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<document id="05447A2F7F4BB52308522C92AE9329A2" ID-CLB-Dataset="31852" ID-DOI="10.11646/zootaxa.4335.1.1" ID-GBIF-Dataset="f2ad3cb1-0bad-4be1-9e36-2170a6a08aaf" ID-ISSN="1175-5326" ID-Zenodo-Dep="1010014" ID-ZooBank="3429592D-23A8-4EBF-8412-BD57B3BC4314" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1507877826645" checkinUser="plazi" docAuthor="Azevedo, Celso O." docDate="2017" docId="3B3687C07C579B01FF67F5369DD0FD80" docLanguage="en" docName="zootaxa.4335.1.1.pdf" docOrigin="Zootaxa 4335 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Dissomphalus culteratus Alencar &amp; Azevedo 2006" docType="treatment" docVersion="15" lastPageNumber="38" masterDocId="C70FFFB87C739B24FFF0F23F9F65FFDF" masterDocTitle="Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species" masterLastPageNumber="72" masterPageNumber="1" pageNumber="37" updateTime="1698455438235" updateUser="plazi">
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<mods:title id="2FF5CC75439BF2C4AC5CB061BCB2992B">Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species</mods:title>
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<mods:namePart id="23208B41FC55F5D9D57E60ACD3B5B950">Azevedo, Celso O.</mods:namePart>
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<mods:date id="663306A51FEB483AFFBBE02B0275A48C">2017</mods:date>
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<treatment id="3B3687C07C579B01FF67F5369DD0FD80" ID-DOI="http://doi.org/10.5281/zenodo.5020400" ID-GBIF-Taxon="135435297" ID-Zenodo-Dep="5020400" LSID="urn:lsid:plazi:treatment:3B3687C07C579B01FF67F5369DD0FD80" httpUri="http://treatment.plazi.org/id/3B3687C07C579B01FF67F5369DD0FD80" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">
<subSubSection id="FB85655D7C579B00FF67F5369C62F8FC" box="[151,775,1801,1827]" pageId="36" pageNumber="37" type="nomenclature">
<paragraph id="B32036D67C579B00FF67F5369C62F8FC" blockId="36.[151,775,1801,1827]" box="[151,775,1801,1827]" pageId="36" pageNumber="37">
<heading id="E86881BA7C579B00FF67F5369C62F8FC" bold="true" box="[151,775,1801,1827]" fontSize="11" level="1" pageId="36" pageNumber="37" reason="1">
<taxonomicName id="749F4D557C579B00FF67F5369C62F8FC" authority="Alencar &amp; Azevedo, 2006" authorityName="Alencar &amp; Azevedo" authorityYear="2006" box="[151,775,1801,1827]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="culteratus">
<emphasis id="81EBEAC47C579B00FF67F5369C62F8FC" bold="true" box="[151,775,1801,1827]" pageId="36" pageNumber="37">
<emphasis id="81EBEAC47C579B00FF67F5369EA4F8FC" bold="true" box="[151,449,1801,1827]" italics="true" pageId="36" pageNumber="37">Dissomphalus culteratus</emphasis>
<bibRefCitation id="D70E4B277C579B00FE39F5369C62F8FC" author="Alencar" box="[457,775,1801,1827]" pageId="36" pageNumber="37" refString="Alencar, I. D. C. C. &amp; Azevedo, C. O. (2006) Definition of Neotropical coronatus species-group (Hymenoptera: Bethylidae, Dissomphalus) with description of thirteen new species. Zootaxa, 1330, 1 - 26." type="journal article" year="2006">Alencar &amp; Azevedo, 2006</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C579B00FF67F56F9BB9F8B9" box="[151,1244,1872,1894]" pageId="36" pageNumber="37" type="reference_group">
<paragraph id="B32036D67C579B00FF67F56F9BB9F8B9" blockId="36.[151,1244,1872,1894]" box="[151,1244,1872,1894]" pageId="36" pageNumber="37">
