" pagination="361 - 443" refId="ref39698" refString="Chevrolat, L. A. A. (1836) [new taxa]. In: Dejean, P. F. M. A. (Ed.), Catalogue des Coleopteres de la collection de M. le Comte Dejean. Livraison 5. Mequignon-Marvis, Paris, pp. 361 - 443." type="book chapter" year="1836">Chevrolat, 1836: 378</bibRefCitation>
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;
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<bibRefCitation id="AD6BE8B13F56E5497213C4F9FD0F47C5" author="Maulik, S." box="[523,710,1983,2007]" pageId="5" pageNumber="6" refId="ref41439" refString="Maulik, S. (1936) The Fauna of British India including Ceylon and Burma. Coleoptera. Chrysomelidae. (Galerucinae). Taylor &amp; Francis, London, xiv + 648 pp., 1 pl." type="book" year="1936">Maulik 1936: 167</bibRefCitation>
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;
<treatmentCitation id="485BB3513F56E54972C9C4F9FC4547C5" author="Wilcox, J. A." box="[721,908,1983,2007]" page="221" pageId="5" pageNumber="6" year="1972">
<bibRefCitation id="AD6BE8B13F56E54972C9C4F9FC4547C5" author="Wilcox, J. A." box="[721,908,1983,2007]" pageId="5" pageNumber="6" pagination="221 - 431" refId="ref43057" refString="Wilcox, J. A. (1972) Chrysomelidae: Galerucinae: Luperini: Aulacophorina, Diabroticina. Coleopterorum Catalogus Supplementa, 78 (2), 221 - 431." type="journal article" year="1972">Wilcox 1972: 221</bibRefCitation>
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;
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<bibRefCitation id="AD6BE8B13F56E549738FC4F9FB4547C5" author="Anand, R. K. &amp; Cox, M. L." box="[919,1164,1983,2007]" pageId="5" pageNumber="6" pagination="80 - 91" refId="ref38733" refString="Anand, R. K. &amp; Cox, M. L. (1986) Taxonomic revision of the genus Aulacophora Chevr. I. Species with yellow elytra. Entomologische Abhandlungen aus dem Museum fur Tierkunde in Dresden, 50 (4), 80 - 91." type="journal article" year="1986">Anand &amp; Cox 1986: 81</bibRefCitation>
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;
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;
<bibRefCitation id="AD6BE8B13F56E549711FC498FE1A47E4" author="Lee, C. - F. &amp; Beenen, R." box="[263,467,2014,2038]" pageId="5" pageNumber="6" pagination="151 - 190" refId="ref41061" refString="Lee, C. - F. &amp; Beenen, R. (2015) Revision of the genus Aulacophora from Taiwan (Coleoptera: Chrysomelidae: Galerucinae). Zootaxa, 3949 (2), 151 - 190. https: // doi. org / 10.11646 / zootaxa. 3949.2.1" type="journal article" year="2015">Lee &amp; Beenen 2015</bibRefCitation>
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=
<treatmentCitation id="485BB3513F55E54A70B4C3D1FDD240BD" author="Chevrolat, L. A. A." box="[172,539,150,175]" page="378" pageId="6" pageNumber="7" year="1836">
<taxonomicName id="0EFAEEC33F55E54A70B4C3D1FDD240BD" authority="Chevrolat, 1836: 378" authorityName="Chevrolat" authorityPageNumber="378" authorityYear="1836" box="[172,539,150,175]" class="Insecta" family="Chrysomelidae" genus="Raphidopalpa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="FB8E49523F55E54A70B4C3D1FEF540BD" box="[172,316,151,175]" italics="true" pageId="6" pageNumber="7">Raphidopalpa</emphasis>
<bibRefCitation id="AD6BE8B13F55E54A715AC3D1FDD240BD" author="Chevrolat, L. A. A." box="[322,539,150,175]" pageId="6" pageNumber="7" pagination="361 - 443" refId="ref39698" refString="Chevrolat, L. A. A. (1836) [new taxa]. In: Dejean, P. F. M. A. (Ed.), Catalogue des Coleopteres de la collection de M. le Comte Dejean. Livraison 5. Mequignon-Marvis, Paris, pp. 361 - 443." type="book chapter" year="1836">Chevrolat, 1836: 378</bibRefCitation>
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;
<bibRefCitation id="AD6BE8B13F55E54A723EC3D1FCF640BC" author="Gemminger, M. &amp; von Harold, E." box="[550,831,150,175]" pageId="6" pageNumber="7" pagination="3479 - 3822" refId="ref40176" refString="Gemminger, M. &amp; von Harold, E. (1876) Catalogus coleopterorum hucusque descriptorum synonymicus et systematicus. Tom XII. Chrysomelidae (Pars II), Languridae, Erotylidae, Endomychidae, Coccinellidae, Corylophidae, Platypsyllidae. Accedit index Generum Universalis. EH Gummi, Monachii, pp. 3479 - 3822." type="book chapter" year="1876">Gemminger &amp; Harold 1876</bibRefCitation>
(synonymy)
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=
<treatmentCitation id="485BB3513F55E54A70B4C3F3FDF240DF" author="Rosenhauer, W. G." box="[172,571,181,205]" page="327" pageId="6" pageNumber="7" year="1856">
<taxonomicName id="0EFAEEC33F55E54A70B4C3F3FDF240DF" authority="Rosenhauer, 1856: 327" authorityName="Rosenhauer" authorityPageNumber="327" authorityYear="1856" box="[172,571,181,205]" class="Insecta" family="Chrysomelidae" genus="Rhaphidopalpa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="FB8E49523F55E54A70B4C3F3FE8140DF" box="[172,328,181,205]" italics="true" pageId="6" pageNumber="7">Rhaphidopalpa</emphasis>
<bibRefCitation id="AD6BE8B13F55E54A7156C3F3FDF240DF" author="Rosenhauer, W. G." box="[334,571,181,205]" pageId="6" pageNumber="7" refId="ref42323" refString="Rosenhauer, W. G. (1856) Thiere Andalusiens nach dem Resultate einer Reise zusammengestellt nebst den Beschreibungen von 249 neuen oder bis jetzt unbeschriebenen Gattungen und Arten. T. Blaesing, Erlangen, viii + 429 pp, 3 pls. https: // doi. org / 10.5962 / bhl. title. 66016" type="book" year="1856">Rosenhauer, 1856: 327</bibRefCitation>
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; Hincks 1949: 621 (synonymy)
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<treatmentCitation id="485BB3513F55E54A70B4C392FDC740FE" author="Rosenhauer, W. G." box="[172,526,212,236]" page="327" pageId="6" pageNumber="7" year="1856">
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<emphasis id="FB8E49523F55E54A70B4C392FED240FE" box="[172,283,212,236]" italics="true" pageId="6" pageNumber="7">Acutipalpa</emphasis>
<bibRefCitation id="AD6BE8B13F55E54A7139C392FDC740FE" author="Rosenhauer, W. G." box="[289,526,212,236]" pageId="6" pageNumber="7" refId="ref42323" refString="Rosenhauer, W. G. (1856) Thiere Andalusiens nach dem Resultate einer Reise zusammengestellt nebst den Beschreibungen von 249 neuen oder bis jetzt unbeschriebenen Gattungen und Arten. T. Blaesing, Erlangen, viii + 429 pp, 3 pls. https: // doi. org / 10.5962 / bhl. title. 66016" type="book" year="1856">Rosenhauer, 1856: 327</bibRefCitation>
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; Joannis 1866: 99 (synonymy)
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=
<taxonomicName id="0EFAEEC33F55E54A70B4C3B5FE014119" ID-CoL="7NX3J" authority="Chapuis, 1876: 100" authorityName="Chapuis" authorityPageNumber="100" authorityYear="1876" box="[172,456,242,267]" class="Gastropoda" family="Iravadiidae" genus="Ceratia" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="7" phylum="Mollusca" rank="genus">
<emphasis id="FB8E49523F55E54A70B4C3B5FF304118" box="[172,249,243,266]" italics="true" pageId="6" pageNumber="7">Ceratia</emphasis>
Chapuis, 1876: 100
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;
<treatmentCitation id="485BB3513F55E54A71CBC3B5FD414118" author="Maulik, S." box="[467,648,242,267]" page="167" pageId="6" pageNumber="7" year="1936">
<bibRefCitation id="AD6BE8B13F55E54A71CBC3B5FD414118" author="Maulik, S." box="[467,648,242,267]" pageId="6" pageNumber="7" refId="ref41439" refString="Maulik, S. (1936) The Fauna of British India including Ceylon and Burma. Coleoptera. Chrysomelidae. (Galerucinae). Taylor &amp; Francis, London, xiv + 648 pp., 1 pl." type="book" year="1936">Maulik 1936: 167</bibRefCitation>
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(synonymy)
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=
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<emphasis id="FB8E49523F55E54A70B4C257FEDA413B" box="[172,275,273,297]" italics="true" pageId="6" pageNumber="7">Triaplatys</emphasis>
Fairmaire, 1879: 113
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;
<treatmentCitation id="485BB3513F55E54A71E2C257FD42413B" author="Weise, J." box="[506,651,273,297]" page="9" pageId="6" pageNumber="7" year="1924">
<bibRefCitation id="AD6BE8B13F55E54A71E2C257FD42413B" author="Weise, J." box="[506,651,273,297]" pageId="6" pageNumber="7" pagination="1 - 225" refId="ref43036" refString="Weise, J. (1924) Chrysomelidae: 13 Galerucinae. Coleopterorum Catalogus, 78, 1 - 225." type="journal article" year="1924">Weise 1924: 9</bibRefCitation>
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(synonymy)
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=
<treatmentCitation id="485BB3513F55E54A70B4C276FE10415A" author="Weise, J." box="[172,473,304,328]" page="396" pageId="6" pageNumber="7" year="1892">
<taxonomicName id="0EFAEEC33F55E54A70B4C276FE10415A" authority="Weise, 1892: 396" authorityName="Weise" authorityPageNumber="396" authorityYear="1892" box="[172,473,304,328]" genus="Orthaulaca" kingdom="Animalia" pageId="6" pageNumber="7" rank="genus">
<emphasis id="FB8E49523F55E54A70B4C276FEE8415A" box="[172,289,304,328]" italics="true" pageId="6" pageNumber="7">Orthaulaca</emphasis>
<bibRefCitation id="AD6BE8B13F55E54A713EC276FE10415A" author="Weise, J." box="[294,473,304,328]" pageId="6" pageNumber="7" pagination="385 - 402" refId="ref42923" refString="Weise, J. (1892) Chrysomeliden und Coccinelliden von der Insel Nias, nebst Bemerkungen uber andere, meistens sudoestasiatische Arten. Deutsche Entomologische Zeitschrift, 1892 (2), 385 - 402. https: // doi. org / 10.1002 / mmnd. 48018920128" type="journal article" year="1892">Weise, 1892: 396</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="485BB3513F55E54A71FCC276FD50415A" author="Maulik, S." box="[484,665,304,328]" page="167" pageId="6" pageNumber="7" year="1936">
<bibRefCitation id="AD6BE8B13F55E54A71FCC276FD50415A" author="Maulik, S." box="[484,665,304,328]" pageId="6" pageNumber="7" refId="ref41439" refString="Maulik, S. (1936) The Fauna of British India including Ceylon and Burma. Coleoptera. Chrysomelidae. (Galerucinae). Taylor &amp; Francis, London, xiv + 648 pp., 1 pl." type="book" year="1936">Maulik 1936: 167</bibRefCitation>
</treatmentCitation>
(synonymy)
</treatmentCitationGroup>
</paragraph>
<paragraph id="C94595403F55E54A708FC209FD314175" blockId="6.[151,959,150,451]" box="[151,760,334,359]" pageId="6" pageNumber="7">
<treatmentCitationGroup id="E9EAB26E3F55E54A708FC209FD314175" box="[151,760,334,359]" pageId="6" pageNumber="7">
=
<treatmentCitation id="485BB3513F55E54A70B4C209FE714174" author="Weise, J." box="[172,440,334,359]" page="396" pageId="6" pageNumber="7" year="1892">
<taxonomicName id="0EFAEEC33F55E54A70B4C209FE714174" authority="Weise, 1892: 396" authorityName="Weise" authorityPageNumber="396" authorityYear="1892" box="[172,440,334,359]" class="Insecta" family="Chrysomelidae" genus="Cerania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="FB8E49523F55E54A70B4C209FF364174" box="[172,255,335,358]" italics="true" pageId="6" pageNumber="7">Cerania</emphasis>
<bibRefCitation id="AD6BE8B13F55E54A711DC209FE714174" author="Weise, J." box="[261,440,334,359]" pageId="6" pageNumber="7" pagination="385 - 402" refId="ref42923" refString="Weise, J. (1892) Chrysomeliden und Coccinelliden von der Insel Nias, nebst Bemerkungen uber andere, meistens sudoestasiatische Arten. Deutsche Entomologische Zeitschrift, 1892 (2), 385 - 402. https: // doi. org / 10.1002 / mmnd. 48018920128" type="journal article" year="1892">Weise, 1892: 396</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="485BB3513F55E54A71DAC209FDB14174" author="Maulik, S." box="[450,632,334,359]" page="167" pageId="6" pageNumber="7" year="1936">
<bibRefCitation id="AD6BE8B13F55E54A71DAC209FDB14174" author="Maulik, S." box="[450,632,334,359]" pageId="6" pageNumber="7" refId="ref41439" refString="Maulik, S. (1936) The Fauna of British India including Ceylon and Burma. Coleoptera. Chrysomelidae. (Galerucinae). Taylor &amp; Francis, London, xiv + 648 pp., 1 pl." type="book" year="1936">Maulik 1936: 167</bibRefCitation>
</treatmentCitation>
(synonymy)
</treatmentCitationGroup>
</paragraph>
<paragraph id="C94595403F55E54A708FC228FCE74197" blockId="6.[151,959,150,451]" box="[151,814,365,389]" pageId="6" pageNumber="7">
<treatmentCitationGroup id="E9EAB26E3F55E54A708FC228FCE74197" box="[151,814,365,389]" pageId="6" pageNumber="7">
=
<taxonomicName id="0EFAEEC33F55E54A70B4C22BFE244197" authority="Weise, 1896: 396" authorityName="Weise" authorityPageNumber="396" authorityYear="1896" box="[172,493,365,389]" genus="Sphaerarthra" kingdom="Animalia" pageId="6" pageNumber="7" rank="genus">
<emphasis id="FB8E49523F55E54A70B4C22BFEFC4197" box="[172,309,365,389]" italics="true" pageId="6" pageNumber="7">Sphaerarthra</emphasis>
Weise, 1896: 396
</taxonomicName>
;
<treatmentCitation id="485BB3513F55E54A71E0C22BFD644197" author="Maulik, S." box="[504,685,365,389]" page="167" pageId="6" pageNumber="7" year="1936">
<bibRefCitation id="AD6BE8B13F55E54A71E0C22BFD644197" author="Maulik, S." box="[504,685,365,389]" pageId="6" pageNumber="7" refId="ref41439" refString="Maulik, S. (1936) The Fauna of British India including Ceylon and Burma. Coleoptera. Chrysomelidae. (Galerucinae). Taylor &amp; Francis, London, xiv + 648 pp., 1 pl." type="book" year="1936">Maulik 1936: 167</bibRefCitation>
</treatmentCitation>
(synonymy)
</treatmentCitationGroup>
</paragraph>
<paragraph id="C94595403F55E54A708FC2CAFCD741B6" blockId="6.[151,959,150,451]" box="[151,798,396,420]" pageId="6" pageNumber="7">
<treatmentCitationGroup id="E9EAB26E3F55E54A708FC2CAFCD741B6" box="[151,798,396,420]" pageId="6" pageNumber="7">
=
<taxonomicName id="0EFAEEC33F55E54A70B4C2CAFE1741B6" authority="Weise, 1896: 392" authorityName="Weise" authorityPageNumber="392" authorityYear="1896" box="[172,478,396,420]" genus="Pachypalpa" kingdom="Animalia" pageId="6" pageNumber="7" rank="genus">
<emphasis id="FB8E49523F55E54A70B4C2CAFEEC41B6" box="[172,293,396,420]" italics="true" pageId="6" pageNumber="7">Pachypalpa</emphasis>
Weise, 1896: 392
</taxonomicName>
;
<treatmentCitation id="485BB3513F55E54A71F0C2CAFD5741B6" author="Maulik, S." box="[488,670,396,420]" page="167" pageId="6" pageNumber="7" year="1936">
<bibRefCitation id="AD6BE8B13F55E54A71F0C2CAFD5741B6" author="Maulik, S." box="[488,670,396,420]" pageId="6" pageNumber="7" refId="ref41439" refString="Maulik, S. (1936) The Fauna of British India including Ceylon and Burma. Coleoptera. Chrysomelidae. (Galerucinae). Taylor &amp; Francis, London, xiv + 648 pp., 1 pl." type="book" year="1936">Maulik 1936: 167</bibRefCitation>
</treatmentCitation>
(synonymy)
</treatmentCitationGroup>
</paragraph>
<paragraph id="C94595403F55E54A708FC2EDFC8C41D1" blockId="6.[151,959,150,451]" box="[151,837,426,451]" pageId="6" pageNumber="7">
<treatmentCitationGroup id="E9EAB26E3F55E54A708FC2EDFC8C41D1" box="[151,837,426,451]" pageId="6" pageNumber="7">
=
<taxonomicName id="0EFAEEC33F55E54A70B4C2EDFDCC41D1" authority="Csiki, 1953: 131" authorityName="Csiki" authorityPageNumber="131" authorityYear="1953" box="[172,517,426,451]" genus="Paraulacophora" kingdom="Animalia" pageId="6" pageNumber="7" rank="genus">
<emphasis id="FB8E49523F55E54A70B4C2EDFE9A41D1" box="[172,339,427,451]" italics="true" pageId="6" pageNumber="7">Paraulacophora</emphasis>
Csiki, 1953: 131
</taxonomicName>
;
<treatmentCitation id="485BB3513F55E54A7217C2EDFD0C41D1" author="Wilcox, J. A." box="[527,709,427,451]" page="222" pageId="6" pageNumber="7" year="1972">
<bibRefCitation id="AD6BE8B13F55E54A7217C2EDFD0C41D1" author="Wilcox, J. A." box="[527,709,427,451]" pageId="6" pageNumber="7" pagination="221 - 431" refId="ref43057" refString="Wilcox, J. A. (1972) Chrysomelidae: Galerucinae: Luperini: Aulacophorina, Diabroticina. Coleopterorum Catalogus Supplementa, 78 (2), 221 - 431." type="journal article" year="1972">Wilcox 1972: 222</bibRefCitation>
</treatmentCitation>
(synonymy)
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="81E0C6CB3F55E54A708FC2AEFA85420C" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="C94595403F55E54A708FC2AEFA85420C" blockId="6.[151,1436,488,543]" pageId="6" pageNumber="7">
<typeStatus id="16412BE23F55E54A708FC2AEFF004212" box="[151,201,488,512]" pageId="6" pageNumber="7">Type</typeStatus>
species:
<taxonomicName id="0EFAEEC33F55E54A713FC2AEFD4A4212" authority="Olivier, 1808" authorityName="Olivier" authorityYear="1808" box="[295,643,488,512]" class="Insecta" family="Chrysomelidae" genus="Galeruca" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="quadraria">
<emphasis id="FB8E49523F55E54A713FC2AEFE3D4212" box="[295,500,488,512]" italics="true" pageId="6" pageNumber="7">Galeruca quadraria</emphasis>
<bibRefCitation id="AD6BE8B13F55E54A71E3C2AEFD4A4212" author="Olivier, A. G." box="[507,643,488,512]" pageId="6" pageNumber="7" pagination="613 - 1104" refId="ref41936" refString="Olivier, A. G. (1808) Entomologie, ou histoire naturelle des insectes, avec leur caracteres generiques et specifiques, leur description, leur synonymie, et leur figure enluminee. Coleopteres. Tome sixieme. Desray, Paris, pp. 613 - 1104, 46 pls." type="book chapter" year="1808">Olivier, 1808</bibRefCitation>
</taxonomicName>
, by subsequent designation (
<bibRefCitation id="AD6BE8B13F55E54A73BFC2AEFB124212" author="Duponchel, P &amp; Chevrolat, L. A. A." box="[935,1243,488,512]" pageId="6" pageNumber="7" pagination="210 - 211" refId="ref39883" refString="Duponchel, P, &amp; Chevrolat, L. A. A. (1842) Aulacophora. In: d'Orbigny, C. (Ed.), Dictionnaire universel d'histoire naturelle. Tome deuxieme. MM Renard, Martinet et Cie, Paris, pp. 210 - 211." type="book chapter" year="1842">Duponchel &amp; Chevrolat 1842</bibRefCitation>
). This name is currently considered a junior synonym of the south-east Asian species
<taxonomicName id="0EFAEEC33F55E54A736AC141FA81420C" authority="(Weber, 1801) (Lee &amp; Beenen 2015)" authorityName="Lee &amp; Beenen" authorityYear="2015" baseAuthorityName="Weber" baseAuthorityYear="1801" box="[882,1352,518,543]" class="Insecta" family="Chrysomelidae" genus="Acutipalpa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="analis">
<emphasis id="FB8E49523F55E54A736AC141FC05420D" box="[882,972,519,543]" italics="true" pageId="6" pageNumber="7">A. analis</emphasis>
(Weber, 1801) (
<bibRefCitation id="AD6BE8B13F55E54A7468C141FA88420C" author="Lee, C. - F. &amp; Beenen, R." box="[1136,1345,518,543]" pageId="6" pageNumber="7" pagination="151 - 190" refId="ref41061" refString="Lee, C. - F. &amp; Beenen, R. (2015) Revision of the genus Aulacophora from Taiwan (Coleoptera: Chrysomelidae: Galerucinae). Zootaxa, 3949 (2), 151 - 190. https: // doi. org / 10.11646 / zootaxa. 3949.2.1" type="journal article" year="2015">Lee &amp; Beenen 2015</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="81E0C6CB3F55E54A708FC10AFD0542BD" pageId="6" pageNumber="7" type="diagnosis">
<paragraph id="C94595403F55E54A708FC10AFE6B4274" blockId="6.[151,1437,588,1983]" box="[151,418,588,614]" pageId="6" pageNumber="7">
<emphasis id="FB8E49523F55E54A708FC10AFE6B4274" bold="true" box="[151,418,588,614]" pageId="6" pageNumber="7">Diagnostic description</emphasis>
</paragraph>
<paragraph id="C94595403F55E54A70DFC137FD0542BD" blockId="6.[151,1437,588,1983]" pageId="6" pageNumber="7">
<emphasis id="FB8E49523F55E54A70DFC137FE534299" box="[199,410,625,651]" italics="true" pageId="6" pageNumber="7">Generic diagnosis.</emphasis>
The following diagnostic description is based on the Australopapuan species and the generic redefinition by
<bibRefCitation id="AD6BE8B13F55E54A715CC1D3FD0142BD" author="Barroga, G. F. &amp; Mohamedsaid, M. S." box="[324,712,661,687]" pageId="6" pageNumber="7" pagination="15 - 194" refId="ref39042" refString="Barroga, G. F. &amp; Mohamedsaid, M. S. (2002) Revision of the genus Aulacophora Chevrolat (Coleoptera: Chrysomelidae: Galerucinae) in Sundaland. Serangga, 7 (1 - 2), 15 - 194." type="journal article" year="2002">Barroga and Mohamedsaid (2002)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="81E0C6CB3F55E54A70DFC1FFFAE747AD" pageId="6" pageNumber="7" type="description">
<paragraph id="C94595403F55E54A70DFC1FFFECE4395" blockId="6.[151,1437,588,1983]" pageId="6" pageNumber="7">
Body
<taxonomicName id="0EFAEEC33F55E54A7113C1FFFE6E42C1" box="[267,423,697,723]" form="elongate" pageId="6" pageNumber="7" rank="form">form elongate</taxonomicName>
pear-shaped in dorsal view, length about 2x width, head, prothorax and elytra at shoulders varying slightly in proportions but generally successively wider by a factor of 1.5; dorsally sparsely and finely punctured (interspaces&gt; puncture diameters), except pronotal anterolateral patch of secretory pits, and dorsally glabrous except trichobothria on frons and pronotal angles, semi-erect setae on clypeus, short erect setae on elytral lateral margins and some males with patch of setae posterior to elytral humeri; closely setose ventrally and on appendages.
</paragraph>
<paragraph id="C94595403F55E54A70DFC0D7FBFB4501" blockId="6.[151,1437,588,1983]" pageId="6" pageNumber="7">
<emphasis id="FB8E49523F55E54A70DFC0D7FECD43B9" box="[199,260,913,939]" italics="true" pageId="6" pageNumber="7">Head</emphasis>
. Vertex smooth and impunctate, except a single trichobothrium above each eye, without longitudinal median groove; row of five pairs of long setae anterior to antennal sockets; frontoclypeus medially ridged and elevated from between antennal sockets towards anterior; anterior edge frontoclypeus truncate, shallowly concave or medially excavate; interantennal space 0.51.0x socket diameter, sockets level with anterior half of eyes and separated from them by 0.50.75x socket widths; eyes large and laterally prominent, width of head across eyes about 1.5x width of head at anterior eye margins; gena 0.50.7x eye length; postantennal calli distinct, slightly convex and approximately triangular, divided by short median groove, bases truncate and defined by deep postcallal groove extending laterally to posterior edge of eyes, sides defined by rim of antennal sockets, internal margins defined by grooves at base of frontoclypeal ridge; labrum with four pairs of discal setae, apical margin truncate; antennae 0.50.8x body length; all antennomeres elongate, 311 narrow and of similar width except some may be expanded in males; apical maxillary palpomere conical, shorter and narrower than preapical.
</paragraph>
<paragraph id="C94595403F55E54A70DFC65BFAB94719" blockId="6.[151,1437,588,1983]" pageId="6" pageNumber="7">
<emphasis id="FB8E49523F55E54A70DFC65BFED14525" box="[199,280,1309,1335]" italics="true" pageId="6" pageNumber="7">Thorax</emphasis>
. Pronotum transverse in dorsal view, widest anterior to middle, basal 2/3 straight edged and slightly contracted to base, anterior 1/3 strongly contracted to obtuse apical angles; pronotal lateral margins sinuate in lateral view; all pronotal sides with raised margins, margination broadest laterally and narrowest anteriorly (may be partly absent); anterior and posterior angles each with a trichobothrium, posterior angles about 90º; pronotum with transverse or sinuate groove at middle; prosternal process triangular, tapering to a thin elevated blade which terminates at about half length of procoxae; procoxae conical; procoxal cavities externally wide open, gap about length of hypomeral process; scutellum flat, approximately triangular with rounded apex; elytra usually at least slightly expanded from base to about 2/3 length, with rounded apex, often leaving pygidium partially exposed; elytra nonstriate, non-carinate, with or without transverse posthumeral depression; epipleura broad at base, abruptly contracted between meso and metacoxae and not defined beyond this point; fully winged; mesoventrite not covered by metaventrite, apex acutely pointed and base without procoxal rests; mesocoxae separated by much less than width of coxa; femora almost parallel-sided; tibiae with or without single apical spur, if present, length about half or slightly less than apical width of tibia metatarsus about 0.8x as long as metatibia; first metatarsomere as long as or slightly longer than metatarsomeres 2+3; tarsal claws bifid, inner teeth convergent and about 2/3 as long as outer teeth.
</paragraph>
<paragraph id="C94595403F55E54A70DFC453FAE747AD" blockId="6.[151,1437,588,1983]" pageId="6" pageNumber="7">
<emphasis id="FB8E49523F55E54A70DFC453FEF8473D" box="[199,305,1813,1839]" italics="true" pageId="6" pageNumber="7">Abdomen</emphasis>
. Ventrites not laterally bordered by raised bead; ventrite I with rounded to broadly triangular intercoxal process; apex of male ventrite V with rectangular lobe defined on either side by slots; apex of female ventrite V truncate or concave; penis symmetrical or slightly asymmetric, apex entire, base without recurved lobes; ovipositor with well-developed apodeme (tignum) at base of sternite VIII; palpi one-segmented; spermathecal receptaculum falcate or C-shaped, transversely grooved at base and with swollen collum at insertion of duct and gland.
