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<document id="79B6439AA738E4A82B144062B5B29391" ID-CLB-Dataset="31852" ID-DOI="10.11646/zootaxa.4335.1.1" ID-GBIF-Dataset="f2ad3cb1-0bad-4be1-9e36-2170a6a08aaf" ID-ISSN="1175-5326" ID-Zenodo-Dep="1010014" ID-ZooBank="3429592D-23A8-4EBF-8412-BD57B3BC4314" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1507877826645" checkinUser="plazi" docAuthor="Azevedo, Celso O." docDate="2017" docId="3B3687C07C529B06FF67F1989B7BFE73" docLanguage="en" docName="zootaxa.4335.1.1.pdf" docOrigin="Zootaxa 4335 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Dissomphalus teraus Azevedo 2017, sp. nov." docType="treatment" docVersion="15" lastPageNumber="35" masterDocId="C70FFFB87C739B24FFF0F23F9F65FFDF" masterDocTitle="Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species" masterLastPageNumber="72" masterPageNumber="1" pageNumber="34" updateTime="1698455438235" updateUser="plazi">
<mods:mods id="25D34315E1370610DAEAB2DBA83F468C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="E9D1DE7C950AD9ED6D9FDBC71FDEF406">Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species</mods:title>
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<mods:namePart id="DDE6B2ECC06ADBEE341FFF2917472CD0">Azevedo, Celso O.</mods:namePart>
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<mods:title id="35E3AAA1FB1ACD51D29FB1B1E55024B9">Zootaxa</mods:title>
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<mods:part id="166B55B652B1964EB23813F6E7A470E3">
<mods:date id="15D812DECEA54C03C4DCCFF94AC8FBE8">2017</mods:date>
<mods:detail id="963687271734A5DAA182D9818314CC18" type="pubDate">
<mods:number id="61E988CF2CB9A67C29368459A4CEF959">2017-10-13</mods:number>
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<mods:number id="7733A12513A2AF00F0DB465B01494CF8">4335</mods:number>
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<mods:identifier id="3589DD21F165B6777B7ABA09F6C2CF26" type="DOI">10.11646/zootaxa.4335.1.1</mods:identifier>
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<mods:identifier id="F20476F54DDA5E38E76B084690327EFC" type="ISSN">1175-5326</mods:identifier>
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<treatment id="3B3687C07C529B06FF67F1989B7BFE73" ID-DOI="http://doi.org/10.5281/zenodo.5020386" ID-GBIF-Taxon="135435285" ID-Zenodo-Dep="5020386" LSID="urn:lsid:plazi:treatment:3B3687C07C529B06FF67F1989B7BFE73" httpUri="http://treatment.plazi.org/id/3B3687C07C529B06FF67F1989B7BFE73" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
<subSubSection id="FB85655D7C529B05FF67F1989EA4FC3C" pageId="33" pageNumber="34" type="nomenclature">
<paragraph id="B32036D67C529B05FF67F1989DBFFC1D" blockId="33.[151,730,935,995]" box="[151,730,935,962]" pageId="33" pageNumber="34">
<heading id="E86881BA7C529B05FF67F1989DBFFC1D" bold="true" box="[151,730,935,962]" fontSize="11" level="1" pageId="33" pageNumber="34" reason="1">
<emphasis id="81EBEAC47C529B05FF67F1989DBFFC1D" bold="true" box="[151,730,935,962]" pageId="33" pageNumber="34">
<taxonomicName id="749F4D557C529B05FF67F1989D0AFC1D" authority="Brito &amp; Azevedo" authorityName="Azevedo" authorityYear="2017" box="[151,623,935,962]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="teraus" status="sp. nov.">
<emphasis id="81EBEAC47C529B05FF67F1989EF1FC1E" bold="true" box="[151,404,935,961]" italics="true" pageId="33" pageNumber="34">Dissomphalus teraus</emphasis>
Brito &amp; Azevedo
</taxonomicName>
,
<taxonomicNameLabel id="9AD857BF7C529B05FD8CF1979DBFFC1D" box="[636,730,936,962]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="B32036D67C529B05FF67F1F59EA4FC3C" blockId="33.[151,730,935,995]" box="[151,449,970,995]" pageId="33" pageNumber="34">
