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<document id="1B0EF396E0079BB80AF3033E9F4028EC" ID-CLB-Dataset="31852" ID-DOI="10.11646/zootaxa.4335.1.1" ID-GBIF-Dataset="f2ad3cb1-0bad-4be1-9e36-2170a6a08aaf" ID-ISSN="1175-5326" ID-Zenodo-Dep="1010014" ID-ZooBank="3429592D-23A8-4EBF-8412-BD57B3BC4314" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1507877826645" checkinUser="plazi" docAuthor="Azevedo, Celso O." docDate="2017" docId="3B3687C07C6D9B3AFF67F3739C1CFAF0" docLanguage="en" docName="zootaxa.4335.1.1.pdf" docOrigin="Zootaxa 4335 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Dissomphalus arteus Azevedo 2017, sp. nov." docType="treatment" docVersion="15" lastPageNumber="31" masterDocId="C70FFFB87C739B24FFF0F23F9F65FFDF" masterDocTitle="Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species" masterLastPageNumber="72" masterPageNumber="1" pageNumber="31" updateTime="1698455438235" updateUser="plazi">
<mods:mods id="6450ABCD63DF7CF5811C97027D9C6717" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="7FBC0E631F59273289C53F493A33BA83">Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species</mods:title>
</mods:titleInfo>
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<mods:namePart id="E4DE0510769498131E4F6B464BE192DB">Azevedo, Celso O.</mods:namePart>
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<mods:title id="EB4C52F455E2E12B7A99C64FDA5D3F15">Zootaxa</mods:title>
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<mods:part id="A32E334E8929A044808CBF1A42A31485">
<mods:date id="AFD8F9A9274B1E5CAB9BCC6FC438A7AF">2017</mods:date>
<mods:detail id="2A990D978882547B219B3E5F5AE31DA4" type="pubDate">
<mods:number id="205239ED7E61FFBDE5D6C4DC96875473">2017-10-13</mods:number>
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<mods:number id="8BA32B3D9762FFF5537D5C76A8B13646">4335</mods:number>
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<mods:classification id="6B036751FD40D8FC41AD7DE4D8BDC36E">journal article</mods:classification>
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<mods:identifier id="41EE85788CDEC85046C9803F3DF438A5" type="DOI">10.11646/zootaxa.4335.1.1</mods:identifier>
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<treatment id="3B3687C07C6D9B3AFF67F3739C1CFAF0" ID-DOI="http://doi.org/10.5281/zenodo.5020376" ID-GBIF-Taxon="135435239" ID-Zenodo-Dep="5020376" LSID="urn:lsid:plazi:treatment:3B3687C07C6D9B3AFF67F3739C1CFAF0" httpUri="http://treatment.plazi.org/id/3B3687C07C6D9B3AFF67F3739C1CFAF0" lastPageNumber="31" pageId="30" pageNumber="31">
<subSubSection id="FB85655D7C6D9B3AFF67F3739EA4FE58" pageId="30" pageNumber="31" type="nomenclature">
<paragraph id="B32036D67C6D9B3AFF67F3739DBFFEB9" blockId="30.[151,730,332,391]" box="[151,730,332,358]" pageId="30" pageNumber="31">
<heading id="E86881BA7C6D9B3AFF67F3739DBFFEB9" bold="true" box="[151,730,332,358]" fontSize="11" level="1" pageId="30" pageNumber="31" reason="1">
<emphasis id="81EBEAC47C6D9B3AFF67F3739DBFFEB9" bold="true" box="[151,730,332,358]" pageId="30" pageNumber="31">
<taxonomicName id="749F4D557C6D9B3AFF67F3739D0AFEB9" authority="Brito &amp; Azevedo" authorityName="Azevedo" authorityYear="2017" box="[151,623,332,358]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="arteus" status="sp. nov.">
<emphasis id="81EBEAC47C6D9B3AFF67F3739EF1FEB9" bold="true" box="[151,404,332,358]" italics="true" pageId="30" pageNumber="31">Dissomphalus arteus</emphasis>
Brito &amp; Azevedo
</taxonomicName>
,
<taxonomicNameLabel id="9AD857BF7C6D9B3AFD8CF3739DBFFEB9" box="[636,730,332,358]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="B32036D67C6D9B3AFF67F3519EA4FE58" blockId="30.[151,730,332,391]" box="[151,449,366,391]" pageId="30" pageNumber="31">
(
<figureCitation id="2BA42A537C6D9B3AFF6FF3519F90FE58" box="[159,245,366,391]" captionStart="FIGURES 1 – 12" captionStartId="10.[151,264,1512,1535]" captionTargetBox="[151,1436,195,1484]" captionTargetId="figure@10.[151,1436,195,1489]" captionTargetPageId="10" captionText="FIGURES 1 – 12. Dissomphalus spp., ♂, head, dorsal view, except 2 lateral view. 1 – 2. D. isteus sp. nov. 3. D. azagus sp. nov. 4. D. asfartus sp. nov. 5. D. iozus sp. nov. 6. D. asitius sp. nov. 7. D. ibocius sp. nov. 8. D. tisinus sp. nov. 9. D. ardonius sp. nov. 10. D. arteus sp. nov. 11. D. ettus sp. nov. 12. D. ibrastus sp. nov.. Scale bars = 20 µm, except fig. 10 = 50 µm and fig. 4 = 100 µm." httpUri="https://zenodo.org/record/1010016/files/figure.png" pageId="30" pageNumber="31">Figs 10</figureCitation>
,
<figureCitation id="2BA42A537C6D9B3AFEF1F3509E7AFE58" box="[257,287,367,391]" captionStart="FIGURES 33 – 44" captionStartId="13.[151,264,1907,1930]" captionTargetBox="[151,1436,553,1886]" captionTargetId="figure@13.[151,1436,553,1886]" captionTargetPageId="13" captionText="FIGURES 33 – 44. Dissomphalus spp., ♂, metasomal tergal process, dorsal view. 33. D. isteus sp. nov. 34. D. azagus sp. nov. 35. D. asfartus sp. nov. 36. D. iozus sp. nov. 37. D. asitius sp. nov. 38. D. ibocius sp. nov. 39. D. tisinus sp. nov. 40. D. ardonius sp. nov. 41. D. arteus sp. nov. 42. D. ettus sp. nov. 43. D. ibrastus sp. nov. 44. D. racarus sp. nov.. Scale bars = 20 µm, except fig. 35 = 200 µm." httpUri="https://zenodo.org/record/1010022/files/figure.png" pageId="30" pageNumber="31">41</figureCitation>
,
<figureCitation id="2BA42A537C6D9B3AFEDBF3509E2FFE58" box="[299,330,367,391]" captionStart="FIGURES 64 – 75" captionStartId="16.[151,264,1938,1961]" captionTargetBox="[175,1408,606,1889]" captionTargetId="figure@16.[151,1436,569,1917]" captionTargetPageId="16" captionText="FIGURES 64 – 75. Dissomphalus spp., ♂, hypopygium, internal view. 64. D. isteus sp. nov. 65. D. azagus sp. nov. 66. D. asfartus sp. nov. 67. D. iozus sp. nov. 68. D. asitius sp. nov. 69. D. ibocius sp. nov. 70. D. tisinus sp. nov. 71. D. ardonius sp. nov. 72. D. arteus sp. nov. 73. D. ettus sp. nov. 74. D. ibrastus sp. nov. 75. D. racarus sp. nov.. Scale bars = 125 µm." httpUri="https://zenodo.org/record/1010028/files/figure.png" pageId="30" pageNumber="31">72</figureCitation>
,
<figureCitation id="2BA42A537C6D9B3AFEA5F3519EDCFE58" box="[341,441,366,391]" captionStart="FIGURES 136 – 147" captionStartId="31.[151,264,1782,1805]" captionTargetBox="[157,1424,384,1751]" captionTargetId="figure@31.[151,1436,373,1762]" captionTargetPageId="31" captionText="FIGURES 136 – 147. Dissomphalus spp., ♂, genitalia, except fig. 147 apical lobes of aedeagal dorsal body. 136 – 137. D. arteus sp. nov. 136. Dorsal view. 137. Ventral view. 138 – 139. D. ettus sp. nov. 138. Dorsal view. 139. Ventral view. 140 – 141. D. ibrastus sp. nov. 140. Dorsal view. 141. Ventral view. 142 – 144. D. racarus sp. nov. 142. Dorsal view. 143. Ventral view. 143. Lateral view. 145 – 147. D. teraus sp. nov. 145. Dorsal view. 146. Ventral view. 147. Apical lobes of aedeagal dorsal body, ventral view. Scale bars = 250 µm." httpUri="https://zenodo.org/record/1010040/files/figure.png" pageId="30" pageNumber="31">136–137</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C6D9B3AFF67F3899B4FFCA0" pageId="30" pageNumber="31" type="description">
<paragraph id="B32036D67C6D9B3AFF67F3899FAEFC84" blockId="30.[151,1437,438,1327]" pageId="30" pageNumber="31">
<emphasis id="81EBEAC47C6D9B3AFF67F3899E4CFE10" bold="true" box="[151,297,438,463]" pageId="30" pageNumber="31">Description.</emphasis>
Male. Body length
<quantity id="74679B337C6D9B3AFDE6F3899DE4FE11" box="[534,641,438,463]" metricMagnitude="-3" metricUnit="m" metricValue="4.96" pageId="30" pageNumber="31" unit="mm" value="4.96">4.96 mm</quantity>
. Color: head and mesosoma dark castaneous; metasoma castaneous. Head (
<figureCitation id="2BA42A537C6D9B3AFF12F3E59E52FE2C" box="[226,311,474,499]" captionStart="FIGURES 1 – 12" captionStartId="10.[151,264,1512,1535]" captionTargetBox="[151,1436,195,1484]" captionTargetId="figure@10.[151,1436,195,1489]" captionTargetPageId="10" captionText="FIGURES 1 – 12. Dissomphalus spp., ♂, head, dorsal view, except 2 lateral view. 1 – 2. D. isteus sp. nov. 3. D. azagus sp. nov. 4. D. asfartus sp. nov. 5. D. iozus sp. nov. 6. D. asitius sp. nov. 7. D. ibocius sp. nov. 8. D. tisinus sp. nov. 9. D. ardonius sp. nov. 10. D. arteus sp. nov. 11. D. ettus sp. nov. 12. D. ibrastus sp. nov.. Scale bars = 20 µm, except fig. 10 = 50 µm and fig. 4 = 100 µm." httpUri="https://zenodo.org/record/1010016/files/figure.png" pageId="30" pageNumber="31">Fig. 10</figureCitation>
). Mandible with two distal teeth. Median clypeal lobe trapezoidal, with one angulate tooth; median clypeal carina high in profile, complete or nearly so, convex in profile. Frons strongly coriaceous and punctures large. Vertex crest weakly concave. Pronotal disc coriaceous; anterior margin coarse. Metasoma (
<figureCitation id="2BA42A537C6D9B3AFB1BF01D9A25FDE4" box="[1259,1344,546,571]" captionStart="FIGURES 33 – 44" captionStartId="13.[151,264,1907,1930]" captionTargetBox="[151,1436,553,1886]" captionTargetId="figure@13.[151,1436,553,1886]" captionTargetPageId="13" captionText="FIGURES 33 – 44. Dissomphalus spp., ♂, metasomal tergal process, dorsal view. 33. D. isteus sp. nov. 34. D. azagus sp. nov. 35. D. asfartus sp. nov. 36. D. iozus sp. nov. 37. D. asitius sp. nov. 38. D. ibocius sp. nov. 39. D. tisinus sp. nov. 40. D. ardonius sp. nov. 41. D. arteus sp. nov. 42. D. ettus sp. nov. 43. D. ibrastus sp. nov. 44. D. racarus sp. nov.. Scale bars = 20 µm, except fig. 35 = 200 µm." httpUri="https://zenodo.org/record/1010022/files/figure.png" pageId="30" pageNumber="31">Fig. 41</figureCitation>
). Tergal process with lateral, deep, subcircular and large depression, with small, low and dorsad tubercle placed on anterior area of depression, pit small, with long and sparse tuft of setae. Posterior hypopygeal margin sinuous (
<figureCitation id="2BA42A537C6D9B3AFAC9F0559AF5FD5C" box="[1337,1424,618,643]" captionStart="FIGURES 64 – 75" captionStartId="16.[151,264,1938,1961]" captionTargetBox="[175,1408,606,1889]" captionTargetId="figure@16.[151,1436,569,1917]" captionTargetPageId="16" captionText="FIGURES 64 – 75. Dissomphalus spp., ♂, hypopygium, internal view. 64. D. isteus sp. nov. 65. D. azagus sp. nov. 66. D. asfartus sp. nov. 67. D. iozus sp. nov. 68. D. asitius sp. nov. 69. D. ibocius sp. nov. 70. D. tisinus sp. nov. 71. D. ardonius sp. nov. 72. D. arteus sp. nov. 73. D. ettus sp. nov. 74. D. ibrastus sp. nov. 75. D. racarus sp. nov.. Scale bars = 125 µm." httpUri="https://zenodo.org/record/1010028/files/figure.png" pageId="30" pageNumber="31">Fig. 72</figureCitation>
). Genitalia (
<figureCitation id="2BA42A537C6D9B3AFEE0F0B19ED6FD78" box="[272,435,654,679]" captionStart="FIGURES 136 – 147" captionStartId="31.[151,264,1782,1805]" captionTargetBox="[157,1424,384,1751]" captionTargetId="figure@31.[151,1436,373,1762]" captionTargetPageId="31" captionText="FIGURES 136 – 147. Dissomphalus spp., ♂, genitalia, except fig. 147 apical lobes of aedeagal dorsal body. 136 – 137. D. arteus sp. nov. 136. Dorsal view. 137. Ventral view. 138 – 139. D. ettus sp. nov. 138. Dorsal view. 139. Ventral view. 140 – 141. D. ibrastus sp. nov. 140. Dorsal view. 141. Ventral view. 142 – 144. D. racarus sp. nov. 142. Dorsal view. 143. Ventral view. 143. Lateral view. 145 – 147. D. teraus sp. nov. 145. Dorsal view. 146. Ventral view. 147. Apical lobes of aedeagal dorsal body, ventral view. Scale bars = 250 µm." httpUri="https://zenodo.org/record/1010040/files/figure.png" pageId="30" pageNumber="31">Figs 136–137</figureCitation>
): paramere with dorsal margin entirely wide; apical margin truncate. Aedeagal ventral ramus shorter than dorsal body, wide, abruptly narrowing apicad; cross section laminar; surface vertical; outer margin and inner margin slightly sinuous; apex long, simple, parallel; additional inner ramus long; basal stub short and narrow. Aedeagal dorsal body with pair of apical lobe; outer lobe long, vertical and very narrow, with apical margin crenulate and ventrad; inner pair stout, membranous and setose. Apodeme not extending beyond genital ring.
</paragraph>
<paragraph id="B32036D67C6D9B3AFF37F1599B4FFCA0" blockId="30.[151,1437,438,1327]" box="[199,1066,870,895]" pageId="30" pageNumber="31">
<emphasis id="81EBEAC47C6D9B3AFF37F1599E59FCA0" bold="true" box="[199,316,870,895]" pageId="30" pageNumber="31">Variation.</emphasis>
Tergal process with very shallow depression and without tubercle.