</paragraph>
</subSubSection>
<subSubSection id="81E0C6CB3F54E54B708FC3D0FD8A422F" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="C94595403F54E54B708FC3D0FCCE40A2" blockId="7.[151,1437,150,2013]" box="[151,775,150,177]" pageId="7" pageNumber="8">
<emphasis id="FB8E49523F54E54B708FC3D0FCCE40A2" bold="true" box="[151,775,150,177]" pageId="7" pageNumber="8">
Diagnosis of the
<taxonomicName id="0EFAEEC33F54E54B7144C3D0FD8B40A3" authorityName="Luzon" baseAuthorityName="Gmelin" baseAuthorityYear="1790" box="[348,578,150,177]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="FB8E49523F54E54B7144C3D0FD8B40A3" bold="true" box="[348,578,150,177]" italics="true" pageId="7" pageNumber="8">Aulacophora indica</emphasis>
</taxonomicName>
species-complex
</emphasis>
</paragraph>
<paragraph id="C94595403F54E54B70DFC3FCFD8A422F" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="8">
Head capsule, pronotum and elytra entirely pale (brownish-yellow when dead, orange or red when alive) (
<figureCitation id="51C189C53F54E54B7572C3FCFF0D40EA" captionStart-0="FIGURES 25" captionStart-1="FIGURES 69" captionStartId-0="15.[151,264,1829,1853]" captionStartId-1="16.[151,264,1957,1981]" captionTargetBox-0="[164,1420,191,1786]" captionTargetBox-1="[174,1418,193,1917]" captionTargetId-0="figure-19@15.[151,1436,181,1805]" captionTargetId-1="figure-22@16.[157,1430,181,1933]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionText-0="FIGURES 25. Aulacophora species, male habitus: 2, A. abdominalis (Fabricius, 1781) (Guadalcanal); 3, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 4, A. cornuta Baly, 1879 (New Ireland); 5, A. indica (Gmelin, 1790) (Luzon)." captionText-1="FIGURES 69. Aulacophora species, male habitus: 6, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 7, A. relicta (Boisduval, 1835) (Mount Molloy); 8, A. wallacii Baly, 1886; 9, A. wilsoni Baly, 1888." figureDoi-0="http://doi.org/10.5281/zenodo.4545452" figureDoi-1="http://doi.org/10.5281/zenodo.4545458" httpUri-0="https://zenodo.org/record/4545452/files/figure.png" httpUri-1="https://zenodo.org/record/4545458/files/figure.png" pageId="7" pageNumber="8">Figs 29</figureCitation>
), one Japanese species,
<taxonomicName id="0EFAEEC33F54E54B71CBC399FCD540EA" authority="(Olivier, 1808)" baseAuthorityName="Olivier" baseAuthorityYear="1808" box="[467,796,222,249]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="bipunctata">
<emphasis id="FB8E49523F54E54B71CBC399FDA240EA" box="[467,619,222,249]" italics="true" pageId="7" pageNumber="8">A. bipunctata</emphasis>
(
<bibRefCitation id="AD6BE8B13F54E54B7262C398FCDA40EA" author="Olivier, A. G." box="[634,787,222,248]" pageId="7" pageNumber="8" pagination="613 - 1104" refId="ref41936" refString="Olivier, A. G. (1808) Entomologie, ou histoire naturelle des insectes, avec leur caracteres generiques et specifiques, leur description, leur synonymie, et leur figure enluminee. Coleopteres. Tome sixieme. Desray, Paris, pp. 613 - 1104, 46 pls." type="book chapter" year="1808">Olivier, 1808</bibRefCitation>
)
</taxonomicName>
, with small black spots; male first antennomere approximately pyramidal in shape, expanded from base to middle, with a flat area on most of its anterior face (
<figureCitation id="51C189C53F54E54B7515C244FA46410E" box="[1293,1423,258,284]" captionStart-0="FIGURES 1015" captionStart-1="FIGURES 1617" captionStartId-0="17.[151,264,1943,1967]" captionStartId-1="18.[151,264,783,807]" captionTargetBox-0="[209,1375,181,1906]" captionTargetBox-1="[158,1427,191,748]" captionTargetId-0="figure-19@17.[209,1379,181,1920]" captionTargetId-1="figure-22@18.[151,1435,181,759]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionText-0="FIGURES 1015. Aulacophora species, male head, anterior view: 10, A. abdominalis (Fabricius, 1781) (Guadalcanal); 11, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 12, A. cornuta Baly, 1879 (New Ireland); 13, A. indica (Gmelin, 1790) (Luzon); 14, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 15, A. relicta (Boisduval, 1835) (Mount Molloy)." captionText-1="FIGURES 1617. Aulacophora species, male head, anterior view: 16, A. wallacii Baly, 1886; 17, A. wilsoni Baly, 1888." figureDoi-0="http://doi.org/10.5281/zenodo.4545460" figureDoi-1="http://doi.org/10.5281/zenodo.4545462" httpUri-0="https://zenodo.org/record/4545460/files/figure.png" httpUri-1="https://zenodo.org/record/4545462/files/figure.png" pageId="7" pageNumber="8">Figs 1017</figureCitation>
); male antennomeres 311 not modified (
<figureCitation id="51C189C53F54E54B7242C260FD164153" box="[602,735,294,321]" captionStart="FIGURES 1825" captionStartId="21.[151,264,1899,1923]" captionTargetBox="[262,1328,190,1869]" captionTargetId="figure-19@21.[250,1338,181,1875]" captionTargetPageId="21" captionText="FIGURES 1825. Aulacophora species, male anterior and antenna: 18, A. abdominalis (Fabricius, 1781) (Guadalcanal); 19, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 20, A. cornuta Baly, 1879 (New Ireland); 21, A. indica (Gmelin, 1790) (Luzon); 22, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 23, A. relicta (Boisduval, 1835) (Mount Molloy); 24, A. wallacii Baly, 1886; 25, A. wilsoni Baly, 1888." figureDoi="http://doi.org/10.5281/zenodo.4545464" httpUri="https://zenodo.org/record/4545464/files/figure.png" pageId="7" pageNumber="8">Figs 1825</figureCitation>
); male with deeper transverse prothoracic groove than female (
<figureCitation id="51C189C53F54E54B7087C20CFEEB4176" box="[159,290,330,356]" captionStart="FIGURES 2631" captionStartId="22.[151,264,1703,1727]" captionTargetBox="[160,1426,191,1670]" captionTargetId="figure-22@22.[151,1436,181,1679]" captionTargetPageId="22" captionText="FIGURES 2631. Aulacophora species, pronotum, lateral view: 26, male A. abdominalis (Fabricius, 1781) (Vanuatu); 27, male A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 28, male A. indica (Gmelin, 1790) (Borneo); 29, female A. indica; 30, male A. relicta (Boisduval, 1835); 31, male A. wallacii Baly, 1886." figureDoi="http://doi.org/10.5281/zenodo.4545466" httpUri="https://zenodo.org/record/4545466/files/figure.png" pageId="7" pageNumber="8">Figs 2829</figureCitation>
); elytra without posthumeral transverse depression, males of some species with lateral patch of erect setae posterior to humeri (
<figureCitation id="51C189C53F54E54B71BBC228FDEC419B" box="[419,549,366,393]" captionStart="FIGURES 1825" captionStartId="21.[151,264,1899,1923]" captionTargetBox="[262,1328,190,1869]" captionTargetId="figure-19@21.[250,1338,181,1875]" captionTargetPageId="21" captionText="FIGURES 1825. Aulacophora species, male anterior and antenna: 18, A. abdominalis (Fabricius, 1781) (Guadalcanal); 19, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 20, A. cornuta Baly, 1879 (New Ireland); 21, A. indica (Gmelin, 1790) (Luzon); 22, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 23, A. relicta (Boisduval, 1835) (Mount Molloy); 24, A. wallacii Baly, 1886; 25, A. wilsoni Baly, 1888." figureDoi="http://doi.org/10.5281/zenodo.4545464" httpUri="https://zenodo.org/record/4545464/files/figure.png" pageId="7" pageNumber="8">Figs 1825</figureCitation>
); all tibiae with single apical spur (
<figureCitation id="51C189C53F54E54B73B3C228FC36419A" box="[939,1023,366,392]" captionStart="FIGURES 1825" captionStartId="21.[151,264,1899,1923]" captionTargetBox="[262,1328,190,1869]" captionTargetId="figure-19@21.[250,1338,181,1875]" captionTargetPageId="21" captionText="FIGURES 1825. Aulacophora species, male anterior and antenna: 18, A. abdominalis (Fabricius, 1781) (Guadalcanal); 19, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 20, A. cornuta Baly, 1879 (New Ireland); 21, A. indica (Gmelin, 1790) (Luzon); 22, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 23, A. relicta (Boisduval, 1835) (Mount Molloy); 24, A. wallacii Baly, 1886; 25, A. wilsoni Baly, 1888." figureDoi="http://doi.org/10.5281/zenodo.4545464" httpUri="https://zenodo.org/record/4545464/files/figure.png" pageId="7" pageNumber="8">Fig. 24</figureCitation>
); female pygidium usually elongated beyond apex of apical ventrite (
<figureCitation id="51C189C53F54E54B71E4C2D4FD4941BF" box="[508,640,402,429]" captionStart-0="FIGURES 4449" captionStart-1="FIGURES 5055" captionStart-2="FIGURES 5657" captionStartId-0="25.[151,264,1798,1822]" captionStartId-1="26.[151,264,1943,1967]" captionStartId-2="27.[151,264,701,725]" captionTargetBox-0="[170,1418,190,1763]" captionTargetBox-1="[198,1388,201,1907]" captionTargetBox-2="[165,1415,191,663]" captionTargetId-0="figure-19@25.[151,1436,181,1774]" captionTargetId-1="figure-22@26.[178,1409,181,1919]" captionTargetId-2="figure-19@27.[151,1436,181,677]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionTargetPageId-2="27" captionText-0="FIGURES 4449. Aulacophora species, female, apex of abdominal venter and pygidium: 44, A. abdominalis (Fabricius, 1781) (Vanuatu); 45, A. abdominalis (New Ireland); 46, A. abdominalis (Mer Island); 47, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 48, A. cornuta Baly, 1879 (Sulawesi); 49, A. indica (Gmelin, 1790) (Laos).Dotted lines indicate depth and width of apical concavity." captionText-1="FIGURES 5055. Aulacophora species, female, apex of abdominal venter: 50, A. indica (Gmelin, 1790) (Biak Island); 51, A. indica (Vietnam); 52, A. indica (Broken Hill); 53, A. mbabaram Reid, Halling &amp; Beatson, sp. nov., Almaden; 54, A. relicta (Boisduval, 1835); 55, A. wallacii Baly, 1886." captionText-2="FIGURES 5657. Aulacophora species, female, apex of abdominal venter: 56, A. wilsoni Baly, 1888 (Morgans Creek); 57, A. wilsoni (Eungella)." figureDoi-0="http://doi.org/10.5281/zenodo.4545472" figureDoi-1="http://doi.org/10.5281/zenodo.4545474" figureDoi-2="http://doi.org/10.5281/zenodo.4545476" httpUri-0="https://zenodo.org/record/4545472/files/figure.png" httpUri-1="https://zenodo.org/record/4545474/files/figure.png" httpUri-2="https://zenodo.org/record/4545476/files/figure.png" pageId="7" pageNumber="8">Figs 4457</figureCitation>
); apex of male last ventrite with elongate-oval cavity on elongate-rectangular middle lobe, which is defined on either side by narrow slots (
<figureCitation id="51C189C53F54E54B73BCC2F0FBE141C2" box="[932,1064,438,464]" captionStart-0="FIGURES 3237" captionStart-1="FIGURES 3843" captionStartId-0="23.[151,264,1840,1864]" captionStartId-1="24.[151,264,1793,1817]" captionTargetBox-0="[167,1416,192,1802]" captionTargetBox-1="[168,1418,191,1757]" captionTargetId-0="figure-19@23.[151,1436,181,1816]" captionTargetId-1="figure-22@24.[151,1437,181,1769]" captionTargetPageId-0="23" captionTargetPageId-1="24" captionText-0="FIGURES 3237. Aulacophora species, male, apex of abdominal venter: 32, A. abdominalis (Fabricius, 1781) (Vanuatu); 33, A. abdominalis (New Ireland); 34, A. abdominalis (Manatuto); 35, A. abdominalis (Metinaro); 36, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 37, A. cornuta Baly, 1879 (Guadalcanal)." captionText-1="FIGURES 3843. Aulacophora species, male, apex of abdominal venter: 38, A. indica (Gmelin, 1790) (Sumatra); 39, A. indica (West Timor); 40, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 41, A. relicta (Boisduval, 1835) (Mount Molloy); 42, A. wallacii Baly, 1886; 43, A. wilsoni Baly, 1888." figureDoi-0="http://doi.org/10.5281/zenodo.4545468" figureDoi-1="http://doi.org/10.5281/zenodo.4545470" httpUri-0="https://zenodo.org/record/4545468/files/figure.png" httpUri-1="https://zenodo.org/record/4545470/files/figure.png" pageId="7" pageNumber="8">Figs 3243</figureCitation>
); male tergite VIII thickened and modified into either a transverse bar or a pair of posteriorly directed prongs (
<figureCitation id="51C189C53F54E54B73FCC29CFBAC41E6" box="[996,1125,474,500]" captionStart-0="FIGURES 8388" captionStart-1="FIGURES 8993" captionStartId-0="42.[151,264,1943,1967]" captionStartId-1="43.[151,264,1944,1968]" captionTargetBox-0="[217,1368,192,1905]" captionTargetBox-1="[196,1391,191,1907]" captionTargetId-0="figure-22@42.[204,1384,181,1919]" captionTargetId-1="figure-19@43.[177,1410,181,1920]" captionTargetPageId-0="42" captionTargetPageId-1="43" captionText-0="FIGURES 8388. Aulacophora species, male sternite 8, ventrolateral view: 83, A. abdominalis (Fabricius, 1781) (Vanuatu); 84, A. abdominalis (New Ireland); 85, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 86, A. cornuta Baly, 1879 (Guadalcanal); 87, A. indica (Gmelin, 1790) (Timor); 88, A. indica (Luzon)." captionText-1="FIGURES 8993. Aulacophora species, male sternite 8, ventrolateral view: 89, A. indica (Gmelin, 1790) (Sumatra); 90, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 91, A. relicta (Boisduval, 1835) (Coonabarabran); 92, A. wallacii Baly, 1886; 93, A. wilsoni Baly, 1888." figureDoi-0="http://doi.org/10.5281/zenodo.4545484" figureDoi-1="http://doi.org/10.5281/zenodo.4545486" httpUri-0="https://zenodo.org/record/4545484/files/figure.png" httpUri-1="https://zenodo.org/record/4545486/files/figure.png" pageId="7" pageNumber="8">Figs 8393</figureCitation>
); penis asymmetricin dorsal view, except
<taxonomicName id="0EFAEEC33F54E54B7133C2B9FE6C420A" authorityName="Baly" authorityYear="1886" box="[299,421,510,537]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="wallacii">
<emphasis id="FB8E49523F54E54B7133C2B9FE6C420A" box="[299,421,510,537]" italics="true" pageId="7" pageNumber="8">A. wallacii</emphasis>
</taxonomicName>
(
<figureCitation id="51C189C53F54E54B71ADC2B8FD81420A" box="[437,584,510,536]" captionStart-0="FIGURES 9497" captionStart-1="FIGURES 98101" captionStart-2="FIGURES 102105" captionStart-3="FIGURES 106107" captionStart-4="FIGURES 108111" captionStartId-0="44.[151,264,1737,1761]" captionStartId-1="45.[151,264,1825,1849]" captionStartId-2="46.[151,264,1777,1801]" captionStartId-3="47.[151,264,1987,2011]" captionStartId-4="48.[151,264,1843,1867]" captionTargetBox-0="[170,1385,198,1674]" captionTargetBox-1="[164,1393,195,1754]" captionTargetBox-2="[166,1388,194,1718]" captionTargetBox-3="[425,1163,195,1932]" captionTargetBox-4="[184,1420,192,1788]" captionTargetId-0="figure-22@44.[151,1435,181,1713]" captionTargetId-1="figure-19@45.[151,1436,181,1801]" captionTargetId-2="figure-22@46.[151,1436,181,1754]" captionTargetId-3="figure-19@47.[405,1183,181,1963]" captionTargetId-4="figure-22@48.[151,1436,181,1820]" captionTargetPageId-0="44" captionTargetPageId-1="45" captionTargetPageId-2="46" captionTargetPageId-3="47" captionTargetPageId-4="48" captionText-0="FIGURES 9497. Aulacophora abdominalis (Fabricius, 1781), penis, dorsal view: 94, Guadalcanal, 95, Metinaro, 96, Saibai, 97, Vanuatu." captionText-1="FIGURES 98101. Aulacophora species, penis, dorsal view: 98, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 99, A. cornuta Baly, 1879 (Guadalcanal); 100, A. indica (Gmelin, 1790) (Laos); 101, A. indica (Luzon)." captionText-2="FIGURES 102105. Aulacophora species, penis, dorsal view: 102, A. indica (Gmelin, 1790) (Timor); 103, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 104, A. relicta (Boisduval, 1835) (Mount Molloy); 105, A. relicta (Mount Mulligan)." captionText-3="FIGURES 106107. Aulacophora species, penis, dorsal view: 106, A. wilsoni Baly, 1888; 107, A. wallacii Baly, 1886." captionText-4="FIGURES 108111. Aulacophora abdominalis (Fabricius, 1781), penis, ventral view: 108, Guadalcanal, 109, Metinaro, 110, Saibai, 111, Vanuatu." figureDoi-0="http://doi.org/10.5281/zenodo.4545488" figureDoi-1="http://doi.org/10.5281/zenodo.4545490" figureDoi-2="http://doi.org/10.5281/zenodo.4545492" figureDoi-3="http://doi.org/10.5281/zenodo.4545494" figureDoi-4="http://doi.org/10.5281/zenodo.4545496" httpUri-0="https://zenodo.org/record/4545488/files/figure.png" httpUri-1="https://zenodo.org/record/4545490/files/figure.png" httpUri-2="https://zenodo.org/record/4545492/files/figure.png" httpUri-3="https://zenodo.org/record/4545494/files/figure.png" httpUri-4="https://zenodo.org/record/4545496/files/figure.png" pageId="7" pageNumber="8">Figs 94108</figureCitation>
). Hostplants where known are predominantly cucurbits (
<taxonomicName id="0EFAEEC33F54E54B74DDC2B8FAA0420B" baseAuthorityName="Reid" baseAuthorityYear="1998" box="[1221,1385,510,537]" class="Magnoliopsida" family="Cucurbitaceae" kingdom="Plantae" order="Cucurbitales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="family">Cucurbitaceae</taxonomicName>
) but may also include legumes (
<taxonomicName id="0EFAEEC33F54E54B71D0C164FDFE422F" box="[456,567,546,573]" class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" order="Fabales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="family">Fabaceae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="81E0C6CB3F54E54570DFC100FE2F47AA" lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8" type="discussion">
<paragraph id="C94595403F54E54B70DFC100FA5F4306" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="8">
The group of species designated here as the Pale Pumpkin Beetles (PPB) or
<taxonomicName id="0EFAEEC33F54E54B740BC101FBB14272" authorityName="Luzon" baseAuthorityName="Gmelin" baseAuthorityYear="1790" box="[1043,1144,582,609]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="FB8E49523F54E54B740BC101FBB14272" box="[1043,1144,582,609]" italics="true" pageId="7" pageNumber="8">A. indica</emphasis>
</taxonomicName>
species-complex (Appendix 1) is a group of externally similar species that are difficult to separate. The following described species belong in this complex, based on examination of material, or descriptions:
<emphasis id="FB8E49523F54E54B7382C1C9FB2442BA" box="[922,1261,654,681]" italics="true" pageId="7" pageNumber="8">
<taxonomicName id="0EFAEEC33F54E54B7382C1C9FB8E42BA" authorityName="Mer" baseAuthorityName="Fabricius" baseAuthorityYear="1781" box="[922,1095,654,681]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="abdominalis">A. abdominalis</taxonomicName>
,
<taxonomicName id="0EFAEEC33F54E54B744DC1C9FB2442BA" baseAuthorityName="Olivier" baseAuthorityYear="1808" box="[1109,1261,654,681]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="bipunctata">A. bipunctata</taxonomicName>
</emphasis>
,
<taxonomicName id="0EFAEEC33F54E54B74E5C1C9FABD42BA" authorityName="Baly" authorityYear="1879" box="[1277,1396,655,681]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="FB8E49523F54E54B74E5C1C9FABD42BA" box="[1277,1396,655,681]" italics="true" pageId="7" pageNumber="8">A. cornuta</emphasis>
</taxonomicName>
,
<taxonomicName id="0EFAEEC33F54E54B759CC1C9FE6442DE" authority="(Lucas, 1849)" baseAuthorityName="Lucas" baseAuthorityYear="1849" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="foveicollis">
<emphasis id="FB8E49523F54E54B759CC1C9FEC342DE" italics="true" pageId="7" pageNumber="8">A. foveicollis</emphasis>
(
<bibRefCitation id="AD6BE8B13F54E54B7100C1F4FE6F42DE" author="Lucas, P. H." box="[280,422,690,716]" pageId="7" pageNumber="8" refId="ref41245" refString="Lucas, P. H. (1849) Cinquieme classe, Insectes. Premier ordre. Les Coleopteres. Pp. 449 - 590, in: Exploration scientifique de l'Algerie pendant les annees 1840, 1841, 1842, publiee par ordre du gouvernement et avec le concours d'une Commission Academique. Sciences physiques zoologie. Vol. II. Histoire naturelle des animaux articules. Imprimerie Nationale, Paris, 590 pp." type="book" year="1849">Lucas, 1849</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="0EFAEEC33F54E54B71A0C1F5FDD542DE" authorityName="Luzon" baseAuthorityName="Gmelin" baseAuthorityYear="1790" box="[440,540,690,717]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="FB8E49523F54E54B71A0C1F5FDD542DE" box="[440,540,690,717]" italics="true" pageId="7" pageNumber="8">A. indica</emphasis>
</taxonomicName>
,
<taxonomicName id="0EFAEEC33F54E54B7231C1F5FC8B42DE" authority="Chujo, 1962" authorityName="Chujo" authorityYear="1962" box="[553,834,690,717]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="kotoensis">
<emphasis id="FB8E49523F54E54B7231C1F5FD6642DE" box="[553,687,690,717]" italics="true" pageId="7" pageNumber="8">A. kotoensis</emphasis>
Chujo, 1962
</taxonomicName>
,
<taxonomicName id="0EFAEEC33F54E54B7354C1F5FC7B42DE" authorityName="Mount Molloy" baseAuthorityName="Boisduval" baseAuthorityYear="1835" box="[844,946,690,717]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="relicta">
<emphasis id="FB8E49523F54E54B7354C1F5FC7B42DE" box="[844,946,690,717]" italics="true" pageId="7" pageNumber="8">A. relicta</emphasis>
</taxonomicName>
,
<taxonomicName id="0EFAEEC33F54E54B73A7C1F5FA8B42DE" authority="Mohamedsaid, 2009" authorityName="Mohamedsaid" authorityYear="2009" box="[959,1346,690,717]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="sulaksonoi">
<emphasis id="FB8E49523F54E54B73A7C1F5FB9F42DE" box="[959,1110,690,717]" italics="true" pageId="7" pageNumber="8">A. sulaksonoi</emphasis>
Mohamedsaid, 2009
</taxonomicName>
,
<taxonomicName id="0EFAEEC33F54E54B7554C1F5FF0042E2" authorityName="Baly" authorityYear="1886" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="wallacii">
<emphasis id="FB8E49523F54E54B7554C1F5FF0042E2" italics="true" pageId="7" pageNumber="8">A. wallacii</emphasis>
</taxonomicName>
and
<taxonomicName id="0EFAEEC33F54E54B7118C191FEB942E2" authorityName="Baly" authorityYear="1888" box="[256,368,726,753]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="wilsoni">
<emphasis id="FB8E49523F54E54B7118C191FEB942E2" box="[256,368,726,753]" italics="true" pageId="7" pageNumber="8">A. wilsoni</emphasis>
</taxonomicName>
. Although the Javanese endemic
<taxonomicName id="0EFAEEC33F54E54B72FFC191FCB742E2" baseAuthorityName="Olivier" baseAuthorityYear="1808" box="[743,894,726,753]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="bipunctata">
<emphasis id="FB8E49523F54E54B72FFC191FCB742E2" box="[743,894,726,753]" italics="true" pageId="7" pageNumber="8">A. bipunctata</emphasis>
</taxonomicName>
has small elytral spots, it otherwise conforms to the definition given above and its male genitalia are similar to those of
<taxonomicName id="0EFAEEC33F54E54B73ACC1BDFA5B4306" authority="(Barroga &amp; Mohamedsaid 2002)" baseAuthorityName="Barroga &amp; Mohamedsaid" baseAuthorityYear="2002" box="[948,1426,762,789]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="FB8E49523F54E54B73ACC1BDFBD04306" box="[948,1049,762,789]" italics="true" pageId="7" pageNumber="8">A. indica</emphasis>
(
<bibRefCitation id="AD6BE8B13F54E54B7430C1BCFA434306" author="Barroga, G. F. &amp; Mohamedsaid, M. S." box="[1064,1418,762,788]" pageId="7" pageNumber="8" pagination="15 - 194" refId="ref39042" refString="Barroga, G. F. &amp; Mohamedsaid, M. S. (2002) Revision of the genus Aulacophora Chevrolat (Coleoptera: Chrysomelidae: Galerucinae) in Sundaland. Serangga, 7 (1 - 2), 15 - 194." type="journal article" year="2002">Barroga &amp; Mohamedsaid 2002</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="C94595403F54E54B70DFC058FDFA4427" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="8">
The
<taxonomicName id="0EFAEEC33F54E54B70E2C059FEA9432A" baseAuthorityName="Barroga &amp; Mohamedsaid" baseAuthorityYear="2002" box="[250,352,798,825]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="FB8E49523F54E54B70E2C059FEA9432A" box="[250,352,798,825]" italics="true" pageId="7" pageNumber="8">A. indica</emphasis>
</taxonomicName>
species-complex is approximately equivalent to the genus
<taxonomicName id="0EFAEEC33F54E54B73E1C058FAB0432B" authority="Rosenhauer, 1856" authorityName="Rosenhauer" authorityYear="1856" box="[1017,1401,798,825]" class="Insecta" family="Chrysomelidae" genus="Rhaphidopalpa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="FB8E49523F54E54B73E1C058FB6F432A" box="[1017,1190,798,824]" italics="true" pageId="7" pageNumber="8">Rhaphidopalpa</emphasis>
<bibRefCitation id="AD6BE8B13F54E54B74B5C058FAB0432B" author="Rosenhauer, W. G." box="[1197,1401,798,825]" pageId="7" pageNumber="8" refId="ref42323" refString="Rosenhauer, W. G. (1856) Thiere Andalusiens nach dem Resultate einer Reise zusammengestellt nebst den Beschreibungen von 249 neuen oder bis jetzt unbeschriebenen Gattungen und Arten. T. Blaesing, Erlangen, viii + 429 pp, 3 pls. https: // doi. org / 10.5962 / bhl. title. 66016" type="book" year="1856">Rosenhauer, 1856</bibRefCitation>
</taxonomicName>
, as defined by
<bibRefCitation id="AD6BE8B13F54E54B710EC004FE65434E" author="Weise, J." box="[278,428,834,860]" pageId="7" pageNumber="8" pagination="385 - 402" refId="ref42923" refString="Weise, J. (1892) Chrysomeliden und Coccinelliden von der Insel Nias, nebst Bemerkungen uber andere, meistens sudoestasiatische Arten. Deutsche Entomologische Zeitschrift, 1892 (2), 385 - 402. https: // doi. org / 10.1002 / mmnd. 48018920128" type="journal article" year="1892">Weise (1892)</bibRefCitation>
using male and female secondary sexual characters. By including only males with setose humeri,
<taxonomicName id="0EFAEEC33F54E54B70EDC020FE6B4392" authorityName="Rosenhauer" authorityYear="1856" box="[245,418,870,896]" class="Insecta" family="Chrysomelidae" genus="Rhaphidopalpa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="FB8E49523F54E54B70EDC020FE6B4392" box="[245,418,870,896]" italics="true" pageId="7" pageNumber="8">Rhaphidopalpa</emphasis>
</taxonomicName>
was constrained to be a smaller group than the definition of PPB included here. Weise erroneously abandoned the name
<taxonomicName id="0EFAEEC33F54E54B71E3C0CCFD4443B6" authorityName="Chevrolat" authorityYear="1836" box="[507,653,906,932]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="FB8E49523F54E54B71E3C0CCFD4443B6" box="[507,653,906,932]" italics="true" pageId="7" pageNumber="8">Aulacophora</emphasis>
</taxonomicName>
because it had been used for a plant genus, described
<taxonomicName id="0EFAEEC33F54E54B74E8C0CDFA9C43B6" baseAuthorityName="Barroga &amp; Mohamedsaid" baseAuthorityYear="2002" box="[1264,1365,906,933]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="FB8E49523F54E54B74E8C0CDFA9C43B6" box="[1264,1365,906,933]" italics="true" pageId="7" pageNumber="8">A. indica</emphasis>
</taxonomicName>
under five different names in
<taxonomicName id="0EFAEEC33F54E54B7183C0E8FD8143DA" authorityName="Rosenhauer" authorityYear="1856" box="[411,584,942,968]" class="Insecta" family="Chrysomelidae" genus="Rhaphidopalpa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="FB8E49523F54E54B7183C0E8FD8143DA" box="[411,584,942,968]" italics="true" pageId="7" pageNumber="8">Rhaphidopalpa</emphasis>
</taxonomicName>
(
<bibRefCitation id="AD6BE8B13F54E54B724EC0E8FC7F43DA" author="Barroga, G. F. &amp; Mohamedsaid, M. S." box="[598,950,942,968]" pageId="7" pageNumber="8" pagination="15 - 194" refId="ref39042" refString="Barroga, G. F. &amp; Mohamedsaid, M. S. (2002) Revision of the genus Aulacophora Chevrolat (Coleoptera: Chrysomelidae: Galerucinae) in Sundaland. Serangga, 7 (1 - 2), 15 - 194." type="journal article" year="2002">Barroga &amp; Mohamedsaid 2002</bibRefCitation>
;
<bibRefCitation id="AD6BE8B13F54E54B73D9C0E9FB6D43DB" author="Lee, C. - F. &amp; Beenen, R." box="[961,1188,942,969]" pageId="7" pageNumber="8" pagination="151 - 190" refId="ref41061" refString="Lee, C. - F. &amp; Beenen, R. (2015) Revision of the genus Aulacophora from Taiwan (Coleoptera: Chrysomelidae: Galerucinae). Zootaxa, 3949 (2), 151 - 190. https: // doi. org / 10.11646 / zootaxa. 3949.2.1" type="journal article" year="2015">Lee &amp; Beenen 2015</bibRefCitation>
), and at the same time made this species (under a sixth name,
<taxonomicName id="0EFAEEC33F54E54B7246C095FC4C43FE" authority="(Olivier, 1808))" authorityName="sensu Maulik" authorityYear="1929" baseAuthorityName="Olivier" baseAuthorityYear="1808" box="[606,901,978,1005]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="FB8E49523F54E54B7246C095FD0043FE" box="[606,713,978,1005]" italics="true" pageId="7" pageNumber="8">A. similis</emphasis>
(
<bibRefCitation id="AD6BE8B13F54E54B72C2C094FCBA43FE" author="Olivier, A. G." box="[730,883,978,1004]" pageId="7" pageNumber="8" pagination="613 - 1104" refId="ref41936" refString="Olivier, A. G. (1808) Entomologie, ou histoire naturelle des insectes, avec leur caracteres generiques et specifiques, leur description, leur synonymie, et leur figure enluminee. Coleopteres. Tome sixieme. Desray, Paris, pp. 613 - 1104, 46 pls." type="book chapter" year="1808">Olivier, 1808</bibRefCitation>
))
</taxonomicName>
the
<typeStatus id="16412BE23F54E54B73A2C095FC2343FF" box="[954,1002,979,1005]" pageId="7" pageNumber="8">type</typeStatus>
of his new genus,
<taxonomicName id="0EFAEEC33F54E54B74D0C094FF1B4402" authority="Weise, 1892" authorityName="Weise" authorityYear="1892" genus="Orthaulaca" kingdom="Animalia" pageId="7" pageNumber="8" rank="genus">
<emphasis id="FB8E49523F54E54B74D0C094FA8043FE" box="[1224,1353,978,1004]" italics="true" pageId="7" pageNumber="8">Orthaulaca</emphasis>
<bibRefCitation id="AD6BE8B13F54E54B754AC094FF1B4402" author="Weise, J." pageId="7" pageNumber="8" pagination="385 - 402" refId="ref42923" refString="Weise, J. (1892) Chrysomeliden und Coccinelliden von der Insel Nias, nebst Bemerkungen uber andere, meistens sudoestasiatische Arten. Deutsche Entomologische Zeitschrift, 1892 (2), 385 - 402. https: // doi. org / 10.1002 / mmnd. 48018920128" type="journal article" year="1892">Weise, 1892</bibRefCitation>
</taxonomicName>
. Using the current nomenclature for all the named species he listed, Weises
<taxonomicName id="0EFAEEC33F54E54B7459C0B0FB274402" authorityName="Rosenhauer" authorityYear="1856" box="[1089,1262,1014,1040]" class="Insecta" family="Chrysomelidae" genus="Rhaphidopalpa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="FB8E49523F54E54B7459C0B0FB274402" box="[1089,1262,1014,1040]" italics="true" pageId="7" pageNumber="8">Rhaphidopalpa</emphasis>
</taxonomicName>
only definitely included
<taxonomicName id="0EFAEEC33F54E54B70E7C75DFE584426" baseAuthorityName="Lucas" baseAuthorityYear="1849" box="[255,401,1050,1077]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="foveicollis">
<emphasis id="FB8E49523F54E54B70E7C75DFE584426" box="[255,401,1050,1077]" italics="true" pageId="7" pageNumber="8">A. foveicollis</emphasis>
</taxonomicName>
and
<taxonomicName id="0EFAEEC33F54E54B71DFC75DFDE54426" baseAuthorityName="Barroga &amp; Mohamedsaid" baseAuthorityYear="2002" box="[455,556,1050,1077]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="FB8E49523F54E54B71DFC75DFDE54426" box="[455,556,1050,1077]" italics="true" pageId="7" pageNumber="8">A. indica</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="C94595403F54E54B70DFC779FDBB461B" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="8">
In a morphological analysis of the nearby Sunda fauna, Barroga &amp; Mohamedsaid (2006) identified a clade which included
<taxonomicName id="0EFAEEC33F54E54B7153C725FE78446E" baseAuthorityName="Barroga &amp; Mohamedsaid" baseAuthorityYear="2002" box="[331,433,1122,1149]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="FB8E49523F54E54B7153C725FE78446E" box="[331,433,1122,1149]" italics="true" pageId="7" pageNumber="8">A. indica</emphasis>
</taxonomicName>
and
<taxonomicName id="0EFAEEC33F54E54B71F0C725FD96446E" authorityName="Baly" authorityYear="1879" box="[488,607,1123,1149]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="FB8E49523F54E54B71F0C725FD96446E" box="[488,607,1123,1149]" italics="true" pageId="7" pageNumber="8">A. cornuta</emphasis>
</taxonomicName>
, together with
<taxonomicName id="0EFAEEC33F54E54B7312C725FC6B446E" baseAuthorityName="Olivier" baseAuthorityYear="1808" box="[778,930,1122,1149]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="bipunctata">
<emphasis id="FB8E49523F54E54B7312C725FC6B446E" box="[778,930,1122,1149]" italics="true" pageId="7" pageNumber="8">A. bipunctata</emphasis>
</taxonomicName>
,
<taxonomicName id="0EFAEEC33F54E54B73A8C725FBB5446E" box="[944,1148,1122,1149]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="flavomarginata">
<emphasis id="FB8E49523F54E54B73A8C725FBB5446E" box="[944,1148,1122,1149]" italics="true" pageId="7" pageNumber="8">A. flavomarginata</emphasis>
</taxonomicName>
,
<taxonomicName id="0EFAEEC33F54E54B7492C725FAA5446E" authority="Baly, 1886" authorityName="Baly" authorityYear="1886" box="[1162,1388,1122,1149]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="lewisi">
<emphasis id="FB8E49523F54E54B7492C725FB22446E" box="[1162,1259,1122,1149]" italics="true" pageId="7" pageNumber="8">A. lewisi</emphasis>
<bibRefCitation id="AD6BE8B13F54E54B74EBC724FAA5446E" author="Baly, J. S." box="[1267,1388,1122,1148]" pageId="7" pageNumber="8" pagination="1 - 27" refId="ref38827" refString="Baly, J. S. (1886) Descriptions of a new genus and of some sp. nov. of Galerucinae, also diagnostic notes on some of the older described species of Aulacophora. Journal of the Linnean Society, Zoology, 20, 1 - 27. https: // doi. org / 10.1111 / j. 1096 - 3642.1886. tb 01432. x" type="journal article" year="1886">Baly, 1886</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="0EFAEEC33F54E54B708FC7C1FE5344B2" authority="Baly, 1886" authorityName="Baly" authorityYear="1886" box="[151,410,1158,1185]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="mouhoti">
<emphasis id="FB8E49523F54E54B708FC7C1FEDD44B2" box="[151,276,1158,1185]" italics="true" pageId="7" pageNumber="8">A. mouhoti</emphasis>
<bibRefCitation id="AD6BE8B13F54E54B7104C7C0FE5344B2" author="Baly, J. S." box="[284,410,1158,1184]" pageId="7" pageNumber="8" pagination="1 - 27" refId="ref38827" refString="Baly, J. S. (1886) Descriptions of a new genus and of some sp. nov. of Galerucinae, also diagnostic notes on some of the older described species of Aulacophora. Journal of the Linnean Society, Zoology, 20, 1 - 27. https: // doi. org / 10.1111 / j. 1096 - 3642.1886. tb 01432. x" type="journal article" year="1886">Baly, 1886</bibRefCitation>
</taxonomicName>
, defined by the thickening of male tergite VIII.
<taxonomicName id="0EFAEEC33F54E54B73D8C7C0FB7E44B2" authorityName="Baly" authorityYear="1886" box="[960,1207,1158,1184]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="mouhoti">
<emphasis id="FB8E49523F54E54B73D8C7C0FB7E44B2" box="[960,1207,1158,1184]" italics="true" pageId="7" pageNumber="8">Aulacophora mouhoti</emphasis>
</taxonomicName>
, described from the Malay Peninsula and
<collectingCountry id="B1EDD5D03F54E54B7194C7ECFE2744D6" box="[396,494,1194,1220]" name="Vietnam" pageId="7" pageNumber="8">Vietnam</collectingCountry>
, is a poorly understood pale species defined by a variable female secondary sexual character and apparently only reliably known from females (
<bibRefCitation id="AD6BE8B13F54E54B7324C788FC7D44FA" author="Baly, J. S." box="[828,948,1230,1256]" pageId="7" pageNumber="8" pagination="1 - 27" refId="ref38827" refString="Baly, J. S. (1886) Descriptions of a new genus and of some sp. nov. of Galerucinae, also diagnostic notes on some of the older described species of Aulacophora. Journal of the Linnean Society, Zoology, 20, 1 - 27. https: // doi. org / 10.1111 / j. 1096 - 3642.1886. tb 01432. x" type="journal article" year="1886">Baly 1886</bibRefCitation>
;
<bibRefCitation id="AD6BE8B13F54E54B73D8C789FAD744FA" author="Barroga, G. F. &amp; Mohamedsaid, M. S." box="[960,1310,1230,1257]" pageId="7" pageNumber="8" pagination="15 - 194" refId="ref39042" refString="Barroga, G. F. &amp; Mohamedsaid, M. S. (2002) Revision of the genus Aulacophora Chevrolat (Coleoptera: Chrysomelidae: Galerucinae) in Sundaland. Serangga, 7 (1 - 2), 15 - 194." type="journal article" year="2002">Barroga &amp; Mohamedsaid 2002</bibRefCitation>
, 2006). We consider it unlikely to be a distinct species and as it is outside the region of interest it is not discussed further.
<taxonomicName id="0EFAEEC33F54E54B756CC7B5FE9A4522" authorityName="Baly" authorityYear="1886" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="lewisi">
<emphasis id="FB8E49523F54E54B756CC7B5FE9A4522" italics="true" pageId="7" pageNumber="8">Aulacophora lewisi</emphasis>
</taxonomicName>
has black elytra, simple male antennae and a short female pygidium (
<bibRefCitation id="AD6BE8B13F54E54B747DC650FA824523" author="Lee, C. - F. &amp; Beenen, R." box="[1125,1355,1302,1329]" pageId="7" pageNumber="8" pagination="151 - 190" refId="ref41061" refString="Lee, C. - F. &amp; Beenen, R. (2015) Revision of the genus Aulacophora from Taiwan (Coleoptera: Chrysomelidae: Galerucinae). Zootaxa, 3949 (2), 151 - 190. https: // doi. org / 10.11646 / zootaxa. 3949.2.1" type="journal article" year="2015">Lee &amp; Beenen 2015</bibRefCitation>
) and is therefore excluded from our definition of the
<taxonomicName id="0EFAEEC33F54E54B7295C67DFD394546" baseAuthorityName="Barroga &amp; Mohamedsaid" baseAuthorityYear="2002" box="[653,752,1338,1365]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="FB8E49523F54E54B7295C67DFD394546" box="[653,752,1338,1365]" italics="true" pageId="7" pageNumber="8">A. indica</emphasis>
</taxonomicName>
species-complex, although there is molecular evidence for its placement in this group (
<figureCitation id="51C189C53F54E54B71B2C618FE22456A" box="[426,491,1374,1400]" captionStart="FIGURE 1" captionStartId="4.[151,250,1814,1838]" captionTargetBox="[226,1352,286,1781]" captionTargetId="figure-81@4.[223,1364,273,1791]" captionTargetPageId="4" captionText="FIGURE 1. Maximum likelihood tree of CO1 sequences of Aulacophora species, based on 100 bootstrap pseudoreplicates. All nodes have a value of one. All sequences labelled LHBP or NAQS newly added in this study." figureDoi="http://doi.org/10.5281/zenodo.4545450" httpUri="https://zenodo.org/record/4545450/files/figure.png" pageId="7" pageNumber="8">Fig. 1</figureCitation>
).
<taxonomicName id="0EFAEEC33F54E54B721AC618FC8F456A" box="[514,838,1374,1400]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="flavomarginata">
<emphasis id="FB8E49523F54E54B721AC618FC8F456A" box="[514,838,1374,1400]" italics="true" pageId="7" pageNumber="8">Aulacophora flavomarginata</emphasis>
</taxonomicName>
has mostly black elytra, modified male antennomeres 35 and the female pygidium is not produced (
<bibRefCitation id="AD6BE8B13F54E54B72B0C6C4FBC4458E" author="Barroga, G. F. &amp; Mohamedsaid, M. S." box="[680,1037,1410,1436]" pageId="7" pageNumber="8" pagination="15 - 194" refId="ref39042" refString="Barroga, G. F. &amp; Mohamedsaid, M. S. (2002) Revision of the genus Aulacophora Chevrolat (Coleoptera: Chrysomelidae: Galerucinae) in Sundaland. Serangga, 7 (1 - 2), 15 - 194." type="journal article" year="2002">Barroga &amp; Mohamedsaid 2002</bibRefCitation>
); this species is also excluded from our definition of the
<taxonomicName id="0EFAEEC33F54E54B719BC6E1FE2045D2" baseAuthorityName="Barroga &amp; Mohamedsaid" baseAuthorityYear="2002" box="[387,489,1446,1473]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="FB8E49523F54E54B719BC6E1FE2045D2" box="[387,489,1446,1473]" italics="true" pageId="7" pageNumber="8">A. indica</emphasis>
</taxonomicName>
species-complex. The thickened and modified male tergite VIII does not define the
<taxonomicName id="0EFAEEC33F54E54B708FC68CFF1445F6" baseAuthorityName="Barroga &amp; Mohamedsaid" baseAuthorityYear="2002" box="[151,221,1482,1508]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="FB8E49523F54E54B708FC68CFF1445F6" box="[151,221,1482,1508]" italics="true" pageId="7" pageNumber="8">indica</emphasis>
</taxonomicName>
species-complex, but may have been a significant precursor for development of the other male and female abdominal characters shared by the group.
</paragraph>
<paragraph id="C94595403F54E54B70DFC554FDE8475E" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="8">
<taxonomicName id="0EFAEEC33F54E54B70DFC554FDB9463E" authority="Baly, 1886" authorityName="Baly" authorityYear="1886" box="[199,624,1554,1580]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="nigroscutata">
<emphasis id="FB8E49523F54E54B70DFC554FE24463E" box="[199,493,1554,1580]" italics="true" pageId="7" pageNumber="8">Aulacophora nigroscutata</emphasis>
<bibRefCitation id="AD6BE8B13F54E54B71ECC554FDB9463E" author="Baly, J. S." box="[500,624,1554,1580]" pageId="7" pageNumber="8" pagination="1 - 27" refId="ref38827" refString="Baly, J. S. (1886) Descriptions of a new genus and of some sp. nov. of Galerucinae, also diagnostic notes on some of the older described species of Aulacophora. Journal of the Linnean Society, Zoology, 20, 1 - 27. https: // doi. org / 10.1111 / j. 1096 - 3642.1886. tb 01432. x" type="journal article" year="1886">Baly, 1886</bibRefCitation>
</taxonomicName>
, described from
<collectingRegion id="0B3E5BA23F54E54B7332C554FC00463E" box="[810,969,1554,1580]" country="Indonesia" name="Maluku Utara" pageId="7" pageNumber="8">North Maluku</collectingRegion>
(eastern
<collectingCountry id="B1EDD5D03F54E54B7437C554FB69463F" box="[1071,1184,1554,1581]" name="Indonesia" pageId="7" pageNumber="8">Indonesia</collectingCountry>
), is dorsally pale, with an excavate male abdomen, but is excluded from the PPB because the female abdomen is structurally quite different (
<bibRefCitation id="AD6BE8B13F54E54B70D2C51CFE8C4666" author="Baly, J. S." box="[202,325,1626,1652]" pageId="7" pageNumber="8" pagination="1 - 27" refId="ref38827" refString="Baly, J. S. (1886) Descriptions of a new genus and of some sp. nov. of Galerucinae, also diagnostic notes on some of the older described species of Aulacophora. Journal of the Linnean Society, Zoology, 20, 1 - 27. https: // doi. org / 10.1111 / j. 1096 - 3642.1886. tb 01432. x" type="journal article" year="1886">Baly 1886</bibRefCitation>
,
<bibRefCitation id="AD6BE8B13F54E54B714AC51CFE474666" author="Baly, J. S." box="[338,398,1626,1652]" pageId="7" pageNumber="8" pagination="297 - 306" refId="ref38948" refString="Baly, J. S. (1889) Notes on Aulacophora and allied genera. Transactions of the entomological Society, London, 1889 (2), 297 - 306. https: // doi. org / 10.1111 / j. 1365 - 2311.1889. tb 02322. x" type="journal article" year="1889">1889</bibRefCitation>
). Two other species of
<taxonomicName id="0EFAEEC33F54E54B7283C51CFCE44666" authorityName="Chevrolat" authorityYear="1836" box="[667,813,1626,1652]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="FB8E49523F54E54B7283C51CFCE44666" box="[667,813,1626,1652]" italics="true" pageId="7" pageNumber="8">Aulacophora</emphasis>
</taxonomicName>
in the region, or nearby,
<taxonomicName id="0EFAEEC33F54E54B744BC51DFAA34666" authority="(Weber 1801)" baseAuthorityName="Weber" baseAuthorityYear="1801" box="[1107,1386,1626,1653]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="coffeae">
<emphasis id="FB8E49523F54E54B744BC51DFB0C4666" box="[1107,1221,1626,1653]" italics="true" pageId="7" pageNumber="8">A. coffeae</emphasis>
(Weber 1801)
</taxonomicName>
and
<taxonomicName id="0EFAEEC33F54E54B708FC539FE66468A" authority="(Weber 1801)" baseAuthorityName="Weber" baseAuthorityYear="1801" box="[151,431,1662,1689]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="FB8E49523F54E54B708FC539FEC0468A" box="[151,265,1662,1689]" italics="true" pageId="7" pageNumber="8">A. bicolor</emphasis>
(Weber 1801)
</taxonomicName>
, may be entirely pale but they have male first antennomere simple, male last abdominal ventrite simple and female without elongated or swollen pygidium. However, these two superficially similar species are included in the key so that they can be separated from the
<taxonomicName id="0EFAEEC33F54E54B739DC581FC2246F2" baseAuthorityName="Barroga &amp; Mohamedsaid" baseAuthorityYear="2002" box="[901,1003,1734,1761]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="FB8E49523F54E54B739DC581FC2246F2" box="[901,1003,1734,1761]" italics="true" pageId="7" pageNumber="8">A. indica</emphasis>
</taxonomicName>
species-complex (PPB). Further away from the region of interest, there is an entirely pale species in
<collectingRegion id="0B3E5BA23F54E54B7351C5ACFC784716" box="[841,945,1770,1796]" country="Indonesia" name="Sulawesi Utara" pageId="7" pageNumber="8">Sulawesi</collectingRegion>
,
<taxonomicName id="0EFAEEC33F54E54B73A4C5ADFB1C4716" authority="Jacoby, 1883" authorityName="Jacoby" authorityYear="1883" box="[956,1237,1770,1797]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="FB8E49523F54E54B73A4C5ADFBF34716" box="[956,1082,1770,1797]" italics="true" pageId="7" pageNumber="8">A. unicolor</emphasis>
Jacoby, 1883
</taxonomicName>
, but this lacks the male characters of the PPB complex and has distinct creamy-white antennae and is therefore also excluded from consideration (
<bibRefCitation id="AD6BE8B13F54E54B7124C474FDDC475E" author="Perkins, P. &amp; Naskrecki, P. &amp; Farrell, B." box="[316,533,1842,1869]" pageId="7" pageNumber="8" refId="ref41989" refString="Perkins, P., Naskrecki, P. &amp; Farrell, B. (2019) MCZ type database @ Harvard Entomology. Available from: http: // 140.247.96.247 / mcz / Species _ record. php? id = 15503 (accessed 1 January 2019)" type="url" year="2019">
Perkins
<emphasis id="FB8E49523F54E54B7181C475FE02475E" box="[409,459,1842,1868]" italics="true" pageId="7" pageNumber="8">et al</emphasis>
. 2019
</bibRefCitation>
).
</paragraph>
<paragraph id="C94595403F54E54470DFC410FF2940EB" blockId="7.[151,1437,150,2013]" lastBlockId="8.[151,1436,151,249]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
The circum-Mediterranean curcurbit pest
<taxonomicName id="0EFAEEC33F54E54B72BFC410FB974762" authority="(Lucas, 1849)" baseAuthorityName="Lucas" baseAuthorityYear="1849" box="[679,1118,1878,1904]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="foveicollis">
<emphasis id="FB8E49523F54E54B72BFC410FC7F4762" box="[679,950,1878,1904]" italics="true" pageId="7" pageNumber="8">Aulacophora foveicollis</emphasis>
(
<bibRefCitation id="AD6BE8B13F54E54B73DFC410FB9F4762" author="Lucas, P. H." box="[967,1110,1878,1904]" pageId="7" pageNumber="8" refId="ref41245" refString="Lucas, P. H. (1849) Cinquieme classe, Insectes. Premier ordre. Les Coleopteres. Pp. 449 - 590, in: Exploration scientifique de l'Algerie pendant les annees 1840, 1841, 1842, publiee par ordre du gouvernement et avec le concours d'une Commission Academique. Sciences physiques zoologie. Vol. II. Histoire naturelle des animaux articules. Imprimerie Nationale, Paris, 590 pp." type="book" year="1849">Lucas, 1849</bibRefCitation>
)
</taxonomicName>
clearly belongs in the PPB complex and is particularly similar to
<taxonomicName id="0EFAEEC33F54E54B7226C43DFD2E4786" authorityName="Mer" baseAuthorityName="Fabricius" baseAuthorityYear="1781" box="[574,743,1914,1941]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis id="FB8E49523F54E54B7226C43DFD2E4786" box="[574,743,1914,1941]" italics="true" pageId="7" pageNumber="8">A. abdominalis</emphasis>
</taxonomicName>
and
<taxonomicName id="0EFAEEC33F54E54B7305C43DFC4B4786" baseAuthorityName="Barroga &amp; Mohamedsaid" baseAuthorityYear="2002" box="[797,898,1914,1941]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="FB8E49523F54E54B7305C43DFC4B4786" box="[797,898,1914,1941]" italics="true" pageId="7" pageNumber="8">A. indica</emphasis>
</taxonomicName>
.
