164 lines
15 KiB
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164 lines
15 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.463.8499" ID-PMC="PMC4294300" ID-Pensoft-Pub="1313-2970-463-21" ID-Pensoft-UUID="3E5CFFC6FFC91628FFF6E355951DFFF8" ID-PubMed="25589861" ID-Zenodo-Dep="578654" ID-ZooBank="71C1AE92B0CE4D26B1E5076E480180AD" ModsDocID="1313-2970-463-21" checkinTime="1451244962410" checkinUser="pensoft" docAuthor="Solodovnikov, Igor A., Zamotajlov, Alexandr S., Fominykh, Dmitriy D. & Titarenko, Andrey Y." docDate="2014" docId="3BECC0580A995D3D937CC3C8F79D6E17" docLanguage="en" docName="ZooKeys 463: 21-56" docOrigin="ZooKeys 463" docPubDate="2014-12-12" docSource="http://dx.doi.org/10.3897/zookeys.463.8499" docTitle="Carabus (Archiplectes) besleticus subsp. adzinbai Retezar 2013" docType="treatment" docVersion="3" id="3E5CFFC6FFC91628FFF6E355951DFFF8" lastPageNumber="31" masterDocId="3E5CFFC6FFC91628FFF6E355951DFFF8" masterDocTitle="Contribution to the knowledge of the Carabus (Archiplectes) satyrus Kurnakov, 1962, species complex in Abkhazia (Coleoptera, Carabidae, Carabini)" masterLastPageNumber="56" masterPageNumber="21" pageNumber="30" updateTime="1668159733340" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Contribution to the knowledge of the Carabus (Archiplectes) satyrus Kurnakov, 1962, species complex in Abkhazia (Coleoptera, Carabidae, Carabini)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Solodovnikov, Igor A.</mods:namePart>
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<mods:affiliation>Vitebsk State P. M. Masherov University, Moskovskyi prospect, 33, Vitebsk 210038, Belarus</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zamotajlov, Alexandr S.</mods:namePart>
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<mods:affiliation>Kuban State Agrarian University, Kalinin St. 13, Krasnodar 350044, Russia</mods:affiliation>
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<mods:nameIdentifier type="email">a_zamotajlov@mail.ru</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fominykh, Dmitriy D.</mods:namePart>
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<mods:affiliation>Russian Entomological Society, Kuban Branch, Kalinin St. 13, Krasnodar 350044, Russia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Titarenko, Andrey Y.</mods:namePart>
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<mods:affiliation>Publicly Traded Company " Morpho Absoloni ", Bolotnikovskaya St. 5 / 3, Moscow 117556, Russia</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2014-12-12</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>463</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>21</mods:start>
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<mods:end>56</mods:end>
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</mods:extent>
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</mods:part>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.463.8499</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.463.8499</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-463-21</mods:identifier>
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<mods:identifier type="ZooBank">71C1AE92B0CE4D26B1E5076E480180AD</mods:identifier>
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<mods:identifier type="Pensoft-UUID">3E5CFFC6FFC91628FFF6E355951DFFF8</mods:identifier>
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<mods:identifier type="Zenodo-Dep">578654</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="182234821" LSID="urn:lsid:plazi:treatment:3BECC0580A995D3D937CC3C8F79D6E17" httpUri="http://treatment.plazi.org/id/3BECC0580A995D3D937CC3C8F79D6E17" lastPageId="10" lastPageNumber="31" pageId="9" pageNumber="30">
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<subSubSection pageId="9" pageNumber="30" type="nomenclature">
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<paragraph pageId="9" pageNumber="30">
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<taxonomicName LSID="3BECC058-0A99-5D3D-937C-C3C8F79D6E17" authority="Retezar, 2013" authorityName="Retezar" authorityYear="2013" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. adzinbai" order="Coleoptera" pageId="9" pageNumber="30" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="adzinbai">
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Carabus (Archiplectes) besleticus adzinbai
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<normalizedToken originalValue="Retezár">Retezar</normalizedToken>
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, 2013
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</taxonomicName>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Distribution of morphometric characters in females of Carabus (Archiplectes) besleticus subspecies constructed using discriminant analysis based on 8 parameters." pageId="9" pageNumber="30">Figs 4</figureCitation>
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Distribution map of Carabus (Archiplectes) satyrus, Carabus (Archiplectes) besleticus, and Carabus (Archiplectes) pseudopshuensis in Abkhazia." pageId="9" pageNumber="30">, 7</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="31" type="reference_group">
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<paragraph pageId="10" pageNumber="31">
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<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) satyrus subsp. adzinbai" order="Coleoptera" pageId="10" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="satyrus" subGenus="Archiplectes" subSpecies="adzinbai">
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<pageBreakToken pageId="10" pageNumber="31" start="start">Carabus</pageBreakToken>