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C6D9B3AFF37F1B59BD7FB88" pageId="30" pageNumber="31" type="discussion">
<paragraph id="B32036D67C6D9B3AFF37F1B59BD7FB88" blockId="30.[151,1437,438,1327]" pageId="30" pageNumber="31">
<emphasis id="81EBEAC47C6D9B3AFF37F1B59E5EFC7C" bold="true" box="[199,315,906,931]" pageId="30" pageNumber="31">Remarks.</emphasis>
This species is similar to the
<taxonomicName id="749F4D557C6D9B3AFD51F1B39C7BFC7C" authority="Azevedo, 2017" authorityName="Azevedo" authorityYear="2017" box="[673,798,908,931]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="racarus" status="sp. nov.">
<emphasis id="81EBEAC47C6D9B3AFD51F1B39C7BFC7C" box="[673,798,908,931]" italics="true" pageId="30" pageNumber="31">D. racarus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9AD857BF7C6D9B3AFCD9F1B59CE3FC7C" box="[809,902,906,931]" pageId="30" pageNumber="31" rank="species">
<emphasis id="81EBEAC47C6D9B3AFCD9F1B59CE3FC7C" bold="true" box="[809,902,906,931]" pageId="30" pageNumber="31">sp. nov.</emphasis>
</taxonomicNameLabel>
by having the dorsal margin of the paramere sinuous, the apex of the basal bar projected forward and the inner margin of the aedeagal ventral ramus crenulate. However, this species has the apex of the basal bar smooth, the ventral margin of the apical lobes of the aedeagal dorsal body not excavated and the apical margin of the aedeagal ventral ramus smooth, whereas
<taxonomicName id="749F4D557C6D9B3AFB05F1C89A14FBD1" authority="Azevedo, 2017" authorityName="Azevedo" authorityYear="2017" box="[1269,1393,1015,1038]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="racarus" status="sp. nov.">
<emphasis id="81EBEAC47C6D9B3AFB05F1C89A14FBD1" box="[1269,1393,1015,1038]" italics="true" pageId="30" pageNumber="31">D. racarus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9AD857BF7C6D9B3AFA8AF1C99FADFBEC" pageId="30" pageNumber="31" rank="species">
<emphasis id="81EBEAC47C6D9B3AFA8AF1C99FADFBEC" bold="true" pageId="30" pageNumber="31">sp. nov.</emphasis>
</taxonomicNameLabel>
has the apex of basal bar crenulate, the ventral margin of the apical lobes of the aedeagal dorsal body deeply excavated and the apical margin of the aedeagal ventral ramus with a small projection dorsad.
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C6D9B3AFF37F65D9D7BFAD4" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="B32036D67C6D9B3AFF37F65D9D7BFAD4" blockId="30.[151,1437,438,1327]" pageId="30" pageNumber="31">
<materialsCitation id="03F73C8B7C6D9B3AFF37F65D9AFCFB40" ID-GBIF-Occurrence="1671715974" collectingDate="1999-10-17" collectingDateMax="1999-10-24" collectingDateMin="1999-10-17" collectionCode="MIUP" collectorName="Teribe, P &amp; Tr &amp; Malaise, A" country="Panama" latitude="9.406944" location="Wekso" longLatPrecision="21" longitude="-82.93889" municipality="Material" pageId="30" pageNumber="31" specimenCount="1" specimenCount-male="1" stateProvince="Bocas del Toro" typeStatus="holotype">
<emphasis id="81EBEAC47C6D9B3AFF37F65D9ED6FBA4" bold="true" box="[199,435,1122,1147]" pageId="30" pageNumber="31">
<collectingMunicipality id="5344ACAC7C6D9B3AFF37F65D9E4AFBA4" box="[199,303,1122,1147]" pageId="30" pageNumber="31">Material</collectingMunicipality>
examined.
</emphasis>
<typeStatus id="6C2488747C6D9B3AFE4DF65D9D40FBA4" box="[445,549,1122,1147]" pageId="30" pageNumber="31" type="holotype">Holotype</typeStatus>
<specimenCount id="A599FD5F7C6D9B3AFDDFF65C9D23FBA5" box="[559,582,1123,1146]" pageId="30" pageNumber="31" type="male">♂</specimenCount>
,
<collectingCountry id="CB8876467C6D9B3AFDA5F65C9DAAFBA4" box="[597,719,1123,1147]" name="Panama" pageId="30" pageNumber="31">PANAMA</collectingCountry>