<taxonomicName id="0EFAEEC33F54E54B7397C43CFB524786" baseAuthorityName="Lucas" baseAuthorityYear="1849" box="[911,1179,1914,1940]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="foveicollis">
<emphasis id="FB8E49523F54E54B7397C43CFB524786" box="[911,1179,1914,1940]" italics="true" pageId="7" pageNumber="8">Aulacophora foveicollis</emphasis>
</taxonomicName>
differs from these species by the following combination of characters: entirely pale, male pronotum with deeply arcuate central depression divided by a median ridge, penis similar to
<taxonomicName id="0EFAEEC33F54E54B72A0C485FCAB47CE" authorityName="Mer" baseAuthorityName="Fabricius" baseAuthorityYear="1781" box="[696,866,1986,2013]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis id="FB8E49523F54E54B72A0C485FCAB47CE" box="[696,866,1986,2013]" italics="true" pageId="7" pageNumber="8">A. abdominalis</emphasis>
</taxonomicName>
but with strongly undulated profile, male tergite 8 elongate with short paired apical projections separated by semicircular concavity, female pygidium produced and sometimes bilobed at apex, apex of female sternite 7 semicircularly excavate (
<bibRefCitation id="AD6BE8B13F5BE544741FC3FDFB5540C7" author="Maulik, S." box="[1031,1180,187,213]" pageId="8" pageNumber="9" refId="ref41439" refString="Maulik, S. (1936) The Fauna of British India including Ceylon and Burma. Coleoptera. Chrysomelidae. (Galerucinae). Taylor &amp; Francis, London, xiv + 648 pp., 1 pl." type="book" year="1936">Maulik 1936</bibRefCitation>
;
<bibRefCitation id="AD6BE8B13F5BE54474B1C3FDFAEA40C7" author="Berti, N." box="[1193,1315,187,213]" pageId="8" pageNumber="9" pagination="109 - 126" refId="ref39202" refString="Berti, N. (1990) Contribution a l'etude des Galerucinae afrotropicaux. IX. - Le genre Aulacophora Chevrolat, description d'un genre nouveau, Chosnia Gen. Nov. (Coleoptera, Chrysomelidae). Journal of African Zoology, 104, 109 - 126." type="journal article" year="1990">Berti 1990</bibRefCitation>
;
<bibRefCitation id="AD6BE8B13F5BE5447537C3FDFF1D40EB" author="Borowiec, L." pageId="8" pageNumber="9" refId="ref39478" refString="Borowiec, L. (2013) Chrysomelidae. The leaf beetles of Europe and the Mediterranean subregion (Checklist and iconography). Available from: http: // www. cassidae. uni. wroc. pl / European % 20 Chrysomelidae / (accessed 1 October 2018)" type="url" year="2013">Borowiec 2013</bibRefCitation>
).
</paragraph>
<paragraph id="C94595403F5BE544708FC261FD914152" blockId="8.[151,1437,294,2013]" box="[151,600,294,321]" pageId="8" pageNumber="9">
<emphasis id="FB8E49523F5BE544708FC261FD914152" bold="true" box="[151,600,294,321]" pageId="8" pageNumber="9">Excluded species names (Appendix 1)</emphasis>
</paragraph>
<paragraph id="C94595403F5BE54470DFC20DFB56420B" blockId="8.[151,1437,294,2013]" pageId="8" pageNumber="9">
There is clearly a problem of identity with the four names coined by
<bibRefCitation id="AD6BE8B13F5BE54473DBC20DFB4F4177" author="Boisduval &amp; J. A" box="[963,1158,330,357]" pageId="8" pageNumber="9" refId="ref39422" refString="Boisduval, [J. A.] (1835) Faune entomologique de l'Oceanie, comprenant les Coleopteres, les Hemipteres, les Nevropteres, les Hymenopteres et les Dipteres. Librairie Encyclopedique du Roret, Paris, 705 pp. https: // doi. org / 10.5962 / bhl. title. 65987" type="book" year="1835">Boisduval (1835)</bibRefCitation>
for pale species of
<taxonomicName id="0EFAEEC33F5BE5447546C20DFF20419B" authorityName="Latreille" authorityYear="1802" class="Insecta" family="Chrysomelidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="subFamily" subFamily="Galerucinae">Galerucinae</taxonomicName>
collected in the region and none have been identified with any certainty by subsequent authors (for example
<bibRefCitation id="AD6BE8B13F5BE544708FC2D5FEE841BF" author="Allard, E." box="[151,289,403,429]" pageId="8" pageNumber="9" pagination="305 - 332" refId="ref38527" refString="Allard, E. (1888) Synopsis des Galerucines a corselet sillonne transversalement. Premiere partie. Annales de la Societe entomologique de France, 8 (6), 305 - 332." type="journal article" year="1888">Allard 1888</bibRefCitation>
;
<bibRefCitation id="AD6BE8B13F5BE5447134C2D5FE7B41BF" author="Weise, J." box="[300,434,403,429]" pageId="8" pageNumber="9" pagination="1 - 225" refId="ref43036" refString="Weise, J. (1924) Chrysomelidae: 13 Galerucinae. Coleopterorum Catalogus, 78, 1 - 225." type="journal article" year="1924">Weise 1924</bibRefCitation>
;
<bibRefCitation id="AD6BE8B13F5BE54471A6C2D5FDE341BF" author="Lea, A. M." box="[446,554,403,429]" pageId="8" pageNumber="9" pagination="45 - 52" refId="ref41026" refString="Lea, A. M. (1924) The Australian species of Aulacophora (Coleoptera, Chrysomelidae). Memoirs of the Queensland Museum, 8 (1), 45 - 52." type="journal article" year="1924">Lea 1924</bibRefCitation>
,
<bibRefCitation id="AD6BE8B13F5BE544722DC2D5FD0041BF" author="Wilcox, J. A." box="[565,713,403,429]" pageId="8" pageNumber="9" pagination="221 - 431" refId="ref43057" refString="Wilcox, J. A. (1972) Chrysomelidae: Galerucinae: Luperini: Aulacophorina, Diabroticina. Coleopterorum Catalogus Supplementa, 78 (2), 221 - 431." type="journal article" year="1972">Wilcox 1972</bibRefCitation>
). We have been unable to find
<typeStatus id="16412BE23F5BE544743AC2D5FB9B41BF" box="[1058,1106,403,429]" pageId="8" pageNumber="9">type</typeStatus>
material. The best described,
<taxonomicName id="0EFAEEC33F5BE544708FC2F1FDD041C3" authority="Boisduval, 1835" authorityName="Boisduval" authorityYear="1835" box="[151,537,438,465]" class="Insecta" family="Chrysomelidae" genus="Galleruca" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="relicta">
<emphasis id="FB8E49523F5BE544708FC2F1FE9141C3" box="[151,344,439,465]" italics="true" pageId="8" pageNumber="9">Galleruca relicta</emphasis>
<bibRefCitation id="AD6BE8B13F5BE5447147C2F1FDD041C3" author="Boisduval &amp; J. A" box="[351,537,438,465]" pageId="8" pageNumber="9" refId="ref39422" refString="Boisduval, [J. A.] (1835) Faune entomologique de l'Oceanie, comprenant les Coleopteres, les Hemipteres, les Nevropteres, les Hymenopteres et les Dipteres. Librairie Encyclopedique du Roret, Paris, 705 pp. https: // doi. org / 10.5962 / bhl. title. 65987" type="book" year="1835">Boisduval, 1835</bibRefCitation>
</taxonomicName>
, is discussed below, under
<taxonomicName id="0EFAEEC33F5BE5447350C2F1FC6641C3" authorityName="Boisduval" authorityYear="1835" box="[840,943,439,465]" class="Insecta" family="Chrysomelidae" genus="Galleruca" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="relicta">
<emphasis id="FB8E49523F5BE5447350C2F1FC6641C3" box="[840,943,439,465]" italics="true" pageId="8" pageNumber="9">A. relicta</emphasis>
</taxonomicName>
. The other three Boisduval names represent poorly described and unrecognisable species from either New
<collectingCountry id="B1EDD5D03F5BE5447341C29DFC6341E7" box="[857,938,475,501]" name="Guinea-Bissau" pageId="8" pageNumber="9">Guinea</collectingCountry>
(
<taxonomicName id="0EFAEEC33F5BE54473A3C29DFA9B41E7" authority="Boisduval, 1835" authorityName="Boisduval" authorityYear="1835" box="[955,1362,474,501]" class="Insecta" family="Chrysomelidae" genus="Galleruca" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="flaveola">
<emphasis id="FB8E49523F5BE54473A3C29DFB4741E7" box="[955,1166,475,501]" italics="true" pageId="8" pageNumber="9">Galleruca flaveola</emphasis>
<bibRefCitation id="AD6BE8B13F5BE544748FC29DFA9B41E7" author="Boisduval &amp; J. A" box="[1175,1362,474,501]" pageId="8" pageNumber="9" refId="ref39422" refString="Boisduval, [J. A.] (1835) Faune entomologique de l'Oceanie, comprenant les Coleopteres, les Hemipteres, les Nevropteres, les Hymenopteres et les Dipteres. Librairie Encyclopedique du Roret, Paris, 705 pp. https: // doi. org / 10.5962 / bhl. title. 65987" type="book" year="1835">Boisduval, 1835</bibRefCitation>
</taxonomicName>
),
<collectingCountry id="B1EDD5D03F5BE544757EC29DFF18420B" name="Australia" pageId="8" pageNumber="9">Australia</collectingCountry>
(
<taxonomicName id="0EFAEEC33F5BE54470F9C2B9FDEC420B" authority="Boisduval, 1835" authorityName="Boisduval" authorityYear="1835" box="[225,549,510,537]" class="Insecta" family="Chrysomelidae" genus="Galleruca" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="FB8E49523F5BE54470F9C2B9FEAC420A" box="[225,357,511,537]" italics="true" pageId="8" pageNumber="9">G. punctata</emphasis>
<bibRefCitation id="AD6BE8B13F5BE5447174C2B9FDEC420B" author="Boisduval &amp; J. A" box="[364,549,510,537]" pageId="8" pageNumber="9" refId="ref39422" refString="Boisduval, [J. A.] (1835) Faune entomologique de l'Oceanie, comprenant les Coleopteres, les Hemipteres, les Nevropteres, les Hymenopteres et les Dipteres. Librairie Encyclopedique du Roret, Paris, 705 pp. https: // doi. org / 10.5962 / bhl. title. 65987" type="book" year="1835">Boisduval, 1835</bibRefCitation>
</taxonomicName>
) or the “Pacific Ocean” (
<taxonomicName id="0EFAEEC33F5BE5447359C2B9FB5A420B" authority="Boisduval, 1835" authorityName="Boisduval" authorityYear="1835" box="[833,1171,510,537]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="scutellata">
<emphasis id="FB8E49523F5BE5447359C2B9FC19420B" box="[833,976,511,537]" italics="true" pageId="8" pageNumber="9">G. scutellata</emphasis>
<bibRefCitation id="AD6BE8B13F5BE54473CFC2B9FB5A420B" author="Boisduval &amp; J. A" box="[983,1171,510,537]" pageId="8" pageNumber="9" refId="ref39422" refString="Boisduval, [J. A.] (1835) Faune entomologique de l'Oceanie, comprenant les Coleopteres, les Hemipteres, les Nevropteres, les Hymenopteres et les Dipteres. Librairie Encyclopedique du Roret, Paris, 705 pp. https: // doi. org / 10.5962 / bhl. title. 65987" type="book" year="1835">Boisduval, 1835</bibRefCitation>
</taxonomicName>
).
</paragraph>
<paragraph id="C94595403F5BE54470DFC165FC014328" blockId="8.[151,1437,294,2013]" pageId="8" pageNumber="9">
<taxonomicName id="0EFAEEC33F5BE54470DFC165FE55422F" authorityName="Boisduval" authorityYear="1835" box="[199,412,547,573]" class="Insecta" family="Chrysomelidae" genus="Galleruca" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="flaveola">
<emphasis id="FB8E49523F5BE54470DFC165FE55422F" box="[199,412,547,573]" italics="true" pageId="8" pageNumber="9">Galleruca flaveola</emphasis>
</taxonomicName>
has been treated as a possible senior synonym of
<taxonomicName id="0EFAEEC33F5BE54473FEC165FB35422F" authority="Weise, 1892" authorityName="Weise" authorityYear="1892" box="[998,1276,547,573]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="aruensis">
<emphasis id="FB8E49523F5BE54473FEC165FBAE422E" box="[998,1127,547,573]" italics="true" pageId="8" pageNumber="9">A. aruensis</emphasis>
<bibRefCitation id="AD6BE8B13F5BE5447468C165FB35422F" author="Weise, J." box="[1136,1276,547,573]" pageId="8" pageNumber="9" pagination="385 - 402" refId="ref42923" refString="Weise, J. (1892) Chrysomeliden und Coccinelliden von der Insel Nias, nebst Bemerkungen uber andere, meistens sudoestasiatische Arten. Deutsche Entomologische Zeitschrift, 1892 (2), 385 - 402. https: // doi. org / 10.1002 / mmnd. 48018920128" type="journal article" year="1892">Weise, 1892</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="AD6BE8B13F5BE5447515C165FA5E422F" author="Weise, J." box="[1293,1431,547,573]" pageId="8" pageNumber="9" pagination="1 - 225" refId="ref43036" refString="Weise, J. (1924) Chrysomelidae: 13 Galerucinae. Coleopterorum Catalogus, 78, 1 - 225." type="journal article" year="1924">Weise 1924</bibRefCitation>
;
<bibRefCitation id="AD6BE8B13F5BE544708FC101FEE54273" author="Wilcox, J. A." box="[151,300,583,609]" pageId="8" pageNumber="9" pagination="221 - 431" refId="ref43057" refString="Wilcox, J. A. (1972) Chrysomelidae: Galerucinae: Luperini: Aulacophorina, Diabroticina. Coleopterorum Catalogus Supplementa, 78 (2), 221 - 431." type="journal article" year="1972">Wilcox 1972</bibRefCitation>
), itself a junior synonym of something else (see below under
<taxonomicName id="0EFAEEC33F5BE54473FFC101FB584273" authorityName="Mer" baseAuthorityName="Fabricius" baseAuthorityYear="1781" box="[999,1169,583,609]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis id="FB8E49523F5BE54473FFC101FB584273" box="[999,1169,583,609]" italics="true" pageId="8" pageNumber="9">A. abdominalis</emphasis>
</taxonomicName>
). The description of
<taxonomicName id="0EFAEEC33F5BE5447599C101FF384297" authorityName="Boisduval" authorityYear="1835" class="Insecta" family="Chrysomelidae" genus="Galleruca" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="flaveola">
<emphasis id="FB8E49523F5BE5447599C101FF384297" italics="true" pageId="8" pageNumber="9">G. flaveola</emphasis>
</taxonomicName>
is so poor (only “lutea; abdomine, pectore pedibusque nigris”:
<bibRefCitation id="AD6BE8B13F5BE54473A6C12DFB604297" author="Boisduval &amp; J. A" box="[958,1193,618,645]" pageId="8" pageNumber="9" refId="ref39422" refString="Boisduval, [J. A.] (1835) Faune entomologique de l'Oceanie, comprenant les Coleopteres, les Hemipteres, les Nevropteres, les Hymenopteres et les Dipteres. Librairie Encyclopedique du Roret, Paris, 705 pp. https: // doi. org / 10.5962 / bhl. title. 65987" type="book" year="1835">Boisduval 1835: 558</bibRefCitation>
) that is impossible to identify to genus. None of the
<taxonomicName id="0EFAEEC33F5BE54471ECC1C9FD4F42BB" authorityName="Chevrolat" authorityYear="1836" box="[500,646,655,681]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="FB8E49523F5BE54471ECC1C9FD4F42BB" box="[500,646,655,681]" italics="true" pageId="8" pageNumber="9">Aulacophora</emphasis>
</taxonomicName>
species in the region have entirely black legs, therefore, if accurately described,
<taxonomicName id="0EFAEEC33F5BE544710AC1F5FE4642DF" authorityName="Boisduval" authorityYear="1835" box="[274,399,691,717]" class="Insecta" family="Chrysomelidae" genus="Galleruca" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="flaveola">
<emphasis id="FB8E49523F5BE544710AC1F5FE4642DF" box="[274,399,691,717]" italics="true" pageId="8" pageNumber="9">G. flaveola</emphasis>
</taxonomicName>
is unlikely to be an
<taxonomicName id="0EFAEEC33F5BE5447262C1F5FCC542DF" authorityName="Chevrolat" authorityYear="1836" box="[634,780,691,717]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="FB8E49523F5BE5447262C1F5FCC542DF" box="[634,780,691,717]" italics="true" pageId="8" pageNumber="9">Aulacophora</emphasis>
</taxonomicName>
.
<taxonomicName id="0EFAEEC33F5BE5447303C1F5FC3E42DE" authorityName="Boisduval" authorityYear="1835" box="[795,1015,691,717]" class="Insecta" family="Chrysomelidae" genus="Galleruca" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="FB8E49523F5BE5447303C1F5FC3E42DE" box="[795,1015,691,717]" italics="true" pageId="8" pageNumber="9">Galleruca punctata</emphasis>
</taxonomicName>
is also not identifiable to genus. If it is an
<taxonomicName id="0EFAEEC33F5BE54470CDC191FEAE42E3" authorityName="Chevrolat" authorityYear="1836" box="[213,359,727,753]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="FB8E49523F5BE54470CDC191FEAE42E3" box="[213,359,727,753]" italics="true" pageId="8" pageNumber="9">Aulacophora</emphasis>
</taxonomicName>
it is most likely a junior synonym of either
<taxonomicName id="0EFAEEC33F5BE5447347C191FBC042E3" authorityName="Mer" baseAuthorityName="Fabricius" baseAuthorityYear="1781" box="[863,1033,727,753]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis id="FB8E49523F5BE5447347C191FBC042E3" box="[863,1033,727,753]" italics="true" pageId="8" pageNumber="9">A. abdominalis</emphasis>
</taxonomicName>
(
<bibRefCitation id="AD6BE8B13F5BE5447402C191FB7942E3" author="Wilcox, J. A." box="[1050,1200,727,753]" pageId="8" pageNumber="9" pagination="221 - 431" refId="ref43057" refString="Wilcox, J. A. (1972) Chrysomelidae: Galerucinae: Luperini: Aulacophorina, Diabroticina. Coleopterorum Catalogus Supplementa, 78 (2), 221 - 431." type="journal article" year="1972">Wilcox 1972</bibRefCitation>
;
<bibRefCitation id="AD6BE8B13F5BE54474A5C191FA9D42E3" author="Kimoto S." box="[1213,1364,727,753]" pageId="8" pageNumber="9" pagination="201 - 237" refId="ref40871" refString="Kimoto S. (1990) Checklist of Chrysomelidae of south-east Asia, south of Thailand and west of Irian Jaya of Indonesia, VI. Galerucinae 2. Kurume University Journal, 39 (2), 201 - 237." type="journal article" year="1990">Kimoto 1990</bibRefCitation>
) or
<taxonomicName id="0EFAEEC33F5BE544759CC191FF164307" authorityName="Boisduval" authorityYear="1835" class="Insecta" family="Chrysomelidae" genus="Galleruca" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="relicta">
<emphasis id="FB8E49523F5BE544759CC191FF164307" italics="true" pageId="8" pageNumber="9">A. relicta</emphasis>
</taxonomicName>
, and therefore of no taxonomic significance. Both
<taxonomicName id="0EFAEEC33F5BE5447306C1BDFC3B4307" authorityName="Boisduval" authorityYear="1835" box="[798,1010,763,789]" class="Insecta" family="Chrysomelidae" genus="Galleruca" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="flaveola">
<emphasis id="FB8E49523F5BE5447306C1BDFC3B4307" box="[798,1010,763,789]" italics="true" pageId="8" pageNumber="9">Galleruca flaveola</emphasis>
</taxonomicName>
and
<taxonomicName id="0EFAEEC33F5BE5447431C1BDFB674306" authorityName="Boisduval" authorityYear="1835" box="[1065,1198,763,789]" class="Insecta" family="Chrysomelidae" genus="Galleruca" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="FB8E49523F5BE5447431C1BDFB674306" box="[1065,1198,763,789]" italics="true" pageId="8" pageNumber="9">G. punctata</emphasis>
</taxonomicName>
should be discarded, as
<emphasis id="FB8E49523F5BE54470AEC059FE98432B" box="[182,337,799,825]" italics="true" pageId="8" pageNumber="9">nomina dubia</emphasis>
, like other unidentifiable Boisduval names (
<bibRefCitation id="AD6BE8B13F5BE5447359C059FC754328" author="Reid, C. A. M." box="[833,956,799,826]" pageId="8" pageNumber="9" pagination="1 - 119" refId="ref42202" refString="Reid, C. A. M. (2006) A taxonomic revision of the Australian Chrysomelinae, with a key to the genera (Coleoptera: Chrysomelidae). Zootaxa, 1292 (1), 1 - 119. https: // doi. org / 10.11646 / zootaxa. 1292.1.1" type="journal article" year="2006">Reid 2006</bibRefCitation>
).
</paragraph>
<paragraph id="C94595403F5BE54470DFC005FB71451F" blockId="8.[151,1437,294,2013]" pageId="8" pageNumber="9">
<taxonomicName id="0EFAEEC33F5BE54470DFC005FE67434F" box="[199,430,835,861]" class="Insecta" family="Chrysomelidae" genus="Galleruca" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="scutellata">
<emphasis id="FB8E49523F5BE54470DFC005FE67434F" box="[199,430,835,861]" italics="true" pageId="8" pageNumber="9">Galleruca scutellata</emphasis>
</taxonomicName>
has been treated as a senior synonym of the single Australian
<taxonomicName id="0EFAEEC33F5BE544746FC005FAC0434F" authorityName="Chevrolat" authorityYear="1836" box="[1143,1289,835,861]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="FB8E49523F5BE544746FC005FAC0434F" box="[1143,1289,835,861]" italics="true" pageId="8" pageNumber="9">Aulacophora</emphasis>
</taxonomicName>
species with a black scutellum,
<taxonomicName id="0EFAEEC33F5BE5447177C021FDAA4393" authority="Baly 1888" authorityName="Baly" authorityYear="1888" box="[367,611,871,897]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="wilsoni">
<emphasis id="FB8E49523F5BE5447177C021FE284393" box="[367,481,871,897]" italics="true" pageId="8" pageNumber="9">A. wilsoni</emphasis>
<bibRefCitation id="AD6BE8B13F5BE54471F2C021FDAA4393" author="Baly, J. S." box="[490,611,871,897]" pageId="8" pageNumber="9" pagination="156 - 188" refId="ref38897" refString="Baly, J. S. (1888) Descriptions of some genera and species of Galerucinae. Journal of the Linnean Society, Zoology, 20, 156 - 188. https: // doi. org / 10.1111 / j. 1096 - 3642.1888. tb 01444. x" type="journal article" year="1888">Baly 1888</bibRefCitation>
</taxonomicName>
, because of Blackburns tentative synonymy of these two names, which ignored the notable difference in
<typeStatus id="16412BE23F5BE544720AC0CDFD8B43B7" box="[530,578,907,933]" pageId="8" pageNumber="9">type</typeStatus>
localities (
<bibRefCitation id="AD6BE8B13F5BE54472D8C0CDFCB343B7" author="Blackburn, T." box="[704,890,907,933]" pageId="8" pageNumber="9" pagination="303 - 366" refId="ref39334" refString="Blackburn, T. (1890) Notes on Australian Coleoptera, with descriptions of sp. nov. Proceedings of the Linnean Society of New South Wales, Series 2, 5, 303 - 366. https: // doi. org / 10.5962 / bhl. part. 18641" type="journal article" year="1890">Blackburn 1890</bibRefCitation>
;
<bibRefCitation id="AD6BE8B13F5BE544739FC0CDFC3D43B7" author="Lea, A. M." box="[903,1012,907,933]" pageId="8" pageNumber="9" pagination="45 - 52" refId="ref41026" refString="Lea, A. M. (1924) The Australian species of Aulacophora (Coleoptera, Chrysomelidae). Memoirs of the Queensland Museum, 8 (1), 45 - 52." type="journal article" year="1924">Lea 1924</bibRefCitation>
;
<bibRefCitation id="AD6BE8B13F5BE5447419C0CDFB5F43B7" author="Wilcox, J. A." box="[1025,1174,907,933]" pageId="8" pageNumber="9" pagination="221 - 431" refId="ref43057" refString="Wilcox, J. A. (1972) Chrysomelidae: Galerucinae: Luperini: Aulacophorina, Diabroticina. Coleopterorum Catalogus Supplementa, 78 (2), 221 - 431." type="journal article" year="1972">Wilcox 1972</bibRefCitation>
;
<bibRefCitation id="AD6BE8B13F5BE54474BBC0CDFAF443B7" author="Kimoto S." box="[1187,1341,907,933]" pageId="8" pageNumber="9" pagination="201 - 237" refId="ref40871" refString="Kimoto S. (1990) Checklist of Chrysomelidae of south-east Asia, south of Thailand and west of Irian Jaya of Indonesia, VI. Galerucinae 2. Kurume University Journal, 39 (2), 201 - 237." type="journal article" year="1990">Kimoto 1990</bibRefCitation>
). Blackburn noted that Boisduvals description was “perhaps unworthy of attention” (
<bibRefCitation id="AD6BE8B13F5BE544741CC0E9FB3F43D8" author="Blackburn, T." box="[1028,1270,943,970]" pageId="8" pageNumber="9" pagination="303 - 366" refId="ref39334" refString="Blackburn, T. (1890) Notes on Australian Coleoptera, with descriptions of sp. nov. Proceedings of the Linnean Society of New South Wales, Series 2, 5, 303 - 366. https: // doi. org / 10.5962 / bhl. part. 18641" type="journal article" year="1890">Blackburn 1890: 361</bibRefCitation>
).
<taxonomicName id="0EFAEEC33F5BE5447512C0E9FF2143FF" authorityName="Baly" authorityYear="1888" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="wilsoni">
<emphasis id="FB8E49523F5BE5447512C0E9FF2143FF" italics="true" pageId="8" pageNumber="9">Aulacophora wilsoni</emphasis>
</taxonomicName>
is a scarce species of high rainfall forests in southeastern
<collectingRegion id="0B3E5BA23F5BE5447365C095FBC843FF" box="[893,1025,979,1005]" country="Australia" name="Queensland" pageId="8" pageNumber="9">Queensland</collectingRegion>
and northeastern
<collectingRegion id="0B3E5BA23F5BE54474D4C095FA5043FF" box="[1228,1433,979,1005]" country="Australia" name="New South Wales" pageId="8" pageNumber="9">New South Wales</collectingRegion>
, a species unlikely to have been encountered by collectors in the early 19
<superScript id="3E8F38083F5BE54473C1C0B0FC2F4416" attach="left" box="[985,998,1014,1028]" fontSize="6" pageId="8" pageNumber="9">th</superScript>
century, and there are no pale species of
<taxonomicName id="0EFAEEC33F5BE54470AEC75DFE814427" authorityName="Chevrolat" authorityYear="1836" box="[182,328,1051,1077]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="FB8E49523F5BE54470AEC75DFE814427" box="[182,328,1051,1077]" italics="true" pageId="8" pageNumber="9">Aulacophora</emphasis>
</taxonomicName>
with a black scutellum in the Pacific region. With this combination of colour and
<typeStatus id="16412BE23F5BE54474F0C75DFAD14427" box="[1256,1304,1051,1077]" pageId="8" pageNumber="9">type</typeStatus>
locality,
<taxonomicName id="0EFAEEC33F5BE5447599C75DFECD444B" class="Insecta" family="Chrysomelidae" genus="Galleruca" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="scutellata">
<emphasis id="FB8E49523F5BE5447599C75DFECD444B" italics="true" pageId="8" pageNumber="9">G. scutellata</emphasis>
</taxonomicName>
is more likely a senior synonym of
<taxonomicName id="0EFAEEC33F5BE54472B7C779FB364448" authority="(Perroud &amp; Montrouzier, 1864)" baseAuthorityName="Perroud &amp; Montrouzier" baseAuthorityYear="1864" box="[687,1279,1087,1114]" class="Insecta" family="Chrysomelidae" genus="Candezea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="palustris">
<emphasis id="FB8E49523F5BE54472B7C779FC43444B" box="[687,906,1087,1113]" italics="true" pageId="8" pageNumber="9">Candezea palustris</emphasis>
(
<bibRefCitation id="AD6BE8B13F5BE5447383C779FB314448" author="Perroud, B. P. &amp; Montrouzier, X." box="[923,1272,1087,1114]" pageId="8" pageNumber="9" pagination="46 - 256" refId="ref42038" refString="Perroud, B. P. &amp; Montrouzier, X. (1864) Essai sur la faune entomologique de Kanala (Nouvelle-Caledonie) et description de quelques especes nouvelles ou peu connues. Annales de la Societe Lineenne de Lyon, Nouvelle Serie, 11, 46 - 256. https: // doi. org / 10.1080 / 00378941.1864.10827340" type="journal article" year="1864">Perroud &amp; Montrouzier, 1864</bibRefCitation>
)
</taxonomicName>
, as originally suggested by
<bibRefCitation id="AD6BE8B13F5BE5447137C725FE01446F" author="Allard, E." box="[303,456,1123,1149]" pageId="8" pageNumber="9" pagination="305 - 332" refId="ref38527" refString="Allard, E. (1888) Synopsis des Galerucines a corselet sillonne transversalement. Premiere partie. Annales de la Societe entomologique de France, 8 (6), 305 - 332." type="journal article" year="1888">Allard (1888)</bibRefCitation>
and discussed by
<bibRefCitation id="AD6BE8B13F5BE544728DC725FCD5446F" author="Baly, J. S." box="[661,796,1123,1149]" pageId="8" pageNumber="9" pagination="297 - 306" refId="ref38948" refString="Baly, J. S. (1889) Notes on Aulacophora and allied genera. Transactions of the entomological Society, London, 1889 (2), 297 - 306. https: // doi. org / 10.1111 / j. 1365 - 2311.1889. tb 02322. x" type="journal article" year="1889">Baly (1889)</bibRefCitation>
and
<bibRefCitation id="AD6BE8B13F5BE544734BC725FC18446F" author="Lea, A. M." box="[851,977,1123,1149]" pageId="8" pageNumber="9" pagination="45 - 52" refId="ref41026" refString="Lea, A. M. (1924) The Australian species of Aulacophora (Coleoptera, Chrysomelidae). Memoirs of the Queensland Museum, 8 (1), 45 - 52." type="journal article" year="1924">Lea (1924)</bibRefCitation>
, the latter accepting
<taxonomicName id="0EFAEEC33F5BE54474A2C725FAE2446F" authorityName="Baly" authorityYear="1888" box="[1210,1323,1123,1149]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="wilsoni">
<emphasis id="FB8E49523F5BE54474A2C725FAE2446F" box="[1210,1323,1123,1149]" italics="true" pageId="8" pageNumber="9">A. wilsoni</emphasis>
</taxonomicName>
as a valid Australian species.
<taxonomicName id="0EFAEEC33F5BE544716CC7C1FD8544B3" baseAuthorityName="Perroud &amp; Montrouzier" baseAuthorityYear="1864" box="[372,588,1159,1185]" class="Insecta" family="Chrysomelidae" genus="Candezea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="palustris">
<emphasis id="FB8E49523F5BE544716CC7C1FD8544B3" box="[372,588,1159,1185]" italics="true" pageId="8" pageNumber="9">Candezea palustris</emphasis>
</taxonomicName>
is an abundant species in the West Pacific (
<bibRefCitation id="AD6BE8B13F5BE5447425C7C1FB1F44B3" author="Beenen, R." box="[1085,1238,1159,1185]" pageId="8" pageNumber="9" pagination="65 - 87" refId="ref39087" refString="Beenen, R. (2008) Contribution to the knowledge of Galerucinae of New Caledonia (Coleoptera: Chrysomelidae). Genus, 19, 65 - 87." type="journal article" year="2008">Beenen 2008</bibRefCitation>
), surprisingly not otherwise noted by Boisduval. The name
<taxonomicName id="0EFAEEC33F5BE544726DC7EDFC8744D7" baseAuthorityName="Perroud &amp; Montrouzier" baseAuthorityYear="1864" box="[629,846,1195,1221]" class="Insecta" family="Chrysomelidae" genus="Candezea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="palustris">
<emphasis id="FB8E49523F5BE544726DC7EDFC8744D7" box="[629,846,1195,1221]" italics="true" pageId="8" pageNumber="9">Candezea palustris</emphasis>
</taxonomicName>
is stable and designates an important pest of sweet potatoes in the region (
<bibRefCitation id="AD6BE8B13F5BE5447186C789FDB444FB" author="Kimoto, S. &amp; Ismay, J. W. &amp; Samuelson, G. A." box="[414,637,1231,1257]" pageId="8" pageNumber="9" pagination="49 - 57" refId="ref40913" refString="Kimoto, S., Ismay, J. W. &amp; Samuelson, G. A. (1984) Distribution of chrysomelid pests associated with certain agricultural plants in Papua New Guinea (Coleoptera). Esakia, 2, 49 - 57." type="journal article" year="1984">
Kimoto
<emphasis id="FB8E49523F5BE54471E6C789FDF844FB" box="[510,561,1231,1257]" italics="true" pageId="8" pageNumber="9">et al</emphasis>
. 1984
</bibRefCitation>
). In the interests of nomenclatural stability, we think it best to discard
<taxonomicName id="0EFAEEC33F5BE544708FC7B5FEB4451F" box="[151,381,1267,1293]" class="Insecta" family="Chrysomelidae" genus="Galleruca" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="scutellata">
<emphasis id="FB8E49523F5BE544708FC7B5FEB4451F" box="[151,381,1267,1293]" italics="true" pageId="8" pageNumber="9">Galleruca scutellata</emphasis>
</taxonomicName>
and also treat this name as a
<emphasis id="FB8E49523F5BE54472DDC7B5FCA3451F" box="[709,874,1267,1293]" italics="true" pageId="8" pageNumber="9">nomen dubium</emphasis>
until it is formally supressed.
</paragraph>
<paragraph id="C94595403F5BE54470DFC651FD3747CF" blockId="8.[151,1437,294,2013]" pageId="8" pageNumber="9">
The names
<taxonomicName id="0EFAEEC33F5BE5447150C651FCF04523" authority="Montrouzier, 1856" authorityName="Montrouzier" authorityYear="1856" box="[328,825,1302,1329]" class="Insecta" family="Chrysomelidae" genus="Galleruca" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="cristovallensis">
<emphasis id="FB8E49523F5BE5447150C651FDAB4523" box="[328,610,1303,1329]" italics="true" pageId="8" pageNumber="9">Galleruca cristovallensis</emphasis>
<bibRefCitation id="AD6BE8B13F5BE5447271C651FCF04523" author="Montrouzier, P." box="[617,825,1302,1329]" pageId="8" pageNumber="9" pagination="1 - 226" refId="ref41755" refString="Montrouzier, P. (1856) Essai sur la faune de l'Ile de Woodlark ou Moiou. Annales de la Societe imperiale d'agriculture, d'histoire naturelle et des arts utiles de Lyon, 7, 1 - 226." type="journal article" year="1856">Montrouzier, 1856</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="0EFAEEC33F5BE5447377C651FB124523" authority="Montrouzier, 1856" authorityName="Montrouzier" authorityYear="1856" box="[879,1243,1302,1329]" class="Insecta" family="Chrysomelidae" genus="Galleruca" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="flavescens">
<emphasis id="FB8E49523F5BE5447377C651FBCD4523" box="[879,1028,1303,1329]" italics="true" pageId="8" pageNumber="9">G. flavescens</emphasis>
<bibRefCitation id="AD6BE8B13F5BE5447413C651FB124523" author="Montrouzier, P." box="[1035,1243,1302,1329]" pageId="8" pageNumber="9" pagination="1 - 226" refId="ref41755" refString="Montrouzier, P. (1856) Essai sur la faune de l'Ile de Woodlark ou Moiou. Annales de la Societe imperiale d'agriculture, d'histoire naturelle et des arts utiles de Lyon, 7, 1 - 226." type="journal article" year="1856">Montrouzier, 1856</bibRefCitation>
</taxonomicName>
are also attached to miserly descriptions, although both are regarded as valid species (
<bibRefCitation id="AD6BE8B13F5BE5447385C67DFBFD4547" author="Wilcox, J. A." box="[925,1076,1339,1365]" pageId="8" pageNumber="9" pagination="221 - 431" refId="ref43057" refString="Wilcox, J. A. (1972) Chrysomelidae: Galerucinae: Luperini: Aulacophorina, Diabroticina. Coleopterorum Catalogus Supplementa, 78 (2), 221 - 431." type="journal article" year="1972">Wilcox 1972</bibRefCitation>
).
<typeStatus id="16412BE23F5BE5447450C67DFB494547" box="[1096,1152,1339,1365]" pageId="8" pageNumber="9">Type</typeStatus>
material of these species is probably no longer extant. The first was described from San Cristoval Island (now
<collectingRegion id="0B3E5BA23F5BE5447453C619FB56456B" box="[1099,1183,1375,1401]" country="Solomon Islands" name="Makira" pageId="8" pageNumber="9">Makira</collectingRegion>
,
<collectingCountry id="B1EDD5D03F5BE54474B1C619FAAF4568" box="[1193,1382,1375,1402]" name="Solomon Islands" pageId="8" pageNumber="9">Solomon Islands</collectingCountry>
) and has a transverse pronotal groove, parallel-sided elytra, is 4.5 “lignes” long (
<quantity id="0E0238A53F5BE54473F9C6C5FBFB458F" box="[993,1074,1411,1437]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="9" unit="mm" value="10.0">10 mm</quantity>
) and is “testacée, lisse” (smooth and brick-red coloured) (
<bibRefCitation id="AD6BE8B13F5BE54471B4C6E1FD6E45D3" author="Montrouzier, P." box="[428,679,1446,1473]" pageId="8" pageNumber="9" pagination="1 - 226" refId="ref41755" refString="Montrouzier, P. (1856) Essai sur la faune de l'Ile de Woodlark ou Moiou. Annales de la Societe imperiale d'agriculture, d'histoire naturelle et des arts utiles de Lyon, 7, 1 - 226." type="journal article" year="1856">Montrouzier 1856: 70</bibRefCitation>
). This species is too large and wrongly coloured for a PPB, so is not discussed further here.