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(Archiplectes) satyrus adzinbai
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</taxonomicName>
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<normalizedToken originalValue="Retezár">Retezar</normalizedToken>
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, 2013: 2
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="31" type="comparative material examined">
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<paragraph pageId="10" pageNumber="31">Comparative material examined.</paragraph>
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<paragraph pageId="10" pageNumber="31">
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15 specimens were examined (12 specimens measured, 1 male genitalia preparations studied): 1 male, 2 females, Abkhazia, Bzybian Mt. Range, N slope of Mt. Akibakhu (=Turetskaya shapka), 2000 m, alpine zone, 14.VI.-09.VIII.1986, leg. A. Koval (cZAM); 1 male, 10 females, Abkhazia, Bzybian Mt. Range, N slope of Mt. Akibakhu (=Turetskaya shapka), 2000 m, alpine zone, 7-10.VII.2010, leg. D. Fominykh, A. Bondarenko (cFDD, cSIA); 1 female, Abkhazia, Gudauta distr., S slope Bzybian Mt.R., NW Mt.Akugra,
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<geoCoordinate degrees="43" direction="north" minutes="18" orientation="latitude" precision="925" value="43.3">43°18'N</geoCoordinate>
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/
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<geoCoordinate degrees="40" direction="east" minutes="43" orientation="longitude" precision="925" value="40.716667">40°43'E</geoCoordinate>
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, h = 2130-1150 m, alpine zone, 08.08.2014, leg. I. Solodovnikov, E. Tatun (cSIA).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="31" type="description">
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<paragraph pageId="10" pageNumber="31">Description.</paragraph>
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<paragraph pageId="10" pageNumber="31">
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Large form, males 31.0-32.0 mm, according to
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<bibRefCitation author="Retezar, I" journalOrPublisher="Folia Entomologica Hungarica" pageId="23" pageNumber="44" pagination="1 - 6" refId="B9" refString="Retezar, I, 2013. Taxonomic and faunistic studies on the subgenus Carabus (Archiplectes) Gottwald, 1982 in Abkhazia (Coleoptera: Carabidae). Folia Entomologica Hungarica 74: 1 - 6" title="Taxonomic and faunistic studies on the subgenus Carabus (Archiplectes) Gottwald, 1982 in Abkhazia (Coleoptera: Carabidae)." volume="74" year="2013">
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<normalizedToken originalValue="Retezár">Retezar</normalizedToken>
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(2013)
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</bibRefCitation>
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32-34 mm long, females 32.0-35.8 mm, according to
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<bibRefCitation author="Retezar, I" journalOrPublisher="Folia Entomologica Hungarica" pageId="23" pageNumber="44" pagination="1 - 6" refId="B9" refString="Retezar, I, 2013. Taxonomic and faunistic studies on the subgenus Carabus (Archiplectes) Gottwald, 1982 in Abkhazia (Coleoptera: Carabidae). Folia Entomologica Hungarica 74: 1 - 6" title="Taxonomic and faunistic studies on the subgenus Carabus (Archiplectes) Gottwald, 1982 in Abkhazia (Coleoptera: Carabidae)." volume="74" year="2013">
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<normalizedToken originalValue="Retezár">Retezar</normalizedToken>
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(2013)
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</bibRefCitation>
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33-37 mm long. Underside black, dorsum usually with bright metallic lustre, dark bronze, greenish-bronze, dark blue, reddish-bronze or black, pronotum often greenish; mandibles, palpi, antennae, and legs black. Habitus see
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<normalizedToken originalValue="Retezár">Retezar</normalizedToken>
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, 2013.
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</paragraph>
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<paragraph pageId="10" pageNumber="31">
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Head normal, frons coarsely, neck moderately wrinkled. Pronotum subquadrate, broadest in anterior one third, PW/PL = 1.14-1.42 (1.25), PW/PB = 1.0-1.26 (1.14), lateral sides of pronotum slightly sinuated before hind angles, the latter strongly protruding backwards and sidewards, pointed apically. Median groove distinct, disk moderately, basal foveae coarsely rugose. Elytra oblong-ovate, broadest behind their middle, EL/EW = 1.61-1.77 (1.69) in males and 1.64-1.85 (1.72) 1.73-1.98 in females. Elytral sculpture somewhat identical in males and females, forming coarse and precise series of links, primary interspaces elevated stronger than secondary ones, regularly interrupted by large foveae. Table
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<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphometric characteristic of males (n = 16) and females (n = 31) of Carabus (Archiplectes) satyrus and males (n = 83) and females (n = 91) of Carabus (Archiplectes) besleticus subspecies." httpUri="http://table.plazi.org/id/E3AA304075ED326595338A0A005BF64B" pageId="10" pageNumber="31" tableUuid="E3AA304075ED326595338A0A005BF64B">2</tableCitation>
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lists the main morphometric measurements of one studied population.