(
<figureCitation id="2BA42A537C529B05FF6FF1F59F90FC3C" box="[159,245,970,995]" captionStart="FIGURES 13 – 24" captionStartId="11.[151,264,1615,1638]" captionTargetBox="[151,1436,306,1594]" captionTargetId="figure@11.[151,1436,303,1594]" captionTargetPageId="11" captionText="FIGURES 13 – 24. Dissomphalus spp., ♂, head, dorsal view. 13. D. racarus sp. nov. 14. D. teraus sp. nov. 15. D. urazius sp. nov. 16. D. afestus sp. nov. 17. D. arbeius sp. nov. 18. D. raziogus sp. nov. 19. D. areius sp. nov. 20. D. aretisus sp. nov. 21. D. citus sp. nov. 22. D. sartus sp. nov. 23. D. teobus sp. nov. 24. D. terbius sp. nov.. Scale bars = 20 µm, except figs 14, 20 = 10 µm and fig. 15 = 50 µm." httpUri="https://zenodo.org/record/1010018/files/figure.png" pageId="33" pageNumber="34">Figs 14</figureCitation>
,
<figureCitation id="2BA42A537C529B05FEF1F1F59E45FC3C" box="[257,288,970,995]" captionStart="FIGURES 45 – 56" captionStartId="14.[151,264,1544,1567]" captionTargetBox="[151,1436,193,1524]" captionTargetId="figure@14.[151,1436,193,1524]" captionTargetPageId="14" captionText="FIGURES 45 – 56. Dissomphalus spp., ♂, metasomal tergal process, dorsal view. 45. D. teraus sp. nov. 46. D. urazius sp. nov. 47. D. afestus sp. nov. 48. D. arbeius sp. nov. 49. D. raziogus sp. nov. 50. D. areius sp. nov. 51. D. aretisus sp. nov. 52. D. citus sp. nov. 53. D. sartus sp. nov. 54. D. teobus sp. nov. 55. D. terbius sp. nov. 56. D. abarus sp. nov.. Scale bars = 20 µm except, figs 45 and 46 = 50 µm." httpUri="https://zenodo.org/record/1010024/files/figure.png" pageId="33" pageNumber="34">45</figureCitation>
,
<figureCitation id="2BA42A537C529B05FEDBF1F59E2FFC3C" box="[299,330,970,995]" captionStart="FIGURES 76 – 87" captionStartId="17.[151,264,1595,1618]" captionTargetBox="[159,1404,219,1523]" captionTargetId="figure@17.[147,1432,209,1558]" captionTargetPageId="17" captionText="FIGURES 76 – 87. Dissomphalus spp., ♂, hypopygium, internal view. 76. D. teraus sp. nov. 77. D. urazius sp. nov. 78. D. afestus sp. nov. 79. D. arbeius sp. nov. 80. D. raziogus sp. nov. 81. D. areius sp. nov. 82. D. aretisus sp. nov. 83. D. citus sp. nov. 84. D. sartus sp. nov. 85. D. teobus sp. nov. 86. D. terbius sp. nov. 87. D. abarus sp. nov.. Scale bars = 125 µm." httpUri="https://zenodo.org/record/1010030/files/figure.png" pageId="33" pageNumber="34">76</figureCitation>
,
<figureCitation id="2BA42A537C529B05FEA5F1F59EDCFC3C" box="[341,441,970,995]" captionStart-0="FIGURES 136 – 147" captionStart-1="FIGURES 148 – 162" captionStartId-0="31.[151,264,1782,1805]" captionStartId-1="35.[151,264,1605,1628]" captionTargetBox-0="[157,1424,384,1751]" captionTargetBox-1="[157,1414,214,1575]" captionTargetId-0="figure@31.[151,1436,373,1762]" captionTargetId-1="figure@35.[151,1436,195,1584]" captionTargetPageId-0="31" captionTargetPageId-1="35" captionText-0="FIGURES 136 – 147. Dissomphalus spp., ♂, genitalia, except fig. 147 apical lobes of aedeagal dorsal body. 136 – 137. D. arteus sp. nov. 136. Dorsal view. 137. Ventral view. 138 – 139. D. ettus sp. nov. 138. Dorsal view. 139. Ventral view. 140 – 141. D. ibrastus sp. nov. 140. Dorsal view. 141. Ventral view. 142 – 144. D. racarus sp. nov. 142. Dorsal view. 143. Ventral view. 143. Lateral view. 145 – 147. D. teraus sp. nov. 145. Dorsal view. 146. Ventral view. 147. Apical lobes of aedeagal dorsal body, ventral view. Scale bars = 250 µm." captionText-1="FIGURES 148 – 162. Dissomphalus spp., ♂, genitalia, except otherwise indicated. 148 – 153. D. teraus sp. nov. 148. Apical lobes of aedeagal dorsal body, dorsal view (var 1). 149. Apical lobes of aedeagal dorsal body, ventral view (var 1). 150. Aedeagal ventral ramus, ventral view (var 1). 