<taxonomicName id="0EFAEEC33F5BE5447183C68DFD6A45F7" box="[411,675,1483,1509]" class="Magnoliopsida" family="Fabaceae" genus="Gallerucella" kingdom="Plantae" order="Fabales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="flavescens">
<emphasis id="FB8E49523F5BE5447183C68DFD6A45F7" box="[411,675,1483,1509]" italics="true" pageId="8" pageNumber="9">Gallerucella flavescens</emphasis>
</taxonomicName>
was described from both Woodlark Island (now eastern
<collectingCountry id="B1EDD5D03F5BE5447504C68DFF234618" name="Papua New Guinea" pageId="8" pageNumber="9">Papua New Guinea</collectingCountry>
) and San Cristoval Island (Solomons) and also has a transverse pronotal groove, but has expanded elytra, is 3 “lignes” long (
<quantity id="0E0238A53F5BE5447157C555FE6E463F" box="[335,423,1555,1581]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="8" pageNumber="9" unit="mm" value="6.8">6.8 mm</quantity>
) and is “entièrement jaune, sauf le dessous du corps qui est rembruni” (entirely yellow except the underside which is darkened) (
<bibRefCitation id="AD6BE8B13F5BE5447250C571FC8E4643" author="Montrouzier, P." box="[584,839,1590,1617]" pageId="8" pageNumber="9" pagination="1 - 226" refId="ref41755" refString="Montrouzier, P. (1856) Essai sur la faune de l'Ile de Woodlark ou Moiou. Annales de la Societe imperiale d'agriculture, d'histoire naturelle et des arts utiles de Lyon, 7, 1 - 226." type="journal article" year="1856">Montrouzier, 1856: 71</bibRefCitation>
). This vague description probably applies to a species of PPB but it is impossible to say which one, and in any case all the evidence from the material to hand is that the currently valid species names for any PPB in the New
<collectingCountry id="B1EDD5D03F5BE54472E9C539FC8B468B" box="[753,834,1663,1689]" name="Guinea-Bissau" pageId="8" pageNumber="9">Guinea</collectingCountry>
and Solomons region are senior to anything described by Montrouzier. In the interests of nomenclatural stability, we think it best to discard
<taxonomicName id="0EFAEEC33F5BE5447452C5E5FAFC46AF" authorityName="Montrouzier" authorityYear="1856" box="[1098,1333,1699,1725]" class="Insecta" family="Chrysomelidae" genus="Galleruca" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="flavescens">
<emphasis id="FB8E49523F5BE5447452C5E5FAFC46AF" box="[1098,1333,1699,1725]" italics="true" pageId="8" pageNumber="9">Galleruca flavescens</emphasis>
</taxonomicName>
and treat this name as a
<emphasis id="FB8E49523F5BE5447158C581FE2E46F3" box="[320,487,1735,1761]" italics="true" pageId="8" pageNumber="9">nomen dubium</emphasis>
. Another Montrouzier name,
<taxonomicName id="0EFAEEC33F5BE544732FC581FB2F46F3" authority="Montrouzier, 1861" authorityName="Montrouzier" authorityYear="1861" box="[823,1254,1735,1761]" class="Insecta" family="Chrysomelidae" genus="Galleruca" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="artensis">
<emphasis id="FB8E49523F5BE544732FC581FBC346F2" box="[823,1034,1735,1761]" italics="true" pageId="8" pageNumber="9">Galleruca artensis</emphasis>
<bibRefCitation id="AD6BE8B13F5BE544740AC581FB2F46F3" author="Montrouzier, X." box="[1042,1254,1735,1761]" pageId="8" pageNumber="9" pagination="265 - 306" refId="ref41795" refString="Montrouzier, X. (1861) Essai sur la faune entomologique de la Nouvelle-Caledonie (Balade) et des iles des Pins, Art, Lifu, etc. Annales de la Societe entomologique de France, 4, 265 - 306." type="journal article" year="1861">Montrouzier, 1861</bibRefCitation>
</taxonomicName>
, described from
<collectingCountry id="B1EDD5D03F5BE544708FC5ADFE8E4717" box="[151,327,1771,1797]" name="New Caledonia" pageId="8" pageNumber="9">New Caledonia</collectingCountry>
and overlooked in revisions of that fauna (
<bibRefCitation id="AD6BE8B13F5BE544732EC5ADFC064717" author="Beenen, R." box="[822,975,1771,1797]" pageId="8" pageNumber="9" pagination="65 - 87" refId="ref39087" refString="Beenen, R. (2008) Contribution to the knowledge of Galerucinae of New Caledonia (Coleoptera: Chrysomelidae). Genus, 19, 65 - 87." type="journal article" year="2008">Beenen 2008</bibRefCitation>
,
<bibRefCitation id="AD6BE8B13F5BE54473C4C5ADFBD14717" author="Beenen, R." box="[988,1048,1771,1797]" pageId="8" pageNumber="9" pagination="65 - 108" refId="ref39117" refString="Beenen, R. (2013) Contribution to the knowledge of Galerucinae of New Caledonia 2 (Coleoptera: Chrysomelidae). Genus, 24 (1), 65 - 108." type="journal article" year="2013">2013</bibRefCitation>
), is equally impossible to identify from its four lines of description. It was originally compared with
<taxonomicName id="0EFAEEC33F5BE544736AC449FB30473B" authority="Montrouzier, 1861" authorityName="Montrouzier" authorityYear="1861" box="[882,1273,1807,1833]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="argyrogaster">
<emphasis id="FB8E49523F5BE544736AC449FBEA473A" box="[882,1059,1807,1833]" italics="true" pageId="8" pageNumber="9">G. argyrogaster</emphasis>
<bibRefCitation id="AD6BE8B13F5BE5447431C449FB30473B" author="Montrouzier, X." box="[1065,1273,1807,1833]" pageId="8" pageNumber="9" pagination="265 - 306" refId="ref41795" refString="Montrouzier, X. (1861) Essai sur la faune entomologique de la Nouvelle-Caledonie (Balade) et des iles des Pins, Art, Lifu, etc. Annales de la Societe entomologique de France, 4, 265 - 306." type="journal article" year="1861">Montrouzier, 1861</bibRefCitation>
</taxonomicName>
(synonymised below with
<taxonomicName id="0EFAEEC33F5BE5447106C475FE00475F" authorityName="Mer" baseAuthorityName="Fabricius" baseAuthorityYear="1781" box="[286,457,1843,1869]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis id="FB8E49523F5BE5447106C475FE00475F" box="[286,457,1843,1869]" italics="true" pageId="8" pageNumber="9">A. abdominalis</emphasis>
</taxonomicName>
), from which it only differed by pale antennae and legs and an unusual golden elytral spot in living specimens (
<bibRefCitation id="AD6BE8B13F5BE54471A9C411FD704763" author="Montrouzier, X." box="[433,697,1879,1905]" pageId="8" pageNumber="9" pagination="265 - 306" refId="ref41795" refString="Montrouzier, X. (1861) Essai sur la faune entomologique de la Nouvelle-Caledonie (Balade) et des iles des Pins, Art, Lifu, etc. Annales de la Societe entomologique de France, 4, 265 - 306." type="journal article" year="1861">Montrouzier 1861: 394</bibRefCitation>
).
<taxonomicName id="0EFAEEC33F5BE54472D4C411FC554762" authorityName="Montrouzier" authorityYear="1861" box="[716,924,1879,1905]" class="Insecta" family="Chrysomelidae" genus="Galleruca" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="artensis">
<emphasis id="FB8E49523F5BE54472D4C411FC554762" box="[716,924,1879,1905]" italics="true" pageId="8" pageNumber="9">Galleruca artensis</emphasis>
</taxonomicName>
is probably a junior synonym of
<taxonomicName id="0EFAEEC33F5BE5447512C411FEE84787" authorityName="Mer" baseAuthorityName="Fabricius" baseAuthorityYear="1781" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis id="FB8E49523F5BE5447512C411FEE84787" italics="true" pageId="8" pageNumber="9">Aulacophora abdominalis</emphasis>
</taxonomicName>
, but it could possibly be a pale form of
<taxonomicName id="0EFAEEC33F5BE54472C0C43DFB1F4784" authority="(Perroud &amp; Montrouzier, 1864)" baseAuthorityName="Perroud &amp; Montrouzier" baseAuthorityYear="1864" box="[728,1238,1915,1942]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="deplanchei">
<emphasis id="FB8E49523F5BE54472C0C43DFCB84787" box="[728,881,1915,1941]" italics="true" pageId="8" pageNumber="9">A. deplanchei</emphasis>
(
<bibRefCitation id="AD6BE8B13F5BE5447366C43DFB064784" author="Perroud, B. P. &amp; Montrouzier, X." box="[894,1231,1915,1942]" pageId="8" pageNumber="9" pagination="46 - 256" refId="ref42038" refString="Perroud, B. P. &amp; Montrouzier, X. (1864) Essai sur la faune entomologique de Kanala (Nouvelle-Caledonie) et description de quelques especes nouvelles ou peu connues. Annales de la Societe Lineenne de Lyon, Nouvelle Serie, 11, 46 - 256. https: // doi. org / 10.1080 / 00378941.1864.10827340" type="journal article" year="1864">Perroud &amp; Montrouzier, 1864</bibRefCitation>
)
</taxonomicName>
, discussed below.
<typeStatus id="16412BE23F5BE544708FC4D9FF0647AB" box="[151,207,1951,1977]" pageId="8" pageNumber="9">Type</typeStatus>
material of
<taxonomicName id="0EFAEEC33F5BE544714EC4D9FE1B47AA" authorityName="Montrouzier" authorityYear="1861" box="[342,466,1951,1977]" class="Insecta" family="Chrysomelidae" genus="Galleruca" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="artensis">
<emphasis id="FB8E49523F5BE544714EC4D9FE1B47AA" box="[342,466,1951,1977]" italics="true" pageId="8" pageNumber="9">G. artensis</emphasis>
</taxonomicName>
is probably lost. Since there is no reasonable way of determining the species, we think this name should also be set aside, as a
<emphasis id="FB8E49523F5BE5447249C485FD3E47CF" box="[593,759,1987,2013]" italics="true" pageId="8" pageNumber="9">nomen dubium</emphasis>
.
</paragraph>
<paragraph id="C94595403F5AE545708FC3D0FB2A40A2" blockId="9.[151,1437,150,1149]" box="[151,1251,150,177]" pageId="9" pageNumber="10">
<emphasis id="FB8E49523F5AE545708FC3D0FB2A40A2" bold="true" box="[151,1251,150,177]" pageId="9" pageNumber="10">
Entirely pale species in the region that do not belong to
<taxonomicName id="0EFAEEC33F5AE5457336C3D0FC5F40A3" baseAuthorityName="Barroga &amp; Mohamedsaid" baseAuthorityYear="2002" box="[814,918,150,177]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="FB8E49523F5AE5457336C3D0FC5F40A3" bold="true" box="[814,918,150,177]" italics="true" pageId="9" pageNumber="10">A. indica</emphasis>
</taxonomicName>
species-group (Appendix 2)
</emphasis>
</paragraph>
<paragraph id="C94595403F5AE545708FC3FCFA4E40C6" blockId="9.[151,1437,150,1149]" box="[151,1415,186,213]" pageId="9" pageNumber="10">
(i) Pale colour morphs of
<taxonomicName id="0EFAEEC33F5AE54571A0C3FCFBFA40C7" authority="(Perroud &amp; Montrouzier, 1864)" baseAuthorityName="Perroud &amp; Montrouzier" baseAuthorityYear="1864" box="[440,1075,186,213]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="deplanchei">
<emphasis id="FB8E49523F5AE54571A0C3FCFD0540C6" box="[440,716,186,212]" italics="true" pageId="9" pageNumber="10">Aulacophora deplanchei</emphasis>
(
<bibRefCitation id="AD6BE8B13F5AE54572C2C3FCFBE340C7" author="Perroud, B. P. &amp; Montrouzier, X." box="[730,1066,186,213]" pageId="9" pageNumber="10" pagination="46 - 256" refId="ref42038" refString="Perroud, B. P. &amp; Montrouzier, X. (1864) Essai sur la faune entomologique de Kanala (Nouvelle-Caledonie) et description de quelques especes nouvelles ou peu connues. Annales de la Societe Lineenne de Lyon, Nouvelle Serie, 11, 46 - 256. https: // doi. org / 10.1080 / 00378941.1864.10827340" type="journal article" year="1864">Perroud &amp; Montrouzier, 1864</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="0EFAEEC33F5AE5457471C3FDFA4D40C6" authority="(Weber, 1801)" baseAuthorityName="Weber" baseAuthorityYear="1801" box="[1129,1412,186,213]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="FB8E49523F5AE5457471C3FDFB1040C6" box="[1129,1241,186,213]" italics="true" pageId="9" pageNumber="10">A. bicolor</emphasis>
(Weber, 1801)
</taxonomicName>
.
</paragraph>
<paragraph id="C94595403F5AE54570DFC399FC0843B7" blockId="9.[151,1437,150,1149]" pageId="9" pageNumber="10">
We have only had access to typical maculate forms of
<taxonomicName id="0EFAEEC33F5AE5457305C399FC7F40EA" baseAuthorityName="Perroud &amp; Montrouzier" baseAuthorityYear="1864" box="[797,950,222,249]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="deplanchei">
<emphasis id="FB8E49523F5AE5457305C399FC7F40EA" box="[797,950,222,249]" italics="true" pageId="9" pageNumber="10">A. deplanchei</emphasis>
</taxonomicName>
, including a photograph of the
<typeStatus id="16412BE23F5AE5457508C398FAB340EA" box="[1296,1402,222,248]" pageId="9" pageNumber="10" type="lectotype">lectotype</typeStatus>
, so the following discussion is partly based on the confusing literature.
<taxonomicName id="0EFAEEC33F5AE545739EC244FECA4153" authority="(Perroud &amp; Montrouzier, 1864)" baseAuthorityName="Perroud &amp; Montrouzier" baseAuthorityYear="1864" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="deplanchei">
<emphasis id="FB8E49523F5AE545739EC244FB53410E" box="[902,1178,258,284]" italics="true" pageId="9" pageNumber="10">Aulacophora deplanchei</emphasis>
(
<bibRefCitation id="AD6BE8B13F5AE54574BFC244FF324153" author="Perroud, B. P. &amp; Montrouzier, X." pageId="9" pageNumber="10" pagination="46 - 256" refId="ref42038" refString="Perroud, B. P. &amp; Montrouzier, X. (1864) Essai sur la faune entomologique de Kanala (Nouvelle-Caledonie) et description de quelques especes nouvelles ou peu connues. Annales de la Societe Lineenne de Lyon, Nouvelle Serie, 11, 46 - 256. https: // doi. org / 10.1080 / 00378941.1864.10827340" type="journal article" year="1864">Perroud &amp; Montrouzier, 1864</bibRefCitation>
)
</taxonomicName>
was described from
<collectingCountry id="B1EDD5D03F5AE54571F4C261FD534152" box="[492,666,294,321]" name="New Caledonia" pageId="9" pageNumber="10">New Caledonia</collectingCountry>
with black maculae on the elytra. It was then redescribed by
<bibRefCitation id="AD6BE8B13F5AE5457550C261FF2B4176" author="Beenen, R." pageId="9" pageNumber="10" pagination="65 - 87" refId="ref39087" refString="Beenen, R. (2008) Contribution to the knowledge of Galerucinae of New Caledonia (Coleoptera: Chrysomelidae). Genus, 19, 65 - 87." type="journal article" year="2008">Beenen (2008)</bibRefCitation>
but he later entirely rejected his own redescription without explanation (
<bibRefCitation id="AD6BE8B13F5AE5457431C20CFB0C4176" author="Beenen, R." box="[1065,1221,330,356]" pageId="9" pageNumber="10" pagination="65 - 108" refId="ref39117" refString="Beenen, R. (2013) Contribution to the knowledge of Galerucinae of New Caledonia 2 (Coleoptera: Chrysomelidae). Genus, 24 (1), 65 - 108." type="journal article" year="2013">Beenen 2013</bibRefCitation>
). His redescription appears to have been an accidental copy and paste of his description of
<taxonomicName id="0EFAEEC33F5AE54573DFC229FADF419A" authority="Beenen, 2008" authorityName="Beenen" authorityYear="2008" box="[967,1302,366,393]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="montrouzieri">
<emphasis id="FB8E49523F5AE54573DFC229FBBF419A" box="[967,1142,367,393]" italics="true" pageId="9" pageNumber="10">A. montrouzieri</emphasis>
<bibRefCitation id="AD6BE8B13F5AE5457464C229FADF419A" author="Beenen, R." box="[1148,1302,366,393]" pageId="9" pageNumber="10" pagination="65 - 87" refId="ref39087" refString="Beenen, R. (2008) Contribution to the knowledge of Galerucinae of New Caledonia (Coleoptera: Chrysomelidae). Genus, 19, 65 - 87." type="journal article" year="2008">
Beenen,
<quantity id="0E0238A53F5AE54574C6C228FADF419A" box="[1246,1302,366,392]" metricMagnitude="1" metricUnit="m" metricValue="5.10032" pageId="9" pageNumber="10" unit="in" value="2008.0">2008</quantity>
</bibRefCitation>
</taxonomicName>
in the same paper (a preoccupied name, later renamed
<taxonomicName id="0EFAEEC33F5AE5457260C2D5FC4441BE" authority="Beenen, 2013" authorityName="Beenen" authorityYear="2013" box="[632,909,402,429]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="xavieri">
<emphasis id="FB8E49523F5AE5457260C2D5FD2F41BE" box="[632,742,403,429]" italics="true" pageId="9" pageNumber="10">A. xavieri</emphasis>
<bibRefCitation id="AD6BE8B13F5AE54572F5C2D5FC4441BE" author="Beenen, R." box="[749,909,402,429]" pageId="9" pageNumber="10" pagination="65 - 108" refId="ref39117" refString="Beenen, R. (2013) Contribution to the knowledge of Galerucinae of New Caledonia 2 (Coleoptera: Chrysomelidae). Genus, 24 (1), 65 - 108." type="journal article" year="2013">Beenen, 2013</bibRefCitation>
</taxonomicName>
).
<bibRefCitation id="AD6BE8B13F5AE54573B8C2D5FB8F41BE" author="Beenen, R." box="[928,1094,402,429]" pageId="9" pageNumber="10" pagination="65 - 87" refId="ref39087" refString="Beenen, R. (2008) Contribution to the knowledge of Galerucinae of New Caledonia (Coleoptera: Chrysomelidae). Genus, 19, 65 - 87." type="journal article" year="2008">Beenen (2008)</bibRefCitation>
noted that some specimens of
<taxonomicName id="0EFAEEC33F5AE545708FC2F1FEFB41C2" baseAuthorityName="Perroud &amp; Montrouzier" baseAuthorityYear="1864" box="[151,306,438,465]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="deplanchei">
<emphasis id="FB8E49523F5AE545708FC2F1FEFB41C2" box="[151,306,438,465]" italics="true" pageId="9" pageNumber="10">A. deplanchei</emphasis>
</taxonomicName>
were entirely pale dorsally. This potentially makes them members of the PPB species-complex. The original description of
<taxonomicName id="0EFAEEC33F5AE545718DC29DFDE741E6" baseAuthorityName="Perroud &amp; Montrouzier" baseAuthorityYear="1864" box="[405,558,474,501]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="deplanchei">
<emphasis id="FB8E49523F5AE545718DC29DFDE741E6" box="[405,558,474,501]" italics="true" pageId="9" pageNumber="10">A. deplanchei</emphasis>
</taxonomicName>
did not mention structures of the first antennomere, male ventrite 5 and female pygidium, but Beenen associated this species with the maculate species
<taxonomicName id="0EFAEEC33F5AE54573A8C2B9FBD5420A" authorityName="Beenen" authorityYear="2013" box="[944,1052,511,537]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="xavieri">
<emphasis id="FB8E49523F5AE54573A8C2B9FBD5420A" box="[944,1052,511,537]" italics="true" pageId="9" pageNumber="10">A. xavieri</emphasis>
</taxonomicName>
, which was described in detail and lacks all of the PPB diagnostic attributes. Our examination of specimens of
<taxonomicName id="0EFAEEC33F5AE54573C0C165FBB8422E" baseAuthorityName="Perroud &amp; Montrouzier" baseAuthorityYear="1864" box="[984,1137,546,573]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="deplanchei">
<emphasis id="FB8E49523F5AE54573C0C165FBB8422E" box="[984,1137,546,573]" italics="true" pageId="9" pageNumber="10">A. deplanchei</emphasis>
</taxonomicName>
shows that the male antennae are simple, the male apical ventrite is short and flat and the female pygidium is not extended. The male genitalia of
<taxonomicName id="0EFAEEC33F5AE54570AEC12DFE984296" baseAuthorityName="Perroud &amp; Montrouzier" baseAuthorityYear="1864" box="[182,337,618,645]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="deplanchei">
<emphasis id="FB8E49523F5AE54570AEC12DFE984296" box="[182,337,618,645]" italics="true" pageId="9" pageNumber="10">A. deplanchei</emphasis>
</taxonomicName>
are certainly similar to
<taxonomicName id="0EFAEEC33F5AE5457278C12DFD074296" authorityName="Beenen" authorityYear="2013" box="[608,718,619,645]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="xavieri">
<emphasis id="FB8E49523F5AE5457278C12DFD074296" box="[608,718,619,645]" italics="true" pageId="9" pageNumber="10">A. xavieri</emphasis>
</taxonomicName>
and dissimilar to any member of the PPB complex treated here (
<bibRefCitation id="AD6BE8B13F5AE5457087C1C8FEF542BA" author="Beenen, R." box="[159,316,654,680]" pageId="9" pageNumber="10" pagination="65 - 87" refId="ref39087" refString="Beenen, R. (2008) Contribution to the knowledge of Galerucinae of New Caledonia (Coleoptera: Chrysomelidae). Genus, 19, 65 - 87." type="journal article" year="2008">Beenen 2008</bibRefCitation>
). We therefore exclude
<taxonomicName id="0EFAEEC33F5AE5457248C1C9FD2442BA" baseAuthorityName="Perroud &amp; Montrouzier" baseAuthorityYear="1864" box="[592,749,654,681]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="deplanchei">
<emphasis id="FB8E49523F5AE5457248C1C9FD2442BA" box="[592,749,654,681]" italics="true" pageId="9" pageNumber="10">A. deplanchei</emphasis>
</taxonomicName>
from the PPB complex, but as it has entirely dorsally pale morphs we include it in our diagnostic key. The presence of rows of long setae on the male antennomeres suggests that
<taxonomicName id="0EFAEEC33F5AE54570DFC191FEA842E2" baseAuthorityName="Perroud &amp; Montrouzier" baseAuthorityYear="1864" box="[199,353,726,753]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="deplanchei">
<emphasis id="FB8E49523F5AE54570DFC191FEA842E2" box="[199,353,726,753]" italics="true" pageId="9" pageNumber="10">A. deplanchei</emphasis>
</taxonomicName>
might be related to both
<taxonomicName id="0EFAEEC33F5AE545726CC190FC4542E2" authority="(Weber, 1801)" baseAuthorityName="Weber" baseAuthorityYear="1801" box="[628,908,726,752]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="bicolor">
A.
<emphasis id="FB8E49523F5AE545728EC190FD2E42E2" box="[662,743,726,752]" italics="true" pageId="9" pageNumber="10">bicolor</emphasis>
(Weber, 1801)
</taxonomicName>
and
<taxonomicName id="0EFAEEC33F5AE54573D9C191FAC942E2" authority="(Hornstedt, 1788)" baseAuthorityName="Hornstedt" baseAuthorityYear="1788" box="[961,1280,726,753]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="coffeae">
<emphasis id="FB8E49523F5AE54573D9C191FBF942E2" box="[961,1072,726,753]" italics="true" pageId="9" pageNumber="10">A. coffeae</emphasis>
(
<bibRefCitation id="AD6BE8B13F5AE5457426C190FB3E42E2" author="Hornstedt, C. F." box="[1086,1271,726,752]" pageId="9" pageNumber="10" pagination="1 - 8" refId="ref40643" refString="Hornstedt, C. F. (1788) Beschreibungen neuer Blattkaferarten. Schriften der Gesellschaft Naturforschender Freunden zu Berlin, 8 (1 - 8), pl. 1." type="journal article" year="1788">Hornstedt, 1788</bibRefCitation>
)
</taxonomicName>
.
<taxonomicName id="0EFAEEC33F5AE5457512C190FEB34306" authority="Baly, 1888" authorityName="Baly" authorityYear="1888" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="perroudi">
<emphasis id="FB8E49523F5AE5457512C190FF314306" italics="true" pageId="9" pageNumber="10">Aulacophora perroudi</emphasis>
<bibRefCitation id="AD6BE8B13F5AE54570E7C1BCFEB34306" author="Baly, J. S." box="[255,378,762,788]" pageId="9" pageNumber="10" pagination="156 - 188" refId="ref38897" refString="Baly, J. S. (1888) Descriptions of some genera and species of Galerucinae. Journal of the Linnean Society, Zoology, 20, 156 - 188. https: // doi. org / 10.1111 / j. 1096 - 3642.1888. tb 01444. x" type="journal article" year="1888">Baly, 1888</bibRefCitation>
</taxonomicName>
, described from
<collectingCountry id="B1EDD5D03F5AE545722BC1BDFD284306" box="[563,737,762,789]" name="New Caledonia" pageId="9" pageNumber="10">New Caledonia</collectingCountry>
and overlooked by subsequent authors, is probably a synonym of
<taxonomicName id="0EFAEEC33F5AE54570AEC059FE98432A" baseAuthorityName="Perroud &amp; Montrouzier" baseAuthorityYear="1864" box="[182,337,798,825]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="deplanchei">
<emphasis id="FB8E49523F5AE54570AEC059FE98432A" box="[182,337,798,825]" italics="true" pageId="9" pageNumber="10">A. deplanchei</emphasis>
</taxonomicName>
as it has dark antennae, dark elytral humeri and female pygidium rounded (
<bibRefCitation id="AD6BE8B13F5AE54574BEC058FA90432A" author="Baly, J. S." box="[1190,1369,798,824]" pageId="9" pageNumber="10" pagination="156 - 188" refId="ref38897" refString="Baly, J. S. (1888) Descriptions of some genera and species of Galerucinae. Journal of the Linnean Society, Zoology, 20, 156 - 188. https: // doi. org / 10.1111 / j. 1096 - 3642.1888. tb 01444. x" type="journal article" year="1888">Baly 1888: 177</bibRefCitation>
). Furthermore, the female pronotal groove is described as absent medially, which is almost true of the small amount of material to hand. However, in the absence of type material and the possibility that there is a valid species on
<collectingCountry id="B1EDD5D03F5AE545757FC021FEC043B6" name="New Caledonia" pageId="9" pageNumber="10">New Caledonia</collectingCountry>
with maculate humeri, we refrain from making this synonym.
</paragraph>
<paragraph id="C94595403F5AE54570DFC0E8FC43446E" blockId="9.[151,1437,150,1149]" pageId="9" pageNumber="10">
<taxonomicName id="0EFAEEC33F5AE54570DFC0E8FE7B43DA" baseAuthorityName="Weber" baseAuthorityYear="1801" box="[199,434,942,968]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="FB8E49523F5AE54570DFC0E8FE7B43DA" box="[199,434,942,968]" italics="true" pageId="9" pageNumber="10">Aulacophora bicolor</emphasis>
</taxonomicName>
is a widespread and variable south-east Asian species, which usually has black markings on the elytra but has rare entirely dorsally pale colour morphs (
<bibRefCitation id="AD6BE8B13F5AE5457363C094FB2F43FE" author="Barroga, G. F. &amp; Mohamedsaid, M. S." box="[891,1254,978,1004]" pageId="9" pageNumber="10" pagination="15 - 194" refId="ref39042" refString="Barroga, G. F. &amp; Mohamedsaid, M. S. (2002) Revision of the genus Aulacophora Chevrolat (Coleoptera: Chrysomelidae: Galerucinae) in Sundaland. Serangga, 7 (1 - 2), 15 - 194." type="journal article" year="2002">Barroga &amp; Mohamedsaid 2002</bibRefCitation>
;
<bibRefCitation id="AD6BE8B13F5AE54574ECC095FF1D4403" author="Lee, C. - F. &amp; Beenen, R." pageId="9" pageNumber="10" pagination="151 - 190" refId="ref41061" refString="Lee, C. - F. &amp; Beenen, R. (2015) Revision of the genus Aulacophora from Taiwan (Coleoptera: Chrysomelidae: Galerucinae). Zootaxa, 3949 (2), 151 - 190. https: // doi. org / 10.11646 / zootaxa. 3949.2.1" type="journal article" year="2015">Lee &amp; Beenen 2015</bibRefCitation>
).
<taxonomicName id="0EFAEEC33F5AE54570FFC0B0FE1B4402" baseAuthorityName="Weber" baseAuthorityYear="1801" box="[231,466,1014,1040]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="FB8E49523F5AE54570FFC0B0FE1B4402" box="[231,466,1014,1040]" italics="true" pageId="9" pageNumber="10">Aulacophora bicolor</emphasis>
</taxonomicName>
has an unexpanded first antennomere, short flat apical male ventrite and short female pygidium and therefore lacks the characters defining the PPB complex (
<bibRefCitation id="AD6BE8B13F5AE54573A7C75CFAEB4426" author="Barroga, G. F. &amp; Mohamedsaid, M. S." box="[959,1314,1050,1076]" pageId="9" pageNumber="10" pagination="15 - 194" refId="ref39042" refString="Barroga, G. F. &amp; Mohamedsaid, M. S. (2002) Revision of the genus Aulacophora Chevrolat (Coleoptera: Chrysomelidae: Galerucinae) in Sundaland. Serangga, 7 (1 - 2), 15 - 194." type="journal article" year="2002">Barroga &amp; Mohamedsaid 2002</bibRefCitation>
). This species is not recorded from the Pacific Region but is associated with cucurbit crops and is recorded from as far east as Timor (Mohamedsaid 2009), so it is included in our diagnostic key.
</paragraph>
<paragraph id="C94595403F5AE545708FC7ECFDB044D6" blockId="9.[151,1437,1194,1833]" box="[151,633,1194,1221]" pageId="9" pageNumber="10">
(ii)
<taxonomicName id="0EFAEEC33F5AE54570D8C7ECFDB044D6" authority="(Hornstedt, 1788)" baseAuthorityName="Hornstedt" baseAuthorityYear="1788" box="[192,633,1194,1220]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="coffeae">
<emphasis id="FB8E49523F5AE54570D8C7ECFE6244D6" box="[192,427,1194,1220]" italics="true" pageId="9" pageNumber="10">Aulacophora coffeae</emphasis>
(
<bibRefCitation id="AD6BE8B13F5AE54571A1C7ECFDB944D6" author="Hornstedt, C. F." box="[441,624,1194,1220]" pageId="9" pageNumber="10" pagination="1 - 8" refId="ref40643" refString="Hornstedt, C. F. (1788) Beschreibungen neuer Blattkaferarten. Schriften der Gesellschaft Naturforschender Freunden zu Berlin, 8 (1 - 8), pl. 1." type="journal article" year="1788">Hornstedt, 1788</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="C94595403F5AE54570DFC788FAFA473A" blockId="9.[151,1437,1194,1833]" pageId="9" pageNumber="10">
<taxonomicName id="0EFAEEC33F5AE54570DFC788FE7944FA" baseAuthorityName="Hornstedt" baseAuthorityYear="1788" box="[199,432,1230,1256]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="coffeae">
<emphasis id="FB8E49523F5AE54570DFC788FE7944FA" box="[199,432,1230,1256]" italics="true" pageId="9" pageNumber="10">Aulacophora coffeae</emphasis>
</taxonomicName>
is another pale species of
<emphasis id="FB8E49523F5AE54572D5C788FCAF44FA" box="[717,870,1230,1256]" italics="true" pageId="9" pageNumber="10">
<taxonomicName id="0EFAEEC33F5AE54572D5C788FCAB44FA" authorityName="Chevrolat" authorityYear="1836" box="[717,866,1230,1256]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">Aulacophora</taxonomicName>
,
</emphasis>
which, like
<taxonomicName id="0EFAEEC33F5AE54573F4C789FB9344FA" baseAuthorityName="Weber" baseAuthorityYear="1801" box="[1004,1114,1230,1257]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="FB8E49523F5AE54573F4C789FB9344FA" box="[1004,1114,1230,1257]" italics="true" pageId="9" pageNumber="10">A. bicolor</emphasis>
</taxonomicName>
, lacks the diagnostic secondary sexual characters of the PPB complex (
<bibRefCitation id="AD6BE8B13F5AE545726DC7B4FC1F451E" author="Barroga, G. F. &amp; Mohamedsaid, M. S." box="[629,982,1266,1292]" pageId="9" pageNumber="10" pagination="15 - 194" refId="ref39042" refString="Barroga, G. F. &amp; Mohamedsaid, M. S. (2002) Revision of the genus Aulacophora Chevrolat (Coleoptera: Chrysomelidae: Galerucinae) in Sundaland. Serangga, 7 (1 - 2), 15 - 194." type="journal article" year="2002">Barroga &amp; Mohamedsaid 2002</bibRefCitation>
).
<taxonomicName id="0EFAEEC33F5AE54573F0C7B4FB18451E" baseAuthorityName="Hornstedt" baseAuthorityYear="1788" box="[1000,1233,1266,1292]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="coffeae">
<emphasis id="FB8E49523F5AE54573F0C7B4FB18451E" box="[1000,1233,1266,1292]" italics="true" pageId="9" pageNumber="10">Aulacophora coffeae</emphasis>
</taxonomicName>
is a pest of cucurbits in southeast Asia and
<collectingCountry id="B1EDD5D03F5AE54571A1C650FDEF4522" box="[441,550,1302,1328]" name="Indonesia" pageId="9" pageNumber="10">Indonesia</collectingCountry>
(Kalshoven 1951;
<bibRefCitation id="AD6BE8B13F5AE54572E2C650FC324522" author="Lewis, P. A. &amp; Metcalf, R. L." box="[762,1019,1302,1328]" pageId="9" pageNumber="10" pagination="150 - 155" refId="ref41192" refString="Lewis, P. A. &amp; Metcalf, R. L. (1996) Behavior and ecology of Old World Luperini beetles of the genus Aulacophora (Coleoptera: Chrysomelidae). Chemoecology, 7, 150 - 155. https: // doi. org / 10.1007 / BF 01245968" type="journal article" year="1996">Lewis &amp; Metcalf 1996</bibRefCitation>
;
<bibRefCitation id="AD6BE8B13F5AE545741EC650FB3D4522" author="Mohamedsaid, M. S." box="[1030,1268,1302,1328]" pageId="9" pageNumber="10" refId="ref41633" refString="Mohamedsaid, M. S. (2009 a) Chrysomelidae of the Lesser Sunda Islands: Wallace's Line and the crossing of worlds. In: Jolivet, P., Santiago-Blay, J. &amp; Schmitt, M. (Eds.), Research on Chrysomelidae. Vol. 2. Brill, Leiden, pp. 57 - 104. https: // doi. org / 10.1163 / ej. 9789004169470.1 - 299.21" type="book" year="2009">Mohamedsaid 2009a</bibRefCitation>
), also recorded from
<taxonomicName id="0EFAEEC33F5AE54570CDC67DFD784546" authority="(Barroga &amp; Mohamedsaid 2002)" baseAuthorityName="Barroga &amp; Mohamedsaid" baseAuthorityYear="2002" box="[213,689,1338,1364]" class="Magnoliopsida" family="Convolvulaceae" genus="Ipomoea" kingdom="Plantae" order="Solanales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="genus">
<emphasis id="FB8E49523F5AE54570CDC67DFEFE4546" box="[213,311,1339,1364]" italics="true" pageId="9" pageNumber="10">Ipomoea</emphasis>
(
<bibRefCitation id="AD6BE8B13F5AE545715EC67CFD604546" author="Barroga, G. F. &amp; Mohamedsaid, M. S." box="[326,681,1338,1364]" pageId="9" pageNumber="10" pagination="15 - 194" refId="ref39042" refString="Barroga, G. F. &amp; Mohamedsaid, M. S. (2002) Revision of the genus Aulacophora Chevrolat (Coleoptera: Chrysomelidae: Galerucinae) in Sundaland. Serangga, 7 (1 - 2), 15 - 194." type="journal article" year="2002">Barroga &amp; Mohamedsaid 2002</bibRefCitation>
)
</taxonomicName>
. In the region under review, it has been noted from
<collectingCountry id="B1EDD5D03F5AE54574EFC67CFAD74546" box="[1271,1310,1338,1364]" name="Fiji" pageId="9" pageNumber="10">Fiji</collectingCountry>
feeding on
<taxonomicName id="0EFAEEC33F5AE545708FC619FF30456B" box="[151,249,1375,1401]" class="Magnoliopsida" family="Cucurbitaceae" genus="Cucumis" kingdom="Plantae" order="Cucurbitales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="genus">
<emphasis id="FB8E49523F5AE545708FC619FF30456B" box="[151,249,1375,1401]" italics="true" pageId="9" pageNumber="10">Cucumis</emphasis>
</taxonomicName>
and
<taxonomicName id="0EFAEEC33F5AE5457135C618FE56456A" box="[301,415,1374,1400]" class="Magnoliopsida" family="Cucurbitaceae" genus="Cucurbita" kingdom="Plantae" order="Cucurbitales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="genus">
<emphasis id="FB8E49523F5AE5457135C618FE56456A" box="[301,415,1374,1400]" italics="true" pageId="9" pageNumber="10">Cucurbita</emphasis>
</taxonomicName>
(
<bibRefCitation id="AD6BE8B13F5AE54571B3C618FDB8456A" author="Greenwood, W." box="[427,625,1374,1400]" pageId="9" pageNumber="10" pagination="211 - 218" refId="ref40303" refString="Greenwood, W. (1940) The food-plants or hosts of some Fijian insects. IV. Proceedings of the Linnean Society of New South Wales, 65, 211 - 218." type="journal article" year="1940">Greenwood 1940</bibRefCitation>
;
<bibRefCitation id="AD6BE8B13F5AE5457263C619FD37456A" author="Lever, R. J. A. W." box="[635,766,1374,1401]" pageId="9" pageNumber="10" pagination="48 - 49" refId="ref41161" refString="Lever, R. J. A. W. (1942) Entomological notes. Fiji Department of Agriculture Journal, 13 (2), 48 - 49." type="journal article" year="1942">Lever 1942</bibRefCitation>
), but these records were queried by
<bibRefCitation id="AD6BE8B13F5AE5457490C619FA55456B" author="Bryant, G. E. &amp; Gressitt, J. L." box="[1160,1436,1374,1401]" pageId="9" pageNumber="10" pagination="3 - 91" refId="ref39598" refString="Bryant, G. E. &amp; Gressitt, J. L. (1957) Chrysomelidae of Fiji (Coleoptera). Pacific Science, 11, 3 - 91." type="journal article" year="1957">Bryant &amp; Gressitt (1957)</bibRefCitation>
who implied that they referred to
<taxonomicName id="0EFAEEC33F5AE5457208C6C5FDB1458E" authorityName="sensu Maulik" authorityYear="1929" baseAuthorityName="Olivier" baseAuthorityYear="1808" box="[528,632,1410,1437]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="FB8E49523F5AE5457208C6C5FDB1458E" box="[528,632,1410,1437]" italics="true" pageId="9" pageNumber="10">A. similis</emphasis>
</taxonomicName>
(itself a misidentification of
<taxonomicName id="0EFAEEC33F5AE54573A6C6C5FBA1458E" authorityName="Mer" baseAuthorityName="Fabricius" baseAuthorityYear="1781" box="[958,1128,1410,1437]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis id="FB8E49523F5AE54573A6C6C5FBA1458E" box="[958,1128,1410,1437]" italics="true" pageId="9" pageNumber="10">A. abdominalis</emphasis>
</taxonomicName>
). In
<collectingCountry id="B1EDD5D03F5AE5457486C6C5FAB0458E" box="[1182,1401,1410,1437]" name="Papua New Guinea" pageId="9" pageNumber="10">Papua New Guinea</collectingCountry>
,
<taxonomicName id="0EFAEEC33F5AE545759CC6C5FF2145D2" baseAuthorityName="Hornstedt" baseAuthorityYear="1788" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="coffeae">
<emphasis id="FB8E49523F5AE545759CC6C5FF2145D2" italics="true" pageId="9" pageNumber="10">A. coffeae</emphasis>
</taxonomicName>
has been recorded feeding on cucurbits and legumes: mung bean (
<taxonomicName id="0EFAEEC33F5AE54573D4C6E1FBC445D2" box="[972,1037,1447,1472]" class="Magnoliopsida" family="Fabaceae" genus="Vigna" kingdom="Plantae" order="Fabales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="genus">
<emphasis id="FB8E49523F5AE54573D4C6E1FBC445D2" box="[972,1037,1447,1472]" italics="true" pageId="9" pageNumber="10">Vigna</emphasis>
</taxonomicName>
), rock melon (
<taxonomicName id="0EFAEEC33F5AE54574A9C6E1FADA45D3" box="[1201,1299,1447,1473]" class="Magnoliopsida" family="Cucurbitaceae" genus="Cucumis" kingdom="Plantae" order="Cucurbitales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="genus">
<emphasis id="FB8E49523F5AE54574A9C6E1FADA45D3" box="[1201,1299,1447,1473]" italics="true" pageId="9" pageNumber="10">Cucumis</emphasis>
</taxonomicName>
) and peanut (
<taxonomicName id="0EFAEEC33F5AE54570B9C68CFF3145F6" box="[161,248,1482,1508]" class="Magnoliopsida" family="Fabaceae" genus="Arachis" kingdom="Plantae" order="Fabales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="genus">
<emphasis id="FB8E49523F5AE54570B9C68CFF3145F6" box="[161,248,1482,1508]" italics="true" pageId="9" pageNumber="10">Arachis</emphasis>
</taxonomicName>
) (
<bibRefCitation id="AD6BE8B13F5AE5457117C68CFDC845F6" author="Greve, J. E. van &amp; Ismay, J. W." box="[271,513,1482,1509]" pageId="9" pageNumber="10" pagination="1 - 4" refId="ref40402" refString="Greve, J. E. van S. &amp; Ismay, J. W. (1983) Crop insect survey of Papua New Guinea from July 1 st 1969 to December 31 st 1978. Papua New Guinea Agricultural Journal, 32 (1 - 4), i-iv + 1 - 123." type="journal article" year="1983">Greve &amp; Ismay 1983</bibRefCitation>
). All specimens of pale
<taxonomicName id="0EFAEEC33F5AE5457314C68CFC5745F6" authorityName="Chevrolat" authorityYear="1836" box="[780,926,1482,1508]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="FB8E49523F5AE5457314C68CFC5745F6" box="[780,926,1482,1508]" italics="true" pageId="9" pageNumber="10">Aulacophora</emphasis>
</taxonomicName>
we have examined from
<collectingCountry id="B1EDD5D03F5AE54574A2C68CFB2845F6" box="[1210,1249,1482,1508]" name="Fiji" pageId="9" pageNumber="10">Fiji</collectingCountry>
belong to
<taxonomicName id="0EFAEEC33F5AE5457540C68DFECC461A" authorityName="Mer" baseAuthorityName="Fabricius" baseAuthorityYear="1781" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis id="FB8E49523F5AE5457540C68DFECC461A" italics="true" pageId="9" pageNumber="10">A. abdominalis</emphasis>
</taxonomicName>
and all pale
<taxonomicName id="0EFAEEC33F5AE545718FC6A8FDE0461A" authorityName="Chevrolat" authorityYear="1836" box="[407,553,1518,1544]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="FB8E49523F5AE545718FC6A8FDE0461A" box="[407,553,1518,1544]" italics="true" pageId="9" pageNumber="10">Aulacophora</emphasis>
</taxonomicName>
we have examined from New
<collectingCountry id="B1EDD5D03F5AE545739CC6A8FC1C461A" box="[900,981,1518,1544]" name="Guinea-Bissau" pageId="9" pageNumber="10">Guinea</collectingCountry>
belong to
<taxonomicName id="0EFAEEC33F5AE5457456C6A9FB31461A" authorityName="Mer" baseAuthorityName="Fabricius" baseAuthorityYear="1781" box="[1102,1272,1518,1545]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis id="FB8E49523F5AE5457456C6A9FB31461A" box="[1102,1272,1518,1545]" italics="true" pageId="9" pageNumber="10">A. abdominalis</emphasis>
</taxonomicName>
,
<taxonomicName id="0EFAEEC33F5AE545751FC6A9FAB4461A" authorityName="Baly" authorityYear="1879" box="[1287,1405,1519,1545]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="FB8E49523F5AE545751FC6A9FAB4461A" box="[1287,1405,1519,1545]" italics="true" pageId="9" pageNumber="10">A. cornuta</emphasis>
</taxonomicName>
or
<taxonomicName id="0EFAEEC33F5AE545708FC555FF34463E" baseAuthorityName="Barroga &amp; Mohamedsaid" baseAuthorityYear="2002" box="[151,253,1554,1581]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="FB8E49523F5AE545708FC555FF34463E" box="[151,253,1554,1581]" italics="true" pageId="9" pageNumber="10">A. indica</emphasis>
</taxonomicName>
.
