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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.44.8693" ID-PMC="PMC4329390" ID-Pensoft-Pub="1314-2003-44-39" ID-Pensoft-UUID="2F7D0B61EB037257FFD5B7587923FFB7" ID-PubMed="25698893" ID-Zenodo-Dep="576261" ModsDocID="1314-2003-44-39" checkinTime="1451251619874" checkinUser="pensoft" docAuthor="Saerkinen, Tiina, Gonzales, Paul &amp; Knapp, Sandra" docDate="2015" docId="3FFB8F97D06E5E629C75CB29F1CF9BDF" docLanguage="en" docName="PhytoKeys 44: 39-64" docOrigin="PhytoKeys 44" docPubDate="2015-01-13" docSource="http://dx.doi.org/10.3897/phytokeys.44.8693" docTitle="Solanum antisuyo Saerkinen &amp; S. Knapp 2015, sp. nov." docType="treatment" docVersion="3" id="2F7D0B61EB037257FFD5B7587923FFB7" lastPageNumber="49" masterDocId="2F7D0B61EB037257FFD5B7587923FFB7" masterDocTitle="Four new non-spiny Solanum (Solanaceae) species from South America" masterLastPageNumber="64" masterPageNumber="39" pageNumber="43" updateTime="1668141277134" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Four new non-spiny Solanum (Solanaceae) species from South America</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Saerkinen, Tiina</mods:namePart>
<mods:affiliation>Royal Botanic Garden Edinburgh, 20 A Inverleith Row, EH 3 5 LR Edinburgh, United Kingdom</mods:affiliation>
<mods:nameIdentifier type="email">tiinasarkinen@yahoo.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gonzales, Paul</mods:namePart>
<mods:affiliation>Laboratorio de Floristica, Departamento de Dicotiledoneas, Museo de Historia Natural - Universidad Nacional Mayor de San Marcos, Avenida Arenales 1256, Apartado Postal 14 - 0434, Lima, Peru</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Knapp, Sandra</mods:namePart>
<mods:affiliation>Department of Life Sciences, Natural History Museum, Cromwell Road, SW 7 5 BD London, United Kingdom</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2015</mods:date>
<mods:detail type="pubDate">
<mods:number>2015-01-13</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>44</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>39</mods:start>
<mods:end>64</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/phytokeys.44.8693</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.44.8693</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-44-39</mods:identifier>
<mods:identifier type="Pensoft-UUID">2F7D0B61EB037257FFD5B7587923FFB7</mods:identifier>
<mods:identifier type="Zenodo-Dep">576261</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="182228482" LSID="urn:lsid:plazi:treatment:3FFB8F97D06E5E629C75CB29F1CF9BDF" httpUri="http://treatment.plazi.org/id/3FFB8F97D06E5E629C75CB29F1CF9BDF" lastPageId="10" lastPageNumber="49" pageId="4" pageNumber="43">
<subSubSection pageId="4" pageNumber="43" type="nomenclature">
<paragraph pageId="4" pageNumber="43">
<taxonomicName LSID="3FFB8F97-D06E-5E62-9C75-CB29F1CF9BDF" authority="Särkinen &amp; S. Knapp" authorityName="Saerkinen &amp; S. Knapp" authorityYear="2015" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum antisuyo" order="Solanales" pageId="4" pageNumber="43" phylum="Tracheophyta" rank="species" species="antisuyo" status="sp. nov.">
Solanum antisuyo
<normalizedToken originalValue="Särkinen">Saerkinen</normalizedToken>
&amp; S.Knapp
</taxonomicName>
<taxonomicNameLabel pageId="4" pageNumber="43">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Photos of Solanum antisuyo. A Buds and flowers, showing the distinct calyx with long tube and minute but thick purple coloured lobes (Knapp et al. 10399) B Habit, growing in a rocky land slide in gravel (Knapp et al. 10399) C Woody pedicels of the infrutescence, distinct character in herbarium specimens (Knapp et al. 10401) D Ellipsoid fruits with reflexed pedicels, and the characteristic appressed calyx lobes that split into two in fruit (Knapp et al. 10435). Scale bars = 2 mm. All photos by S. Knapp." pageId="4" pageNumber="43">Figs 3</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Distribution map of Solanum antisuyo, endemic species to Peru." pageId="4" pageNumber="43">-4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="45" pageId="5" pageNumber="44" type="diagnosis">
<paragraph pageId="5" pageNumber="44">
<pageBreakToken pageId="5" pageNumber="44" start="start">Diagnosis</pageBreakToken>
.
</paragraph>
<paragraph lastPageId="6" lastPageNumber="45" pageId="5" pageNumber="44">
Like
<taxonomicName authorityName="Bitter" authorityYear="1912" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum polytrichostylum" order="Solanales" pageId="6" pageNumber="45" phylum="Tracheophyta" rank="species" species="polytrichostylum">
<emphasis italics="true" pageId="6" pageNumber="45">
<pageBreakToken pageId="6" pageNumber="45" start="start">Solanum</pageBreakToken>
polytrichostylum
</emphasis>
</taxonomicName>
Bitter, but differing in having either simple or once-branched inflorescences with pedicels spaced c. 1-3 mm apart along the rachis, more reduced, minute calyx lobes, ellipsoid rather than spherical fruits, and larger brown-coloured seeds.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="45" type="type">
<paragraph pageId="6" pageNumber="45">Type.</paragraph>
<paragraph pageId="6" pageNumber="45">
<emphasis bold="true" pageId="6" pageNumber="45">Peru. Cusco:</emphasis>
Prov. Paucartambo, 1 km from Puesto de Vigilancia of Parque Nacional de Manu on rd from Paucartambo to Pilcopata coming from Puesto,
<geoCoordinate degrees="13" direction="south" minutes="12" orientation="latitude" precision="15" seconds="05" value="-13.201388">13°12'05&quot;S</geoCoordinate>
,
<geoCoordinate degrees="71" direction="west" minutes="37" orientation="longitude" precision="15" seconds="21" value="-71.622505">71°37'21&quot;W</geoCoordinate>
, 3480 m, 15 Mar 2012 (fl,fr),
<emphasis italics="true" pageId="6" pageNumber="45">
S. Knapp, P.
<normalizedToken originalValue="Gonzáles">Gonzales</normalizedToken>
, A. Matthews &amp; T.
<normalizedToken originalValue="Särkinen">Saerkinen</normalizedToken>
10435
</emphasis>
(holotype: USM; isotypes: BM [BM001114929], F, HUSA, HUT, MO).
