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<mods:title id="D4302C794D42A577F2C834D26F9FB446">Generic revision of the staphylinid beetle tribe Hoplandriini (Insecta: Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<taxonomicName id="708EEC94504EFFF9C2E6F9F7FB6AF989" ID-CoL="8VRR4" authority="KRAATZ, 1857" authorityName="Kraatz" authorityYear="1857" box="[941,1258,1627,1651]" class="Insecta" family="Staphylinidae" genus="Hoplandria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="genus">
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<figureCitation id="2FB58B92504EFFF9C5B0F9F0FAF0F989" box="[1275,1392,1627,1651]" captionStart-0="Figure 2" captionStart-1="Figure 3" captionStart-2="Figure 4" captionStart-3="Figure 5" captionStart-4="Figure 6" captionStart-5="Figure 7" captionStart-6="Figure 8" captionStart-7="Figure 9" captionStart-8="Figure 10" captionStart-9="Figure 11" captionStartId-0="9.[155,233,1736,1755]" captionStartId-1="10.[142,220,1355,1374]" captionStartId-2="12.[142,220,1775,1794]" captionStartId-3="13.[155,233,1367,1386]" captionStartId-4="15.[163,241,1761,1780]" captionStartId-5="16.[142,220,1153,1172]" captionStartId-6="18.[142,220,1788,1807]" captionStartId-7="19.[155,233,1370,1389]" captionStartId-8="21.[155,233,1778,1797]" captionStartId-9="22.[142,220,1298,1317]" captionTargetBox-0="[165,1419,207,1701]" captionTargetBox-1="[142,1420,198,1318]" captionTargetBox-2="[165,1412,194,1729]" captionTargetBox-3="[158,1441,195,1329]" captionTargetBox-4="[158,1440,200,1725]" captionTargetBox-5="[164,1401,200,1115]" captionTargetBox-6="[171,1406,215,1740]" captionTargetBox-7="[167,1437,210,1332]" captionTargetBox-8="[158,1443,205,1739]" captionTargetBox-9="[148,1421,210,1263]" captionTargetId-0="figure-52@9.[155,1435,194,1715]" captionTargetId-1="figure-275@10.[142,1421,196,1322]" captionTargetId-2="figure-53@12.[142,1432,183,1753]" captionTargetId-3="figure-246@13.[154,1446,191,1337]" captionTargetId-4="figure-61@15.[154,1449,191,1734]" captionTargetId-5="figure-378@16.[153,1410,194,1120]" captionTargetId-6="figure-53@18.[146,1418,194,1755]" captionTargetId-7="figure-265@19.[155,1453,191,1341]" captionTargetId-8="figure-60@21.[154,1446,193,1750]" captionTargetId-9="figure-328@22.[133,1431,194,1271]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="12" captionTargetPageId-3="13" captionTargetPageId-4="15" captionTargetPageId-5="16" captionTargetPageId-6="18" captionTargetPageId-7="19" captionTargetPageId-8="21" captionTargetPageId-9="22" captionText-0="Figure 2. Hoplandria (Arrenhandria) laeviventris Casey. (A) Labrum. (B) Epipharynx. (C) Mandible, dorsal view. (D) Mandible, ventral view. (E) Maxilla. (F) Labium. (G) Hypopharynx." captionText-1="Figure 3. Hoplandria (Arrhenandria) laeviventris Casey. (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Aedeagus, lateral view. (D) Paramere, lateral view. (E) Spermatheca. (F) Left elytron, male (arrow to denticle)." captionText-2="Figure 4. Hoplandria (Genosema) pulchra Kraatz. (A) Labrum. (B) Epipharynx. (C) Mandible, dorsal view. (D) Mandible, ventral view. (E) Maxilla. (F) Labium. (G) Hypopharynx." captionText-3="Figure 5. Hoplandria (Genosema) pulchra Kraatz. (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Aedeagus, lateral view. (D) Paramere, lateral view. (E) Spermatheca. (F) Left elytron, male (arrow to denticle)." captionText-4="Figure 6. Hoplandria (Hoplandria) lateralis Melsheimer. (A) Habitus, dorsal view. (B) Labrum. (C) Epipharynx. (D) Mandible, dorsal view. (E) Mandible, ventral view. (F) Maxilla. (G) Labium. (H) Hypopharynx." captionText-5="Figure 7. Hoplandria (Hoplandria) lateralis Melsheimer. (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Tarsus of hind leg. (D) Aedeagus, lateral view. (E) Paramere, lateral view. (F) Spermatheca." captionText-6="Figure 8. Hoplandria (Lophomucter) laevicollis (Notman). (A) Labrum. (B) Epipharynx. (C) Mandible, dorsal view. (D) Mandible, ventral view. (E) Maxilla. (F) Labium. (G) Hypopharynx." captionText-7="Figure 9. Hoplandria (Lophomucter) laevicollis (Notman). (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Aedeagus, lateral view. (D) Paramere, lateral view. (E) Spermatheca." captionText-8="Figure 10. Hoplandria (Platonica) major (Sharp). (A) Habitus, dorsal view. (B) Labrum. (C) Epipharynx. (D) Mandible, dorsal view. (E) Mandible, ventral view. (F) Maxilla. (G) Labium. (H) Hypopharynx." captionText-9="Figure 11. Hoplandria (Platonica) major (Sharp). (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Aedeagus, lateral view. (D) Paramere, lateral view. (E) Spermatheca." figureDoi-0="http://doi.org/10.5281/zenodo.5437796" figureDoi-1="http://doi.org/10.5281/zenodo.5437798" figureDoi-2="http://doi.org/10.5281/zenodo.5437800" figureDoi-3="http://doi.org/10.5281/zenodo.5437802" figureDoi-4="http://doi.org/10.5281/zenodo.5437804" figureDoi-5="http://doi.org/10.5281/zenodo.5437808" figureDoi-6="http://doi.org/10.5281/zenodo.5437810" figureDoi-7="http://doi.org/10.5281/zenodo.5437812" figureDoi-8="http://doi.org/10.5281/zenodo.5437814" figureDoi-9="http://doi.org/10.5281/zenodo.5437816" httpUri-0="https://zenodo.org/record/5437796/files/figure.png" httpUri-1="https://zenodo.org/record/5437798/files/figure.png" httpUri-2="https://zenodo.org/record/5437800/files/figure.png" httpUri-3="https://zenodo.org/record/5437802/files/figure.png" httpUri-4="https://zenodo.org/record/5437804/files/figure.png" httpUri-5="https://zenodo.org/record/5437808/files/figure.png" httpUri-6="https://zenodo.org/record/5437810/files/figure.png" httpUri-7="https://zenodo.org/record/5437812/files/figure.png" httpUri-8="https://zenodo.org/record/5437814/files/figure.png" httpUri-9="https://zenodo.org/record/5437816/files/figure.png" pageId="3" pageNumber="86">FIGS 2–11</figureCitation>
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<bibRefCitation id="D31FEAE6504EFFF9C2FFF897FB02F8AB" author="Bradley JC" box="[948,1154,1852,1874]" pageId="3" pageNumber="86" refId="ref36895" refString="Bradley JC. 1930. A manual of the genera of beetles of America, north of Mexico. Ithaca, New York: Daw, Illston." type="book" year="1930">Bradley, 1930:84</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="362FB106504EFFF9C5D1F897FA17F8AB" author="Scheerpeltz O" box="[1178,1431,1852,1874]" page="20" pageId="3" pageNumber="86" year="1929">
<bibRefCitation id="D31FEAE6504EFFF9C5D1F897FA17F8AB" author="Scheerpeltz O" box="[1178,1431,1852,1874]" pageId="3" pageNumber="86" pagination="1 - 21" refId="ref39225" refString="Scheerpeltz O. 1929. Eine neue Staphyliniden-Gattung von der Insel Korfu nebst einer Bestimmungstabelle der palaarktischen Gattungen der Tribus Myrmedoniini. (14. Beitrag zur Kenntnis der palaarktischen Staphylinidenfauna). Koleopterologische Rundschau 15: 1 - 21." type="journal article" year="1929">Scheerpeltz, 1929:20</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="362FB106504EFFF9C278F8F0FBBCF88B" author="Scheerpeltz O" box="[819,1084,1883,1905]" page="1667" pageId="3" pageNumber="86" year="1934">
<bibRefCitation id="D31FEAE6504EFFF9C278F8F0FBBCF88B" author="Scheerpeltz O" box="[819,1084,1883,1905]" pageId="3" pageNumber="86" pagination="1501 - 1881" refId="ref39266" refString="Scheerpeltz O. 1934. Staphylinidae VIII: Supplementum II In: Schenkling S, ed. Coleopterorum Catalogus, Pars. 129. Berlin: Dr W. Junk, 1501 - 1881." type="book chapter" year="1934">Scheerpeltz, 1934:1667</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="362FB106504EFFF9C50DF8F0FAC8F88A" author="Blackwelder RE" box="[1094,1352,1882,1905]" page="557" pageId="3" pageNumber="86" year="1943">
<bibRefCitation id="D31FEAE6504EFFF9C50DF8F0FAC8F88A" author="Blackwelder RE" box="[1094,1352,1882,1905]" pageId="3" pageNumber="86" pagination="1 - 658" refId="ref36723" refString="Blackwelder RE. 1943. Monograph of the West Indian beetles of the family Staphylinidae. Bulletin of the United States National Museum 182: 1 - 658." type="journal article" year="1943">Blackwelder, 1943:557</bibRefCitation>
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. Black-
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</paragraph>
<caption id="E3F1C79F5049FFFEC1C5FF6EFC0AFED5" pageId="4" pageNumber="87" startId="4.[142,206,197,216]">
<paragraph id="B73197175049FFFEC1C5FF6EFC0AFED5" blockId="4.[142,1422,196,304]" pageId="4" pageNumber="87">
<emphasis id="85FA4B055049FFFEC1C5FF6EFF68FF2D" bold="true" box="[142,232,196,216]" pageId="4" pageNumber="87">Table 1.</emphasis>
Diagnostic characters that were used in the descriptions of genera and subgenera of
<taxonomicName id="708EEC945049FFFEC525FF6FFB79FF2D" ID-CoL="8VR8M" authorityName="CASEY" authorityYear="1910" box="[1134,1273,196,215]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="87" phylum="Arthropoda" rank="tribe" tribe="Hoplandriini">Hoplandriini</taxonomicName>
. The numbers in brackets refer to specific characters and are uniform from one description to the next in order to facilitate comparisons among taxa. In a few instances where there was a significant modification to a listed character or a character specific to a particular taxon, a lower case letter was added to the character number
</paragraph>
</caption>
<paragraph id="B73197175049FFFEC1C5FEFEFAACFE9C" blockId="4.[142,1423,341,1788]" box="[142,1324,341,359]" pageId="4" pageNumber="87">[1] Length of adult. [46] Width of apex of ligula of labium. [88] Presence of macrosetae on</paragraph>
</subSubSection>
<subSubSection id="FF94C49C5049FFFCC1C5FEC1FD02FE05" lastPageId="6" lastPageNumber="89" pageId="4" pageNumber="87" type="description">
<paragraph id="B73197175049FFFEC1C5FEC1FB55FE86" blockId="4.[142,1423,341,1788]" box="[142,1237,362,380]" pageId="4" pageNumber="87">[2] General body shape. [47] Shape of apex of ligula of labium. metasternum.</paragraph>
<paragraph id="B73197175049FFFEC1C5FE2BFAACFE6B" blockId="4.[142,1423,341,1788]" box="[142,1324,384,402]" pageId="4" pageNumber="87">[3] General reflectivity of body. [48] Presence of spines at apex/subapex of [89] Presence of macrosetae on</paragraph>
<paragraph id="B73197175049FFFEC1C5FE3EFB55FE5C" blockId="4.[142,1423,341,1788]" box="[142,1237,405,423]" pageId="4" pageNumber="87">[4] General punctation and ligula of labium. mesosternum.</paragraph>
<paragraph id="B73197175049FFFEC1A1FE00FB4EFE46" blockId="4.[142,1423,341,1788]" box="[234,1230,426,445]" pageId="4" pageNumber="87">microsculpturing of body. [49] Presence and length of medial setae [90] Tarsal formula.</paragraph>
<paragraph id="B73197175049FFFEC1C5FE6BFA86FE28" blockId="4.[142,1423,341,1788]" box="[142,1286,448,466]" pageId="4" pageNumber="87">[5] Pubescence concentration on head. on prementum of labium. [91] Shape of tarsal claws.</paragraph>
<paragraph id="B73197175049FFFEC1C5FE7EFAC2FE1D" blockId="4.[142,1423,341,1788]" box="[142,1346,469,487]" pageId="4" pageNumber="87">[6] Pubescence on abdominal terga. [50] Insertion of medial setae of labium. [92] Presence of empodial bristle.</paragraph>
<paragraph id="B73197175049FFFEC1C5FE41FAB3FE06" blockId="4.[142,1423,341,1788]" box="[142,1331,490,509]" pageId="4" pageNumber="87">[7] Pubescence on pronotum. [51] Presence of real pores in lateral pore [93] Length of empodial bristle.</paragraph>
<paragraph id="B73197175049FFFEC1C5FDABFAB7FDE8" blockId="4.[142,1423,341,1788]" box="[142,1335,512,530]" pageId="4" pageNumber="87">[8] Pubescence on elytra. field of labium. [94] Length of tarsal segment 1.</paragraph>
<paragraph id="B73197175049FFFEC1C5FDBEFAD6FDDD" blockId="4.[142,1423,341,1788]" box="[142,1366,533,551]" pageId="4" pageNumber="87">[9] Pubescence on abdominal sterna. [52] Presence of setal pores in lateral pore [95] Length of tarsal segments 2–4.</paragraph>
<paragraph id="B73197175049FFFEC1C5FD81FAB8FDC7" blockId="4.[142,1423,341,1788]" box="[142,1336,554,573]" pageId="4" pageNumber="87">[10] Head shape, comparison of width to field of labium. [96] Length of tarsal segment 5.</paragraph>
<paragraph id="B73197175049FFFEC1A1FDEAFB63FDA8" blockId="4.[142,1423,341,1788]" box="[234,1251,576,594]" pageId="4" pageNumber="87">length. [53] Size and shape of median pseudopore [97] Abdominal shape.</paragraph>
<paragraph id="B73197175049FFFEC1C5FDFEFABCFD86" blockId="4.[142,1423,341,1788]" pageId="4" pageNumber="87">[11] Eye size, comparison to length of field of labium. [98] Presence and location of basal head. [54] Composition of lateral pore fields. transverse depressions on</paragraph>
<paragraph id="B73197175049FFFEC1C5FD2BFB6FFD6B" blockId="4.[142,1423,341,1788]" box="[142,1263,640,658]" pageId="4" pageNumber="87">[12] Neck (present or absent). [55] Length of hypoglossal lobe of abdominal terga.</paragraph>
<paragraph id="B73197175049FFFEC1C5FD3EFA07FD5D" blockId="4.[142,1423,341,1788]" box="[142,1415,661,679]" pageId="4" pageNumber="87">[13] Infraorbital carina development. labium. [99] Presence of macrosetae on abdomen.</paragraph>
<paragraph id="B73197175049FFFEC1C5FD01FAF8FD46" blockId="4.[142,1423,341,1788]" box="[142,1400,682,701]" pageId="4" pageNumber="87">[14] Number of antennomeres. [56] Size of setae of hypoglossal lobe of [100] Presence of differentiated region of</paragraph>
<paragraph id="B73197175049FFFEC1C5FD6BFA0AFD2B" blockId="4.[142,1423,341,1788]" box="[142,1418,704,722]" pageId="4" pageNumber="87">[15] Shape of antennal segments 1–3. labium. microsculpturing on anterior corner</paragraph>
<paragraph id="B73197175049FFFEC1C5FD7EFABDFD1C" blockId="4.[142,1423,341,1788]" box="[142,1341,725,743]" pageId="4" pageNumber="87">[16] Shape of antennal segment 4. [57] Composition of setae of hypoglossal of abdominal sternum III.</paragraph>
<paragraph id="B73197175049FFFEC1C5FD41FAFBFD06" blockId="4.[142,1423,341,1788]" box="[142,1403,746,765]" pageId="4" pageNumber="87">[17] Shape of antennal segments 5–10. lobe of labium. [101] Setal pattern on abdominal tergum</paragraph>
<paragraph id="B73197175049FFFEC1C5FCABFBF5FCEB" blockId="4.[142,1423,341,1788]" box="[142,1141,768,786]" pageId="4" pageNumber="87">[18] Coeloconical sensilla (present or [58] Size of each labial palpus. IX.</paragraph>
<paragraph id="B73197175049FFFEC1A1FCBEFA04FCDD" blockId="4.[142,1423,341,1788]" box="[234,1412,789,807]" pageId="4" pageNumber="87">absent). [59] Number of segments of each labial [102] Presence of humeral carina or bump</paragraph>
<paragraph id="B73197175049FFFEC1C5FC81FA9AFCA8" blockId="4.[142,1423,341,1788]" pageId="4" pageNumber="87">[19] Medial pores of epipharyngeal area of palpus. on each elytron (male secondary labrum, size and distribution. [60] Composition of distal pore field. sexual characteristic).</paragraph>
<paragraph id="B73197175049FFFEC1C5FCFEFADCFC86" blockId="4.[142,1423,341,1788]" pageId="4" pageNumber="87">[20] Definition of longitudinal sensory [61] Length of article 1 of each labial [103] Presence of carina or conical process field. palpus (compared to article 2). on medioapical region of each</paragraph>
<paragraph id="B73197175049FFFEC1C5FC2BFB5CFC5D" blockId="4.[142,1423,341,1788]" pageId="4" pageNumber="87">[21] Pores between longitudinal sensory [62] Length of article 3 of each labial elytron (male secondary sexual field and lateral sclerotized areas. palpus (compared to article 2). characteristic).</paragraph>
<paragraph id="B73197175049FFFEC1C5FC01FAA5FC1D" blockId="4.[142,1423,341,1788]" pageId="4" pageNumber="87">[22] Mandibular asymmetry, right [63] Presence of twin pores of each labial [104] Presence of denticle, carina or bump mandible with distinct median palpus. on outer apical third of each elytron tooth. [64] Presence and number of median pores (male secondary sexual</paragraph>
<paragraph id="B73197175049FFFEC1C5FC41FB5CFC07" blockId="4.[142,1423,341,1788]" box="[142,1244,1002,1021]" pageId="4" pageNumber="87">[23] Shape of mandibular apex. of each labial palpus. characteristic).</paragraph>
<paragraph id="B73197175049FFFEC1C5FBABFA05FBEB" blockId="4.[142,1423,341,1788]" box="[142,1413,1024,1042]" pageId="4" pageNumber="87">[24] Size of condylar molar patch. [65] Shape of apical margin of mentum. [105] Presence of spine at posterior corner</paragraph>
<paragraph id="B73197175049FFFEC1C5FBBEFAEEFBDD" blockId="4.[142,1423,341,1788]" box="[142,1390,1045,1063]" pageId="4" pageNumber="87">[25] Composition of condylar molar patch. [66] Shape of antero-lateral angles of of abdominal sternum III (male</paragraph>
<paragraph id="B73197175049FFFEC1C5FB81FAF7FBC7" blockId="4.[142,1423,341,1788]" box="[142,1399,1066,1085]" pageId="4" pageNumber="87">[26] Arrangement of condylar molar patch. mentum. secondary sexual characteristic).</paragraph>
<paragraph id="B73197175049FFFEC1C5FBEBFAECFB9D" blockId="4.[142,1423,341,1788]" pageId="4" pageNumber="87">[27] Subcondylar molar patch (present or [67] General distribution of sensory pores [106] Presence of spine at posterior corner absent). of mentum. of abdominal sternum IV (male</paragraph>
<paragraph id="B73197175049FFFEC1C5FBC1FAF7FB87" blockId="4.[142,1423,341,1788]" box="[142,1399,1130,1149]" pageId="4" pageNumber="87">[28] Size of mandibular ‘velvety patch. [68] General shape of pronotum (dorsal secondary sexual characteristic).</paragraph>
<paragraph id="B73197175049FFFEC1C5FB2BFACCFB5D" blockId="4.[142,1423,341,1788]" pageId="4" pageNumber="87">[29] Composition of mandibular ‘velvety view). [107] Presence of longitudinal carina on patch. [69] Specific size of pronotum. abdominal tergum IV (male</paragraph>
<paragraph id="B73197175049FFFEC1C5FB01FAF1FB2B" blockId="4.[142,1423,341,1788]" pageId="4" pageNumber="87">[30] Setae on ventral aspect of outer basal [70] General shape of pronotum (lateral secondary sexual characteristic). angle of each mandible. view). [108] Presence of longitudinal carina on</paragraph>
<paragraph id="B73197175049FFFEC1C5FB7EFAD3FB1D" blockId="4.[142,1423,341,1788]" box="[142,1363,1237,1255]" pageId="4" pageNumber="87">[31] Length of each lacinia. [71] Setation of pronotum. abdominal tergum VII (male</paragraph>
<paragraph id="B73197175049FFFEC1C5FB41FAF7FB07" blockId="4.[142,1423,341,1788]" box="[142,1399,1258,1277]" pageId="4" pageNumber="87">[32] Shape of apex of each lacinia. [72] Orientation of setation of pronotum. secondary sexual characteristic).</paragraph>
<paragraph id="B73197175049FFFEC1C5FAABFA09FADD" blockId="4.[142,1423,341,1788]" pageId="4" pageNumber="87">[33] Length of teeth on adoral margin of [73] Visibility of hypomeron of pronotum [109] Presence of two transverse rows of each lacinia. in lateral view. macrosetae near apex of abdominal</paragraph>
<paragraph id="B73197175049FFFEC1C5FA81FB5CFAA8" blockId="4.[142,1423,341,1788]" pageId="4" pageNumber="87">[34] Placement of teeth/hairs on adoral [74] General size of elytra. tergum VIII (male secondary sexual margin. [75] Shape of margin of apico-lateral angle characteristic).</paragraph>
<paragraph id="B73197175049FFFEC1C5FAFEFAB6FA9D" blockId="4.[142,1423,341,1788]" box="[142,1334,1365,1383]" pageId="4" pageNumber="87">[35] Number of distinct regions of teeth/ of elytron. [110] General shape of aedeagus.</paragraph>
<paragraph id="B73197175049FFFEC1A1FAC0FAE4FA86" blockId="4.[142,1423,341,1788]" box="[234,1380,1386,1405]" pageId="4" pageNumber="87">hairs on adoral margin. [76] Size of elytra. [111] Presence of ventral projection on</paragraph>
<paragraph id="B73197175049FFFEC1C5FA2BFB30FA68" blockId="4.[142,1423,341,1788]" box="[142,1200,1408,1426]" pageId="4" pageNumber="87">[36] Number of spinose setae on dorsal [77] Presence of microsetae on each aedeagus.</paragraph>
<paragraph id="B73197175049FFFEC1A1FA3DFADFFA5C" blockId="4.[142,1423,341,1788]" box="[234,1375,1429,1447]" pageId="4" pageNumber="87">surface of each lacinea. elytron. [112] Presence of spinules/sclerites on</paragraph>
<paragraph id="B73197175049FFFEC1C5FA01FB47FA46" blockId="4.[142,1423,341,1788]" box="[142,1223,1450,1468]" pageId="4" pageNumber="87">[37] Width of each galea. [78] General distribution of microsetae on internal sac.</paragraph>
<paragraph id="B73197175049FFFEC1C5FA6BFB34FA1C" blockId="4.[142,1423,341,1788]" pageId="4" pageNumber="87">[38] Size of membranous area at apex of each elytron. [113] Length of apical lobe of each each galea. [79] Orientation of microsetation on each paramere.</paragraph>
<paragraph id="B73197175049FFFEC1C5FA41FB34F9EB" blockId="4.[142,1423,341,1788]" pageId="4" pageNumber="87">[39] Composition of setae at apex of each elytron. [114] Setation at apex of apical lobe of each galea. [80] Presence of medial carina on paramere.</paragraph>
<paragraph id="B73197175049FFFEC1C5F9BEFB41F9C6" blockId="4.[142,1423,341,1788]" pageId="4" pageNumber="87">[40] Number of articles of each maxillary mesosternum. [115] Shape of anterior margin of palpus (including pseudosegment). [81] Width of separation of mesocoxal paramerite.</paragraph>
<paragraph id="B73197175049FFFEC1C5F9EBFB41F99D" blockId="4.[142,1423,341,1788]" pageId="4" pageNumber="87">[41] Setation of articles 2–3 of each cavities. [116] Modifications to apex of condylite of maxillary palpus. [82] Length of mesocoxal process. paramerite.</paragraph>
<paragraph id="B73197175049FFFEC1C5F9C1FB74F986" blockId="4.[142,1423,341,1788]" box="[142,1268,1642,1661]" pageId="4" pageNumber="87">[42] Setation of article 1 of each maxillary [83] Presence of isthmus between meso- [117] Length of condylite.</paragraph>
<paragraph id="B73197175049FFFEC1A1F92AFAD5F968" blockId="4.[142,1423,341,1788]" box="[234,1365,1664,1682]" pageId="4" pageNumber="87">palpus. and metasternal processes. [118] General shape of spermatheca.</paragraph>
<footnote id="D4958B195049FFFEC1C5F93EFAD1F946" pageId="4" pageNumber="87">
<paragraph id="B73197175049FFFEC1C5F93EFAD1F946" blockId="4.[142,1423,341,1788]" pageId="4" pageNumber="87">[43] Setation of article 4 of each maxillary [84] Shape of apex of mesosternal process. [119] Shape of basal bulb of spermatheca. palpus. [85] Length of metasternal process. [120] Shape of neck of spermatheca.</paragraph>
</footnote>
<footnote id="D4958B195049FFFEC1C5F96BFAD5F91D" pageId="4" pageNumber="87">
<paragraph id="B73197175049FFFEC1C5F96BFAD5F91D" blockId="4.[142,1423,341,1788]" pageId="4" pageNumber="87">[44] Setation of pseudosegment of each [86] Presence of medial carina on [121] Sclerotization of tube of spermatheca. maxillary palpus. metasternal process. [122] Degree of coils/bends of tube of</paragraph>
</footnote>
<footnote id="D4958B195049FFFEC1C5F941FB4FF906" box="[142,1231,1770,1788]" pageId="4" pageNumber="87">
<paragraph id="B73197175049FFFEC1C5F941FB4FF906" blockId="4.[142,1423,341,1788]" box="[142,1231,1770,1788]" pageId="4" pageNumber="87">[45] Length of ligula of labium. [87] Shape of apex of metasternal process. spermatheca.</paragraph>
</footnote>
<table id="C58E65B750480005C1D5FF68FA1FF8A3" box="[158,1439,195,1881]" pageId="5" pageNumber="88">
<paragraph id="B73197175048FFFFC0F1FF79FB30FF13" blockId="5.[442,1200,210,234]" box="[442,1200,210,234]" pageId="5" pageNumber="88">
KEY TO THE GENERA OF THE
<collectionCode id="D19F0FD25048FFFFC247FF7DFCCEFF13" box="[780,846,214,233]" pageId="5" pageNumber="88">TRIBE</collectionCode>
<taxonomicName id="708EEC945048FFFFC21DFF79FB30FF13" authority="OF THE WORLD" authorityName="OF THE WORLD" box="[854,1200,210,234]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Hoplandriini">HOPLANDRIINI OF THE WORLD</taxonomicName>
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<emphasis id="85FA4B055048FFFFC1E6FF57FF40FEF5" bold="true" box="[173,192,252,271]" pageId="5" pageNumber="88">1.</emphasis>