,
<collectingRegion id="715BF8347C6D9B3AFD2DF65B9CEEFBA4" box="[733,907,1122,1147]" country="Panama" name="Bocas del Toro" pageId="30" pageNumber="31">
<emphasis id="81EBEAC47C6D9B3AFD2DF65B9CEEFBA4" box="[733,907,1122,1147]" italics="true" pageId="30" pageNumber="31">Bocas del Toro</emphasis>
</collectingRegion>
:
<location id="B640600D7C6D9B3AFC6CF65D9C8EFBA4" LSID="urn:lsid:plazi:treatment:3B3687C07C6D9B3AFF67F3739C1CFAF0:B640600D7C6D9B3AFC6CF65D9C8EFBA4" box="[924,1003,1122,1147]" country="Panama" latitude="9.406944" longLatPrecision="21" longitude="-82.93889" municipality="Material" name="Wekso" pageId="30" pageNumber="31" stateProvince="Bocas del Toro">Wekso</location>
,
<collectorName id="1E6A53007C6D9B3AFC09F65D9B04FBA5" box="[1017,1121,1122,1147]" pageId="30" pageNumber="31">Teribe, P</collectorName>
[arque] I[nternacional] L[a] A[mistad], 50 msnm, [
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,
<geoCoordinate id="D6AB50117C6D9B3AFDC4F6B99DA4FB40" box="[564,705,1158,1183]" direction="west" orientation="longitude" pageId="30" pageNumber="31" precision="15" value="-82.93889">82º56'20&quot;W</geoCoordinate>
],
<collectingDate id="D765E9FE7C6D9B3AFD27F6B89CF2FB40" box="[727,919,1159,1183]" pageId="30" pageNumber="31" value="1999-10-17" valueMax="1999-10-24" valueMin="1999-10-17">
<date id="C72110167C6D9B3AFD27F6B89CF2FB40" box="[727,919,1159,1183]" pageId="30" pageNumber="31" value="1999-10-17" valueMax="1999-10-24" valueMin="1999-10-17">17–24 oct. 1999</date>
</collectingDate>
,
<collectorName id="1E6A53007C6D9B3AFC57F6B89CA6FB41" box="[935,963,1159,1182]" pageId="30" pageNumber="31">Tr</collectorName>
[ampa]
<collectorName id="1E6A53007C6D9B3AFBEAF6B99BF9FB40" box="[1050,1180,1158,1183]" pageId="30" pageNumber="31">Malaise, A</collectorName>
. Santos[col.] (
<collectionCode id="D58EAE137C6D9B3AFAB4F6B89AF5FB40" box="[1348,1424,1159,1183]" pageId="30" pageNumber="31">MIUP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="03F73C8B7C6D9B3AFF67F6949D20FB1C" ID-GBIF-Occurrence="1671715995" box="[151,581,1194,1219]" collectingDate="1999-10-17" collectingDateMax="1999-10-24" collectingDateMin="1999-10-17" collectionCode="MIUP" collectorName="Teribe, P &amp; Tr &amp; Malaise, A" country="Panama" latitude="9.406944" location="Wekso" longLatPrecision="21" longitude="-82.93889" municipality="Material" pageId="30" pageNumber="31" specimenCount="7" specimenCount-male="7" stateProvince="Bocas del Toro" typeStatus="paratype">
<typeStatus id="6C2488747C6D9B3AFF67F6949E6CFB1C" box="[151,265,1195,1219]" pageId="30" pageNumber="31" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="A599FD5F7C6D9B3AFEE4F6949E5CFB1D" box="[276,313,1195,1218]" pageId="30" pageNumber="31" type="male">7♂</specimenCount>
, same data as holotype
</materialsCitation>
;
<materialsCitation id="03F73C8B7C6D9B3AFDA2F6959D7FFAD4" ID-GBIF-Occurrence="1671715983" collectingDate="1999-09-26" collectingDateMax="1999-09-30" collectingDateMin="1999-09-26" collectionCode="MIUP" collectorName="El Cope &amp; de Division &amp; Omar Torrijos H &amp; Tr &amp; Malaise, A &amp; Santos, P" country="Panama" county="El Cope" elevation="620" latitude="8.698889" location="Nac" longLatPrecision="21" longitude="-80.654724" municipality="Par" pageId="30" pageNumber="31" specimenCount="1" specimenCount-male="1" stateProvince="Cocle Pr" typeStatus="paratype">
<emphasis id="81EBEAC47C6D9B3AFDA2F6959C45FB1D" box="[594,800,1194,1219]" italics="true" pageId="30" pageNumber="31">
<collectingRegion id="715BF8347C6D9B3AFDA2F6959DDDFB1D" box="[594,696,1194,1219]" isEnumeration="true" pageId="30" pageNumber="31">
