treatments-xml/data/3B/79/54/3B7954795D769F3D63F0FE81FEB665F1.xml
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<mods:namePart id="492D9F754479BE7FB84EADF9F7BD658E">Adrain, Jonathan M.</mods:namePart>
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<treatment id="3B7954795D769F3D63F0FE81FEB665F1" ID-DOI="http://doi.org/10.5281/zenodo.5256924" ID-Zenodo-Dep="5256924" LSID="urn:lsid:plazi:treatment:3B7954795D769F3D63F0FE81FEB665F1" httpUri="http://treatment.plazi.org/id/3B7954795D769F3D63F0FE81FEB665F1" lastPageNumber="30" pageId="29" pageNumber="30">
<subSubSection id="FBCAB6E45D769F3D63F0FE81FE226171" pageId="29" pageNumber="30" type="nomenclature">
<paragraph id="B36FE56F5D769F3D63F0FE81FD8C6150" blockId="29.[151,590,294,353]" box="[151,590,294,320]" pageId="29" pageNumber="30">
<heading id="E82752035D769F3D63F0FE81FD8C6150" bold="true" box="[151,590,294,320]" fontSize="11" level="1" pageId="29" pageNumber="30" reason="1">
<emphasis id="81A4397D5D769F3D63F0FE81FD8C6150" bold="true" box="[151,590,294,320]" pageId="29" pageNumber="30">
<taxonomicName id="74D09EEC5D769F3D63F0FE81FDC66150" authority="Adrain &amp; Karim, 2012" authorityName="Adrain &amp; Karim" authorityYear="2012" box="[151,516,294,320]" class="Trilobita" family="Bathyuridae" genus="Pseudoolenoides" kingdom="Animalia" order="Proetida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="oilcreekensis" status="sp. nov.">
<emphasis id="81A4397D5D769F3D63F0FE81FDC66150" bold="true" box="[151,516,294,320]" italics="true" pageId="29" pageNumber="30">Pseudoolenoides oilcreekensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9A9784065D769F3D616BFE81FD8C6150" box="[524,590,294,320]" pageId="29" pageNumber="30" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="B36FE56F5D769F3D63F0FEEEFE226171" blockId="29.[151,590,294,353]" box="[151,480,329,353]" pageId="29" pageNumber="30">Plate 12, figs 15, 20, Plate 20</paragraph>
</subSubSection>
<subSubSection id="FBCAB6E45D769F3D63A0FE2DFC8461AF" pageId="29" pageNumber="30" type="description">
<paragraph id="B36FE56F5D769F3D63A0FE2DFBD561B0" blockId="29.[199,1047,394,447]" box="[199,1047,394,416]" pageId="29" pageNumber="30">
1974
<taxonomicName id="74D09EEC5D769F3D6234FE2DFD7B61B0" authority="Hintze" authorityName="Hintze" authorityYear="1953" box="[339,697,394,416]" class="Trilobita" family="Bathyuridae" genus="Pseudoolenoides" kingdom="Animalia" order="Proetida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="acicaudus">
<emphasis id="81A4397D5D769F3D6234FE2DFDAE61B0" box="[339,620,394,416]" italics="true" pageId="29" pageNumber="30">Pseudoolenoides acicaudus</emphasis>
Hintze
</taxonomicName>
; Shaw, p. 22, pl. 1, figs 1322, 24.
</paragraph>
<paragraph id="B36FE56F5D769F3D63A0FE0EFC8461AF" blockId="29.[199,1047,394,447]" box="[199,838,425,447]" pageId="29" pageNumber="30">
1975
<taxonomicName id="74D09EEC5D769F3D6234FE0EFD7B61AF" authority="Hintze" authorityName="Hintze" authorityYear="1953" box="[339,697,425,447]" class="Trilobita" family="Bathyuridae" genus="Pseudoolenoides" kingdom="Animalia" order="Proetida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="acicaudus">
<emphasis id="81A4397D5D769F3D6234FE0EFDAE61AF" box="[339,620,425,447]" italics="true" pageId="29" pageNumber="30">Pseudoolenoides acicaudus</emphasis>
Hintze
</taxonomicName>
; Derby, p. 26.