151. Apical lobes of aedeagal dorsal body, dorsal view (var 2). 152. Apical lobes of aedeagal dorsal body, ventral view (var 2). 153. Aedeagal ventral ramus, ventral view (var 2). 154 – 156. D. urazius sp. nov. 154. Dorsal view. 155. Ventral view. 156. Lateral view. 157 – 158. D. afestus sp. nov. 157. Dorsal view. 158. Ventral view. 159 – 162. D. curvifoveatus. 159. Dorsal view. 160. Ventral lobes of aedeagal dorsal body, dorsal view. 161. Outer lobes of aedeagal dorsal body, dorsal view. 162. Dorsal lobes of aedeagal dorsal body, dorsal view. Scale bars = 250 µm." httpUri-0="https://zenodo.org/record/1010040/files/figure.png" httpUri-1="https://zenodo.org/record/1010042/files/figure.png" pageId="33" pageNumber="34">145–153</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C529B05FF67F6309E0BF90A" pageId="33" pageNumber="34" type="description">
<paragraph id="B32036D67C529B05FF67F6309E1DFA6A" blockId="33.[151,1437,1039,2037]" pageId="33" pageNumber="34">
<emphasis id="81EBEAC47C529B05FF67F6309E4CFBF7" bold="true" box="[151,297,1039,1064]" pageId="33" pageNumber="34">Description.</emphasis>
Male. Body length
<quantity id="74679B337C529B05FDF5F62F9D0EFBF8" box="[517,619,1040,1064]" metricMagnitude="-3" metricUnit="m" metricValue="4.33" pageId="33" pageNumber="34" unit="mm" value="4.33">4.33 mm</quantity>
. Color: head and mesosoma dark castaneous; metasoma light castaneous. Head (
<figureCitation id="2BA42A537C529B05FF12F60B9E52FB93" box="[226,311,1076,1101]" captionStart="FIGURES 13 – 24" captionStartId="11.[151,264,1615,1638]" captionTargetBox="[151,1436,306,1594]" captionTargetId="figure@11.[151,1436,303,1594]" captionTargetPageId="11" captionText="FIGURES 13 – 24. Dissomphalus spp., ♂, head, dorsal view. 13. D. racarus sp. nov. 14. D. teraus sp. nov. 15. D. urazius sp. nov. 16. D. afestus sp. nov. 17. D. arbeius sp. nov. 18. D. raziogus sp. nov. 19. D. areius sp. nov. 20. D. aretisus sp. nov. 21. D. citus sp. nov. 22. D. sartus sp. nov. 23. D. teobus sp. nov. 24. D. terbius sp. nov.. Scale bars = 20 µm, except figs 14, 20 = 10 µm and fig. 15 = 50 µm." httpUri="https://zenodo.org/record/1010018/files/figure.png" pageId="33" pageNumber="34">Fig. 14</figureCitation>
). Mandible with two distal teeth. Median clypeal lobe trapezoidal, with one angulate tooth; median clypeal carina high in profile, incomplete apically, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest weakly convex. Pronotal disc coriaceous; anterior margin coarse. Metasoma (
<figureCitation id="2BA42A537C529B05FA9FF6439FD2FB67" captionStart="FIGURES 45 – 56" captionStartId="14.[151,264,1544,1567]" captionTargetBox="[151,1436,193,1524]" captionTargetId="figure@14.[151,1436,193,1524]" captionTargetPageId="14" captionText="FIGURES 45 – 56. Dissomphalus spp., ♂, metasomal tergal process, dorsal view. 45. D. teraus sp. nov. 46. D. urazius sp. nov. 47. D. afestus sp. nov. 48. D. arbeius sp. nov. 49. D. raziogus sp. nov. 50. D. areius sp. nov. 51. D. aretisus sp. nov. 52. D. citus sp. nov. 53. D. sartus sp. nov. 54. D. teobus sp. nov. 55. D. terbius sp. nov. 56. D. abarus sp. nov.. Scale bars = 20 µm except, figs 45 and 46 = 50 µm." httpUri="https://zenodo.org/record/1010024/files/figure.png" pageId="33" pageNumber="34">Fig. 45</figureCitation>
). Tergal process with lateral, shallow, circular and large depression, with small, low and laterad tubercle placed on inner area of depression, pit small, with short tuft of setae. Posterior hypopygeal margin sinuous (