<collectingRegion id="715BF8347C6D9B3AFDA2F6959DF6FB1C" box="[594,659,1194,1219]" country="Panama" name="Cocle" pageId="30" pageNumber="31">Coclé</collectingRegion>
<collectingRegion id="715BF8347C6D9B3AFD6CF6949DDDFB1D" box="[668,696,1195,1218]" country="Brazil" name="Para" pageId="30" pageNumber="31">Pr</collectingRegion>
</collectingRegion>
[ovincia]
</emphasis>
:
<specimenCount id="A599FD5F7C6D9B3AFCC0F6949C31FB1C" box="[816,852,1195,1219]" pageId="30" pageNumber="31" type="male">1♂</specimenCount>
,
<collectorName id="1E6A53007C6D9B3AFC92F6959CA7FB1C" box="[866,962,1194,1219]" pageId="30" pageNumber="31">
<collectingCounty id="5A414E5A7C6D9B3AFC92F6959CA7FB1C" box="[866,962,1194,1219]" pageId="30" pageNumber="31">El Copé</collectingCounty>
</collectorName>
,
<collectingMunicipality id="5344ACAC7C6D9B3AFC3EF6949C92FB1D" box="[974,1015,1195,1218]" pageId="30" pageNumber="31">Par</collectingMunicipality>
[que]
<location id="B640600D7C6D9B3AFBC7F6959B03FB1C" LSID="urn:lsid:plazi:treatment:3B3687C07C6D9B3AFF67F3739C1CFAF0:B640600D7C6D9B3AFBC7F6959B03FB1C" box="[1079,1126,1194,1219]" country="Panama" county="El Cope" latitude="8.698889" longLatPrecision="21" longitude="-80.654724" municipality="Par" name="Nac" pageId="30" pageNumber="31" stateProvince="Cocle Pr">Nac</location>
[ional
<location id="B640600D7C6D9B3AFB5EF6959AF0FB1C" LSID="urn:lsid:plazi:treatment:3B3687C07C6D9B3AFF67F3739C1CFAF0:B640600D7C6D9B3AFB5EF6959AF0FB1C" box="[1198,1429,1194,1219]" country="Panama" county="El Cope" latitude="8.698889" longLatPrecision="21" longitude="-80.654724" municipality="Par" name="General de Division" pageId="30" pageNumber="31" stateProvince="Cocle Pr">
General
<collectorName id="1E6A53007C6D9B3AFAFFF6959AF0FB1C" box="[1295,1429,1194,1219]" pageId="30" pageNumber="31">de División</collectorName>
</location>
]
<collectorName id="1E6A53007C6D9B3AFF67F6F09E3DFB39" box="[151,344,1230,1255]" pageId="30" pageNumber="31">Omar Torrijos H</collectorName>
[errera],
<elevation id="38B2D1E57C6D9B3AFE4CF6F19D27FB39" box="[444,578,1230,1255]" metricMagnitude="2" metricUnit="m" metricValue="6.2" metricValueMax="6.7" metricValueMin="5.7" pageId="30" pageNumber="31" unit="m" value="620.0" valueMax="670.0" valueMin="570.0">
<quantity id="74679B337C6D9B3AFE4CF6F19D27FB39" box="[444,578,1230,1255]" metricMagnitude="2" metricUnit="m" metricValue="6.2" metricValueMax="6.7" metricValueMin="5.7" pageId="30" pageNumber="31" unit="m" value="620.0" valueMax="670.0" valueMin="570.0">570–670 m</quantity>
</elevation>
.s.n.m., [
<geoCoordinate id="D6AB50117C6D9B3AFD50F6F19C43FB38" box="[672,806,1230,1255]" direction="north" orientation="latitude" pageId="30" pageNumber="31" precision="15" value="8.698889">08º41'56&quot;N</geoCoordinate>
,
<geoCoordinate id="D6AB50117C6D9B3AFCC4F6F09CA5FB38" box="[820,960,1231,1255]" direction="west" orientation="longitude" pageId="30" pageNumber="31" precision="15" value="-80.654724">80º39'17&quot;W</geoCoordinate>
],
<collectingDate id="D765E9FE7C6D9B3AFC26F6F19BF2FB38" box="[982,1175,1230,1255]" pageId="30" pageNumber="31" value="1999-09-26" valueMax="1999-09-30" valueMin="1999-09-26">
<date id="C72110167C6D9B3AFC26F6F19BF2FB38" box="[982,1175,1230,1255]" pageId="30" pageNumber="31" value="1999-09-26" valueMax="1999-09-30" valueMin="1999-09-26">26–30 sep. 1999</date>
</collectingDate>
,
<collectorName id="1E6A53007C6D9B3AFB55F6F09BA4FB39" box="[1189,1217,1231,1254]" pageId="30" pageNumber="31">Tr</collectorName>
[ampa]
<collectorName id="1E6A53007C6D9B3AFAE7F6F19AFDFB38" box="[1303,1432,1230,1255]" pageId="30" pageNumber="31">Malaise, A</collectorName>
.