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="46" pageId="6" pageNumber="45" type="description">
<paragraph pageId="6" pageNumber="45">Description.</paragraph>
<paragraph lastPageId="7" lastPageNumber="46" pageId="6" pageNumber="45">
Stout herb to a shrub up to 1.5 m tall, much branching at base, the individual branches up to 1m long. Stems 2-ridged or slightly winged especially towards base, 0.4-0.6 cm in diameter, purple-coloured especially at leaf nodes, nearly glabrous, sparsely pubescent with simple uniseriate, much reduced 1-3-celled trichomes
<pageBreakToken pageId="7" pageNumber="46" start="start">especially</pageBreakToken>
on the often purple coloured young growth. Sympodial units difoliate, not geminate. Leaves simple, 2-17 cm long, 1.2-8.4 cm wide, broadly ovate-lanceolate, membranous to somewhat fleshy; adaxial and abaxial surfaces sparsely pubescent with more or less appressed 1-3-celled simple uniseriate trichomes 0.1-0.2 mm long; primary veins 7-10 pairs; base rounded, decurrent on the petiole; margins entire, often purple tinged; apex acute to acuminate; petiole 0.3-1.2 cm long, occasionally narrowly winged, sparsely pubescent with simple uniseriate trichomes like those of the stems and leaves. Inflorescences 1.4-4.0 cm long, lateral and internodal, simple or once-branched, with 5-14 flowers arising very close together, sparsely pubescent with appressed 1-2-celled simple uniseriate trichomes similar to those on stem and leaves; peduncle 1.0-3.3 cm long, if the inflorescence branched then the peduncle rachis 0.2-0.4 cm long, short and congested; pedicels 1.0-1.2 cm long, 0.5-0.6 mm in diameter at the base tapering gradually to 1.0-1.2 mm in diameter at apex, straight and spreading at anthesis, recurving and becoming woody in fruit, not dehiscing; pedicel scars spaced 0-2 mm apart. Buds conical-ellipsoid, cream-coloured, the corolla strongly exerted from the calyx tube before anthesis. Flowers 5-merous, all perfect; calyx tube 1.5-2.0 mm long, green, the lobes 0.7-0.9 mm long, broadly deltate with rounded apices, purple coloured, sparsely pubescent with 1-celled simple uniseriate trichomes; corolla 12-24 mm in diameter, stellate, white or rarely lilac with a yellow to yellow-green central star at the base, lobed slightly less than halfway to the base, the lobes ca. 9-15 mm long, 4-5 mm wide, spreading to reflexed at anthesis, pubescent abaxially with 1-3-celled simple uniseriate trichomes shorter than the trichomes of the stems and leaves, sparsely pubescent adaxially at base near the filaments with 5-7-celled simple uniseriate trichomes; filament tube ca. 2 mm long, adaxially pubescent with 5-7-celled simple uniseriate trichomes; free portion of the filaments ca. 2 mm long, sometimes slightly longer in two lowermost anthers at anthesis (elongating after anthesis?), pubescent like the tube; anthers ca. (2.8)3.0-3.4 mm long, 1 mm wide, ellipsoid, yellow, poricidal at the tips, the pores lengthening to slits with age; ovary cylindrical, pubescent 2/3 from the base with 2-3-celled simple uniseriate trichomes; style 6 mm long, exerted (0.5)1-2 mm beyond the anther cone, densely pubescent up to 2/3 of the length with 2-3-celled simple uniseriate trichomes at the base; stigma globose, minutely papillate, pale yellow in live plants. Fruit an ellipsoid berry, 8-11 mm in diameter, green turning translucent green-orange when ripe (purple-black in
<emphasis italics="true" pageId="7" pageNumber="46">Knapp et al. 10404</emphasis>
but these affected by pathogens?), the surface smooth and shiny when young, with relatively thick pericarp ca. 0.1 mm; fruiting peduncle woody; fruiting pedicels 11-22 mm long, purple coloured, ca. 1 mm in diameter at the base and 1.5 mm at apex, reflexed and woody in fruit, remaining on the plant after fruit drops; fruiting calyx lobes tightly appressed to the berry, purple-coloured, calyx often splitting into two larger lobes. Seeds 35-45 per berry, ca. 1.1 mm long, ca. 1.7 mm wide, concave-reniform, narrower at one end, brown, the hilum positioned sub-laterally towards the narrower end, the testal cells pentagonal in outline; stone cells few per fruit.
</paragraph>
<caption pageId="7" pageNumber="46" start="Figure 3" startId="F3">
<paragraph pageId="7" pageNumber="46">
<emphasis bold="true" pageId="7" pageNumber="46">Figure 3.</emphasis>
Photos of
<taxonomicName authorityName="Saerkinen &amp; S. Knapp" authorityYear="2015" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum antisuyo" order="Solanales" pageId="7" pageNumber="46" phylum="Tracheophyta" rank="species" species="antisuyo">
<emphasis italics="true" pageId="7" pageNumber="46">Solanum antisuyo</emphasis>
</taxonomicName>
.
<emphasis bold="true" pageId="7" pageNumber="46">A</emphasis>
Buds and flowers, showing the distinct calyx with long tube and minute but thick purple coloured lobes (
<emphasis italics="true" pageId="7" pageNumber="46">Knapp et al.</emphasis>
10399)
<emphasis bold="true" pageId="7" pageNumber="46">B</emphasis>
Habit, growing in a rocky land slide in gravel (
<emphasis italics="true" pageId="7" pageNumber="46">Knapp et al.</emphasis>
10399)
<emphasis bold="true" pageId="7" pageNumber="46">C</emphasis>
Woody pedicels of the infrutescence, distinct character in herbarium specimens (
<emphasis italics="true" pageId="7" pageNumber="46">Knapp et al.</emphasis>
10401)
<emphasis bold="true" pageId="7" pageNumber="46">D</emphasis>
Ellipsoid fruits with reflexed pedicels, and the characteristic appressed calyx lobes that split into two in fruit (
<emphasis italics="true" pageId="7" pageNumber="46">Knapp et al.</emphasis>
10435). Scale bars = 2 mm. All photos by S. Knapp.