<figureCitation id="2BA42A537C529B05FAC8F6FB9AF5FB02" box="[1336,1424,1220,1245]" captionStart="FIGURES 76 – 87" captionStartId="17.[151,264,1595,1618]" captionTargetBox="[159,1404,219,1523]" captionTargetId="figure@17.[147,1432,209,1558]" captionTargetPageId="17" captionText="FIGURES 76 – 87. Dissomphalus spp., ♂, hypopygium, internal view. 76. D. teraus sp. nov. 77. D. urazius sp. nov. 78. D. afestus sp. nov. 79. D. arbeius sp. nov. 80. D. raziogus sp. nov. 81. D. areius sp. nov. 82. D. aretisus sp. nov. 83. D. citus sp. nov. 84. D. sartus sp. nov. 85. D. teobus sp. nov. 86. D. terbius sp. nov. 87. D. abarus sp. nov.. Scale bars = 125 µm." httpUri="https://zenodo.org/record/1010030/files/figure.png" pageId="33" pageNumber="34">Fig. 76</figureCitation>
). Genitalia (
<figureCitation id="2BA42A537C529B05FEE0F6D89ED6FADF" box="[272,435,1255,1280]" captionStart-0="FIGURES 136 – 147" captionStart-1="FIGURES 148 – 162" captionStartId-0="31.[151,264,1782,1805]" captionStartId-1="35.[151,264,1605,1628]" captionTargetBox-0="[157,1424,384,1751]" captionTargetBox-1="[157,1414,214,1575]" captionTargetId-0="figure@31.[151,1436,373,1762]" captionTargetId-1="figure@35.[151,1436,195,1584]" captionTargetPageId-0="31" captionTargetPageId-1="35" captionText-0="FIGURES 136 – 147. Dissomphalus spp., ♂, genitalia, except fig. 147 apical lobes of aedeagal dorsal body. 136 – 137. D. arteus sp. nov. 136. Dorsal view. 137. Ventral view. 138 – 139. D. ettus sp. nov. 138. Dorsal view. 139. Ventral view. 140 – 141. D. ibrastus sp. nov. 140. Dorsal view. 141. Ventral view. 142 – 144. D. racarus sp. nov. 142. Dorsal view. 143. Ventral view. 143. Lateral view. 145 – 147. D. teraus sp. nov. 145. Dorsal view. 146. Ventral view. 147. Apical lobes of aedeagal dorsal body, ventral view. Scale bars = 250 µm." captionText-1="FIGURES 148 – 162. Dissomphalus spp., ♂, genitalia, except otherwise indicated. 148 – 153. D. teraus sp. nov. 148. Apical lobes of aedeagal dorsal body, dorsal view (var 1). 149. Apical lobes of aedeagal dorsal body, ventral view (var 1). 150. Aedeagal ventral ramus, ventral view (var 1). 151. Apical lobes of aedeagal dorsal body, dorsal view (var 2). 152. Apical lobes of aedeagal dorsal body, ventral view (var 2). 153. Aedeagal ventral ramus, ventral view (var 2). 154 – 156. D. urazius sp. nov. 154. Dorsal view. 155. Ventral view. 156. Lateral view. 157 – 158. D. afestus sp. nov. 157. Dorsal view. 158. Ventral view. 159 – 162. D. curvifoveatus. 159. Dorsal view. 160. Ventral lobes of aedeagal dorsal body, dorsal view. 161. Outer lobes of aedeagal dorsal body, dorsal view. 162. Dorsal lobes of aedeagal dorsal body, dorsal view. Scale bars = 250 µm." httpUri-0="https://zenodo.org/record/1010040/files/figure.png" httpUri-1="https://zenodo.org/record/1010042/files/figure.png" pageId="33" pageNumber="34">Figs 145–153</figureCitation>
): paramere with dorsal margin entirely wide; apical margin truncate. Aedeagal ventral ramus higher than dorsal body, wide, progressively narrowing apicad; cross section laminar; ramous bifid; surface horizontal; inner margin sinuous; outer margin sinuous; apex long, simple, curved outward; additional inner ramus long; basal stub absent. Aedeagal dorsal body with two pairs of apical lobes; outer lobe long, horizontal and narrow, with apical margin rounded and mesad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring.
</paragraph>
<paragraph id="B32036D67C529B05FF37F7809ED2F99A" blockId="33.[151,1437,1039,2037]" pageId="33" pageNumber="34">
<emphasis id="81EBEAC47C529B05FF37F7809E22FA07" bold="true" box="[199,327,1471,1496]" pageId="33" pageNumber="34">Variations.</emphasis>
Head dark castaneous; metassoma castaneous; frons coriaceous; median clypeal lobe subtrapezoidal; depression of tergal process very shallow or without depression, with very low, very small and without tubercle; apical lobe narrow in sickle-shaped with teeth in inner margin and apical lobes with acute apex and inner margin smooth.