Mesocoxal cavities broadly separated by broad meso- and metasternal processes (
<figureCitation id="2FB58B925048FFFFC577FF57FB09FEF5" box="[1084,1161,252,271]" captionStart="Figure 3" captionStartId="10.[142,220,1355,1374]" captionTargetBox="[142,1420,198,1318]" captionTargetId="figure-275@10.[142,1421,196,1322]" captionTargetPageId="10" captionText="Figure 3. Hoplandria (Arrhenandria) laeviventris Casey. (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Aedeagus, lateral view. (D) Paramere, lateral view. (E) Spermatheca. (F) Left elytron, male (arrow to denticle)." figureDoi="http://doi.org/10.5281/zenodo.5437798" httpUri="https://zenodo.org/record/5437798/files/figure.png" pageId="5" pageNumber="88">Fig. 3A</figureCitation>
) ..................................... 2
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<paragraph id="B73197175048FFFFC1E6FEBEFA0DFEB8" blockId="5.[173,1421,252,1869]" pageId="5" pageNumber="88">
<keyLead id="0C7F34225048FFFFC1E6FEBEFA0DFEB8" pageId="5" pageNumber="88">
<emphasis id="85FA4B055048FFFFC1E6FEBEFF45FED2" bold="true" box="[173,197,277,296]" pageId="5" pageNumber="88">1¢.</emphasis>
Mesocoxal cavities continuous, only partially separated by meso- and metasternal processes; subtribe
<taxonomicName id="708EEC945048FFFFC070FE85FE41FEBB" authorityName="HANLEY" authorityYear="2002" box="[315,449,302,321]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="88" phylum="Arthropoda" rank="subTribe" subTribe="Platandriina">Platandriina</taxonomicName>
(part) (
<figureCitation id="2FB58B925048FFFFC358FE85FDF2FEBB" box="[531,626,302,321]" captionStart="Figure 16" captionStartId="30.[142,220,1773,1792]" captionTargetBox="[146,1423,222,1737]" captionTargetId="figure-50@30.[129,1433,194,1756]" captionTargetPageId="30" captionText="Figure 16. Platandria mormonica Casey. (A) Labrum. (B) Epipharynx. (C) Mandible, dorsal view. (D) Mandible, ventral view. (E) Maxilla. (F) Labium. (G) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437826" httpUri="https://zenodo.org/record/5437826/files/figure.png" pageId="5" pageNumber="88">Figs 16A</figureCitation>
,
<figureCitation id="2FB58B925048FFFFC335FE84FD28FEB8" box="[638,680,303,322]" captionStart="Figure 21" captionStartId="40.[142,220,1652,1671]" captionTargetBox="[160,1422,207,1592]" captionTargetId="figure-136@40.[142,1448,194,1619]" captionTargetPageId="40" captionText="Figure 21. Microlia meticola (Casey). (A) Labrum. (B) Epipharynx. (C) Mandible, dorsal view. (D) Mandible, ventral view. (E) Maxilla. (F) Labium. (G) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437836" httpUri="https://zenodo.org/record/5437836/files/figure.png" pageId="5" pageNumber="88">21A</figureCitation>
) ................................................................................................................. 4
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<emphasis id="85FA4B055048FFFFC1E6FEE3FF40FEA1" bold="true" box="[173,192,328,347]" pageId="5" pageNumber="88">2.</emphasis>
Apex of galea of maxilla with rows of long to very long hairs, typically giving appearance of flowing hairs; subtribe
<taxonomicName id="708EEC945048FFFFC099FECAFDD8FE8E" authorityName="HANLEY" authorityYear="2002" box="[466,600,353,372]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="88" phylum="Arthropoda" rank="subTribe" subTribe="Platandriina">Platandriina</taxonomicName>
(part) (
<figureCitation id="2FB58B925048FFFFC3E1FECAFC81FE8E" box="[682,769,353,372]" captionStart="Figure 19" captionStartId="37.[155,233,1640,1659]" captionTargetBox="[183,1411,211,1606]" captionTargetId="figure-126@37.[176,1412,196,1608]" captionTargetPageId="37" captionText="Figure 19. Ligulata hansoni Hanley &amp; Ashe. (A) Habitus, dorsal view. (B) Labrum. (C) Epipharynx. (D) Mandible, dorsal view. (E) Mandible, ventral view. (F) Maxilla. (G) Labium. (H) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437832" httpUri="https://zenodo.org/record/5437832/files/figure.png" pageId="5" pageNumber="88">Fig. 19F</figureCitation>
) ................................................................................................... 3
</keyLead>
</paragraph>
<paragraph id="B73197175048FFFFC1E6FED1FA0DFE5D" blockId="5.[173,1421,252,1869]" pageId="5" pageNumber="88">
<keyLead id="0C7F34225048FFFFC1E6FED1FA0DFE5D" pageId="5" pageNumber="88">
<emphasis id="85FA4B055048FFFFC1E6FED1FF45FE74" bold="true" box="[173,197,378,398]" pageId="5" pageNumber="88">2¢.</emphasis>
Apex of galea of maxilla with rows of short to very short hairs, giving appearance of stubble (
<figureCitation id="2FB58B925048FFFFC1AFFE3FFEADFE5D" box="[228,301,404,423]" captionStart="Figure 6" captionStartId="15.[163,241,1761,1780]" captionTargetBox="[158,1440,200,1725]" captionTargetId="figure-61@15.[154,1449,191,1734]" captionTargetPageId="15" captionText="Figure 6. Hoplandria (Hoplandria) lateralis Melsheimer. (A) Habitus, dorsal view. (B) Labrum. (C) Epipharynx. (D) Mandible, dorsal view. (E) Mandible, ventral view. (F) Maxilla. (G) Labium. (H) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437804" httpUri="https://zenodo.org/record/5437804/files/figure.png" pageId="5" pageNumber="88">Fig. 6F</figureCitation>
) ............................................................................................................................................................................. 7
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</keyStep>
<keyStep id="0C7A8FB25048FFFFC1E6FE06FA0CFD8B" pageId="5" pageNumber="88">
<paragraph id="B73197175048FFFFC1E6FE06FA0CFDF6" blockId="5.[173,1421,252,1869]" pageId="5" pageNumber="88">
<keyLead id="0C7F34225048FFFFC1E6FE06FA0CFDF6" pageId="5" pageNumber="88">
<emphasis id="85FA4B055048FFFFC1E6FE06FF40FE3A" bold="true" box="[173,192,429,448]" pageId="5" pageNumber="88">3.</emphasis>
Mesosternum with a distinct medial carina, not reaching apex of mesosternal process (
<figureCitation id="2FB58B925048FFFFC532FE06FB53FE3A" box="[1145,1235,429,448]" captionStart="Figure 25" captionStartId="46.[142,220,1201,1220]" captionTargetBox="[162,1403,194,1160]" captionTargetId="figure-327@46.[159,1405,188,1175]" captionTargetPageId="46" captionText="Figure 25. Paroplandria typica Cameron. (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Aedeagus (with air bubble near middle), lateral view. (D) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5437844" httpUri="https://zenodo.org/record/5437844/files/figure.png" pageId="5" pageNumber="88">Fig. 25A</figureCitation>
); apices of abdominal terga each with 2–4 very large, black, heavy setae (
<figureCitation id="2FB58B925048FFFFC291FE6DFBB3FE23" box="[986,1075,454,473]" captionStart="Figure 25" captionStartId="46.[142,220,1201,1220]" captionTargetBox="[162,1403,194,1160]" captionTargetId="figure-327@46.[159,1405,188,1175]" captionTargetPageId="46" captionText="Figure 25. Paroplandria typica Cameron. (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Aedeagus (with air bubble near middle), lateral view. (D) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5437844" httpUri="https://zenodo.org/record/5437844/files/figure.png" pageId="5" pageNumber="88">Fig. 25B</figureCitation>
); abdominal tergum IX without complete crescent-shaped setal pattern; males with abdominal tergum VII with two narrow denticles along midline; known only from the Oriental Region (mainland of
<collectingCountry id="CF99D7875048FFFFC51FFE52FB36FDF6" box="[1108,1206,505,524]" name="Malaysia" pageId="5" pageNumber="88">Malaysia</collectingCountry>
) ..........
<taxonomicName id="708EEC945048FFFFC44BFE52FA0CFDF6" authorityName="CAMERON" authorityYear="1936" box="[1280,1420,505,524]" class="Insecta" family="Staphylinidae" genus="Paroplandria" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="88" phylum="Arthropoda" rank="genus">
<emphasis id="85FA4B055048FFFFC44BFE52FA0CFDF6" box="[1280,1420,505,524]" italics="true" pageId="5" pageNumber="88">Paroplandria</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B73197175048FFFFC1E6FDB9FA0CFD8B" blockId="5.[173,1421,252,1869]" pageId="5" pageNumber="88">
<keyLead id="0C7F34225048FFFFC1E6FDB9FA0CFD8B" pageId="5" pageNumber="88">
<emphasis id="85FA4B055048FFFFC1E6FDB9FF45FDDC" bold="true" box="[173,197,530,550]" pageId="5" pageNumber="88">3¢.</emphasis>
Mesosternum without a medial carina (
<figureCitation id="2FB58B925048FFFFC3CAFDB9FD5CFDDC" box="[641,732,530,550]" captionStart="Figure 20" captionStartId="38.[142,220,1403,1422]" captionTargetBox="[148,1420,206,1370]" captionTargetId="figure-216@38.[135,1437,194,1382]" captionTargetPageId="38" captionText="Figure 20. Ligulata hansoni Hanley &amp; Ashe. (A) Meso- and metathorax. (B) Abdominal tergum IX. (C) Aedeagus, lateral view. (D) Paramere, lateral view. (E) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5437834" httpUri="https://zenodo.org/record/5437834/files/figure.png" pageId="5" pageNumber="88">Fig. 20A</figureCitation>
); apices of abdominal terga with typical macrosetae, not large, black, heavy setae; abdominal tergum IX with distinct crescent-shaped setal pattern (
<figureCitation id="2FB58B925048FFFFC03FFDEEFE4DFDA2" box="[372,461,581,600]" captionStart="Figure 20" captionStartId="38.[142,220,1403,1422]" captionTargetBox="[148,1420,206,1370]" captionTargetId="figure-216@38.[135,1437,194,1382]" captionTargetPageId="38" captionText="Figure 20. Ligulata hansoni Hanley &amp; Ashe. (A) Meso- and metathorax. (B) Abdominal tergum IX. (C) Aedeagus, lateral view. (D) Paramere, lateral view. (E) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5437834" httpUri="https://zenodo.org/record/5437834/files/figure.png" pageId="5" pageNumber="88">Fig. 20B</figureCitation>
); males without secondary sexual features; known only from the Neotropical Region (
<collectingCountry id="CF99D7875048FFFFC0FFFDF4FDA7FD8B" box="[436,551,606,626]" name="Costa Rica" pageId="5" pageNumber="88">Costa Rica</collectingCountry>
)..........................................................................................................................
<taxonomicName id="708EEC945048FFFFC479FDF5FA0CFD8B" baseAuthorityName="Hanley &amp; Ashe" baseAuthorityYear="1998" box="[1330,1420,606,625]" class="Insecta" family="Staphylinidae" genus="Ligulata" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="88" phylum="Arthropoda" rank="genus">
<emphasis id="85FA4B055048FFFFC479FDF5FA0CFD8B" box="[1330,1420,606,625]" italics="true" pageId="5" pageNumber="88">Ligulata</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="0C7A8FB25048FFFFC1E6FDD3FA0CFCC6" pageId="5" pageNumber="88">
<paragraph id="B73197175048FFFFC1E6FDD3FA0DFD5E" blockId="5.[173,1421,252,1869]" pageId="5" pageNumber="88">
<keyLead id="0C7F34225048FFFFC1E6FDD3FA0DFD5E" pageId="5" pageNumber="88">
<emphasis id="85FA4B055048FFFFC1E6FDD3FF40FD71" bold="true" box="[173,192,632,651]" pageId="5" pageNumber="88">4.</emphasis>
Mesosternum without a distinct medial ridge (
<figureCitation id="2FB58B925048FFFFC387FDD3FCA7FD71" box="[716,807,632,651]" captionStart="Figure 22" captionStartId="41.[155,233,1630,1649]" captionTargetBox="[160,1434,201,1581]" captionTargetId="figure-156@41.[136,1452,194,1597]" captionTargetPageId="41" captionText="Figure 22. Microlia meticola (Casey). (A) Meso- and metathorax. (B) Abdominal tergum IX. (C) Aedeagus, external, lateral view. (D) Aedeagus, internal, lateral view. (E) Paramere, lateral view. (F) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5437838" httpUri="https://zenodo.org/record/5437838/files/figure.png" pageId="5" pageNumber="88">Fig. 22A</figureCitation>
); each labial palpus with three distinct articles; male secondary features not as below .............................................................................................................. 5
</keyLead>
</paragraph>
<paragraph id="B73197175048FFFFC1E6FD01FA0CFCC6" blockId="5.[173,1421,252,1869]" pageId="5" pageNumber="88">
<keyLead id="0C7F34225048FFFFC1E6FD01FA0CFCC6" pageId="5" pageNumber="88">
<emphasis id="85FA4B055048FFFFC1E6FD01FF45FD44" bold="true" box="[173,197,682,702]" pageId="5" pageNumber="88">4¢.</emphasis>
Mesosternum with a distinct, heavy, medial ridge extending to apex of mesosternal process, ridge with numerous, small hairs (
<figureCitation id="2FB58B925048FFFFC35BFD6FFDE9FD2D" box="[528,617,708,727]" captionStart="Figure 18" captionStartId="35.[155,233,1144,1163]" captionTargetBox="[155,1431,196,1111]" captionTargetId="figure-402@35.[154,1431,196,1111]" captionTargetPageId="35" captionText="Figure 18. Ditropandria (Pedandria) ambrensis Pace. (A) Mandible, ventral view. (B) Antenna. (C) Pronotum. (D) Left elytron. (E) Meso- and metathoracic region. (F) Abdominal tergum VII (male) (arrows to denticles). (G) Abdominal tergum IX. (H) Aedeagus, lateral view. (I) Paramere, lateral view. (J) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5437830" httpUri="https://zenodo.org/record/5437830/files/figure.png" pageId="5" pageNumber="88">Fig. 18E</figureCitation>
); each labial palpus with articles 1 + 2 fused, only two recognizable segments present (males with secondary sexual features of abdominal tergum VII with four small, narrow denticles along midline and tergum
<collectionCode id="D19F0FD25048FFFFC22BFD5CFC0CFCF0" box="[864,908,759,778]" pageId="5" pageNumber="88">VIII</collectionCode>
with distinct longitudinal row of setiferous denticles) (
<figureCitation id="2FB58B925048FFFFC091FCBBFDB1FCD9" box="[474,561,784,803]" captionStart="Figure 18" captionStartId="35.[155,233,1144,1163]" captionTargetBox="[155,1431,196,1111]" captionTargetId="figure-402@35.[154,1431,196,1111]" captionTargetPageId="35" captionText="Figure 18. Ditropandria (Pedandria) ambrensis Pace. (A) Mandible, ventral view. (B) Antenna. (C) Pronotum. (D) Left elytron. (E) Meso- and metathoracic region. (F) Abdominal tergum VII (male) (arrows to denticles). (G) Abdominal tergum IX. (H) Aedeagus, lateral view. (I) Paramere, lateral view. (J) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5437830" httpUri="https://zenodo.org/record/5437830/files/figure.png" pageId="5" pageNumber="88">Fig. 18F</figureCitation>
); known only from Afrotropical Region (
<collectingCountry id="CF99D7875048FFFFC290FCBBFBDCFCD9" box="[987,1116,784,803]" name="Madagascar" pageId="5" pageNumber="88">Madagascar</collectingCountry>
)
<taxonomicName id="708EEC945048FFFFC196FC82FEE8FCC6" authorityName="PACE" authorityYear="1984" box="[221,360,809,828]" class="Insecta" family="Staphylinidae" genus="Ditropandria" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="88" phylum="Arthropoda" rank="genus">
<emphasis id="85FA4B055048FFFFC196FC82FEE8FCC6" box="[221,360,809,828]" italics="true" pageId="5" pageNumber="88">Ditropandria</emphasis>
</taxonomicName>
subgenus .....................................................................................................................................
<taxonomicName id="708EEC945048FFFFC454FC82FA0CFCC6" authorityName="Pace" authorityYear="1991" box="[1311,1420,809,828]" class="Insecta" family="Staphylinidae" genus="Ditropandria" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="119" phylum="Arthropoda" rank="subGenus" subGenus="Pedandria">
<emphasis id="85FA4B055048FFFFC454FC82FA0CFCC6" box="[1311,1420,809,828]" italics="true" pageId="5" pageNumber="88">Pedandria</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="0C7A8FB25048FFFFC1E6FCE8FA0DFC2E" pageId="5" pageNumber="88">
<paragraph id="B73197175048FFFFC1E6FCE8FA0DFC72" blockId="5.[173,1421,252,1869]" pageId="5" pageNumber="88">
<keyLead id="0C7F34225048FFFFC1E6FCE8FA0DFC72" pageId="5" pageNumber="88">
<emphasis id="85FA4B055048FFFFC1E6FCE8FF40FCAC" bold="true" box="[173,192,835,854]" pageId="5" pageNumber="88">5.</emphasis>
Males with secondary sexual features somewhat variable, but without a medial, longitudinal carina on either abdominal terga VII or
<collectionCode id="D19F0FD25048FFFFC3C1FCF7FD38FC95" box="[650,696,860,879]" pageId="5" pageNumber="88">VIII</collectionCode>
; body very small, length
<quantity id="70763AF25048FFFFC28FFCF7FBC0FC95" box="[964,1088,860,879]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.3" metricValueMin="1.2" pageId="5" pageNumber="88" unit="mm" value="1.75" valueMax="2.3" valueMin="1.2">1.2–2.3 mm</quantity>
; eyes average, 0.4–0.6 times length of head............................................................................................................................................ 6
</keyLead>
</paragraph>
<paragraph id="B73197175048FFFFC1E6FC24FA0DFC2E" blockId="5.[173,1421,252,1869]" pageId="5" pageNumber="88">
<keyLead id="0C7F34225048FFFFC1E6FC24FA0DFC2E" pageId="5" pageNumber="88">
<emphasis id="85FA4B055048FFFFC1E6FC24FF45FC58" bold="true" box="[173,197,911,930]" pageId="5" pageNumber="88">5¢.</emphasis>
Males with abdominal tergum VII with narrow median longitudinal carina and tergum
<collectionCode id="D19F0FD25048FFFFC5C8FC24FB2FFC58" box="[1155,1199,911,930]" pageId="5" pageNumber="88">VIII</collectionCode>
with more or less wide median longitudinal carina; body size average, length
<quantity id="70763AF25048FFFFC2E8FC03FB9FFC41" box="[931,1055,936,955]" metricMagnitude="-3" metricUnit="m" metricValue="3.3499999999999996" metricValueMax="4.3" metricValueMin="2.4" pageId="5" pageNumber="88" unit="mm" value="3.3499999999999996" valueMax="4.3" valueMin="2.4">2.4–4.3 mm</quantity>
; eyes small, 0.2–0.4 times length of head; known from Nearctic Region; frequently found in flowers of various shrubs ........
<taxonomicName id="708EEC945048FFFFC451FC6AFA0DFC2E" authorityName="Casey" authorityYear="1893" box="[1306,1421,961,980]" class="Insecta" family="Staphylinidae" genus="Platandria" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="88" phylum="Arthropoda" rank="genus">
<emphasis id="85FA4B055048FFFFC451FC6AFA0DFC2E" box="[1306,1421,961,980]" italics="true" pageId="5" pageNumber="88">Platandria</emphasis>
</taxonomicName>
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</paragraph>
</keyStep>
<keyStep id="0C7A8FB25048FFFFC1E6FC71FA0DFB65" pageId="5" pageNumber="88">
<paragraph id="B73197175048FFFFC1E6FC71FA0CFBC3" blockId="5.[173,1421,252,1869]" pageId="5" pageNumber="88">
<keyLead id="0C7F34225048FFFFC1E6FC71FA0CFBC3" pageId="5" pageNumber="88">
<emphasis id="85FA4B055048FFFFC1E6FC71FF40FC17" bold="true" box="[173,192,986,1005]" pageId="5" pageNumber="88">6.</emphasis>
Abdominal tergum IX without complete crescent-shaped setal pattern, setae absent basally along midline; eyes average, 0.5 times length of head; males without secondary sexual features; known from Nearctic Region (Arizona, North Carolina, Texas, Virginia, northern
<collectingCountry id="CF99D7875048FFFFC2A6FBA6FBBAFBDA" box="[1005,1082,1037,1056]" name="Mexico" pageId="5" pageNumber="88">Mexico</collectingCountry>
); likely found in association with flowers .......................................................................................................................................
<taxonomicName id="708EEC945048FFFFC47FFB8DFA0CFBC3" authorityName="Casey" authorityYear="1910" box="[1332,1420,1062,1081]" class="Insecta" family="Staphylinidae" genus="Microlia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="88" phylum="Arthropoda" rank="genus">
<emphasis id="85FA4B055048FFFFC47FFB8DFA0CFBC3" box="[1332,1420,1062,1081]" italics="true" pageId="5" pageNumber="88">Microlia</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B73197175048FFFFC1E6FBEBFA0DFB65" blockId="5.[173,1421,252,1869]" pageId="5" pageNumber="88">
<keyLead id="0C7F34225048FFFFC1E6FBEBFA0DFB65" pageId="5" pageNumber="88">
<emphasis id="85FA4B055048FFFFC1E6FBEBFF45FBA9" bold="true" box="[173,197,1088,1107]" pageId="5" pageNumber="88">6¢.</emphasis>
Abdominal tergum IX with a distinct crescent-shaped setal pattern; eyes average to large, 0.5–0.7 times length of head; males tergum IV with two small denticles on each side of midline (
<figureCitation id="2FB58B925048FFFFC5C6FBF2FB64FB96" box="[1165,1252,1113,1132]" captionStart="Figure 23" captionStartId="43.[155,233,1232,1251]" captionTargetBox="[156,1432,195,1200]" captionTargetId="figure-294@43.[156,1433,195,1200]" captionTargetPageId="43" captionText="Figure 23. Omoplandria fuscipennis Cameron. (A) Apical antennal segment featuring coeloconical sensilla. (B) Mandible, dorsal view. (C) Mandible, ventral view. (D) Maxilla. (E) Meso- and metathoracic region. (F) Abdominal tergum IV (male). (G) Abdominal tergum VIII. (H) Paramere, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5437840" httpUri="https://zenodo.org/record/5437840/files/figure.png" pageId="5" pageNumber="88">Fig. 23F</figureCitation>
) and terga VII–
<collectionCode id="D19F0FD25048FFFFC031FBD8FE25FB7C" box="[378,421,1139,1158]" pageId="5" pageNumber="88">VIII</collectionCode>
with prominent small, narrow setiferous denticles (
<figureCitation id="2FB58B925048FFFFC284FBD9FBA9FB7C" box="[975,1065,1138,1158]" captionStart="Figure 23" captionStartId="43.[155,233,1232,1251]" captionTargetBox="[156,1432,195,1200]" captionTargetId="figure-294@43.[156,1433,195,1200]" captionTargetPageId="43" captionText="Figure 23. Omoplandria fuscipennis Cameron. (A) Apical antennal segment featuring coeloconical sensilla. (B) Mandible, dorsal view. (C) Mandible, ventral view. (D) Maxilla. (E) Meso- and metathoracic region. (F) Abdominal tergum IV (male). (G) Abdominal tergum VIII. (H) Paramere, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5437840" httpUri="https://zenodo.org/record/5437840/files/figure.png" pageId="5" pageNumber="88">Fig. 23G</figureCitation>
); known from the Oriental Region (Taiwan) and the Palaearctic Region (
<collectingCountry id="CF99D7875048FFFFC256FB27FCDEFB65" box="[797,862,1164,1183]" name="Japan" pageId="5" pageNumber="88">Japan</collectingCountry>
) ................................................................
<taxonomicName id="708EEC945048FFFFC44BFB27FA0DFB65" authorityName="CAMERON" authorityYear="1949" box="[1280,1421,1164,1183]" class="Insecta" family="Staphylinidae" genus="Omoplandria" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="88" phylum="Arthropoda" rank="genus">
<emphasis id="85FA4B055048FFFFC44BFB27FA0DFB65" box="[1280,1421,1164,1183]" italics="true" pageId="5" pageNumber="88">Omoplandria</emphasis>
</taxonomicName>
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</paragraph>
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<keyStep id="0C7A8FB25048FFFFC1E6FB0EFA0DFAAA" pageId="5" pageNumber="88">
<paragraph id="B73197175048FFFFC1E6FB0EFA0CFAFE" blockId="5.[173,1421,252,1869]" pageId="5" pageNumber="88">
<keyLead id="0C7F34225048FFFFC1E6FB0EFA0CFAFE" pageId="5" pageNumber="88">
<emphasis id="85FA4B055048FFFFC1E6FB0EFF40FB42" bold="true" box="[173,192,1189,1208]" pageId="5" pageNumber="88">7.</emphasis>
Mesosternum with median longitudinal carina extending to apex of mesosternal process; tarsal formula 5-5-5; males with secondary sexual features, including sinuate areas of apico-lateral angles of elytra often produced into heavy carina-like structures; known from Oriental Region; subtribe
<taxonomicName id="708EEC945048FFFFC070FB5AFE74FAFE" authorityName="HANLEY" authorityYear="2002" box="[315,500,1265,1284]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="88" phylum="Arthropoda" rank="subTribe" subTribe="Pseudoplandriina">Pseudoplandriina</taxonomicName>
.......................................................................................................................