</paragraph>
</caption>
</subSubSection>
<subSubSection lastPageId="8" lastPageNumber="47" pageId="7" pageNumber="46" type="distribution">
<paragraph pageId="7" pageNumber="46">Distribution.</paragraph>
<paragraph lastPageId="8" lastPageNumber="47" pageId="7" pageNumber="46">
Andean Ecuador, Peru, and Bolivia; growing in secondary vegetation, disturbed roadsides, landslides, and gravely slopes in ceja de Selva, montane cloud
<pageBreakToken pageId="8" pageNumber="47" start="start">forest</pageBreakToken>
and
<taxonomicName class="Magnoliopsida" family="Rosaceae" genus="Polylepis" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Polylepis" order="Rosales" pageId="8" pageNumber="47" phylum="Magnoliophyta" rank="genus">
<emphasis italics="true" pageId="8" pageNumber="47">Polylepis</emphasis>
</taxonomicName>
forests, associated with
<taxonomicName class="Magnoliopsida" family="Poaceae" genus="Chusquea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Chusquea" order="Poales" pageId="8" pageNumber="47" phylum="Angiospermae" rank="genus">
<emphasis italics="true" pageId="8" pageNumber="47">Chusquea</emphasis>
</taxonomicName>
(
<taxonomicName class="Magnoliopsida" family="Poaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Poales" pageId="8" pageNumber="47" phylum="Angiospermae" rank="family">Poaceae</taxonomicName>
).
<taxonomicName class="Magnoliopsida" family="Gunneraceae" genus="Gunnera" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Gunnera" order="Gunnerales" pageId="8" pageNumber="47" phylum="Magnoliophyta" rank="genus">
<emphasis italics="true" pageId="8" pageNumber="47">Gunnera</emphasis>
</taxonomicName>
(
<taxonomicName class="Magnoliopsida" family="Gunneraceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Gunnerales" pageId="8" pageNumber="47" phylum="Magnoliophyta" rank="family">Gunneraceae</taxonomicName>
),
<taxonomicName class="Magnoliopsida" family="Urticaceae" genus="Cecropia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cecropia" order="Rosales" pageId="8" pageNumber="47" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="8" pageNumber="47">Cecropia</emphasis>
</taxonomicName>
(
<taxonomicName class="Magnoliopsida" family="Urticaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Rosales" pageId="8" pageNumber="47" phylum="Tracheophyta" rank="family">Urticaceae</taxonomicName>
) and
<taxonomicName class="Magnoliopsida" family="Cunoniaceae" genus="Weinmannia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Weinmannia" order="Oxalidales" pageId="8" pageNumber="47" phylum="Magnoliophyta" rank="genus">
<emphasis italics="true" pageId="8" pageNumber="47">Weinmannia</emphasis>
</taxonomicName>
(
<taxonomicName class="Magnoliopsida" family="Cunoniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Oxalidales" pageId="8" pageNumber="47" phylum="Magnoliophyta" rank="family">Cunoniaceae</taxonomicName>
); 2,000-3,600 (-3,900) m in elevation.
</paragraph>
<caption pageId="8" pageNumber="47" start="Figure 4" startId="F4">
<paragraph pageId="8" pageNumber="47">
<emphasis bold="true" pageId="8" pageNumber="47">Figure 4.</emphasis>
Distribution map of
<taxonomicName authorityName="Saerkinen &amp; S. Knapp" authorityYear="2015" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum antisuyo" order="Solanales" pageId="8" pageNumber="47" phylum="Tracheophyta" rank="species" species="antisuyo">
<emphasis italics="true" pageId="8" pageNumber="47">Solanum antisuyo</emphasis>
</taxonomicName>
, endemic species to Peru.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="8" pageNumber="47" type="ecology">
<paragraph pageId="8" pageNumber="47">Ecology.</paragraph>
<paragraph pageId="8" pageNumber="47">Flowering and fruiting throughout the year, peak in March and June-July.</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="47" type="etymology">
<paragraph pageId="8" pageNumber="47">Etymology.</paragraph>
<paragraph pageId="8" pageNumber="47">
The species name refers to Antisuyo (also Antisuyu), Quechua, for the eastern (
<emphasis italics="true" pageId="8" pageNumber="47">anti</emphasis>
) region (
<emphasis italics="true" pageId="8" pageNumber="47">suyo/suyu</emphasis>
) of the Inca territory, that referred to the lands on the eastern Andean slopes. The species is most diverse along the eastern flanks of the Andes in southern Peru, and the name is chosen to reflect this.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="47" type="conservation status">
<paragraph pageId="8" pageNumber="47">Conservation status.</paragraph>
<paragraph pageId="8" pageNumber="47">
The preliminary conservation status (
<bibRefCitation author="IUCN" journalOrPublisher="Taxon" pageId="18" pageNumber="57" publicationUrl="http://jr.iucnredlist.org/documents/redlist_cats_crit_en.pdf" refId="B15" refString="IUCN, 2010. Guidelines for Using the IUCN Red List Categories and Criteria. Version 8.0. [March 2010]., http://jr.iucnredlist.org/documents/redlist_cats_crit_en.pdf" title="Guidelines for Using the IUCN Red List Categories and Criteria. Version 8.0. [March 2010]" url="http://jr.iucnredlist.org/documents/redlist_cats_crit_en.pdf" year="2010">IUCN 2010</bibRefCitation>
) of
<taxonomicName authorityName="Saerkinen &amp; S. Knapp" authorityYear="2015" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum antisuyo" order="Solanales" pageId="8" pageNumber="47" phylum="Tracheophyta" rank="species" species="antisuyo">
<emphasis italics="true" pageId="8" pageNumber="47">Solanum antisuyo</emphasis>
</taxonomicName>
is here considered of least concern (LC) based on the relatively large EOO (ca. 692,500 km2), although the AOO (136 km2) would merit listing as endangered (EN). The species grows readily in disturbed sites and combined with its wide range, it appears to have relatively low threat status despite the generally increasing human pressure and habitat destruction in the Andes.
