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<mods:title id="65588244B138FB9E63EBE8D66DDA4928">Redescription of Palaemon antennarius H. Milne Edwards, 1837 and Palaemon migratorius (Heller, 1862) (Crustacea, Decapoda, Palaemonidae) and description of two new species of the genus from the circum-Mediterranean area</mods:title>
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Key to identification of the inland
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species from the circum-Mediterranean area
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1. Fifth pleuron distal end not rounded (
<figureCitation id="2FF62A5FFFACFFFB1613FF6A95ABEBBA" box="[583,674,222,243]" captionStart="FIGURE 2" captionStartId="5.[151,250,1915,1937]" captionTargetBox="[156,1426,206,1914]" captionTargetId="figure@5.[156,1436,177,1916]" captionTargetPageId="5" captionText="FIGURE 2. Palaemon antennarius H. Milne Edwards, 1837. A. Whole animal, lateral view. B. Eye, dorso-mesial view. C. Frontal region. D. Antennula. E. Fifth pleuron. F. Uropod. G. Scaphocerite. Lectotype ♀ (pocl 7.7 mm), MNHN IU 2013 9296 (= MNHN Na 1500), Trasimeno Lake, Italy. Scale bar indicates 5 mm (A), 1 mm (B), 3 mm (C, D, F, G), 2 mm (E)." httpUri="https://zenodo.org/record/287659/files/figure.png" pageId="22" pageNumber="49">Figures 2</figureCitation>
E, 7B, 10A)..................................................... 2
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- Fifth pleuron distal end rounded (
<figureCitation id="2FF62A5FFFACFFFB1676FF4F957CEA59" box="[546,629,251,272]" captionStart="FIGURE 6" captionStartId="12.[151,250,1921,1943]" captionTargetBox="[182,1402,201,1889]" captionTargetId="figure@12.[178,1402,193,1898]" captionTargetPageId="12" captionText="FIGURE 6. Palaemon migratorius (Heller, 1862). A. Whole animal, lateral view. B. Fifth pleuron. C. First maxilliped. D. Thoracic sternal armature. E. Abdominal sternal armature. F. Telson distal part. A B, D F lectotype ♀ (pocl 8.9 mm), NHMW 25559, Egypt; C ♀ paralectotype (pocl 6.9 mm), NHMW 7733, Egypt. Scale bar indicates 2 mm (A), 1 mm (B, C), 1.5 mm (D, E), 0.5 mm (F)." httpUri="https://zenodo.org/record/287663/files/figure.png" pageId="22" pageNumber="49">Figure 6</figureCitation>
B, 9B)............................................................. 7
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2. Fifth pleuron distal end strongly pointed (
<figureCitation id="2FF62A5FFFACFFFB1624FEAD95CBEA67" box="[624,706,281,302]" captionStart="FIGURE 2" captionStartId="5.[151,250,1915,1937]" captionTargetBox="[156,1426,206,1914]" captionTargetId="figure@5.[156,1436,177,1916]" captionTargetPageId="5" captionText="FIGURE 2. Palaemon antennarius H. Milne Edwards, 1837. A. Whole animal, lateral view. B. Eye, dorso-mesial view. C. Frontal region. D. Antennula. E. Fifth pleuron. F. Uropod. G. Scaphocerite. Lectotype ♀ (pocl 7.7 mm), MNHN IU 2013 9296 (= MNHN Na 1500), Trasimeno Lake, Italy. Scale bar indicates 5 mm (A), 1 mm (B), 3 mm (C, D, F, G), 2 mm (E)." httpUri="https://zenodo.org/record/287659/files/figure.png" pageId="22" pageNumber="49">Figure 2</figureCitation>
E, 7B)....................................................... 3
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- Fifth pleuron distal end subquadrate and not pointed (
<figureCitation id="2FF62A5FFFACFFFB16B6FE82945AEA02" box="[738,851,310,331]" captionStart="FIGURE 10. A" captionStartId="20.[151,250,1556,1578]" captionTargetBox="[201,1401,224,1513]" captionTargetId="figure@20.[180,1412,209,1518]" captionTargetPageId="20" captionText="FIGURE 10. A. Fifth pleuron of P. zariquieyi. B. Maxillula of P. varians. C. Telson distal part of P. v a r i an s. D. Telson distal part of P. mesopotamicus. E. Telson distal part of P. zariquieyi. F. First maxilliped of P. varians. G. Third maxilliped of P. mesopotamicus. H. Scaphocerite of P. mesopotamicus. A, E, ♀ (pocl 7.0 mm), OUMNH. ZC. 2004 11 001, Las Turberas, Prat de Cabamas, Torreblanca, Spain; B C, F, ♀ (pocl 7.2 mm), NHM 1959.10.17.72 82, Roose Cote, Power Station, Morecambe Bay, United Kingdom; D, G H, ♀ (pocl 6.9 mm), SMF TSH 30 / 88, Khabur river, Syria. Scale bar indicates 3 mm (A), 1 mm (B E), 1.5 mm (F), 2 mm (G), 5 mm (H)." httpUri="https://zenodo.org/record/287667/files/figure.png" pageId="22" pageNumber="49">Figure 10A</figureCitation>
)............................................... 6
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3. Telson with two plumose setae on posterior margin (