<taxonomicName id="708EEC945048FFFFC5ADFB5AFA0CFAFE" authorityName="Fenyes" authorityYear="1921" box="[1254,1420,1265,1284]" class="Insecta" family="Staphylinidae" genus="Pseudoplandria" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="88" phylum="Arthropoda" rank="genus">
<emphasis id="85FA4B055048FFFFC5ADFB5AFA0CFAFE" box="[1254,1420,1265,1284]" italics="true" pageId="5" pageNumber="88">Pseudoplandria</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B73197175048FFFFC1E6FAA0FA0DFAAA" blockId="5.[173,1421,252,1869]" pageId="5" pageNumber="88">
<keyLead id="0C7F34225048FFFFC1E6FAA0FA0DFAAA" pageId="5" pageNumber="88">
<emphasis id="85FA4B055048FFFFC1E6FAA0FF45FAE4" bold="true" box="[173,197,1291,1310]" pageId="5" pageNumber="88">7¢.</emphasis>
Mesosternum without median longitudinal carina; tarsal formula 4-5-5 (or 4-4-4 rarely); males without sinuate areas of apico-lateral angles of elytra produced into heavy carina-like structures; subtribe
<taxonomicName id="708EEC945048FFFFC0FFFA96FDC1FAAA" authorityName="CASEY" authorityYear="1910" box="[436,577,1341,1360]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="88" phylum="Arthropoda" rank="subTribe" subTribe="Hoplandriina">Hoplandriina</taxonomicName>
................................................................................................................................... 8
</keyLead>
</paragraph>
</keyStep>
<keyStep id="0C7A8FB25048FFFFC1E6FAFDFA0DF9CE" pageId="5" pageNumber="88">
<paragraph id="B73197175048FFFFC1E6FAFDFA0DFA12" blockId="5.[173,1421,252,1869]" pageId="5" pageNumber="88">
<keyLead id="0C7F34225048FFFFC1E6FAFDFA0DFA12" pageId="5" pageNumber="88">
<emphasis id="85FA4B055048FFFFC1E6FAFDFF40FA93" bold="true" box="[173,192,1366,1385]" pageId="5" pageNumber="88">8.</emphasis>
Pronotum with outer apical angles not sinuate, apical area along midline not extended posteriorly; males with secondary sexual features highly variable, including various combinations of elytra with humeral area with carina or bump, medioapical region with carina or conical process, outer apical third with denticle, carina or bump; abdomen with posterior corners of sterna
<collectionCode id="D19F0FD25048FFFFC196FA17FF79FA35" box="[221,249,1468,1487]" pageId="5" pageNumber="88">III</collectionCode>
and IV produced into spines, tergum IV with raised area or carina, and tergum VII with a narrow posteriomedian longitudinal carina; known from throughout the New World
<emphasis id="85FA4B055048FFFFC525FA7DFA0DFA12" box="[1134,1421,1493,1512]" italics="true" pageId="5" pageNumber="88">
....................
<taxonomicName id="708EEC945048FFFFC5A5FA7EFAE8FA12" authorityName="Kraatz" authorityYear="1857" box="[1262,1384,1493,1512]" class="Insecta" family="Staphylinidae" genus="Hoplandria" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="88" phylum="Arthropoda" rank="genus" sensu="lato">Hoplandria</taxonomicName>
<taxonomicNameLabel id="9EC9F67E5048FFFFC424FA7EFA0DFA12" box="[1391,1421,1493,1512]" pageId="5" pageNumber="88" sensu="lato">s.l.</taxonomicNameLabel>
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</keyLead>
</paragraph>
<paragraph id="B73197175048FFFFC1E6FA45FA0DF9CE" blockId="5.[173,1421,252,1869]" pageId="5" pageNumber="88">
<keyLead id="0C7F34225048FFFFC1E6FA45FA0DF9CE" pageId="5" pageNumber="88">
<emphasis id="85FA4B055048FFFFC1E6FA45FF45F9F8" bold="true" box="[173,197,1518,1538]" pageId="5" pageNumber="88">8¢.</emphasis>
Pronotum with outer apical angles sinuate, apical area along midline extended posteriorly; males with variable secondary sexual features, but without corners of abdominal sternum IV produced into spines and without modifications to tergum IV..................................................................................... 9
</keyLead>
</paragraph>
</keyStep>
<keyStep id="0C7A8FB25048FFFFC1E6F991FA0DF8B6" pageId="5" pageNumber="88">
<paragraph id="B73197175048FFFFC1E6F991FA0CF936" blockId="5.[173,1421,252,1869]" pageId="5" pageNumber="88">
<keyLead id="0C7F34225048FFFFC1E6F991FA0CF936" pageId="5" pageNumber="88">
<emphasis id="85FA4B055048FFFFC1E6F991FF40F9B7" bold="true" box="[173,192,1594,1613]" pageId="5" pageNumber="88">9.</emphasis>
Extended area along midline apex of pronotum with rounded angles (
<figureCitation id="2FB58B925048FFFFC28BF991FB9CF9B4" box="[960,1052,1594,1614]" captionStart="Figure 12" captionStartId="24.[142,220,1251,1270]" captionTargetBox="[143,1421,194,1218]" captionTargetId="figure-324@24.[143,1421,194,1218]" captionTargetPageId="24" captionText="Figure 12. Leptandria ecitophila Hanley. (A) Antenna. (B) Labrum. (C) Mandible, dorsal view. (D) Mandible, ventral view. (E) Maxilla. (F) Labium. (G) Hypopharynx. (H) Pronotum. (I) Right elytron. (J) Tarsus, hind leg. (K) Aedeagus, lateral view. (L) Paramere, lateral view. (M) Spermatheca. (N) Abdominal tergum VII, male (arrow to conical carina). (O) Male abdominal sternum VII." figureDoi="http://doi.org/10.5281/zenodo.5437818" httpUri="https://zenodo.org/record/5437818/files/figure.png" pageId="5" pageNumber="88">Fig. 12H</figureCitation>
); medial setae of prementum with insertions widely separated (
<figureCitation id="2FB58B925048FFFFC382F9FFFCA1F99D" box="[713,801,1620,1639]" captionStart="Figure 12" captionStartId="24.[142,220,1251,1270]" captionTargetBox="[143,1421,194,1218]" captionTargetId="figure-324@24.[143,1421,194,1218]" captionTargetPageId="24" captionText="Figure 12. Leptandria ecitophila Hanley. (A) Antenna. (B) Labrum. (C) Mandible, dorsal view. (D) Mandible, ventral view. (E) Maxilla. (F) Labium. (G) Hypopharynx. (H) Pronotum. (I) Right elytron. (J) Tarsus, hind leg. (K) Aedeagus, lateral view. (L) Paramere, lateral view. (M) Spermatheca. (N) Abdominal tergum VII, male (arrow to conical carina). (O) Male abdominal sternum VII." figureDoi="http://doi.org/10.5281/zenodo.5437818" httpUri="https://zenodo.org/record/5437818/files/figure.png" pageId="5" pageNumber="88">Fig. 12F</figureCitation>
); males with secondary sexual features exclusively limited to each elytron, with small denticle on medioapical area (
<figureCitation id="2FB58B925048FFFFC546F9C6FBDEF97A" box="[1037,1118,1645,1664]" captionStart="Figure 12" captionStartId="24.[142,220,1251,1270]" captionTargetBox="[143,1421,194,1218]" captionTargetId="figure-324@24.[143,1421,194,1218]" captionTargetPageId="24" captionText="Figure 12. Leptandria ecitophila Hanley. (A) Antenna. (B) Labrum. (C) Mandible, dorsal view. (D) Mandible, ventral view. (E) Maxilla. (F) Labium. (G) Hypopharynx. (H) Pronotum. (I) Right elytron. (J) Tarsus, hind leg. (K) Aedeagus, lateral view. (L) Paramere, lateral view. (M) Spermatheca. (N) Abdominal tergum VII, male (arrow to conical carina). (O) Male abdominal sternum VII." figureDoi="http://doi.org/10.5281/zenodo.5437818" httpUri="https://zenodo.org/record/5437818/files/figure.png" pageId="5" pageNumber="88">Fig. 12I</figureCitation>
), and abdominal tergum VII with narrow posteromedian longitudinal carina or conical carina (
<figureCitation id="2FB58B925048FFFFC1AFF90BFEBFF949" box="[228,319,1696,1715]" captionStart="Figure 12" captionStartId="24.[142,220,1251,1270]" captionTargetBox="[143,1421,194,1218]" captionTargetId="figure-324@24.[143,1421,194,1218]" captionTargetPageId="24" captionText="Figure 12. Leptandria ecitophila Hanley. (A) Antenna. (B) Labrum. (C) Mandible, dorsal view. (D) Mandible, ventral view. (E) Maxilla. (F) Labium. (G) Hypopharynx. (H) Pronotum. (I) Right elytron. (J) Tarsus, hind leg. (K) Aedeagus, lateral view. (L) Paramere, lateral view. (M) Spermatheca. (N) Abdominal tergum VII, male (arrow to conical carina). (O) Male abdominal sternum VII." figureDoi="http://doi.org/10.5281/zenodo.5437818" httpUri="https://zenodo.org/record/5437818/files/figure.png" pageId="5" pageNumber="88">Fig. 12N</figureCitation>
); elytra without modifications of humeral areas; spines absent from abdominal sterna
<collectionCode id="D19F0FD25048FFFFC196F911FF79F937" box="[221,249,1722,1741]" pageId="5" pageNumber="88">III</collectionCode>
and IV; tergum IV without modification; known from Neotropical Region (
<collectingCountry id="CF99D7875048FFFFC544F912FBE8F936" box="[1039,1128,1721,1740]" name="Ecuador" pageId="5" pageNumber="88">Ecuador</collectingCountry>
)
<emphasis id="85FA4B055048FFFFC53FF911FA0CF936" box="[1140,1420,1721,1740]" italics="true" pageId="5" pageNumber="88">
.........................
<taxonomicName id="708EEC945048FFFFC45CF912FA0CF936" authorityName="HANLEY, IN" authorityYear="2003" box="[1303,1420,1721,1740]" class="Insecta" family="Staphylinidae" genus="Leptandria" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="88" phylum="Arthropoda" rank="genus">Leptandria</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="B73197175048FFFFC1E6F97FFA0DF8B6" blockId="5.[173,1421,252,1869]" pageId="5" pageNumber="88">
<keyLead id="0C7F34225048FFFFC1E6F97FFA0DF8B6" pageId="5" pageNumber="88">
<emphasis id="85FA4B055048FFFFC1E6F97FFF45F91D" bold="true" box="[173,197,1748,1767]" pageId="5" pageNumber="88">9¢.</emphasis>
Extended area along midline apex of pronotum with distinctly sharp angles (
<figureCitation id="2FB58B925048FFFFC55BF97FFBEBF91D" box="[1040,1131,1748,1767]" captionStart="Figure 15" captionStartId="28.[142,220,1228,1247]" captionTargetBox="[147,1417,194,1196]" captionTargetId="figure-339@28.[147,1417,194,1196]" captionTargetPageId="28" captionText="Figure 15. Heliconandria peoechma Hanley. (A) Pronotum. (B) Meso- and metathorax. (C) Tarsus, hind leg. (E) Abdominal tergum III, male. Heliconandria gynochela Hanley. (D) Genitalic tract, female." figureDoi="http://doi.org/10.5281/zenodo.5437824" httpUri="https://zenodo.org/record/5437824/files/figure.png" pageId="5" pageNumber="88">Fig. 15A</figureCitation>
); medial setae of prementum with insertions nearly adjacent (
<figureCitation id="2FB58B925048FFFFC259F946FCEDF8FA" box="[786,877,1773,1792]" captionStart="Figure 13" captionStartId="26.[142,220,1813,1832]" captionTargetBox="[163,1395,198,1779]" captionTargetId="figure-55@26.[163,797,196,1145]" captionTargetPageId="26" captionText="Figure 13. Heliconandria wettersteniana Hanley. (A) Habitus, dorsal view. (B) Labrum. (C) Epipharynx. (D) Mandible, dorsal view. (E) Mandible, ventral view. (F) Maxilla. (G) Labium. (H) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437820" httpUri="https://zenodo.org/record/5437820/files/figure.png" pageId="5" pageNumber="88">Fig. 13G</figureCitation>
); males with secondary sexual features variable, including medioapical region of elytra with a short, wide carina (
<figureCitation id="2FB58B925048FFFFC519F8ADFB2BF8E0" box="[1106,1195,1798,1818]" captionStart="Figure 13" captionStartId="26.[142,220,1813,1832]" captionTargetBox="[163,1395,198,1779]" captionTargetId="figure-55@26.[163,797,196,1145]" captionTargetPageId="26" captionText="Figure 13. Heliconandria wettersteniana Hanley. (A) Habitus, dorsal view. (B) Labrum. (C) Epipharynx. (D) Mandible, dorsal view. (E) Mandible, ventral view. (F) Maxilla. (G) Labium. (H) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437820" httpUri="https://zenodo.org/record/5437820/files/figure.png" pageId="5" pageNumber="88">Fig. 13A</figureCitation>
), and corners of abdominal sternum
<collectionCode id="D19F0FD25048FFFFC319F88BFDEEF8C9" box="[594,622,1824,1843]" pageId="5" pageNumber="88">III</collectionCode>
produced into spines (
<figureCitation id="2FB58B925048FFFFC215F88BFC38F8C9" box="[862,952,1824,1843]" captionStart="Figure 15" captionStartId="28.[142,220,1228,1247]" captionTargetBox="[147,1417,194,1196]" captionTargetId="figure-339@28.[147,1417,194,1196]" captionTargetPageId="28" captionText="Figure 15. Heliconandria peoechma Hanley. (A) Pronotum. (B) Meso- and metathorax. (C) Tarsus, hind leg. (E) Abdominal tergum III, male. Heliconandria gynochela Hanley. (D) Genitalic tract, female." figureDoi="http://doi.org/10.5281/zenodo.5437824" httpUri="https://zenodo.org/record/5437824/files/figure.png" pageId="5" pageNumber="88">Fig. 15E</figureCitation>
); known from Neotropical Region (
<collectingCountry id="CF99D7875048FFFFC078F892FE0BF8B6" box="[307,395,1849,1868]" name="Ecuador" pageId="5" pageNumber="88">Ecuador</collectingCountry>
and
<collectingCountry id="CF99D7875048FFFFC089F892FE74F8B6" box="[450,500,1849,1868]" name="Peru" pageId="5" pageNumber="88">Peru</collectingCountry>
)........................................................................................................................
<taxonomicName id="708EEC945048FFFFC5BDF892FA0DF8B6" authorityName="HANLEY" authorityYear="2002" box="[1270,1421,1849,1868]" class="Insecta" family="Staphylinidae" genus="Heliconandria" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="88" phylum="Arthropoda" rank="genus">Heliconandria</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</table>
<paragraph id="B7319717504BFFFCC1C5FF6FFEE4FE68" blockId="6.[142,758,196,402]" pageId="6" pageNumber="89">
welder, 1952:191.
<bibRefCitation id="D31FEAE6504BFFFCC010FF6FFD0CFF20" author="Moore I &amp; Legner EF" box="[347,652,196,218]" pageId="6" pageNumber="89" pagination="1 - 514" refId="ref38699" refString="Moore I, Legner EF. 1975. A catalogue of the Staphylinidae of America north of Mexico (Coleoptera). Special Publication of the Division of Agriculture Science, University of California 3015: 1 - 514." type="journal article" year="1975">Moore &amp; Legner, 1975:438</bibRefCitation>
.
<bibRefCitation id="D31FEAE6504BFFFCC3DCFF6FFF77FF02" author="Seevers CH" pageId="6" pageNumber="89" pagination="1 - 289" refId="ref39301" refString="Seevers CH. 1978. A generic and tribal revision of the North American Aleocharinae (Coleoptera: Staphylinidae). Fieldiana: Zoology 71: 1 - 289." type="journal article" year="1978">Seevers, 1978:141</bibRefCitation>
.
<bibRefCitation id="D31FEAE6504BFFFCC049FF48FD8AFF02" author="Klimaszewski J" box="[258,522,227,249]" pageId="6" pageNumber="89" pagination="1 - 211" refId="ref38245" refString="Klimaszewski J. 1984. A revision of the genus Aleochara Gravenhorst of America north of Mexico (Coleoptera, Staphylinidae, Aleocharinae). Memoirs of the Entomological Society of Canada 129: 1 - 211." type="journal article" year="1984">Klimaszewski, 1984:10</bibRefCitation>
.
<bibRefCitation id="D31FEAE6504BFFFCC35FFF48FEC9FEED" author="Genier F &amp; Klimaszewski J" pageId="6" pageNumber="89" pagination="201 - 216" refId="ref37611" refString="Genier F, Klimaszewski J. 1986. Review of the types of the genus Platandria Casey with a key to the species (Coleoptera: Staphylinidae: Aleocharinae). Coleopterists Bulletin 40: 201 - 216." type="journal article" year="1986">Génier &amp; Klimaszewski, 1986:202</bibRefCitation>
.
<bibRefCitation id="D31FEAE6504BFFFCC028FEAAFD73FEED" author="Klimaszewski J &amp; Peck SB" box="[355,755,257,279]" pageId="6" pageNumber="89" pagination="51 - 113" refId="ref38347" refString="Klimaszewski J, Peck SB. 1986. A review of the cavernicolous Staphylinidae (Coleoptera) of eastern North America: part I. Aleocharinae. Quaestiones Entomologicae 22: 51 - 113." type="journal article" year="1986">Klimaszewski &amp; Peck, 1986:83</bibRefCitation>
.
<bibRefCitation id="D31FEAE6504BFFFCC1C5FE8BFEB5FECF" author="Genier F" box="[142,309,288,310]" pageId="6" pageNumber="89" pagination="3 - 59" refId="ref37573" refString="Genier F. 1989. A revision of the genus Hoplandria Kraatz of America north of Mexico (Coleoptera: Staphylinidae, Aleocharinae). Memoirs of the Entomological Society of Canada 150: 3 - 59." type="journal article" year="1989">Génier, 1989:8</bibRefCitation>
.
<bibRefCitation id="D31FEAE6504BFFFCC00BFE8BFDFCFECC" author="Downie NM &amp; Arnett RH Jr." box="[320,636,288,310]" pageId="6" pageNumber="89" refId="ref37282" refString="Downie NM, Arnett RH Jr. 1996. The beetles of the northeastern North America. Vol. I. Introduction; suborders Archostemata, Adephaga, and Polyphaga, thru superfamily Cantheroidea. Gainesville, Florida: Sandhill Crane Press." type="book" year="1996">Downie &amp; Arnett, 1996:536</bibRefCitation>
.
<bibRefCitation id="D31FEAE6504BFFFCC3CCFE8BFED3FEAE" author="Hanley RS &amp; Ashe JS" pageId="6" pageNumber="89" pagination="183 - 191" refId="ref37925" refString="Hanley RS, Ashe JS. 1998. A new genus and species of aleocharine rove beetle (Coleoptera Staphylinidae Aleocharinae Hoplandriini) from the New World. Tropical Zoology 11: 183 - 191." type="journal article" year="1998">Hanley &amp; Ashe, 1998:184</bibRefCitation>
. Klimaszewski, 2000:49.
<bibRefCitation id="D31FEAE6504BFFFCC3FCFE94FF77FE88" author="Ashe JS" pageId="6" pageNumber="89" pagination="272 - 418" refId="ref36458" refString="Ashe JS. 2001. Aleocharinae, in Newton AF Jr, Thayer MK, Ashe JS, Chandler DS, 22. Staphylinidae Latreille, 1802. In: Thomas MC, Arnett RH Jr, eds. American beetles. Vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. Boca Raton, Florida: CRC Press, 272 - 418." type="book chapter" year="2001">Ashe, 2001:304</bibRefCitation>
.
<bibRefCitation id="D31FEAE6504BFFFCC05AFEF6FE74FE88" author="Hanley RS" box="[273,500,349,371]" pageId="6" pageNumber="89" pagination="301 - 321" refId="ref37747" refString="Hanley RS. 2002 a. Phylogeny and higher classification of Hoplandriini (Coleoptera: Staphylinidae: Aleocharinae). Systematic Entomology 27: 301 - 321." type="journal article" year="2002">Hanley, 2002a:301</bibRefCitation>
.
<bibRefCitation id="D31FEAE6504BFFFCC345FEF6FD73FE89" author="Hanley RS" box="[526,755,349,371]" pageId="6" pageNumber="89" pagination="121 - 132" refId="ref37775" refString="Hanley RS. 2002 b. Heliconandria, a new genus of aleocharine staphylinid beetle from Ecuador (Coleoptera: Staphylinidae). Insect Evolution and Systematics 33: 121 - 132." type="journal article" year="2002">Hanley, 2002b:121</bibRefCitation>
. Hanley, in press b.
</paragraph>
<paragraph id="B7319717504BFFFCC1C5FE06FD02FE05" blockId="6.[142,758,427,511]" pageId="6" pageNumber="89">
<emphasis id="85FA4B05504BFFFCC1C5FE06FE9FFE38" box="[142,287,429,450]" italics="true" pageId="6" pageNumber="89">
<typeStatus id="683529B5504BFFFCC1C5FE06FF43FE38" box="[142,195,429,450]" pageId="6" pageNumber="89">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="708EEC94504BFFFCC067FE07FD47FE38" authority="Kraatz, 1857" authorityName="Kraatz" authorityYear="1857" box="[300,711,428,450]" class="Insecta" family="Staphylinidae" genus="Hoplandria" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="89" phylum="Arthropoda" rank="species" species="ochracea">
<emphasis id="85FA4B05504BFFFCC067FE07FDA2FE3B" box="[300,546,428,449]" italics="true" pageId="6" pageNumber="89">Hoplandria ochracea</emphasis>
<bibRefCitation id="D31FEAE6504BFFFCC365FE06FD47FE38" author="Kraatz G" box="[558,711,428,450]" pageId="6" pageNumber="89" pagination="1 - 43" refId="ref38415" refString="Kraatz G. 1857. Genera aleocharinorum illustrata. Linnaea Entomologica 11: 1 - 43." type="journal article" year="1857">Kraatz, 1857</bibRefCitation>
</taxonomicName>
[=
<taxonomicName id="708EEC94504BFFFCC1C5FE60FDE2FE1B" authority="(Melsheimer, 1844)" baseAuthorityName="Melsheimer" baseAuthorityYear="1844" box="[142,610,458,481]" class="Insecta" family="Staphylinidae" genus="Hoplandria" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="89" phylum="Arthropoda" rank="species" species="lateralis">
<emphasis id="85FA4B05504BFFFCC1C5FE60FEFBFE1A" box="[142,379,459,480]" italics="true" pageId="6" pageNumber="89">Hoplandria lateralis</emphasis>
(
<bibRefCitation id="D31FEAE6504BFFFCC0C7FE60FDDAFE1B" author="Melsheimer FE" box="[396,602,458,481]" pageId="6" pageNumber="89" pagination="26 - 43" refId="ref38668" refString="Melsheimer FE. 1844. Description of new species of Coleoptera of the United States. Proceedings of the Academy of Natural Sciences of Philadelphia 2: 26 - 43." type="journal article" year="1844">Melsheimer, 1844</bibRefCitation>
)
</taxonomicName>
] (designated by
<bibRefCitation id="D31FEAE6504BFFFCC1FAFE41FEB8FE05" author="Casey TL" box="[177,312,490,511]" pageId="6" pageNumber="89" pagination="1 - 183" refId="ref37202" refString="Casey TL. 1910. New species of the staphylinid tribe Myrmedoniini. Memoirs on the Coleoptera 1: 1 - 183." type="journal article" year="1910">Casey, 1910</bibRefCitation>
by subsequent designation).