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</paragraph>
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<paragraph pageId="10" pageNumber="31">
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The shape of the male genitalia is practically identical to that of the other
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<taxonomicName authorityName="Kurnakov" authorityYear="1972" baseAuthorityName="besleticus Kurnakov" baseAuthorityYear="1972" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus" order="Coleoptera" pageId="10" pageNumber="31" phylum="Arthropoda" rank="species" species="besleticus" subGenus="Archiplectes">Carabus (Archiplectes) besleticus</taxonomicName>
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taxa.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="31" type="differential diagnosis">
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<paragraph pageId="10" pageNumber="31">Differential diagnosis.</paragraph>
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<paragraph pageId="10" pageNumber="31">
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This form recently described by
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<bibRefCitation author="Retezar, I" journalOrPublisher="Folia Entomologica Hungarica" pageId="23" pageNumber="44" pagination="1 - 6" refId="B9" refString="Retezar, I, 2013. Taxonomic and faunistic studies on the subgenus Carabus (Archiplectes) Gottwald, 1982 in Abkhazia (Coleoptera: Carabidae). Folia Entomologica Hungarica 74: 1 - 6" title="Taxonomic and faunistic studies on the subgenus Carabus (Archiplectes) Gottwald, 1982 in Abkhazia (Coleoptera: Carabidae)." volume="74" year="2013">
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<normalizedToken originalValue="Retezár">Retezar</normalizedToken>
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(2013)
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</bibRefCitation>
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, resembles habitually some populations of
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<taxonomicName authorityName="Kurnakov" authorityYear="1972" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. mtsaranus" order="Coleoptera" pageId="10" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="mtsaranus">Carabus (Archiplectes) besleticus mtsaranus</taxonomicName>
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, but unlike them represents somewhat different combination of features. However, the significant and stable difference is not still ascertained, accumulation of the further material from the upper forest belt will apparently make possible unequivocal precision of its taxonomic relationships with other neighboring populations.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="31" type="distribution">
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<paragraph pageId="10" pageNumber="31">Distribution.</paragraph>
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<paragraph pageId="10" pageNumber="31">Populates the alpine belt of Mt. Khipsta, Mt. Akugra, and Mt. Akibakhu at an altitude of 2000-2300 m.</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="31" type="habitat">
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<paragraph pageId="10" pageNumber="31">Habitat.</paragraph>
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<paragraph pageId="10" pageNumber="31">
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Prefers alpine herb meadows. Activity of imago proceeds from May to August, beetles being active at melting of snow cover.
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<taxonomicName authorityName="Zamotajlov & Koval" authorityYear="1989" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Procechenochilus) gusevi" order="Coleoptera" pageId="10" pageNumber="31" phylum="Arthropoda" rank="species" species="gusevi" subGenus="Procechenochilus">Carabus (Procechenochilus) gusevi</taxonomicName>
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,
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<taxonomicName authority="circassicus" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Tribax) circassicus subsp. circassicus" order="Coleoptera" pageId="10" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="circassicus" subGenus="Tribax" subSpecies="circassicus">Carabus (Tribax) circassicus circassicus</taxonomicName>
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(natio
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<taxonomicName class="Insecta" family="Carabidae" form="abasinus" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="31" phylum="Arthropoda" rank="form">
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<emphasis italics="true" pageId="10" pageNumber="31">abasinus</emphasis>
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</taxonomicName>
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),
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<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Tribax) constantinovi subsp. otcharensis" order="Coleoptera" pageId="10" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="constantinovi" subGenus="Tribax" subSpecies="otcharensis">Carabus (Tribax) constantinovi otcharensis</taxonomicName>
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,
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<taxonomicName baseAuthorityName="Ganglbauer" baseAuthorityYear="1889" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Tribax) agnatus" order="Coleoptera" pageId="10" pageNumber="31" phylum="Arthropoda" rank="species" species="agnatus" subGenus="Tribax">Carabus (Tribax) agnatus</taxonomicName>
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,
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<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Lipaster) stjervalli subsp. humbolti" order="Coleoptera" pageId="10" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="stjervalli" subGenus="Lipaster" subSpecies="humbolti">Carabus (Lipaster) stjervalli humbolti</taxonomicName>
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Faldermann, 1835, and
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<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Pachycarabus) imitator subsp. katharinae" order="Coleoptera" pageId="10" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="imitator" subGenus="Pachycarabus" subSpecies="katharinae">Carabus (Pachycarabus) imitator katharinae</taxonomicName>
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Reitter, 1896 occur together with this subspecies.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |