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<document id="990FEC8F760B36F82FECA7A26E6961B0" ID-CLB-Dataset="31852" ID-DOI="10.11646/zootaxa.4335.1.1" ID-GBIF-Dataset="f2ad3cb1-0bad-4be1-9e36-2170a6a08aaf" ID-ISSN="1175-5326" ID-Zenodo-Dep="1010014" ID-ZooBank="3429592D-23A8-4EBF-8412-BD57B3BC4314" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1507877826645" checkinUser="plazi" docAuthor="Azevedo, Celso O." docDate="2017" docId="3B3687C07C6E9B3AFF67F6BE9C64FF27" docLanguage="en" docName="zootaxa.4335.1.1.pdf" docOrigin="Zootaxa 4335 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Dissomphalus ardonius Azevedo 2017, sp. nov." docType="treatment" docVersion="15" lastPageNumber="31" masterDocId="C70FFFB87C739B24FFF0F23F9F65FFDF" masterDocTitle="Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species" masterLastPageNumber="72" masterPageNumber="1" pageNumber="30" updateTime="1698455438235" updateUser="plazi">
<mods:mods id="9BC235DD889907243004F073C463FB5D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="A9E3A8A7A9F4F86690976A0666F11A55">Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species</mods:title>
</mods:titleInfo>
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<mods:namePart id="9C51EA501390DE77C6B51992E579E6B3">Azevedo, Celso O.</mods:namePart>
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<mods:typeOfResource id="27AF2ECBEAE5F62AC377B18601DF4445">text</mods:typeOfResource>
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<mods:title id="9BDF6FEDBD799B99245DD7CA879CDB6C">Zootaxa</mods:title>
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<mods:part id="DDCA25CC1FB445D1157FF62A4F756840">
<mods:date id="F58DAFC44B65968CCFE9F5FE6009F484">2017</mods:date>
<mods:detail id="1AFE8678A470FB8882EA059DFE7CF7A8" type="pubDate">
<mods:number id="7A91AAD4EC4CCD3CF1BD0E5359259A2C">2017-10-13</mods:number>
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<mods:number id="DB6AF3195F61C1CC48FD91F9F91C81C2">4335</mods:number>
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<mods:classification id="D88320D7D7C335AD7635B133C340A7AE">journal article</mods:classification>
<mods:identifier id="6D50B2AAFA5232E137C59AFE5DDE3B41" type="CLB-Dataset">31852</mods:identifier>
<mods:identifier id="D0590E5B481B7E918FF5E155840ED273" type="DOI">10.11646/zootaxa.4335.1.1</mods:identifier>
<mods:identifier id="B5DFE5DF54BCDFBD09ED86D4C21BF649" type="GBIF-Dataset">f2ad3cb1-0bad-4be1-9e36-2170a6a08aaf</mods:identifier>
<mods:identifier id="54F26D08BD4276D5E0F842549A760D70" type="ISSN">1175-5326</mods:identifier>
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<treatment id="3B3687C07C6E9B3AFF67F6BE9C64FF27" ID-DOI="http://doi.org/10.5281/zenodo.5020374" ID-GBIF-Taxon="135435237" ID-Zenodo-Dep="5020374" LSID="urn:lsid:plazi:treatment:3B3687C07C6E9B3AFF67F6BE9C64FF27" httpUri="http://treatment.plazi.org/id/3B3687C07C6E9B3AFF67F6BE9C64FF27" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
<subSubSection id="FB85655D7C6E9B39FF67F6BE9ED6FB62" pageId="29" pageNumber="30" type="nomenclature">
<paragraph id="B32036D67C6E9B39FF67F6BE9D9FFB44" blockId="29.[151,762,1153,1213]" box="[151,762,1153,1179]" pageId="29" pageNumber="30">
<heading id="E86881BA7C6E9B39FF67F6BE9D9FFB44" bold="true" box="[151,762,1153,1179]" fontSize="11" level="1" pageId="29" pageNumber="30" reason="1">
<emphasis id="81EBEAC47C6E9B39FF67F6BE9D9FFB44" bold="true" box="[151,762,1153,1179]" pageId="29" pageNumber="30">
<taxonomicName id="749F4D557C6E9B39FF67F6BE9DEAFB44" authority="Brito &amp; Azevedo" authorityName="Azevedo" authorityYear="2017" box="[151,655,1153,1179]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="ardonius" status="sp. nov.">
<emphasis id="81EBEAC47C6E9B39FF67F6BE9ED0FB44" bold="true" box="[151,437,1153,1179]" italics="true" pageId="29" pageNumber="30">Dissomphalus ardonius</emphasis>
Brito &amp; Azevedo
</taxonomicName>
,
<taxonomicNameLabel id="9AD857BF7C6E9B39FD6DF6BE9D9FFB44" box="[669,762,1153,1179]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="B32036D67C6E9B39FF67F69B9ED6FB62" blockId="29.[151,762,1153,1213]" box="[151,435,1188,1213]" pageId="29" pageNumber="30">
(
<figureCitation id="2BA42A537C6E9B39FF6FF69B9F82FB63" box="[159,231,1188,1213]" captionStart="FIGURES 1 – 12" captionStartId="10.[151,264,1512,1535]" captionTargetBox="[151,1436,195,1484]" captionTargetId="figure@10.[151,1436,195,1489]" captionTargetPageId="10" captionText="FIGURES 1 – 12. Dissomphalus spp., ♂, head, dorsal view, except 2 lateral view. 1 – 2. D. isteus sp. nov. 3. D. azagus sp. nov. 4. D. asfartus sp. nov. 5. D. iozus sp. nov. 6. D. asitius sp. nov. 7. D. ibocius sp. nov. 8. D. tisinus sp. nov. 9. D. ardonius sp. nov. 10. D. arteus sp. nov. 11. D. ettus sp. nov. 12. D. ibrastus sp. nov.. Scale bars = 20 µm, except fig. 10 = 50 µm and fig. 4 = 100 µm." httpUri="https://zenodo.org/record/1010016/files/figure.png" pageId="29" pageNumber="30">Figs 9</figureCitation>
,
<figureCitation id="2BA42A537C6E9B39FF03F69B9E77FB63" box="[243,274,1188,1212]" captionStart="FIGURES 33 – 44" captionStartId="13.[151,264,1907,1930]" captionTargetBox="[151,1436,553,1886]" captionTargetId="figure@13.[151,1436,553,1886]" captionTargetPageId="13" captionText="FIGURES 33 – 44. Dissomphalus spp., ♂, metasomal tergal process, dorsal view. 33. D. isteus sp. nov. 34. D. azagus sp. nov. 35. D. asfartus sp. nov. 36. D. iozus sp. nov. 37. D. asitius sp. nov. 38. D. ibocius sp. nov. 39. D. tisinus sp. nov. 40. D. ardonius sp. nov. 41. D. arteus sp. nov. 42. D. ettus sp. nov. 43. D. ibrastus sp. nov. 44. D. racarus sp. nov.. Scale bars = 20 µm, except fig. 35 = 200 µm." httpUri="https://zenodo.org/record/1010022/files/figure.png" pageId="29" pageNumber="30">40</figureCitation>
,
<figureCitation id="2BA42A537C6E9B39FEEDF69B9E5EFB63" box="[285,315,1188,1212]" captionStart="FIGURES 64 – 75" captionStartId="16.[151,264,1938,1961]" captionTargetBox="[175,1408,606,1889]" captionTargetId="figure@16.[151,1436,569,1917]" captionTargetPageId="16" captionText="FIGURES 64 – 75. Dissomphalus spp., ♂, hypopygium, internal view. 64. D. isteus sp. nov. 65. D. azagus sp. nov. 66. D. asfartus sp. nov. 67. D. iozus sp. nov. 68. D. asitius sp. nov. 69. D. ibocius sp. nov. 70. D. tisinus sp. nov. 71. D. ardonius sp. nov. 72. D. arteus sp. nov. 73. D. ettus sp. nov. 74. D. ibrastus sp. nov. 75. D. racarus sp. nov.. Scale bars = 125 µm." httpUri="https://zenodo.org/record/1010028/files/figure.png" pageId="29" pageNumber="30">71</figureCitation>
,
<figureCitation id="2BA42A537C6E9B39FEB7F69B9ECEFB62" box="[327,427,1188,1213]" captionStart="FIGURES 123 – 135" captionStartId="26.[151,264,1820,1843]" captionTargetBox="[170,1430,415,1784]" captionTargetId="figure@26.[151,1436,411,1799]" captionTargetPageId="26" captionText="FIGURES 123 – 135. Dissomphalus spp., ♂, genitália. 123 – 124. D. angulatus 123. Ventral view. 124. Lateral view. 125 – 126. D. asitius sp. nov. 125. Dorsal view. 126. Ventral view. 127 – 128. D. ibocius sp. nov. 127. Dorsal view. 128. Ventral view. 129 – 130. D. tisinus sp. nov. 129. Dorsal view. 130. Ventral view. 131 – 132. D. cervoides 131. Dorsal view. 132. Ventral view. 133 – 135. D. ardonius sp. nov. 133. Dorsal view. 134. Ventral view. 135. Lateral view. Scale bars = 250 µm." httpUri="https://zenodo.org/record/1010038/files/figure.png" pageId="29" pageNumber="30">133–135</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C6E9B39FF67F6D39D19F950" pageId="29" pageNumber="30" type="description">
<paragraph id="B32036D67C6E9B39FF67F6D39A57F9B2" blockId="29.[151,1437,1260,2005]" pageId="29" pageNumber="30">
<emphasis id="81EBEAC47C6E9B39FF67F6D39E4CFADA" bold="true" box="[151,297,1260,1285]" pageId="29" pageNumber="30">Description.</emphasis>
Male. Body length
<quantity id="74679B337C6E9B39FDFAF6D39D17FADB" box="[522,626,1260,1284]" metricMagnitude="-3" metricUnit="m" metricValue="3.22" pageId="29" pageNumber="30" unit="mm" value="3.22">3.22 mm</quantity>
. Color: body castaneous. Head (
<figureCitation id="2BA42A537C6E9B39FC10F6D39B40FADB" box="[992,1061,1260,1285]" captionStart="FIGURES 1 – 12" captionStartId="10.[151,264,1512,1535]" captionTargetBox="[151,1436,195,1484]" captionTargetId="figure@10.[151,1436,195,1489]" captionTargetPageId="10" captionText="FIGURES 1 – 12. Dissomphalus spp., ♂, head, dorsal view, except 2 lateral view. 1 – 2. D. isteus sp. nov. 3. D. azagus sp. nov. 4. D. asfartus sp. nov. 5. D. iozus sp. nov. 6. D. asitius sp. nov. 7. D. ibocius sp. nov. 8. D. tisinus sp. nov. 9. D. ardonius sp. nov. 10. D. arteus sp. nov. 11. D. ettus sp. nov. 12. D. ibrastus sp. nov.. Scale bars = 20 µm, except fig. 10 = 50 µm and fig. 4 = 100 µm." httpUri="https://zenodo.org/record/1010016/files/figure.png" pageId="29" pageNumber="30">Fig. 9</figureCitation>
). Mandible with two distal teeth. Median clypeal lobe trapezoidal, with one angulate tooth; median clypeal carina high in profile, complete or nearly so, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest weakly concave. Pronotal disc coriaceous; anterior margin coarse. Metasoma (