</paragraph>
</subSubSection>
<subSubSection id="FF94C49C504BFFFCC1C5FDB1FEDEFDB7" pageId="6" pageNumber="89" type="reference_group">
<paragraph id="B7319717504BFFFCC1C5FDB1FEDEFDB7" blockId="6.[142,758,538,590]" pageId="6" pageNumber="89">
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<treatmentCitation id="362FB106504BFFFCC1C5FDB1FE5DFDCA" author="Bierig A" box="[142,477,538,560]" page="26" pageId="6" pageNumber="89" year="1939">
<taxonomicName id="708EEC94504BFFFCC1C5FDB1FE5DFDCA" ID-CoL="8XKWC" authority="Bierig, 1939: 26" authorityName="Bierig" authorityPageNumber="26" authorityYear="1939" box="[142,477,538,560]" class="Insecta" family="Staphylinidae" genus="Diaboligenus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="85FA4B05504BFFFCC1C5FDB1FEA3FDD5" box="[142,291,538,559]" italics="true" pageId="6" pageNumber="89">Diaboligenus</emphasis>
<bibRefCitation id="D31FEAE6504BFFFCC067FDB1FE5DFDCA" author="Bierig A" box="[300,477,538,560]" pageId="6" pageNumber="89" pagination="16 - 31" refId="ref36697" refString="Bierig A. 1939. Neue neotropische Staphylinidae der Subfamilie Aleocharinae. Arbeiten uber Morphologische und Taxonomische Entomologie aus Berlin-Dahlem 6: 16 - 31." type="journal article" year="1939">Bierig, 1939:26</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<treatmentCitation id="362FB106504BFFFCC0A2FDB1FD16FDD5" author="Pace R" box="[489,662,538,559]" page="174" pageId="6" pageNumber="89" year="1990">
<bibRefCitation id="D31FEAE6504BFFFCC0A2FDB1FD16FDD5" author="Pace R" box="[489,662,538,559]" pageId="6" pageNumber="89" pagination="157 - 180" refId="ref38961" refString="Pace R. 1990. Nuovi Falagriini, Hoplandriini ed Aleocharini della Regione Neotropica (Coleoptera, Staphylinidae) (LXXX contributo alla conoscenza delle Aleocharinae). Giornale Italiano di Entomologia 5: 157 - 180." type="journal article" year="1990">Pace, 1990:174</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="362FB106504BFFFCC3E8FDB1FEDBFDB7" author="Hanley RS &amp; Ashe JS" page="184" pageId="6" pageNumber="89" year="1998">
<bibRefCitation id="D31FEAE6504BFFFCC3E8FDB1FEDBFDB7" author="Hanley RS &amp; Ashe JS" pageId="6" pageNumber="89" pagination="183 - 191" refId="ref37925" refString="Hanley RS, Ashe JS. 1998. A new genus and species of aleocharine rove beetle (Coleoptera Staphylinidae Aleocharinae Hoplandriini) from the New World. Tropical Zoology 11: 183 - 191." type="journal article" year="1998">Hanley &amp; Ashe, 1998:184</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="FF94C49C504BFFF2C1C5FDC2FE7AFA02" lastPageId="8" lastPageNumber="91" pageId="6" pageNumber="89" type="materials_examined">
<paragraph id="B7319717504BFFFCC1C5FDC2FE83FD67" blockId="6.[142,758,616,669]" pageId="6" pageNumber="89">
<emphasis id="85FA4B05504BFFFCC1C5FDC2FE9FFD84" box="[142,287,617,638]" italics="true" pageId="6" pageNumber="89">
<typeStatus id="683529B5504BFFFCC1C5FDC2FF43FD84" box="[142,195,617,638]" pageId="6" pageNumber="89">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="708EEC94504BFFFCC067FDC3FD4DFD85" authority="Bierig (1939)" authorityName="Bierig" authorityYear="1939" box="[300,717,616,639]" class="Insecta" family="Staphylinidae" genus="Diaboligenus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="89" phylum="Arthropoda" rank="species" species="primus">
<emphasis id="85FA4B05504BFFFCC067FDC3FDA1FD84" box="[300,545,616,638]" italics="true" pageId="6" pageNumber="89">Diaboligenus primus</emphasis>
<bibRefCitation id="D31FEAE6504BFFFCC37BFDC3FD4DFD85" author="Bierig A" box="[560,717,616,639]" pageId="6" pageNumber="89" pagination="16 - 31" refId="ref36697" refString="Bierig A. 1939. Neue neotropische Staphylinidae der Subfamilie Aleocharinae. Arbeiten uber Morphologische und Taxonomische Entomologie aus Berlin-Dahlem 6: 16 - 31." type="journal article" year="1939">Bierig (1939)</bibRefCitation>
</taxonomicName>
by monotypy.
</paragraph>
<paragraph id="B7319717504BFFFCC1C5FD13FDDCFB42" blockId="6.[142,758,695,1208]" pageId="6" pageNumber="89">
<emphasis id="85FA4B05504BFFFCC1C5FD13FE85FD37" box="[142,261,696,717]" italics="true" pageId="6" pageNumber="89">Diagnosis:</emphasis>
This genus is distinguishable from the other genera of
<taxonomicName id="708EEC94504BFFFCC004FD7DFE68FD16" authorityName="CASEY" authorityYear="1910" box="[335,488,726,748]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="89" phylum="Arthropoda" rank="subTribe" subTribe="Hoplandriina">Hoplandriina</taxonomicName>
by the following combination of characters: body small to very large,
<quantity id="70763AF2504BFFFCC1C5FCB8FEA1FCD3" box="[142,289,787,809]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" metricValueMax="13.5" metricValueMin="1.7" pageId="6" pageNumber="89" unit="mm" value="7.6" valueMax="13.5" valueMin="1.7">1.7–13.5 mm</quantity>
in length; eyes small to large, 0.3–0.7 times length of head; insertion of medial setae of prementum nearly adjacent to widely separated; pronotum with outer apical angles not sinuate, apical area along midline not extended posteriorly, generally distinctly oval in shape; tarsal formula 4-5-5; males with secondary sexual features highly variable, including various combinations of each elytron with humeral area with carina or bump, medioapical region with carina or conical process, outer apical third with denticle, carina or bump; abdomen with posterior corners of sterna
<collectionCode id="D19F0FD2504BFFFCC1B5FBCEFE9CFB80" box="[254,284,1125,1146]" pageId="6" pageNumber="89">III</collectionCode>
and
<collectionCode id="D19F0FD2504BFFFCC012FBCEFEF5FB80" box="[345,373,1125,1146]" pageId="6" pageNumber="89">IV</collectionCode>
produced into spines, tergum
<collectionCode id="D19F0FD2504BFFFCC391FBCEFD76FB80" box="[730,758,1125,1146]" pageId="6" pageNumber="89">IV</collectionCode>
with raised area or carina, and tergum VII with a narrow posteriomedian longitudinal carina.
</paragraph>
<paragraph id="B7319717504BFFFCC1C5FB78FF5EFA65" blockId="6.[142,758,1233,1439]" pageId="6" pageNumber="89">
<emphasis id="85FA4B05504BFFFCC1C5FB78FE96FB12" box="[142,278,1235,1256]" italics="true" pageId="6" pageNumber="89">Description:</emphasis>
[1] Lengths of adults
<quantity id="70763AF2504BFFFCC37BFB79FD47FB1D" box="[560,711,1234,1256]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" metricValueMax="13.5" metricValueMin="1.7" pageId="6" pageNumber="89" unit="mm" value="7.6" valueMax="13.5" valueMin="1.7">1.7–13.5 mm</quantity>
. [2] Body (
<figureCitation id="2FB58B92504BFFFCC192FB5AFEB3FAFD" box="[217,307,1265,1287]" captionStart="Figure 6" captionStartId="15.[163,241,1761,1780]" captionTargetBox="[158,1440,200,1725]" captionTargetId="figure-61@15.[154,1449,191,1734]" captionTargetPageId="15" captionText="Figure 6. Hoplandria (Hoplandria) lateralis Melsheimer. (A) Habitus, dorsal view. (B) Labrum. (C) Epipharynx. (D) Mandible, dorsal view. (E) Mandible, ventral view. (F) Maxilla. (G) Labium. (H) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437804" httpUri="https://zenodo.org/record/5437804/files/figure.png" pageId="6" pageNumber="89">Figs 6A</figureCitation>
,
<figureCitation id="2FB58B92504BFFFCC07CFB5AFEE6FAFD" box="[311,358,1265,1287]" captionStart="Figure 10" captionStartId="21.[155,233,1778,1797]" captionTargetBox="[158,1443,205,1739]" captionTargetId="figure-60@21.[154,1446,193,1750]" captionTargetPageId="21" captionText="Figure 10. Hoplandria (Platonica) major (Sharp). (A) Habitus, dorsal view. (B) Labrum. (C) Epipharynx. (D) Mandible, dorsal view. (E) Mandible, ventral view. (F) Maxilla. (G) Labium. (H) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437814" httpUri="https://zenodo.org/record/5437814/files/figure.png" pageId="6" pageNumber="89">10A</figureCitation>
) fusiform to broadly oval; [3] surface typically glossy with [4] punctation and microsculpturing in many species; [5] pubescence generally infrequent on head and [6] abdominal terga, variably dense on [7] pronotum, [8] elytra, and [9] abdominal sterna.
</paragraph>
<paragraph id="B7319717504BFFFCC1C5FA11FED4F93F" blockId="6.[142,758,1465,1733]" pageId="6" pageNumber="89">
<emphasis id="85FA4B05504BFFFCC1C5FA11FF52FA35" box="[142,210,1466,1487]" italics="true" pageId="6" pageNumber="89">Head:</emphasis>
(
<figureCitation id="2FB58B92504BFFFCC1ADFA11FEC0FA2A" box="[230,320,1466,1488]" captionStart="Figure 6" captionStartId="15.[163,241,1761,1780]" captionTargetBox="[158,1440,200,1725]" captionTargetId="figure-61@15.[154,1449,191,1734]" captionTargetPageId="15" captionText="Figure 6. Hoplandria (Hoplandria) lateralis Melsheimer. (A) Habitus, dorsal view. (B) Labrum. (C) Epipharynx. (D) Mandible, dorsal view. (E) Mandible, ventral view. (F) Maxilla. (G) Labium. (H) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437804" httpUri="https://zenodo.org/record/5437804/files/figure.png" pageId="6" pageNumber="89">Figs 6A</figureCitation>
,
<figureCitation id="2FB58B92504BFFFCC00FFA11FEF3FA2A" box="[324,371,1466,1488]" captionStart="Figure 10" captionStartId="21.[155,233,1778,1797]" captionTargetBox="[158,1443,205,1739]" captionTargetId="figure-60@21.[154,1446,193,1750]" captionTargetPageId="21" captionText="Figure 10. Hoplandria (Platonica) major (Sharp). (A) Habitus, dorsal view. (B) Labrum. (C) Epipharynx. (D) Mandible, dorsal view. (E) Mandible, ventral view. (F) Maxilla. (G) Labium. (H) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437814" httpUri="https://zenodo.org/record/5437814/files/figure.png" pageId="6" pageNumber="89">10A</figureCitation>
) [10] Distinctly broader than long. [11] Eyes small to large, 0.3–0.7 times length of head. [12] Neck absent. [13] Infraorbital carina strongly developed and complete to absent. Antenna with [14] 11 antennomeres; [15] articles 1–3 elongate, 1.5–2.5 times longer than wide; [16] article 4 quadrate; [17] articles 5–10 short, 1.3–2.0 times wider than long, [18] coeloconical sensilla present or absent within article 11.
</paragraph>
<paragraph id="B7319717504BFFFCC1C5F94BFB93F88B" blockId="6.[142,758,1760,1904]" lastBlockId="6.[806,1422,196,1905]" pageId="6" pageNumber="89">
<emphasis id="85FA4B05504BFFFCC1C5F94BFE9AF90F" box="[142,282,1760,1781]" italics="true" pageId="6" pageNumber="89">Mouthparts:</emphasis>
Labrum (
<figureCitation id="2FB58B92504BFFFCC0D8F94BFE6DF90F" box="[403,493,1760,1782]" captionStart="Figure 2" captionStartId="9.[155,233,1736,1755]" captionTargetBox="[165,1419,207,1701]" captionTargetId="figure-52@9.[155,1435,194,1715]" captionTargetPageId="9" captionText="Figure 2. Hoplandria (Arrenhandria) laeviventris Casey. (A) Labrum. (B) Epipharynx. (C) Mandible, dorsal view. (D) Mandible, ventral view. (E) Maxilla. (F) Labium. (G) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437796" httpUri="https://zenodo.org/record/5437796/files/figure.png" pageId="6" pageNumber="89">Figs 2A</figureCitation>
,
<figureCitation id="2FB58B92504BFFFCC0B1F94BFD9BF90F" box="[506,539,1760,1781]" captionStart="Figure 4" captionStartId="12.[142,220,1775,1794]" captionTargetBox="[165,1412,194,1729]" captionTargetId="figure-53@12.[142,1432,183,1753]" captionTargetPageId="12" captionText="Figure 4. Hoplandria (Genosema) pulchra Kraatz. (A) Labrum. (B) Epipharynx. (C) Mandible, dorsal view. (D) Mandible, ventral view. (E) Maxilla. (F) Labium. (G) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437800" httpUri="https://zenodo.org/record/5437800/files/figure.png" pageId="6" pageNumber="89">4A</figureCitation>
,
<figureCitation id="2FB58B92504BFFFCC363F94BFDC9F90C" box="[552,585,1760,1782]" captionStart="Figure 6" captionStartId="15.[163,241,1761,1780]" captionTargetBox="[158,1440,200,1725]" captionTargetId="figure-61@15.[154,1449,191,1734]" captionTargetPageId="15" captionText="Figure 6. Hoplandria (Hoplandria) lateralis Melsheimer. (A) Habitus, dorsal view. (B) Labrum. (C) Epipharynx. (D) Mandible, dorsal view. (E) Mandible, ventral view. (F) Maxilla. (G) Labium. (H) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437804" httpUri="https://zenodo.org/record/5437804/files/figure.png" pageId="6" pageNumber="89">6B</figureCitation>
,
<figureCitation id="2FB58B92504BFFFCC31DF94BFDF7F90F" box="[598,631,1760,1781]" captionStart="Figure 8" captionStartId="18.[142,220,1788,1807]" captionTargetBox="[171,1406,215,1740]" captionTargetId="figure-53@18.[146,1418,194,1755]" captionTargetPageId="18" captionText="Figure 8. Hoplandria (Lophomucter) laevicollis (Notman). (A) Labrum. (B) Epipharynx. (C) Mandible, dorsal view. (D) Mandible, ventral view. (E) Maxilla. (F) Labium. (G) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437810" httpUri="https://zenodo.org/record/5437810/files/figure.png" pageId="6" pageNumber="89">8A</figureCitation>
,
<figureCitation id="2FB58B92504BFFFCC3CFF94BFD31F90C" box="[644,689,1760,1782]" captionStart="Figure 10" captionStartId="21.[155,233,1778,1797]" captionTargetBox="[158,1443,205,1739]" captionTargetId="figure-60@21.[154,1446,193,1750]" captionTargetPageId="21" captionText="Figure 10. Hoplandria (Platonica) major (Sharp). (A) Habitus, dorsal view. (B) Labrum. (C) Epipharynx. (D) Mandible, dorsal view. (E) Mandible, ventral view. (F) Maxilla. (G) Labium. (H) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437814" httpUri="https://zenodo.org/record/5437814/files/figure.png" pageId="6" pageNumber="89">10B</figureCitation>
) with epipharyngeal area with [19] medial pores large, numerous, more or less uniformly distributed in [20] typically poorly delimited, longitudinal sensory field; [21] pores between longitudinal sensory field and lateral sclerotized areas absent (
<figureCitation id="2FB58B92504BFFFCC5DEFF6FFB6FFF23" box="[1173,1263,196,218]" captionStart="Figure 2" captionStartId="9.[155,233,1736,1755]" captionTargetBox="[165,1419,207,1701]" captionTargetId="figure-52@9.[155,1435,194,1715]" captionTargetPageId="9" captionText="Figure 2. Hoplandria (Arrenhandria) laeviventris Casey. (A) Labrum. (B) Epipharynx. (C) Mandible, dorsal view. (D) Mandible, ventral view. (E) Maxilla. (F) Labium. (G) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437796" httpUri="https://zenodo.org/record/5437796/files/figure.png" pageId="6" pageNumber="89">Figs 2B</figureCitation>
,
<figureCitation id="2FB58B92504BFFFCC44AFF6FFAA2FF23" box="[1281,1314,196,217]" captionStart="Figure 4" captionStartId="12.[142,220,1775,1794]" captionTargetBox="[165,1412,194,1729]" captionTargetId="figure-53@12.[142,1432,183,1753]" captionTargetPageId="12" captionText="Figure 4. Hoplandria (Genosema) pulchra Kraatz. (A) Labrum. (B) Epipharynx. (C) Mandible, dorsal view. (D) Mandible, ventral view. (E) Maxilla. (F) Labium. (G) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437800" httpUri="https://zenodo.org/record/5437800/files/figure.png" pageId="6" pageNumber="89">4B</figureCitation>
,
<figureCitation id="2FB58B92504BFFFCC47FFF6FFAD5FF20" box="[1332,1365,196,218]" captionStart="Figure 6" captionStartId="15.[163,241,1761,1780]" captionTargetBox="[158,1440,200,1725]" captionTargetId="figure-61@15.[154,1449,191,1734]" captionTargetPageId="15" captionText="Figure 6. Hoplandria (Hoplandria) lateralis Melsheimer. (A) Habitus, dorsal view. (B) Labrum. (C) Epipharynx. (D) Mandible, dorsal view. (E) Mandible, ventral view. (F) Maxilla. (G) Labium. (H) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437804" httpUri="https://zenodo.org/record/5437804/files/figure.png" pageId="6" pageNumber="89">6C</figureCitation>
,
<figureCitation id="2FB58B92504BFFFCC42CFF6FFA09FF23" box="[1383,1417,196,217]" captionStart="Figure 8" captionStartId="18.[142,220,1788,1807]" captionTargetBox="[171,1406,215,1740]" captionTargetId="figure-53@18.[146,1418,194,1755]" captionTargetPageId="18" captionText="Figure 8. Hoplandria (Lophomucter) laevicollis (Notman). (A) Labrum. (B) Epipharynx. (C) Mandible, dorsal view. (D) Mandible, ventral view. (E) Maxilla. (F) Labium. (G) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437810" httpUri="https://zenodo.org/record/5437810/files/figure.png" pageId="6" pageNumber="89">8B</figureCitation>
,
<figureCitation id="2FB58B92504BFFFCC26DFF48FCD6FF03" box="[806,854,227,249]" captionStart="Figure 10" captionStartId="21.[155,233,1778,1797]" captionTargetBox="[158,1443,205,1739]" captionTargetId="figure-60@21.[154,1446,193,1750]" captionTargetPageId="21" captionText="Figure 10. Hoplandria (Platonica) major (Sharp). (A) Habitus, dorsal view. (B) Labrum. (C) Epipharynx. (D) Mandible, dorsal view. (E) Mandible, ventral view. (F) Maxilla. (G) Labium. (H) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437814" httpUri="https://zenodo.org/record/5437814/files/figure.png" pageId="6" pageNumber="89">10C</figureCitation>
). Mandible (
<figureCitation id="2FB58B92504BFFFCC2ACFF48FBDAFF02" box="[999,1114,227,249]" captionStart="Figure 2" captionStartId="9.[155,233,1736,1755]" captionTargetBox="[165,1419,207,1701]" captionTargetId="figure-52@9.[155,1435,194,1715]" captionTargetPageId="9" captionText="Figure 2. Hoplandria (Arrenhandria) laeviventris Casey. (A) Labrum. (B) Epipharynx. (C) Mandible, dorsal view. (D) Mandible, ventral view. (E) Maxilla. (F) Labium. (G) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437796" httpUri="https://zenodo.org/record/5437796/files/figure.png" pageId="6" pageNumber="89">Figs 2C,D</figureCitation>
,
<figureCitation id="2FB58B92504BFFFCC523FF48FB22FF02" box="[1128,1186,227,248]" captionStart="Figure 4" captionStartId="12.[142,220,1775,1794]" captionTargetBox="[165,1412,194,1729]" captionTargetId="figure-53@12.[142,1432,183,1753]" captionTargetPageId="12" captionText="Figure 4. Hoplandria (Genosema) pulchra Kraatz. (A) Labrum. (B) Epipharynx. (C) Mandible, dorsal view. (D) Mandible, ventral view. (E) Maxilla. (F) Labium. (G) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437800" httpUri="https://zenodo.org/record/5437800/files/figure.png" pageId="6" pageNumber="89">4C,D</figureCitation>
,
<figureCitation id="2FB58B92504BFFFCC5E4FF48FB6AFF03" box="[1199,1258,227,249]" captionStart="Figure 6" captionStartId="15.[163,241,1761,1780]" captionTargetBox="[158,1440,200,1725]" captionTargetId="figure-61@15.[154,1449,191,1734]" captionTargetPageId="15" captionText="Figure 6. Hoplandria (Hoplandria) lateralis Melsheimer. (A) Habitus, dorsal view. (B) Labrum. (C) Epipharynx. (D) Mandible, dorsal view. (E) Mandible, ventral view. (F) Maxilla. (G) Labium. (H) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437804" httpUri="https://zenodo.org/record/5437804/files/figure.png" pageId="6" pageNumber="89">6D,E</figureCitation>
,
<figureCitation id="2FB58B92504BFFFCC5BCFF48FAB2FF02" box="[1271,1330,227,248]" captionStart="Figure 8" captionStartId="18.[142,220,1788,1807]" captionTargetBox="[171,1406,215,1740]" captionTargetId="figure-53@18.[146,1418,194,1755]" captionTargetPageId="18" captionText="Figure 8. Hoplandria (Lophomucter) laevicollis (Notman). (A) Labrum. (B) Epipharynx. (C) Mandible, dorsal view. (D) Mandible, ventral view. (E) Maxilla. (F) Labium. (G) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437810" httpUri="https://zenodo.org/record/5437810/files/figure.png" pageId="6" pageNumber="89">8C,D</figureCitation>
,
<figureCitation id="2FB58B92504BFFFCC474FF48FA05FF03" box="[1343,1413,227,249]" captionStart="Figure 10" captionStartId="21.[155,233,1778,1797]" captionTargetBox="[158,1443,205,1739]" captionTargetId="figure-60@21.[154,1446,193,1750]" captionTargetPageId="21" captionText="Figure 10. Hoplandria (Platonica) major (Sharp). (A) Habitus, dorsal view. (B) Labrum. (C) Epipharynx. (D) Mandible, dorsal view. (E) Mandible, ventral view. (F) Maxilla. (G) Labium. (H) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437814" httpUri="https://zenodo.org/record/5437814/files/figure.png" pageId="6" pageNumber="89">10D,E</figureCitation>
) [22] asymmetrical, right mandible with a distinct median tooth; [23] apex more or less acute and curved adorally; [24] condylar molar patch small, width less than 1/5 of basal mandibular width, [25] composed of very small denticles, [26] densely arranged in irregular transverse rows; [27] subcondylar molar patch absent; dorsobasal ‘velvety patch’ [28] absent to large, size up to wider than half of mandibular base, generally composed of [29] numerous, very fine hairs or spinules; [30] ventral aspect with outer basal angle with 0–4 setae. Maxilla (
<figureCitation id="2FB58B92504BFFFCC513FD9FFB32FDB3" box="[1112,1202,564,586]" captionStart="Figure 2" captionStartId="9.[155,233,1736,1755]" captionTargetBox="[165,1419,207,1701]" captionTargetId="figure-52@9.[155,1435,194,1715]" captionTargetPageId="9" captionText="Figure 2. Hoplandria (Arrenhandria) laeviventris Casey. (A) Labrum. (B) Epipharynx. (C) Mandible, dorsal view. (D) Mandible, ventral view. (E) Maxilla. (F) Labium. (G) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437796" httpUri="https://zenodo.org/record/5437796/files/figure.png" pageId="6" pageNumber="89">Figs 2E</figureCitation>
,
<figureCitation id="2FB58B92504BFFFCC588FD9FFB65FDB3" box="[1219,1253,564,585]" captionStart="Figure 4" captionStartId="12.[142,220,1775,1794]" captionTargetBox="[165,1412,194,1729]" captionTargetId="figure-53@12.[142,1432,183,1753]" captionTargetPageId="12" captionText="Figure 4. Hoplandria (Genosema) pulchra Kraatz. (A) Labrum. (B) Epipharynx. (C) Mandible, dorsal view. (D) Mandible, ventral view. (E) Maxilla. (F) Labium. (G) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437800" httpUri="https://zenodo.org/record/5437800/files/figure.png" pageId="6" pageNumber="89">4E</figureCitation>
,
<figureCitation id="2FB58B92504BFFFCC5BDFD9FFA95FDB0" box="[1270,1301,564,586]" captionStart="Figure 6" captionStartId="15.[163,241,1761,1780]" captionTargetBox="[158,1440,200,1725]" captionTargetId="figure-61@15.[154,1449,191,1734]" captionTargetPageId="15" captionText="Figure 6. Hoplandria (Hoplandria) lateralis Melsheimer. (A) Habitus, dorsal view. (B) Labrum. (C) Epipharynx. (D) Mandible, dorsal view. (E) Mandible, ventral view. (F) Maxilla. (G) Labium. (H) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437804" httpUri="https://zenodo.org/record/5437804/files/figure.png" pageId="6" pageNumber="89">6F</figureCitation>
,
<figureCitation id="2FB58B92504BFFFCC46CFD9FFAC8FDB3" box="[1319,1352,564,585]" captionStart="Figure 8" captionStartId="18.[142,220,1788,1807]" captionTargetBox="[171,1406,215,1740]" captionTargetId="figure-53@18.[146,1418,194,1755]" captionTargetPageId="18" captionText="Figure 8. Hoplandria (Lophomucter) laevicollis (Notman). (A) Labrum. (B) Epipharynx. (C) Mandible, dorsal view. (D) Mandible, ventral view. (E) Maxilla. (F) Labium. (G) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437810" httpUri="https://zenodo.org/record/5437810/files/figure.png" pageId="6" pageNumber="89">8E</figureCitation>
,
<figureCitation id="2FB58B92504BFFFCC411FD9FFA06FDB0" box="[1370,1414,564,586]" captionStart="Figure 10" captionStartId="21.[155,233,1778,1797]" captionTargetBox="[158,1443,205,1739]" captionTargetId="figure-60@21.[154,1446,193,1750]" captionTargetPageId="21" captionText="Figure 10. Hoplandria (Platonica) major (Sharp). (A) Habitus, dorsal view. (B) Labrum. (C) Epipharynx. (D) Mandible, dorsal view. (E) Mandible, ventral view. (F) Maxilla. (G) Labium. (H) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437814" httpUri="https://zenodo.org/record/5437814/files/figure.png" pageId="6" pageNumber="89">10F</figureCitation>
) with [31] length of lacinia generally shorter than galea, [32] lacinia acute apically, [33] teeth on adoral margin short to moderate in length, about 3–5 times longer than wide, and [34] more or less closely placed, into [35] two or three regions and [36] 2 large spinose setae on dorsal surface; [37] galea narrow to wide, narrower to wider than lacinia at widest area, and rounded apically, [38] membranous in apical 1/2–1/3, [39] densely covered with rows of very short, fine hairs giving stubble-like appearance; [40] maxillary palpus with 4 articles and a moderately defined apical pseudosegment, [41] articles 2 and 3 with numerous long setae, [42] articles 1, [43] 4 and [44] pseudosegment without setae. Labium (
<figureCitation id="2FB58B92504BFFFCC57EFC4AFB0DFC0C" box="[1077,1165,993,1015]" captionStart="Figure 2" captionStartId="9.[155,233,1736,1755]" captionTargetBox="[165,1419,207,1701]" captionTargetId="figure-52@9.[155,1435,194,1715]" captionTargetPageId="9" captionText="Figure 2. Hoplandria (Arrenhandria) laeviventris Casey. (A) Labrum. (B) Epipharynx. (C) Mandible, dorsal view. (D) Mandible, ventral view. (E) Maxilla. (F) Labium. (G) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437796" httpUri="https://zenodo.org/record/5437796/files/figure.png" pageId="6" pageNumber="89">Figs 2F</figureCitation>
,
<figureCitation id="2FB58B92504BFFFCC5D3FC4AFB38FC0C" box="[1176,1208,993,1014]" captionStart="Figure 4" captionStartId="12.[142,220,1775,1794]" captionTargetBox="[165,1412,194,1729]" captionTargetId="figure-53@12.[142,1432,183,1753]" captionTargetPageId="12" captionText="Figure 4. Hoplandria (Genosema) pulchra Kraatz. (A) Labrum. (B) Epipharynx. (C) Mandible, dorsal view. (D) Mandible, ventral view. (E) Maxilla. (F) Labium. (G) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437800" httpUri="https://zenodo.org/record/5437800/files/figure.png" pageId="6" pageNumber="89">4F</figureCitation>
,
<figureCitation id="2FB58B92504BFFFCC588FC4AFB66FC0D" box="[1219,1254,993,1015]" captionStart="Figure 6" captionStartId="15.[163,241,1761,1780]" captionTargetBox="[158,1440,200,1725]" captionTargetId="figure-61@15.[154,1449,191,1734]" captionTargetPageId="15" captionText="Figure 6. Hoplandria (Hoplandria) lateralis Melsheimer. (A) Habitus, dorsal view. (B) Labrum. (C) Epipharynx. (D) Mandible, dorsal view. (E) Mandible, ventral view. (F) Maxilla. (G) Labium. (H) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437804" httpUri="https://zenodo.org/record/5437804/files/figure.png" pageId="6" pageNumber="89">6G</figureCitation>