</paragraph>
</subSubSection>
<subSubSection lastPageId="9" lastPageNumber="48" pageId="8" pageNumber="47" type="specimens examined">
<paragraph pageId="8" pageNumber="47">Specimens examined.</paragraph>
<paragraph lastPageId="9" lastPageNumber="48" pageId="8" pageNumber="47">
<emphasis bold="true" pageId="8" pageNumber="47">BOLIVIA. Cochabamba:</emphasis>
Chapare: along highway from Villa Tunari to Cochabamba, at Rio San Jacinto bridge, 54.5 km SW of bridge over Rio Espirito Santo,
<geoCoordinate degrees="17" direction="south" minutes="11" orientation="latitude" precision="15" seconds="09" value="-17.185833">17°11'09&quot;S</geoCoordinate>
,
<geoCoordinate degrees="65" direction="west" minutes="44" orientation="longitude" precision="15" seconds="49" value="-65.74694">65°44'49&quot;W</geoCoordinate>
, 1915m, 1 May 2007,
<emphasis italics="true" pageId="9" pageNumber="48">
<pageBreakToken pageId="9" pageNumber="48" start="start">M</pageBreakToken>
. Nee 55288
</emphasis>
(NY); Chapare: New highway to the Chapare, ca. 4 km beyond Laguna Corani, 3140m, 15 Jul 1994,
<emphasis italics="true" pageId="9" pageNumber="48">N. Ritter 1231</emphasis>
(NY); Chapare: El Limbo,
<geoCoordinate degrees="17" direction="south" minutes="09" orientation="latitude" precision="15" seconds="38" value="-17.160555">17°09'38&quot;S</geoCoordinate>
,
<geoCoordinate degrees="65" direction="west" minutes="38" orientation="longitude" precision="15" seconds="22" value="-65.63944">65°38'22&quot;W</geoCoordinate>
, 2900m, 18 Dec 2007,
<emphasis italics="true" pageId="9" pageNumber="48">
J.
<normalizedToken originalValue="Terán">Teran</normalizedToken>
1757
</emphasis>
(NY);
<emphasis bold="true" pageId="9" pageNumber="48">La Paz:</emphasis>
Franz Tamayo: Madidi, Tanhuara Area Natural de Manejo Integrado Apolobamba, bajando del campamento Tanhuara cruzando el
<normalizedToken originalValue="Río">Rio</normalizedToken>
Pelechuco,
<geoCoordinate degrees="14" direction="south" minutes="44" orientation="latitude" precision="15" seconds="47" value="-14.746389">14°44'47&quot;S</geoCoordinate>
,
<geoCoordinate degrees="68" direction="west" minutes="56" orientation="longitude" precision="15" seconds="38" value="-68.94389">68°56'38&quot;W</geoCoordinate>
, 2291m, 11 Jul 2009,
<emphasis italics="true" pageId="9" pageNumber="48">I. Loza 1086</emphasis>
(NY); Inquisivi: Following the slopes E of Communidad Micayani to the
<normalizedToken originalValue="Río">Rio</normalizedToken>
Khokhoni more or less to the junction with a fork flowing down from Communidad Yamora, and following the
<normalizedToken originalValue="Río">Rio</normalizedToken>
Khokhoni upstream 1 km from this point, ca. 4 km SE from Inquisivi, 16°91'67&quot;S,
<geoCoordinate degrees="67" direction="west" minutes="10" orientation="longitude" precision="15" seconds="00" value="-67.166664">67°10'00&quot;W</geoCoordinate>
, 14 Jan 1989,
<emphasis italics="true" pageId="9" pageNumber="48">M. Lewis 35076</emphasis>
(G,MO); Nor Yungas: Unduavi, 3200m, Feb 1914,
<emphasis italics="true" pageId="9" pageNumber="48">O. Buchtien 463</emphasis>
(BM, G, US).
<emphasis bold="true" pageId="9" pageNumber="48">ECUADOR. Loja:</emphasis>
Loja: Parroquia Yangana, rd descending from Cerro Toledo antennas toward Yangana,
<geoCoordinate degrees="4" direction="south" minutes="23" orientation="latitude" precision="15" seconds="02" value="-4.3838887">4°23'02&quot;S</geoCoordinate>
,
<geoCoordinate degrees="79" direction="west" minutes="06" orientation="longitude" precision="15" seconds="58" value="-79.11611">79°06'58&quot;W</geoCoordinate>
, 3050m, 27 Mar 2005,
<emphasis italics="true" pageId="9" pageNumber="48">L. Bohs 3321</emphasis>
(BM, LOJA, NY, QCNE, UT); Loja: Rd from Yangana to boundary of Zamora-Chinchipe, 2560m, 13 Jul 1984,
<emphasis italics="true" pageId="9" pageNumber="48">
W.G.
<normalizedToken originalValue="DArcy">D'Arcy</normalizedToken>
15728
</emphasis>
(NY);
<emphasis bold="true" pageId="9" pageNumber="48">Napo:</emphasis>
Napo-Pastasa, E.N.E of Cayambe Mountain, Oriente trail, 3261m, 6 Dec 1961,
<emphasis italics="true" pageId="9" pageNumber="48">P.C.D. Cazalet 5523</emphasis>
(K); between Tena and Papallacta, 12 Jan 1981,
<emphasis italics="true" pageId="9" pageNumber="48">
W.G.
<normalizedToken originalValue="DArcy">D'Arcy</normalizedToken>
14108
</emphasis>
(NY);
<emphasis bold="true" pageId="9" pageNumber="48">Pichincha:</emphasis>
Quito: Termas de Papallacta Hot Springs, 67 km E of Quito nr the Hwy,
<geoCoordinate degrees="0" direction="south" minutes="20" orientation="latitude" precision="925" value="-0.33333334">0°20'S</geoCoordinate>
,
<geoCoordinate degrees="78" direction="west" minutes="10" orientation="longitude" precision="925" value="-78.166664">78°10'W</geoCoordinate>
, 3300m, 18 Aug 2005,
<emphasis italics="true" pageId="9" pageNumber="48">J.L. Clark 9444</emphasis>
(BM, NY, QCNE, US); Quito: Parroquia, Nono, Reserva Yanacocha, Fundacion de Conservacion Jocotoco,
<geoCoordinate degrees="0" direction="south" minutes="07" orientation="latitude" precision="15" seconds="00" value="-0.11666667">0°07'00&quot;S</geoCoordinate>
,
<geoCoordinate degrees="78" direction="west" minutes="35" orientation="longitude" precision="15" seconds="10" value="-78.58611">78°35'10&quot;W</geoCoordinate>
, 3500m, 13 May 2007,
<emphasis italics="true" pageId="9" pageNumber="48">J.L. Clark 9534</emphasis>
(NY);
<emphasis bold="true" pageId="9" pageNumber="48">Zamora-Chinchipe:</emphasis>
Rd from Yangana to Zumba, 2438m, 13 Jul 1984,
<emphasis italics="true" pageId="9" pageNumber="48">
W.G.