<figureCitation id="2FF62A5FFFACFFFB168BFEE79432EA21" box="[735,827,339,360]" captionStart="FIGURE 5" captionStartId="8.[151,250,1080,1102]" captionTargetBox="[242,1344,193,1051]" captionTargetId="figure@8.[242,1344,193,1059]" captionTargetPageId="8" captionText="FIGURE 5. Palaemon antennarius H. Milne Edwards, 1837. A. Telson. B. Telson distal part. Lectotype ♀ (pocl 7.7 mm), MNHN IU 2013 9296 (= MNHN Na 1500), Trasimeno Lake, Italy. Scale bar indicates 3 mm (A), 1 mm (B)." httpUri="https://zenodo.org/record/287662/files/figure.png" pageId="22" pageNumber="49">Figures 5</figureCitation>
A, 5B, 8C, 10C); third maxilliped exopod equal or longer than isciomerus (
<figureCitation id="2FF62A5FFFACFFFB1502FEC596A3EACF" box="[342,426,369,390]" captionStart="FIGURE 3" captionStartId="6.[151,250,1850,1872]" captionTargetBox="[165,1424,204,1818]" captionTargetId="figure@6.[152,1435,193,1828]" captionTargetPageId="6" captionText="FIGURE 3. Palaemon antennarius H. Milne Edwards, 1837. A. Thoracic sternal armature. B. Abdominal sternal armature. C. Mandible. D. Maxillula. E. Maxilla. F. First maxilliped. G. Second maxilliped. H. Third maxilliped. A H ♀ paralectotype (pocl 8.0 mm), MNHN IU 2013 4023 (= MNHN Na 1500), Trasimeno Lake, Italy; A B ♂ (pocl 6.6 mm), Trasimeno Lake, 6.9. 1974, coll. Froglia C. &amp; E. Peloni. Scale bar indicates 1.5 mm (A, B, E H), 1 mm (C, D)." httpUri="https://zenodo.org/record/287660/files/figure.png" pageId="22" pageNumber="49">Figure 3</figureCitation>
H); endopod of first pleopod of male clearly shorter than exopod (
<figureCitation id="2FF62A5FFFACFFFB1066FEC59386EACF" box="[1074,1167,369,390]" captionStart="FIGURE 4" captionStartId="7.[151,250,1802,1824]" captionTargetBox="[177,1406,203,1736]" captionTargetId="figure@7.[164,1423,193,1749]" captionTargetPageId="7" captionText="FIGURE 4. Palaemon antennarius H. Milne Edwards, 1837. A. First pereiopod. B. Second pereiopod. C. Third pereiopod. D. Fourth pereiopod. E. Fifth pereiopod. F. First pleopod male. G. Second pleopod male. H. First pleopod female. I. Second pleopod female. J. Third pleopod female. K. Fifth pleopod female. A E, H K ♀ paralectotype (pocl 8.0 mm), MNHN IU 2013 4023 (= MNHN Na 1500), Trasimeno Lake, Italy; F G ♂ (pocl 6.6 mm), Trasimeno Lake, 6.9. 1974, coll. Froglia C. &amp; E. Peloni. Scale bar indicates 2 mm." httpUri="https://zenodo.org/record/287661/files/figure.png" pageId="22" pageNumber="49">Figures 4</figureCitation>
F, 8A); appendix masculina shorter than the endopod of the second pleopod (
<figureCitation id="2FF62A5FFFACFFFB16E0FE3A940FEAEA" box="[692,774,398,419]" captionStart="FIGURE 4" captionStartId="7.[151,250,1802,1824]" captionTargetBox="[177,1406,203,1736]" captionTargetId="figure@7.[164,1423,193,1749]" captionTargetPageId="7" captionText="FIGURE 4. Palaemon antennarius H. Milne Edwards, 1837. A. First pereiopod. B. Second pereiopod. C. Third pereiopod. D. Fourth pereiopod. E. Fifth pereiopod. F. First pleopod male. G. Second pleopod male. H. First pleopod female. I. Second pleopod female. J. Third pleopod female. K. Fifth pleopod female. A E, H K ♀ paralectotype (pocl 8.0 mm), MNHN IU 2013 4023 (= MNHN Na 1500), Trasimeno Lake, Italy; F G ♂ (pocl 6.6 mm), Trasimeno Lake, 6.9. 1974, coll. Froglia C. &amp; E. Peloni. Scale bar indicates 2 mm." httpUri="https://zenodo.org/record/287661/files/figure.png" pageId="22" pageNumber="49">Figure 4</figureCitation>
G).................................................... 4
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- Telson with 26 plumose setae on posterior margin (Figure
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); third maxilliped exopod shorter than isciomerus (
<figureCitation id="2FF62A5FFFACFFFB1561FE7D9680EA97" box="[309,393,457,478]" captionStart="FIGURE 7" captionStartId="14.[151,250,1865,1887]" captionTargetBox="[181,1418,204,1829]" captionTargetId="figure@14.[168,1418,193,1829]" captionTargetPageId="14" captionText="FIGURE 7. Palaemon colossus sp. nov. A. Whole animal, lateral view. B. Fifth pleuron. C. Maxillula. D. Thoracic sternal armature. E. Abdominal sternal armature. A B, D E holotype ♀ (pocl 8.5 mm), OUMNH. ZC. 2014 13 001, Epta Piges, Rhodes Island, Greece; C, D E, ♂ (pocl 6.0 mm), ZMAUTH G 3 30, Epta Piges, Rhodes Island, Greece, 2002, coll. Anastasiadou Ch. Scale bar indicates 4 mm (A), 2 mm (B), 0.5 mm (C), 1.5 mm (D, E)." httpUri="https://zenodo.org/record/287664/files/figure.png" pageId="22" pageNumber="49">
Figure
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in
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); endopod of first pleopod of male slightly shorter than exopod (Figure
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); appendix masculina longer than the endopod of the second pleopod (Figure
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).....