</paragraph>
<paragraph id="B32036D67C529B05FF37F4709E0BF90A" blockId="33.[151,1437,1039,2037]" pageId="33" pageNumber="34">This species has two different aedeagal ventral ramus and apical lobes of the aedeagal dorsal body. In one kind, the aedeagal ventral ramus can be lower than the aedeagal dorsal body; the apical margin entirely wide with rounded and dorsad apex; in the other kind the aedeagal ventral ramus can have the apex narrow and two small and laterad projections.</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C529B05FF37F4E09FB0F872" pageId="33" pageNumber="34" type="discussion">
<paragraph id="B32036D67C529B05FF37F4E09FB0F872" blockId="33.[151,1437,1039,2037]" pageId="33" pageNumber="34">
<emphasis id="81EBEAC47C529B05FF37F4E09E5EF927" bold="true" box="[199,315,1759,1784]" pageId="33" pageNumber="34">Remarks.</emphasis>
This species is similar to
<taxonomicName id="749F4D557C529B05FDAEF4DE9D9BF927" authorityName="Azevedo" authorityYear="2001" box="[606,766,1761,1784]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="piscicercus">
<emphasis id="81EBEAC47C529B05FDAEF4DE9D9BF927" box="[606,766,1761,1784]" italics="true" pageId="33" pageNumber="34">D. piscicercus</emphasis>
</taxonomicName>
by having the paramere entirely wide and the basiparamere very wide. However, this species has the tergal process with tubercle, small and shallow depression, the aedeagal ventral ramus with simple apex and the apex of the basal bar straight with two laterad teeth, whereas
<taxonomicName id="749F4D557C529B05FB0DF5169AF9F89F" authorityName="Azevedo" authorityYear="2001" box="[1277,1436,1833,1856]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="piscicercus">
<emphasis id="81EBEAC47C529B05FB0DF5169AF9F89F" box="[1277,1436,1833,1856]" italics="true" pageId="33" pageNumber="34">D. piscicercus</emphasis>
</taxonomicName>
has the tergal process with ovoid, deep, large depressions nearly touching the posterior margin of the tergite I, with pit, tuft of long setae, the aedeagal ventral ramus with two large apical teeth and the basal bar without two laterad teeth.
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C529B06FF37F5879B53FE57" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34" type="materials_examined">
<paragraph id="B32036D67C529B06FF37F5879B53FE57" blockId="33.[151,1437,1039,2037]" lastBlockId="34.[151,1437,151,429]" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
<materialsCitation id="03F73C8B7C529B05FF37F5879AFCF80F" ID-GBIF-Occurrence="1671715810" box="[199,1433,1976,2001]" collectingDate="1981-06-28" collectingDateMax="1981-07-03" collectingDateMin="1981-06-28" collectionCode="PMAE" collectorName="Chi &amp; Hartmans Finca" country="Panama" location="Material" pageId="33" pageNumber="34" specimenCount="1" specimenCount-male="1" stateProvince="PANAMA" typeStatus="holotype">
<emphasis id="81EBEAC47C529B05FF37F5879EDCF80E" bold="true" box="[199,441,1976,2001]" pageId="33" pageNumber="34">
<location id="B640600D7C529B05FF37F5879E4AF80E" LSID="urn:lsid:plazi:treatment:3B3687C07C529B06FF67F1989B7BFE73:B640600D7C529B05FF37F5879E4AF80E" box="[199,303,1976,2001]" country="Panama" name="Material" pageId="33" pageNumber="34" stateProvince="PANAMA">Material</location>
examined.
</emphasis>
<typeStatus id="6C2488747C529B05FE38F5879D56F80E" box="[456,563,1976,2001]" pageId="33" pageNumber="34" type="holotype">Holotype</typeStatus>
,
<specimenCount id="A599FD5F7C529B05FDB6F5869D38F80F" box="[582,605,1977,2000]" pageId="33" pageNumber="34" type="male">♂</specimenCount>
,
<collectingRegion id="715BF8347C529B05FD82F5869D89F80E" box="[626,748,1977,2001]" country="Panama" name="Panama" pageId="33" pageNumber="34">
<collectingCountry id="CB8876467C529B05FD82F5869D89F80E" box="[626,748,1977,2001]" name="Panama" pageId="33" pageNumber="34">PANAMA</collectingCountry>