</paragraph>
</subSubSection>
<subSubSection id="FBCAB6E45D769F3D63F0FE43FDC76254" pageId="29" pageNumber="30" type="materials_examined">
<paragraph id="B36FE56F5D769F3D63F0FE43FDC76254" blockId="29.[151,1440,484,1505]" pageId="29" pageNumber="30">
<emphasis id="81A4397D5D769F3D63F0FE43FEC461ED" bold="true" box="[151,262,484,509]" inLineHeading="true" pageId="29" pageNumber="30" reason="1">Material.</emphasis>
<materialsCitation id="03B8EF325D769F3D6272FE42FDC76254" collectionCode="OU" collectorName="Section SP" country="United States of America" county="Carter County" elevation="127" location="Highway" municipality="Oil Creek" pageId="29" pageNumber="30" specimenCode="OU 7939, OU 7940-7947" specimenCount="1" stateProvince="Oklahoma" typeStatus="holotype">
<typeStatus id="6C6B5BCD5D769F3D6272FE42FE4261ED" box="[277,384,485,509]" pageId="29" pageNumber="30" type="holotype">Holotype</typeStatus>
, cranidium,
<specimenCode id="E3764D145D769F3D616AFE42FDB761ED" box="[525,629,485,509]" collectionCode="OU" country="New Zealand" name="Fossil Catalgoue in the Geology Museum" pageId="29" pageNumber="30">OU 7939</specimenCode>
(Pl. 20, figs 1, 4), from
<collectorName id="1E2580B95D769F3D60EDFE42FBC761ED" box="[906,1029,485,509]" pageId="29" pageNumber="30">Section SP</collectorName>
1
<quantity id="7428488A5D769F3D6744FE42FB4061EC" box="[1059,1154,485,509]" metricMagnitude="2" metricUnit="m" metricValue="1.268" pageId="29" pageNumber="30" unit="m" value="126.8">
<elevation id="38FD025C5D769F3D6744FE42FB4061EC" box="[1059,1154,485,509]" metricMagnitude="2" metricUnit="m" metricValue="1.268" pageId="29" pageNumber="30" unit="m" value="126.8">126.8 m</elevation>
</quantity>
, and assigned specimens
<specimenCode id="E3764D145D769F3D63F0FDAFFE856230" box="[151,327,520,544]" collectionCode="OU" country="New Zealand" name="Fossil Catalgoue in the Geology Museum" pageId="29" pageNumber="30">OU 79407947</specimenCode>
, from
<collectorName id="1E2580B95D769F3D62F7FDAFFDC86230" box="[400,522,520,544]" pageId="29" pageNumber="30">Section SP</collectorName>
1
<elevation id="38FD025C5D769F3D6141FDAFFCD46230" box="[550,790,520,544]" isEnumeration="true" pageId="29" pageNumber="30">
<quantity id="7428488A5D769F3D6141FDAFFD406230" box="[550,642,520,544]" metricMagnitude="2" metricUnit="m" metricValue="1.045" pageId="29" pageNumber="30" unit="m" value="104.5">104.5 m</quantity>
and
<quantity id="7428488A5D769F3D61DFFDAFFCD46230" box="[696,790,520,544]" metricMagnitude="2" metricUnit="m" metricValue="1.268" pageId="29" pageNumber="30" unit="m" value="126.8">126.8 m</quantity>
</elevation>
,
<location id="B60FB3B45D769F3D6045FDAFFC4B6230" LSID="urn:lsid:plazi:treatment:3B7954795D769F3D63F0FE81FEB665F1:B60FB3B45D769F3D6045FDAFFC4B6230" box="[802,905,520,544]" country="United States of America" county="Carter County" municipality="Oil Creek" name="Highway" pageId="29" pageNumber="30" stateProvince="Oklahoma">Highway</location>
77 section,
<location id="B60FB3B45D769F3D6776FDAFFB386230" LSID="urn:lsid:plazi:treatment:3B7954795D769F3D63F0FE81FEB665F1:B60FB3B45D769F3D6776FDAFFB386230" box="[1041,1274,520,544]" country="United States of America" county="Carter County" municipality="Oil Creek" name="Oil Creek Formation" pageId="29" pageNumber="30" stateProvince="Oklahoma">
<collectingMunicipality id="530B7F155D769F3D6776FDAFFBBC6230" box="[1041,1150,520,544]" pageId="29" pageNumber="30">Oil Creek</collectingMunicipality>
Formation
</location>
(Darriwilian),
<collectingCounty id="5A0E9DE35D769F3D63F0FD8BFEF56254" box="[151,311,556,580]" pageId="29" pageNumber="30">Carter County</collectingCounty>
,
<collectingRegion id="71142B8D5D769F3D6225FD8BFE786254" box="[322,442,556,580]" country="United States of America" name="Oklahoma" pageId="29" pageNumber="30">Oklahoma</collectingRegion>
,
<collectingCountry id="CBC7A5FF5D769F3D62A2FD8BFDC36254" box="[453,513,556,580]" name="United States of America" pageId="29" pageNumber="30">USA</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="FBCAB6E45D769F3D63A0FDF7FD7B6277" box="[199,697,591,616]" pageId="29" pageNumber="30" type="etymology">
<paragraph id="B36FE56F5D769F3D63A0FDF7FD7B6277" blockId="29.[151,1440,484,1505]" box="[199,697,591,616]" pageId="29" pageNumber="30">
<emphasis id="81A4397D5D769F3D63A0FDF7FE8F6278" bold="true" box="[199,333,592,616]" inLineHeading="true" pageId="29" pageNumber="30" reason="1">Etymology.</emphasis>
From the Oil Creek Formation.