<materialsCitation id="03F73C8B7C579B00FF67F56F9BB9F8B9" ID-GBIF-Occurrence="1671715872" box="[151,1244,1872,1894]" collectionCode="PMAE" country="Panama" location="Panama" pageId="36" pageNumber="37" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<taxonomicName id="749F4D557C579B00FF67F56F9DC3F8B9" authority="Alencar &amp; Azevedo, 2006" authorityName="Alencar &amp; Azevedo" authorityYear="2006" box="[151,678,1872,1894]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="culteratus">
<emphasis id="81EBEAC47C579B00FF67F56F9EF7F8B9" box="[151,402,1872,1894]" italics="true" pageId="36" pageNumber="37">Dissomphalus culteratus</emphasis>
<bibRefCitation id="D70E4B277C579B00FE68F56F9DC3F8B9" author="Alencar" box="[408,678,1872,1894]" pageId="36" pageNumber="37" refString="Alencar, I. D. C. C. &amp; Azevedo, C. O. (2006) Definition of Neotropical coronatus species-group (Hymenoptera: Bethylidae, Dissomphalus) with description of thirteen new species. Zootaxa, 1330, 1 - 26." type="journal article" year="2006">Alencar &amp; Azevedo, 2006</bibRefCitation>
</taxonomicName>
: 0911 (
<specimenCount id="A599FD5F7C579B00FD0CF56F9C74F8B9" box="[764,785,1872,1894]" pageId="36" pageNumber="37" type="male"></specimenCount>
,
<typeStatus id="6C2488747C579B00FCEBF56F9C11F8B9" box="[795,884,1872,1894]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
from
<collectingCountry id="CB8876467C579B00FC42F56E9B60F8B9" box="[946,1029,1873,1894]" name="Panama" pageId="36" pageNumber="37">Panama</collectingCountry>
,
<collectionCode id="D58EAE137C579B00FBFFF56F9B3DF8BA" box="[1039,1112,1872,1893]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15193" name="Royal Alberta Museum" pageId="36" pageNumber="37">PMAE</collectionCode>
, figs 1011).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C579B01FF67F5AD9BE5FEC2" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37" type="diagnosis">
<paragraph id="B32036D67C579B01FF67F5AD9BE5FEC2" blockId="36.[151,1436,1938,2035]" lastBlockId="37.[151,1436,151,608]" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">
<emphasis id="81EBEAC47C579B00FF67F5AD9E74F874" bold="true" box="[151,273,1938,1963]" pageId="36" pageNumber="37">Diagnosis.</emphasis>
Mandible tridentate. Clypeus trapezoidal. Tergal process with pair of circular, large and deep lateral depressions with dense tuft of convergent setae directed backward. Hypopygium with posterior margin with median convexity. Genitalia: paramere with dorsal margin straight and ventral margin concave; aedeagal ventral ramus shorter than aedeagal dorsal body, laminar, surface horizontal and arched ventrally; basal half wide; inner margin sinuous; apical margin with outer margin straight and inner margin microserrate; aedeagal dorsal body wide, narrowing to apex; apical part with concavity mesad, apex with two arms; crown-process with long lateral prolongation in middle of aedeagal dorsal body. Apodeme extending beyond genital ring.
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C569B01FF37F3189B47FEBA" pageId="37" pageNumber="38" type="discussion">
<paragraph id="B32036D67C569B01FF37F3189B47FEBA" blockId="37.[151,1436,151,608]" pageId="37" pageNumber="38">
<emphasis id="81EBEAC47C569B01FF37F3189E5EFE9F" bold="true" box="[199,315,295,320]" pageId="37" pageNumber="38">Remarks.</emphasis>
This species is known to
<collectingRegion id="715BF8347C569B01FD83F3179DA9FEE0" box="[627,716,296,319]" country="Panama" name="Panama" pageId="37" pageNumber="38">
<collectingCountry id="CB8876467C569B01FD83F3179DA9FEE0" box="[627,716,296,319]" name="Panama" pageId="37" pageNumber="38">Panama</collectingCountry>
</collectingRegion>
(Alencar &amp; Azevedo). In this series, the specimens are quite similar to those previously studied, without any variation that worths to mention.