</collectingRegion>
,
<emphasis id="81EBEAC47C529B05FD0FF5879C16F80E" box="[767,883,1976,2001]" italics="true" pageId="33" pageNumber="34">
<collectorName id="1E6A53007C529B05FD0FF5879C42F80E" box="[767,807,1976,2001]" pageId="33" pageNumber="34">Chi</collectorName>
[riqui]
</emphasis>
:
<collectorName id="1E6A53007C529B05FC7BF5869B2DF80E" box="[907,1096,1976,2001]" pageId="33" pageNumber="34">Hartmans Finca</collectorName>
,
<collectingDate id="D765E9FE7C529B05FBACF5879A40F80F" box="[1116,1317,1976,2000]" pageId="33" pageNumber="34" value="1981-06-28" valueMax="1981-07-03" valueMin="1981-06-28">
<date id="C72110167C529B05FBACF5879A40F80F" box="[1116,1317,1976,2000]" pageId="33" pageNumber="34" value="1981-06-28" valueMax="1981-07-03" valueMin="1981-06-28">28.VI–3.VII.1981</date>
</collectingDate>
(
<collectionCode id="D58EAE137C529B05FACCF5879AF5F80F" box="[1340,1424,1976,2000]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15193" name="Royal Alberta Museum" pageId="33" pageNumber="34">PMAE</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="03F73C8B7C529B06FF67F5E29C8FFF6F" ID-GBIF-Occurrence="1671715819" collectingDate="1999-10-17" collectingDateMax="1999-10-24" collectingDateMin="1999-10-17" collectionCode="MIUP" collectorName="Tr &amp; Malaise, A &amp; Santos" country="Panama" lastPageId="34" lastPageNumber="35" latitude="9.406944" location="Teribe" longLatPrecision="21" longitude="-82.93889" municipality="Wekso" pageId="33" pageNumber="34" specimenCount="7" specimenCount-male="7" stateProvince="Bocas del Toro" typeStatus="paratype">
<typeStatus id="6C2488747C529B05FF67F5E29E6CF82A" box="[151,265,2013,2037]" pageId="33" pageNumber="34" type="paratype">Paratypes</typeStatus>
:
<collectingRegion id="715BF8347C529B05FEE4F5E29EA5F82B" box="[276,448,2012,2037]" country="Panama" name="Bocas del Toro" pageId="33" pageNumber="34">
<emphasis id="81EBEAC47C529B05FEE4F5E29EA5F82B" box="[276,448,2012,2037]" italics="true" pageId="33" pageNumber="34">Bocas del Toro</emphasis>
</collectingRegion>
:
<specimenCount id="A599FD5F7C529B05FE20F5E29E9BF82B" box="[464,510,2013,2036]" pageId="33" pageNumber="34" type="male">7 ♂</specimenCount>
,
<collectingMunicipality id="5344ACAC7C529B05FDFCF5E39D3EF82A" box="[524,603,2012,2037]" pageId="33" pageNumber="34">Wekso</collectingMunicipality>
,
<location id="B640600D7C529B05FD97F5E39DD4F82A" LSID="urn:lsid:plazi:treatment:3B3687C07C529B06FF67F1989B7BFE73:B640600D7C529B05FD97F5E39DD4F82A" box="[615,689,2012,2037]" country="Panama" latitude="9.406944" longLatPrecision="21" longitude="-82.93889" municipality="Wekso" name="Teribe" pageId="33" pageNumber="34" stateProvince="Bocas del Toro">Teribe</location>
, P[arque] I[nternacional] L[a] A[mistad], 50 msnm, [
<geoCoordinate id="D6AB50117C529B05FAE1F5E39AF2F82A" box="[1297,1431,2012,2037]" direction="north" orientation="latitude" pageId="33" pageNumber="34" precision="15" value="9.406944">09º24'25&quot;N</geoCoordinate>
,
<geoCoordinate id="D6AB50117C519B06FF67F2A89E41FF6F" box="[151,292,151,176]" direction="west" orientation="longitude" pageId="34" pageNumber="35" precision="15" value="-82.93889">82º56'20&quot;W</geoCoordinate>
],
<collectingDate id="D765E9FE7C519B06FEC7F2A79E96FF6F" box="[311,499,152,176]" pageId="34" pageNumber="35" value="1999-10-17" valueMax="1999-10-24" valueMin="1999-10-17">
<date id="C72110167C519B06FEC7F2A79E96FF6F" box="[311,499,152,176]" pageId="34" pageNumber="35" value="1999-10-17" valueMax="1999-10-24" valueMin="1999-10-17">17–24 oct. 1999</date>
</collectingDate>
,
<collectorName id="1E6A53007C519B06FDF0F2A79D79FF70" box="[512,540,152,175]" pageId="34" pageNumber="35">Tr</collectorName>
[ampa]
<collectorName id="1E6A53007C519B06FD82F2A89D94FF6F" box="[626,753,151,176]" pageId="34" pageNumber="35">Malaise, A</collectorName>
.