<bibRefCitation id="AD6BE8B13F5AE5457113C554FDC3463E" author="Greve, J. E. van &amp; Ismay, J. W." box="[267,522,1554,1581]" pageId="9" pageNumber="10" pagination="1 - 4" refId="ref40402" refString="Greve, J. E. van S. &amp; Ismay, J. W. (1983) Crop insect survey of Papua New Guinea from July 1 st 1969 to December 31 st 1978. Papua New Guinea Agricultural Journal, 32 (1 - 4), i-iv + 1 - 123." type="journal article" year="1983">Greve &amp; Ismay (1983)</bibRefCitation>
did not indicate where their material was deposited but at the time of their report both were based in Port Moresby. Our colleague Al Samuelson has searched the largest holding of New Guinean
<taxonomicName id="0EFAEEC33F5AE545708FC51CFE884666" box="[151,321,1626,1652]" class="Insecta" family="Chrysomelidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="family">Chrysomelidae</taxonomicName>
in the world, in BPBM, and has failed to find any specimens of
<taxonomicName id="0EFAEEC33F5AE545740BC51DFB4A4666" baseAuthorityName="Hornstedt" baseAuthorityYear="1788" box="[1043,1155,1626,1653]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="coffeae">
<emphasis id="FB8E49523F5AE545740BC51DFB4A4666" box="[1043,1155,1626,1653]" italics="true" pageId="9" pageNumber="10">A. coffeae</emphasis>
</taxonomicName>
from New
<collectingCountry id="B1EDD5D03F5AE545751CC51CFA9C4666" box="[1284,1365,1626,1652]" name="Guinea-Bissau" pageId="9" pageNumber="10">Guinea</collectingCountry>
or the Pacific (Samuelson, pers. com.
<date id="BD44B3803F5AE54571E3C538FD71468A" box="[507,696,1662,1688]" pageId="9" pageNumber="10" value="2018-09">September 2018</date>
). We therefore conclude that all records of
<taxonomicName id="0EFAEEC33F5AE5457485C539FAC7468A" baseAuthorityName="Hornstedt" baseAuthorityYear="1788" box="[1181,1294,1662,1689]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="coffeae">
<emphasis id="FB8E49523F5AE5457485C539FAC7468A" box="[1181,1294,1662,1689]" italics="true" pageId="9" pageNumber="10">A. coffeae</emphasis>
</taxonomicName>
from the region of interest have been misidentifications, which may have been caused by early misidentification of
<taxonomicName id="0EFAEEC33F5AE5457534C5E5FA5546AE" baseAuthorityName="Hornstedt" baseAuthorityYear="1788" box="[1324,1436,1698,1725]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="coffeae">
<emphasis id="FB8E49523F5AE5457534C5E5FA5546AE" box="[1324,1436,1698,1725]" italics="true" pageId="9" pageNumber="10">A. coffeae</emphasis>
</taxonomicName>
by
<bibRefCitation id="AD6BE8B13F5AE54570A0C580FE9D46F2" author="Allard, E." box="[184,340,1734,1760]" pageId="9" pageNumber="10" pagination="305 - 332" refId="ref38527" refString="Allard, E. (1888) Synopsis des Galerucines a corselet sillonne transversalement. Premiere partie. Annales de la Societe entomologique de France, 8 (6), 305 - 332." type="journal article" year="1888">Allard (1888)</bibRefCitation>
, as noted by
<bibRefCitation id="AD6BE8B13F5AE54571FDC580FDA646F2" author="Baly, J. S." box="[485,623,1734,1760]" pageId="9" pageNumber="10" pagination="297 - 306" refId="ref38948" refString="Baly, J. S. (1889) Notes on Aulacophora and allied genera. Transactions of the entomological Society, London, 1889 (2), 297 - 306. https: // doi. org / 10.1111 / j. 1365 - 2311.1889. tb 02322. x" type="journal article" year="1889">Baly (1889)</bibRefCitation>
. The nearest accurate record of
<taxonomicName id="0EFAEEC33F5AE54573CAC581FB8B46F2" baseAuthorityName="Hornstedt" baseAuthorityYear="1788" box="[978,1090,1734,1761]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="coffeae">
<emphasis id="FB8E49523F5AE54573CAC581FB8B46F2" box="[978,1090,1734,1761]" italics="true" pageId="9" pageNumber="10">A. coffeae</emphasis>
</taxonomicName>
is from Sumba (
<bibRefCitation id="AD6BE8B13F5AE54574E4C580FF294716" author="Mohamedsaid, M. S." pageId="9" pageNumber="10" refId="ref41633" refString="Mohamedsaid, M. S. (2009 a) Chrysomelidae of the Lesser Sunda Islands: Wallace's Line and the crossing of worlds. In: Jolivet, P., Santiago-Blay, J. &amp; Schmitt, M. (Eds.), Research on Chrysomelidae. Vol. 2. Brill, Leiden, pp. 57 - 104. https: // doi. org / 10.1163 / ej. 9789004169470.1 - 299.21" type="book" year="2009">Mohamedsaid 2009a</bibRefCitation>
), only
<quantity id="0E0238A53F5AE5457132C5ACFEB74716" box="[298,382,1770,1796]" metricMagnitude="5" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="10" unit="km" value="300.0">300 km</quantity>
west of Timor, so it may spread further east, as little agricultural biosecurity is practiced within the Indonesian Archipelago. The species is included in our diagnostic key in case it appears in the region.
</paragraph>
<subSubSection id="81E0C6CB3F5AE545708FC43CFE2F47AA" pageId="9" pageNumber="10" type="key">
<paragraph id="C94595403F5AE545708FC43CFE2F47AA" blockId="9.[151,1436,1914,1976]" pageId="9" pageNumber="10">
<emphasis id="FB8E49523F5AE545708FC43CFE2F47AA" bold="true" pageId="9" pageNumber="10">
Key to species of plain pumpkin-beetles,
<taxonomicName id="0EFAEEC33F5AE545728EC43CFC4F4787" baseAuthorityName="Barroga &amp; Mohamedsaid" baseAuthorityYear="2002" box="[662,902,1914,1941]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="FB8E49523F5AE545728EC43CFC4F4787" bold="true" box="[662,902,1914,1941]" italics="true" pageId="9" pageNumber="10">Aulacophora indica</emphasis>
</taxonomicName>
species-complex, in Australopapua, Timor and the west Pacific region
</emphasis>
</paragraph>
</subSubSection>
</subSubSection>
<subSubSection id="81E0C6CB3F5AE540708FC4A5FA554131" lastPageId="12" lastPageNumber="13" pageId="9" pageNumber="10" type="key">
<key id="BD6BC2713F5AE540708FC4A5FA554131" lastPageId="12" lastPageNumber="13" pageId="9" pageNumber="10">
<keyStep id="720E8DE53F5AE546708FC4A5FA55414F" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<paragraph id="C94595403F5AE546708FC4A5FA5540D8" blockId="9.[151,1436,2019,2041]" lastBlockId="10.[151,1437,151,2021]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<keyLead id="720B36753F5AE546708FC4A5FA5540D8" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<specimenCount id="DFFC5EC93F5AE545708FC4A5FEDA47EB" box="[151,275,2019,2041]" pageId="9" pageNumber="10" type="male">1 Male</specimenCount>
without enlarged first antennomere, as female; male antennomeres 411 with longitudinal row of erect setae on inner margins; male last ventrite with short flat apical median projection; penis symmetrical; female pygidium neither dorsally swollen nor with apex extended beyond apical ventrite (pygidium dark brown to black)................................. 2
</keyLead>
</paragraph>
<paragraph id="C94595403F59E546708FC395FA55414F" blockId="10.[151,1437,151,2021]" pageId="10" pageNumber="11">
<keyLead id="720B36753F59E546708FC395FA55414F" pageId="10" pageNumber="11">
- Male first antennomere enlarged, like an inverted pyramid or almost cuboid, with one side partly flattened (
<figureCitation id="51C189C53F59E54674CEC394FAF140FA" box="[1238,1336,210,232]" captionStart-0="FIGURES 25" captionStart-1="FIGURES 69" captionStart-2="FIGURES 1015" captionStart-3="FIGURES 1617" captionStartId-0="15.[151,264,1829,1853]" captionStartId-1="16.[151,264,1957,1981]" captionStartId-2="17.[151,264,1943,1967]" captionStartId-3="18.[151,264,783,807]" captionTargetBox-0="[164,1420,191,1786]" captionTargetBox-1="[174,1418,193,1917]" captionTargetBox-2="[209,1375,181,1906]" captionTargetBox-3="[158,1427,191,748]" captionTargetId-0="figure-19@15.[151,1436,181,1805]" captionTargetId-1="figure-22@16.[157,1430,181,1933]" captionTargetId-2="figure-19@17.[209,1379,181,1920]" captionTargetId-3="figure-22@18.[151,1435,181,759]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionTargetPageId-2="17" captionTargetPageId-3="18" captionText-0="FIGURES 25. Aulacophora species, male habitus: 2, A. abdominalis (Fabricius, 1781) (Guadalcanal); 3, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 4, A. cornuta Baly, 1879 (New Ireland); 5, A. indica (Gmelin, 1790) (Luzon)." captionText-1="FIGURES 69. Aulacophora species, male habitus: 6, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 7, A. relicta (Boisduval, 1835) (Mount Molloy); 8, A. wallacii Baly, 1886; 9, A. wilsoni Baly, 1888." captionText-2="FIGURES 1015. Aulacophora species, male head, anterior view: 10, A. abdominalis (Fabricius, 1781) (Guadalcanal); 11, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 12, A. cornuta Baly, 1879 (New Ireland); 13, A. indica (Gmelin, 1790) (Luzon); 14, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 15, A. relicta (Boisduval, 1835) (Mount Molloy)." captionText-3="FIGURES 1617. Aulacophora species, male head, anterior view: 16, A. wallacii Baly, 1886; 17, A. wilsoni Baly, 1888." figureDoi-0="http://doi.org/10.5281/zenodo.4545452" figureDoi-1="http://doi.org/10.5281/zenodo.4545458" figureDoi-2="http://doi.org/10.5281/zenodo.4545460" figureDoi-3="http://doi.org/10.5281/zenodo.4545462" httpUri-0="https://zenodo.org/record/4545452/files/figure.png" httpUri-1="https://zenodo.org/record/4545458/files/figure.png" httpUri-2="https://zenodo.org/record/4545460/files/figure.png" httpUri-3="https://zenodo.org/record/4545462/files/figure.png" pageId="10" pageNumber="11">Figs 217</figureCitation>
), contrasting with almost cylindrical segment of female; male antennomeres 411 with 46 scattered erect setae on inner margin between base and apex, not forming a single longitudinal row; male last ventrite with a deeply and elongately hollowed median lobe (
<figureCitation id="51C189C53F59E54670F1C26CFE924152" box="[233,347,298,320]" captionStart-0="FIGURES 3237" captionStart-1="FIGURES 3843" captionStartId-0="23.[151,264,1840,1864]" captionStartId-1="24.[151,264,1793,1817]" captionTargetBox-0="[167,1416,192,1802]" captionTargetBox-1="[168,1418,191,1757]" captionTargetId-0="figure-19@23.[151,1436,181,1816]" captionTargetId-1="figure-22@24.[151,1437,181,1769]" captionTargetPageId-0="23" captionTargetPageId-1="24" captionText-0="FIGURES 3237. Aulacophora species, male, apex of abdominal venter: 32, A. abdominalis (Fabricius, 1781) (Vanuatu); 33, A. abdominalis (New Ireland); 34, A. abdominalis (Manatuto); 35, A. abdominalis (Metinaro); 36, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 37, A. cornuta Baly, 1879 (Guadalcanal)." captionText-1="FIGURES 3843. Aulacophora species, male, apex of abdominal venter: 38, A. indica (Gmelin, 1790) (Sumatra); 39, A. indica (West Timor); 40, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 41, A. relicta (Boisduval, 1835) (Mount Molloy); 42, A. wallacii Baly, 1886; 43, A. wilsoni Baly, 1888." figureDoi-0="http://doi.org/10.5281/zenodo.4545468" figureDoi-1="http://doi.org/10.5281/zenodo.4545470" httpUri-0="https://zenodo.org/record/4545468/files/figure.png" httpUri-1="https://zenodo.org/record/4545470/files/figure.png" pageId="10" pageNumber="11">Figs 3243</figureCitation>
); penis usually asymmetric in dorsal view (
<figureCitation id="51C189C53F59E546731EC26CFC4D4152" box="[774,900,298,320]" captionStart-0="FIGURES 9497" captionStart-1="FIGURES 98101" captionStart-2="FIGURES 102105" captionStart-3="FIGURES 106107" captionStartId-0="44.[151,264,1737,1761]" captionStartId-1="45.[151,264,1825,1849]" captionStartId-2="46.[151,264,1777,1801]" captionStartId-3="47.[151,264,1987,2011]" captionTargetBox-0="[170,1385,198,1674]" captionTargetBox-1="[164,1393,195,1754]" captionTargetBox-2="[166,1388,194,1718]" captionTargetBox-3="[425,1163,195,1932]" captionTargetId-0="figure-22@44.[151,1435,181,1713]" captionTargetId-1="figure-19@45.[151,1436,181,1801]" captionTargetId-2="figure-22@46.[151,1436,181,1754]" captionTargetId-3="figure-19@47.[405,1183,181,1963]" captionTargetPageId-0="44" captionTargetPageId-1="45" captionTargetPageId-2="46" captionTargetPageId-3="47" captionText-0="FIGURES 9497. Aulacophora abdominalis (Fabricius, 1781), penis, dorsal view: 94, Guadalcanal, 95, Metinaro, 96, Saibai, 97, Vanuatu." captionText-1="FIGURES 98101. Aulacophora species, penis, dorsal view: 98, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 99, A. cornuta Baly, 1879 (Guadalcanal); 100, A. indica (Gmelin, 1790) (Laos); 101, A. indica (Luzon)." captionText-2="FIGURES 102105. Aulacophora species, penis, dorsal view: 102, A. indica (Gmelin, 1790) (Timor); 103, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 104, A. relicta (Boisduval, 1835) (Mount Molloy); 105, A. relicta (Mount Mulligan)." captionText-3="FIGURES 106107. Aulacophora species, penis, dorsal view: 106, A. wilsoni Baly, 1888; 107, A. wallacii Baly, 1886." figureDoi-0="http://doi.org/10.5281/zenodo.4545488" figureDoi-1="http://doi.org/10.5281/zenodo.4545490" figureDoi-2="http://doi.org/10.5281/zenodo.4545492" figureDoi-3="http://doi.org/10.5281/zenodo.4545494" httpUri-0="https://zenodo.org/record/4545488/files/figure.png" httpUri-1="https://zenodo.org/record/4545490/files/figure.png" httpUri-2="https://zenodo.org/record/4545492/files/figure.png" httpUri-3="https://zenodo.org/record/4545494/files/figure.png" pageId="10" pageNumber="11">Figs 94107</figureCitation>
); female pygidium with apex usually extended beyond apex of ventrite 5 and often dorsally swollen (
<figureCitation id="51C189C53F59E5467283C201FCC0414F" box="[667,777,327,349]" captionStart-0="FIGURES 4449" captionStart-1="FIGURES 5055" captionStartId-0="25.[151,264,1798,1822]" captionStartId-1="26.[151,264,1943,1967]" captionTargetBox-0="[170,1418,190,1763]" captionTargetBox-1="[198,1388,201,1907]" captionTargetId-0="figure-19@25.[151,1436,181,1774]" captionTargetId-1="figure-22@26.[178,1409,181,1919]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionText-0="FIGURES 4449. Aulacophora species, female, apex of abdominal venter and pygidium: 44, A. abdominalis (Fabricius, 1781) (Vanuatu); 45, A. abdominalis (New Ireland); 46, A. abdominalis (Mer Island); 47, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 48, A. cornuta Baly, 1879 (Sulawesi); 49, A. indica (Gmelin, 1790) (Laos).Dotted lines indicate depth and width of apical concavity." captionText-1="FIGURES 5055. Aulacophora species, female, apex of abdominal venter: 50, A. indica (Gmelin, 1790) (Biak Island); 51, A. indica (Vietnam); 52, A. indica (Broken Hill); 53, A. mbabaram Reid, Halling &amp; Beatson, sp. nov., Almaden; 54, A. relicta (Boisduval, 1835); 55, A. wallacii Baly, 1886." figureDoi-0="http://doi.org/10.5281/zenodo.4545472" figureDoi-1="http://doi.org/10.5281/zenodo.4545474" httpUri-0="https://zenodo.org/record/4545472/files/figure.png" httpUri-1="https://zenodo.org/record/4545474/files/figure.png" pageId="10" pageNumber="11">Figs 4455</figureCitation>
) (
<taxonomicName id="0EFAEEC33F59E5467306C20EFCBD414E" baseAuthorityName="Barroga &amp; Mohamedsaid" baseAuthorityYear="2002" box="[798,884,327,349]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="FB8E49523F59E5467306C20EFCBD414E" box="[798,884,327,349]" italics="true" pageId="10" pageNumber="11">A. indica</emphasis>
</taxonomicName>
species-complex).............................. 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="720E8DE53F59E546708FC223FA5541E2" pageId="10" pageNumber="11">
<paragraph id="C94595403F59E546708FC223FA554185" blockId="10.[151,1437,151,2021]" pageId="10" pageNumber="11">
<keyLead id="720B36753F59E546708FC223FA554185" pageId="10" pageNumber="11">2(1) Elytra dull, finely microreticulate; antennae entirely pale; pygidium clothed in dense silvery recumbent setae; labrum black; pronotal transverse depression deep at middle............................................................... 3</keyLead>
</paragraph>
<paragraph id="C94595403F59E546708FC2E6FA5541E2" blockId="10.[151,1437,151,2021]" pageId="10" pageNumber="11">
<keyLead id="720B36753F59E546708FC2E6FA5541E2" pageId="10" pageNumber="11">
- Elytra shiny, without microreticulation; antennomeres 411 dark brown to black; pygidium with sparser semi-erect brownish setae; labrum pale brown; pronotal depression shallow at middle (elytra usually blotched or maculate; endemic to
<collectingCountry id="B1EDD5D03F59E546752CC2FBFED041E2" name="New Caledonia" pageId="10" pageNumber="11">New Caledonia</collectingCountry>
)...............................................................
<taxonomicName id="0EFAEEC33F59E5467438C29CFA5541E2" authority="(Perroud &amp; Montrouzier)" baseAuthorityName="Perroud &amp; Montrouzier" baseAuthorityYear="1864" box="[1056,1436,474,496]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="deplanchei">
<emphasis id="FB8E49523F59E5467438C29CFB6A41FD" box="[1056,1187,474,495]" italics="true" pageId="10" pageNumber="11">A. deplanchei</emphasis>
(Perroud &amp; Montrouzier)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="720E8DE53F59E546708FC2BEFA5542C9" pageId="10" pageNumber="11">
<paragraph id="C94595403F59E546708FC2BEFA554277" blockId="10.[151,1437,151,2021]" pageId="10" pageNumber="11">
<keyLead id="720B36753F59E546708FC2BEFA554277" pageId="10" pageNumber="11">
3(2) Base of head and labrum usually pale; male antennomeres 311 each with longitudinal row of about 10 erect setae along one side, setae about as long as width of antennomere, and antennomeres 37 parallel-sided; female ventrite V with narrowly notched apex; venter of prothorax partly dark brown (elytra usually spotted or at least partly black; only recorded from Timor in region, widespread in SE Asia)..........................................................
<taxonomicName id="0EFAEEC33F59E54674F0C116FA554277" authority="(Weber)" baseAuthorityName="Weber" baseAuthorityYear="1801" box="[1256,1436,591,613]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="FB8E49523F59E54674F0C116FA814276" box="[1256,1352,591,613]" italics="true" pageId="10" pageNumber="11">A. bicolor</emphasis>
(Weber)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C94595403F59E546708FC12BFA5542C9" blockId="10.[151,1437,151,2021]" pageId="10" pageNumber="11">
<keyLead id="720B36753F59E546708FC12BFA5542C9" pageId="10" pageNumber="11">
- Base of head and labrum dark brown to black; Male antennomeres 311 each with longitudinal row of 56 short erect setae, setae less than width of antennomere, and antennomeres 37 slightly expanded apically; female ventrite V with shallowly concave to truncate apical margin; venter of prothorax pale (not confirmed in region but likely to be adventive, widespread in SE Asia as far east as
<collectingCountry id="B1EDD5D03F59E546718CC183FE1842C9" box="[404,465,709,731]" name="Indonesia" pageId="10" pageNumber="11">Flores</collectingCountry>
)..............................................................
<taxonomicName id="0EFAEEC33F59E54674D0C183FA5542C9" authority="(Hornstedt)" baseAuthorityName="Hornstedt" baseAuthorityYear="1788" box="[1224,1436,709,731]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="coffeae">
<emphasis id="FB8E49523F59E54674D0C183FAEE42C8" box="[1224,1319,709,730]" italics="true" pageId="10" pageNumber="11">A. coffeae</emphasis>
(Hornstedt)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="720E8DE53F59E546708FC1A4FA55437F" pageId="10" pageNumber="11">
<paragraph id="C94595403F59E546708FC1A4FA554321" blockId="10.[151,1437,151,2021]" pageId="10" pageNumber="11">
<keyLead id="720B36753F59E546708FC1A4FA554321" pageId="10" pageNumber="11">
4(1) First antennomere enlarged apically, like an inverted elongate pyramid, with one side partly flattened (
<figureCitation id="51C189C53F59E54674AAC1A4FADF42EA" box="[1202,1302,738,760]" captionStart-0="FIGURES 25" captionStart-1="FIGURES 69" captionStart-2="FIGURES 1015" captionStart-3="FIGURES 1617" captionStartId-0="15.[151,264,1829,1853]" captionStartId-1="16.[151,264,1957,1981]" captionStartId-2="17.[151,264,1943,1967]" captionStartId-3="18.[151,264,783,807]" captionTargetBox-0="[164,1420,191,1786]" captionTargetBox-1="[174,1418,193,1917]" captionTargetBox-2="[209,1375,181,1906]" captionTargetBox-3="[158,1427,191,748]" captionTargetId-0="figure-19@15.[151,1436,181,1805]" captionTargetId-1="figure-22@16.[157,1430,181,1933]" captionTargetId-2="figure-19@17.[209,1379,181,1920]" captionTargetId-3="figure-22@18.[151,1435,181,759]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionTargetPageId-2="17" captionTargetPageId-3="18" captionText-0="FIGURES 25. Aulacophora species, male habitus: 2, A. abdominalis (Fabricius, 1781) (Guadalcanal); 3, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 4, A. cornuta Baly, 1879 (New Ireland); 5, A. indica (Gmelin, 1790) (Luzon)." captionText-1="FIGURES 69. Aulacophora species, male habitus: 6, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 7, A. relicta (Boisduval, 1835) (Mount Molloy); 8, A. wallacii Baly, 1886; 9, A. wilsoni Baly, 1888." captionText-2="FIGURES 1015. Aulacophora species, male head, anterior view: 10, A. abdominalis (Fabricius, 1781) (Guadalcanal); 11, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 12, A. cornuta Baly, 1879 (New Ireland); 13, A. indica (Gmelin, 1790) (Luzon); 14, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 15, A. relicta (Boisduval, 1835) (Mount Molloy)." captionText-3="FIGURES 1617. Aulacophora species, male head, anterior view: 16, A. wallacii Baly, 1886; 17, A. wilsoni Baly, 1888." figureDoi-0="http://doi.org/10.5281/zenodo.4545452" figureDoi-1="http://doi.org/10.5281/zenodo.4545458" figureDoi-2="http://doi.org/10.5281/zenodo.4545460" figureDoi-3="http://doi.org/10.5281/zenodo.4545462" httpUri-0="https://zenodo.org/record/4545452/files/figure.png" httpUri-1="https://zenodo.org/record/4545458/files/figure.png" httpUri-2="https://zenodo.org/record/4545460/files/figure.png" httpUri-3="https://zenodo.org/record/4545462/files/figure.png" pageId="10" pageNumber="11">Figs 217</figureCitation>
); pronotal lateral groove deep (
<figureCitation id="51C189C53F59E5467196C1B9FE1F4304" box="[398,470,767,790]" captionStart="FIGURES 2631" captionStartId="22.[151,264,1703,1727]" captionTargetBox="[160,1426,191,1670]" captionTargetId="figure-22@22.[151,1436,181,1679]" captionTargetPageId="22" captionText="FIGURES 2631. Aulacophora species, pronotum, lateral view: 26, male A. abdominalis (Fabricius, 1781) (Vanuatu); 27, male A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 28, male A. indica (Gmelin, 1790) (Borneo); 29, female A. indica; 30, male A. relicta (Boisduval, 1835); 31, male A. wallacii Baly, 1886." figureDoi="http://doi.org/10.5281/zenodo.4545466" httpUri="https://zenodo.org/record/4545466/files/figure.png" pageId="10" pageNumber="11">Fig. 28</figureCitation>
); apical ventrite deeply and elongately hollowed (
<figureCitation id="51C189C53F59E54673AAC1B9FBEA4304" box="[946,1059,767,790]" captionStart-0="FIGURES 3237" captionStart-1="FIGURES 3843" captionStartId-0="23.[151,264,1840,1864]" captionStartId-1="24.[151,264,1793,1817]" captionTargetBox-0="[167,1416,192,1802]" captionTargetBox-1="[168,1418,191,1757]" captionTargetId-0="figure-19@23.[151,1436,181,1816]" captionTargetId-1="figure-22@24.[151,1437,181,1769]" captionTargetPageId-0="23" captionTargetPageId-1="24" captionText-0="FIGURES 3237. Aulacophora species, male, apex of abdominal venter: 32, A. abdominalis (Fabricius, 1781) (Vanuatu); 33, A. abdominalis (New Ireland); 34, A. abdominalis (Manatuto); 35, A. abdominalis (Metinaro); 36, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 37, A. cornuta Baly, 1879 (Guadalcanal)." captionText-1="FIGURES 3843. Aulacophora species, male, apex of abdominal venter: 38, A. indica (Gmelin, 1790) (Sumatra); 39, A. indica (West Timor); 40, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 41, A. relicta (Boisduval, 1835) (Mount Molloy); 42, A. wallacii Baly, 1886; 43, A. wilsoni Baly, 1888." figureDoi-0="http://doi.org/10.5281/zenodo.4545468" figureDoi-1="http://doi.org/10.5281/zenodo.4545470" httpUri-0="https://zenodo.org/record/4545468/files/figure.png" httpUri-1="https://zenodo.org/record/4545470/files/figure.png" pageId="10" pageNumber="11">Figs 3243</figureCitation>
); basitarsomeres of anterior and middle legs slightly expanded, with pads of discoidal setae (males).................................................... 5
</keyLead>
</paragraph>
<paragraph id="C94595403F59E546708FC07DFA55437F" blockId="10.[151,1437,151,2021]" pageId="10" pageNumber="11">
<keyLead id="720B36753F59E546708FC07DFA55437F" pageId="10" pageNumber="11">
- First antennomere simple, fusiform; pronotal lateral groove shallow (
<figureCitation id="51C189C53F59E5467363C07CFC0A4342" box="[891,963,826,848]" captionStart="FIGURES 2631" captionStartId="22.[151,264,1703,1727]" captionTargetBox="[160,1426,191,1670]" captionTargetId="figure-22@22.[151,1436,181,1679]" captionTargetPageId="22" captionText="FIGURES 2631. Aulacophora species, pronotum, lateral view: 26, male A. abdominalis (Fabricius, 1781) (Vanuatu); 27, male A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 28, male A. indica (Gmelin, 1790) (Borneo); 29, female A. indica; 30, male A. relicta (Boisduval, 1835); 31, male A. wallacii Baly, 1886." figureDoi="http://doi.org/10.5281/zenodo.4545466" httpUri="https://zenodo.org/record/4545466/files/figure.png" pageId="10" pageNumber="11">Fig. 29</figureCitation>
); apical abdominal ventrite without elongate hollow but apex truncate to strongly notched (
<figureCitation id="51C189C53F59E5467265C011FD27437F" box="[637,750,855,877]" captionStart-0="FIGURES 4449" captionStart-1="FIGURES 5055" captionStart-2="FIGURES 5657" captionStartId-0="25.[151,264,1798,1822]" captionStartId-1="26.[151,264,1943,1967]" captionStartId-2="27.[151,264,701,725]" captionTargetBox-0="[170,1418,190,1763]" captionTargetBox-1="[198,1388,201,1907]" captionTargetBox-2="[165,1415,191,663]" captionTargetId-0="figure-19@25.[151,1436,181,1774]" captionTargetId-1="figure-22@26.[178,1409,181,1919]" captionTargetId-2="figure-19@27.[151,1436,181,677]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionTargetPageId-2="27" captionText-0="FIGURES 4449. Aulacophora species, female, apex of abdominal venter and pygidium: 44, A. abdominalis (Fabricius, 1781) (Vanuatu); 45, A. abdominalis (New Ireland); 46, A. abdominalis (Mer Island); 47, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 48, A. cornuta Baly, 1879 (Sulawesi); 49, A. indica (Gmelin, 1790) (Laos).Dotted lines indicate depth and width of apical concavity." captionText-1="FIGURES 5055. Aulacophora species, female, apex of abdominal venter: 50, A. indica (Gmelin, 1790) (Biak Island); 51, A. indica (Vietnam); 52, A. indica (Broken Hill); 53, A. mbabaram Reid, Halling &amp; Beatson, sp. nov., Almaden; 54, A. relicta (Boisduval, 1835); 55, A. wallacii Baly, 1886." captionText-2="FIGURES 5657. Aulacophora species, female, apex of abdominal venter: 56, A. wilsoni Baly, 1888 (Morgans Creek); 57, A. wilsoni (Eungella)." figureDoi-0="http://doi.org/10.5281/zenodo.4545472" figureDoi-1="http://doi.org/10.5281/zenodo.4545474" figureDoi-2="http://doi.org/10.5281/zenodo.4545476" httpUri-0="https://zenodo.org/record/4545472/files/figure.png" httpUri-1="https://zenodo.org/record/4545474/files/figure.png" httpUri-2="https://zenodo.org/record/4545476/files/figure.png" pageId="10" pageNumber="11">Figs 4457</figureCitation>
); basitarsomeres straight sided, without discoidal setae (females)... 12
</keyLead>
</paragraph>
</keyStep>
<keyStep id="720E8DE53F59E546708FC033FA5543F1" pageId="10" pageNumber="11">
<paragraph id="C94595403F59E546708FC033FA554399" blockId="10.[151,1437,151,2021]" box="[151,1436,885,907]" pageId="10" pageNumber="11">
<keyLead id="720B36753F59E546708FC033FA554399" box="[151,1436,885,907]" pageId="10" pageNumber="11">
5(4) Frontoclypeus with median ridge and without pits or paired lateral ridges (
<figureCitation id="51C189C53F59E54673B4C033FBA84399" box="[940,1121,885,907]" captionStart-0="FIGURES 1015" captionStart-1="FIGURES 1617" captionStartId-0="17.[151,264,1943,1967]" captionStartId-1="18.[151,264,783,807]" captionTargetBox-0="[209,1375,181,1906]" captionTargetBox-1="[158,1427,191,748]" captionTargetId-0="figure-19@17.[209,1379,181,1920]" captionTargetId-1="figure-22@18.[151,1435,181,759]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionText-0="FIGURES 1015. Aulacophora species, male head, anterior view: 10, A. abdominalis (Fabricius, 1781) (Guadalcanal); 11, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 12, A. cornuta Baly, 1879 (New Ireland); 13, A. indica (Gmelin, 1790) (Luzon); 14, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 15, A. relicta (Boisduval, 1835) (Mount Molloy)." captionText-1="FIGURES 1617. Aulacophora species, male head, anterior view: 16, A. wallacii Baly, 1886; 17, A. wilsoni Baly, 1888." figureDoi-0="http://doi.org/10.5281/zenodo.4545460" figureDoi-1="http://doi.org/10.5281/zenodo.4545462" httpUri-0="https://zenodo.org/record/4545460/files/figure.png" httpUri-1="https://zenodo.org/record/4545462/files/figure.png" pageId="10" pageNumber="11">Figs 1011, 1317</figureCitation>
)........................ 6
</keyLead>
</paragraph>
<paragraph id="C94595403F59E546708FC0D5FA5543F1" blockId="10.[151,1437,151,2021]" pageId="10" pageNumber="11">
<keyLead id="720B36753F59E546708FC0D5FA5543F1" pageId="10" pageNumber="11">
- Frontoclypeus without median ridge, with pair of arcuate ridges below antennal sockets and median patch of dense short golden setae above these (
<figureCitation id="51C189C53F59E546718EC0E9FE1743D4" box="[406,478,943,966]" captionStart="FIGURES 1015" captionStartId="17.[151,264,1943,1967]" captionTargetBox="[209,1375,181,1906]" captionTargetId="figure-19@17.[209,1379,181,1920]" captionTargetPageId="17" captionText="FIGURES 1015. Aulacophora species, male head, anterior view: 10, A. abdominalis (Fabricius, 1781) (Guadalcanal); 11, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 12, A. cornuta Baly, 1879 (New Ireland); 13, A. indica (Gmelin, 1790) (Luzon); 14, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 15, A. relicta (Boisduval, 1835) (Mount Molloy)." figureDoi="http://doi.org/10.5281/zenodo.4545460" httpUri="https://zenodo.org/record/4545460/files/figure.png" pageId="10" pageNumber="11">Fig. 12</figureCitation>
); (length
<quantity id="0E0238A53F59E5467222C0E9FD7843D7" box="[570,689,943,965]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.5" metricValueMin="7.5" pageId="10" pageNumber="11" unit="mm" value="8.5" valueMax="9.5" valueMin="7.5">7.59.5 mm</quantity>
; elytral humeri without setae (
<figureCitation id="51C189C53F59E54673CDC0E9FBD743D4" box="[981,1054,943,966]" captionStart="FIGURES 1825" captionStartId="21.[151,264,1899,1923]" captionTargetBox="[262,1328,190,1869]" captionTargetId="figure-19@21.[250,1338,181,1875]" captionTargetPageId="21" captionText="FIGURES 1825. Aulacophora species, male anterior and antenna: 18, A. abdominalis (Fabricius, 1781) (Guadalcanal); 19, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 20, A. cornuta Baly, 1879 (New Ireland); 21, A. indica (Gmelin, 1790) (Luzon); 22, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 23, A. relicta (Boisduval, 1835) (Mount Molloy); 24, A. wallacii Baly, 1886; 25, A. wilsoni Baly, 1888." figureDoi="http://doi.org/10.5281/zenodo.4545464" httpUri="https://zenodo.org/record/4545464/files/figure.png" pageId="10" pageNumber="11">Fig. 20</figureCitation>
); penis symmetrical, sagittate (
<figureCitation id="51C189C53F59E5467550C0E9FA5843D4" box="[1352,1425,943,966]" captionStart="FIGURES 98101" captionStartId="45.[151,264,1825,1849]" captionTargetBox="[164,1393,195,1754]" captionTargetId="figure-19@45.[151,1436,181,1801]" captionTargetPageId="45" captionText="FIGURES 98101. Aulacophora species, penis, dorsal view: 98, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 99, A. cornuta Baly, 1879 (Guadalcanal); 100, A. indica (Gmelin, 1790) (Laos); 101, A. indica (Luzon)." figureDoi="http://doi.org/10.5281/zenodo.4545490" httpUri="https://zenodo.org/record/4545490/files/figure.png" pageId="10" pageNumber="11">Fig. 99</figureCitation>
); SE Asia to
<collectingCountry id="B1EDD5D03F59E5467155C08BFE2643F1" box="[333,495,973,995]" name="Solomon Islands" pageId="10" pageNumber="11">Solomon Islands</collectingCountry>
)................................................................
<taxonomicName id="0EFAEEC33F59E546751CC08BFA5543F1" authority="Baly" authorityName="Baly" authorityYear="1879" box="[1284,1436,973,995]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="FB8E49523F59E546751CC08BFAA043F0" box="[1284,1385,973,994]" italics="true" pageId="10" pageNumber="11">A. cornuta</emphasis>
Baly
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="720E8DE53F59E546708FC0ACFA5544A2" pageId="10" pageNumber="11">
<paragraph id="C94595403F59E546708FC0ACFA554412" blockId="10.[151,1437,151,2021]" box="[151,1436,1002,1024]" pageId="10" pageNumber="11">
<keyLead id="720B36753F59E546708FC0ACFA554412" box="[151,1436,1002,1024]" pageId="10" pageNumber="11">
6(5) Elytral humeri with oval patch of erect setae (
<figureCitation id="51C189C53F59E5467280C0ACFC5E4412" box="[664,919,1002,1024]" captionStart="FIGURES 1825" captionStartId="21.[151,264,1899,1923]" captionTargetBox="[262,1328,190,1869]" captionTargetId="figure-19@21.[250,1338,181,1875]" captionTargetPageId="21" captionText="FIGURES 1825. Aulacophora species, male anterior and antenna: 18, A. abdominalis (Fabricius, 1781) (Guadalcanal); 19, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 20, A. cornuta Baly, 1879 (New Ireland); 21, A. indica (Gmelin, 1790) (Luzon); 22, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 23, A. relicta (Boisduval, 1835) (Mount Molloy); 24, A. wallacii Baly, 1886; 25, A. wilsoni Baly, 1888." figureDoi="http://doi.org/10.5281/zenodo.4545464" httpUri="https://zenodo.org/record/4545464/files/figure.png" pageId="10" pageNumber="11">Figs 1819, 2122, 2425</figureCitation>
)......................................... 7
</keyLead>
</paragraph>
<paragraph id="C94595403F59E546708FC74EFA5544A2" blockId="10.[151,1437,151,2021]" pageId="10" pageNumber="11">
<keyLead id="720B36753F59E546708FC74EFA5544A2" pageId="10" pageNumber="11">
- Elytral humeri glabrous (
<figureCitation id="51C189C53F59E54671CDC741FDD7440C" box="[469,542,1031,1054]" captionStart="FIGURES 1825" captionStartId="21.[151,264,1899,1923]" captionTargetBox="[262,1328,190,1869]" captionTargetId="figure-19@21.[250,1338,181,1875]" captionTargetPageId="21" captionText="FIGURES 1825. Aulacophora species, male anterior and antenna: 18, A. abdominalis (Fabricius, 1781) (Guadalcanal); 19, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 20, A. cornuta Baly, 1879 (New Ireland); 21, A. indica (Gmelin, 1790) (Luzon); 22, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 23, A. relicta (Boisduval, 1835) (Mount Molloy); 24, A. wallacii Baly, 1886; 25, A. wilsoni Baly, 1888." figureDoi="http://doi.org/10.5281/zenodo.4545464" httpUri="https://zenodo.org/record/4545464/files/figure.png" pageId="10" pageNumber="11">Fig. 23</figureCitation>
); (length
<quantity id="0E0238A53F59E5467263C741FD29440F" box="[635,736,1031,1053]" metricMagnitude="-3" metricUnit="m" metricValue="6.75" metricValueMax="7.5" metricValueMin="6.0" pageId="10" pageNumber="11" unit="mm" value="6.75" valueMax="7.5" valueMin="6.0">67.5 mm</quantity>
; femora pale, with meso- and metatibiae darkened, or legs entirely pale; apical segments of antennae usually slightly darkened; pygidium entirely brownish-yellow; pronotum with pair of large pits anterior to transverse groove, anterior half of pronotum more convex than posterior half, pronotal depression in lateral view with anterior slope steeper than posterior (
<figureCitation id="51C189C53F59E5467264C719FD0C4464" box="[636,709,1119,1142]" captionStart="FIGURES 2631" captionStartId="22.[151,264,1703,1727]" captionTargetBox="[160,1426,191,1670]" captionTargetId="figure-22@22.[151,1436,181,1679]" captionTargetPageId="22" captionText="FIGURES 2631. Aulacophora species, pronotum, lateral view: 26, male A. abdominalis (Fabricius, 1781) (Vanuatu); 27, male A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 28, male A. indica (Gmelin, 1790) (Borneo); 29, female A. indica; 30, male A. relicta (Boisduval, 1835); 31, male A. wallacii Baly, 1886." figureDoi="http://doi.org/10.5281/zenodo.4545466" httpUri="https://zenodo.org/record/4545466/files/figure.png" pageId="10" pageNumber="11">Fig. 30</figureCitation>
); last ventrite with apical excavation asymmetric, deeper and more sharply bordered on left (
<figureCitation id="51C189C53F59E5467190C73BFE194481" box="[392,464,1149,1171]" captionStart="FIGURES 3843" captionStartId="24.[151,264,1793,1817]" captionTargetBox="[168,1418,191,1757]" captionTargetId="figure-22@24.[151,1437,181,1769]" captionTargetPageId="24" captionText="FIGURES 3843. Aulacophora species, male, apex of abdominal venter: 38, A. indica (Gmelin, 1790) (Sumatra); 39, A. indica (West Timor); 40, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 41, A. relicta (Boisduval, 1835) (Mount Molloy); 42, A. wallacii Baly, 1886; 43, A. wilsoni Baly, 1888." figureDoi="http://doi.org/10.5281/zenodo.4545470" httpUri="https://zenodo.org/record/4545470/files/figure.png" pageId="10" pageNumber="11">Fig. 41</figureCitation>
); penis in dorsal view slightly constricted at middle (
<figureCitation id="51C189C53F59E54673D4C73BFB9C4481" box="[972,1109,1149,1171]" captionStart="FIGURES 102105" captionStartId="46.[151,264,1777,1801]" captionTargetBox="[166,1388,194,1718]" captionTargetId="figure-22@46.[151,1436,181,1754]" captionTargetPageId="46" captionText="FIGURES 102105. Aulacophora species, penis, dorsal view: 102, A. indica (Gmelin, 1790) (Timor); 103, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 104, A. relicta (Boisduval, 1835) (Mount Molloy); 105, A. relicta (Mount Mulligan)." figureDoi="http://doi.org/10.5281/zenodo.4545492" httpUri="https://zenodo.org/record/4545492/files/figure.png" pageId="10" pageNumber="11">Figs 104105</figureCitation>
);
<collectingCountry id="B1EDD5D03F59E546747CC73BFB774481" box="[1124,1214,1149,1171]" name="Australia" pageId="10" pageNumber="11">Australia</collectingCountry>
).....................................................................................................
<taxonomicName id="0EFAEEC33F59E54674C4C7DCFA5544A2" authority="Boisduval" authorityName="Boisduval" authorityYear="1835" box="[1244,1436,1178,1200]" class="Insecta" family="Chrysomelidae" genus="Galleruca" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="relicta">
<emphasis id="FB8E49523F59E54674C4C7DCFAFC44BD" box="[1244,1333,1178,1199]" italics="true" pageId="10" pageNumber="11">A. relicta</emphasis>
Boisduval
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</paragraph>
</keyStep>
<keyStep id="720E8DE53F59E546708FC7F1FA5545A5" pageId="10" pageNumber="11">
<paragraph id="C94595403F59E546708FC7F1FA554537" blockId="10.[151,1437,151,2021]" pageId="10" pageNumber="11">
<keyLead id="720B36753F59E546708FC7F1FA554537" pageId="10" pageNumber="11">
7(6) Meso- and metafemora usually mostly darkened, but if femora mostly to entirely pale, meso- and metatibiae also mostly to entirely pale (
<figureCitation id="51C189C53F59E546717DC793FE6944F8" box="[357,416,1237,1259]" captionStart="FIGURES 69" captionStartId="16.[151,264,1957,1981]" captionTargetBox="[174,1418,193,1917]" captionTargetId="figure-22@16.[157,1430,181,1933]" captionTargetPageId="16" captionText="FIGURES 69. Aulacophora species, male habitus: 6, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 7, A. relicta (Boisduval, 1835) (Mount Molloy); 8, A. wallacii Baly, 1886; 9, A. wilsoni Baly, 1888." figureDoi="http://doi.org/10.5281/zenodo.4545458" httpUri="https://zenodo.org/record/4545458/files/figure.png" pageId="10" pageNumber="11">Fig. 7</figureCitation>
); groove in last ventrite usually assymmetric, with swelling at one side and/or deepened groove at one side (
<figureCitation id="51C189C53F59E54670F1C7B4FE90451A" box="[233,345,1266,1288]" captionStart="FIGURES 3237" captionStartId="23.[151,264,1840,1864]" captionTargetBox="[167,1416,192,1802]" captionTargetId="figure-19@23.[151,1436,181,1816]" captionTargetPageId="23" captionText="FIGURES 3237. Aulacophora species, male, apex of abdominal venter: 32, A. abdominalis (Fabricius, 1781) (Vanuatu); 33, A. abdominalis (New Ireland); 34, A. abdominalis (Manatuto); 35, A. abdominalis (Metinaro); 36, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 37, A. cornuta Baly, 1879 (Guadalcanal)." figureDoi="http://doi.org/10.5281/zenodo.4545468" httpUri="https://zenodo.org/record/4545468/files/figure.png" pageId="10" pageNumber="11">Figs 3235</figureCitation>
); pronotal groove narrower at middle (
<figureCitation id="51C189C53F59E54672DFC7B4FCAC451A" box="[711,869,1266,1288]" captionStart-0="FIGURES 25" captionStart-1="FIGURES 69" captionStartId-0="15.[151,264,1829,1853]" captionStartId-1="16.[151,264,1957,1981]" captionTargetBox-0="[164,1420,191,1786]" captionTargetBox-1="[174,1418,193,1917]" captionTargetId-0="figure-19@15.[151,1436,181,1805]" captionTargetId-1="figure-22@16.[157,1430,181,1933]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionText-0="FIGURES 25. Aulacophora species, male habitus: 2, A. abdominalis (Fabricius, 1781) (Guadalcanal); 3, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 4, A. cornuta Baly, 1879 (New Ireland); 5, A. indica (Gmelin, 1790) (Luzon)." captionText-1="FIGURES 69. Aulacophora species, male habitus: 6, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 7, A. relicta (Boisduval, 1835) (Mount Molloy); 8, A. wallacii Baly, 1886; 9, A. wilsoni Baly, 1888." figureDoi-0="http://doi.org/10.5281/zenodo.4545452" figureDoi-1="http://doi.org/10.5281/zenodo.4545458" httpUri-0="https://zenodo.org/record/4545452/files/figure.png" httpUri-1="https://zenodo.org/record/4545458/files/figure.png" pageId="10" pageNumber="11">Figs 2, 56, 89</figureCitation>
); penis without median tooth in lateral view (
<figureCitation id="51C189C53F59E546750AC7B4FA514515" box="[1298,1432,1266,1288]" captionStart="FIGURES 121124" captionStartId="52.[151,264,1872,1896]" captionTargetBox="[166,1410,190,1822]" captionTargetId="figure-22@52.[151,1436,181,1848]" captionTargetPageId="52" captionText="FIGURES 121124. Aulacophora abdominalis (Fabricius, 1781), penis, lateral view: 121, Guadalcanal, 122, Metinaro, 123, Saibai, 124, Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.4545502" httpUri="https://zenodo.org/record/4545502/files/figure.png" pageId="10" pageNumber="11">Figs 121124</figureCitation>
,
<figureCitation id="51C189C53F59E54670FAC656FEF14537" box="[226,312,1296,1317]" captionStart-0="FIGURES 125128" captionStart-1="FIGURES 129132" captionStartId-0="53.[151,264,1722,1746]" captionStartId-1="54.[151,264,1767,1791]" captionTargetBox-0="[165,1400,194,1671]" captionTargetBox-1="[167,1400,192,1716]" captionTargetId-0="figure-19@53.[151,1436,181,1699]" captionTargetId-1="figure-22@54.[151,1436,181,1743]" captionTargetPageId-0="53" captionTargetPageId-1="54" captionText-0="FIGURES 125128. Aulacophora species, penis, lateral view: 125, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 126, A. cornuta Baly, 1879; 127, A. indica (Gmelin, 1790) (Laos); 128, A. indica (Luzon). Arrow indicates small spine on dorsal surface." captionText-1="FIGURES 129132. Aulacophora species, penis, lateral view: 129, A. indica (Gmelin, 1790) (W Timor); 130, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 131, A. relicta (Boisduval, 1835) (Mount Molloy); 132, A. relicta (Mount Mulligan)." figureDoi-0="http://doi.org/10.5281/zenodo.4545504" figureDoi-1="http://doi.org/10.5281/zenodo.4545506" httpUri-0="https://zenodo.org/record/4545504/files/figure.png" httpUri-1="https://zenodo.org/record/4545506/files/figure.png" pageId="10" pageNumber="11">127130</figureCitation>
,
<figureCitation id="51C189C53F59E546715AC656FE5E4534" box="[322,407,1296,1318]" captionStart="FIGURES 133134" captionStartId="55.[151,264,1996,2020]" captionTargetBox="[288,1299,196,1952]" captionTargetId="figure-19@55.[271,1317,181,1973]" captionTargetPageId="55" captionText="FIGURES 133134. Aulacophora species, penis, lateral view: 133, A. wallacii Baly, 1886; 134, A. wilsoni Baly, 1888." figureDoi="http://doi.org/10.5281/zenodo.4545508" httpUri="https://zenodo.org/record/4545508/files/figure.png" pageId="10" pageNumber="11">133134</figureCitation>
).................................................................................... 8
</keyLead>
</paragraph>
<paragraph id="C94595403F59E546708FC66BFA5545A5" blockId="10.[151,1437,151,2021]" pageId="10" pageNumber="11">
<keyLead id="720B36753F59E546708FC66BFA5545A5" pageId="10" pageNumber="11">
- Femora pale except meso- and metafemora with dark apices, meso- and metatibiae dark brown to black; apical groove on ventrite 5 symmetrical (
<figureCitation id="51C189C53F59E54671D1C60CFDD84572" box="[457,529,1354,1376]" captionStart="FIGURES 3237" captionStartId="23.[151,264,1840,1864]" captionTargetBox="[167,1416,192,1802]" captionTargetId="figure-19@23.[151,1436,181,1816]" captionTargetPageId="23" captionText="FIGURES 3237. Aulacophora species, male, apex of abdominal venter: 32, A. abdominalis (Fabricius, 1781) (Vanuatu); 33, A. abdominalis (New Ireland); 34, A. abdominalis (Manatuto); 35, A. abdominalis (Metinaro); 36, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 37, A. cornuta Baly, 1879 (Guadalcanal)." figureDoi="http://doi.org/10.5281/zenodo.4545468" httpUri="https://zenodo.org/record/4545468/files/figure.png" pageId="10" pageNumber="11">Fig. 36</figureCitation>
); pronotal groove greatly expanded at middle with well-defined sinuate anterior and posterior edges (
<figureCitation id="51C189C53F59E546713DC621FEAA456F" box="[293,355,1383,1405]" captionStart="FIGURES 25" captionStartId="15.[151,264,1829,1853]" captionTargetBox="[164,1420,191,1786]" captionTargetId="figure-19@15.[151,1436,181,1805]" captionTargetPageId="15" captionText="FIGURES 25. Aulacophora species, male habitus: 2, A. abdominalis (Fabricius, 1781) (Guadalcanal); 3, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 4, A. cornuta Baly, 1879 (New Ireland); 5, A. indica (Gmelin, 1790) (Luzon)." figureDoi="http://doi.org/10.5281/zenodo.4545452" httpUri="https://zenodo.org/record/4545452/files/figure.png" pageId="10" pageNumber="11">Figs 3</figureCitation>
,
<figureCitation id="51C189C53F59E5467177C62EFE40456C" box="[367,393,1384,1406]" captionStart="FIGURES 1825" captionStartId="21.[151,264,1899,1923]" captionTargetBox="[262,1328,190,1869]" captionTargetId="figure-19@21.[250,1338,181,1875]" captionTargetPageId="21" captionText="FIGURES 1825. Aulacophora species, male anterior and antenna: 18, A. abdominalis (Fabricius, 1781) (Guadalcanal); 19, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 20, A. cornuta Baly, 1879 (New Ireland); 21, A. indica (Gmelin, 1790) (Luzon); 22, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 23, A. relicta (Boisduval, 1835) (Mount Molloy); 24, A. wallacii Baly, 1886; 25, A. wilsoni Baly, 1888." figureDoi="http://doi.org/10.5281/zenodo.4545464" httpUri="https://zenodo.org/record/4545464/files/figure.png" pageId="10" pageNumber="11">19</figureCitation>
); penis with minute median tooth in lateral view (
<figureCitation id="51C189C53F59E5467372C621FC74456F" box="[874,957,1383,1405]" captionStart="FIGURES 125128" captionStartId="53.[151,264,1722,1746]" captionTargetBox="[165,1400,194,1671]" captionTargetId="figure-19@53.[151,1436,181,1699]" captionTargetPageId="53" captionText="FIGURES 125128. Aulacophora species, penis, lateral view: 125, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 126, A. cornuta Baly, 1879; 127, A. indica (Gmelin, 1790) (Laos); 128, A. indica (Luzon). Arrow indicates small spine on dorsal surface." figureDoi="http://doi.org/10.5281/zenodo.4545504" httpUri="https://zenodo.org/record/4545504/files/figure.png" pageId="10" pageNumber="11">Fig. 125</figureCitation>
) (length
<quantity id="0E0238A53F59E546740CC621FBB0456F" box="[1044,1145,1383,1405]" metricMagnitude="-3" metricUnit="m" metricValue="6.75" metricValueMax="7.0" metricValueMin="6.5" pageId="10" pageNumber="11" unit="mm" value="6.75" valueMax="7.0" valueMin="6.5">6.57 mm</quantity>
; tergite 8 medially lobed (
<figureCitation id="51C189C53F59E546756DC621FF344589" captionStart="FIGURES 8388" captionStartId="42.[151,264,1943,1967]" captionTargetBox="[217,1368,192,1905]" captionTargetId="figure-22@42.[204,1384,181,1919]" captionTargetPageId="42" captionText="FIGURES 8388. Aulacophora species, male sternite 8, ventrolateral view: 83, A. abdominalis (Fabricius, 1781) (Vanuatu); 84, A. abdominalis (New Ireland); 85, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 86, A. cornuta Baly, 1879 (Guadalcanal); 87, A. indica (Gmelin, 1790) (Timor); 88, A. indica (Luzon)." figureDoi="http://doi.org/10.5281/zenodo.4545484" httpUri="https://zenodo.org/record/4545484/files/figure.png" pageId="10" pageNumber="11">Fig. 85</figureCitation>
); pronotum without pits anterior to median groove, in lateral view groove with posterior angle greater than curving anterior angle (
<figureCitation id="51C189C53F59E546713BC6E4FEA245AA" box="[291,363,1442,1464]" captionStart="FIGURES 2631" captionStartId="22.[151,264,1703,1727]" captionTargetBox="[160,1426,191,1670]" captionTargetId="figure-22@22.[151,1436,181,1679]" captionTargetPageId="22" captionText="FIGURES 2631. Aulacophora species, pronotum, lateral view: 26, male A. abdominalis (Fabricius, 1781) (Vanuatu); 27, male A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 28, male A. indica (Gmelin, 1790) (Borneo); 29, female A. indica; 30, male A. relicta (Boisduval, 1835); 31, male A. wallacii Baly, 1886." figureDoi="http://doi.org/10.5281/zenodo.4545466" httpUri="https://zenodo.org/record/4545466/files/figure.png" pageId="10" pageNumber="11">Fig. 27</figureCitation>
); coastal NT and WA)..................................
<taxonomicName id="0EFAEEC33F59E54673C5C6E4FA8F45AA" authority="Reid, Halling &amp; Beatson" authorityName="Reid, Halling &amp; Beatson" authorityYear="2021" box="[989,1350,1442,1464]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="barrogae" status="sp. nov.">
<emphasis id="FB8E49523F59E54673C5C6E4FB9945A5" box="[989,1104,1442,1463]" italics="true" pageId="10" pageNumber="11">A. barrogae</emphasis>
Reid, Halling &amp; Beatson
</taxonomicName>
,
<emphasis id="FB8E49523F59E5467548C6E4FA5545A5" bold="true" box="[1360,1436,1442,1463]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="E0BDF4293F59E5467548C6E4FA5545A5" box="[1360,1436,1442,1463]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
</keyStep>
<keyStep id="720E8DE53F59E546708FC686FA5546B1" pageId="10" pageNumber="11">
<paragraph id="C94595403F59E546708FC686FA55461D" blockId="10.[151,1437,151,2021]" pageId="10" pageNumber="11">
<keyLead id="720B36753F59E546708FC686FA55461D" pageId="10" pageNumber="11">
8(7) Abdomen with at least apices of both pygidium and ventrite V pale (
<figureCitation id="51C189C53F59E5467366C6F9FC2745C7" box="[894,1006,1471,1493]" captionStart="FIGURES 3237" captionStartId="23.[151,264,1840,1864]" captionTargetBox="[167,1416,192,1802]" captionTargetId="figure-19@23.[151,1436,181,1816]" captionTargetPageId="23" captionText="FIGURES 3237. Aulacophora species, male, apex of abdominal venter: 32, A. abdominalis (Fabricius, 1781) (Vanuatu); 33, A. abdominalis (New Ireland); 34, A. abdominalis (Manatuto); 35, A. abdominalis (Metinaro); 36, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 37, A. cornuta Baly, 1879 (Guadalcanal)." figureDoi="http://doi.org/10.5281/zenodo.4545468" httpUri="https://zenodo.org/record/4545468/files/figure.png" pageId="10" pageNumber="11">Figs 3235</figureCitation>
,
<figureCitation id="51C189C53F59E54673E1C686FB9445C4" box="[1017,1117,1472,1494]" captionStart="FIGURES 3843" captionStartId="24.[151,264,1793,1817]" captionTargetBox="[168,1418,191,1757]" captionTargetId="figure-22@24.[151,1437,181,1769]" captionTargetPageId="24" captionText="FIGURES 3843. Aulacophora species, male, apex of abdominal venter: 38, A. indica (Gmelin, 1790) (Sumatra); 39, A. indica (West Timor); 40, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 41, A. relicta (Boisduval, 1835) (Mount Molloy); 42, A. wallacii Baly, 1886; 43, A. wilsoni Baly, 1888." figureDoi="http://doi.org/10.5281/zenodo.4545470" httpUri="https://zenodo.org/record/4545470/files/figure.png" pageId="10" pageNumber="11">3840, 42</figureCitation>
); antennomeres longer, length of 5th more than 2.3x width (
<figureCitation id="51C189C53F59E54671FFC69BFD6A45E1" box="[487,675,1501,1523]" captionStart="FIGURES 1825" captionStartId="21.[151,264,1899,1923]" captionTargetBox="[262,1328,190,1869]" captionTargetId="figure-19@21.[250,1338,181,1875]" captionTargetPageId="21" captionText="FIGURES 1825. Aulacophora species, male anterior and antenna: 18, A. abdominalis (Fabricius, 1781) (Guadalcanal); 19, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 20, A. cornuta Baly, 1879 (New Ireland); 21, A. indica (Gmelin, 1790) (Luzon); 22, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 23, A. relicta (Boisduval, 1835) (Mount Molloy); 24, A. wallacii Baly, 1886; 25, A. wilsoni Baly, 1888." figureDoi="http://doi.org/10.5281/zenodo.4545464" httpUri="https://zenodo.org/record/4545464/files/figure.png" pageId="10" pageNumber="11">Figs 18, 2122, 24</figureCitation>
); apex of penis usually hooked in lateral view (
<figureCitation id="51C189C53F59E546746DC69BFB3445E0" box="[1141,1277,1501,1523]" captionStart="FIGURES 121124" captionStartId="52.[151,264,1872,1896]" captionTargetBox="[166,1410,190,1822]" captionTargetId="figure-22@52.[151,1436,181,1848]" captionTargetPageId="52" captionText="FIGURES 121124. Aulacophora abdominalis (Fabricius, 1781), penis, lateral view: 121, Guadalcanal, 122, Metinaro, 123, Saibai, 124, Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.4545502" httpUri="https://zenodo.org/record/4545502/files/figure.png" pageId="10" pageNumber="11">Figs 121124</figureCitation>
,
<figureCitation id="51C189C53F59E5467510C69BFA9645E0" box="[1288,1375,1501,1522]" captionStart-0="FIGURES 125128" captionStart-1="FIGURES 129132" captionStartId-0="53.[151,264,1722,1746]" captionStartId-1="54.[151,264,1767,1791]" captionTargetBox-0="[165,1400,194,1671]" captionTargetBox-1="[167,1400,192,1716]" captionTargetId-0="figure-19@53.[151,1436,181,1699]" captionTargetId-1="figure-22@54.[151,1436,181,1743]" captionTargetPageId-0="53" captionTargetPageId-1="54" captionText-0="FIGURES 125128. Aulacophora species, penis, lateral view: 125, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 126, A. cornuta Baly, 1879; 127, A. indica (Gmelin, 1790) (Laos); 128, A. indica (Luzon). Arrow indicates small spine on dorsal surface." captionText-1="FIGURES 129132. Aulacophora species, penis, lateral view: 129, A. indica (Gmelin, 1790) (W Timor); 130, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 131, A. relicta (Boisduval, 1835) (Mount Molloy); 132, A. relicta (Mount Mulligan)." figureDoi-0="http://doi.org/10.5281/zenodo.4545504" figureDoi-1="http://doi.org/10.5281/zenodo.4545506" httpUri-0="https://zenodo.org/record/4545504/files/figure.png" httpUri-1="https://zenodo.org/record/4545506/files/figure.png" pageId="10" pageNumber="11">127130</figureCitation>
,
<figureCitation id="51C189C53F59E5467572C69BFA5845E1" box="[1386,1425,1501,1523]" captionStart="FIGURES 133134" captionStartId="55.[151,264,1996,2020]" captionTargetBox="[288,1299,196,1952]" captionTargetId="figure-19@55.[271,1317,181,1973]" captionTargetPageId="55" captionText="FIGURES 133134. Aulacophora species, penis, lateral view: 133, A. wallacii Baly, 1886; 134, A. wilsoni Baly, 1888." figureDoi="http://doi.org/10.5281/zenodo.4545508" httpUri="https://zenodo.org/record/4545508/files/figure.png" pageId="10" pageNumber="11">133</figureCitation>
); scutellum usually same colour as elytra, rarely darker brown (
<figureCitation id="51C189C53F59E546732BC6BCFC404602" box="[819,905,1530,1552]" captionStart="FIGURES 25" captionStartId="15.[151,264,1829,1853]" captionTargetBox="[164,1420,191,1786]" captionTargetId="figure-19@15.[151,1436,181,1805]" captionTargetPageId="15" captionText="FIGURES 25. Aulacophora species, male habitus: 2, A. abdominalis (Fabricius, 1781) (Guadalcanal); 3, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 4, A. cornuta Baly, 1879 (New Ireland); 5, A. indica (Gmelin, 1790) (Luzon)." figureDoi="http://doi.org/10.5281/zenodo.4545452" httpUri="https://zenodo.org/record/4545452/files/figure.png" pageId="10" pageNumber="11">Figs 2, 5</figureCitation>
,
<figureCitation id="51C189C53F59E546738CC6BCFC694602" box="[916,928,1530,1552]" captionStart="FIGURES 69" captionStartId="16.[151,264,1957,1981]" captionTargetBox="[174,1418,193,1917]" captionTargetId="figure-22@16.[157,1430,181,1933]" captionTargetPageId="16" captionText="FIGURES 69. Aulacophora species, male habitus: 6, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 7, A. relicta (Boisduval, 1835) (Mount Molloy); 8, A. wallacii Baly, 1886; 9, A. wilsoni Baly, 1888." figureDoi="http://doi.org/10.5281/zenodo.4545458" httpUri="https://zenodo.org/record/4545458/files/figure.png" pageId="10" pageNumber="11">8</figureCitation>
) (mesepimeron usually pale)................. 9
</keyLead>
</paragraph>
<paragraph id="C94595403F59E546708FC55EFA5546B1" blockId="10.[151,1437,151,2021]" pageId="10" pageNumber="11">
<keyLead id="720B36753F59E546708FC55EFA5546B1" pageId="10" pageNumber="11">
- Abdomen entirely black (
<figureCitation id="51C189C53F59E54671CEC551FDD7463C" box="[470,542,1559,1582]" captionStart="FIGURES 3843" captionStartId="24.[151,264,1793,1817]" captionTargetBox="[168,1418,191,1757]" captionTargetId="figure-22@24.[151,1437,181,1769]" captionTargetPageId="24" captionText="FIGURES 3843. Aulacophora species, male, apex of abdominal venter: 38, A. indica (Gmelin, 1790) (Sumatra); 39, A. indica (West Timor); 40, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 41, A. relicta (Boisduval, 1835) (Mount Molloy); 42, A. wallacii Baly, 1886; 43, A. wilsoni Baly, 1888." figureDoi="http://doi.org/10.5281/zenodo.4545470" httpUri="https://zenodo.org/record/4545470/files/figure.png" pageId="10" pageNumber="11">Fig. 43</figureCitation>
); antennomeres shorter, length of 5th about 2.2x width (
<figureCitation id="51C189C53F59E546742FC551FBB7463F" box="[1079,1150,1559,1581]" captionStart="FIGURES 1825" captionStartId="21.[151,264,1899,1923]" captionTargetBox="[262,1328,190,1869]" captionTargetId="figure-19@21.[250,1338,181,1875]" captionTargetPageId="21" captionText="FIGURES 1825. Aulacophora species, male anterior and antenna: 18, A. abdominalis (Fabricius, 1781) (Guadalcanal); 19, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 20, A. cornuta Baly, 1879 (New Ireland); 21, A. indica (Gmelin, 1790) (Luzon); 22, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 23, A. relicta (Boisduval, 1835) (Mount Molloy); 24, A. wallacii Baly, 1886; 25, A. wilsoni Baly, 1888." figureDoi="http://doi.org/10.5281/zenodo.4545464" httpUri="https://zenodo.org/record/4545464/files/figure.png" pageId="10" pageNumber="11">Fig. 25</figureCitation>
); penis strongly curved in lateral view, without hooked tip (
<figureCitation id="51C189C53F59E546721AC573FD9D4659" box="[514,596,1589,1611]" captionStart="FIGURES 133134" captionStartId="55.[151,264,1996,2020]" captionTargetBox="[288,1299,196,1952]" captionTargetId="figure-19@55.[271,1317,181,1973]" captionTargetPageId="55" captionText="FIGURES 133134. Aulacophora species, penis, lateral view: 133, A. wallacii Baly, 1886; 134, A. wilsoni Baly, 1888." figureDoi="http://doi.org/10.5281/zenodo.4545508" httpUri="https://zenodo.org/record/4545508/files/figure.png" pageId="10" pageNumber="11">Fig. 134</figureCitation>
); scutellum always much darker than elytra, brown to almost black (
<figureCitation id="51C189C53F59E54674C1C573FADB4659" box="[1241,1298,1589,1611]" captionStart="FIGURES 69" captionStartId="16.[151,264,1957,1981]" captionTargetBox="[174,1418,193,1917]" captionTargetId="figure-22@16.[157,1430,181,1933]" captionTargetPageId="16" captionText="FIGURES 69. Aulacophora species, male habitus: 6, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 7, A. relicta (Boisduval, 1835) (Mount Molloy); 8, A. wallacii Baly, 1886; 9, A. wilsoni Baly, 1888." figureDoi="http://doi.org/10.5281/zenodo.4545458" httpUri="https://zenodo.org/record/4545458/files/figure.png" pageId="10" pageNumber="11">Fig. 9</figureCitation>
) (length
<quantity id="0E0238A53F59E546757EC573FEC3467A" metricMagnitude="-3" metricUnit="m" metricValue="6.75" metricValueMax="7.0" metricValueMin="6.5" pageId="10" pageNumber="11" unit="mm" value="6.75" valueMax="7.0" valueMin="6.5">6.57 mm</quantity>
; mesepimeron black or dark brown; meso- and metatibiae black and anterior tibiae with black outer edges; antennomeres 411 black; last ventrite with apical excavation slightly asymmetric; tergite 8 medially lobed (
<figureCitation id="51C189C53F59E5467441C529FB564694" box="[1113,1183,1647,1670]" captionStart="FIGURES 8993" captionStartId="43.[151,264,1944,1968]" captionTargetBox="[196,1391,191,1907]" captionTargetId="figure-19@43.[177,1410,181,1920]" captionTargetPageId="43" captionText="FIGURES 8993. Aulacophora species, male sternite 8, ventrolateral view: 89, A. indica (Gmelin, 1790) (Sumatra); 90, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 91, A. relicta (Boisduval, 1835) (Coonabarabran); 92, A. wallacii Baly, 1886; 93, A. wilsoni Baly, 1888." figureDoi="http://doi.org/10.5281/zenodo.4545486" httpUri="https://zenodo.org/record/4545486/files/figure.png" pageId="10" pageNumber="11">Fig. 93</figureCitation>
); pronotum without paired pits anterior to median groove; high rainfall areas of NE NSW &amp; SE Qld).............................