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<emphasis id="85B9EAC8FFACFFFB1159FE529292EAB2" box="[1293,1435,486,507]" italics="true" pageId="22" pageNumber="49">P. mesogenitor</emphasis>
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4. Telson plumose setae fail to overpass the inner spines (
<figureCitation id="2FF62A5FFFACFFFB1754FDB7945CE951" box="[768,853,515,536]" captionStart="FIGURE 5" captionStartId="8.[151,250,1080,1102]" captionTargetBox="[242,1344,193,1051]" captionTargetId="figure@8.[242,1344,193,1059]" captionTargetPageId="8" captionText="FIGURE 5. Palaemon antennarius H. Milne Edwards, 1837. A. Telson. B. Telson distal part. Lectotype ♀ (pocl 7.7 mm), MNHN IU 2013 9296 (= MNHN Na 1500), Trasimeno Lake, Italy. Scale bar indicates 3 mm (A), 1 mm (B)." httpUri="https://zenodo.org/record/287662/files/figure.png" pageId="22" pageNumber="49">Figure 5</figureCitation>
B); first maxilliped endites separated with their inner edges forming an obtuse angle (
<figureCitation id="2FF62A5FFFACFFFB1581FD95952EE97F" box="[469,551,545,566]" captionStart="FIGURE 3" captionStartId="6.[151,250,1850,1872]" captionTargetBox="[165,1424,204,1818]" captionTargetId="figure@6.[152,1435,193,1828]" captionTargetPageId="6" captionText="FIGURE 3. Palaemon antennarius H. Milne Edwards, 1837. A. Thoracic sternal armature. B. Abdominal sternal armature. C. Mandible. D. Maxillula. E. Maxilla. F. First maxilliped. G. Second maxilliped. H. Third maxilliped. A H ♀ paralectotype (pocl 8.0 mm), MNHN IU 2013 4023 (= MNHN Na 1500), Trasimeno Lake, Italy; A B ♂ (pocl 6.6 mm), Trasimeno Lake, 6.9. 1974, coll. Froglia C. &amp; E. Peloni. Scale bar indicates 1.5 mm (A, B, E H), 1 mm (C, D)." httpUri="https://zenodo.org/record/287660/files/figure.png" pageId="22" pageNumber="49">Figure 3</figureCitation>
F); branchiostegal tooth originating just before anterior margin of carapace (
<figureCitation id="2FF62A5FFFACFFFB1156FD95925DE97F" box="[1282,1364,545,566]" captionStart="FIGURE 2" captionStartId="5.[151,250,1915,1937]" captionTargetBox="[156,1426,206,1914]" captionTargetId="figure@5.[156,1436,177,1916]" captionTargetPageId="5" captionText="FIGURE 2. Palaemon antennarius H. Milne Edwards, 1837. A. Whole animal, lateral view. B. Eye, dorso-mesial view. C. Frontal region. D. Antennula. E. Fifth pleuron. F. Uropod. G. Scaphocerite. Lectotype ♀ (pocl 7.7 mm), MNHN IU 2013 9296 (= MNHN Na 1500), Trasimeno Lake, Italy. Scale bar indicates 5 mm (A), 1 mm (B), 3 mm (C, D, F, G), 2 mm (E)." httpUri="https://zenodo.org/record/287659/files/figure.png" pageId="22" pageNumber="49">Figure 2</figureCitation>
C)... 5
</keyLead>
</paragraph>
<paragraph id="B77236DAFFACFFFB14C3FD8A9295E9C7" blockId="22.[151,1436,222,1154]" pageId="22" pageNumber="49">
<keyLead id="0C3C95EFFFACFFFB14C3FD8A9295E9C7" pageId="22" pageNumber="49">
- Telson plumose setae overpass the inner spines (
<figureCitation id="2FF62A5FFFACFFFB1694FD8A9428E91A" box="[704,801,574,595]" captionStart="FIGURE 10. A" captionStartId="20.[151,250,1556,1578]" captionTargetBox="[201,1401,224,1513]" captionTargetId="figure@20.[180,1412,209,1518]" captionTargetPageId="20" captionText="FIGURE 10. A. Fifth pleuron of P. zariquieyi. B. Maxillula of P. varians. C. Telson distal part of P. v a r i an s. D. Telson distal part of P. mesopotamicus. E. Telson distal part of P. zariquieyi. F. First maxilliped of P. varians. G. Third maxilliped of P. mesopotamicus. H. Scaphocerite of P. mesopotamicus. A, E, ♀ (pocl 7.0 mm), OUMNH. ZC. 2004 11 001, Las Turberas, Prat de Cabamas, Torreblanca, Spain; B C, F, ♀ (pocl 7.2 mm), NHM 1959.10.17.72 82, Roose Cote, Power Station, Morecambe Bay, United Kingdom; D, G H, ♀ (pocl 6.9 mm), SMF TSH 30 / 88, Khabur river, Syria. Scale bar indicates 3 mm (A), 1 mm (B E), 1.5 mm (F), 2 mm (G), 5 mm (H)." httpUri="https://zenodo.org/record/287667/files/figure.png" pageId="22" pageNumber="49">Figure 10</figureCitation>
C); first maxilliped endites are arranged almost in a straight line forming just a notch between them (
<figureCitation id="2FF62A5FFFACFFFB1612FDE895AEE938" box="[582,679,604,625]" captionStart="FIGURE 10. A" captionStartId="20.[151,250,1556,1578]" captionTargetBox="[201,1401,224,1513]" captionTargetId="figure@20.[180,1412,209,1518]" captionTargetPageId="20" captionText="FIGURE 10. A. Fifth pleuron of P. zariquieyi. B. Maxillula of P. varians. C. Telson distal part of P. v a r i an s. D. Telson distal part of P. mesopotamicus. E. Telson distal part of P. zariquieyi. F. First maxilliped of P. varians. G. Third maxilliped of P. mesopotamicus. H. Scaphocerite of P. mesopotamicus. A, E, ♀ (pocl 7.0 mm), OUMNH. ZC. 2004 11 001, Las Turberas, Prat de Cabamas, Torreblanca, Spain; B C, F, ♀ (pocl 7.2 mm), NHM 1959.10.17.72 82, Roose Cote, Power Station, Morecambe Bay, United Kingdom; D, G H, ♀ (pocl 6.9 mm), SMF TSH 30 / 88, Khabur river, Syria. Scale bar indicates 3 mm (A), 1 mm (B E), 1.5 mm (F), 2 mm (G), 5 mm (H)." httpUri="https://zenodo.org/record/287667/files/figure.png" pageId="22" pageNumber="49">Figure 10</figureCitation>
F); branchiostegal tooth marginal (
<figureCitation id="2FF62A5FFFACFFFB17ADFDE89345E938" box="[1017,1100,604,625]" captionStart="FIGURE 2" captionStartId="5.[151,250,1915,1937]" captionTargetBox="[156,1426,206,1914]" captionTargetId="figure@5.[156,1436,177,1916]" captionTargetPageId="5" captionText="FIGURE 2. Palaemon antennarius H. Milne Edwards, 1837. A. Whole animal, lateral view. B. Eye, dorso-mesial view. C. Frontal region. D. Antennula. E. Fifth pleuron. F. Uropod. G. Scaphocerite. Lectotype ♀ (pocl 7.7 mm), MNHN IU 2013 9296 (= MNHN Na 1500), Trasimeno Lake, Italy. Scale bar indicates 5 mm (A), 1 mm (B), 3 mm (C, D, F, G), 2 mm (E)." httpUri="https://zenodo.org/record/287659/files/figure.png" pageId="22" pageNumber="49">Figure 2</figureCitation>
A in
<bibRefCitation id="D35C4B2BFFACFFFB10D5FDE8961AE9C7" author="Gonzalez-Ortegon" pageId="22" pageNumber="49" refString="Gonzalez-Ortegon, E. &amp; Cuesta, J. A. (2006) An illustrated key to species of Palaemon and Palaemonetes (Crustacea: Decapoda: Caridea) from European waters, including the alien species Palaemon macrodactylus. Journal of the Marine Biological Association of the United Kingdom, 86, 93 - 102. http: // dx. doi. org / 10.1017 / s 0025315406012896" type="journal article" year="2006">González-Ortegón &amp; Cuesta, 2006</bibRefCitation>
)........................................................................................