<collectorName id="1E6A53007C519B06FD0FF2A89C35FF6F" box="[767,848,151,176]" pageId="34" pageNumber="35">Santos</collectorName>
[col.] (
<collectionCode id="D58EAE137C519B06FC66F2A79C87FF6F" box="[918,994,152,176]" pageId="34" pageNumber="35">MIUP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="03F73C8B7C519B06FC07F2A89A4FFF6F" ID-GBIF-Occurrence="1671715802" box="[1015,1322,151,176]" collectingDate="1981-06-28" collectingDateMax="1981-07-03" collectingDateMin="1981-06-28" collectionCode="PMAE" collectorName="Chi &amp; Hartmans Finca" country="Panama" location="Material" pageId="34" pageNumber="35" specimenCount="5" specimenCount-male="5" stateProvince="PANAMA" typeStatus="paratype">
<specimenCount id="A599FD5F7C519B06FC07F2A89B79FF6F" box="[1015,1052,151,176]" pageId="34" pageNumber="35" type="male">5♂</specimenCount>
, same data as holotype
</materialsCitation>
;
<materialsCitation id="03F73C8B7C519B06FAC7F2A89C52FF0B" ID-GBIF-Occurrence="1671715841" collectingDate="1981-07-03" collectingDateMax="1981-07-03" collectingDateMin="1981-06-28" collectionCode="PMAE" collectorName="B. Gill" country="Panama" location="Material" pageId="34" pageNumber="35" specimenCount="6" specimenCount-male="6" stateProvince="PANAMA" typeStatus="paratype">
<specimenCount id="A599FD5F7C519B06FAC7F2A89A3FFF6F" box="[1335,1370,151,176]" pageId="34" pageNumber="35" type="male">6♂</specimenCount>
same data as holotype except
<collectingDate id="D765E9FE7C519B06FE5CF2839D4FFF0B" box="[428,554,188,212]" pageId="34" pageNumber="35" value="1981-07-03">
<date id="C72110167C519B06FE5CF2839D4FFF0B" box="[428,554,188,212]" pageId="34" pageNumber="35" value="1981-07-03">3–VII1981</date>
</collectingDate>
,
<collectorName id="1E6A53007C519B06FDC7F2829DE0FF0A" box="[567,645,188,213]" pageId="34" pageNumber="35">B. Gill</collectorName>
[col.] (
<collectionCode id="D58EAE137C519B06FD2BF2839C4AFF0B" box="[731,815,188,212]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15193" name="Royal Alberta Museum" pageId="34" pageNumber="35">PMAE</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="03F73C8B7C519B06FCB4F2839A01FF27" ID-GBIF-Occurrence="1671715824" collectingDate="2000-06-02" collectingDateMax="2000-06-06" collectingDateMin="2000-06-02" collectionCode="CNCI" collectorName="H. &amp; A. Howden" country="Switzerland" latitude="8.666667" location="Hornitos" longLatPrecision="1300" longitude="-82.23333" pageId="34" pageNumber="35" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<specimenCount id="A599FD5F7C519B06FCB4F2839C0DFF0B" box="[836,872,188,212]" pageId="34" pageNumber="35" type="male">1♂</specimenCount>
, same province of
<typeStatus id="6C2488747C519B06FBB5F2839BCFFF0A" box="[1093,1194,188,213]" pageId="34" pageNumber="35" type="holotype">holotype</typeStatus>
, Finca La
<collectingCountry id="CB8876467C519B06FAD5F2839A06FF0A" box="[1317,1379,188,213]" name="Switzerland" pageId="34" pageNumber="35">Suiza</collectingCountry>
near
<location id="B640600D7C519B06FF67F2E09F99FF27" LSID="urn:lsid:plazi:treatment:3B3687C07C529B06FF67F1989B7BFE73:B640600D7C519B06FF67F2E09F99FF27" box="[151,252,223,248]" country="Switzerland" latitude="8.666667" longLatPrecision="1300" longitude="-82.23333" name="Hornitos" pageId="34" pageNumber="35">Hornitos</location>
, [
<geoCoordinate id="D6AB50117C519B06FEE0F2DF9E17FF27" box="[272,370,224,248]" direction="north" orientation="latitude" pageId="34" pageNumber="35" precision="925" value="8.666667">08°40'N</geoCoordinate>
,
<geoCoordinate id="D6AB50117C519B06FE70F2DF9E82FF27" box="[384,487,224,248]" direction="west" orientation="longitude" pageId="34" pageNumber="35" precision="925" value="-82.23333">82°14'W</geoCoordinate>
],
<collectingDate id="D765E9FE7C519B06FE0CF2E09DE8FF27" box="[508,653,223,248]" pageId="34" pageNumber="35" value="2000-06-02" valueMax="2000-06-06" valueMin="2000-06-02">
<date id="C72110167C519B06FE0CF2E09DE8FF27" box="[508,653,223,248]" pageId="34" pageNumber="35" value="2000-06-02" valueMax="2000-06-06" valueMin="2000-06-02">2–6.VI.2000</date>
</collectingDate>
, F[light] I[nterception] T[rap],
<collectorName id="1E6A53007C519B06FC05F2DF9BD8FF27" box="[1013,1213,223,248]" pageId="34" pageNumber="35">H. &amp; A. Howden</collectorName>