<figureCitation id="2BA42A537C6E9B39FCB6F7689CFFFAAF" box="[838,922,1367,1392]" captionStart="FIGURES 33 – 44" captionStartId="13.[151,264,1907,1930]" captionTargetBox="[151,1436,553,1886]" captionTargetId="figure@13.[151,1436,553,1886]" captionTargetPageId="13" captionText="FIGURES 33 – 44. Dissomphalus spp., ♂, metasomal tergal process, dorsal view. 33. D. isteus sp. nov. 34. D. azagus sp. nov. 35. D. asfartus sp. nov. 36. D. iozus sp. nov. 37. D. asitius sp. nov. 38. D. ibocius sp. nov. 39. D. tisinus sp. nov. 40. D. ardonius sp. nov. 41. D. arteus sp. nov. 42. D. ettus sp. nov. 43. D. ibrastus sp. nov. 44. D. racarus sp. nov.. Scale bars = 20 µm, except fig. 35 = 200 µm." httpUri="https://zenodo.org/record/1010022/files/figure.png" pageId="29" pageNumber="30">Fig. 40</figureCitation>
). Tergal process with lateral, shallow, circular and small depression, with large, very low and dorsad tubercle placed on center of depression, pit large, with short and sparse tuft of setae. Posterior hypopygeal margin median straight with angled corners (
<figureCitation id="2BA42A537C6E9B39FB7EF7A09B84FA67" box="[1166,1249,1439,1464]" captionStart="FIGURES 64 – 75" captionStartId="16.[151,264,1938,1961]" captionTargetBox="[175,1408,606,1889]" captionTargetId="figure@16.[151,1436,569,1917]" captionTargetPageId="16" captionText="FIGURES 64 – 75. Dissomphalus spp., ♂, hypopygium, internal view. 64. D. isteus sp. nov. 65. D. azagus sp. nov. 66. D. asfartus sp. nov. 67. D. iozus sp. nov. 68. D. asitius sp. nov. 69. D. ibocius sp. nov. 70. D. tisinus sp. nov. 71. D. ardonius sp. nov. 72. D. arteus sp. nov. 73. D. ettus sp. nov. 74. D. ibrastus sp. nov. 75. D. racarus sp. nov.. Scale bars = 125 µm." httpUri="https://zenodo.org/record/1010028/files/figure.png" pageId="29" pageNumber="30">Fig. 71</figureCitation>
). Genitalia (
<figureCitation id="2BA42A537C6E9B39FA9AF7A09F98FA02" captionStart="FIGURES 123 – 135" captionStartId="26.[151,264,1820,1843]" captionTargetBox="[170,1430,415,1784]" captionTargetId="figure@26.[151,1436,411,1799]" captionTargetPageId="26" captionText="FIGURES 123 – 135. Dissomphalus spp., ♂, genitália. 123 – 124. D. angulatus 123. Ventral view. 124. Lateral view. 125 – 126. D. asitius sp. nov. 125. Dorsal view. 126. Ventral view. 127 – 128. D. ibocius sp. nov. 127. Dorsal view. 128. Ventral view. 129 – 130. D. tisinus sp. nov. 129. Dorsal view. 130. Ventral view. 131 – 132. D. cervoides 131. Dorsal view. 132. Ventral view. 133 – 135. D. ardonius sp. nov. 133. Dorsal view. 134. Ventral view. 135. Lateral view. Scale bars = 250 µm." httpUri="https://zenodo.org/record/1010038/files/figure.png" pageId="29" pageNumber="30">Figs 133–135</figureCitation>
): paramere with dorsal margin entirely wide; apical margin truncate. Aedeagal ventral ramus shorter than dorsal body, entirely wide, cross section laminar; surface vertical; inner margin strongly sinuous; outer margin slightly sinuous; apex long, slightly crenulate, parallel; additional inner ramus long; basal stub long and narrow. Aedeagal dorsal body with two pairs of apical lobes, outer lobe long, vertical and very narrow, with apical margin acute and ventrad; inner pair stout, membranous and setose. Apodeme not extending beyond genital ring.