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C569B01FF37F3509D91FDE3" pageId="37" pageNumber="38" type="materials_examined">
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<emphasis id="81EBEAC47C569B01FF37F3509ED6FE57" bold="true" box="[199,435,367,392]" pageId="37" pageNumber="38">Material examined.</emphasis>
<typeStatus id="6C2488747C569B01FE4DF3509D4DFE57" box="[445,552,367,392]" pageId="37" pageNumber="38" type="holotype">Holotype</typeStatus>
,
<specimenCount id="A599FD5F7C569B01FDC6F34F9D28FE58" box="[566,589,368,391]" pageId="37" pageNumber="38" type="male"></specimenCount>
,
<collectingRegion id="715BF8347C569B01FDACF34F9DB3FE57" box="[604,726,368,392]" country="Panama" name="Panama" pageId="37" pageNumber="38">
<collectingCountry id="CB8876467C569B01FDACF34F9DB3FE57" box="[604,726,368,392]" name="Panama" pageId="37" pageNumber="38">PANAMA</collectingCountry>
</collectingRegion>
,
<collectingRegion id="715BF8347C569B01FD14F3509C23FE57" box="[740,838,367,392]" country="Panama" name="Chiriqui" pageId="37" pageNumber="38">Chiriquí</collectingRegion>
: Hatmans Farm, [
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],
<date id="C72110167C569B01FB07F34F9A11FE57" box="[1271,1396,368,392]" pageId="37" pageNumber="38" value="1981-07-03">3.VII.1981</date>
, B. Gill col (
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);
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<collectingCountry id="CB8876467C569B01FE94F3AB9EBBFE73" box="[356,478,404,428]" name="Panama" pageId="37" pageNumber="38">PANAMA</collectingCountry>
</collectingRegion>
,
<emphasis id="81EBEAC47C569B01FE1AF3AB9DB0FE73" box="[490,725,404,429]" italics="true" pageId="37" pageNumber="38">
<collectingRegion id="715BF8347C569B01FE1AF3AB9D2CFE72" box="[490,585,404,429]" country="Panama" name="Chiriqui" pageId="37" pageNumber="38">Chiriqui</collectingRegion>
<collectingRegion id="715BF8347C569B01FDA1F3AA9D08FE73" box="[593,621,405,428]" country="Brazil" name="Para" pageId="37" pageNumber="38">Pr</collectingRegion>
[ovincia]
</emphasis>
:
<specimenCount id="A599FD5F7C569B01FD14F3AB9C6FFE73" box="[740,778,404,428]" pageId="37" pageNumber="38" type="male">2♂</specimenCount>
, P[arque] I[nternacional] L[a] A[mistad], Jurutungo,
<quantity id="74679B337C569B01FA93F3AB9FCAFE10" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="37" pageNumber="38" unit="m" value="1800.0">1800 m</quantity>
, [
<geoCoordinate id="D6AB50117C569B01FF32F3889E46FE0F" box="[194,291,439,464]" direction="north" orientation="latitude" pageId="37" pageNumber="38" precision="925" value="8.9">08°54'N</geoCoordinate>
,
<geoCoordinate id="D6AB50117C569B01FEDFF3879EF2FE0F" box="[303,407,440,464]" direction="west" orientation="longitude" pageId="37" pageNumber="38" precision="925" value="-82.73333">82°44'W</geoCoordinate>
], 30oci.
<date id="C72110167C569B01FE1FF3889D03FE0F" box="[495,614,439,464]" pageId="37" pageNumber="38" value="1994-11-05">5nov.1994</date>
, Tr[ampa] Malaise, A. Rodriguez [col.] (
<collectionCode id="D58EAE137C569B01FBC5F3879BE4FE0F" box="[1077,1153,440,464]" pageId="37" pageNumber="38">MIUP</collectionCode>
);
<specimenCount id="A599FD5F7C569B01FB65F3879BDFFE0F" box="[1173,1210,440,464]" pageId="37" pageNumber="38" type="male">9♂</specimenCount>
,
<quantity id="74679B337C569B01FB36F3889A70FE0F" box="[1222,1301,439,464]" metricMagnitude="3" metricUnit="m" metricValue="5.6" pageId="37" pageNumber="38" unit="km" value="5.6">5.6 km</quantity>
N Boquete, La
<collectingRegion id="715BF8347C569B01FF4FF3E39E7DFE2A" box="[191,280,476,501]" country="Puerto Rico" name="Culebra" pageId="37" pageNumber="38">Culebra</collectingRegion>
trail, [
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,
<geoCoordinate id="D6AB50117C569B01FDF0F3E39DEAFE2A" box="[512,655,476,501]" direction="west" orientation="longitude" pageId="37" pageNumber="38" precision="15" value="-82.42166">82°25'18&quot;W</geoCoordinate>
], (
<specimenCount id="A599FD5F7C569B01FD5EF3E39DB7FE2B" box="[686,722,476,500]" pageId="37" pageNumber="38" type="male">8♂</specimenCount>
) and [
<geoCoordinate id="D6AB50117C569B01FCD0F3E39CC4FE2A" box="[800,929,476,501]" direction="north" orientation="latitude" pageId="37" pageNumber="38" precision="92" value="8.911667">08° 54,7'N</geoCoordinate>
, 82° 47,3 W],
<date id="C72110167C569B01FBA3F3E39B92FE2A" box="[1107,1271,476,501]" pageId="37" pageNumber="38" value="1996-06-17" valueMax="1996-06-18" valueMin="1996-06-17">
1718.vi.