,
<figureCitation id="2FB58B92504BFFFCC5BAFC4AFA90FC0C" box="[1265,1296,993,1014]" captionStart="Figure 8" captionStartId="18.[142,220,1788,1807]" captionTargetBox="[171,1406,215,1740]" captionTargetId="figure-53@18.[146,1418,194,1755]" captionTargetPageId="18" captionText="Figure 8. Hoplandria (Lophomucter) laevicollis (Notman). (A) Labrum. (B) Epipharynx. (C) Mandible, dorsal view. (D) Mandible, ventral view. (E) Maxilla. (F) Labium. (G) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437810" httpUri="https://zenodo.org/record/5437810/files/figure.png" pageId="6" pageNumber="89">8F</figureCitation>
,
<figureCitation id="2FB58B92504BFFFCC457FC4AFACCFC0D" box="[1308,1356,993,1015]" captionStart="Figure 10" captionStartId="21.[155,233,1778,1797]" captionTargetBox="[158,1443,205,1739]" captionTargetId="figure-60@21.[154,1446,193,1750]" captionTargetPageId="21" captionText="Figure 10. Hoplandria (Platonica) major (Sharp). (A) Habitus, dorsal view. (B) Labrum. (C) Epipharynx. (D) Mandible, dorsal view. (E) Mandible, ventral view. (F) Maxilla. (G) Labium. (H) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437814" httpUri="https://zenodo.org/record/5437814/files/figure.png" pageId="6" pageNumber="89">10G</figureCitation>
) with ligula [45] elongate, as long or longer than labial palpal segments 1 + 2, [46] slightly broadened apically with [47] shallow to deeply forked apex, [48] most commonly with two apical and two subapical spines; [49] two short or long medial setae of prementum present, generally longer to shorter than ligula, [50] insertions of setae almost adjacent to widely separated; [51] real pores and [52] setal pores present; [53] median pseudopore field narrow and more or less linear, [54] lateral pore field most commonly with a single setose pore and two asetose pores; [55] hypoglossal lobes (
<figureCitation id="2FB58B92504BFFFCC265FAFAFC09FA9C" box="[814,905,1361,1383]" captionStart="Figure 2" captionStartId="9.[155,233,1736,1755]" captionTargetBox="[165,1419,207,1701]" captionTargetId="figure-52@9.[155,1435,194,1715]" captionTargetPageId="9" captionText="Figure 2. Hoplandria (Arrenhandria) laeviventris Casey. (A) Labrum. (B) Epipharynx. (C) Mandible, dorsal view. (D) Mandible, ventral view. (E) Maxilla. (F) Labium. (G) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437796" httpUri="https://zenodo.org/record/5437796/files/figure.png" pageId="6" pageNumber="89">Figs 2G</figureCitation>
,
<figureCitation id="2FB58B92504BFFFCC2D3FAFAFC3BFA9C" box="[920,955,1361,1382]" captionStart="Figure 4" captionStartId="12.[142,220,1775,1794]" captionTargetBox="[165,1412,194,1729]" captionTargetId="figure-53@12.[142,1432,183,1753]" captionTargetPageId="12" captionText="Figure 4. Hoplandria (Genosema) pulchra Kraatz. (A) Labrum. (B) Epipharynx. (C) Mandible, dorsal view. (D) Mandible, ventral view. (E) Maxilla. (F) Labium. (G) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437800" httpUri="https://zenodo.org/record/5437800/files/figure.png" pageId="6" pageNumber="89">4G</figureCitation>
,
<figureCitation id="2FB58B92504BFFFCC281FAFAFC6EFA9D" box="[970,1006,1361,1383]" captionStart="Figure 6" captionStartId="15.[163,241,1761,1780]" captionTargetBox="[158,1440,200,1725]" captionTargetId="figure-61@15.[154,1449,191,1734]" captionTargetPageId="15" captionText="Figure 6. Hoplandria (Hoplandria) lateralis Melsheimer. (A) Habitus, dorsal view. (B) Labrum. (C) Epipharynx. (D) Mandible, dorsal view. (E) Mandible, ventral view. (F) Maxilla. (G) Labium. (H) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437804" httpUri="https://zenodo.org/record/5437804/files/figure.png" pageId="6" pageNumber="89">6H</figureCitation>
,
<figureCitation id="2FB58B92504BFFFCC2B5FAFAFBA0FA9C" box="[1022,1056,1361,1382]" captionStart="Figure 8" captionStartId="18.[142,220,1788,1807]" captionTargetBox="[171,1406,215,1740]" captionTargetId="figure-53@18.[146,1418,194,1755]" captionTargetPageId="18" captionText="Figure 8. Hoplandria (Lophomucter) laevicollis (Notman). (A) Labrum. (B) Epipharynx. (C) Mandible, dorsal view. (D) Mandible, ventral view. (E) Maxilla. (F) Labium. (G) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437810" httpUri="https://zenodo.org/record/5437810/files/figure.png" pageId="6" pageNumber="89">8G</figureCitation>
,
<figureCitation id="2FB58B92504BFFFCC57BFAFAFBE0FA9D" box="[1072,1120,1361,1383]" captionStart="Figure 10" captionStartId="21.[155,233,1778,1797]" captionTargetBox="[158,1443,205,1739]" captionTargetId="figure-60@21.[154,1446,193,1750]" captionTargetPageId="21" captionText="Figure 10. Hoplandria (Platonica) major (Sharp). (A) Habitus, dorsal view. (B) Labrum. (C) Epipharynx. (D) Mandible, dorsal view. (E) Mandible, ventral view. (F) Maxilla. (G) Labium. (H) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437814" httpUri="https://zenodo.org/record/5437814/files/figure.png" pageId="6" pageNumber="89">10H</figureCitation>
) generally long, typically reaching beyond base of ligula, [56] with very long (about five times longer than width of lobe), comb-like internally curved setae along [57] entire length of adoral margin. Labial palpus (
<figureCitation id="2FB58B92504BFFFCC5D1FA67FB72FA1B" box="[1178,1266,1484,1506]" captionStart="Figure 2" captionStartId="9.[155,233,1736,1755]" captionTargetBox="[165,1419,207,1701]" captionTargetId="figure-52@9.[155,1435,194,1715]" captionTargetPageId="9" captionText="Figure 2. Hoplandria (Arrenhandria) laeviventris Casey. (A) Labrum. (B) Epipharynx. (C) Mandible, dorsal view. (D) Mandible, ventral view. (E) Maxilla. (F) Labium. (G) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437796" httpUri="https://zenodo.org/record/5437796/files/figure.png" pageId="6" pageNumber="89">Figs 2F</figureCitation>
,
<figureCitation id="2FB58B92504BFFFCC44FFA67FAA3FA1B" box="[1284,1315,1484,1505]" captionStart="Figure 4" captionStartId="12.[142,220,1775,1794]" captionTargetBox="[165,1412,194,1729]" captionTargetId="figure-53@12.[142,1432,183,1753]" captionTargetPageId="12" captionText="Figure 4. Hoplandria (Genosema) pulchra Kraatz. (A) Labrum. (B) Epipharynx. (C) Mandible, dorsal view. (D) Mandible, ventral view. (E) Maxilla. (F) Labium. (G) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437800" httpUri="https://zenodo.org/record/5437800/files/figure.png" pageId="6" pageNumber="89">4F</figureCitation>
,
<figureCitation id="2FB58B92504BFFFCC47EFA67FAD7FA18" box="[1333,1367,1484,1506]" captionStart="Figure 6" captionStartId="15.[163,241,1761,1780]" captionTargetBox="[158,1440,200,1725]" captionTargetId="figure-61@15.[154,1449,191,1734]" captionTargetPageId="15" captionText="Figure 6. Hoplandria (Hoplandria) lateralis Melsheimer. (A) Habitus, dorsal view. (B) Labrum. (C) Epipharynx. (D) Mandible, dorsal view. (E) Mandible, ventral view. (F) Maxilla. (G) Labium. (H) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437804" httpUri="https://zenodo.org/record/5437804/files/figure.png" pageId="6" pageNumber="89">6G</figureCitation>
,
<figureCitation id="2FB58B92504BFFFCC422FA67FA09FA1B" box="[1385,1417,1484,1505]" captionStart="Figure 8" captionStartId="18.[142,220,1788,1807]" captionTargetBox="[171,1406,215,1740]" captionTargetId="figure-53@18.[146,1418,194,1755]" captionTargetPageId="18" captionText="Figure 8. Hoplandria (Lophomucter) laevicollis (Notman). (A) Labrum. (B) Epipharynx. (C) Mandible, dorsal view. (D) Mandible, ventral view. (E) Maxilla. (F) Labium. (G) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437810" httpUri="https://zenodo.org/record/5437810/files/figure.png" pageId="6" pageNumber="89">8F</figureCitation>
,
<figureCitation id="2FB58B92504BFFFCC26DFA40FCD5F9FB" box="[806,853,1515,1537]" captionStart="Figure 10" captionStartId="21.[155,233,1778,1797]" captionTargetBox="[158,1443,205,1739]" captionTargetId="figure-60@21.[154,1446,193,1750]" captionTargetPageId="21" captionText="Figure 10. Hoplandria (Platonica) major (Sharp). (A) Habitus, dorsal view. (B) Labrum. (C) Epipharynx. (D) Mandible, dorsal view. (E) Mandible, ventral view. (F) Maxilla. (G) Labium. (H) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437814" httpUri="https://zenodo.org/record/5437814/files/figure.png" pageId="6" pageNumber="89">10G</figureCitation>
) [58] elongate, overall length 4–15 times longer than greatest width, with [59] 3 distinct articles, [60] with distal pore field composed of one spine and two large pores, with small sensory pegs inside in
<emphasis id="85FA4B05504BFFFCC47AF9ECFC06F981" italics="true" pageId="6" pageNumber="89">
<taxonomicName id="708EEC94504BFFFCC47AF9ECFCD6F980" authorityName="Kraatz" authorityYear="1857" class="Insecta" family="Staphylinidae" genus="Hoplandria" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="89" phylum="Arthropoda" rank="genus" sensu="stricto">Hoplandria</taxonomicName>
<taxonomicNameLabel id="9EC9F67E504BFFFCC215F9CDFC06F981" box="[862,902,1638,1659]" pageId="6" pageNumber="89" sensu="stricto">s. s.</taxonomicNameLabel>
</emphasis>
; [61] article 1 2.0–7.0 times longer than article 2, [62] article 3 about 2.3–7.0 times longer than article 2; [63] twin pores and [64] two median pores present.
<collectingRegion id="754A59F5504BFFFCC2C1F969FC6BF92D" box="[906,1003,1730,1751]" country="Myanmar" name="Chin" pageId="6" pageNumber="89">Mentum</collectingRegion>
with [65] apical margin flattened to slightly concave with [66] antero-lateral angles obtusely rounded and often extended anteriorly; [67] many sensory pores typically distributed in basal 2/3, however, numerous taxa with pores distrubuted along each side of midline.
</paragraph>
<paragraph id="B7319717504AFFFDC1D0FF6FFF4BFB54" blockId="7.[155,771,195,1198]" pageId="7" pageNumber="90">
<emphasis id="85FA4B05504AFFFDC1D0FF6FFF73FF23" box="[155,243,196,217]" italics="true" pageId="7" pageNumber="90">Thorax:</emphasis>
Pronotum (
<figureCitation id="2FB58B92504AFFFDC0CBFF6FFE59FF20" box="[384,473,196,218]" captionStart="Figure 6" captionStartId="15.[163,241,1761,1780]" captionTargetBox="[158,1440,200,1725]" captionTargetId="figure-61@15.[154,1449,191,1734]" captionTargetPageId="15" captionText="Figure 6. Hoplandria (Hoplandria) lateralis Melsheimer. (A) Habitus, dorsal view. (B) Labrum. (C) Epipharynx. (D) Mandible, dorsal view. (E) Mandible, ventral view. (F) Maxilla. (G) Labium. (H) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437804" httpUri="https://zenodo.org/record/5437804/files/figure.png" pageId="7" pageNumber="90">Figs 6A</figureCitation>
,
<figureCitation id="2FB58B92504AFFFDC0AEFF6FFD93FF20" box="[485,531,196,218]" captionStart="Figure 10" captionStartId="21.[155,233,1778,1797]" captionTargetBox="[158,1443,205,1739]" captionTargetId="figure-60@21.[154,1446,193,1750]" captionTargetPageId="21" captionText="Figure 10. Hoplandria (Platonica) major (Sharp). (A) Habitus, dorsal view. (B) Labrum. (C) Epipharynx. (D) Mandible, dorsal view. (E) Mandible, ventral view. (F) Maxilla. (G) Labium. (H) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437814" httpUri="https://zenodo.org/record/5437814/files/figure.png" pageId="7" pageNumber="90">10A</figureCitation>
) [68] subquadrate to transverse, variable in size, [69] typically 1.2–1.4 times wider than long, [70] more or less convex to flattened. Setae [71] generally densely distributed, [72] typically directed posteriorly at midline on disc, often with differentiated macrosetae near lateral margins. [73] Hypomera strongly inflexed, not visible in lateral aspect. Each elytron [74] moderately broad, commonly wider apically than basally; [75] apico-lateral angles sinuate; [76] elytra together about 1.3–1.8 times as wide as long; [77] microsetae numerous, [78] generally uniformly distributed and [79] directed posteriorly. Mesosternum (
<figureCitation id="2FB58B92504AFFFDC00CFD9FFE20FDB3" box="[327,416,564,586]" captionStart="Figure 3" captionStartId="10.[142,220,1355,1374]" captionTargetBox="[142,1420,198,1318]" captionTargetId="figure-275@10.[142,1421,196,1322]" captionTargetPageId="10" captionText="Figure 3. Hoplandria (Arrhenandria) laeviventris Casey. (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Aedeagus, lateral view. (D) Paramere, lateral view. (E) Spermatheca. (F) Left elytron, male (arrow to denticle)." figureDoi="http://doi.org/10.5281/zenodo.5437798" httpUri="https://zenodo.org/record/5437798/files/figure.png" pageId="7" pageNumber="90">Figs 3A</figureCitation>
,
<figureCitation id="2FB58B92504AFFFDC0E6FD9FFE4EFDB0" box="[429,462,564,586]" captionStart="Figure 5" captionStartId="13.[155,233,1367,1386]" captionTargetBox="[158,1441,195,1329]" captionTargetId="figure-246@13.[154,1446,191,1337]" captionTargetPageId="13" captionText="Figure 5. Hoplandria (Genosema) pulchra Kraatz. (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Aedeagus, lateral view. (D) Paramere, lateral view. (E) Spermatheca. (F) Left elytron, male (arrow to denticle)." figureDoi="http://doi.org/10.5281/zenodo.5437802" httpUri="https://zenodo.org/record/5437802/files/figure.png" pageId="7" pageNumber="90">5A</figureCitation>
,
<figureCitation id="2FB58B92504AFFFDC090FD9FFE7DFDB3" box="[475,509,564,585]" captionStart="Figure 7" captionStartId="16.[142,220,1153,1172]" captionTargetBox="[164,1401,200,1115]" captionTargetId="figure-378@16.[153,1410,194,1120]" captionTargetPageId="16" captionText="Figure 7. Hoplandria (Hoplandria) lateralis Melsheimer. (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Tarsus of hind leg. (D) Aedeagus, lateral view. (E) Paramere, lateral view. (F) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5437808" httpUri="https://zenodo.org/record/5437808/files/figure.png" pageId="7" pageNumber="90">7A</figureCitation>
,
<figureCitation id="2FB58B92504AFFFDC342FD9FFDABFDB3" box="[521,555,564,585]" captionStart="Figure 9" captionStartId="19.[155,233,1370,1389]" captionTargetBox="[167,1437,210,1332]" captionTargetId="figure-265@19.[155,1453,191,1341]" captionTargetPageId="19" captionText="Figure 9. Hoplandria (Lophomucter) laevicollis (Notman). (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Aedeagus, lateral view. (D) Paramere, lateral view. (E) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5437812" httpUri="https://zenodo.org/record/5437812/files/figure.png" pageId="7" pageNumber="90">9A</figureCitation>
,
<figureCitation id="2FB58B92504AFFFDC37CFD9FFDE4FDB0" box="[567,612,564,586]" captionStart="Figure 11" captionStartId="22.[142,220,1298,1317]" captionTargetBox="[148,1421,210,1263]" captionTargetId="figure-328@22.[133,1431,194,1271]" captionTargetPageId="22" captionText="Figure 11. Hoplandria (Platonica) major (Sharp). (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Aedeagus, lateral view. (D) Paramere, lateral view. (E) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5437816" httpUri="https://zenodo.org/record/5437816/files/figure.png" pageId="7" pageNumber="90">11A</figureCitation>
) [80] without medial carina. Mesocoxal cavities (
<figureCitation id="2FB58B92504AFFFDC374FDF8FD18FD92" box="[575,664,595,617]" captionStart="Figure 3" captionStartId="10.[142,220,1355,1374]" captionTargetBox="[142,1420,198,1318]" captionTargetId="figure-275@10.[142,1421,196,1322]" captionTargetPageId="10" captionText="Figure 3. Hoplandria (Arrhenandria) laeviventris Casey. (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Aedeagus, lateral view. (D) Paramere, lateral view. (E) Spermatheca. (F) Left elytron, male (arrow to denticle)." figureDoi="http://doi.org/10.5281/zenodo.5437798" httpUri="https://zenodo.org/record/5437798/files/figure.png" pageId="7" pageNumber="90">Figs 3A</figureCitation>
,
<figureCitation id="2FB58B92504AFFFDC3E1FDF8FD4BFD93" box="[682,715,595,617]" captionStart="Figure 5" captionStartId="13.[155,233,1367,1386]" captionTargetBox="[158,1441,195,1329]" captionTargetId="figure-246@13.[154,1446,191,1337]" captionTargetPageId="13" captionText="Figure 5. Hoplandria (Genosema) pulchra Kraatz. (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Aedeagus, lateral view. (D) Paramere, lateral view. (E) Spermatheca. (F) Left elytron, male (arrow to denticle)." figureDoi="http://doi.org/10.5281/zenodo.5437802" httpUri="https://zenodo.org/record/5437802/files/figure.png" pageId="7" pageNumber="90">5A</figureCitation>
,
<figureCitation id="2FB58B92504AFFFDC397FDF8FD7EFD92" box="[732,766,595,616]" captionStart="Figure 7" captionStartId="16.[142,220,1153,1172]" captionTargetBox="[164,1401,200,1115]" captionTargetId="figure-378@16.[153,1410,194,1120]" captionTargetPageId="16" captionText="Figure 7. Hoplandria (Hoplandria) lateralis Melsheimer. (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Tarsus of hind leg. (D) Aedeagus, lateral view. (E) Paramere, lateral view. (F) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5437808" httpUri="https://zenodo.org/record/5437808/files/figure.png" pageId="7" pageNumber="90">7A</figureCitation>
,
<figureCitation id="2FB58B92504AFFFDC1D0FDDAFF3CFD7C" box="[155,188,625,646]" captionStart="Figure 9" captionStartId="19.[155,233,1370,1389]" captionTargetBox="[167,1437,210,1332]" captionTargetId="figure-265@19.[155,1453,191,1341]" captionTargetPageId="19" captionText="Figure 9. Hoplandria (Lophomucter) laevicollis (Notman). (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Aedeagus, lateral view. (D) Paramere, lateral view. (E) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5437812" httpUri="https://zenodo.org/record/5437812/files/figure.png" pageId="7" pageNumber="90">9A</figureCitation>
,
<figureCitation id="2FB58B92504AFFFDC182FDDAFF76FD7D" box="[201,246,625,647]" captionStart="Figure 11" captionStartId="22.[142,220,1298,1317]" captionTargetBox="[148,1421,210,1263]" captionTargetId="figure-328@22.[133,1431,194,1271]" captionTargetPageId="22" captionText="Figure 11. Hoplandria (Platonica) major (Sharp). (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Aedeagus, lateral view. (D) Paramere, lateral view. (E) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5437816" httpUri="https://zenodo.org/record/5437816/files/figure.png" pageId="7" pageNumber="90">11A</figureCitation>
) [81] widely separated by meso- and metasternal processes by about 1/5 length of coxal cavities. Mesosternal process [82] longer than metasternal process, extended to basal 1/5 of coxal cavities; [83] meso- and metasternal processes typically separated by very short isthmus or contiguous, isthmus absent; [84] mesosternal process rounded or flattened at apex. Metasternum [85] shorter to longer than width of mesocoxae, [86] without medial carina; [87] metasternal process generally flattened at apex. Macrosetae [88] present or absent on metasternum; [89] absent from mesosternum. Legs [90] with tarsal formula 4-5- 5; [91] tarsal claws (
<figureCitation id="2FB58B92504AFFFDC0DCFC4AFE69FC0D" box="[407,489,993,1015]" captionStart="Figure 7" captionStartId="16.[142,220,1153,1172]" captionTargetBox="[164,1401,200,1115]" captionTargetId="figure-378@16.[153,1410,194,1120]" captionTargetPageId="16" captionText="Figure 7. Hoplandria (Hoplandria) lateralis Melsheimer. (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Tarsus of hind leg. (D) Aedeagus, lateral view. (E) Paramere, lateral view. (F) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5437808" httpUri="https://zenodo.org/record/5437808/files/figure.png" pageId="7" pageNumber="90">Fig. 7C</figureCitation>
) long and slender, [92] with single empodial bristle, [93] empodial bristle distinctly shorter to distinctly longer than tarsal claws; [94] article 1 of hind tarsus about 1.3 times length of article 2; [95] articles 2–4 subequal in length; [96] article 5 subequal in length to combined lengths of articles 2–4.
</paragraph>
<paragraph id="B7319717504AFFFDC1D0FB6CFE0FFA0B" blockId="7.[155,771,1223,1521]" pageId="7" pageNumber="90">
<emphasis id="85FA4B05504AFFFDC1D0FB6CFE8BFB26" box="[155,267,1223,1244]" italics="true" pageId="7" pageNumber="90">Abdomen:</emphasis>
(
<figureCitation id="2FB58B92504AFFFDC057FB6CFEF6FB26" box="[284,374,1223,1245]" captionStart="Figure 3" captionStartId="10.[142,220,1355,1374]" captionTargetBox="[142,1420,198,1318]" captionTargetId="figure-275@10.[142,1421,196,1322]" captionTargetPageId="10" captionText="Figure 3. Hoplandria (Arrhenandria) laeviventris Casey. (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Aedeagus, lateral view. (D) Paramere, lateral view. (E) Spermatheca. (F) Left elytron, male (arrow to denticle)." figureDoi="http://doi.org/10.5281/zenodo.5437798" httpUri="https://zenodo.org/record/5437798/files/figure.png" pageId="7" pageNumber="90">Figs 3B</figureCitation>
,
<figureCitation id="2FB58B92504AFFFDC0CCFB6CFE28FB27" box="[391,424,1223,1245]" captionStart="Figure 5" captionStartId="13.[155,233,1367,1386]" captionTargetBox="[158,1441,195,1329]" captionTargetId="figure-246@13.[154,1446,191,1337]" captionTargetPageId="13" captionText="Figure 5. Hoplandria (Genosema) pulchra Kraatz. (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Aedeagus, lateral view. (D) Paramere, lateral view. (E) Spermatheca. (F) Left elytron, male (arrow to denticle)." figureDoi="http://doi.org/10.5281/zenodo.5437802" httpUri="https://zenodo.org/record/5437802/files/figure.png" pageId="7" pageNumber="90">5B</figureCitation>
,
<figureCitation id="2FB58B92504AFFFDC0F1FB6CFE5CFB27" box="[442,476,1223,1245]" captionStart="Figure 6" captionStartId="15.[163,241,1761,1780]" captionTargetBox="[158,1440,200,1725]" captionTargetId="figure-61@15.[154,1449,191,1734]" captionTargetPageId="15" captionText="Figure 6. Hoplandria (Hoplandria) lateralis Melsheimer. (A) Habitus, dorsal view. (B) Labrum. (C) Epipharynx. (D) Mandible, dorsal view. (E) Mandible, ventral view. (F) Maxilla. (G) Labium. (H) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437804" httpUri="https://zenodo.org/record/5437804/files/figure.png" pageId="7" pageNumber="90">6A</figureCitation>
,
<figureCitation id="2FB58B92504AFFFDC0A6FB63FD8EFB27" box="[493,526,1224,1245]" captionStart="Figure 7" captionStartId="16.[142,220,1153,1172]" captionTargetBox="[164,1401,200,1115]" captionTargetId="figure-378@16.[153,1410,194,1120]" captionTargetPageId="16" captionText="Figure 7. Hoplandria (Hoplandria) lateralis Melsheimer. (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Tarsus of hind leg. (D) Aedeagus, lateral view. (E) Paramere, lateral view. (F) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5437808" httpUri="https://zenodo.org/record/5437808/files/figure.png" pageId="7" pageNumber="90">7B</figureCitation>
,
<figureCitation id="2FB58B92504AFFFDC36BFB6CFDC1FB26" box="[544,577,1223,1244]" captionStart="Figure 9" captionStartId="19.[155,233,1370,1389]" captionTargetBox="[167,1437,210,1332]" captionTargetId="figure-265@19.[155,1453,191,1341]" captionTargetPageId="19" captionText="Figure 9. Hoplandria (Lophomucter) laevicollis (Notman). (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Aedeagus, lateral view. (D) Paramere, lateral view. (E) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5437812" httpUri="https://zenodo.org/record/5437812/files/figure.png" pageId="7" pageNumber="90">9B</figureCitation>
,
<figureCitation id="2FB58B92504AFFFDC318FB6CFD02FB26" box="[595,642,1223,1244]" captionStart="Figure 10" captionStartId="21.[155,233,1778,1797]" captionTargetBox="[158,1443,205,1739]" captionTargetId="figure-60@21.[154,1446,193,1750]" captionTargetPageId="21" captionText="Figure 10. Hoplandria (Platonica) major (Sharp). (A) Habitus, dorsal view. (B) Labrum. (C) Epipharynx. (D) Mandible, dorsal view. (E) Mandible, ventral view. (F) Maxilla. (G) Labium. (H) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437814" httpUri="https://zenodo.org/record/5437814/files/figure.png" pageId="7" pageNumber="90">10A</figureCitation>
,
<figureCitation id="2FB58B92504AFFFDC3D8FB6CFD40FB27" box="[659,704,1223,1245]" captionStart="Figure 11" captionStartId="22.[142,220,1298,1317]" captionTargetBox="[148,1421,210,1263]" captionTargetId="figure-328@22.[133,1431,194,1271]" captionTargetPageId="22" captionText="Figure 11. Hoplandria (Platonica) major (Sharp). (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Aedeagus, lateral view. (D) Paramere, lateral view. (E) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5437816" httpUri="https://zenodo.org/record/5437816/files/figure.png" pageId="7" pageNumber="90">11B</figureCitation>
) [97] Fusiform, tapering apically to broadly pointed apex; [98] terga
<collectionCode id="D19F0FD2504AFFFDC05CFAAEFEB2FAE0" box="[279,306,1285,1306]" pageId="7" pageNumber="90">III</collectionCode>
–
<collectionCode id="D19F0FD2504AFFFDC00AFAAEFED5FAE0" box="[321,341,1285,1306]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="7" pageNumber="90" type="Museum">V</collectionCode>
(
<collectionCode id="D19F0FD2504AFFFDC02CFAAEFE04FAE0" box="[359,388,1285,1306]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="7" pageNumber="90">VI</collectionCode>
slightly) with moderate to deep transverse basal depressions on anterior portion. [99] Terga and sterna with prominent macrosetae. [100] Anterior corner of sternum
<collectionCode id="D19F0FD2504AFFFDC090FACAFE79FA8C" box="[475,505,1377,1398]" pageId="7" pageNumber="90">III</collectionCode>
with distinct region of differentiated microsculpturing, generally delimited by very fine carina, area appearing irridescent in many species. [101] Tergum IX with distinct crescentshaped setal pattern.