<taxonomicName id="70CD4D59FFACFFFB1168FDCD9295E9C7" ID-CoL="cb001304-7f39-4d6d-b4e7-84d5ff5241a0" box="[1340,1436,633,654]" class="Malacostraca" family="Palaemonidae" genus="Palaemon" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="49" phylum="Arthropoda" rank="species" species="varians">
<emphasis id="85B9EAC8FFACFFFB1168FDCD9295E9C7" box="[1340,1436,633,654]" italics="true" pageId="22" pageNumber="49">P. varians</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="0C392E7FFFACFFFB14C3FD229292E877" pageId="22" pageNumber="49">
<paragraph id="B77236DAFFACFFFB14C3FD229295E9AF" blockId="22.[151,1436,222,1154]" pageId="22" pageNumber="49">
<keyLead id="0C3C95EFFFACFFFB14C3FD229295E9AF" pageId="22" pageNumber="49">
5. Telson with 4050 pairs of marginal lanceolate setae (
<figureCitation id="2FF62A5FFFACFFFB16BCFD22944AE9E2" box="[744,835,662,683]" captionStart="FIGURE 8" captionStartId="15.[151,250,1102,1124]" captionTargetBox="[156,1433,180,1075]" captionTargetId="figure@15.[156,1437,177,1081]" captionTargetPageId="15" captionText="FIGURE 8. Palaemon colossus sp. nov. A. First pleopod male. B. First pleopod female. C. Telson. D. Marginal lanceolate setae of telson. A ♂ (pocl 6.0 mm), ZMAUTH G 3 30, Epta Piges, Rhodes Island, Greece, 2002, coll. Anastasiadou Ch.; B D holotype ♀ (pocl 8.5 mm), OUMNH. ZC. 2014 13 001, Epta Piges, Rhodes Island, Greece. Scale bar indicates 1 mm (A C), 0.4 mm (D)." httpUri="https://zenodo.org/record/287665/files/figure.png" pageId="22" pageNumber="49">Figures 8</figureCitation>
C, 8D); in males, endopod of the first pleopod deeply concave on the mesial portion of inner margin (
<figureCitation id="2FF62A5FFFACFFFB160BFD0095BAE980" box="[607,691,692,713]" captionStart="FIGURE 8" captionStartId="15.[151,250,1102,1124]" captionTargetBox="[156,1433,180,1075]" captionTargetId="figure@15.[156,1437,177,1081]" captionTargetPageId="15" captionText="FIGURE 8. Palaemon colossus sp. nov. A. First pleopod male. B. First pleopod female. C. Telson. D. Marginal lanceolate setae of telson. A ♂ (pocl 6.0 mm), ZMAUTH G 3 30, Epta Piges, Rhodes Island, Greece, 2002, coll. Anastasiadou Ch.; B D holotype ♀ (pocl 8.5 mm), OUMNH. ZC. 2014 13 001, Epta Piges, Rhodes Island, Greece. Scale bar indicates 1 mm (A C), 0.4 mm (D)." httpUri="https://zenodo.org/record/287665/files/figure.png" pageId="22" pageNumber="49">Figure 8</figureCitation>
A); in females, endopod of the first pleopod with concave inner margin and rounded tip (
<figureCitation id="2FF62A5FFFACFFFB150FFD6596A5E9AF" box="[347,428,721,742]" captionStart="FIGURE 8" captionStartId="15.[151,250,1102,1124]" captionTargetBox="[156,1433,180,1075]" captionTargetId="figure@15.[156,1437,177,1081]" captionTargetPageId="15" captionText="FIGURE 8. Palaemon colossus sp. nov. A. First pleopod male. B. First pleopod female. C. Telson. D. Marginal lanceolate setae of telson. A ♂ (pocl 6.0 mm), ZMAUTH G 3 30, Epta Piges, Rhodes Island, Greece, 2002, coll. Anastasiadou Ch.; B D holotype ♀ (pocl 8.5 mm), OUMNH. ZC. 2014 13 001, Epta Piges, Rhodes Island, Greece. Scale bar indicates 1 mm (A C), 0.4 mm (D)." httpUri="https://zenodo.org/record/287665/files/figure.png" pageId="22" pageNumber="49">Figure 8</figureCitation>
B)..................................................................