</paragraph>
<paragraph id="B32036D67C6E9B39FF37F4489D19F950" blockId="29.[151,1437,1260,2005]" box="[199,636,1655,1680]" pageId="29" pageNumber="30">
<emphasis id="81EBEAC47C6E9B39FF37F4489E59F94F" bold="true" box="[199,316,1655,1680]" pageId="29" pageNumber="30">Variation.</emphasis>
Metasoma light castaneous.
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C6E9B39FF37F4A39D68F86E" pageId="29" pageNumber="30" type="discussion">
<paragraph id="B32036D67C6E9B39FF37F4A39D68F86E" blockId="29.[151,1437,1260,2005]" pageId="29" pageNumber="30">
<emphasis id="81EBEAC47C6E9B39FF37F4A39E5EF96A" bold="true" box="[199,315,1692,1717]" pageId="29" pageNumber="30">Remarks.</emphasis>
This species is easily recognized by having two spines in the ventral margin of the apical lobes of the aedeagal dorsal body with apical spine shorter than distal one, the aedeagal ventral ramus vertical and wide in the median portion with wavy apex. This species is similar to
<taxonomicName id="749F4D557C6E9B39FC92F4DA9CBCF923" authority="Azevedo, 2017" authorityName="Azevedo" authorityYear="2017" box="[866,985,1765,1788]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="urazius" status="sp. nov.">
<emphasis id="81EBEAC47C6E9B39FC92F4DA9CBCF923" box="[866,985,1765,1788]" italics="true" pageId="29" pageNumber="30">D. urazius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9AD857BF7C6E9B39FC12F4DB9B58F922" box="[994,1085,1764,1789]" pageId="29" pageNumber="30" rank="species">
<emphasis id="81EBEAC47C6E9B39FC12F4DB9B58F922" bold="true" box="[994,1085,1764,1789]" pageId="29" pageNumber="30">sp. nov.</emphasis>
</taxonomicNameLabel>
by having spine in the ventral margin of apical lobes of the aedeagal dorsal body and the aedeagal ventral ramus entirely wide with the inner margin sinuous. However, this species has the dorsal margin of the paramere sinuous, the apex of the basal bar not projected forward, and the apical margin of the aedeagal dorsal body wavy, whereas
<taxonomicName id="749F4D557C6E9B39FB97F56E9BBBF8B7" authority="Azevedo, 2017" authorityName="Azevedo" authorityYear="2017" box="[1127,1246,1873,1896]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="urazius" status="sp. nov.">
<emphasis id="81EBEAC47C6E9B39FB97F56E9BBBF8B7" box="[1127,1246,1873,1896]" italics="true" pageId="29" pageNumber="30">D. urazius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9AD857BF7C6E9B39FB16F5709A27F8B7" box="[1254,1346,1871,1896]" pageId="29" pageNumber="30" rank="species">
<emphasis id="81EBEAC47C6E9B39FB16F5709A27F8B7" bold="true" box="[1254,1346,1871,1896]" pageId="29" pageNumber="30">sp. nov.</emphasis>
</taxonomicNameLabel>
has the dorsal margin of the paramere not sinuous, the apex of the basal bar projected forward and the apical margin of the aedeagal ventral ramus not wavy.