</paragraph>
</subSubSection>
<subSubSection id="FBCAB6E45D769F3D63A0FDD4FD3E62BF" pageId="29" pageNumber="30" type="diagnosis">
<paragraph id="B36FE56F5D769F3D63A0FDD4FD3E62BF" blockId="29.[151,1440,484,1505]" pageId="29" pageNumber="30">
<emphasis id="81A4397D5D769F3D63A0FDD4FE83629C" bold="true" box="[199,321,627,652]" inLineHeading="true" pageId="29" pageNumber="30" reason="1">Diagnosis.</emphasis>
Eye ridge with highly oblique course; fixigena relatively narrow; pygidium with raised pleural regions; anterior pleural band with distinct distal node.
</paragraph>
</subSubSection>
<subSubSection id="FBCAB6E45D769F3D63A0FD1DFC116403" pageId="29" pageNumber="30" type="description">
<paragraph id="B36FE56F5D769F3D63A0FD1DFC116403" blockId="29.[151,1440,484,1505]" pageId="29" pageNumber="30">
<emphasis id="81A4397D5D769F3D63A0FD1DFE9A62C3" bold="true" box="[199,344,698,723]" inLineHeading="true" pageId="29" pageNumber="30" reason="1">Description.</emphasis>
This species is very similar in cephalic morphology to
<taxonomicName id="74D09EEC5D769F3D60AFFD1CFB9462C3" authorityName="Hintze" authorityYear="1953" box="[968,1110,699,723]" class="Trilobita" family="Bathyuridae" genus="Pseudoolenoides" kingdom="Animalia" order="Proetida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="acicaudus">
<emphasis id="81A4397D5D769F3D60AFFD1CFB9462C3" box="[968,1110,699,723]" italics="true" pageId="29" pageNumber="30">P. acicaudus</emphasis>
</taxonomicName>
, with which it was originally identified (
<bibRefCitation id="D741989E5D769F3D6276FD79FE5B62E6" author="Shaw, F. C." box="[273,409,734,758]" pageId="29" pageNumber="30" pagination="1 - 54" refId="ref35027" refString="Shaw, F. C. (1974) Simpson Group (Middle Ordovician) trilobites of Oklahoma. Paleontological Society Memoir, 6, 1 - 54." type="journal article" year="1974">Shaw, 1974</bibRefCitation>
). Hence, it is described via comparison with that species, noting all observable differences. Cranidium with eye ridge set more obliquely (very clear in Pl. 20, figs 1, 2, but Pl. 20, fig. 3 approaches the attitude seen in
<taxonomicName id="74D09EEC5D769F3D6393FC81FE47632D" authorityName="Hintze" authorityYear="1953" box="[244,389,805,829]" class="Trilobita" family="Bathyuridae" genus="Pseudoolenoides" kingdom="Animalia" order="Proetida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="acicaudus">
<emphasis id="81A4397D5D769F3D6393FC81FE47632D" box="[244,389,805,829]" italics="true" pageId="29" pageNumber="30">P. acicaudus</emphasis>
</taxonomicName>
); interocular fixigenae distinctly narrower (transverse line from maximum lateral point of glabella to exsagittal line running through Ύ is 2731% sagittal distance from front of LO to front of glabella in
<taxonomicName id="74D09EEC5D769F3D66E0FCEDFEEA6395" authorityName="Adrain &amp; Karim" authorityYear="2012" class="Trilobita" family="Bathyuridae" genus="Pseudoolenoides" kingdom="Animalia" order="Proetida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="oilcreekensis">
<emphasis id="81A4397D5D769F3D66E0FCEDFEEA6395" italics="true" pageId="29" pageNumber="30">P. oilcreekensis</emphasis>
</taxonomicName>
, versus 3540% in
<taxonomicName id="74D09EEC5D769F3D616DFCCAFD5A6395" authorityName="Hintze" authorityYear="1953" box="[522,664,877,901]" class="Trilobita" family="Bathyuridae" genus="Pseudoolenoides" kingdom="Animalia" order="Proetida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="acicaudus">