<taxonomicName id="70CD4D59FFACFFFB10B4FD659240E9AF" ID-CoL="d24dc7f6-ed58-414c-9b2e-46851b317568" box="[1248,1353,721,742]" class="Malacostraca" family="Palaemonidae" genus="Palaemon" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="49" phylum="Arthropoda" rank="species" species="colossus" status="sp. nov.">
<emphasis id="85B9EAC8FFACFFFB10B4FD659240E9AF" box="[1248,1353,721,742]" italics="true" pageId="22" pageNumber="49">P. colossus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9E8A57B3FFACFFFB111BFD649295E9AF" box="[1359,1436,720,742]" pageId="22" pageNumber="49" rank="species">
<emphasis id="85B9EAC8FFACFFFB111BFD649295E9AF" bold="true" box="[1359,1436,720,742]" pageId="22" pageNumber="49">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="B77236DAFFACFFFB14C3FD5A9292E877" blockId="22.[151,1436,222,1154]" pageId="22" pageNumber="49">
<keyLead id="0C3C95EFFFACFFFB14C3FD5A9292E877" pageId="22" pageNumber="49">
- Telson with 06 pairs of marginal lanceolate setae just proximally (
<figureCitation id="2FF62A5FFFACFFFB1733FD5A94B3E84A" box="[871,954,750,771]" captionStart="FIGURE 5" captionStartId="8.[151,250,1080,1102]" captionTargetBox="[242,1344,193,1051]" captionTargetId="figure@8.[242,1344,193,1059]" captionTargetPageId="8" captionText="FIGURE 5. Palaemon antennarius H. Milne Edwards, 1837. A. Telson. B. Telson distal part. Lectotype ♀ (pocl 7.7 mm), MNHN IU 2013 9296 (= MNHN Na 1500), Trasimeno Lake, Italy. Scale bar indicates 3 mm (A), 1 mm (B)." httpUri="https://zenodo.org/record/287662/files/figure.png" pageId="22" pageNumber="49">Figure 5</figureCitation>
A); in males, endopod of the first pleopod slightly concave on the mesial portion of inner margin (
<figureCitation id="2FF62A5FFFACFFFB16F3FCB895F3E868" box="[679,762,780,801]" captionStart="FIGURE 4" captionStartId="7.[151,250,1802,1824]" captionTargetBox="[177,1406,203,1736]" captionTargetId="figure@7.[164,1423,193,1749]" captionTargetPageId="7" captionText="FIGURE 4. Palaemon antennarius H. Milne Edwards, 1837. A. First pereiopod. B. Second pereiopod. C. Third pereiopod. D. Fourth pereiopod. E. Fifth pereiopod. F. First pleopod male. G. Second pleopod male. H. First pleopod female. I. Second pleopod female. J. Third pleopod female. K. Fifth pleopod female. A E, H K ♀ paralectotype (pocl 8.0 mm), MNHN IU 2013 4023 (= MNHN Na 1500), Trasimeno Lake, Italy; F G ♂ (pocl 6.6 mm), Trasimeno Lake, 6.9. 1974, coll. Froglia C. &amp; E. Peloni. Scale bar indicates 2 mm." httpUri="https://zenodo.org/record/287661/files/figure.png" pageId="22" pageNumber="49">Figure 4</figureCitation>
F); in females, endopod of the first pleopod lanceolate (
<figureCitation id="2FF62A5FFFACFFFB1159FCB89269E868" box="[1293,1376,780,801]" captionStart="FIGURE 4" captionStartId="7.[151,250,1802,1824]" captionTargetBox="[177,1406,203,1736]" captionTargetId="figure@7.[164,1423,193,1749]" captionTargetPageId="7" captionText="FIGURE 4. Palaemon antennarius H. Milne Edwards, 1837. A. First pereiopod. B. Second pereiopod. C. Third pereiopod. D. Fourth pereiopod. E. Fifth pereiopod. F. First pleopod male. G. Second pleopod male. H. First pleopod female. I. Second pleopod female. J. Third pleopod female. K. Fifth pleopod female. A E, H K ♀ paralectotype (pocl 8.0 mm), MNHN IU 2013 4023 (= MNHN Na 1500), Trasimeno Lake, Italy; F G ♂ (pocl 6.6 mm), Trasimeno Lake, 6.9. 1974, coll. Froglia C. &amp; E. Peloni. Scale bar indicates 2 mm." httpUri="https://zenodo.org/record/287661/files/figure.png" pageId="22" pageNumber="49">Figure 4</figureCitation>
H).............................................................................................