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C6E9B3AFF37F5839D27FF0A" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30" type="materials_examined">
<paragraph id="B32036D67C6E9B3AFF37F5839D27FF0A" blockId="29.[151,1437,1260,2005]" lastBlockId="30.[151,1437,151,248]" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
<materialsCitation id="03F73C8B7C6E9B3AFF37F5839AFCFF6F" ID-GBIF-Occurrence="1671715986" collectingDate="1999-10-17" collectingDateMax="1999-10-24" collectingDateMin="1999-10-17" collectionCode="MIUP" collectorName="Teribe, P &amp; Tr &amp; Malaise, A" country="Panama" lastPageId="30" lastPageNumber="31" latitude="9.406944" location="Material" longLatPrecision="21" longitude="-82.93889" municipality="Wekso" pageId="29" pageNumber="30" specimenCount="1" specimenCount-male="1" stateProvince="Bocas del Toro" typeStatus="holotype">
<emphasis id="81EBEAC47C6E9B39FF37F5839ED6F80A" bold="true" box="[199,435,1980,2005]" pageId="29" pageNumber="30">
<location id="B640600D7C6E9B39FF37F5839E4AF80A" LSID="urn:lsid:plazi:treatment:3B3687C07C6E9B3AFF67F6BE9C64FF27:B640600D7C6E9B39FF37F5839E4AF80A" box="[199,303,1980,2005]" country="Panama" latitude="9.406944" longLatPrecision="21" longitude="-82.93889" municipality="Wekso" name="Material" pageId="29" pageNumber="30" stateProvince="Bocas del Toro">Material</location>
examined.
</emphasis>
<typeStatus id="6C2488747C6E9B39FE4CF5839D42F80A" box="[444,551,1980,2005]" pageId="29" pageNumber="30" type="holotype">Holotype</typeStatus>
,
<specimenCount id="A599FD5F7C6E9B39FDC5F5829D29F80B" box="[565,588,1981,2004]" pageId="29" pageNumber="30" type="male">♂</specimenCount>
,
<collectingCountry id="CB8876467C6E9B39FDAAF5839DB1F80B" box="[602,724,1980,2004]" name="Panama" pageId="29" pageNumber="30">PANAMA</collectingCountry>
,
<collectingRegion id="715BF8347C6E9B39FD11F5829CEBF80B" box="[737,910,1980,2005]" country="Panama" name="Bocas del Toro" pageId="29" pageNumber="30">
<emphasis id="81EBEAC47C6E9B39FD11F5829CEBF80B" box="[737,910,1980,2005]" italics="true" pageId="29" pageNumber="30">Bocas del Toro</emphasis>
</collectingRegion>
:
<collectingMunicipality id="5344ACAC7C6E9B39FC6FF5839C8BF80A" box="[927,1006,1980,2005]" pageId="29" pageNumber="30">Wekso</collectingMunicipality>
,
<collectorName id="1E6A53007C6E9B39FC0BF5839B07F80B" box="[1019,1122,1980,2005]" pageId="29" pageNumber="30">Teribe, P</collectorName>
[arque] I[nternacional] L[a] A[mistad], 50 msnm, [
<geoCoordinate id="D6AB50117C6D9B3AFE6FF2A89D41FF6F" box="[415,548,151,176]" direction="north" orientation="latitude" pageId="30" pageNumber="31" precision="15" value="9.406944">09º24'25&quot;N</geoCoordinate>
,
<geoCoordinate id="D6AB50117C6D9B3AFDC4F2A89DA4FF6F" box="[564,705,151,176]" direction="west" orientation="longitude" pageId="30" pageNumber="31" precision="15" value="-82.93889">82º56'20&quot;W</geoCoordinate>
],
<collectingDate id="D765E9FE7C6D9B3AFD27F2A79CF2FF6F" box="[727,919,152,176]" pageId="30" pageNumber="31" value="1999-10-17" valueMax="1999-10-24" valueMin="1999-10-17">
<date id="C72110167C6D9B3AFD27F2A79CF2FF6F" box="[727,919,152,176]" pageId="30" pageNumber="31" value="1999-10-17" valueMax="1999-10-24" valueMin="1999-10-17">17–24 oct. 1999</date>
</collectingDate>
,
<collectorName id="1E6A53007C6D9B3AFC57F2A79CA6FF70" box="[935,963,152,175]" pageId="30" pageNumber="31">Tr</collectorName>