<taxonomicName id="0EFAEEC33F59E5467511C5CBFA5546B1" authority="Baly" authorityName="Baly" authorityYear="1888" box="[1289,1436,1677,1699]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="wilsoni">
<emphasis id="FB8E49523F59E5467511C5CBFAA046B0" box="[1289,1385,1677,1698]" italics="true" pageId="10" pageNumber="11">A. wilsoni</emphasis>
Baly
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="720E8DE53F59E546708FC5ECFA55479F" pageId="10" pageNumber="11">
<paragraph id="C94595403F59E546708FC5ECFA5546E8" blockId="10.[151,1437,151,2021]" pageId="10" pageNumber="11">
<keyLead id="720B36753F59E546708FC5ECFA5546E8" pageId="10" pageNumber="11">
9(8) Pronotal groove sinuate or almost straight, deep at middle and with pair of pits anterior to median groove (
<figureCitation id="51C189C53F59E54674F9C5ECFAFE46D2" box="[1249,1335,1706,1728]" captionStart="FIGURES 25" captionStartId="15.[151,264,1829,1853]" captionTargetBox="[164,1420,191,1786]" captionTargetId="figure-19@15.[151,1436,181,1805]" captionTargetPageId="15" captionText="FIGURES 25. Aulacophora species, male habitus: 2, A. abdominalis (Fabricius, 1781) (Guadalcanal); 3, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 4, A. cornuta Baly, 1879 (New Ireland); 5, A. indica (Gmelin, 1790) (Luzon)." figureDoi="http://doi.org/10.5281/zenodo.4545452" httpUri="https://zenodo.org/record/4545452/files/figure.png" pageId="10" pageNumber="11">Figs 2, 5</figureCitation>
,
<figureCitation id="51C189C53F59E546755AC5ECFA8646AD" box="[1346,1359,1706,1727]" captionStart="FIGURES 69" captionStartId="16.[151,264,1957,1981]" captionTargetBox="[174,1418,193,1917]" captionTargetId="figure-22@16.[157,1430,181,1933]" captionTargetPageId="16" captionText="FIGURES 69. Aulacophora species, male habitus: 6, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 7, A. relicta (Boisduval, 1835) (Mount Molloy); 8, A. wallacii Baly, 1886; 9, A. wilsoni Baly, 1888." figureDoi="http://doi.org/10.5281/zenodo.4545458" httpUri="https://zenodo.org/record/4545458/files/figure.png" pageId="10" pageNumber="11">8</figureCitation>
,
<figureCitation id="51C189C53F59E5467542C5ECFF3446CC" captionStart="FIGURES 1825" captionStartId="21.[151,264,1899,1923]" captionTargetBox="[262,1328,190,1869]" captionTargetId="figure-19@21.[250,1338,181,1875]" captionTargetPageId="21" captionText="FIGURES 1825. Aulacophora species, male anterior and antenna: 18, A. abdominalis (Fabricius, 1781) (Guadalcanal); 19, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 20, A. cornuta Baly, 1879 (New Ireland); 21, A. indica (Gmelin, 1790) (Luzon); 22, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 23, A. relicta (Boisduval, 1835) (Mount Molloy); 24, A. wallacii Baly, 1886; 25, A. wilsoni Baly, 1888." figureDoi="http://doi.org/10.5281/zenodo.4545464" httpUri="https://zenodo.org/record/4545464/files/figure.png" pageId="10" pageNumber="11">18, 21, 24</figureCitation>
); elytral setal patch usually large and dense (
<figureCitation id="51C189C53F59E54672B9C581FCFB46CC" box="[673,818,1735,1758]" captionStart="FIGURES 1825" captionStartId="21.[151,264,1899,1923]" captionTargetBox="[262,1328,190,1869]" captionTargetId="figure-19@21.[250,1338,181,1875]" captionTargetPageId="21" captionText="FIGURES 1825. Aulacophora species, male anterior and antenna: 18, A. abdominalis (Fabricius, 1781) (Guadalcanal); 19, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 20, A. cornuta Baly, 1879 (New Ireland); 21, A. indica (Gmelin, 1790) (Luzon); 22, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 23, A. relicta (Boisduval, 1835) (Mount Molloy); 24, A. wallacii Baly, 1886; 25, A. wilsoni Baly, 1888." figureDoi="http://doi.org/10.5281/zenodo.4545464" httpUri="https://zenodo.org/record/4545464/files/figure.png" pageId="10" pageNumber="11">Figs 18, 21, 24</figureCitation>
); tergite 8 strongly bilobed (
<figureCitation id="51C189C53F59E5467424C581FADD46CC" box="[1084,1300,1735,1758]" captionStart-0="FIGURES 8388" captionStart-1="FIGURES 8993" captionStartId-0="42.[151,264,1943,1967]" captionStartId-1="43.[151,264,1944,1968]" captionTargetBox-0="[217,1368,192,1905]" captionTargetBox-1="[196,1391,191,1907]" captionTargetId-0="figure-22@42.[204,1384,181,1919]" captionTargetId-1="figure-19@43.[177,1410,181,1920]" captionTargetPageId-0="42" captionTargetPageId-1="43" captionText-0="FIGURES 8388. Aulacophora species, male sternite 8, ventrolateral view: 83, A. abdominalis (Fabricius, 1781) (Vanuatu); 84, A. abdominalis (New Ireland); 85, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 86, A. cornuta Baly, 1879 (Guadalcanal); 87, A. indica (Gmelin, 1790) (Timor); 88, A. indica (Luzon)." captionText-1="FIGURES 8993. Aulacophora species, male sternite 8, ventrolateral view: 89, A. indica (Gmelin, 1790) (Sumatra); 90, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 91, A. relicta (Boisduval, 1835) (Coonabarabran); 92, A. wallacii Baly, 1886; 93, A. wilsoni Baly, 1888." figureDoi-0="http://doi.org/10.5281/zenodo.4545484" figureDoi-1="http://doi.org/10.5281/zenodo.4545486" httpUri-0="https://zenodo.org/record/4545484/files/figure.png" httpUri-1="https://zenodo.org/record/4545486/files/figure.png" pageId="10" pageNumber="11">Figs 8384, 8789, 92</figureCitation>
); penis almost straight in lateral view (
<figureCitation id="51C189C53F59E54671DEC5A3FD8546E8" box="[454,588,1765,1787]" captionStart="FIGURES 121124" captionStartId="52.[151,264,1872,1896]" captionTargetBox="[166,1410,190,1822]" captionTargetId="figure-22@52.[151,1436,181,1848]" captionTargetPageId="52" captionText="FIGURES 121124. Aulacophora abdominalis (Fabricius, 1781), penis, lateral view: 121, Guadalcanal, 122, Metinaro, 123, Saibai, 124, Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.4545502" httpUri="https://zenodo.org/record/4545502/files/figure.png" pageId="10" pageNumber="11">Figs 121124</figureCitation>
,
<figureCitation id="51C189C53F59E546724EC5A3FD6546E8" box="[598,684,1765,1786]" captionStart-0="FIGURES 125128" captionStart-1="FIGURES 129132" captionStartId-0="53.[151,264,1722,1746]" captionStartId-1="54.[151,264,1767,1791]" captionTargetBox-0="[165,1400,194,1671]" captionTargetBox-1="[167,1400,192,1716]" captionTargetId-0="figure-19@53.[151,1436,181,1699]" captionTargetId-1="figure-22@54.[151,1436,181,1743]" captionTargetPageId-0="53" captionTargetPageId-1="54" captionText-0="FIGURES 125128. Aulacophora species, penis, lateral view: 125, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 126, A. cornuta Baly, 1879; 127, A. indica (Gmelin, 1790) (Laos); 128, A. indica (Luzon). Arrow indicates small spine on dorsal surface." captionText-1="FIGURES 129132. Aulacophora species, penis, lateral view: 129, A. indica (Gmelin, 1790) (W Timor); 130, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 131, A. relicta (Boisduval, 1835) (Mount Molloy); 132, A. relicta (Mount Mulligan)." figureDoi-0="http://doi.org/10.5281/zenodo.4545504" figureDoi-1="http://doi.org/10.5281/zenodo.4545506" httpUri-0="https://zenodo.org/record/4545504/files/figure.png" httpUri-1="https://zenodo.org/record/4545506/files/figure.png" pageId="10" pageNumber="11">127129</figureCitation>
,
<figureCitation id="51C189C53F59E54672AEC5A3FD1246E9" box="[694,731,1765,1787]" captionStart="FIGURES 133134" captionStartId="55.[151,264,1996,2020]" captionTargetBox="[288,1299,196,1952]" captionTargetId="figure-19@55.[271,1317,181,1973]" captionTargetPageId="55" captionText="FIGURES 133134. Aulacophora species, penis, lateral view: 133, A. wallacii Baly, 1886; 134, A. wilsoni Baly, 1888." figureDoi="http://doi.org/10.5281/zenodo.4545508" httpUri="https://zenodo.org/record/4545508/files/figure.png" pageId="10" pageNumber="11">133</figureCitation>
)........................................................ 10
</keyLead>
</paragraph>
<paragraph id="C94595403F59E546708FC445FA55479F" blockId="10.[151,1437,151,2021]" pageId="10" pageNumber="11">
<keyLead id="720B36753F59E546708FC445FA55479F" pageId="10" pageNumber="11">
- Pronotal groove almost straight, and shallow throughout, anterior and posterior slopes shallow and symmetrical in lateral view, and without pits anterior to median groove (
<figureCitation id="51C189C53F59E546729FC459FD0C4727" box="[647,709,1823,1845]" captionStart="FIGURES 69" captionStartId="16.[151,264,1957,1981]" captionTargetBox="[174,1418,193,1917]" captionTargetId="figure-22@16.[157,1430,181,1933]" captionTargetPageId="16" captionText="FIGURES 69. Aulacophora species, male habitus: 6, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 7, A. relicta (Boisduval, 1835) (Mount Molloy); 8, A. wallacii Baly, 1886; 9, A. wilsoni Baly, 1888." figureDoi="http://doi.org/10.5281/zenodo.4545458" httpUri="https://zenodo.org/record/4545458/files/figure.png" pageId="10" pageNumber="11">Figs 6</figureCitation>
,
<figureCitation id="51C189C53F59E54672D7C466FD234724" box="[719,746,1824,1846]" captionStart="FIGURES 1825" captionStartId="21.[151,264,1899,1923]" captionTargetBox="[262,1328,190,1869]" captionTargetId="figure-19@21.[250,1338,181,1875]" captionTargetPageId="21" captionText="FIGURES 1825. Aulacophora species, male anterior and antenna: 18, A. abdominalis (Fabricius, 1781) (Guadalcanal); 19, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 20, A. cornuta Baly, 1879 (New Ireland); 21, A. indica (Gmelin, 1790) (Luzon); 22, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 23, A. relicta (Boisduval, 1835) (Mount Molloy); 24, A. wallacii Baly, 1886; 25, A. wilsoni Baly, 1888." figureDoi="http://doi.org/10.5281/zenodo.4545464" httpUri="https://zenodo.org/record/4545464/files/figure.png" pageId="10" pageNumber="11">22</figureCitation>
); elytral setal patch small, sparse, not dense (
<figureCitation id="51C189C53F59E5467482C459FB2B4724" box="[1178,1250,1823,1846]" captionStart="FIGURES 1825" captionStartId="21.[151,264,1899,1923]" captionTargetBox="[262,1328,190,1869]" captionTargetId="figure-19@21.[250,1338,181,1875]" captionTargetPageId="21" captionText="FIGURES 1825. Aulacophora species, male anterior and antenna: 18, A. abdominalis (Fabricius, 1781) (Guadalcanal); 19, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 20, A. cornuta Baly, 1879 (New Ireland); 21, A. indica (Gmelin, 1790) (Luzon); 22, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 23, A. relicta (Boisduval, 1835) (Mount Molloy); 24, A. wallacii Baly, 1886; 25, A. wilsoni Baly, 1888." figureDoi="http://doi.org/10.5281/zenodo.4545464" httpUri="https://zenodo.org/record/4545464/files/figure.png" pageId="10" pageNumber="11">Fig. 22</figureCitation>
); tergite 8 medially lobed (
<figureCitation id="51C189C53F59E546713DC47BFEA44741" box="[293,365,1853,1875]" captionStart="FIGURES 8993" captionStartId="43.[151,264,1944,1968]" captionTargetBox="[196,1391,191,1907]" captionTargetId="figure-19@43.[177,1410,181,1920]" captionTargetPageId="43" captionText="FIGURES 8993. Aulacophora species, male sternite 8, ventrolateral view: 89, A. indica (Gmelin, 1790) (Sumatra); 90, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 91, A. relicta (Boisduval, 1835) (Coonabarabran); 92, A. wallacii Baly, 1886; 93, A. wilsoni Baly, 1888." figureDoi="http://doi.org/10.5281/zenodo.4545486" httpUri="https://zenodo.org/record/4545486/files/figure.png" pageId="10" pageNumber="11">Fig. 90</figureCitation>
); penis strongly curved in lateral view (
<figureCitation id="51C189C53F59E54672F5C47BFCF64741" box="[749,831,1853,1875]" captionStart="FIGURES 129132" captionStartId="54.[151,264,1767,1791]" captionTargetBox="[167,1400,192,1716]" captionTargetId="figure-22@54.[151,1436,181,1743]" captionTargetPageId="54" captionText="FIGURES 129132. Aulacophora species, penis, lateral view: 129, A. indica (Gmelin, 1790) (W Timor); 130, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 131, A. relicta (Boisduval, 1835) (Mount Molloy); 132, A. relicta (Mount Mulligan)." figureDoi="http://doi.org/10.5281/zenodo.4545506" httpUri="https://zenodo.org/record/4545506/files/figure.png" pageId="10" pageNumber="11">Fig. 130</figureCitation>
) (length
<quantity id="0E0238A53F59E546738FC47BFC324740" box="[919,1019,1853,1875]" metricMagnitude="-3" metricUnit="m" metricValue="7.25" metricValueMax="7.5" metricValueMin="7.0" pageId="10" pageNumber="11" unit="mm" value="7.25" valueMax="7.5" valueMin="7.0">77.5 mm</quantity>
; meso- and metatibiae dark brown to black and anterior tibiae often with dark outer edges; antennomeres 411 dark brown to black; inland N Qld)................................................................................
<taxonomicName id="0EFAEEC33F59E54673D6C43EFA8F479F" authority="Reid, Halling &amp; Beatson" authorityName="Reid, Halling &amp; Beatson" authorityYear="2021" box="[974,1350,1911,1933]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="mbabaram" status="sp. nov.">
<emphasis id="FB8E49523F59E54673D6C43EFB99479E" box="[974,1104,1911,1933]" italics="true" pageId="10" pageNumber="11">A. mbabaram</emphasis>
Reid, Halling &amp; Beatson
</taxonomicName>
,
<emphasis id="FB8E49523F59E5467548C43EFA55479F" bold="true" box="[1360,1436,1912,1933]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="E0BDF4293F59E5467548C43EFA55479F" box="[1360,1436,1912,1933]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="720E8DE53F59E547708FC4D3FA544131" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<paragraph id="C94595403F59E546708FC4D3FA5547D5" blockId="10.[151,1437,151,2021]" pageId="10" pageNumber="11">
<keyLead id="720B36753F59E546708FC4D3FA5547D5" pageId="10" pageNumber="11">
10(9) Pronotal median depression medially deeper, in lateral view with anterior slope steeper than posterior (
<figureCitation id="51C189C53F59E54674CAC4D3FA8D47B8" box="[1234,1348,1941,1963]" captionStart="FIGURES 1825" captionStartId="21.[151,264,1899,1923]" captionTargetBox="[262,1328,190,1869]" captionTargetId="figure-19@21.[250,1338,181,1875]" captionTargetPageId="21" captionText="FIGURES 1825. Aulacophora species, male anterior and antenna: 18, A. abdominalis (Fabricius, 1781) (Guadalcanal); 19, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 20, A. cornuta Baly, 1879 (New Ireland); 21, A. indica (Gmelin, 1790) (Luzon); 22, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 23, A. relicta (Boisduval, 1835) (Mount Molloy); 24, A. wallacii Baly, 1886; 25, A. wilsoni Baly, 1888." figureDoi="http://doi.org/10.5281/zenodo.4545464" httpUri="https://zenodo.org/record/4545464/files/figure.png" pageId="10" pageNumber="11">Figs 18, 24</figureCitation>
,
<figureCitation id="51C189C53F59E5467548C4D3FA5947B9" box="[1360,1424,1941,1963]" captionStart="FIGURES 2631" captionStartId="22.[151,264,1703,1727]" captionTargetBox="[160,1426,191,1670]" captionTargetId="figure-22@22.[151,1436,181,1679]" captionTargetPageId="22" captionText="FIGURES 2631. Aulacophora species, pronotum, lateral view: 26, male A. abdominalis (Fabricius, 1781) (Vanuatu); 27, male A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 28, male A. indica (Gmelin, 1790) (Borneo); 29, female A. indica; 30, male A. relicta (Boisduval, 1835); 31, male A. wallacii Baly, 1886." figureDoi="http://doi.org/10.5281/zenodo.4545466" httpUri="https://zenodo.org/record/4545466/files/figure.png" pageId="10" pageNumber="11">26, 31</figureCitation>
); penis sinuately margined in dorsal view, with acute apex (
<figureCitation id="51C189C53F59E5467315C4F4FCB547D5" box="[781,892,1970,1992]" captionStart="FIGURES 9497" captionStartId="44.[151,264,1737,1761]" captionTargetBox="[170,1385,198,1674]" captionTargetId="figure-22@44.[151,1435,181,1713]" captionTargetPageId="44" captionText="FIGURES 9497. Aulacophora abdominalis (Fabricius, 1781), penis, dorsal view: 94, Guadalcanal, 95, Metinaro, 96, Saibai, 97, Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.4545488" httpUri="https://zenodo.org/record/4545488/files/figure.png" pageId="10" pageNumber="11">Figs 9497</figureCitation>
,
<figureCitation id="51C189C53F59E546739EC4F4FC6247DA" box="[902,939,1970,1992]" captionStart="FIGURES 106107" captionStartId="47.[151,264,1987,2011]" captionTargetBox="[425,1163,195,1932]" captionTargetId="figure-19@47.[405,1183,181,1963]" captionTargetPageId="47" captionText="FIGURES 106107. Aulacophora species, penis, dorsal view: 106, A. wilsoni Baly, 1888; 107, A. wallacii Baly, 1886." figureDoi="http://doi.org/10.5281/zenodo.4545494" httpUri="https://zenodo.org/record/4545494/files/figure.png" pageId="10" pageNumber="11">107</figureCitation>
)...................................... 11
</keyLead>
</paragraph>
<paragraph id="C94595403F59E547708FC496FA544131" blockId="10.[151,1437,151,2021]" lastBlockId="11.[151,1437,151,2022]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<keyLead id="720B36753F59E547708FC496FA544131" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
- Pronotal median depression medially shallower, in lateral view with anterior and posterior slopes about equally steep (
<figureCitation id="51C189C53F59E5467556C489FA5147F7" box="[1358,1432,1999,2021]" captionStart="FIGURES 1825" captionStartId="21.[151,264,1899,1923]" captionTargetBox="[262,1328,190,1869]" captionTargetId="figure-19@21.[250,1338,181,1875]" captionTargetPageId="21" captionText="FIGURES 1825. Aulacophora species, male anterior and antenna: 18, A. abdominalis (Fabricius, 1781) (Guadalcanal); 19, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 20, A. cornuta Baly, 1879 (New Ireland); 21, A. indica (Gmelin, 1790) (Luzon); 22, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 23, A. relicta (Boisduval, 1835) (Mount Molloy); 24, A. wallacii Baly, 1886; 25, A. wilsoni Baly, 1888." figureDoi="http://doi.org/10.5281/zenodo.4545464" httpUri="https://zenodo.org/record/4545464/files/figure.png" pageId="10" pageNumber="11">Figs 21</figureCitation>
,
<figureCitation id="51C189C53F58E54770FAC3DEFF3440BC" box="[226,253,152,174]" captionStart="FIGURES 2631" captionStartId="22.[151,264,1703,1727]" captionTargetBox="[160,1426,191,1670]" captionTargetId="figure-22@22.[151,1436,181,1679]" captionTargetPageId="22" captionText="FIGURES 2631. Aulacophora species, pronotum, lateral view: 26, male A. abdominalis (Fabricius, 1781) (Vanuatu); 27, male A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 28, male A. indica (Gmelin, 1790) (Borneo); 29, female A. indica; 30, male A. relicta (Boisduval, 1835); 31, male A. wallacii Baly, 1886." figureDoi="http://doi.org/10.5281/zenodo.4545466" httpUri="https://zenodo.org/record/4545466/files/figure.png" pageId="11" pageNumber="12">27</figureCitation>
); penis almost parallel-sided in dorsal view, with blunt apex (
<figureCitation id="51C189C53F58E547735CC3D1FC0340BC" box="[836,970,151,174]" captionStart-0="FIGURES 98101" captionStart-1="FIGURES 102105" captionStartId-0="45.[151,264,1825,1849]" captionStartId-1="46.[151,264,1777,1801]" captionTargetBox-0="[164,1393,195,1754]" captionTargetBox-1="[166,1388,194,1718]" captionTargetId-0="figure-19@45.[151,1436,181,1801]" captionTargetId-1="figure-22@46.[151,1436,181,1754]" captionTargetPageId-0="45" captionTargetPageId-1="46" captionText-0="FIGURES 98101. Aulacophora species, penis, dorsal view: 98, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 99, A. cornuta Baly, 1879 (Guadalcanal); 100, A. indica (Gmelin, 1790) (Laos); 101, A. indica (Luzon)." captionText-1="FIGURES 102105. Aulacophora species, penis, dorsal view: 102, A. indica (Gmelin, 1790) (Timor); 103, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 104, A. relicta (Boisduval, 1835) (Mount Molloy); 105, A. relicta (Mount Mulligan)." figureDoi-0="http://doi.org/10.5281/zenodo.4545490" figureDoi-1="http://doi.org/10.5281/zenodo.4545492" httpUri-0="https://zenodo.org/record/4545490/files/figure.png" httpUri-1="https://zenodo.org/record/4545492/files/figure.png" pageId="11" pageNumber="12">Figs 100102</figureCitation>
) (length
<quantity id="0E0238A53F58E5477407C3D1FBB840BF" box="[1055,1137,151,173]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="11" pageNumber="12" unit="mm" value="7.0" valueMax="8.0" valueMin="6.0">68 mm</quantity>
; antennae usually entirely pale, rarely darkened from antennomere 4 to apex; scutellum usually pale, dark brown in some specimens from Borneo; mesepimeron pale; legs entirely pale or with darkened meso- and metafemora and external edges of meso- and metatibiae; last ventrite with apical excavation asymmetric, more sharply bordered on left; tergite 8 arcuate between narrowly triangular prongs (
<figureCitation id="51C189C53F58E547756AC3A9FED64131" captionStart-0="FIGURES 8388" captionStart-1="FIGURES 8993" captionStartId-0="42.[151,264,1943,1967]" captionStartId-1="43.[151,264,1944,1968]" captionTargetBox-0="[217,1368,192,1905]" captionTargetBox-1="[196,1391,191,1907]" captionTargetId-0="figure-22@42.[204,1384,181,1919]" captionTargetId-1="figure-19@43.[177,1410,181,1920]" captionTargetPageId-0="42" captionTargetPageId-1="43" captionText-0="FIGURES 8388. Aulacophora species, male sternite 8, ventrolateral view: 83, A. abdominalis (Fabricius, 1781) (Vanuatu); 84, A. abdominalis (New Ireland); 85, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 86, A. cornuta Baly, 1879 (Guadalcanal); 87, A. indica (Gmelin, 1790) (Timor); 88, A. indica (Luzon)." captionText-1="FIGURES 8993. Aulacophora species, male sternite 8, ventrolateral view: 89, A. indica (Gmelin, 1790) (Sumatra); 90, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 91, A. relicta (Boisduval, 1835) (Coonabarabran); 92, A. wallacii Baly, 1886; 93, A. wilsoni Baly, 1888." figureDoi-0="http://doi.org/10.5281/zenodo.4545484" figureDoi-1="http://doi.org/10.5281/zenodo.4545486" httpUri-0="https://zenodo.org/record/4545484/files/figure.png" httpUri-1="https://zenodo.org/record/4545486/files/figure.png" pageId="11" pageNumber="12">Figs 8789</figureCitation>
) (S &amp; E Asia to Timor,
<collectingCountry id="B1EDD5D03F58E547721CC24BFDBA4131" box="[516,627,269,291]" name="Micronesia" pageId="11" pageNumber="12">Micronesia</collectingCountry>
,
<collectingCountry id="B1EDD5D03F58E5477264C24BFD704130" box="[636,697,269,290]" name="Guam" pageId="11" pageNumber="12">Guam</collectingCountry>
, and intercepted in
<collectingCountry id="B1EDD5D03F58E547736BC24BFC024131" box="[883,971,269,291]" name="Australia" pageId="11" pageNumber="12">Australia</collectingCountry>
but not established).......
<taxonomicName id="0EFAEEC33F58E54774F0C24BFA544131" authority="(Gmelin)" authorityName="Luzon" baseAuthorityName="Gmelin" baseAuthorityYear="1790" box="[1256,1437,269,291]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="FB8E49523F58E54774F0C24BFAF74130" box="[1256,1342,269,290]" italics="true" pageId="11" pageNumber="12">A. indica</emphasis>
(Gmelin)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="720E8DE53F58E547708FC26CFA554277" pageId="11" pageNumber="12">
<paragraph id="C94595403F58E547708FC26CFA5541C1" blockId="11.[151,1437,151,2022]" pageId="11" pageNumber="12">
<keyLead id="720B36753F58E547708FC26CFA5541C1" pageId="11" pageNumber="12">
11(10) Antennae usually darkened apically (
<figureCitation id="51C189C53F58E5477253C26CFD5A4152" box="[587,659,298,320]" captionStart="FIGURES 1825" captionStartId="21.[151,264,1899,1923]" captionTargetBox="[262,1328,190,1869]" captionTargetId="figure-19@21.[250,1338,181,1875]" captionTargetPageId="21" captionText="FIGURES 1825. Aulacophora species, male anterior and antenna: 18, A. abdominalis (Fabricius, 1781) (Guadalcanal); 19, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 20, A. cornuta Baly, 1879 (New Ireland); 21, A. indica (Gmelin, 1790) (Luzon); 22, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 23, A. relicta (Boisduval, 1835) (Mount Molloy); 24, A. wallacii Baly, 1886; 25, A. wilsoni Baly, 1888." figureDoi="http://doi.org/10.5281/zenodo.4545464" httpUri="https://zenodo.org/record/4545464/files/figure.png" pageId="11" pageNumber="12">Fig. 18</figureCitation>
), rarely entirely pale (some Timorese specimens); last ventrite with asymmetric deep apical excavation, more sharply bordered on left (
<figureCitation id="51C189C53F58E54772F4C201FC95414F" box="[748,860,327,349]" captionStart="FIGURES 3237" captionStartId="23.[151,264,1840,1864]" captionTargetBox="[167,1416,192,1802]" captionTargetId="figure-19@23.[151,1436,181,1816]" captionTargetPageId="23" captionText="FIGURES 3237. Aulacophora species, male, apex of abdominal venter: 32, A. abdominalis (Fabricius, 1781) (Vanuatu); 33, A. abdominalis (New Ireland); 34, A. abdominalis (Manatuto); 35, A. abdominalis (Metinaro); 36, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 37, A. cornuta Baly, 1879 (Guadalcanal)." figureDoi="http://doi.org/10.5281/zenodo.4545468" httpUri="https://zenodo.org/record/4545468/files/figure.png" pageId="11" pageNumber="12">Figs 3235</figureCitation>
); scutellum pale, as elytra; mesepimeron pale; tergite 8 arcuate between elongate triangular prongs (
<figureCitation id="51C189C53F58E547727CC223FD1D4169" box="[612,724,357,379]" captionStart="FIGURES 8388" captionStartId="42.[151,264,1943,1967]" captionTargetBox="[217,1368,192,1905]" captionTargetId="figure-22@42.[204,1384,181,1919]" captionTargetPageId="42" captionText="FIGURES 8388. Aulacophora species, male sternite 8, ventrolateral view: 83, A. abdominalis (Fabricius, 1781) (Vanuatu); 84, A. abdominalis (New Ireland); 85, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 86, A. cornuta Baly, 1879 (Guadalcanal); 87, A. indica (Gmelin, 1790) (Timor); 88, A. indica (Luzon)." figureDoi="http://doi.org/10.5281/zenodo.4545484" httpUri="https://zenodo.org/record/4545484/files/figure.png" pageId="11" pageNumber="12">Figs 8384</figureCitation>
); penis broader, asymmetric, in dorsal view slightly constricted at middle, without ventral ridge (
<figureCitation id="51C189C53F58E54771A0C2C4FDEE4185" box="[440,551,386,408]" captionStart="FIGURES 9497" captionStartId="44.[151,264,1737,1761]" captionTargetBox="[170,1385,198,1674]" captionTargetId="figure-22@44.[151,1435,181,1713]" captionTargetPageId="44" captionText="FIGURES 9497. Aulacophora abdominalis (Fabricius, 1781), penis, dorsal view: 94, Guadalcanal, 95, Metinaro, 96, Saibai, 97, Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.4545488" httpUri="https://zenodo.org/record/4545488/files/figure.png" pageId="11" pageNumber="12">Figs 9497</figureCitation>
,
<figureCitation id="51C189C53F58E5477229C2C4FD4D418A" box="[561,644,386,408]" captionStart="FIGURES 108111" captionStartId="48.[151,264,1843,1867]" captionTargetBox="[184,1420,192,1788]" captionTargetId="figure-22@48.[151,1436,181,1820]" captionTargetPageId="48" captionText="FIGURES 108111. Aulacophora abdominalis (Fabricius, 1781), penis, ventral view: 108, Guadalcanal, 109, Metinaro, 110, Saibai, 111, Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.4545496" httpUri="https://zenodo.org/record/4545496/files/figure.png" pageId="11" pageNumber="12">108111</figureCitation>
) (length
<quantity id="0E0238A53F58E54772C3C2C4FC89418A" box="[731,832,386,408]" metricMagnitude="-3" metricUnit="m" metricValue="6.75" metricValueMax="7.5" metricValueMin="6.0" pageId="11" pageNumber="12" unit="mm" value="6.75" valueMax="7.5" valueMin="6.0">67.5 mm</quantity>
; meso- and metafemora and tibiae usually darkened; pygidium with base darker than apex; Timor to
<collectingCountry id="B1EDD5D03F58E5477251C2E6FD4C41A7" box="[585,645,416,437]" name="Tonga" pageId="11" pageNumber="12">Tonga</collectingCountry>
and probably
<collectingCountry id="B1EDD5D03F58E5477316C2E6FC9B41A7" box="[782,850,416,437]" name="Samoa" pageId="11" pageNumber="12">Samoa</collectingCountry>
, including Torres Strait Islands but not Australian mainland)...............................................................................
<taxonomicName id="0EFAEEC33F58E5477484C2FBFA5541C1" authority="(Fabricius)" authorityName="Mer" baseAuthorityName="Fabricius" baseAuthorityYear="1781" box="[1180,1436,445,467]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis id="FB8E49523F58E5477484C2FBFAE441C0" box="[1180,1325,445,466]" italics="true" pageId="11" pageNumber="12">A. abdominalis</emphasis>
(Fabricius)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C94595403F58E547708FC29DFA554277" blockId="11.[151,1437,151,2022]" pageId="11" pageNumber="12">
<keyLead id="720B36753F58E547708FC29DFA554277" pageId="11" pageNumber="12">
- Antennae entirely pale (
<figureCitation id="51C189C53F58E54771DFC29CFDC641E2" box="[455,527,474,496]" captionStart="FIGURES 1825" captionStartId="21.[151,264,1899,1923]" captionTargetBox="[262,1328,190,1869]" captionTargetId="figure-19@21.[250,1338,181,1875]" captionTargetPageId="21" captionText="FIGURES 1825. Aulacophora species, male anterior and antenna: 18, A. abdominalis (Fabricius, 1781) (Guadalcanal); 19, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 20, A. cornuta Baly, 1879 (New Ireland); 21, A. indica (Gmelin, 1790) (Luzon); 22, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 23, A. relicta (Boisduval, 1835) (Mount Molloy); 24, A. wallacii Baly, 1886; 25, A. wilsoni Baly, 1888." figureDoi="http://doi.org/10.5281/zenodo.4545464" httpUri="https://zenodo.org/record/4545464/files/figure.png" pageId="11" pageNumber="12">Fig. 24</figureCitation>
); last ventrite with symmetrical broad shallow apical excavation, without sharply ridged lateral margins (
<figureCitation id="51C189C53F58E5477126C2B1FE4E421C" box="[318,391,503,526]" captionStart="FIGURES 3843" captionStartId="24.[151,264,1793,1817]" captionTargetBox="[168,1418,191,1757]" captionTargetId="figure-22@24.[151,1437,181,1769]" captionTargetPageId="24" captionText="FIGURES 3843. Aulacophora species, male, apex of abdominal venter: 38, A. indica (Gmelin, 1790) (Sumatra); 39, A. indica (West Timor); 40, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 41, A. relicta (Boisduval, 1835) (Mount Molloy); 42, A. wallacii Baly, 1886; 43, A. wilsoni Baly, 1888." figureDoi="http://doi.org/10.5281/zenodo.4545470" httpUri="https://zenodo.org/record/4545470/files/figure.png" pageId="11" pageNumber="12">Fig. 42</figureCitation>
); scutellum often darkened, brown or dark brown; mesepimeron usually dark brown; tergite 8 with pair of long needle-like processes, truncate between (
<figureCitation id="51C189C53F58E5477285C153FD2F4239" box="[669,742,533,555]" captionStart="FIGURES 8993" captionStartId="43.[151,264,1944,1968]" captionTargetBox="[196,1391,191,1907]" captionTargetId="figure-19@43.[177,1410,181,1920]" captionTargetPageId="43" captionText="FIGURES 8993. Aulacophora species, male sternite 8, ventrolateral view: 89, A. indica (Gmelin, 1790) (Sumatra); 90, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 91, A. relicta (Boisduval, 1835) (Coonabarabran); 92, A. wallacii Baly, 1886; 93, A. wilsoni Baly, 1888." figureDoi="http://doi.org/10.5281/zenodo.4545486" httpUri="https://zenodo.org/record/4545486/files/figure.png" pageId="11" pageNumber="12">Fig. 92</figureCitation>
); penis thinner, symmetrical, constricted before apex in dorsal view and venter longitudinally ridged (
<figureCitation id="51C189C53F58E5477218C174FD914255" box="[512,600,562,584]" captionStart="FIGURES 106107" captionStartId="47.[151,264,1987,2011]" captionTargetBox="[425,1163,195,1932]" captionTargetId="figure-19@47.[405,1183,181,1963]" captionTargetPageId="47" captionText="FIGURES 106107. Aulacophora species, penis, dorsal view: 106, A. wilsoni Baly, 1888; 107, A. wallacii Baly, 1886." figureDoi="http://doi.org/10.5281/zenodo.4545494" httpUri="https://zenodo.org/record/4545494/files/figure.png" pageId="11" pageNumber="12">Figs 107</figureCitation>
,
<figureCitation id="51C189C53F58E547727CC174FD41425A" box="[612,648,562,584]" captionStart="FIGURES 116119" captionStartId="50.[151,264,1850,1874]" captionTargetBox="[164,1411,194,1790]" captionTargetId="figure-22@50.[151,1436,181,1826]" captionTargetPageId="50" captionText="FIGURES 116119. Aulacophora species, penis, ventral view: 116, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 117, A. relicta (Boisduval, 1835) (Mount Molloy); 118, A. relicta (Mount Mulligan); 119, A. wallacii Baly, 1886." figureDoi="http://doi.org/10.5281/zenodo.4545500" httpUri="https://zenodo.org/record/4545500/files/figure.png" pageId="11" pageNumber="12">119</figureCitation>
) (length
<quantity id="0E0238A53F58E54772FBC174FCF8425A" box="[739,817,562,584]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="11" pageNumber="12" unit="mm" value="7.5">7.5 mm</quantity>
; meso- and metafemora usually dark brown or black but tibiae pale); (endemic to Timor)...................................................................
<taxonomicName id="0EFAEEC33F58E5477519C116FA554277" authority="Baly" authorityName="Baly" authorityYear="1886" box="[1281,1436,591,613]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="wallacii">
<emphasis id="FB8E49523F58E5477519C116FAA04276" box="[1281,1385,591,613]" italics="true" pageId="11" pageNumber="12">A. wallacii</emphasis>
Baly
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</keyStep>
<keyStep id="720E8DE53F58E547708FC12BFA5542EA" pageId="11" pageNumber="12">
<paragraph id="C94595403F58E547708FC12BFA55428D" blockId="11.[151,1437,151,2022]" pageId="11" pageNumber="12">
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12(4) Frontoclypeus as male (
<figureCitation id="51C189C53F58E54771DFC12BFDC74291" box="[455,526,621,643]" captionStart="FIGURES 1015" captionStartId="17.[151,264,1943,1967]" captionTargetBox="[209,1375,181,1906]" captionTargetId="figure-19@17.[209,1379,181,1920]" captionTargetPageId="17" captionText="FIGURES 1015. Aulacophora species, male head, anterior view: 10, A. abdominalis (Fabricius, 1781) (Guadalcanal); 11, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 12, A. cornuta Baly, 1879 (New Ireland); 13, A. indica (Gmelin, 1790) (Luzon); 14, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 15, A. relicta (Boisduval, 1835) (Mount Molloy)." figureDoi="http://doi.org/10.5281/zenodo.4545460" httpUri="https://zenodo.org/record/4545460/files/figure.png" pageId="11" pageNumber="12">Fig. 10</figureCitation>
), with glabrous median keel from postantennal calli to labrum; apex of pygidium strongly produced, and apex truncate, narrowly rounded, or angulate (
<figureCitation id="51C189C53F58E5477314C1CCFCB3428D" box="[780,890,650,672]" captionStart="FIGURES 4449" captionStartId="25.[151,264,1798,1822]" captionTargetBox="[170,1418,190,1763]" captionTargetId="figure-19@25.[151,1436,181,1774]" captionTargetPageId="25" captionText="FIGURES 4449. Aulacophora species, female, apex of abdominal venter and pygidium: 44, A. abdominalis (Fabricius, 1781) (Vanuatu); 45, A. abdominalis (New Ireland); 46, A. abdominalis (Mer Island); 47, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 48, A. cornuta Baly, 1879 (Sulawesi); 49, A. indica (Gmelin, 1790) (Laos).Dotted lines indicate depth and width of apical concavity." figureDoi="http://doi.org/10.5281/zenodo.4545472" httpUri="https://zenodo.org/record/4545472/files/figure.png" pageId="11" pageNumber="12">Figs 4447</figureCitation>
,
<figureCitation id="51C189C53F58E547739CC1CCFC0B42B2" box="[900,962,650,672]" captionStart="FIGURES 5055" captionStartId="26.[151,264,1943,1967]" captionTargetBox="[198,1388,201,1907]" captionTargetId="figure-22@26.[178,1409,181,1919]" captionTargetPageId="26" captionText="FIGURES 5055. Aulacophora species, female, apex of abdominal venter: 50, A. indica (Gmelin, 1790) (Biak Island); 51, A. indica (Vietnam); 52, A. indica (Broken Hill); 53, A. mbabaram Reid, Halling &amp; Beatson, sp. nov., Almaden; 54, A. relicta (Boisduval, 1835); 55, A. wallacii Baly, 1886." figureDoi="http://doi.org/10.5281/zenodo.4545474" httpUri="https://zenodo.org/record/4545474/files/figure.png" pageId="11" pageNumber="12">5055</figureCitation>
).................................... 13
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</paragraph>
<paragraph id="C94595403F58E547708FC1EEFA5542EA" blockId="11.[151,1437,151,2022]" pageId="11" pageNumber="12">
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- Frontoclypeus without median keel, with elongate patch of dense setae at base between antennae; apex of pygidium slightly produced, and with broadly rounded apical margin (
<figureCitation id="51C189C53F58E54772CAC183FCDE42C9" box="[722,791,709,731]" captionStart="FIGURES 4449" captionStartId="25.[151,264,1798,1822]" captionTargetBox="[170,1418,190,1763]" captionTargetId="figure-19@25.[151,1436,181,1774]" captionTargetPageId="25" captionText="FIGURES 4449. Aulacophora species, female, apex of abdominal venter and pygidium: 44, A. abdominalis (Fabricius, 1781) (Vanuatu); 45, A. abdominalis (New Ireland); 46, A. abdominalis (Mer Island); 47, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 48, A. cornuta Baly, 1879 (Sulawesi); 49, A. indica (Gmelin, 1790) (Laos).Dotted lines indicate depth and width of apical concavity." figureDoi="http://doi.org/10.5281/zenodo.4545472" httpUri="https://zenodo.org/record/4545472/files/figure.png" pageId="11" pageNumber="12">Fig. 48</figureCitation>
) (length
<quantity id="0E0238A53F58E5477376C183FC7642C8" box="[878,959,709,730]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="11" pageNumber="12" unit="mm" value="8.5" valueMax="9.0" valueMin="8.0">89 mm</quantity>
; abdomen dark brown to black; apical swelling of pygidium with median carina)...............................................................
<taxonomicName id="0EFAEEC33F58E547751CC1A4FA5542EA" authority="Baly" authorityName="Baly" authorityYear="1879" box="[1284,1436,738,760]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="FB8E49523F58E547751CC1A4FAA042EA" box="[1284,1385,738,760]" italics="true" pageId="11" pageNumber="12">A. cornuta</emphasis>
Baly
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<keyStep id="720E8DE53F58E547708FC046FA5543D7" pageId="11" pageNumber="12">
<paragraph id="C94595403F58E547708FC046FA55435D" blockId="11.[151,1437,151,2022]" pageId="11" pageNumber="12">
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13(12) Apex of abdomen pale, at least apical margins of both pygidium and ventrite 5 (
<figureCitation id="51C189C53F58E54773C1C1B9FB584307" box="[985,1169,767,789]" captionStart-0="FIGURES 4449" captionStart-1="FIGURES 5055" captionStartId-0="25.[151,264,1798,1822]" captionStartId-1="26.[151,264,1943,1967]" captionTargetBox-0="[170,1418,190,1763]" captionTargetBox-1="[198,1388,201,1907]" captionTargetId-0="figure-19@25.[151,1436,181,1774]" captionTargetId-1="figure-22@26.[178,1409,181,1919]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionText-0="FIGURES 4449. Aulacophora species, female, apex of abdominal venter and pygidium: 44, A. abdominalis (Fabricius, 1781) (Vanuatu); 45, A. abdominalis (New Ireland); 46, A. abdominalis (Mer Island); 47, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 48, A. cornuta Baly, 1879 (Sulawesi); 49, A. indica (Gmelin, 1790) (Laos).Dotted lines indicate depth and width of apical concavity." captionText-1="FIGURES 5055. Aulacophora species, female, apex of abdominal venter: 50, A. indica (Gmelin, 1790) (Biak Island); 51, A. indica (Vietnam); 52, A. indica (Broken Hill); 53, A. mbabaram Reid, Halling &amp; Beatson, sp. nov., Almaden; 54, A. relicta (Boisduval, 1835); 55, A. wallacii Baly, 1886." figureDoi-0="http://doi.org/10.5281/zenodo.4545472" figureDoi-1="http://doi.org/10.5281/zenodo.4545474" httpUri-0="https://zenodo.org/record/4545472/files/figure.png" httpUri-1="https://zenodo.org/record/4545474/files/figure.png" pageId="11" pageNumber="12">Figs 4447, 4955</figureCitation>
); apical margin of ventrite 5 medially deeply excavate to shallowly concave; antennomeres longer, length 5th 2.53.2x width, 411 usually pale or slightly darkened, rarely dark brown to black; scutellum usually pale................................................. 14
</keyLead>
</paragraph>
<paragraph id="C94595403F58E547708FC01EFA5543D7" blockId="11.[151,1437,151,2022]" pageId="11" pageNumber="12">
<keyLead id="720B36753F58E547708FC01EFA5543D7" pageId="11" pageNumber="12">
- Abdomen entirely black (
<figureCitation id="51C189C53F58E54771CEC011FD8E437F" box="[470,583,855,877]" captionStart="FIGURES 5657" captionStartId="27.[151,264,701,725]" captionTargetBox="[165,1415,191,663]" captionTargetId="figure-19@27.[151,1436,181,677]" captionTargetPageId="27" captionText="FIGURES 5657. Aulacophora species, female, apex of abdominal venter: 56, A. wilsoni Baly, 1888 (Morgans Creek); 57, A. wilsoni (Eungella)." figureDoi="http://doi.org/10.5281/zenodo.4545476" httpUri="https://zenodo.org/record/4545476/files/figure.png" pageId="11" pageNumber="12">Figs 5657</figureCitation>
); apical margin of ventrite 5 truncate; antennomeres shorter, length 5th about 2.4x width, 411 black; scutellum dark brown to black (length
<quantity id="0E0238A53F58E54772D3C033FCE64398" box="[715,815,885,907]" metricMagnitude="-3" metricUnit="m" metricValue="6.75" metricValueMax="7.0" metricValueMin="6.5" pageId="11" pageNumber="12" unit="mm" value="6.75" valueMax="7.0" valueMin="6.5">6.57 mm</quantity>
; mesepimeron black or dark brown; mid and hind legs black and anterior tibiae with black outer edges; apex of pygidium more or less narrowly truncate with apical median tubercle)...............................................................................................