<taxonomicName id="70CD4D59FFACFFFB1146FC9D9292E877" ID-CoL="63b91d48-a5f3-4939-8b96-863444859bda" box="[1298,1435,809,830]" class="Malacostraca" family="Palaemonidae" genus="Palaemonetes" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="49" phylum="Arthropoda" rank="species" species="antennarius">
<emphasis id="85B9EAC8FFACFFFB1146FC9D9292E877" box="[1298,1435,809,830]" italics="true" pageId="22" pageNumber="49">P. antennarius</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="0C392E7FFFACFFFB14C3FCF29292E898" pageId="22" pageNumber="49">
<paragraph id="B77236DAFFACFFFB14C3FCF29291E8DF" blockId="22.[151,1436,222,1154]" pageId="22" pageNumber="49">
<keyLead id="0C3C95EFFFACFFFB14C3FCF29291E8DF" pageId="22" pageNumber="49">
6. Telson with two plumose setae on posterior margin (
<figureCitation id="2FF62A5FFFACFFFB168FFCF29427E812" box="[731,814,838,859]" captionStart="FIGURE 6" captionStartId="12.[151,250,1921,1943]" captionTargetBox="[182,1402,201,1889]" captionTargetId="figure@12.[178,1402,193,1898]" captionTargetPageId="12" captionText="FIGURE 6. Palaemon migratorius (Heller, 1862). A. Whole animal, lateral view. B. Fifth pleuron. C. First maxilliped. D. Thoracic sternal armature. E. Abdominal sternal armature. F. Telson distal part. A B, D F lectotype ♀ (pocl 8.9 mm), NHMW 25559, Egypt; C ♀ paralectotype (pocl 6.9 mm), NHMW 7733, Egypt. Scale bar indicates 2 mm (A), 1 mm (B, C), 1.5 mm (D, E), 0.5 mm (F)." httpUri="https://zenodo.org/record/287663/files/figure.png" pageId="22" pageNumber="49">Figure 6</figureCitation>
c in
<bibRefCitation id="D35C4B2BFFACFFFB170DFCF294E2E812" author="Holthuis" box="[857,1003,838,859]" pageId="22" pageNumber="49" refString="Holthuis, L. B. (1961) Report on a collection of Crustacea Decapoda and Stomatopoda from Turkey and Balkans. Zoologische Verhandelingen, Leiden, 47, 1 - 67." type="journal article" year="1961">Holthuis, 1961</bibRefCitation>
); endopod of first pleopod of male almost as long as exopod (
<figureCitation id="2FF62A5FFFACFFFB1529FCD096C4E830" box="[381,461,868,889]" captionStart="FIGURE 8" captionStartId="15.[151,250,1102,1124]" captionTargetBox="[156,1433,180,1075]" captionTargetId="figure@15.[156,1437,177,1081]" captionTargetPageId="15" captionText="FIGURE 8. Palaemon colossus sp. nov. A. First pleopod male. B. First pleopod female. C. Telson. D. Marginal lanceolate setae of telson. A ♂ (pocl 6.0 mm), ZMAUTH G 3 30, Epta Piges, Rhodes Island, Greece, 2002, coll. Anastasiadou Ch.; B D holotype ♀ (pocl 8.5 mm), OUMNH. ZC. 2014 13 001, Epta Piges, Rhodes Island, Greece. Scale bar indicates 1 mm (A C), 0.4 mm (D)." httpUri="https://zenodo.org/record/287665/files/figure.png" pageId="22" pageNumber="49">Figure 8</figureCitation>
c in
<bibRefCitation id="D35C4B2BFFACFFFB15A3FCD0958EE830" author="Holthuis" box="[503,647,868,889]" pageId="22" pageNumber="49" refString="Holthuis, L. B. (1961) Report on a collection of Crustacea Decapoda and Stomatopoda from Turkey and Balkans. Zoologische Verhandelingen, Leiden, 47, 1 - 67." type="journal article" year="1961">Holthuis, 1961</bibRefCitation>
); appendix masculina longer than the endopod of the second pleopod (
<figureCitation id="2FF62A5FFFACFFFB1172FCD0927EE830" box="[1318,1399,868,889]" captionStart="FIGURE 8" captionStartId="15.[151,250,1102,1124]" captionTargetBox="[156,1433,180,1075]" captionTargetId="figure@15.[156,1437,177,1081]" captionTargetPageId="15" captionText="FIGURE 8. Palaemon colossus sp. nov. A. First pleopod male. B. First pleopod female. C. Telson. D. Marginal lanceolate setae of telson. A ♂ (pocl 6.0 mm), ZMAUTH G 3 30, Epta Piges, Rhodes Island, Greece, 2002, coll. Anastasiadou Ch.; B D holotype ♀ (pocl 8.5 mm), OUMNH. ZC. 2014 13 001, Epta Piges, Rhodes Island, Greece. Scale bar indicates 1 mm (A C), 0.4 mm (D)." httpUri="https://zenodo.org/record/287665/files/figure.png" pageId="22" pageNumber="49">Figure 8</figureCitation>
d in
<bibRefCitation id="D35C4B2BFFACFFFB14B5FC359679E8DF" author="Holthuis" box="[225,368,897,918]" pageId="22" pageNumber="49" refString="Holthuis, L. B. (1961) Report on a collection of Crustacea Decapoda and Stomatopoda from Turkey and Balkans. Zoologische Verhandelingen, Leiden, 47, 1 - 67." type="journal article" year="1961">Holthuis, 1961</bibRefCitation>
)..............................................................................
<emphasis id="85B9EAC8FFACFFFB1178FC359291E8DF" box="[1324,1432,897,918]" italics="true" pageId="22" pageNumber="49">P. t u rc o r u m</emphasis>
</keyLead>
</paragraph>
<paragraph id="B77236DAFFACFFFB14C3FC2A9292E898" blockId="22.[151,1436,222,1154]" pageId="22" pageNumber="49">
<keyLead id="0C3C95EFFFACFFFB14C3FC2A9292E898" pageId="22" pageNumber="49">
- Telson with 46 plumose setae on posterior margin (
<figureCitation id="2FF62A5FFFACFFFB1687FC2A9438E8FA" box="[723,817,926,947]" captionStart="FIGURE 10. A" captionStartId="20.[151,250,1556,1578]" captionTargetBox="[201,1401,224,1513]" captionTargetId="figure@20.[180,1412,209,1518]" captionTargetPageId="20" captionText="FIGURE 10. A. Fifth pleuron of P. zariquieyi. B. Maxillula of P. varians. C. Telson distal part of P. v a r i an s. D. Telson distal part of P. mesopotamicus. E. Telson distal part of P. zariquieyi. F. First maxilliped of P. varians. G. Third maxilliped of P. mesopotamicus. H. Scaphocerite of P. mesopotamicus. A, E, ♀ (pocl 7.0 mm), OUMNH. ZC. 2004 11 001, Las Turberas, Prat de Cabamas, Torreblanca, Spain; B C, F, ♀ (pocl 7.2 mm), NHM 1959.10.17.72 82, Roose Cote, Power Station, Morecambe Bay, United Kingdom; D, G H, ♀ (pocl 6.9 mm), SMF TSH 30 / 88, Khabur river, Syria. Scale bar indicates 3 mm (A), 1 mm (B E), 1.5 mm (F), 2 mm (G), 5 mm (H)." httpUri="https://zenodo.org/record/287667/files/figure.png" pageId="22" pageNumber="49">Figure 10</figureCitation>
E); endopod of first pleopod of male clearly shorter than the exopod; appendix masculina shorter than the endopod of the second pleopod................................