<normalizedToken originalValue="DArcy">D'Arcy</normalizedToken>
15734
</emphasis>
(NY); Rd from Vilcabamba to Valladolid, 27 km from Yangana, 2682m, 12 Jul 1986,
<emphasis italics="true" pageId="9" pageNumber="48">
W.G.
<normalizedToken originalValue="DArcy">D'Arcy</normalizedToken>
16468
</emphasis>
(BM,NY); Parque Nacional Podocarpus, km 26 on rd Yangana-Valladolid,
<geoCoordinate degrees="4" direction="south" minutes="29" orientation="latitude" precision="925" value="-4.483333">4°29'S</geoCoordinate>
,
<geoCoordinate degrees="79" direction="west" minutes="09" orientation="longitude" precision="925" value="-79.15">79°09'W</geoCoordinate>
, 2550m, 2 Dec 1988,
<emphasis italics="true" pageId="9" pageNumber="48">J.E. Madsen 75741</emphasis>
(BM).
<emphasis bold="true" pageId="9" pageNumber="48">PERU. Amazonas:</emphasis>
Prov. Chachapoyas, Dist. Leymebamba, Cordillera Yasgolga, slopes W of summit El Rayo,
<geoCoordinate degrees="6" direction="south" minutes="40" orientation="latitude" precision="15" seconds="01" value="-6.6669445">6°40'01&quot;S</geoCoordinate>
,
<geoCoordinate degrees="77" direction="west" minutes="43" orientation="longitude" precision="15" seconds="05" value="-77.718056">77°43'05&quot;W</geoCoordinate>
, 3287m, 24 Jun 2009,
<emphasis italics="true" pageId="9" pageNumber="48">R.W. Bussmann et al. 15721</emphasis>
(HUT); Prov. Bagua, SE of la Peca, 2362-2461m, 16 Oct 1978,
<emphasis italics="true" pageId="9" pageNumber="48">P. Barbour 4088</emphasis>
(USM);
<emphasis bold="true" pageId="9" pageNumber="48">Cusco:</emphasis>
Prov. La
<normalizedToken originalValue="Convención">Convencion</normalizedToken>
, Dist. Echarate, Quebrada Lorohuachana, sector Laco, Santuario Nacional Megantoni, 3400m, 21 Jun 2008,
<emphasis italics="true" pageId="9" pageNumber="48">L. Hernani A 1037</emphasis>
(HUSA); Dist. Echarate, Quebrada Lorohuachana, sector Laco, Santuario Nacional Megantoni, 3533m, 18 Jun 2008,
<emphasis italics="true" pageId="9" pageNumber="48">L. Hernani A 968</emphasis>
(HUSA); Dist. Echarate, Quebrada Lorohuachana, sector Laco, Santuario Nacional Megantoni, 3400m, 21 Jun 2008,
<emphasis italics="true" pageId="9" pageNumber="48">L. Hernani A 1033</emphasis>
(HUSA); Bosque del Chuyapi,
<geoCoordinate degrees="12" direction="south" minutes="56" orientation="latitude" precision="15" seconds="33" value="-12.9425">12°56'33&quot;S</geoCoordinate>
,
<geoCoordinate degrees="72" direction="west" minutes="47" orientation="longitude" precision="15" seconds="04" value="-72.78445">72°47'04&quot;W</geoCoordinate>
, 2100m, 19 Jul 2006,
<emphasis italics="true" pageId="9" pageNumber="48">L. Valenzuela et al. 7282</emphasis>
(USM); Bosque del Chuyapi,
<geoCoordinate degrees="12" direction="south" minutes="56" orientation="latitude" precision="15" seconds="33" value="-12.9425">12°56'33&quot;S</geoCoordinate>
,
<geoCoordinate degrees="72" direction="west" minutes="47" orientation="longitude" precision="15" seconds="04" value="-72.78445">72°47'04&quot;W</geoCoordinate>
, 2100m, 19 Jul 2006,
<emphasis italics="true" pageId="9" pageNumber="48">L. Valenzuela et al. 7284</emphasis>
(USM); Ca. 51 km from Ollantaytambo on rd over Abra Malaga to Quillabamba and Quelluno, ca. 10 km below Abra Malaga, Amazon slope,
<geoCoordinate degrees="13" direction="south" minutes="07" orientation="latitude" precision="15" seconds="15" value="-13.120833">13°07'15&quot;S</geoCoordinate>
,
<geoCoordinate degrees="72" direction="west" minutes="19" orientation="longitude" precision="15" seconds="34" value="-72.32611">72°19'34&quot;W</geoCoordinate>
, 3877m, 13 Mar 2012,
<emphasis italics="true" pageId="9" pageNumber="48">S. Knapp et al. 10399</emphasis>
(BM, USM); Ca. 57 km from Ollantaytambo on rd over Abra Malaga to Quillabamba and Quelluno, ca. 16 km below Abra Malaga, Amazon slope,
<geoCoordinate degrees="13" direction="south" minutes="06" orientation="latitude" precision="15" seconds="30" value="-13.108334">13°06'30&quot;S</geoCoordinate>
,
<geoCoordinate degrees="72" direction="west" minutes="21" orientation="longitude" precision="15" seconds="04" value="-72.35111">72°21'04&quot;W</geoCoordinate>
, 3579m, 13 Mar 2012,
<emphasis italics="true" pageId="9" pageNumber="48">S. Knapp et al. 10401</emphasis>
(BM, USM); Ca. 61 km from Ollantaytambo on rd over Abra Malaga to Quillabamba and Quelluno, ca. 21 km below Abra Malaga, Amazon slope,
<geoCoordinate degrees="13" direction="south" minutes="06" orientation="latitude" precision="15" seconds="15" value="-13.104167">13°06'15&quot;S</geoCoordinate>
,
<geoCoordinate degrees="72" direction="west" minutes="22" orientation="longitude" precision="15" seconds="03" value="-72.3675">72°22'03&quot;W</geoCoordinate>
, 3448m, 13 Mar 2012,
<emphasis italics="true" pageId="9" pageNumber="48">S. Knapp et al. 10404</emphasis>
(BM, USM); Ca. 61 km from Ollantaytambo on rd over Abra Malaga to Quillabamba and Quelluno, ca. 21 km below Abra Malaga, Amazon slope,
<geoCoordinate degrees="13" direction="south" minutes="06" orientation="latitude" precision="15" seconds="15" value="-13.104167">13°06'15&quot;S</geoCoordinate>
,
<geoCoordinate degrees="72" direction="west" minutes="22" orientation="longitude" precision="15" seconds="03" value="-72.3675">72°22'03&quot;W</geoCoordinate>
, 3448m, 13 Mar 2012,
<emphasis italics="true" pageId="9" pageNumber="48">S. Knapp et al. 10406</emphasis>
(BM, USM); Prov. Paucartambo, 3500-3600m, 14 Jul 1990,
<emphasis italics="true" pageId="9" pageNumber="48">A. Cano 3636</emphasis>
(USM); Parque Nacional de Manu, 3500-3600m, 11 Jul 1990,
<emphasis italics="true" pageId="9" pageNumber="48">A. Cano 4327</emphasis>
(USM); Parque Nacional de Manu, 3600-3700m, 6 Mar 1991,
<emphasis italics="true" pageId="9" pageNumber="48">A. Cano 4593</emphasis>
(USM); Prov. Urubamba, Aguas Calientes, Quebrada Alccamayo,
<geoCoordinate degrees="13" direction="south" minutes="09" orientation="latitude" precision="15" seconds="01" value="-13.150277">13°09'01&quot;S</geoCoordinate>
,
<geoCoordinate degrees="72" direction="west" minutes="30" orientation="longitude" precision="15" seconds="17" value="-72.50472">72°30'17&quot;W</geoCoordinate>
, 2050-2200m, 29 Aug 2002,
<emphasis italics="true" pageId="9" pageNumber="48">I. Huamantupa &amp; G. Calatayud 2258</emphasis>
(MO, USM);
<emphasis bold="true" pageId="9" pageNumber="48">
<normalizedToken originalValue="Huánuco">Huanuco</normalizedToken>
:
</emphasis>
Prov.
<normalizedToken originalValue="Huánuco">Huanuco</normalizedToken>
, along rd between
<normalizedToken originalValue="Huánuco">Huanuco</normalizedToken>
and Tingo
<normalizedToken originalValue="María">Maria</normalizedToken>
, 1.1 km N of Carpish Tunnel, ca. km 455,
<geoCoordinate degrees="9" direction="south" minutes="40" orientation="latitude" precision="925" value="-9.666667">9°40'S</geoCoordinate>
,
<geoCoordinate degrees="76" direction="west" minutes="04" orientation="longitude" precision="925" value="-76.066666">76°04'W</geoCoordinate>
, 2680m, 1 Jun 1998,
<emphasis italics="true" pageId="9" pageNumber="48">T.B.Croat &amp; M. Sizemore 81568</emphasis>
(BM, MO, USM); Carpish, above Acomayo, lower ceja, 2800m, 17 Jul 1964,
<emphasis italics="true" pageId="9" pageNumber="48">P.C. Hutchison et al. 5931</emphasis>
(MO); Carpish, carretera Huanuco-Tingo Maria, 2700-2900m, 3 Oct 1950,
<emphasis italics="true" pageId="9" pageNumber="48">R. Ferreyra 8164</emphasis>
(MOL, USM); North of Carpish Pass, rd from
<normalizedToken originalValue="Huánuco">Huanuco</normalizedToken>
to Tingo Maria, 50-52 km NE of
<normalizedToken originalValue="Huánuco">Huanuco</normalizedToken>
, 2350-2430m, 6 Dec 1981,
<emphasis italics="true" pageId="9" pageNumber="48">T. Plowman &amp; P.M. Rury 11141</emphasis>
(HB,USM); Prov. Pachitea, 2450m, 24 Dec 1979,
<emphasis italics="true" pageId="9" pageNumber="48">HHV 001519</emphasis>
(USM); Prov. Tingo Maria, 2900m, 21 Jun 1980,
<emphasis italics="true" pageId="9" pageNumber="48">HHV 3050</emphasis>
(USM);
<emphasis bold="true" pageId="9" pageNumber="48">
<normalizedToken originalValue="Junín">Junin</normalizedToken>
:
</emphasis>
Prov. Huancayo, 3200m, 17 Jun 1972,
<emphasis italics="true" pageId="9" pageNumber="48">D. Tovar T. 5</emphasis>
(USM);
<emphasis bold="true" pageId="9" pageNumber="48">Pasco:</emphasis>
Prov. Oxapampa, Dist. Oxapampa, Parque Nacional
<normalizedToken originalValue="Yanachaga-Chemillén">Yanachaga-Chemillen</normalizedToken>
, sector San Alberto, alrededores del Refugio el Cedro,
<geoCoordinate degrees="10" direction="south" minutes="32" orientation="latitude" precision="15" seconds="26" value="-10.540556">10°32'26&quot;S</geoCoordinate>
,
<geoCoordinate degrees="75" direction="west" minutes="21" orientation="longitude" precision="15" seconds="18" value="-75.354996">75°21'18&quot;W</geoCoordinate>
, 2483m, 26 Apr 2009,
<emphasis italics="true" pageId="9" pageNumber="48">M. Cueva &amp; R. Rivera 625</emphasis>
(HUSA, USM); Dist. Huancabamba, Parque Nacional
<normalizedToken originalValue="Yanachaga-Chemillén">Yanachaga-Chemillen</normalizedToken>
, sector San Daniel,
<geoCoordinate degrees="10" direction="south" minutes="26" orientation="latitude" precision="15" seconds="09" value="-10.435833">10°26'09&quot;S</geoCoordinate>
,
<geoCoordinate degrees="75" direction="west" minutes="27" orientation="longitude" precision="15" seconds="07" value="-75.45194">75°27'07&quot;W</geoCoordinate>
, 2192m, 29 Feb 2008,
<emphasis italics="true" pageId="9" pageNumber="48">
R.
<normalizedToken originalValue="Vásquez">Vasquez</normalizedToken>
et al. 33798
</emphasis>
(USM); Parque Nacional
<normalizedToken originalValue="Yanachaga-Chemillén">Yanachaga-Chemillen</normalizedToken>
, quebrada diablo fuerte, trocha hacia la parcela Oso-Playa,
<geoCoordinate degrees="10" direction="south" minutes="18" orientation="latitude" precision="15" seconds="14" value="-10.303889">10°18'14&quot;S</geoCoordinate>
,
<geoCoordinate degrees="75" direction="west" minutes="36" orientation="longitude" precision="15" seconds="14" value="-75.60389">75°36'14&quot;W</geoCoordinate>
, 2398-2500m, 23 Jun 2008,
<emphasis italics="true" pageId="9" pageNumber="48">A. Monteagudo et al. 16477</emphasis>
(USM); Dist. Huancabamba, Zona de amortiquamiento del P.N.