</paragraph>
<paragraph id="B7319717504AFFFDC1D0F9A2FC3BFE89" blockId="7.[155,771,1545,1905]" lastBlockId="7.[819,1435,195,371]" pageId="7" pageNumber="90">
<emphasis id="85FA4B05504AFFFDC1D0F9A2FD9DF9E4" box="[155,541,1545,1566]" italics="true" pageId="7" pageNumber="90">Secondary sexual characteristics:</emphasis>
Males highly variable, including more or less distinct major and minor forms. Minor forms generally similar to females and typically indistinct externally. Major forms highly variable among species with morphological modifications on elytra in form of [102] humeral carina or bump; denticle, [103] carina or conical process on the medioapical region (
<figureCitation id="2FB58B92504AFFFDC0C7F94BFE66F90C" box="[396,486,1760,1782]" captionStart="Figure 6" captionStartId="15.[163,241,1761,1780]" captionTargetBox="[158,1440,200,1725]" captionTargetId="figure-61@15.[154,1449,191,1734]" captionTargetPageId="15" captionText="Figure 6. Hoplandria (Hoplandria) lateralis Melsheimer. (A) Habitus, dorsal view. (B) Labrum. (C) Epipharynx. (D) Mandible, dorsal view. (E) Mandible, ventral view. (F) Maxilla. (G) Labium. (H) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437804" httpUri="https://zenodo.org/record/5437804/files/figure.png" pageId="7" pageNumber="90">Figs 6A</figureCitation>
,
<figureCitation id="2FB58B92504AFFFDC0B3F94BFDA7F90C" box="[504,551,1760,1782]" captionStart="Figure 10" captionStartId="21.[155,233,1778,1797]" captionTargetBox="[158,1443,205,1739]" captionTargetId="figure-60@21.[154,1446,193,1750]" captionTargetPageId="21" captionText="Figure 10. Hoplandria (Platonica) major (Sharp). (A) Habitus, dorsal view. (B) Labrum. (C) Epipharynx. (D) Mandible, dorsal view. (E) Mandible, ventral view. (F) Maxilla. (G) Labium. (H) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437814" httpUri="https://zenodo.org/record/5437814/files/figure.png" pageId="7" pageNumber="90">10A</figureCitation>
); [104] well-developed denticle, carina or bump on the outer apical third (
<figureCitation id="2FB58B92504AFFFDC1E9F8B6FF74F8C9" box="[162,244,1821,1843]" captionStart="Figure 5" captionStartId="13.[155,233,1367,1386]" captionTargetBox="[158,1441,195,1329]" captionTargetId="figure-246@13.[154,1446,191,1337]" captionTargetPageId="13" captionText="Figure 5. Hoplandria (Genosema) pulchra Kraatz. (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Aedeagus, lateral view. (D) Paramere, lateral view. (E) Spermatheca. (F) Left elytron, male (arrow to denticle)." figureDoi="http://doi.org/10.5281/zenodo.5437802" httpUri="https://zenodo.org/record/5437802/files/figure.png" pageId="7" pageNumber="90">Fig. 5F</figureCitation>
). Abdomen: posterior corners of [105] sternum
<collectionCode id="D19F0FD2504AFFFDC1D0F897FF39F8AB" box="[155,185,1852,1873]" pageId="7" pageNumber="90">III</collectionCode>
and [106]
<collectionCode id="D19F0FD2504AFFFDC07BF897FECCF8AB" box="[304,332,1852,1873]" pageId="7" pageNumber="90">IV</collectionCode>
produced into long spines often reaching the posterior quarter of tergum
<collectionCode id="D19F0FD2504AFFFDC377F8F0FDD8F88A" box="[572,600,1883,1904]" pageId="7" pageNumber="90">IV</collectionCode>
and
<collectionCode id="D19F0FD2504AFFFDC3DFF8F0FD25F88A" box="[660,677,1883,1904]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="7" pageNumber="90" type="Museum">V</collectionCode>
, respectively; [107] tergum
<collectionCode id="D19F0FD2504AFFFDC563FF6FFBC4FF23" box="[1064,1092,196,217]" pageId="7" pageNumber="90">IV</collectionCode>
with more or less U-shaped (
<figureCitation id="2FB58B92504AFFFDC271FF48FC0DFF03" box="[826,909,227,249]" captionStart="Figure 6" captionStartId="15.[163,241,1761,1780]" captionTargetBox="[158,1440,200,1725]" captionTargetId="figure-61@15.[154,1449,191,1734]" captionTargetPageId="15" captionText="Figure 6. Hoplandria (Hoplandria) lateralis Melsheimer. (A) Habitus, dorsal view. (B) Labrum. (C) Epipharynx. (D) Mandible, dorsal view. (E) Mandible, ventral view. (F) Maxilla. (G) Labium. (H) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437804" httpUri="https://zenodo.org/record/5437804/files/figure.png" pageId="7" pageNumber="90">Fig. 6A</figureCitation>
) or narrow longtitudinal carina, often appearing iridescent; [108] tergum VII with narrow posteromedian longtitudinal carina (
<figureCitation id="2FB58B92504AFFFDC534FE8BFB59FECC" box="[1151,1241,288,310]" captionStart="Figure 6" captionStartId="15.[163,241,1761,1780]" captionTargetBox="[158,1440,200,1725]" captionTargetId="figure-61@15.[154,1449,191,1734]" captionTargetPageId="15" captionText="Figure 6. Hoplandria (Hoplandria) lateralis Melsheimer. (A) Habitus, dorsal view. (B) Labrum. (C) Epipharynx. (D) Mandible, dorsal view. (E) Mandible, ventral view. (F) Maxilla. (G) Labium. (H) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437804" httpUri="https://zenodo.org/record/5437804/files/figure.png" pageId="7" pageNumber="90">Figs 6A</figureCitation>
,
<figureCitation id="2FB58B92504AFFFDC5A8FE8BFA93FECC" box="[1251,1299,288,310]" captionStart="Figure 10" captionStartId="21.[155,233,1778,1797]" captionTargetBox="[158,1443,205,1739]" captionTargetId="figure-60@21.[154,1446,193,1750]" captionTargetPageId="21" captionText="Figure 10. Hoplandria (Platonica) major (Sharp). (A) Habitus, dorsal view. (B) Labrum. (C) Epipharynx. (D) Mandible, dorsal view. (E) Mandible, ventral view. (F) Maxilla. (G) Labium. (H) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437814" httpUri="https://zenodo.org/record/5437814/files/figure.png" pageId="7" pageNumber="90">10A</figureCitation>
); [109] apex of tergum
<collectionCode id="D19F0FD2504AFFFDC2FCFE94FC67FEAE" box="[951,999,319,340]" pageId="7" pageNumber="90">VIII</collectionCode>
with two transverse rows of large macrosetae.
</paragraph>
<paragraph id="B7319717504AFFFDC278FE3BFA0CFD23" blockId="7.[819,1435,400,913]" pageId="7" pageNumber="90">
<emphasis id="85FA4B05504AFFFDC278FE3BFC25FE5F" box="[819,933,400,421]" italics="true" pageId="7" pageNumber="90">Aedeagus:</emphasis>
(
<figureCitation id="2FB58B92504AFFFDC2F1FE3BFB93FE5C" box="[954,1043,400,422]" captionStart="Figure 3" captionStartId="10.[142,220,1355,1374]" captionTargetBox="[142,1420,198,1318]" captionTargetId="figure-275@10.[142,1421,196,1322]" captionTargetPageId="10" captionText="Figure 3. Hoplandria (Arrhenandria) laeviventris Casey. (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Aedeagus, lateral view. (D) Paramere, lateral view. (E) Spermatheca. (F) Left elytron, male (arrow to denticle)." figureDoi="http://doi.org/10.5281/zenodo.5437798" httpUri="https://zenodo.org/record/5437798/files/figure.png" pageId="7" pageNumber="90">Figs 3C</figureCitation>
,
<figureCitation id="2FB58B92504AFFFDC555FE3BFBBFFE5C" box="[1054,1087,400,422]" captionStart="Figure 5" captionStartId="13.[155,233,1367,1386]" captionTargetBox="[158,1441,195,1329]" captionTargetId="figure-246@13.[154,1446,191,1337]" captionTargetPageId="13" captionText="Figure 5. Hoplandria (Genosema) pulchra Kraatz. (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Aedeagus, lateral view. (D) Paramere, lateral view. (E) Spermatheca. (F) Left elytron, male (arrow to denticle)." figureDoi="http://doi.org/10.5281/zenodo.5437802" httpUri="https://zenodo.org/record/5437802/files/figure.png" pageId="7" pageNumber="90">5C</figureCitation>
,
<figureCitation id="2FB58B92504AFFFDC500FE3AFBEDFE5C" box="[1099,1133,401,422]" captionStart="Figure 7" captionStartId="16.[142,220,1153,1172]" captionTargetBox="[164,1401,200,1115]" captionTargetId="figure-378@16.[153,1410,194,1120]" captionTargetPageId="16" captionText="Figure 7. Hoplandria (Hoplandria) lateralis Melsheimer. (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Tarsus of hind leg. (D) Aedeagus, lateral view. (E) Paramere, lateral view. (F) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5437808" httpUri="https://zenodo.org/record/5437808/files/figure.png" pageId="7" pageNumber="90">7D</figureCitation>
,
<figureCitation id="2FB58B92504AFFFDC533FE3AFB19FE5C" box="[1144,1177,401,422]" captionStart="Figure 9" captionStartId="19.[155,233,1370,1389]" captionTargetBox="[167,1437,210,1332]" captionTargetId="figure-265@19.[155,1453,191,1341]" captionTargetPageId="19" captionText="Figure 9. Hoplandria (Lophomucter) laevicollis (Notman). (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Aedeagus, lateral view. (D) Paramere, lateral view. (E) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5437812" httpUri="https://zenodo.org/record/5437812/files/figure.png" pageId="7" pageNumber="90">9C</figureCitation>
,
<figureCitation id="2FB58B92504AFFFDC5EEFE3AFB52FE5D" box="[1189,1234,401,423]" captionStart="Figure 11" captionStartId="22.[142,220,1298,1317]" captionTargetBox="[148,1421,210,1263]" captionTargetId="figure-328@22.[133,1431,194,1271]" captionTargetPageId="22" captionText="Figure 11. Hoplandria (Platonica) major (Sharp). (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Aedeagus, lateral view. (D) Paramere, lateral view. (E) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5437816" httpUri="https://zenodo.org/record/5437816/files/figure.png" pageId="7" pageNumber="90">11C</figureCitation>
) Highly variable, with bulb of median lobe [110] more or less elongate, subequal to or greater than tube, [111] with distinct ventral projection; internal sac [112] typically with large spinules; [113] paramere with apical lobe of parmerite not greatly extended beyond velum, [114] typically with 2–3 setae at apex, 2 most commonly; [115] paramerite with anterior margin generally slightly to distinctly concave; condylite with [116] apex unmodified to heavily sclerotized, [117] generally shorter to subequal in length to apex of paramerite.
</paragraph>
<paragraph id="B7319717504AFFFDC200FD49FB2FFC6B" blockId="7.[819,1435,400,913]" pageId="7" pageNumber="90">
<emphasis id="85FA4B05504AFFFDC200FD49FC66FD0D" box="[843,998,738,759]" italics="true" pageId="7" pageNumber="90">Spermatheca:</emphasis>
(
<figureCitation id="2FB58B92504AFFFDC2B1FD49FBD3FD0D" box="[1018,1107,738,760]" captionStart="Figure 3" captionStartId="10.[142,220,1355,1374]" captionTargetBox="[142,1420,198,1318]" captionTargetId="figure-275@10.[142,1421,196,1322]" captionTargetPageId="10" captionText="Figure 3. Hoplandria (Arrhenandria) laeviventris Casey. (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Aedeagus, lateral view. (D) Paramere, lateral view. (E) Spermatheca. (F) Left elytron, male (arrow to denticle)." figureDoi="http://doi.org/10.5281/zenodo.5437798" httpUri="https://zenodo.org/record/5437798/files/figure.png" pageId="7" pageNumber="90">Figs 3E</figureCitation>
,
<figureCitation id="2FB58B92504AFFFDC528FD49FB03FD02" box="[1123,1155,738,760]" captionStart="Figure 5" captionStartId="13.[155,233,1367,1386]" captionTargetBox="[158,1441,195,1329]" captionTargetId="figure-246@13.[154,1446,191,1337]" captionTargetPageId="13" captionText="Figure 5. Hoplandria (Genosema) pulchra Kraatz. (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Aedeagus, lateral view. (D) Paramere, lateral view. (E) Spermatheca. (F) Left elytron, male (arrow to denticle)." figureDoi="http://doi.org/10.5281/zenodo.5437802" httpUri="https://zenodo.org/record/5437802/files/figure.png" pageId="7" pageNumber="90">5E</figureCitation>
,
<figureCitation id="2FB58B92504AFFFDC5D8FD49FB30FD0D" box="[1171,1200,738,759]" captionStart="Figure 7" captionStartId="16.[142,220,1153,1172]" captionTargetBox="[164,1401,200,1115]" captionTargetId="figure-378@16.[153,1410,194,1120]" captionTargetPageId="16" captionText="Figure 7. Hoplandria (Hoplandria) lateralis Melsheimer. (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Tarsus of hind leg. (D) Aedeagus, lateral view. (E) Paramere, lateral view. (F) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5437808" httpUri="https://zenodo.org/record/5437808/files/figure.png" pageId="7" pageNumber="90">7F</figureCitation>
,
<figureCitation id="2FB58B92504AFFFDC5F4FD49FB5FFD0D" box="[1215,1247,738,759]" captionStart="Figure 9" captionStartId="19.[155,233,1370,1389]" captionTargetBox="[167,1437,210,1332]" captionTargetId="figure-265@19.[155,1453,191,1341]" captionTargetPageId="19" captionText="Figure 9. Hoplandria (Lophomucter) laevicollis (Notman). (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Aedeagus, lateral view. (D) Paramere, lateral view. (E) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5437812" httpUri="https://zenodo.org/record/5437812/files/figure.png" pageId="7" pageNumber="90">9E</figureCitation>
and
<figureCitation id="2FB58B92504AFFFDC46BFD49FACDFD02" box="[1312,1357,738,760]" captionStart="Figure 11" captionStartId="22.[142,220,1298,1317]" captionTargetBox="[148,1421,210,1263]" captionTargetId="figure-328@22.[133,1431,194,1271]" captionTargetPageId="22" captionText="Figure 11. Hoplandria (Platonica) major (Sharp). (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Aedeagus, lateral view. (D) Paramere, lateral view. (E) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5437816" httpUri="https://zenodo.org/record/5437816/files/figure.png" pageId="7" pageNumber="90">11E</figureCitation>
) [118] Generally L- shaped with three primary divisions: basal bulb, [119] generally simple and rounded at base; neck, [120] significantly bent, commonly at 90∞ angle or greater; and tube, [121] typically membranous, [122] more or less straight.
</paragraph>
<paragraph id="B7319717504AFFFDC278FC05FA1AFA37" blockId="7.[819,1435,942,1485]" pageId="7" pageNumber="90">
<emphasis id="85FA4B05504AFFFDC278FC05FC11FC39" box="[819,913,942,963]" italics="true" pageId="7" pageNumber="90">Habitat:</emphasis>
<bibRefCitation id="D31FEAE6504AFFFDC2D5FC05FBBDFC3E" author="Genier F" box="[926,1085,942,964]" pageId="7" pageNumber="90" pagination="3 - 59" refId="ref37573" refString="Genier F. 1989. A revision of the genus Hoplandria Kraatz of America north of Mexico (Coleoptera: Staphylinidae, Aleocharinae). Memoirs of the Entomological Society of Canada 150: 3 - 59." type="journal article" year="1989">Génier (1989)</bibRefCitation>
reported that species of
<taxonomicName id="708EEC94504AFFFDC428FC04FC09FC18" authorityName="Kraatz" authorityYear="1857" class="Insecta" family="Staphylinidae" genus="Hoplandria" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="85FA4B05504AFFFDC428FC04FC09FC18" italics="true" pageId="7" pageNumber="90">Hoplandria</emphasis>
</taxonomicName>
are mostly saprophytic, with some records of adults collected at light traps. Anecdotal collection data suggests that adults of species of
<emphasis id="85FA4B05504AFFFDC5A5FBA1FA1AFBE5" box="[1262,1434,1034,1055]" italics="true" pageId="7" pageNumber="90">
<taxonomicName id="708EEC94504AFFFDC5A5FBA1FAF3FBE5" authorityName="Kraatz" authorityYear="1857" box="[1262,1395,1034,1055]" class="Insecta" family="Staphylinidae" genus="Hoplandria" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="90" phylum="Arthropoda" rank="genus" sensu="lato">Hoplandria</taxonomicName>
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</emphasis>
feed on dipteran larvae within various decaying organic materials, especially dung and carrion. Recently, the late instar larvae of
<taxonomicName id="708EEC94504AFFFDC586FBCCFBA8FB61" authority="Genier" authorityName="Genier" authorityYear="1989" class="Insecta" family="Staphylinidae" genus="Hoplandria" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="90" phylum="Arthropoda" rank="species" species="klimaszewskii" subGenus="Lophomucter">
<emphasis id="85FA4B05504AFFFDC586FBCCFB62FB86" box="[1229,1250,1127,1148]" italics="true" pageId="7" pageNumber="90">H</emphasis>
. (
<emphasis id="85FA4B05504AFFFDC5B6FBCDFA12FB81" box="[1277,1426,1126,1147]" italics="true" pageId="7" pageNumber="90">Lophomucter</emphasis>
)
<emphasis id="85FA4B05504AFFFDC278FB2EFC53FB60" box="[819,979,1157,1178]" italics="true" pageId="7" pageNumber="90">klimaszewskii</emphasis>
Génier
</taxonomicName>
were described from the
<collectingCountry id="CF99D787504AFFFDC403FB2EFAECFB60" box="[1352,1388,1157,1178]" name="United States of America" pageId="7" pageNumber="90">US</collectingCountry>
(M. Thayer, J. Ashe &amp;
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. Hanley, pers. comm.). These morphologically distinct larvae were generally collected from under oak logs where they typically were very active and fast-moving. Reared individuals were offered
<taxonomicName id="708EEC94504AFFFDC2C0FAB5FB89FAC9" box="[907,1033,1310,1331]" class="Agaricomycetes" family="Psathyrellaceae" genus="Drosophila" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="90" phylum="Basidiomycota" rank="genus">
<emphasis id="85FA4B05504AFFFDC2C0FAB5FB89FAC9" box="[907,1033,1310,1331]" italics="true" pageId="7" pageNumber="90">Drosophila</emphasis>
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rearing medium as a food source, but tended to display little interest in eating it. After several days,
<specimenCount id="A1885C9E504AFFFDC280FAF7FBB5FA8B" box="[971,1077,1371,1393]" pageId="7" pageNumber="90" type="larva">one larva</specimenCount>
began to eat
<taxonomicName id="708EEC94504AFFFDC585FAF0FACCFA8A" box="[1230,1356,1371,1392]" class="Agaricomycetes" family="Psathyrellaceae" genus="Drosophila" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="90" phylum="Basidiomycota" rank="genus">
<emphasis id="85FA4B05504AFFFDC585FAF0FACCFA8A" box="[1230,1356,1371,1392]" italics="true" pageId="7" pageNumber="90">Drosophila</emphasis>
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adults before finally pupating. No other immature stage of any other species of
<taxonomicName id="708EEC94504AFFFDC561FA32FB2FFA54" authorityName="Kraatz" authorityYear="1857" box="[1066,1199,1433,1454]" class="Insecta" family="Staphylinidae" genus="Hoplandria" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="85FA4B05504AFFFDC561FA32FB2FFA54" box="[1066,1199,1433,1454]" italics="true" pageId="7" pageNumber="90">Hoplandria</emphasis>
</taxonomicName>
has been described and the mating behaviour of adults remains unknown.
</paragraph>
<paragraph id="B7319717504AFFF2C278FA40FE47FCB5" blockId="7.[819,1434,1514,1905]" lastBlockId="8.[142,757,794,847]" lastPageId="8" lastPageNumber="91" pageId="7" pageNumber="90">
<emphasis id="85FA4B05504AFFFDC278FA40FBEFFA05" box="[819,1135,1514,1536]" italics="true" pageId="7" pageNumber="90">
Comments:
<taxonomicName id="708EEC94504AFFFDC2F5FA41FBC3FA05" authorityName="Kraatz" authorityYear="1857" box="[958,1091,1514,1535]" class="Insecta" family="Staphylinidae" genus="Hoplandria" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="90" phylum="Arthropoda" rank="genus" sensu="lato">Hoplandria</taxonomicName>
<taxonomicNameLabel id="9EC9F67E504AFFFDC504FA41FBEFFA05" box="[1103,1135,1514,1535]" pageId="7" pageNumber="90" sensu="lato">s.l.</taxonomicNameLabel>
</emphasis>
is a taxonomically huge genus with hundreds or even thousands of undescribed species especially from the Neotropical Region; 95 species are currently described. Even though some species of
<taxonomicName id="708EEC94504AFFFDC2E1F9CEFBAFF980" authorityName="Kraatz" authorityYear="1857" box="[938,1071,1637,1658]" class="Insecta" family="Staphylinidae" genus="Hoplandria" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="85FA4B05504AFFFDC2E1F9CEFBAFF980" box="[938,1071,1637,1658]" italics="true" pageId="7" pageNumber="90">Hoplandria</emphasis>
</taxonomicName>
are described from outside the New World, my examination of many of these species indicates that they represent other genera, or other aleocharine tribes in some instances. The morphological variation represented within
<emphasis id="85FA4B05504AFFFDC582F94BFAFAF90F" box="[1225,1402,1760,1781]" italics="true" pageId="7" pageNumber="90">
<taxonomicName id="708EEC94504AFFFDC582F94BFACEF90F" authorityName="Kraatz" authorityYear="1857" box="[1225,1358,1760,1781]" class="Insecta" family="Staphylinidae" genus="Hoplandria" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="90" phylum="Arthropoda" rank="genus" sensu="lato">Hoplandria</taxonomicName>
<taxonomicNameLabel id="9EC9F67E504AFFFDC411F94BFAFAF90F" box="[1370,1402,1760,1781]" pageId="7" pageNumber="90" sensu="lato">s.l.</taxonomicNameLabel>
</emphasis>
is extremely variable, especially the secondary sexual features in the males. The function of these features is unknown; however, it seems likely that many structures are used in either male–male competition or resource guarding, while others are associated with glands or outlets of glands.
</paragraph>
<table id="C58E65B750450005C1C6FF68FA0EFD31" box="[141,1422,195,715]" pageId="8" pageNumber="91">
<paragraph id="B73197175045FFF2C092FF79FAFCFEB8" blockId="8.[473,1087,210,234]" lastBlockId="8.[156,1404,252,703]" pageId="8" pageNumber="91">
KEY TO THE SUBGENERA OF THE
<collectionCode id="D19F0FD25045FFF2C21CFF7DFC23FF13" box="[855,931,214,233]" pageId="8" pageNumber="91">GENUS</collectionCode>
<taxonomicName id="708EEC945045FFF2C2E0FF78FBBFFF13" authorityName="Kraatz" authorityYear="1857" box="[939,1087,211,234]" class="Insecta" family="Staphylinidae" genus="Hoplandria" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="91" phylum="Arthropoda" rank="genus">
<emphasis id="85FA4B055045FFF2C2E0FF78FBBFFF13" box="[939,1087,211,234]" italics="true" pageId="8" pageNumber="91">HOPLANDRIA</emphasis>
</taxonomicName>
<keyLead id="0C7F34225045FFF2C1D7FF57FAFCFEB8" pageId="8" pageNumber="91">
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<key id="C31FC0265045FFF2C1D7FF57FAFCFEB8" pageId="8" pageNumber="91">
<emphasis id="85FA4B055045FFF2C1D7FF57FF2FFEF5" bold="true" box="[156,175,252,271]" pageId="8" pageNumber="91">1.</emphasis>
Pronotum scarcely to moderately pubescent, often with bare longitudinal area on disc (
<figureCitation id="2FB58B925045FFF2C521FF57FB44FEF5" box="[1130,1220,252,271]" captionStart="Figure 10" captionStartId="21.[155,233,1778,1797]" captionTargetBox="[158,1443,205,1739]" captionTargetId="figure-60@21.[154,1446,193,1750]" captionTargetPageId="21" captionText="Figure 10. Hoplandria (Platonica) major (Sharp). (A) Habitus, dorsal view. (B) Labrum. (C) Epipharynx. (D) Mandible, dorsal view. (E) Mandible, ventral view. (F) Maxilla. (G) Labium. (H) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437814" httpUri="https://zenodo.org/record/5437814/files/figure.png" pageId="8" pageNumber="91">Fig. 10A</figureCitation>
); pronotum most often with 6 macrosetae on disc (
<figureCitation id="2FB58B925045FFF2C398FEBEFCADFED2" box="[723,813,277,296]" captionStart="Figure 10" captionStartId="21.[155,233,1778,1797]" captionTargetBox="[158,1443,205,1739]" captionTargetId="figure-60@21.[154,1446,193,1750]" captionTargetPageId="21" captionText="Figure 10. Hoplandria (Platonica) major (Sharp). (A) Habitus, dorsal view. (B) Labrum. (C) Epipharynx. (D) Mandible, dorsal view. (E) Mandible, ventral view. (F) Maxilla. (G) Labium. (H) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437814" httpUri="https://zenodo.org/record/5437814/files/figure.png" pageId="8" pageNumber="91">Fig. 10A</figureCitation>
); major males with humeral carina on each elytron (
<figureCitation id="2FB58B925045FFF2C016FE85FE35FEB8" box="[349,437,302,322]" captionStart="Figure 10" captionStartId="21.[155,233,1778,1797]" captionTargetBox="[158,1443,205,1739]" captionTargetId="figure-60@21.[154,1446,193,1750]" captionTargetPageId="21" captionText="Figure 10. Hoplandria (Platonica) major (Sharp). (A) Habitus, dorsal view. (B) Labrum. (C) Epipharynx. (D) Mandible, dorsal view. (E) Mandible, ventral view. (F) Maxilla. (G) Labium. (H) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437814" httpUri="https://zenodo.org/record/5437814/files/figure.png" pageId="8" pageNumber="91">Fig. 10A</figureCitation>
)..................................................................................................................................................... 2
</key>
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</keyLead>
</paragraph>
<paragraph id="B73197175045FFF2C1D7FEE3FAFCFE74" blockId="8.[156,1404,252,703]" pageId="8" pageNumber="91">
<keyLead id="0C7F34225045FFF2C1D7FEE3FAFCFE74" pageId="8" pageNumber="91">
<emphasis id="85FA4B055045FFF2C1D7FEE3FF35FEA1" bold="true" box="[156,181,328,347]" pageId="8" pageNumber="91">1¢.</emphasis>
Pronotum moderately to densely pubescent, pubsence generally evenly distributed (
<figureCitation id="2FB58B925045FFF2C50DFEE3FB13FEA1" box="[1094,1171,328,347]" captionStart="Figure 6" captionStartId="15.[163,241,1761,1780]" captionTargetBox="[158,1440,200,1725]" captionTargetId="figure-61@15.[154,1449,191,1734]" captionTargetPageId="15" captionText="Figure 6. Hoplandria (Hoplandria) lateralis Melsheimer. (A) Habitus, dorsal view. (B) Labrum. (C) Epipharynx. (D) Mandible, dorsal view. (E) Mandible, ventral view. (F) Maxilla. (G) Labium. (H) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437804" httpUri="https://zenodo.org/record/5437804/files/figure.png" pageId="8" pageNumber="91">Fig. 6A</figureCitation>
); disc of pronotum typically wthout macrosetae (
<figureCitation id="2FB58B925045FFF2C3F4FECAFC8CFE8E" box="[703,780,353,372]" captionStart="Figure 6" captionStartId="15.[163,241,1761,1780]" captionTargetBox="[158,1440,200,1725]" captionTargetId="figure-61@15.[154,1449,191,1734]" captionTargetPageId="15" captionText="Figure 6. Hoplandria (Hoplandria) lateralis Melsheimer. (A) Habitus, dorsal view. (B) Labrum. (C) Epipharynx. (D) Mandible, dorsal view. (E) Mandible, ventral view. (F) Maxilla. (G) Labium. (H) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437804" httpUri="https://zenodo.org/record/5437804/files/figure.png" pageId="8" pageNumber="91">Fig. 6A</figureCitation>
); major males without humeral carina on elytra (
<figureCitation id="2FB58B925045FFF2C071FED1FE05FE77" box="[314,389,378,397]" captionStart="Figure 6" captionStartId="15.[163,241,1761,1780]" captionTargetBox="[158,1440,200,1725]" captionTargetId="figure-61@15.[154,1449,191,1734]" captionTargetPageId="15" captionText="Figure 6. Hoplandria (Hoplandria) lateralis Melsheimer. (A) Habitus, dorsal view. (B) Labrum. (C) Epipharynx. (D) Mandible, dorsal view. (E) Mandible, ventral view. (F) Maxilla. (G) Labium. (H) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437804" httpUri="https://zenodo.org/record/5437804/files/figure.png" pageId="8" pageNumber="91">Fig. 6A</figureCitation>
) ............................................................................................................................................................ 3
</keyLead>
</paragraph>
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<paragraph id="B73197175045FFF2C1D7FE3FFAFCFE5D" blockId="8.[156,1404,252,703]" box="[156,1404,404,423]" pageId="8" pageNumber="91">
<keyLead id="0C7F34225045FFF2C1D7FE3FFAFCFE5D" box="[156,1404,404,423]" pageId="8" pageNumber="91">
<emphasis id="85FA4B055045FFF2C1D7FE3FFF2FFE5D" bold="true" box="[156,175,404,423]" pageId="8" pageNumber="91">2.</emphasis>
Bulb of aedeagus shorter than tube (
<figureCitation id="2FB58B925045FFF2C319FE3FFD1EFE5D" box="[594,670,404,423]" captionStart="Figure 9" captionStartId="19.[155,233,1370,1389]" captionTargetBox="[167,1437,210,1332]" captionTargetId="figure-265@19.[155,1453,191,1341]" captionTargetPageId="19" captionText="Figure 9. Hoplandria (Lophomucter) laevicollis (Notman). (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Aedeagus, lateral view. (D) Paramere, lateral view. (E) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5437812" httpUri="https://zenodo.org/record/5437812/files/figure.png" pageId="8" pageNumber="91">Fig. 9C</figureCitation>
) ..............................................................................
<taxonomicName id="708EEC945045FFF2C5DDFE3FFAFCFE5D" authority="Notman" authorityName="Notman" authorityYear="1920" box="[1174,1404,404,423]" class="Insecta" family="Staphylinidae" genus="Lophomucter" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="91" phylum="Arthropoda" rank="genus">
<emphasis id="85FA4B055045FFF2C5DDFE3FFA9FFE5D" box="[1174,1311,404,423]" italics="true" pageId="8" pageNumber="91">Lophomucter</emphasis>
Notman
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B73197175045FFF2C1D7FE06FAFCFE3A" blockId="8.[156,1404,252,703]" box="[156,1404,429,449]" pageId="8" pageNumber="91">
<keyLead id="0C7F34225045FFF2C1D7FE06FAFCFE3A" box="[156,1404,429,449]" pageId="8" pageNumber="91">
<emphasis id="85FA4B055045FFF2C1D7FE06FF35FE3A" bold="true" box="[156,181,429,448]" pageId="8" pageNumber="91">2¢.</emphasis>
Bulb of aedeagus subequal to or longer than, tube (
<figureCitation id="2FB58B925045FFF2C3A6FE06FCC5FE3A" box="[749,837,429,448]" captionStart="Figure 11" captionStartId="22.[142,220,1298,1317]" captionTargetBox="[148,1421,210,1263]" captionTargetId="figure-328@22.[133,1431,194,1271]" captionTargetPageId="22" captionText="Figure 11. Hoplandria (Platonica) major (Sharp). (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Aedeagus, lateral view. (D) Paramere, lateral view. (E) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5437816" httpUri="https://zenodo.org/record/5437816/files/figure.png" pageId="8" pageNumber="91">Fig. 11C</figureCitation>
) ............................................................
<taxonomicName id="708EEC945045FFF2C59AFE06FAFCFE3A" authority="Sharp" authorityName="Sharp" authorityYear="1883" box="[1233,1404,429,448]" class="Insecta" family="Staphylinidae" genus="Platonica" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="91" phylum="Arthropoda" rank="genus">
<emphasis id="85FA4B055045FFF2C59AFE06FAB4FE3A" box="[1233,1332,429,448]" italics="true" pageId="8" pageNumber="91">Platonica</emphasis>
Sharp
</taxonomicName>
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</paragraph>
</keyStep>
<keyStep id="0C7A8FB25045FFF2C1D7FE6DFAFCFDDC" pageId="8" pageNumber="91">
<paragraph id="B73197175045FFF2C1D7FE6DFAFCFE09" blockId="8.[156,1404,252,703]" pageId="8" pageNumber="91">
<keyLead id="0C7F34225045FFF2C1D7FE6DFAFCFE09" pageId="8" pageNumber="91">
<emphasis id="85FA4B055045FFF2C1D7FE6DFF2FFE23" bold="true" box="[156,175,454,473]" pageId="8" pageNumber="91">3.</emphasis>
Tergum
<collectionCode id="D19F0FD25045FFF2C06EFE6CFEC1FE20" box="[293,321,455,474]" pageId="8" pageNumber="91">III</collectionCode>
3.0–4.5 times wider than length along midline (
<figureCitation id="2FB58B925045FFF2C208FE6DFC11FE23" box="[835,913,454,473]" captionStart="Figure 6" captionStartId="15.[163,241,1761,1780]" captionTargetBox="[158,1440,200,1725]" captionTargetId="figure-61@15.[154,1449,191,1734]" captionTargetPageId="15" captionText="Figure 6. Hoplandria (Hoplandria) lateralis Melsheimer. (A) Habitus, dorsal view. (B) Labrum. (C) Epipharynx. (D) Mandible, dorsal view. (E) Mandible, ventral view. (F) Maxilla. (G) Labium. (H) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437804" httpUri="https://zenodo.org/record/5437804/files/figure.png" pageId="8" pageNumber="91">Fig. 6A</figureCitation>
); discs of terga IV–
<collectionCode id="D19F0FD25045FFF2C515FE6CFBF8FE20" box="[1118,1144,455,474]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="8" pageNumber="91">VI</collectionCode>
typically pubescent with fine round punctures (
<figureCitation id="2FB58B925045FFF2C32AFE4BFD2EFE09" box="[609,686,480,499]" captionStart="Figure 6" captionStartId="15.[163,241,1761,1780]" captionTargetBox="[158,1440,200,1725]" captionTargetId="figure-61@15.[154,1449,191,1734]" captionTargetPageId="15" captionText="Figure 6. Hoplandria (Hoplandria) lateralis Melsheimer. (A) Habitus, dorsal view. (B) Labrum. (C) Epipharynx. (D) Mandible, dorsal view. (E) Mandible, ventral view. (F) Maxilla. (G) Labium. (H) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437804" httpUri="https://zenodo.org/record/5437804/files/figure.png" pageId="8" pageNumber="91">Fig. 6A</figureCitation>
); body typically yellowish to light brown ...................
<taxonomicName id="708EEC945045FFF2C592FE4BFAD3FE09" authorityName="Kraatz" authorityYear="1857" box="[1241,1363,480,499]" class="Insecta" family="Staphylinidae" genus="Hoplandria" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="91" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="85FA4B055045FFF2C592FE4BFAD3FE09" box="[1241,1363,480,499]" italics="true" pageId="8" pageNumber="91">Hoplandria</emphasis>
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<emphasis id="85FA4B055045FFF2C410FE4BFAE5FE08" box="[1371,1381,480,498]" italics="true" pageId="8" pageNumber="91">s</emphasis>
.
<emphasis id="85FA4B055045FFF2C427FE4BFAF6FE08" box="[1388,1398,480,498]" italics="true" pageId="8" pageNumber="91">s</emphasis>
.
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</keyLead>
</paragraph>
<paragraph id="B73197175045FFF2C1D7FE52FAFCFDDC" blockId="8.[156,1404,252,703]" pageId="8" pageNumber="91">
<keyLead id="0C7F34225045FFF2C1D7FE52FAFCFDDC" pageId="8" pageNumber="91">
<emphasis id="85FA4B055045FFF2C1D7FE52FF35FDF6" bold="true" box="[156,181,505,524]" pageId="8" pageNumber="91">3¢.</emphasis>
Tergum
<collectionCode id="D19F0FD25045FFF2C06EFE51FEC1FDF7" box="[293,321,506,525]" pageId="8" pageNumber="91">III</collectionCode>
2.0–2.5 times wider than length along midline; discs of terga IV–
<collectionCode id="D19F0FD25045FFF2C2B7FE51FB96FDF7" box="[1020,1046,506,525]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="8" pageNumber="91">VI</collectionCode>
typically bare and without fine round punctures; body usually darker, light brown to dark brown ........................................................ 4
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<paragraph id="B73197175045FFF2C1D7FD87FAFCFD88" blockId="8.[156,1404,252,703]" pageId="8" pageNumber="91">
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<emphasis id="85FA4B055045FFF2C1D7FD87FF2FFDC5" bold="true" box="[156,175,556,575]" pageId="8" pageNumber="91">4.</emphasis>
Size generally small (
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); general body shape narrowly elongate; major males with a single prominent denticle on outer third of each elytron (
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), without posterolateral carina (
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); abdominal tergum IV unmodified ......................................................................................
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Notman
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<keyLead id="0C7F34225045FFF2C1D7FDD2FAFBFD45" pageId="8" pageNumber="91">
<emphasis id="85FA4B055045FFF2C1D7FDD2FF35FD76" bold="true" box="[156,181,633,652]" pageId="8" pageNumber="91">4¢.</emphasis>
Size large (3.5–5.0 mm); general body shape fusiform; major males with a prominent denticle on apical area along the midline of each elytron (
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), and with a posterolateral carina on each elytron (
<figureCitation id="2FB58B925045FFF2C06EFD07FEEFFD45" box="[293,367,684,703]" captionStart="Figure 3" captionStartId="10.[142,220,1355,1374]" captionTargetBox="[142,1420,198,1318]" captionTargetId="figure-275@10.[142,1421,196,1322]" captionTargetPageId="10" captionText="Figure 3. Hoplandria (Arrhenandria) laeviventris Casey. (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Aedeagus, lateral view. (D) Paramere, lateral view. (E) Spermatheca. (F) Left elytron, male (arrow to denticle)." figureDoi="http://doi.org/10.5281/zenodo.5437798" httpUri="https://zenodo.org/record/5437798/files/figure.png" pageId="8" pageNumber="91">Fig. 3F</figureCitation>
); abdominal tergum IV with a prominent raised area along midline ..............
<emphasis id="85FA4B055045FFF2C5D0FD07FAACFD45" box="[1179,1324,684,703]" italics="true" pageId="8" pageNumber="91">Arrhenandria</emphasis>
Génier
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,
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(
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TAL
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[People’s Republic] (
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[Fukien],
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</paragraph>
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SUBGENUS
<emphasis id="85FA4B055045FFF2C01DFA6BFD8AFA2D" box="[342,522,1472,1496]" italics="true" pageId="8" pageNumber="91">ARRHENANDRIA</emphasis>
<bibRefCitation id="D31FEAE65045FFF2C359FA6BFD31FA22" author="Genier F" box="[530,689,1472,1496]" pageId="8" pageNumber="91" pagination="3 - 59" refId="ref37573" refString="Genier F. 1989. A revision of the genus Hoplandria Kraatz of America north of Mexico (Coleoptera: Staphylinidae, Aleocharinae). Memoirs of the Entomological Society of Canada 150: 3 - 59." type="journal article" year="1989">GÉNIER, 1989</bibRefCitation>
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(
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2,3)
</paragraph>
</subSubSection>
<subSubSection id="FF94C49C5045FFF2C1C5F9A2FE68F9C4" pageId="8" pageNumber="91" type="reference_group">
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<emphasis id="85FA4B055045FFF2C1C5F9A2FE4CF9E4" box="[142,460,1545,1566]" italics="true" pageId="8" pageNumber="91">Hoplandria (Arrhenandria)</emphasis>
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<bibRefCitation id="D31FEAE65045FFF2C3D4F9A2FE86F9C7" author="Hanley RS" pageId="8" pageNumber="91" pagination="301 - 321" refId="ref37747" refString="Hanley RS. 2002 a. Phylogeny and higher classification of Hoplandriini (Coleoptera: Staphylinidae: Aleocharinae). Systematic Entomology 27: 301 - 321." type="journal article" year="2002">Hanley, 2002a:304</bibRefCitation>
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. Hanley, in press b.
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<paragraph id="B73197175045FFF2C1C5F9FCFDFAF970" blockId="8.[142,758,1622,1675]" pageId="8" pageNumber="91">
<emphasis id="85FA4B055045FFF2C1C5F9FCFE9FF996" box="[142,287,1623,1644]" italics="true" pageId="8" pageNumber="91">
<typeStatus id="683529B55045FFF2C1C5F9FCFF43F996" box="[142,195,1623,1644]" pageId="8" pageNumber="91">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="708EEC945045FFF2C063F9FDFD76F991" authority="Casey, 1910" authorityName="Casey" authorityYear="1910" box="[296,758,1622,1643]" class="Insecta" family="Staphylinidae" genus="Hoplandria" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="91" phylum="Arthropoda" rank="species" species="laeviventris">
<emphasis id="85FA4B055045FFF2C063F9FDFDC8F991" box="[296,584,1622,1643]" italics="true" pageId="8" pageNumber="91">Hoplandria laeviventris</emphasis>
<bibRefCitation id="D31FEAE65045FFF2C32BF9FDFD76F991" author="Casey TL" box="[608,758,1622,1643]" pageId="8" pageNumber="91" pagination="1 - 183" refId="ref37202" refString="Casey TL. 1910. New species of the staphylinid tribe Myrmedoniini. Memoirs on the Coleoptera 1: 1 - 183." type="journal article" year="1910">Casey, 1910</bibRefCitation>
</taxonomicName>
(designated by
<bibRefCitation id="D31FEAE65045FFF2C075F9DEFE4FF970" author="Genier F" box="[318,463,1653,1675]" pageId="8" pageNumber="91" pagination="3 - 59" refId="ref37573" refString="Genier F. 1989. A revision of the genus Hoplandria Kraatz of America north of Mexico (Coleoptera: Staphylinidae, Aleocharinae). Memoirs of the Entomological Society of Canada 150: 3 - 59." type="journal article" year="1989">Génier, 1989</bibRefCitation>
by monotypy).
</paragraph>
<paragraph id="B73197175045FFF2C1C5F908FA81FC97" blockId="8.[142,758,1699,1905]" lastBlockId="8.[806,1421,794,877]" pageId="8" pageNumber="91">
<emphasis id="85FA4B055045FFF2C1C5F908FE85F942" box="[142,261,1699,1720]" italics="true" pageId="8" pageNumber="91">Diagnosis:</emphasis>
This subgenus is distinguishable from the other subgenera of
<taxonomicName id="708EEC945045FFF2C03BF96AFE75F92C" authorityName="Kraatz" authorityYear="1857" box="[368,501,1729,1750]" class="Insecta" family="Staphylinidae" genus="Hoplandria" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="91" phylum="Arthropoda" rank="genus">
<emphasis id="85FA4B055045FFF2C03BF96AFE75F92C" box="[368,501,1729,1750]" italics="true" pageId="8" pageNumber="91">Hoplandria</emphasis>
</taxonomicName>
by the following combination of characters: body moderate to large (3.5–5.0 mm); body colouration dark throughout; punctation on abdominal terga
<collectionCode id="D19F0FD25045FFF2C0BDF8B5FD92F8C9" box="[502,530,1822,1843]" pageId="8" pageNumber="91">III</collectionCode>
–
<collectionCode id="D19F0FD25045FFF2C36AF8B5FDBEF8C9" box="[545,574,1822,1843]" pageId="8" pageNumber="91">IV</collectionCode>
with lanceolate ridges impressed laterally and more or less widely separated; males with elytra with prominent denticles on the medioapical margins and prominent posterolateral carina; abdominal tergum
<collectionCode id="D19F0FD25045FFF2C5DFFC92FB30FCB4" box="[1172,1200,825,846]" pageId="8" pageNumber="91">IV</collectionCode>
of males with distinct swelling or flat iridescent elevation.
</paragraph>
<paragraph id="B73197175045FFF2C26DFC2DFB49FB8B" blockId="8.[806,1422,901,1138]" pageId="8" pageNumber="91">
<emphasis id="85FA4B055045FFF2C26DFC2DFC2EFC61" box="[806,942,902,923]" italics="true" pageId="8" pageNumber="91">Description:</emphasis>
In agreement with
<taxonomicName id="708EEC945045FFF2C5E8FC2EFAA8FC60" authorityName="Kraatz" authorityYear="1857" box="[1187,1320,901,922]" class="Insecta" family="Staphylinidae" genus="Hoplandria" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="91" phylum="Arthropoda" rank="genus">
<emphasis id="85FA4B055045FFF2C5E8FC2EFAA8FC60" box="[1187,1320,901,922]" italics="true" pageId="8" pageNumber="91">Hoplandria</emphasis>
</taxonomicName>
description, except for the following characters. [1] Lengths of adults 3.5–5.0 mm. [2] Body elongate with sides broadly arcuate; [3] surface more or less glossy with [4] prominent punctation and microsculpturing; [5] pubescence moderately frequent on head and [6] abdominal terga, more or less dense on [7] pronotum, [8] elytra, and [9] abdominal sterna.
</paragraph>
<paragraph id="B73197175045FFF2C26DFB21FB27FAE0" blockId="8.[806,1422,1161,1307]" pageId="8" pageNumber="91">
<emphasis id="85FA4B055045FFF2C26DFB21FCEAFB65" box="[806,874,1162,1183]" italics="true" pageId="8" pageNumber="91">Head:</emphasis>
[10] Slightly broader than long. [11] Eyes moderate in size, 0.3–0.5 times length of head. [13] Infraorbital carina weakly developed. Antenna with [16] article 4 slightly longer than wide; [18] coeloconical sensilla present in article 11.
</paragraph>
<paragraph id="B73197175045FFF0C26DFA98FD86F9A6" blockId="8.[806,1422,1331,1905]" lastBlockId="10.[142,758,1453,1629]" lastPageId="10" lastPageNumber="93" pageId="8" pageNumber="91">
<emphasis id="85FA4B055045FFF2C26DFA98FC32FAB2" box="[806,946,1331,1352]" italics="true" pageId="8" pageNumber="91">Mouthparts:</emphasis>
Labrum (
<figureCitation id="2FB58B925045FFF2C57DFA98FB09FAB3" box="[1078,1161,1331,1353]" captionStart="Figure 2" captionStartId="9.[155,233,1736,1755]" captionTargetBox="[165,1419,207,1701]" captionTargetId="figure-52@9.[155,1435,194,1715]" captionTargetPageId="9" captionText="Figure 2. Hoplandria (Arrenhandria) laeviventris Casey. (A) Labrum. (B) Epipharynx. (C) Mandible, dorsal view. (D) Mandible, ventral view. (E) Maxilla. (F) Labium. (G) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437796" httpUri="https://zenodo.org/record/5437796/files/figure.png" pageId="8" pageNumber="91">Fig. 2A</figureCitation>
) with epipharyngeal area with [19] medial pores large, numerous; [21] pores between longitudinal sensory field and lateral sclerotized areas few, irregularly arranged (
<figureCitation id="2FB58B925045FFF2C466FA24FA01FA5F" box="[1325,1409,1423,1445]" captionStart="Figure 2" captionStartId="9.[155,233,1736,1755]" captionTargetBox="[165,1419,207,1701]" captionTargetId="figure-52@9.[155,1435,194,1715]" captionTargetPageId="9" captionText="Figure 2. Hoplandria (Arrenhandria) laeviventris Casey. (A) Labrum. (B) Epipharynx. (C) Mandible, dorsal view. (D) Mandible, ventral view. (E) Maxilla. (F) Labium. (G) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437796" httpUri="https://zenodo.org/record/5437796/files/figure.png" pageId="8" pageNumber="91">Fig. 2B</figureCitation>
). Mandible (
<figureCitation id="2FB58B925045FFF2C2E1FA06FB95FA39" box="[938,1045,1453,1475]" captionStart="Figure 2" captionStartId="9.[155,233,1736,1755]" captionTargetBox="[165,1419,207,1701]" captionTargetId="figure-52@9.[155,1435,194,1715]" captionTargetPageId="9" captionText="Figure 2. Hoplandria (Arrenhandria) laeviventris Casey. (A) Labrum. (B) Epipharynx. (C) Mandible, dorsal view. (D) Mandible, ventral view. (E) Maxilla. (F) Labium. (G) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437796" httpUri="https://zenodo.org/record/5437796/files/figure.png" pageId="8" pageNumber="91">Fig. 2C,D</figureCitation>
) with [23] apex more or less rounded with slight curve adorally; condylar molar patch with [26] densely, irregularly arranged denticles; dorsobasal ‘velvety patch’ [28,29] absent; [30] ventral aspect with outer basal angle with two small setae; [33] teeth of adoral margin of lacinia short, each about 3 times longer than wide. Maxilla (
<figureCitation id="2FB58B925045FFF2C5B1F9CEFACCF981" box="[1274,1356,1637,1659]" captionStart="Figure 2" captionStartId="9.[155,233,1736,1755]" captionTargetBox="[165,1419,207,1701]" captionTargetId="figure-52@9.[155,1435,194,1715]" captionTargetPageId="9" captionText="Figure 2. Hoplandria (Arrenhandria) laeviventris Casey. (A) Labrum. (B) Epipharynx. (C) Mandible, dorsal view. (D) Mandible, ventral view. (E) Maxilla. (F) Labium. (G) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437796" httpUri="https://zenodo.org/record/5437796/files/figure.png" pageId="8" pageNumber="91">Fig. 2E</figureCitation>
) with teeth on adoral margin divided into [35] three more or less distinct regions; [37] galea moderately narrow, subequal to width of lacinea at widest area, and rounded apically. Labium (
<figureCitation id="2FB58B925045FFF2C514F94BFB47F90C" box="[1119,1223,1760,1782]" captionStart="Figure 2" captionStartId="9.[155,233,1736,1755]" captionTargetBox="[165,1419,207,1701]" captionTargetId="figure-52@9.[155,1435,194,1715]" captionTargetPageId="9" captionText="Figure 2. Hoplandria (Arrenhandria) laeviventris Casey. (A) Labrum. (B) Epipharynx. (C) Mandible, dorsal view. (D) Mandible, ventral view. (E) Maxilla. (F) Labium. (G) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437796" httpUri="https://zenodo.org/record/5437796/files/figure.png" pageId="8" pageNumber="91">Fig. 2F,G</figureCitation>
) with ligula [45] elongate, longer than labial palpus 1 + 2, with [47] moderately forked apex; [49] two long medial setae of prementum present, typically longer length of ligula, [50] insertions of setae almost adjacent. Labial palpus (
<figureCitation id="2FB58B925047FFF0C1DDFA06FF67FA39" box="[150,231,1453,1475]" captionStart="Figure 2" captionStartId="9.[155,233,1736,1755]" captionTargetBox="[165,1419,207,1701]" captionTargetId="figure-52@9.[155,1435,194,1715]" captionTargetPageId="9" captionText="Figure 2. Hoplandria (Arrenhandria) laeviventris Casey. (A) Labrum. (B) Epipharynx. (C) Mandible, dorsal view. (D) Mandible, ventral view. (E) Maxilla. (F) Labium. (G) Hypopharynx." figureDoi="http://doi.org/10.5281/zenodo.5437796" httpUri="https://zenodo.org/record/5437796/files/figure.png" pageId="10" pageNumber="93">Fig. 2F</figureCitation>
) [58] elongate, overall length 5–6 times longer than greatest width, [60] with distal pore field composed of two large pores and small sensory pegs; [61] article 1 about 4.0 times longer than article 2, [62] article 3 about 3.0 times longer than article 2.