<emphasis id="81A4397D5D769F3D616DFCCAFD5A6395" box="[522,664,877,901]" italics="true" pageId="29" pageNumber="30">P. acicaudus</emphasis>
</taxonomicName>
); anteromedian portion of LO set forward as distinct subtrapezoidal protrusion of anterior margin versus margin nearly transversely straight with only slight anterior projection in a few specimens (e.g., Pl. 17, fig. 17); pygidium with four versus five unreleased segments; raised pleural regions, anterior pleural band with swollen node on distal part, axis tapers more rapidly, rear of axis much narrower; posteromedian spine much more robust proximally and apparently longer.
</paragraph>
</subSubSection>
<subSubSection id="FBCAB6E45D769F3D63A0FBB9FEB665F1" pageId="29" pageNumber="30" type="discussion">
<paragraph id="B36FE56F5D769F3D63A0FBB9FEB665F1" blockId="29.[151,1440,484,1505]" pageId="29" pageNumber="30">
<emphasis id="81A4397D5D769F3D63A0FBB9FE8E6427" bold="true" box="[199,332,1054,1079]" inLineHeading="true" pageId="29" pageNumber="30" reason="1">Discussion.</emphasis>
In cephalic morphology,
<taxonomicName id="74D09EEC5D769F3D611BFBB8FCE96426" authorityName="Adrain &amp; Karim" authorityYear="2012" box="[636,811,1054,1078]" class="Trilobita" family="Bathyuridae" genus="Pseudoolenoides" kingdom="Animalia" order="Proetida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="oilcreekensis">
<emphasis id="81A4397D5D769F3D611BFBB8FCE96426" box="[636,811,1054,1078]" italics="true" pageId="29" pageNumber="30">P. oilcreekensis</emphasis>
</taxonomicName>
is differentiated from
<taxonomicName id="74D09EEC5D769F3D6748FBB8FB7C6426" authorityName="Hintze" authorityYear="1953" box="[1071,1214,1054,1078]" class="Trilobita" family="Bathyuridae" genus="Pseudoolenoides" kingdom="Animalia" order="Proetida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="acicaudus">
<emphasis id="81A4397D5D769F3D6748FBB8FB7C6426" box="[1071,1214,1054,1078]" italics="true" pageId="29" pageNumber="30">P. acicaudus</emphasis>
</taxonomicName>
to an extent which leaves no doubt it is a distinct species. The differentiation is still very subtle, and given the very close similarity of the cranidia and pygidia one would expect the pygidia also to be close. This is not the case, as the single pygidium occurring in the Oil Creek Formation with the cephalic material (Pl. 20, fig. 15) is quite radically different from that of
<taxonomicName id="74D09EEC5D769F3D6380FB09FEB764D5" authorityName="Hintze" authorityYear="1953" box="[231,373,1197,1221]" class="Trilobita" family="Bathyuridae" genus="Pseudoolenoides" kingdom="Animalia" order="Proetida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="acicaudus">
<emphasis id="81A4397D5D769F3D6380FB09FEB764D5" box="[231,373,1197,1221]" italics="true" pageId="29" pageNumber="30">P. acicaudus</emphasis>
</taxonomicName>
, and instead more closely resembles those of
<taxonomicName id="74D09EEC5D769F3D601DFB09FBD364D5" authorityName="Shaw" authorityYear="1974" box="[890,1041,1198,1221]" class="Trilobita" family="Bathyuridae" genus="Pseudoolenoides" kingdom="Animalia" order="Proetida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="carterensis">
<emphasis id="81A4397D5D769F3D601DFB09FBD364D5" box="[890,1041,1198,1221]" italics="true" pageId="29" pageNumber="30">P. carterensis</emphasis>
</taxonomicName>
and
<taxonomicName id="74D09EEC5D769F3D672EFB09FA8064D5" authorityName="Adrain &amp; Karim" authorityYear="2012" box="[1097,1346,1197,1221]" class="Trilobita" family="Bathyuridae" genus="Pseudoolenoides" kingdom="Animalia" order="Proetida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="fossilmountainensis">