<normalizedToken originalValue="Yanachaga-Chemillén">Yanachaga-Chemillen</normalizedToken>
, sector Milpo,
<geoCoordinate degrees="10" direction="south" minutes="22" orientation="latitude" precision="15" seconds="20" value="-10.372222">10°22'20&quot;S</geoCoordinate>
,
<geoCoordinate degrees="75" direction="west" minutes="36" orientation="longitude" precision="15" seconds="29" value="-75.608055">75°36'29&quot;W</geoCoordinate>
, 2800m, 23 Sep 2004,
<emphasis italics="true" pageId="9" pageNumber="48">A.Monteagudo et al. 7326</emphasis>
(K); Dist. Huancabamba, Parque Nacional
<normalizedToken originalValue="Yanachaga-Chemillén">Yanachaga-Chemillen</normalizedToken>
, sector San Daniel,
<geoCoordinate degrees="10" direction="south" minutes="25" orientation="latitude" precision="15" seconds="48" value="-10.43">10°25'48&quot;S</geoCoordinate>
,
<geoCoordinate degrees="75" direction="west" minutes="27" orientation="longitude" precision="15" seconds="00" value="-75.45">75°27'00&quot;W</geoCoordinate>
, 2500-3500m, 1 Mar 2009,
<emphasis italics="true" pageId="9" pageNumber="48">M. Cueva et al. 492</emphasis>
(USM);
<emphasis bold="true" pageId="9" pageNumber="48">Puno:</emphasis>
Prov. Carabaya, margen derecha del rio que pasa frente al campamento Chacayage,
<geoCoordinate degrees="13" direction="south" minutes="45" orientation="latitude" precision="15" seconds="48" value="-13.763333">13°45'48&quot;S</geoCoordinate>
,
<geoCoordinate degrees="70" direction="west" minutes="13" orientation="longitude" precision="15" seconds="07" value="-70.21861">70°13'07&quot;W</geoCoordinate>
, 2600m, 10 Mar 2004,
<emphasis italics="true" pageId="9" pageNumber="48">S. Vilca C. et al. 71</emphasis>
(HUSA); km 238-239 on rd to Juliaca from San Gaban, ca. 8-9 km before Ollachea and Puente Chillichaca,
<geoCoordinate degrees="13" direction="south" minutes="45" orientation="latitude" precision="15" seconds="30" value="-13.758333">13°45'30&quot;S</geoCoordinate>
,
<geoCoordinate degrees="70" direction="west" minutes="28" orientation="longitude" precision="15" seconds="09" value="-70.46917">70°28'09&quot;W</geoCoordinate>
, 2451m, 19 Mar 2012,
<emphasis italics="true" pageId="9" pageNumber="48">
T.
<normalizedToken originalValue="Särkinen">Saerkinen</normalizedToken>
et al. 4035
</emphasis>
(USM); Prov. Sandia, 8 km from Sandia on rd to Cuyocuyo,
<geoCoordinate degrees="14" direction="south" minutes="22" orientation="latitude" precision="15" seconds="08" value="-14.368889">14°22'08&quot;S</geoCoordinate>
,
<geoCoordinate degrees="69" direction="west" minutes="28" orientation="longitude" precision="15" seconds="23" value="-69.47305">69°28'23&quot;W</geoCoordinate>
, 2709m, 21 Mar 2012,
<emphasis italics="true" pageId="9" pageNumber="48">
T.
<normalizedToken originalValue="Särkinen">Saerkinen</normalizedToken>
et al. 4048
</emphasis>
(BM, USM); 16 km from Sandia on rd to Cuyocuyo,
<geoCoordinate degrees="14" direction="south" minutes="24" orientation="latitude" precision="15" seconds="06" value="-14.401667">14°24'06&quot;S</geoCoordinate>
,
<geoCoordinate degrees="69" direction="west" minutes="28" orientation="longitude" precision="15" seconds="25" value="-69.47361">69°28'25&quot;W</geoCoordinate>
, 2990m, 21 Mar 2012,
<emphasis italics="true" pageId="9" pageNumber="48">
T.
<normalizedToken originalValue="Särkinen">Saerkinen</normalizedToken>
et al. 4049
</emphasis>
(BM, USM); in Cuyocuyo outside of a house on main rd to Sandia,
<geoCoordinate degrees="14" direction="south" minutes="27" orientation="latitude" precision="15" seconds="09" value="-14.452499">14°27'09&quot;S</geoCoordinate>
,
<geoCoordinate degrees="69" direction="west" minutes="32" orientation="longitude" precision="15" seconds="03" value="-69.534164">69°32'03&quot;W</geoCoordinate>
, 3364m, 21 Mar 2012,
<emphasis italics="true" pageId="9" pageNumber="48">
T.
<normalizedToken originalValue="Särkinen">Saerkinen</normalizedToken>
et al. 4053
</emphasis>
(BM, USM).
</paragraph>
</subSubSection>
<subSubSection lastPageId="10" lastPageNumber="49" pageId="9" pageNumber="48" type="discussion">
<paragraph pageId="9" pageNumber="48">Discussion.</paragraph>
<paragraph pageId="10" pageNumber="49">
<taxonomicName authorityName="Saerkinen &amp; S. Knapp" authorityYear="2015" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum antisuyo" order="Solanales" pageId="10" pageNumber="49" phylum="Tracheophyta" rank="species" species="antisuyo">
<emphasis italics="true" pageId="10" pageNumber="49">
<pageBreakToken pageId="10" pageNumber="49" start="start">Solanum</pageBreakToken>
antisuyo
</emphasis>
</taxonomicName>
is morphologically most similar to
<taxonomicName authorityName="Bitter" authorityYear="1912" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum polytrichostylum" order="Solanales" pageId="10" pageNumber="49" phylum="Tracheophyta" rank="species" species="polytrichostylum">
<emphasis italics="true" pageId="10" pageNumber="49">Solanum polytrichostylum</emphasis>
</taxonomicName>
Bitter with which it has been conflated in the past. It can be distinguished by simple or once-branched inflorescences where pedicels are spaced ca. 1-3 mm apart along the short rachis compared to consistently branched inflorescences with the flowers congested at the branch tips in
<taxonomicName authorityName="Bitter" authorityYear="1912" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum polytrichostylum" order="Solanales" pageId="10" pageNumber="49" phylum="Tracheophyta" rank="species" species="polytrichostylum">
<emphasis italics="true" pageId="10" pageNumber="49">Solanum polytrichostylum</emphasis>
</taxonomicName>
, and ellipsoid fruits as compared to the spherical fruits of
<taxonomicName authorityName="Bitter" authorityYear="1912" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum polytrichostylum" order="Solanales" pageId="10" pageNumber="49" phylum="Tracheophyta" rank="species" species="polytrichostylum">
<emphasis italics="true" pageId="10" pageNumber="49">Solanum polytrichostylum</emphasis>
</taxonomicName>
.
<taxonomicName authorityName="Saerkinen &amp; S. Knapp" authorityYear="2015" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum antisuyo" order="Solanales" pageId="10" pageNumber="49" phylum="Tracheophyta" rank="species" species="antisuyo">
<emphasis italics="true" pageId="10" pageNumber="49">Solanum antisuyo</emphasis>
</taxonomicName>
has a longer calyx tube with more reduced, poorly developed purple-tinged calyx lobes compared to the shorter calyx tubes with slightly larger, triangular calyx lobes in
<taxonomicName authorityName="Bitter" authorityYear="1912" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum polytrichostylum" order="Solanales" pageId="10" pageNumber="49" phylum="Tracheophyta" rank="species" species="polytrichostylum">
<emphasis italics="true" pageId="10" pageNumber="49">Solanum polytrichostylum</emphasis>
</taxonomicName>
, and larger brown coloured seeds compared to smaller yellow seeds in
<taxonomicName authorityName="Bitter" authorityYear="1912" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum polytrichostylum" order="Solanales" pageId="10" pageNumber="49" phylum="Tracheophyta" rank="species" species="polytrichostylum">
<emphasis italics="true" pageId="10" pageNumber="49">Solanum polytrichostylum</emphasis>
</taxonomicName>
. Furthermore, styles are always more exerted (2-4 mm versus 1-2 mm beyond the anther cone) in
<taxonomicName authorityName="Bitter" authorityYear="1912" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum polytrichostylum" order="Solanales" pageId="10" pageNumber="49" phylum="Tracheophyta" rank="species" species="polytrichostylum">
<emphasis italics="true" pageId="10" pageNumber="49">Solanum polytrichostylum</emphasis>
</taxonomicName>
. The two species are also ecologically somewhat distinct, with
<taxonomicName authorityName="Bitter" authorityYear="1912" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum polytrichostylum" order="Solanales" pageId="10" pageNumber="49" phylum="Tracheophyta" rank="species" species="polytrichostylum">
<emphasis italics="true" pageId="10" pageNumber="49">Solanum polytrichostylum</emphasis>
</taxonomicName>
restricted to streams and moist road sides, and
<taxonomicName authorityName="Saerkinen &amp; S. Knapp" authorityYear="2015" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum antisuyo" order="Solanales" pageId="10" pageNumber="49" phylum="Tracheophyta" rank="species" species="antisuyo">
<emphasis italics="true" pageId="10" pageNumber="49">Solanum antisuyo</emphasis>
</taxonomicName>
is found in drier areas in gravel, disturbed areas, and landslides. Other members of the Morelloid clade in Peru without glandular trichomes with which
<taxonomicName authorityName="Saerkinen &amp; S. Knapp" authorityYear="2015" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum antisuyo" order="Solanales" pageId="10" pageNumber="49" phylum="Tracheophyta" rank="species" species="antisuyo">
<emphasis italics="true" pageId="10" pageNumber="49">Solanum antisuyo</emphasis>
</taxonomicName>
could be confused include
<taxonomicName authorityName="Bitter" authorityYear="1916" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum probolospermum" order="Solanales" pageId="10" pageNumber="49" phylum="Tracheophyta" rank="species" species="probolospermum">
<emphasis italics="true" pageId="10" pageNumber="49">Solanum probolospermum</emphasis>
</taxonomicName>
Bitter that has smaller, spherical fruits, larger violet corollas that are more rotate in outline, and denser indumentum with longer 3-7-celled simple hairs, and
<taxonomicName authorityName="Rusby" authorityYear="1895" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum pallidum" order="Solanales" pageId="10" pageNumber="49" phylum="Tracheophyta" rank="species" species="pallidum">
<emphasis italics="true" pageId="10" pageNumber="49">Solanum pallidum</emphasis>
</taxonomicName>
Rusby (incl.
<taxonomicName authorityName="Bitter" authorityYear="1912" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum planifurcum" order="Solanales" pageId="10" pageNumber="49" phylum="Tracheophyta" rank="species" species="planifurcum">
<emphasis italics="true" pageId="10" pageNumber="49">Solanum planifurcum</emphasis>
</taxonomicName>
Bitter) that is similar to
<taxonomicName authorityName="Bitter" authorityYear="1916" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum probolospermum" order="Solanales" pageId="10" pageNumber="49" phylum="Tracheophyta" rank="species" species="probolospermum">
<emphasis italics="true" pageId="10" pageNumber="49">Solanum probolospermum</emphasis>
</taxonomicName>
but has branched rather than simple hairs.
</paragraph>
<paragraph pageId="10" pageNumber="49">
Variation in growth form and flower colour can be observed in the field, where individuals growing in more humid conditions grow into stout herbs to ca. 1.5 m tall, whilst individuals in drier, higher elevation habitats in rocky landslides are stunted herbs reaching only ca. 40 cm in height. Colour variation in corolla is common within Morelloids and
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum" order="Solanales" pageId="10" pageNumber="49" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="10" pageNumber="49">Solanum</emphasis>
</taxonomicName>
species in general: most specimens of
<taxonomicName authorityName="Saerkinen &amp; S. Knapp" authorityYear="2015" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum antisuyo" order="Solanales" pageId="10" pageNumber="49" phylum="Tracheophyta" rank="species" species="antisuyo">
<emphasis italics="true" pageId="10" pageNumber="49">Solanum antisuyo</emphasis>
</taxonomicName>
have creamy white petals, but occasional specimens with lilac corollas are known (e.g.,
<emphasis italics="true" pageId="10" pageNumber="49">
<normalizedToken originalValue="Särkinen">Saerkinen</normalizedToken>
et al. 4048, 4049
</emphasis>
, and
<emphasis italics="true" pageId="10" pageNumber="49">4053</emphasis>
).
</paragraph>
</subSubSection>
</treatment>
</document>