<taxonomicName id="70CD4D59FFACFFFB1170FC089292E898" ID-CoL="f08751a0-ee04-4598-ad24-1799de201b75" box="[1316,1435,956,977]" class="Malacostraca" family="Palaemonidae" genus="Palaemon" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="49" phylum="Arthropoda" rank="species" species="zariquieyi">
<emphasis id="85B9EAC8FFACFFFB1170FC089292E898" box="[1316,1435,956,977]" italics="true" pageId="22" pageNumber="49">P. zariquieyi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="0C392E7FFFACFFFB14C3FC6E9292EF45" pageId="22" pageNumber="49">
<paragraph id="B77236DAFFACFFFB14C3FC6E9292E8A7" blockId="22.[151,1436,222,1154]" box="[151,1435,985,1007]" pageId="22" pageNumber="49">
<keyLead id="0C3C95EFFFACFFFB14C3FC6E9292E8A7" box="[151,1435,985,1007]" pageId="22" pageNumber="49">
7. Telson always with two plumose setae on posterior margin (
<figureCitation id="2FF62A5FFFACFFFB1774FC6E947BE8A6" box="[800,882,986,1007]" captionStart="FIGURE 9" captionStartId="19.[151,250,1877,1899]" captionTargetBox="[163,1435,197,1846]" captionTargetId="figure@19.[151,1435,193,1847]" captionTargetPageId="19" captionText="FIGURE 9. Palaemon minos sp. nov. A. Whole animal, lateral view. B. Fifth pleuron. C. Telson distal part. D. Thoracic sternal armature. E. Abdominal sternal armature. A E holotype ♂ (pocl 4.5 mm), OUMNH. ZC. 2014 13 003, Kourna Lake, Crete, Greece; D E ♀ (pocl 7.0 mm), ZMAUTH G 3 31, Kourna Lake, Crete, Greece; Scale bar indicates 4 mm (A), 1.5 mm (B, D, E), 0.5 mm (C)." httpUri="https://zenodo.org/record/287666/files/figure.png" pageId="22" pageNumber="49">Figure 9</figureCitation>
C)..............................
<taxonomicName id="70CD4D59FFACFFFB10A3FC6E9241E8A6" ID-CoL="c479a912-02d0-4a87-b296-e4a6f05a954d" box="[1271,1352,986,1007]" class="Malacostraca" family="Palaemonidae" genus="Palaemon" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="49" phylum="Arthropoda" rank="species" species="minos" status="sp. nov.">
<emphasis id="85B9EAC8FFACFFFB10A3FC6E9241E8A6" box="[1271,1352,986,1007]" italics="true" pageId="22" pageNumber="49">P. minos</emphasis>
</taxonomicName>
<emphasis id="85B9EAC8FFACFFFB111BFC6D9292E8A7" bold="true" box="[1359,1435,985,1007]" pageId="22" pageNumber="49">
<taxonomicNameLabel id="9E8A57B3FFACFFFB111BFC6D9292E8A7" box="[1359,1435,985,1007]" pageId="22" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
<paragraph id="B77236DAFFACFFFB14C3FC439292EF45" blockId="22.[151,1436,222,1154]" box="[151,1435,1015,1036]" pageId="22" pageNumber="49">
<keyLead id="0C3C95EFFFACFFFB14C3FC439292EF45" box="[151,1435,1015,1036]" pageId="22" pageNumber="49">
- Telson with 412 plumose setae on posterior margin (
<figureCitation id="2FF62A5FFFACFFFB16B6FC439437EF45" box="[738,830,1015,1036]" captionStart="FIGURE 6" captionStartId="12.[151,250,1921,1943]" captionTargetBox="[182,1402,201,1889]" captionTargetId="figure@12.[178,1402,193,1898]" captionTargetPageId="12" captionText="FIGURE 6. Palaemon migratorius (Heller, 1862). A. Whole animal, lateral view. B. Fifth pleuron. C. First maxilliped. D. Thoracic sternal armature. E. Abdominal sternal armature. F. Telson distal part. A B, D F lectotype ♀ (pocl 8.9 mm), NHMW 25559, Egypt; C ♀ paralectotype (pocl 6.9 mm), NHMW 7733, Egypt. Scale bar indicates 2 mm (A), 1 mm (B, C), 1.5 mm (D, E), 0.5 mm (F)." httpUri="https://zenodo.org/record/287663/files/figure.png" pageId="22" pageNumber="49">Figures 6</figureCitation>
F, 10D)........................................... 8
</keyLead>
</paragraph>
</keyStep>
<keyStep id="0C392E7FFFACFFFB14C3FBA19292EFCB" pageId="22" pageNumber="49">
<paragraph id="B77236DAFFACFFFB14C3FBA19292EF0E" blockId="22.[151,1436,222,1154]" pageId="22" pageNumber="49">
<keyLead id="0C3C95EFFFACFFFB14C3FBA19292EF0E" pageId="22" pageNumber="49">
8. Telson with 1012 plumose setae on posterior margin (
<figureCitation id="2FF62A5FFFACFFFB16A5FBA19459EF63" box="[753,848,1045,1066]" captionStart="FIGURE 10. A" captionStartId="20.[151,250,1556,1578]" captionTargetBox="[201,1401,224,1513]" captionTargetId="figure@20.[180,1412,209,1518]" captionTargetPageId="20" captionText="FIGURE 10. A. Fifth pleuron of P. zariquieyi. B. Maxillula of P. varians. C. Telson distal part of P. v a r i an s. D. Telson distal part of P. mesopotamicus. E. Telson distal part of P. zariquieyi. F. First maxilliped of P. varians. G. Third maxilliped of P. mesopotamicus. H. Scaphocerite of P. mesopotamicus. A, E, ♀ (pocl 7.0 mm), OUMNH. ZC. 2004 11 001, Las Turberas, Prat de Cabamas, Torreblanca, Spain; B C, F, ♀ (pocl 7.2 mm), NHM 1959.10.17.72 82, Roose Cote, Power Station, Morecambe Bay, United Kingdom; D, G H, ♀ (pocl 6.9 mm), SMF TSH 30 / 88, Khabur river, Syria. Scale bar indicates 3 mm (A), 1 mm (B E), 1.5 mm (F), 2 mm (G), 5 mm (H)." httpUri="https://zenodo.org/record/287667/files/figure.png" pageId="22" pageNumber="49">Figure 10</figureCitation>