<emphasis id="81A4397D5D769F3D672EFB09FA8064D5" box="[1097,1346,1197,1221]" italics="true" pageId="29" pageNumber="30">P. fossilmountainensis</emphasis>
</taxonomicName>
. Given this, one must at least consider the possibility of misassociation. There is apparently no cephalic material associated with the pygidium other than figured herein, but the overall sample size is very small and the possibility exists that the pygidium belongs to a second, rare, species. For this reason we have selected one of the larger heads (Pl. 20, fig. 1) which displays the cranidial differentia well as the
<typeStatus id="6C6B5BCD5D769F3D601DFA9CFC1D6543" box="[890,991,1339,1363]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
, even though if correctly associated the pygidium is the much more distinctive sclerite.
<taxonomicName id="74D09EEC5D769F3D61BBFAF8FBF56567" authorityName="Adrain &amp; Karim" authorityYear="2012" box="[732,1079,1375,1399]" class="Trilobita" family="Bathyuridae" genus="Pseudoolenoides" kingdom="Animalia" order="Proetida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="oilcreekensis">
<emphasis id="81A4397D5D769F3D61BBFAF8FBF56567" box="[732,1079,1375,1399]" italics="true" pageId="29" pageNumber="30">Pseudoolenoides oilcreekensis</emphasis>
</taxonomicName>
occurs a comparable distance beneath
<taxonomicName id="74D09EEC5D769F3D6393FA24FE49658A" authorityName="Shaw" authorityYear="1974" box="[244,395,1411,1434]" class="Trilobita" family="Bathyuridae" genus="Pseudoolenoides" kingdom="Animalia" order="Proetida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="carterensis">
<emphasis id="81A4397D5D769F3D6393FA24FE49658A" box="[244,395,1411,1434]" italics="true" pageId="29" pageNumber="30">P. carterensis</emphasis>
</taxonomicName>
in the Oil Creek Formation (
<quantity id="7428488A5D769F3D61A8FA25FCDD658A" box="[719,799,1410,1434]" metricMagnitude="1" metricUnit="m" metricValue="4.88" pageId="29" pageNumber="30" unit="m" value="48.8">48.8 m</quantity>
at the Highway 77 section) as
<taxonomicName id="74D09EEC5D769F3D6710FA24FAC1658B" authorityName="Hintze" authorityYear="1953" box="[1143,1283,1411,1435]" class="Trilobita" family="Bathyuridae" genus="Pseudoolenoides" kingdom="Animalia" order="Proetida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="acicaudus">
<emphasis id="81A4397D5D769F3D6710FA24FAC1658B" box="[1143,1283,1411,1435]" italics="true" pageId="29" pageNumber="30">P. acicaudus</emphasis>
</taxonomicName>
does beneath
<taxonomicName id="74D09EEC5D769F3D63F0FA00FE5065AE" authorityName="Adrain &amp; Karim" authorityYear="2012" box="[151,402,1446,1470]" class="Trilobita" family="Bathyuridae" genus="Pseudoolenoides" kingdom="Animalia" order="Proetida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="fossilmountainensis">
<emphasis id="81A4397D5D769F3D63F0FA00FE5065AE" box="[151,402,1446,1470]" italics="true" pageId="29" pageNumber="30">P. fossilmountainensis</emphasis>
</taxonomicName>
in the Kanosh Formation (approximately
<quantity id="7428488A5D769F3D601AFA01FC7865AE" box="[893,954,1446,1470]" metricMagnitude="1" metricUnit="m" metricValue="6.8" pageId="29" pageNumber="30" unit="m" value="68.0">68 m</quantity>
) and the pairs of species are each other's closest comparison.
</paragraph>
</subSubSection>
</treatment>
</document>