<taxonomicName id="0EFAEEC33F58E5477511C0F6FA5543D7" authority="Baly" authorityName="Baly" authorityYear="1888" box="[1289,1436,943,965]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="wilsoni">
<emphasis id="FB8E49523F58E5477511C0F6FAA043D6" box="[1289,1385,943,965]" italics="true" pageId="11" pageNumber="12">A. wilsoni</emphasis>
Baly
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<keyStep id="720E8DE53F58E547708FC08BFA54451A" pageId="11" pageNumber="12">
<paragraph id="C94595403F58E547708FC08BFA55444A" blockId="11.[151,1437,151,2022]" pageId="11" pageNumber="12">
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14(13) Apical excision of ventrite 5 shallowly arcuate to broadly U-, V-, or W-shaped, often asymmetric or irregular in shape, width at least 1.8x depth (deepest in
<collectingRegion id="0B3E5BA23F58E547721DC0ADFDB54412" box="[517,636,1002,1024]" country="Papua New Guinea" name="New Ireland" pageId="11" pageNumber="12">New Ireland</collectingRegion>
specimens of
<taxonomicName id="0EFAEEC33F58E547731EC0ACFC5143ED" authorityName="Mer" baseAuthorityName="Fabricius" baseAuthorityYear="1781" box="[774,920,1002,1023]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis id="FB8E49523F58E547731EC0ACFC5143ED" box="[774,920,1002,1023]" italics="true" pageId="11" pageNumber="12">A. abdominalis</emphasis>
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), usually much greater, and apex of ventrite 5 usually with strongly reflexed edge (
<figureCitation id="51C189C53F58E54771ECC741FDAA440F" box="[500,611,1031,1053]" captionStart="FIGURES 4449" captionStartId="25.[151,264,1798,1822]" captionTargetBox="[170,1418,190,1763]" captionTargetId="figure-19@25.[151,1436,181,1774]" captionTargetPageId="25" captionText="FIGURES 4449. Aulacophora species, female, apex of abdominal venter and pygidium: 44, A. abdominalis (Fabricius, 1781) (Vanuatu); 45, A. abdominalis (New Ireland); 46, A. abdominalis (Mer Island); 47, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 48, A. cornuta Baly, 1879 (Sulawesi); 49, A. indica (Gmelin, 1790) (Laos).Dotted lines indicate depth and width of apical concavity." figureDoi="http://doi.org/10.5281/zenodo.4545472" httpUri="https://zenodo.org/record/4545472/files/figure.png" pageId="11" pageNumber="12">Figs 4547</figureCitation>
,
<figureCitation id="51C189C53F58E5477273C741FD60440F" box="[619,681,1031,1053]" captionStart="FIGURES 5055" captionStartId="26.[151,264,1943,1967]" captionTargetBox="[198,1388,201,1907]" captionTargetId="figure-22@26.[178,1409,181,1919]" captionTargetPageId="26" captionText="FIGURES 5055. Aulacophora species, female, apex of abdominal venter: 50, A. indica (Gmelin, 1790) (Biak Island); 51, A. indica (Vietnam); 52, A. indica (Broken Hill); 53, A. mbabaram Reid, Halling &amp; Beatson, sp. nov., Almaden; 54, A. relicta (Boisduval, 1835); 55, A. wallacii Baly, 1886." figureDoi="http://doi.org/10.5281/zenodo.4545474" httpUri="https://zenodo.org/record/4545474/files/figure.png" pageId="11" pageNumber="12">5355</figureCitation>
,
<figureCitation id="51C189C53F58E54772ABC741FD3B440F" box="[691,754,1031,1053]" captionStart="FIGURES 5869" captionStartId="39.[151,264,1916,1940]" captionTargetBox="[360,1224,192,1879]" captionTargetId="figure-19@39.[347,1241,181,1893]" captionTargetPageId="39" captionText="FIGURES 5869. Apical margin of female sternite VII, Aulacophora abdominalis (Fabricius, 1781) (locality sequence from west to east): 58, East Timor, 59, East Timor, 60, west New Guinea, 61, east New Guinea, 62, New Ireland, 63, New Ireland, 64, Torres Strait, 65, Torres Strait, 66, Guadalcanal, 67, Vanuatu, 68, Fiji, 69, Tonga." figureDoi="http://doi.org/10.5281/zenodo.4545478" httpUri="https://zenodo.org/record/4545478/files/figure.png" pageId="11" pageNumber="12">5869</figureCitation>
,
<figureCitation id="51C189C53F58E54772E2C74EFCF3440C" box="[762,826,1032,1054]" captionStart="FIGURES 7782" captionStartId="41.[151,264,1928,1952]" captionTargetBox="[261,1330,194,1884]" captionTargetId="figure-19@41.[242,1346,181,1905]" captionTargetPageId="41" captionText="FIGURES 7782. Apical margin of female sternite VII, Aulacophora relicta (Boisduval, 1835) (locality sequence from north to south): 77, Adelaide River, NT, 78, Davies Creek, north QLD, 79, Hidden Valley, north QLD, 80, Blackdown Tableland, south QLD, 81, Carnarvon Gorge, south QLD, 82, Ashford, NSW." figureDoi="http://doi.org/10.5281/zenodo.4545482" httpUri="https://zenodo.org/record/4545482/files/figure.png" pageId="11" pageNumber="12">7782</figureCitation>
), with subapical transversely arcuate depression or paired transverse depressions not bounded by lateral ridges, apex of pygidium usually thick in lateral view with small apicodorsal tubercle; pygidium usually less or not acutely produced, apical angle more than 70°, or rounded............................. 15
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</paragraph>
<paragraph id="C94595403F58E547708FC726FA54451A" blockId="11.[151,1437,151,2022]" pageId="11" pageNumber="12">
<keyLead id="720B36753F58E547708FC726FA54451A" pageId="11" pageNumber="12">
- Apical excision of ventrite 5 deep, width at most 1.8x depth (shallowest in Vietnamese specimens), almost symmetrically U- or V-shaped and apical margin with flat edge and pair of preapical ovate depressions, bounded laterally by a ridge which fuses apically with margin of ventrite (
<figureCitation id="51C189C53F58E547723FC7DCFD5144A2" box="[551,664,1178,1200]" captionStart-0="FIGURES 4449" captionStart-1="FIGURES 5055" captionStartId-0="25.[151,264,1798,1822]" captionStartId-1="26.[151,264,1943,1967]" captionTargetBox-0="[170,1418,190,1763]" captionTargetBox-1="[198,1388,201,1907]" captionTargetId-0="figure-19@25.[151,1436,181,1774]" captionTargetId-1="figure-22@26.[178,1409,181,1919]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionText-0="FIGURES 4449. Aulacophora species, female, apex of abdominal venter and pygidium: 44, A. abdominalis (Fabricius, 1781) (Vanuatu); 45, A. abdominalis (New Ireland); 46, A. abdominalis (Mer Island); 47, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 48, A. cornuta Baly, 1879 (Sulawesi); 49, A. indica (Gmelin, 1790) (Laos).Dotted lines indicate depth and width of apical concavity." captionText-1="FIGURES 5055. Aulacophora species, female, apex of abdominal venter: 50, A. indica (Gmelin, 1790) (Biak Island); 51, A. indica (Vietnam); 52, A. indica (Broken Hill); 53, A. mbabaram Reid, Halling &amp; Beatson, sp. nov., Almaden; 54, A. relicta (Boisduval, 1835); 55, A. wallacii Baly, 1886." figureDoi-0="http://doi.org/10.5281/zenodo.4545472" figureDoi-1="http://doi.org/10.5281/zenodo.4545474" httpUri-0="https://zenodo.org/record/4545472/files/figure.png" httpUri-1="https://zenodo.org/record/4545474/files/figure.png" pageId="11" pageNumber="12">Figs 4952</figureCitation>
,
<figureCitation id="51C189C53F58E54772BBC7DCFD2A44A2" box="[675,739,1178,1200]" captionStart="FIGURES 7076" captionStartId="40.[151,264,1960,1984]" captionTargetBox="[269,1316,190,1923]" captionTargetId="figure-22@40.[254,1334,181,1937]" captionTargetPageId="40" captionText="FIGURES 7076. Apical margin of female sternite VII, Aulacophora indica (Gmelin, 1790) (locality sequence from west to east): 70, Laos, 71, Vietnam, 72, Luzon, 73, east Borneo, 74, west Timor, 75, east Timor, 76, west New Guinea (Biak)." figureDoi="http://doi.org/10.5281/zenodo.4545480" httpUri="https://zenodo.org/record/4545480/files/figure.png" pageId="11" pageNumber="12">7076</figureCitation>
); pygidial apex usually thin in lateral view, without tubercle; pygidium usually produced as an elongate straight-sided triangular lobe, apical angle usually 5075°, rarely 7680° (
<figureCitation id="51C189C53F58E54774C6C7F1FA8544DF" box="[1246,1356,1207,1229]" captionStart-0="FIGURES 5869" captionStart-1="FIGURES 7076" captionStartId-0="39.[151,264,1916,1940]" captionStartId-1="40.[151,264,1960,1984]" captionTargetBox-0="[360,1224,192,1879]" captionTargetBox-1="[269,1316,190,1923]" captionTargetId-0="figure-19@39.[347,1241,181,1893]" captionTargetId-1="figure-22@40.[254,1334,181,1937]" captionTargetPageId-0="39" captionTargetPageId-1="40" captionText-0="FIGURES 5869. Apical margin of female sternite VII, Aulacophora abdominalis (Fabricius, 1781) (locality sequence from west to east): 58, East Timor, 59, East Timor, 60, west New Guinea, 61, east New Guinea, 62, New Ireland, 63, New Ireland, 64, Torres Strait, 65, Torres Strait, 66, Guadalcanal, 67, Vanuatu, 68, Fiji, 69, Tonga." captionText-1="FIGURES 7076. Apical margin of female sternite VII, Aulacophora indica (Gmelin, 1790) (locality sequence from west to east): 70, Laos, 71, Vietnam, 72, Luzon, 73, east Borneo, 74, west Timor, 75, east Timor, 76, west New Guinea (Biak)." figureDoi-0="http://doi.org/10.5281/zenodo.4545478" figureDoi-1="http://doi.org/10.5281/zenodo.4545480" httpUri-0="https://zenodo.org/record/4545478/files/figure.png" httpUri-1="https://zenodo.org/record/4545480/files/figure.png" pageId="11" pageNumber="12">Figs 6972</figureCitation>
) (length
<quantity id="0E0238A53F58E54770FAC793FEFD44F8" box="[226,308,1237,1259]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="11" pageNumber="12" unit="mm" value="7.0" valueMax="8.0" valueMin="6.0">68 mm</quantity>
; antennae usually entirely pale, rarely darkened to almost black from antennomere 4 to apex; legs variable in colour; abdominal ventrite V with basal 1/32/3 black)................................................
<taxonomicName id="0EFAEEC33F58E54774F0C7B4FA54451A" authority="(Gmelin)" authorityName="Luzon" baseAuthorityName="Gmelin" baseAuthorityYear="1790" box="[1256,1437,1266,1288]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="FB8E49523F58E54774F0C7B4FAF74515" box="[1256,1342,1266,1287]" italics="true" pageId="11" pageNumber="12">A. indica</emphasis>
(Gmelin)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="720E8DE53F58E547708FC649FA55454D" pageId="11" pageNumber="12">
<paragraph id="C94595403F58E547708FC649FA554551" blockId="11.[151,1437,151,2022]" pageId="11" pageNumber="12">
<keyLead id="720B36753F58E547708FC649FA554551" pageId="11" pageNumber="12">
15(14) Metafemora entirely pale (
<figureCitation id="51C189C53F58E54771F0C649FDEC4537" box="[488,549,1295,1317]" captionStart="FIGURES 69" captionStartId="16.[151,264,1957,1981]" captionTargetBox="[174,1418,193,1917]" captionTargetId="figure-22@16.[157,1430,181,1933]" captionTargetPageId="16" captionText="FIGURES 69. Aulacophora species, male habitus: 6, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 7, A. relicta (Boisduval, 1835) (Mount Molloy); 8, A. wallacii Baly, 1886; 9, A. wilsoni Baly, 1888." figureDoi="http://doi.org/10.5281/zenodo.4545458" httpUri="https://zenodo.org/record/4545458/files/figure.png" pageId="11" pageNumber="12">Fig. 7</figureCitation>
); pygidium produced as a broadly obtuse to narrowly acute triangle (
<figureCitation id="51C189C53F58E54774CCC649FA8F4537" box="[1236,1350,1295,1317]" captionStart="FIGURES 4449" captionStartId="25.[151,264,1798,1822]" captionTargetBox="[170,1418,190,1763]" captionTargetId="figure-19@25.[151,1436,181,1774]" captionTargetPageId="25" captionText="FIGURES 4449. Aulacophora species, female, apex of abdominal venter and pygidium: 44, A. abdominalis (Fabricius, 1781) (Vanuatu); 45, A. abdominalis (New Ireland); 46, A. abdominalis (Mer Island); 47, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 48, A. cornuta Baly, 1879 (Sulawesi); 49, A. indica (Gmelin, 1790) (Laos).Dotted lines indicate depth and width of apical concavity." figureDoi="http://doi.org/10.5281/zenodo.4545472" httpUri="https://zenodo.org/record/4545472/files/figure.png" pageId="11" pageNumber="12">Figs 4446</figureCitation>
,
<figureCitation id="51C189C53F58E547754AC649FA5B4537" box="[1362,1426,1295,1317]" captionStart="FIGURES 5055" captionStartId="26.[151,264,1943,1967]" captionTargetBox="[198,1388,201,1907]" captionTargetId="figure-22@26.[178,1409,181,1919]" captionTargetPageId="26" captionText="FIGURES 5055. Aulacophora species, female, apex of abdominal venter: 50, A. indica (Gmelin, 1790) (Biak Island); 51, A. indica (Vietnam); 52, A. indica (Broken Hill); 53, A. mbabaram Reid, Halling &amp; Beatson, sp. nov., Almaden; 54, A. relicta (Boisduval, 1835); 55, A. wallacii Baly, 1886." figureDoi="http://doi.org/10.5281/zenodo.4545474" httpUri="https://zenodo.org/record/4545474/files/figure.png" pageId="11" pageNumber="12">5354</figureCitation>
); (antennae pale at base including antennomere 4, usually slightly darkened apically)................................ 16
</keyLead>
</paragraph>
<paragraph id="C94595403F58E547708FC60DFA55454D" blockId="11.[151,1437,151,2022]" box="[151,1436,1354,1376]" pageId="11" pageNumber="12">
<keyLead id="720B36753F58E547708FC60DFA55454D" box="[151,1436,1354,1376]" pageId="11" pageNumber="12">
- At least apical third of metafemora dark brown to black (
<figureCitation id="51C189C53F58E547731CC60CFC8A454D" box="[772,835,1354,1376]" captionStart="FIGURES 25" captionStartId="15.[151,264,1829,1853]" captionTargetBox="[164,1420,191,1786]" captionTargetId="figure-19@15.[151,1436,181,1805]" captionTargetPageId="15" captionText="FIGURES 25. Aulacophora species, male habitus: 2, A. abdominalis (Fabricius, 1781) (Guadalcanal); 3, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 4, A. cornuta Baly, 1879 (New Ireland); 5, A. indica (Gmelin, 1790) (Luzon)." figureDoi="http://doi.org/10.5281/zenodo.4545452" httpUri="https://zenodo.org/record/4545452/files/figure.png" pageId="11" pageNumber="12">Figs 2</figureCitation>
,
<figureCitation id="51C189C53F58E5477355C60CFC934572" box="[845,858,1354,1376]" captionStart="FIGURES 69" captionStartId="16.[151,264,1957,1981]" captionTargetBox="[174,1418,193,1917]" captionTargetId="figure-22@16.[157,1430,181,1933]" captionTargetPageId="16" captionText="FIGURES 69. Aulacophora species, male habitus: 6, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 7, A. relicta (Boisduval, 1835) (Mount Molloy); 8, A. wallacii Baly, 1886; 9, A. wilsoni Baly, 1888." figureDoi="http://doi.org/10.5281/zenodo.4545458" httpUri="https://zenodo.org/record/4545458/files/figure.png" pageId="11" pageNumber="12">8</figureCitation>
); pygidium variable in shape...................... 17
</keyLead>
</paragraph>
</keyStep>
<keyStep id="720E8DE53F58E547708FC621FA554602" pageId="11" pageNumber="12">
<paragraph id="C94595403F58E547708FC621FA5445AA" blockId="11.[151,1437,151,2022]" pageId="11" pageNumber="12">
<keyLead id="720B36753F58E547708FC621FA5445AA" pageId="11" pageNumber="12">
16(15) Abdominal tergites brownish-yellow, including whole of pygidium; ventrite 5 with brownish-yellow middle of base (
<figureCitation id="51C189C53F58E5477557C621FA5C456F" box="[1359,1429,1383,1405]" captionStart="FIGURES 5055" captionStartId="26.[151,264,1943,1967]" captionTargetBox="[198,1388,201,1907]" captionTargetId="figure-22@26.[178,1409,181,1919]" captionTargetPageId="26" captionText="FIGURES 5055. Aulacophora species, female, apex of abdominal venter: 50, A. indica (Gmelin, 1790) (Biak Island); 51, A. indica (Vietnam); 52, A. indica (Broken Hill); 53, A. mbabaram Reid, Halling &amp; Beatson, sp. nov., Almaden; 54, A. relicta (Boisduval, 1835); 55, A. wallacii Baly, 1886." figureDoi="http://doi.org/10.5281/zenodo.4545474" httpUri="https://zenodo.org/record/4545474/files/figure.png" pageId="11" pageNumber="12">Fig. 54</figureCitation>
) (length
<quantity id="0E0238A53F58E5477134C6C3FE6B4588" box="[300,418,1413,1435]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.5" metricValueMin="5.5" pageId="11" pageNumber="12" unit="mm" value="6.5" valueMax="7.5" valueMin="5.5">5.57.5 mm</quantity>
; pygidium usually produced as ± 90º triangle, rarely rounded, usually with acute apicodorsal tubercle).....................................................................................
<taxonomicName id="0EFAEEC33F58E54774D4C6E4FA5445AA" authority="(Boisduval)" baseAuthorityName="Boisduval" box="[1228,1437,1442,1464]" class="Insecta" family="Chrysomelidae" genus="Galleruca" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="relicta">
<emphasis id="FB8E49523F58E54774D4C6E4FAEC45A5" box="[1228,1317,1442,1463]" italics="true" pageId="11" pageNumber="12">A. relicta</emphasis>
(Boisduval)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C94595403F58E547708FC686FA554602" blockId="11.[151,1437,151,2022]" pageId="11" pageNumber="12">
<keyLead id="720B36753F58E547708FC686FA554602" pageId="11" pageNumber="12">
- Base of pygidium darker than brownish-yellow apex and remainder of abdominal dorsum black or dark brown; ventrite 5 with brown base (
<figureCitation id="51C189C53F58E5477147C69BFE0745E1" box="[351,462,1501,1523]" captionStart="FIGURES 4449" captionStartId="25.[151,264,1798,1822]" captionTargetBox="[170,1418,190,1763]" captionTargetId="figure-19@25.[151,1436,181,1774]" captionTargetPageId="25" captionText="FIGURES 4449. Aulacophora species, female, apex of abdominal venter and pygidium: 44, A. abdominalis (Fabricius, 1781) (Vanuatu); 45, A. abdominalis (New Ireland); 46, A. abdominalis (Mer Island); 47, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 48, A. cornuta Baly, 1879 (Sulawesi); 49, A. indica (Gmelin, 1790) (Laos).Dotted lines indicate depth and width of apical concavity." figureDoi="http://doi.org/10.5281/zenodo.4545472" httpUri="https://zenodo.org/record/4545472/files/figure.png" pageId="11" pageNumber="12">Figs 4446</figureCitation>
) (length
<quantity id="0E0238A53F58E547723FC69BFDB345E0" box="[551,634,1501,1522]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="11" pageNumber="12" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">78 mm</quantity>
; apex pygidium usually produced as slightly elongate triangular lobe, apical angle usually 7690̊, rarely 90100º, acute tubercle, if present, usually apical).
<taxonomicName id="0EFAEEC33F58E54773B8C6BCFB694602" authority="(Fabricius)" authorityName="Mer" baseAuthorityName="Fabricius" baseAuthorityYear="1781" box="[928,1184,1530,1552]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis id="FB8E49523F58E54773B8C6BCFBFB461D" box="[928,1074,1530,1551]" italics="true" pageId="11" pageNumber="12">A. abdominalis</emphasis>
(Fabricius)
</taxonomicName>
(Timorese colour variety)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="720E8DE53F58E547708FC551FA5546E8" pageId="11" pageNumber="12">
<paragraph id="C94595403F58E547708FC551FA554697" blockId="11.[151,1437,151,2022]" pageId="11" pageNumber="12">
<keyLead id="720B36753F58E547708FC551FA554697" pageId="11" pageNumber="12">
17(15) Antennomeres 13 pale and 411 dark brown to black (
<figureCitation id="51C189C53F58E54772E6C551FCA6463C" box="[766,879,1559,1582]" captionStart="FIGURES 1825" captionStartId="21.[151,264,1899,1923]" captionTargetBox="[262,1328,190,1869]" captionTargetId="figure-19@21.[250,1338,181,1875]" captionTargetPageId="21" captionText="FIGURES 1825. Aulacophora species, male anterior and antenna: 18, A. abdominalis (Fabricius, 1781) (Guadalcanal); 19, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 20, A. cornuta Baly, 1879 (New Ireland); 21, A. indica (Gmelin, 1790) (Luzon); 22, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 23, A. relicta (Boisduval, 1835) (Mount Molloy); 24, A. wallacii Baly, 1886; 25, A. wilsoni Baly, 1888." figureDoi="http://doi.org/10.5281/zenodo.4545464" httpUri="https://zenodo.org/record/4545464/files/figure.png" pageId="11" pageNumber="12">Figs 19, 22</figureCitation>
); pygidium narrowly produced, apical margin rounded or almost truncate but with small median tubercle (
<figureCitation id="51C189C53F58E54772ADC573FCC94658" box="[693,768,1589,1611]" captionStart="FIGURES 4449" captionStartId="25.[151,264,1798,1822]" captionTargetBox="[170,1418,190,1763]" captionTargetId="figure-19@25.[151,1436,181,1774]" captionTargetPageId="25" captionText="FIGURES 4449. Aulacophora species, female, apex of abdominal venter and pygidium: 44, A. abdominalis (Fabricius, 1781) (Vanuatu); 45, A. abdominalis (New Ireland); 46, A. abdominalis (Mer Island); 47, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 48, A. cornuta Baly, 1879 (Sulawesi); 49, A. indica (Gmelin, 1790) (Laos).Dotted lines indicate depth and width of apical concavity." figureDoi="http://doi.org/10.5281/zenodo.4545472" httpUri="https://zenodo.org/record/4545472/files/figure.png" pageId="11" pageNumber="12">Figs 47</figureCitation>
,
<figureCitation id="51C189C53F58E5477313C573FCEF4659" box="[779,806,1589,1611]" captionStart="FIGURES 5055" captionStartId="26.[151,264,1943,1967]" captionTargetBox="[198,1388,201,1907]" captionTargetId="figure-22@26.[178,1409,181,1919]" captionTargetPageId="26" captionText="FIGURES 5055. Aulacophora species, female, apex of abdominal venter: 50, A. indica (Gmelin, 1790) (Biak Island); 51, A. indica (Vietnam); 52, A. indica (Broken Hill); 53, A. mbabaram Reid, Halling &amp; Beatson, sp. nov., Almaden; 54, A. relicta (Boisduval, 1835); 55, A. wallacii Baly, 1886." figureDoi="http://doi.org/10.5281/zenodo.4545474" httpUri="https://zenodo.org/record/4545474/files/figure.png" pageId="11" pageNumber="12">53</figureCitation>
); apex of ventrite 5 shallowly arcuate and not or weakly reflexed (meso- and metafemora usually with basal third pale and apical third dark brown, middle third variable; meso- and metatibiae dark brown)........................................................................................ 18
</keyLead>
</paragraph>
<paragraph id="C94595403F58E547708FC5CBFA5546E8" blockId="11.[151,1437,151,2022]" pageId="11" pageNumber="12">
<keyLead id="720B36753F58E547708FC5CBFA5546E8" pageId="11" pageNumber="12">
- Antennae entirely pale or gradually apically darkened, sometimes with antennomeres 611 brown to dark brown but at least inner surface of 4 pale (
<figureCitation id="51C189C53F58E54771D9C5ECFDE746D2" box="[449,558,1706,1728]" captionStart="FIGURES 1825" captionStartId="21.[151,264,1899,1923]" captionTargetBox="[262,1328,190,1869]" captionTargetId="figure-19@21.[250,1338,181,1875]" captionTargetPageId="21" captionText="FIGURES 1825. Aulacophora species, male anterior and antenna: 18, A. abdominalis (Fabricius, 1781) (Guadalcanal); 19, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 20, A. cornuta Baly, 1879 (New Ireland); 21, A. indica (Gmelin, 1790) (Luzon); 22, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 23, A. relicta (Boisduval, 1835) (Mount Molloy); 24, A. wallacii Baly, 1886; 25, A. wilsoni Baly, 1888." figureDoi="http://doi.org/10.5281/zenodo.4545464" httpUri="https://zenodo.org/record/4545464/files/figure.png" pageId="11" pageNumber="12">Figs 18, 24</figureCitation>
); pygidium triangularly produced, or apex almost rounded (
<figureCitation id="51C189C53F58E547747DC5ECFB1A46D2" box="[1125,1235,1706,1728]" captionStart="FIGURES 4449" captionStartId="25.[151,264,1798,1822]" captionTargetBox="[170,1418,190,1763]" captionTargetId="figure-19@25.[151,1436,181,1774]" captionTargetPageId="25" captionText="FIGURES 4449. Aulacophora species, female, apex of abdominal venter and pygidium: 44, A. abdominalis (Fabricius, 1781) (Vanuatu); 45, A. abdominalis (New Ireland); 46, A. abdominalis (Mer Island); 47, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 48, A. cornuta Baly, 1879 (Sulawesi); 49, A. indica (Gmelin, 1790) (Laos).Dotted lines indicate depth and width of apical concavity." figureDoi="http://doi.org/10.5281/zenodo.4545472" httpUri="https://zenodo.org/record/4545472/files/figure.png" pageId="11" pageNumber="12">Figs 4446</figureCitation>
,
<figureCitation id="51C189C53F58E54774C4C5ECFB3E46D2" box="[1244,1271,1706,1728]" captionStart="FIGURES 5055" captionStartId="26.[151,264,1943,1967]" captionTargetBox="[198,1388,201,1907]" captionTargetId="figure-22@26.[178,1409,181,1919]" captionTargetPageId="26" captionText="FIGURES 5055. Aulacophora species, female, apex of abdominal venter: 50, A. indica (Gmelin, 1790) (Biak Island); 51, A. indica (Vietnam); 52, A. indica (Broken Hill); 53, A. mbabaram Reid, Halling &amp; Beatson, sp. nov., Almaden; 54, A. relicta (Boisduval, 1835); 55, A. wallacii Baly, 1886." figureDoi="http://doi.org/10.5281/zenodo.4545474" httpUri="https://zenodo.org/record/4545474/files/figure.png" pageId="11" pageNumber="12">55</figureCitation>
); apex of ventrite 5 variably excised, usually strongly reflexed (metafemora usually entirely dark brown to black; ventrite 5 with at least basal third dark brown or black)............................................................................. 19
</keyLead>
</paragraph>
</keyStep>
<keyStep id="720E8DE53F58E547708FC444FA5547B8" pageId="11" pageNumber="12">
<paragraph id="C94595403F58E547708FC444FA554740" blockId="11.[151,1437,151,2022]" pageId="11" pageNumber="12">
<keyLead id="720B36753F58E547708FC444FA554740" pageId="11" pageNumber="12">
18(17) Pronotal groove laterally deepened; abdominal ventrites 14 with basal half yellowish-brown, ventrite 5 yellow except for laterobasal dark patches (
<figureCitation id="51C189C53F58E54771CEC459FDD44724" box="[470,541,1823,1846]" captionStart="FIGURES 4449" captionStartId="25.[151,264,1798,1822]" captionTargetBox="[170,1418,190,1763]" captionTargetId="figure-19@25.[151,1436,181,1774]" captionTargetPageId="25" captionText="FIGURES 4449. Aulacophora species, female, apex of abdominal venter and pygidium: 44, A. abdominalis (Fabricius, 1781) (Vanuatu); 45, A. abdominalis (New Ireland); 46, A. abdominalis (Mer Island); 47, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 48, A. cornuta Baly, 1879 (Sulawesi); 49, A. indica (Gmelin, 1790) (Laos).Dotted lines indicate depth and width of apical concavity." figureDoi="http://doi.org/10.5281/zenodo.4545472" httpUri="https://zenodo.org/record/4545472/files/figure.png" pageId="11" pageNumber="12">Fig. 47</figureCitation>
); pygidium brownish-yellow (length
<quantity id="0E0238A53F58E547739CC466FC1C4727" box="[900,981,1824,1845]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="11" pageNumber="12" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">78 mm</quantity>
)....................................................................................................
<taxonomicName id="0EFAEEC33F58E54773C5C47BFA8F4740" authority="Reid, Halling &amp; Beatson" authorityName="Reid, Halling &amp; Beatson" authorityYear="2021" box="[989,1350,1853,1875]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="barrogae" status="sp. nov.">
<emphasis id="FB8E49523F58E54773C5C47BFB994740" box="[989,1104,1853,1874]" italics="true" pageId="11" pageNumber="12">A. barrogae</emphasis>
Reid, Halling &amp; Beatson
</taxonomicName>
,
<emphasis id="FB8E49523F58E5477548C47BFA554740" bold="true" box="[1360,1436,1853,1874]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="E0BDF4293F58E5477548C47BFA554740" box="[1360,1436,1853,1874]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="C94595403F58E547708FC41DFA5547B8" blockId="11.[151,1437,151,2022]" pageId="11" pageNumber="12">
<keyLead id="720B36753F58E547708FC41DFA5547B8" pageId="11" pageNumber="12">
- Pronotal groove entirely shallow; abdominal ventrites 14 black with brown apical margins, basal half or more of ventrite 5 dark brown to black (
<figureCitation id="51C189C53F58E54771A8C431FE3E479F" box="[432,503,1911,1933]" captionStart="FIGURES 5055" captionStartId="26.[151,264,1943,1967]" captionTargetBox="[198,1388,201,1907]" captionTargetId="figure-22@26.[178,1409,181,1919]" captionTargetPageId="26" captionText="FIGURES 5055. Aulacophora species, female, apex of abdominal venter: 50, A. indica (Gmelin, 1790) (Biak Island); 51, A. indica (Vietnam); 52, A. indica (Broken Hill); 53, A. mbabaram Reid, Halling &amp; Beatson, sp. nov., Almaden; 54, A. relicta (Boisduval, 1835); 55, A. wallacii Baly, 1886." figureDoi="http://doi.org/10.5281/zenodo.4545474" httpUri="https://zenodo.org/record/4545474/files/figure.png" pageId="11" pageNumber="12">Fig. 53</figureCitation>
); pygidium partly dark brown (length
<quantity id="0E0238A53F58E547737FC431FC03479F" box="[871,970,1911,1933]" metricMagnitude="-3" metricUnit="m" metricValue="7.25" metricValueMax="7.5" metricValueMin="7.0" pageId="11" pageNumber="12" unit="mm" value="7.25" valueMax="7.5" valueMin="7.0">77.5 mm</quantity>
)....................................................................................................
<taxonomicName id="0EFAEEC33F58E54773D6C4D3FA8F47B8" authority="Reid, Halling &amp; Beatson" authorityName="Reid, Halling &amp; Beatson" authorityYear="2021" box="[974,1350,1941,1963]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="mbabaram" status="sp. nov.">
<emphasis id="FB8E49523F58E54773D6C4D3FB9947B8" box="[974,1104,1941,1962]" italics="true" pageId="11" pageNumber="12">A. mbabaram</emphasis>
Reid, Halling &amp; Beatson
</taxonomicName>
,
<emphasis id="FB8E49523F58E5477548C4D3FA5547B8" bold="true" box="[1360,1436,1941,1962]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="E0BDF4293F58E5477548C4D3FA5547B8" box="[1360,1436,1941,1962]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="720E8DE53F58E540708FC4F4FA554131" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<paragraph id="C94595403F58E540708FC4F4FA5540D9" blockId="11.[151,1437,151,2022]" lastBlockId="12.[151,1437,151,291]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<keyLead id="720B36753F58E540708FC4F4FA5540D9" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
19(15) Venter of pygidial apex flat or shallowly concave, and apex pygidium produced as a triangular lobe, apical angle usually 7690̊, rarely 90100º (
<figureCitation id="51C189C53F58E5477164C489FE2547F7" box="[380,492,1999,2021]" captionStart="FIGURES 4449" captionStartId="25.[151,264,1798,1822]" captionTargetBox="[170,1418,190,1763]" captionTargetId="figure-19@25.[151,1436,181,1774]" captionTargetPageId="25" captionText="FIGURES 4449. Aulacophora species, female, apex of abdominal venter and pygidium: 44, A. abdominalis (Fabricius, 1781) (Vanuatu); 45, A. abdominalis (New Ireland); 46, A. abdominalis (Mer Island); 47, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 48, A. cornuta Baly, 1879 (Sulawesi); 49, A. indica (Gmelin, 1790) (Laos).Dotted lines indicate depth and width of apical concavity." figureDoi="http://doi.org/10.5281/zenodo.4545472" httpUri="https://zenodo.org/record/4545472/files/figure.png" pageId="11" pageNumber="12">Figs 4446</figureCitation>
); metafemora and tibiae dark brown, or tibiae slightly paler (
<figureCitation id="51C189C53F58E547742AC489FBA447F4" box="[1074,1133,1999,2022]" captionStart="FIGURES 25" captionStartId="15.[151,264,1829,1853]" captionTargetBox="[164,1420,191,1786]" captionTargetId="figure-19@15.[151,1436,181,1805]" captionTargetPageId="15" captionText="FIGURES 25. Aulacophora species, male habitus: 2, A. abdominalis (Fabricius, 1781) (Guadalcanal); 3, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 4, A. cornuta Baly, 1879 (New Ireland); 5, A. indica (Gmelin, 1790) (Luzon)." figureDoi="http://doi.org/10.5281/zenodo.4545452" httpUri="https://zenodo.org/record/4545452/files/figure.png" pageId="11" pageNumber="12">Fig. 2</figureCitation>
); scutellum same colour as ely- tra; ventrite 5 marginal excavation of variable depth (
<figureCitation id="51C189C53F5FE54072C0C3D1FC8C40BF" box="[728,837,151,173]" captionStart="FIGURES 5869" captionStartId="39.[151,264,1916,1940]" captionTargetBox="[360,1224,192,1879]" captionTargetId="figure-19@39.[347,1241,181,1893]" captionTargetPageId="39" captionText="FIGURES 5869. Apical margin of female sternite VII, Aulacophora abdominalis (Fabricius, 1781) (locality sequence from west to east): 58, East Timor, 59, East Timor, 60, west New Guinea, 61, east New Guinea, 62, New Ireland, 63, New Ireland, 64, Torres Strait, 65, Torres Strait, 66, Guadalcanal, 67, Vanuatu, 68, Fiji, 69, Tonga." figureDoi="http://doi.org/10.5281/zenodo.4545478" httpUri="https://zenodo.org/record/4545478/files/figure.png" pageId="12" pageNumber="13">Figs 5869</figureCitation>
) and apical margin not or weakly reflexed (
<figureCitation id="51C189C53F5FE54074F8C3D1FA8440BF" box="[1248,1357,151,173]" captionStart="FIGURES 4449" captionStartId="25.[151,264,1798,1822]" captionTargetBox="[170,1418,190,1763]" captionTargetId="figure-19@25.[151,1436,181,1774]" captionTargetPageId="25" captionText="FIGURES 4449. Aulacophora species, female, apex of abdominal venter and pygidium: 44, A. abdominalis (Fabricius, 1781) (Vanuatu); 45, A. abdominalis (New Ireland); 46, A. abdominalis (Mer Island); 47, A. barrogae Reid, Halling &amp; Beatson, sp. nov.; 48, A. cornuta Baly, 1879 (Sulawesi); 49, A. indica (Gmelin, 1790) (Laos).Dotted lines indicate depth and width of apical concavity." figureDoi="http://doi.org/10.5281/zenodo.4545472" httpUri="https://zenodo.org/record/4545472/files/figure.png" pageId="12" pageNumber="13">Figs 4446</figureCitation>
) (length
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)......................................................
<taxonomicName id="0EFAEEC33F5FE54073D4C3F3FB0440D9" authority="(Fabricius)" authorityName="Mer" baseAuthorityName="Fabricius" baseAuthorityYear="1781" box="[972,1229,181,203]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis id="FB8E49523F5FE54073D4C3F3FB9740D8" box="[972,1118,181,202]" italics="true" pageId="12" pageNumber="13">A. abdominalis</emphasis>
(Fabricius)
</taxonomicName>
(typical colour form)
</keyLead>
</paragraph>
<paragraph id="C94595403F5FE540708FC395FA554131" blockId="12.[151,1437,151,291]" pageId="12" pageNumber="13">
<keyLead id="720B36753F5FE540708FC395FA554131" pageId="12" pageNumber="13">
- Venter of pygidial apex deeply notched, and apex of pygidium almost rounded (
<figureCitation id="51C189C53F5FE54073FDC394FBE740FA" box="[997,1070,210,232]" captionStart="FIGURES 5055" captionStartId="26.[151,264,1943,1967]" captionTargetBox="[198,1388,201,1907]" captionTargetId="figure-22@26.[178,1409,181,1919]" captionTargetPageId="26" captionText="FIGURES 5055. Aulacophora species, female, apex of abdominal venter: 50, A. indica (Gmelin, 1790) (Biak Island); 51, A. indica (Vietnam); 52, A. indica (Broken Hill); 53, A. mbabaram Reid, Halling &amp; Beatson, sp. nov., Almaden; 54, A. relicta (Boisduval, 1835); 55, A. wallacii Baly, 1886." figureDoi="http://doi.org/10.5281/zenodo.4545474" httpUri="https://zenodo.org/record/4545474/files/figure.png" pageId="12" pageNumber="13">Fig. 55</figureCitation>
); metafemora black strongly contrasting with yellow metatibiae (
<figureCitation id="51C189C53F5FE54071ECC3A9FDE74114" box="[500,558,239,262]" captionStart="FIGURES 69" captionStartId="16.[151,264,1957,1981]" captionTargetBox="[174,1418,193,1917]" captionTargetId="figure-22@16.[157,1430,181,1933]" captionTargetPageId="16" captionText="FIGURES 69. Aulacophora species, male habitus: 6, A. mbabaram Reid, Halling &amp; Beatson, sp. nov.; 7, A. relicta (Boisduval, 1835) (Mount Molloy); 8, A. wallacii Baly, 1886; 9, A. wilsoni Baly, 1888." figureDoi="http://doi.org/10.5281/zenodo.4545458" httpUri="https://zenodo.org/record/4545458/files/figure.png" pageId="12" pageNumber="13">Fig. 8</figureCitation>
); scutellum usually darker than elytra; apex of ventrite 5 narrowly excavate at middle and strongly reflexed (
<figureCitation id="51C189C53F5FE540718AC24BFE114131" box="[402,472,269,291]" captionStart="FIGURES 5055" captionStartId="26.[151,264,1943,1967]" captionTargetBox="[198,1388,201,1907]" captionTargetId="figure-22@26.[178,1409,181,1919]" captionTargetPageId="26" captionText="FIGURES 5055. Aulacophora species, female, apex of abdominal venter: 50, A. indica (Gmelin, 1790) (Biak Island); 51, A. indica (Vietnam); 52, A. indica (Broken Hill); 53, A. mbabaram Reid, Halling &amp; Beatson, sp. nov., Almaden; 54, A. relicta (Boisduval, 1835); 55, A. wallacii Baly, 1886." figureDoi="http://doi.org/10.5281/zenodo.4545474" httpUri="https://zenodo.org/record/4545474/files/figure.png" pageId="12" pageNumber="13">Fig. 55</figureCitation>
) (length
<quantity id="0E0238A53F5FE5407237C24BFD4B4130" box="[559,642,269,290]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="12" pageNumber="13" unit="mm" value="8.5" valueMax="9.0" valueMin="8.0">89 mm</quantity>
;)....................................................
<taxonomicName id="0EFAEEC33F5FE5407519C24BFA554131" authority="Baly" authorityName="Baly" authorityYear="1886" box="[1281,1436,269,291]" class="Insecta" family="Aphididae" genus="Aulacophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="wallacii">
<emphasis id="FB8E49523F5FE5407519C24BFAA04130" box="[1281,1385,269,290]" italics="true" pageId="12" pageNumber="13">A. wallacii</emphasis>
Baly
</taxonomicName>
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</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>