D); third maxilliped exopod shorter than ischiomerus (
<figureCitation id="2FF62A5FFFACFFFB110FFBA197F3EF0E" captionStart="FIGURE 10. A" captionStartId="20.[151,250,1556,1578]" captionTargetBox="[201,1401,224,1513]" captionTargetId="figure@20.[180,1412,209,1518]" captionTargetPageId="20" captionText="FIGURE 10. A. Fifth pleuron of P. zariquieyi. B. Maxillula of P. varians. C. Telson distal part of P. v a r i an s. D. Telson distal part of P. mesopotamicus. E. Telson distal part of P. zariquieyi. F. First maxilliped of P. varians. G. Third maxilliped of P. mesopotamicus. H. Scaphocerite of P. mesopotamicus. A, E, ♀ (pocl 7.0 mm), OUMNH. ZC. 2004 11 001, Las Turberas, Prat de Cabamas, Torreblanca, Spain; B C, F, ♀ (pocl 7.2 mm), NHM 1959.10.17.72 82, Roose Cote, Power Station, Morecambe Bay, United Kingdom; D, G H, ♀ (pocl 6.9 mm), SMF TSH 30 / 88, Khabur river, Syria. Scale bar indicates 3 mm (A), 1 mm (B E), 1.5 mm (F), 2 mm (G), 5 mm (H)." httpUri="https://zenodo.org/record/287667/files/figure.png" pageId="22" pageNumber="49">Figure 10</figureCitation>
G); outer margin of scaphocerite concave (
<figureCitation id="2FF62A5FFFACFFFB16C6FB8695F9EF0E" box="[658,752,1074,1095]" captionStart="FIGURE 10. A" captionStartId="20.[151,250,1556,1578]" captionTargetBox="[201,1401,224,1513]" captionTargetId="figure@20.[180,1412,209,1518]" captionTargetPageId="20" captionText="FIGURE 10. A. Fifth pleuron of P. zariquieyi. B. Maxillula of P. varians. C. Telson distal part of P. v a r i an s. D. Telson distal part of P. mesopotamicus. E. Telson distal part of P. zariquieyi. F. First maxilliped of P. varians. G. Third maxilliped of P. mesopotamicus. H. Scaphocerite of P. mesopotamicus. A, E, ♀ (pocl 7.0 mm), OUMNH. ZC. 2004 11 001, Las Turberas, Prat de Cabamas, Torreblanca, Spain; B C, F, ♀ (pocl 7.2 mm), NHM 1959.10.17.72 82, Roose Cote, Power Station, Morecambe Bay, United Kingdom; D, G H, ♀ (pocl 6.9 mm), SMF TSH 30 / 88, Khabur river, Syria. Scale bar indicates 3 mm (A), 1 mm (B E), 1.5 mm (F), 2 mm (G), 5 mm (H)." httpUri="https://zenodo.org/record/287667/files/figure.png" pageId="22" pageNumber="49">Figure 10</figureCitation>
H).........................................
<taxonomicName id="70CD4D59FFACFFFB10B8FB869292EF0E" ID-CoL="e65726af-e30b-45d3-80b0-7073d8e2f66b" box="[1260,1435,1074,1095]" class="Malacostraca" family="Palaemonidae" genus="Palaemon" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="49" phylum="Arthropoda" rank="species" species="mesopotamicus">
<emphasis id="85B9EAC8FFACFFFB10B8FB869292EF0E" box="[1260,1435,1074,1095]" italics="true" pageId="22" pageNumber="49">P. mesopotamicus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B77236DAFFACFFFB14C3FBFB9292EFCB" blockId="22.[151,1436,222,1154]" pageId="22" pageNumber="49">
<keyLead id="0C3C95EFFFACFFFB14C3FBFB9292EFCB" pageId="22" pageNumber="49">
- Telson with 47 plumose setae on posterior margin (
<figureCitation id="2FF62A5FFFACFFFB16BCFBFB9434EF2D" box="[744,829,1103,1124]" captionStart="FIGURE 6" captionStartId="12.[151,250,1921,1943]" captionTargetBox="[182,1402,201,1889]" captionTargetId="figure@12.[178,1402,193,1898]" captionTargetPageId="12" captionText="FIGURE 6. Palaemon migratorius (Heller, 1862). A. Whole animal, lateral view. B. Fifth pleuron. C. First maxilliped. D. Thoracic sternal armature. E. Abdominal sternal armature. F. Telson distal part. A B, D F lectotype ♀ (pocl 8.9 mm), NHMW 25559, Egypt; C ♀ paralectotype (pocl 6.9 mm), NHMW 7733, Egypt. Scale bar indicates 2 mm (A), 1 mm (B, C), 1.5 mm (D, E), 0.5 mm (F)." httpUri="https://zenodo.org/record/287663/files/figure.png" pageId="22" pageNumber="49">Figure 6</figureCitation>
F); third maxilliped exopod equal or longer than ischiomerus; outer margin of scaphocerite straight...........................................................
<taxonomicName id="70CD4D59FFACFFFB1146FBD99292EFCB" ID-CoL="38a39fca-9780-4a94-8e00-1bed815876db" box="[1298,1435,1133,1154]" class="Malacostraca" family="Palaemonidae" genus="Palaemon" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="49" phylum="Arthropoda" rank="species" species="migratorius">
<emphasis id="85B9EAC8FFACFFFB1146FBD99292EFCB" box="[1298,1435,1133,1154]" italics="true" pageId="22" pageNumber="49">P. migratorius</emphasis>
</taxonomicName>
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</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>