[ampa]
<collectorName id="1E6A53007C6D9B3AFBEAF2A89BF9FF6F" box="[1050,1180,151,176]" pageId="30" pageNumber="31">Malaise, A</collectorName>
. Santos[col.] (
<collectionCode id="D58EAE137C6D9B3AFAB4F2A79AF5FF6F" box="[1348,1424,152,176]" pageId="30" pageNumber="31">MIUP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="03F73C8B7C6D9B3AFF67F2829D5BFF0A" ID-GBIF-Occurrence="1671715978" box="[151,574,188,213]" collectingDate="1999-10-17" collectingDateMax="1999-10-24" collectingDateMin="1999-10-17" collectionCode="MIUP" collectorName="Teribe, P &amp; Tr &amp; Malaise, A" country="Panama" latitude="9.406944" location="Material" longLatPrecision="21" longitude="-82.93889" municipality="Wekso" pageId="30" pageNumber="31" specimenCount="9" specimenCount-male="9" stateProvince="Bocas del Toro" typeStatus="paratype">
<typeStatus id="6C2488747C6D9B3AFF67F2829E6CFF0A" box="[151,265,189,213]" pageId="30" pageNumber="31" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="A599FD5F7C6D9B3AFEE3F2839E5DFF0B" box="[275,312,188,212]" pageId="30" pageNumber="31" type="male">9♂</specimenCount>
, same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C6D9B3AFF37F2E09C64FF27" box="[199,769,223,248]" pageId="30" pageNumber="31" type="distribution">
<paragraph id="B32036D67C6D9B3AFF37F2E09C64FF27" blockId="30.[151,1437,151,248]" box="[199,769,223,248]" pageId="30" pageNumber="31">
<emphasis id="81EBEAC47C6D9B3AFF37F2E09E3CFF27" bold="true" box="[199,345,223,248]" pageId="30" pageNumber="31">Distribution</emphasis>
(
<figureCitation id="2BA42A537C6D9B3AFE98F2E09EAEFF27" box="[360,459,223,248]" captionStart="FIGURES 224 – 233" captionStartId="66.[151,264,1814,1837]" captionTargetBox="[182,1404,199,1768]" captionTargetId="figure@66.[169,1413,193,1773]" captionTargetPageId="66" captionText="FIGURES 224 – 233. Known distribution of Dissomphalus in Panama. 224. Provinces and Counties of Panama. 1. Bocas del Toro. 2. Chiriquí. 3. Ngöbe-Buglé County. 4. Veraguas. 5. Herrera. 6. Los Santos. 7. Colón. 8. Coclé. 9. Panama. 10. Guna Yala County. 11. Darién. 12. Emberá Woonan County. 225. D. abarus sp. nov., D. afestus sp. nov. and D. altivolans. 226. D. angulatus sp. nov., D. ardonius sp. nov. and D. aretisus sp. nov.. 227. D. apertus, D. areius sp. nov. and D. arteobius sp. nov.. 228. D. arbeius sp. nov., D. asitius sp. nov. and D. brasiliensis. 229. D. arteus sp. nov., D. cervoides and D. culteratus. 230. D. asfartus sp. nov.. 231. D. azagus sp. nov., D. bilobatus and D. coronatus. 232. D. borus sp. nov., D. citus sp. nov. and D. curvifoveatus. 233. D. declinatus, D. dilatatus and D. ettus sp. nov.." httpUri="https://zenodo.org/record/1010056/files/figure.png" pageId="30" pageNumber="31">Fig. 226</figureCitation>
).
<collectingRegion id="715BF8347C6D9B3AFE2EF2DF9D52FF28" box="[478,567,224,247]" country="Panama" name="Panama" pageId="30" pageNumber="31">
<collectingCountry id="CB8876467C6D9B3AFE2EF2DF9D52FF28" box="[478,567,224,247]" name="Panama" pageId="30" pageNumber="31">Panama</collectingCountry>
</collectingRegion>
(
<collectingRegion id="715BF8347C6D9B3AFDB6F2DF9D93FF27" box="[582,758,223,248]" country="Panama" name="Bocas del Toro" pageId="30" pageNumber="31">Bocas del Toro</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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