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</date>
, 1875m (
<specimenCount id="A599FD5F7C569B01FA99F3E39AE9FE2B" box="[1385,1420,476,500]" pageId="37" pageNumber="38" type="male">1♂</specimenCount>
), F[light] I[nterception] T[rap], J. Ashe &amp; R. Brooks [col.] (
<collectionCode id="D58EAE137C569B01FCB9F03F9CF7FDC7" box="[841,914,512,536]" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="37" pageNumber="38">CNCI</collectionCode>
);
<emphasis id="81EBEAC47C569B01FC59F03F9B58FDC8" box="[937,1085,512,535]" italics="true" pageId="37" pageNumber="38">
[
<collectingRegion id="715BF8347C569B01FC44F03F9B57FDC8" box="[948,1074,512,535]" country="Panama" name="Los Santos" pageId="37" pageNumber="38">Los Santos</collectingRegion>
]
</emphasis>
:
<specimenCount id="A599FD5F7C569B01FBA0F03F9B10FDC7" box="[1104,1141,512,536]" pageId="37" pageNumber="38" type="male">1♂</specimenCount>
, Ojo de Agua, [
<geoCoordinate id="D6AB50117C569B01FAC6F03F9AFDFDC7" box="[1334,1432,512,536]" direction="north" orientation="latitude" pageId="37" pageNumber="38" precision="925" value="7.5166664">07°31'N</geoCoordinate>
,
<geoCoordinate id="D6AB50117C569B01FF67F01B9F9AFDE2" box="[151,255,548,573]" direction="west" orientation="longitude" pageId="37" pageNumber="38" precision="925" value="-80.416664">80°25'W</geoCoordinate>
],
<date id="C72110167C569B01FEE1F01B9E8EFDE3" box="[273,491,548,572]" pageId="37" pageNumber="38" value="1982-06-30" valueMax="1982-07-14" valueMin="1982-06-30">30.VI14.VII.1982</date>
, B. Gill [col.] (
<collectionCode id="D58EAE137C569B01FD64F01B9D8CFDE3" box="[660,745,548,572]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15193" name="Royal Alberta Museum" pageId="37" pageNumber="38">PMAE</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C569B01FF37F0789DD0FD80" box="[199,693,583,608]" pageId="37" pageNumber="38" type="distribution">
<paragraph id="B32036D67C569B01FF37F0789DD0FD80" blockId="37.[151,1436,151,608]" box="[199,693,583,608]" pageId="37" pageNumber="38">
<emphasis id="81EBEAC47C569B01FF37F0789E3CFDBF" bold="true" box="[199,345,583,608]" pageId="37" pageNumber="38">Distribution</emphasis>
(
<figureCitation id="2BA42A537C569B01FE98F0789EAEFDBF" box="[360,459,583,608]" captionStart="FIGURES 224 233" captionStartId="66.[151,264,1814,1837]" captionTargetBox="[182,1404,199,1768]" captionTargetId="figure@66.[169,1413,193,1773]" captionTargetPageId="66" captionText="FIGURES 224 233. Known distribution of Dissomphalus in Panama. 224. Provinces and Counties of Panama. 1. Bocas del Toro. 2. Chiriquí. 3. Ngöbe-Buglé County. 4. Veraguas. 5. Herrera. 6. Los Santos. 7. Colón. 8. Coclé. 9. Panama. 10. Guna Yala County. 11. Darién. 12. Emberá Woonan County. 225. D. abarus sp. nov., D. afestus sp. nov. and D. altivolans. 226. D. angulatus sp. nov., D. ardonius sp. nov. and D. aretisus sp. nov.. 227. D. apertus, D. areius sp. nov. and D. arteobius sp. nov.. 228. D. arbeius sp. nov., D. asitius sp. nov. and D. brasiliensis. 229. D. arteus sp. nov., D. cervoides and D. culteratus. 230. D. asfartus sp. nov.. 231. D. azagus sp. nov., D. bilobatus and D. coronatus. 232. D. borus sp. nov., D. citus sp. nov. and D. curvifoveatus. 233. D. declinatus, D. dilatatus and D. ettus sp. nov.." httpUri="https://zenodo.org/record/1010056/files/figure.png" pageId="37" pageNumber="38">Fig. 229</figureCitation>
).
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and
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<collectingCountry id="CB8876467C569B01FDA4F0779DD5FD80" box="[596,688,584,607]" name="Panama" pageId="37" pageNumber="38">Panama</collectingCountry>
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.
</paragraph>
</subSubSection>
</treatment>
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