<collectorName id="1E6A53007C6D9B3AFF67F6CC9F9AFAD5" box="[151,255,1267,1291]" pageId="30" pageNumber="31">Santos, P</collectorName>
. González [col.] (
<collectionCode id="D58EAE137C6D9B3AFE35F6CC9D77FAD4" box="[453,530,1267,1291]" pageId="30" pageNumber="31">MIUP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C6D9B3AFF37F7299C1CFAF0" box="[199,889,1302,1327]" pageId="30" pageNumber="31" type="distribution">
<paragraph id="B32036D67C6D9B3AFF37F7299C1CFAF0" blockId="30.[151,1437,438,1327]" box="[199,889,1302,1327]" pageId="30" pageNumber="31">
<emphasis id="81EBEAC47C6D9B3AFF37F7299E3CFAF0" bold="true" box="[199,345,1302,1327]" pageId="30" pageNumber="31">Distribution</emphasis>
(
<figureCitation id="2BA42A537C6D9B3AFE98F7299EAEFAF0" box="[360,459,1302,1327]" captionStart="FIGURES 224 – 233" captionStartId="66.[151,264,1814,1837]" captionTargetBox="[182,1404,199,1768]" captionTargetId="figure@66.[169,1413,193,1773]" captionTargetPageId="66" captionText="FIGURES 224 – 233. Known distribution of Dissomphalus in Panama. 224. Provinces and Counties of Panama. 1. Bocas del Toro. 2. Chiriquí. 3. Ngöbe-Buglé County. 4. Veraguas. 5. Herrera. 6. Los Santos. 7. Colón. 8. Coclé. 9. Panama. 10. Guna Yala County. 11. Darién. 12. Emberá Woonan County. 225. D. abarus sp. nov., D. afestus sp. nov. and D. altivolans. 226. D. angulatus sp. nov., D. ardonius sp. nov. and D. aretisus sp. nov.. 227. D. apertus, D. areius sp. nov. and D. arteobius sp. nov.. 228. D. arbeius sp. nov., D. asitius sp. nov. and D. brasiliensis. 229. D. arteus sp. nov., D. cervoides and D. culteratus. 230. D. asfartus sp. nov.. 231. D. azagus sp. nov., D. bilobatus and D. coronatus. 232. D. borus sp. nov., D. citus sp. nov. and D. curvifoveatus. 233. D. declinatus, D. dilatatus and D. ettus sp. nov.." httpUri="https://zenodo.org/record/1010056/files/figure.png" pageId="30" pageNumber="31">Fig. 229</figureCitation>
).
<collectingRegion id="715BF8347C6D9B3AFE2EF7289D52FAF1" box="[478,567,1303,1326]" country="Panama" name="Panama" pageId="30" pageNumber="31">
<collectingCountry id="CB8876467C6D9B3AFE2EF7289D52FAF1" box="[478,567,1303,1326]" name="Panama" pageId="30" pageNumber="31">Panama</collectingCountry>
</collectingRegion>
(
<collectingRegion id="715BF8347C6D9B3AFDB6F7289D95FAF1" box="[582,752,1302,1327]" country="Panama" name="Bocas del Toro" pageId="30" pageNumber="31">Bocas del Toro</collectingRegion>
and
<collectingRegion id="715BF8347C6D9B3AFCD6F7299C09FAF0" box="[806,876,1302,1327]" country="Panama" name="Cocle" pageId="30" pageNumber="31">Coclé</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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