<collectingRegion id="754A59F55047FFF0C3DFF983FD75F9C7" box="[660,757,1576,1597]" country="Myanmar" name="Chin" pageId="10" pageNumber="93">Mentum</collectingRegion>
with [65] apical margin concave.
</paragraph>
<caption id="E3F1C79F5044FFF3C1D0F963FCE0F902" ID-DOI="http://doi.org/10.5281/zenodo.5437796" ID-Zenodo-Dep="5437796" httpUri="https://zenodo.org/record/5437796/files/figure.png" pageId="9" pageNumber="92" startId="9.[155,233,1736,1755]" targetBox="[165,1419,207,1701]" targetPageId="9">
<paragraph id="B73197175044FFF3C1D0F963FCE0F902" blockId="9.[155,1434,1736,1784]" pageId="9" pageNumber="92">
<emphasis id="85FA4B055044FFF3C1D0F963FE83F921" bold="true" box="[155,259,1736,1755]" pageId="9" pageNumber="92">Figure 2.</emphasis>
<taxonomicName id="708EEC945044FFF3C045F963FC85F921" authority="Casey." authorityName="Casey." box="[270,773,1736,1755]" class="Insecta" family="Staphylinidae" genus="Hoplandria" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="92" phylum="Arthropoda" rank="species" species="laeviventris" subGenus="Arrenhandria">
<emphasis id="85FA4B055044FFF3C045F963FE08F921" box="[270,392,1736,1755]" italics="true" pageId="9" pageNumber="92">Hoplandria</emphasis>
(
<emphasis id="85FA4B055044FFF3C0D0F963FDACF921" box="[411,556,1736,1755]" italics="true" pageId="9" pageNumber="92">Arrenhandria</emphasis>
)
<emphasis id="85FA4B055044FFF3C375F963FD37F921" box="[574,695,1736,1755]" italics="true" pageId="9" pageNumber="92">laeviventris</emphasis>
Casey.
</taxonomicName>
(A) Labrum. (B) Epipharynx. (C) Mandible, dorsal view. (D) Mandible, ventral view. (E) Maxilla. (F) Labium. (G) Hypopharynx.
</paragraph>
</caption>
<caption id="E3F1C79F5047FFF0C1C5FAE0FB6BFA81" ID-DOI="http://doi.org/10.5281/zenodo.5437798" ID-Zenodo-Dep="5437798" httpUri="https://zenodo.org/record/5437798/files/figure.png" pageId="10" pageNumber="93" startId="10.[142,220,1355,1374]" targetBox="[142,1420,198,1318]" targetPageId="10">
<paragraph id="B73197175047FFF0C1C5FAE0FB6BFA81" blockId="10.[142,1422,1355,1403]" pageId="10" pageNumber="93">
<emphasis id="85FA4B055047FFF0C1C5FAE0FF76FAA4" bold="true" box="[142,246,1355,1374]" pageId="10" pageNumber="93">Figure 3.</emphasis>
<taxonomicName id="708EEC945047FFF0C049FAE0FD69FAA4" authority="Casey." authorityName="Casey." box="[258,745,1355,1374]" class="Insecta" family="Staphylinidae" genus="Hoplandria" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="93" phylum="Arthropoda" rank="species" species="laeviventris" subGenus="Arrhenandria">
<emphasis id="85FA4B055047FFF0C049FAE0FEFCFAA4" box="[258,380,1355,1374]" italics="true" pageId="10" pageNumber="93">Hoplandria</emphasis>
(
<emphasis id="85FA4B055047FFF0C0C2FAE0FD9AFAA4" box="[393,538,1355,1374]" italics="true" pageId="10" pageNumber="93">Arrhenandria</emphasis>
)
<emphasis id="85FA4B055047FFF0C36CFAE0FD20FAA4" box="[551,672,1355,1374]" italics="true" pageId="10" pageNumber="93">laeviventris</emphasis>
Casey.
</taxonomicName>
(A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Aedeagus, lateral view. (D) Paramere, lateral view. (E) Spermatheca. (F) Left elytron, male (arrow to denticle).
</paragraph>
</caption>
<paragraph id="B73197175047FFF0C1C5F92FFBF8F9C4" blockId="10.[142,758,1667,1905]" lastBlockId="10.[806,1422,1453,1598]" pageId="10" pageNumber="93">
<emphasis id="85FA4B055047FFF0C1C5F92FFF66F963" box="[142,230,1668,1689]" italics="true" pageId="10" pageNumber="93">Thorax:</emphasis>
Pronotum [68] distinctly transverse. Elytra with [75] apico-lateral angles distinctly sinuate; [76] elytra together about 1.3 times wider than long. Mesocoxal cavities (
<figureCitation id="2FB58B925047FFF0C07EF94BFE07F90C" box="[309,391,1760,1782]" captionStart="Figure 3" captionStartId="10.[142,220,1355,1374]" captionTargetBox="[142,1420,198,1318]" captionTargetId="figure-275@10.[142,1421,196,1322]" captionTargetPageId="10" captionText="Figure 3. Hoplandria (Arrhenandria) laeviventris Casey. (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Aedeagus, lateral view. (D) Paramere, lateral view. (E) Spermatheca. (F) Left elytron, male (arrow to denticle)." figureDoi="http://doi.org/10.5281/zenodo.5437798" httpUri="https://zenodo.org/record/5437798/files/figure.png" pageId="10" pageNumber="93">Fig. 3A</figureCitation>
) [81] widely separated by meso- and metasternal processes by about 1/4 length of coxal cavities. Mesosternal process length [82] subequal to length of metasternal process; [83] meso- and metasternal processes generally contiguous, isthmus absent; [84] mesosternal process generally rounded at apex, [85] shorter than width of mesocoxa. Macrosetae [88] present on metasternum. Legs with [93] empodial bristle distinctly shorter than tarsal claws, but over half of length of tarsal claws.
</paragraph>
<paragraph id="B73197175047FFF0C26DF9FDFBA4F932" blockId="10.[806,1422,1621,1736]" pageId="10" pageNumber="93">
<emphasis id="85FA4B055047FFF0C26DF9FDFC17F991" box="[806,919,1622,1643]" italics="true" pageId="10" pageNumber="93">Abdomen:</emphasis>
(
<figureCitation id="2FB58B925047FFF0C2E7F9FDFC7DF996" box="[940,1021,1622,1644]" captionStart="Figure 3" captionStartId="10.[142,220,1355,1374]" captionTargetBox="[142,1420,198,1318]" captionTargetId="figure-275@10.[142,1421,196,1322]" captionTargetPageId="10" captionText="Figure 3. Hoplandria (Arrhenandria) laeviventris Casey. (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Aedeagus, lateral view. (D) Paramere, lateral view. (E) Spermatheca. (F) Left elytron, male (arrow to denticle)." figureDoi="http://doi.org/10.5281/zenodo.5437798" httpUri="https://zenodo.org/record/5437798/files/figure.png" pageId="10" pageNumber="93">Fig. 3B</figureCitation>
) [100] Anterior corner of sternum
<collectionCode id="D19F0FD25047FFF0C26DF9DEFCC4F970" box="[806,836,1653,1674]" pageId="10" pageNumber="93">III</collectionCode>
with distinct region of differentiated microsculpturing, generally delimited by very fine carina, area appearing irridescent.
</paragraph>
<paragraph id="B73197175047FFF1C26DF94BFE41FDD1" blockId="10.[806,1422,1760,1905]" lastBlockId="11.[155,771,195,555]" lastPageId="11" lastPageNumber="94" pageId="10" pageNumber="93">
<emphasis id="85FA4B055047FFF0C26DF94BFB20F90F" box="[806,1184,1760,1781]" italics="true" pageId="10" pageNumber="93">Secondary sexual characteristics:</emphasis>
Males highly variable, including more or less distinct major and minor forms. Minor forms generally similar to females and typically indistinct externally. Major forms highly variable among species with morphological modifications of elytra in form of [102] without humeral carina or bump; [103] prominent denticle, carina or conical process on the medioapical region; [104] without denticle, carina or bump on the outer apical third; [104a] prominent posterolateral carina on each elytron. Abdomen: posterior corners of [105] sterna
<collectionCode id="D19F0FD25046FFF1C3E0FEF5FD49FE89" box="[683,713,350,371]" pageId="11" pageNumber="94">III</collectionCode>
and [106]
<collectionCode id="D19F0FD25046FFF1C191FED7FF76FE6B" box="[218,246,380,401]" pageId="11" pageNumber="94">IV</collectionCode>
produced into long, bent spines often reaching posterior quarter of terga
<collectionCode id="D19F0FD25046FFF1C088FE30FE5FFE4A" box="[451,479,411,432]" pageId="11" pageNumber="94">IV</collectionCode>
and
<collectionCode id="D19F0FD25046FFF1C353FE30FDA9FE4A" box="[536,553,411,432]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="11" pageNumber="94" type="Museum">V</collectionCode>
, respectively; [107] tergum
<collectionCode id="D19F0FD25046FFF1C1BEFE11FE91FE35" box="[245,273,442,463]" pageId="11" pageNumber="94">IV</collectionCode>
with more or less large swollen area; [108] tergum VII with highly elevated and acute narrow carina; [109] apex of tergum
<collectionCode id="D19F0FD25046FFF1C08AFE5CFE71FDF6" box="[449,497,503,524]" pageId="11" pageNumber="94">VIII</collectionCode>
without two transverse rows of large macrosetae.
</paragraph>
<paragraph id="B73197175046FFF1C1D0FDE8FEA1FD2E" blockId="11.[155,771,579,724]" pageId="11" pageNumber="94">
<emphasis id="85FA4B055046FFF1C1D0FDE8FE8DFDA2" box="[155,269,579,600]" italics="true" pageId="11" pageNumber="94">Aedeagus:</emphasis>
(
<figureCitation id="2FB58B925046FFF1C055FDE8FEEFFDA3" box="[286,367,579,601]" captionStart="Figure 3" captionStartId="10.[142,220,1355,1374]" captionTargetBox="[142,1420,198,1318]" captionTargetId="figure-275@10.[142,1421,196,1322]" captionTargetPageId="10" captionText="Figure 3. Hoplandria (Arrhenandria) laeviventris Casey. (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Aedeagus, lateral view. (D) Paramere, lateral view. (E) Spermatheca. (F) Left elytron, male (arrow to denticle)." figureDoi="http://doi.org/10.5281/zenodo.5437798" httpUri="https://zenodo.org/record/5437798/files/figure.png" pageId="11" pageNumber="94">Fig. 3C</figureCitation>
) Bulb of median lobe [110] more or less irregularly shaped, shorter than tube, [111] with moderately developed ventral projection; condylite (
<figureCitation id="2FB58B925046FFF1C1E9FD34FF75FD4F" box="[162,245,671,693]" captionStart="Figure 3" captionStartId="10.[142,220,1355,1374]" captionTargetBox="[142,1420,198,1318]" captionTargetId="figure-275@10.[142,1421,196,1322]" captionTargetPageId="10" captionText="Figure 3. Hoplandria (Arrhenandria) laeviventris Casey. (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Aedeagus, lateral view. (D) Paramere, lateral view. (E) Spermatheca. (F) Left elytron, male (arrow to denticle)." figureDoi="http://doi.org/10.5281/zenodo.5437798" httpUri="https://zenodo.org/record/5437798/files/figure.png" pageId="11" pageNumber="94">Fig. 3D</figureCitation>
) [117] generally subequal in length to apex of paramerite.
</paragraph>
<paragraph id="B73197175046FFF1C1D0FD47FF77FCC5" blockId="11.[155,769,747,831]" pageId="11" pageNumber="94">
<emphasis id="85FA4B055046FFF1C1D0FD47FEB6FCFB" box="[155,310,748,769]" italics="true" pageId="11" pageNumber="94">Spermatheca:</emphasis>
(
<figureCitation id="2FB58B925046FFF1C000FD47FE1CFCF8" box="[331,412,748,770]" captionStart="Figure 3" captionStartId="10.[142,220,1355,1374]" captionTargetBox="[142,1420,198,1318]" captionTargetId="figure-275@10.[142,1421,196,1322]" captionTargetPageId="10" captionText="Figure 3. Hoplandria (Arrhenandria) laeviventris Casey. (A) Meso- and metathorax. (B) Abdominal tergum VIII. (C) Aedeagus, lateral view. (D) Paramere, lateral view. (E) Spermatheca. (F) Left elytron, male (arrow to denticle)." figureDoi="http://doi.org/10.5281/zenodo.5437798" httpUri="https://zenodo.org/record/5437798/files/figure.png" pageId="11" pageNumber="94">Fig. 3E</figureCitation>
) Basal bulb [119] generally simple and rounded at base with a distinct depression at apex.
</paragraph>
<paragraph id="B73197175046FFF1C1D0FCFCFDBBFB99" blockId="11.[155,771,855,1123]" pageId="11" pageNumber="94">
<emphasis id="85FA4B055046FFF1C1D0FCFCFF79FC96" box="[155,249,855,876]" italics="true" pageId="11" pageNumber="94">Habitat:</emphasis>
Specimens of
<taxonomicName id="708EEC945046FFF1C0EAFCF3FD7BFC96" box="[417,763,855,877]" class="Insecta" family="Staphylinidae" genus="Hoplandria" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="94" phylum="Arthropoda" rank="species" species="laeviventris" subGenus="Arrhenandria">
<emphasis id="85FA4B055046FFF1C0EAFCF3FE36FC97" box="[417,438,856,877]" italics="true" pageId="11" pageNumber="94">H</emphasis>
. (
<emphasis id="85FA4B055046FFF1C081FCFCFDE8FC96" box="[458,616,855,876]" italics="true" pageId="11" pageNumber="94">Arrhenandria</emphasis>
)
<emphasis id="85FA4B055046FFF1C33CFCFCFD7BFC96" box="[631,763,855,876]" italics="true" pageId="11" pageNumber="94">laeviventris</emphasis>
</taxonomicName>
, the only species in this subgenus, have been collected from various sources including: dead mice, carrionbaited pitfall traps, and through sifting unspecified leaf litter, rhododendron litter, mushroom litter, leaf litter in deep crevices and berlese samples or leaf debris, oak-hickory debris, dry oak litter, or mixed hardwood forest litter. The collecting period for this species is from February to October.
</paragraph>
<paragraph id="B73197175046FFF1C1D0FBD0FDBCFA1B" blockId="11.[155,771,1147,1506]" pageId="11" pageNumber="94">
<emphasis id="85FA4B055046FFF1C1D0FBD0FE99FB6A" box="[155,281,1147,1168]" italics="true" pageId="11" pageNumber="94">Comments:</emphasis>
As
<bibRefCitation id="D31FEAE65046FFF1C006FBD0FE6EFB6B" author="Genier F" box="[333,494,1147,1169]" pageId="11" pageNumber="94" pagination="3 - 59" refId="ref37573" refString="Genier F. 1989. A revision of the genus Hoplandria Kraatz of America north of Mexico (Coleoptera: Staphylinidae, Aleocharinae). Memoirs of the Entomological Society of Canada 150: 3 - 59." type="journal article" year="1989">Génier (1989)</bibRefCitation>
pointed out, the establishment of
<emphasis id="85FA4B055046FFF1C06DFB32FE44FB54" box="[294,452,1177,1198]" italics="true" pageId="11" pageNumber="94">Arrhenandria</emphasis>
as a subgenus of
<taxonomicName id="708EEC945046FFF1C3DDFB32FF3CFB34" authorityName="Kraatz" authorityYear="1857" class="Insecta" family="Staphylinidae" genus="Hoplandria" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="85FA4B055046FFF1C3DDFB32FF3CFB34" italics="true" pageId="11" pageNumber="94">Hoplandria</emphasis>
</taxonomicName>
is based on the distinctive secondary sexual characters, especially the presence of a lateral carina on each elytron and a raised swelling on tergum
<collectionCode id="D19F0FD25046FFF1C3AAFB5DFD7FFAF1" box="[737,767,1270,1291]" pageId="11" pageNumber="94">IV</collectionCode>
. Both of these structures are absent in the subgenus
<taxonomicName id="708EEC945046FFF1C1D0FA98FE8FFAB2" box="[155,271,1331,1352]" class="Insecta" family="Staphylinidae" genus="Hoplandria" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="94" phylum="Arthropoda" rank="subGenus" subGenus="Genosema">
<emphasis id="85FA4B055046FFF1C1D0FA98FE8FFAB2" box="[155,271,1331,1352]" italics="true" pageId="11" pageNumber="94">Genosema</emphasis>
</taxonomicName>
, the sister group to
<emphasis id="85FA4B055046FFF1C0B1FA98FD18FAB2" box="[506,664,1331,1352]" italics="true" pageId="11" pageNumber="94">Arrhenandria</emphasis>
. In addition, the absence of microsculpure on the swollen area of tergum
<collectionCode id="D19F0FD25046FFF1C051FADBFEB8FA7F" box="[282,312,1392,1413]" pageId="11" pageNumber="94">III</collectionCode>
of the abdomen also supports a close relationship with
<taxonomicName id="708EEC945046FFF1C03AFA24FE65FA5E" box="[369,485,1423,1444]" class="Insecta" family="Staphylinidae" genus="Hoplandria" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="94" phylum="Arthropoda" rank="subGenus" subGenus="Genosema">
<emphasis id="85FA4B055046FFF1C03AFA24FE65FA5E" box="[369,485,1423,1444]" italics="true" pageId="11" pageNumber="94">Genosema</emphasis>
</taxonomicName>
.
<emphasis id="85FA4B055046FFF1C0BCFA24FD15FA5E" box="[503,661,1423,1444]" italics="true" pageId="11" pageNumber="94">Arrhenandria</emphasis>
contains only one species,
<taxonomicName id="708EEC945046FFF1C017FA05FC83FA38" authority="Casey" authorityName="Casey" box="[348,771,1453,1475]" class="Insecta" family="Staphylinidae" genus="Hoplandria" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="94" phylum="Arthropoda" rank="species" species="laeviventris" subGenus="Arrhenandria">
<emphasis id="85FA4B055046FFF1C017FA05FEF1FA39" box="[348,369,1454,1475]" italics="true" pageId="11" pageNumber="94">H</emphasis>
. (
<emphasis id="85FA4B055046FFF1C0CDFA06FDA4FA38" box="[390,548,1453,1474]" italics="true" pageId="11" pageNumber="94">Arrhenandria</emphasis>
)
<emphasis id="85FA4B055046FFF1C378FA06FD37FA38" box="[563,695,1453,1474]" italics="true" pageId="11" pageNumber="94">laeviventris</emphasis>
Casey
</taxonomicName>
from the midwestern
<collectingCountry id="CF99D7875046FFF1C0DFFA67FDB8FA1B" box="[404,568,1484,1506]" name="United States of America" pageId="11" pageNumber="94">United States</collectingCountry>
.
</paragraph>
<paragraph id="B73197175046FFF1C1D0FA51FE0FF953" blockId="11.[155,771,1530,1705]" pageId="11" pageNumber="94">
<emphasis id="85FA4B055046FFF1C1D0FA51FEACF9F5" box="[155,300,1530,1551]" italics="true" pageId="11" pageNumber="94">Distribution:</emphasis>
<collectionCode id="D19F0FD25046FFF1C072FA55FD86F9F5" box="[313,518,1534,1551]" pageId="11" pageNumber="94">NEARCTIC REGION</collectionCode>
.
<collectingCountry id="CF99D7875046FFF1C351FA51FD3DF9F5" box="[538,701,1530,1552]" name="United States of America" pageId="11" pageNumber="94">United States</collectingCountry>
(
<collectingRegion id="754A59F55046FFF1C39AFA51FF5CF9D5" country="United States of America" name="Alabama" pageId="11" pageNumber="94">Alabama</collectingRegion>
,
<collectingRegion id="754A59F55046FFF1C1A7F9B2FEDAF9D5" box="[236,346,1561,1583]" country="United States of America" name="Arkansas" pageId="11" pageNumber="94">Arkansas</collectingRegion>
,
<collectingRegion id="754A59F55046FFF1C020F9B2FE76F9D5" box="[363,502,1561,1583]" country="United States of America" name="Connecticut" pageId="11" pageNumber="94">Connecticut</collectingRegion>
,
<collectingRegion id="754A59F55046FFF1C34CF9B2FD7EF9D5" box="[519,766,1561,1583]" country="United States of America" name="District of Columbia" pageId="11" pageNumber="94">District of Columbia</collectingRegion>
,
<collectingRegion id="754A59F55046FFF1C1D0F99CFF77F9B7" box="[155,247,1591,1613]" country="United States of America" name="Georgia" pageId="11" pageNumber="94">Georgia</collectingRegion>
,
<collectingRegion id="754A59F55046FFF1C049F99CFED3F9B7" box="[258,339,1591,1613]" country="United States of America" name="Illinois" pageId="11" pageNumber="94">Illinois</collectingRegion>
,
<collectingRegion id="754A59F55046FFF1C015F99CFE39F9B7" box="[350,441,1591,1613]" country="United States of America" name="Indiana" pageId="11" pageNumber="94">Indiana</collectingRegion>
,
<collectingRegion id="754A59F55046FFF1C08EF99CFDB2F9B7" box="[453,562,1591,1613]" country="United States of America" name="Kentucky" pageId="11" pageNumber="94">Kentucky</collectingRegion>
,
<collectingRegion id="754A59F55046FFF1C375F99CFD31F9B7" box="[574,689,1591,1613]" country="United States of America" name="Louisiana" pageId="11" pageNumber="94">Louisiana</collectingRegion>
,
<collectingRegion id="754A59F55046FFF1C3F6F993FF4FF996" country="United States of America" name="Maryland" pageId="11" pageNumber="94">Maryland</collectingRegion>
,
<collectingRegion id="754A59F55046FFF1C1ABF9FDFE0AF996" box="[224,394,1622,1644]" country="United States of America" name="Massachusetts" pageId="11" pageNumber="94">Massachusetts</collectingRegion>
,
<collectingRegion id="754A59F55046FFF1C0D7F9FDFDA6F991" box="[412,550,1622,1643]" country="United States of America" name="New Jersey" pageId="11" pageNumber="94">New Jersey</collectingRegion>
,
<collectingRegion id="754A59F55046FFF1C37DF9FDFD2DF996" box="[566,685,1622,1644]" country="United States of America" name="New York" pageId="11" pageNumber="94">New York</collectingRegion>
,
<collectingRegion id="754A59F55046FFF1C3F5F9FDFE80F971" country="United States of America" name="North Carolina" pageId="11" pageNumber="94">North Carolina</collectingRegion>
,
<collectingRegion id="754A59F55046FFF1C044F9DEFEC7F971" box="[271,327,1653,1675]" country="United States of America" name="Ohio" pageId="11" pageNumber="94">Ohio</collectingRegion>
,
<collectingRegion id="754A59F55046FFF1C01DF9DEFE72F971" box="[342,498,1653,1675]" country="United States of America" name="Pennsylvania" pageId="11" pageNumber="94">Pennsylvania</collectingRegion>
,
<collectingRegion id="754A59F55046FFF1C34BF9DEFDF7F970" box="[512,631,1653,1674]" country="United States of America" name="Tennessee" pageId="11" pageNumber="94">Tennessee</collectingRegion>
,
<collectingRegion id="754A59F55046FFF1C3CDF9DEFD49F970" box="[646,713,1653,1674]" country="United States of America" name="Texas" pageId="11" pageNumber="94">Texas</collectingRegion>
,
<collectingRegion id="754A59F55046FFF1C39CF9DEFF56F953" country="United States of America" name="Virginia" pageId="11" pageNumber="94">Virginia</collectingRegion>
,
<collectingRegion id="754A59F55046FFF1C1A9F93FFE03F953" box="[226,387,1683,1705]" country="United States of America" name="West Virginia" pageId="11" pageNumber="94">West Virginia</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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