[col.] (
<collectionCode id="D58EAE137C519B06FAE3F2DF9A39FF27" box="[1299,1372,224,248]" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="34" pageNumber="35">CNCI</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="03F73C8B7C519B06FA82F2DF9DFAFE9F" ID-GBIF-Occurrence="1671715793" collectionCode="CNCI" collectorName="J. Ashe &amp; R. Brooks" country="Switzerland" latitude="8.823055" location="Boquete" longLatPrecision="21" longitude="-82.42166" pageId="34" pageNumber="35" specimenCount="1" specimenCount-male="1" stateProvince="Culebra" typeStatus="holotype">
<specimenCount id="A599FD5F7C519B06FA82F2DF9AF3FF27" box="[1394,1430,224,248]" pageId="34" pageNumber="35" type="male">1♂</specimenCount>
, same province of
<typeStatus id="6C2488747C519B06FEAEF33B9EA6FEC2" box="[350,451,260,285]" pageId="34" pageNumber="35" type="holotype">holotype</typeStatus>
,
<quantity id="74679B337C519B06FE3FF33B9D79FEC2" box="[463,540,260,285]" metricMagnitude="3" metricUnit="m" metricValue="5.6" pageId="34" pageNumber="35" unit="km" value="5.6">5,6 km</quantity>
N
<location id="B640600D7C519B06FDCFF33A9DFAFEC3" LSID="urn:lsid:plazi:treatment:3B3687C07C529B06FF67F1989B7BFE73:B640600D7C519B06FDCFF33A9DFAFEC3" box="[575,671,261,284]" country="Switzerland" latitude="8.823055" longLatPrecision="21" longitude="-82.42166" name="Boquete" pageId="34" pageNumber="35" stateProvince="Culebra">Boquete</location>
, La
<collectingRegion id="715BF8347C519B06FD3EF33B9C42FEC2" box="[718,807,260,285]" country="Puerto Rico" name="Culebra" pageId="34" pageNumber="35">Culebra</collectingRegion>
trail, [
<geoCoordinate id="D6AB50117C519B06FC80F33B9C9CFEC3" box="[880,1017,260,284]" direction="north" orientation="latitude" pageId="34" pageNumber="35" precision="15" value="8.823055">08°49'23&quot;N</geoCoordinate>
,
<geoCoordinate id="D6AB50117C519B06FBF4F33B9BF6FEC2" box="[1028,1171,260,285]" direction="west" orientation="longitude" pageId="34" pageNumber="35" precision="15" value="-82.42166">82°25'18&quot;W</geoCoordinate>
], F[light] I[nterception] T[rap],
<collectorName id="1E6A53007C519B06FF02F3179E2EFE9F" box="[242,331,295,320]" pageId="34" pageNumber="35">J. Ashe</collectorName>
&amp;
<collectorName id="1E6A53007C519B06FE8AF3179E94FE9F" box="[378,497,295,320]" pageId="34" pageNumber="35">R. Brooks</collectorName>
[col.] (
<collectionCode id="D58EAE137C519B06FDBEF3179DF2FE9F" box="[590,663,296,320]" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="34" pageNumber="35">CNCI</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="03F73C8B7C519B06FD40F3179B93FEBB" ID-GBIF-Occurrence="1671715858" collectingDate="2000-06-06" collectingDateMax="2000-06-25" collectingDateMin="2000-06-06" collectionCode="CNCI" collectorName="H. &amp; A. Howden" country="Switzerland" latitude="8.666667" location="Hornitos" longLatPrecision="1300" longitude="-82.23333" pageId="34" pageNumber="35" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
<specimenCount id="A599FD5F7C519B06FD40F3179DB3FE9F" box="[688,726,296,320]" pageId="34" pageNumber="35" type="male">2♂</specimenCount>
, same province of
<typeStatus id="6C2488747C519B06FC30F3189B40FE9F" box="[960,1061,295,320]" pageId="34" pageNumber="35" type="holotype">holotype</typeStatus>
, Finca La
<collectingCountry id="CB8876467C519B06FB5BF3189B8CFE9F" box="[1195,1257,295,320]" name="Switzerland" pageId="34" pageNumber="35">Suiza</collectingCountry>
near
<location id="B640600D7C519B06FAC3F3189AFDFE9F" LSID="urn:lsid:plazi:treatment:3B3687C07C529B06FF67F1989B7BFE73:B640600D7C519B06FAC3F3189AFDFE9F" box="[1331,1432,295,320]" country="Switzerland" latitude="8.666667" longLatPrecision="1300" longitude="-82.23333" name="Hornitos" pageId="34" pageNumber="35">Hornitos</location>
, [
<geoCoordinate id="D6AB50117C519B06FF6EF3739F9AFEBB" box="[158,255,332,356]" direction="north" orientation="latitude" pageId="34" pageNumber="35" precision="925" value="8.666667">08°40'N</geoCoordinate>
,
<geoCoordinate id="D6AB50117C519B06FEFCF3739E16FEBB" box="[268,371,332,356]" direction="west" orientation="longitude" pageId="34" pageNumber="35" precision="925" value="-82.23333">82°14'W</geoCoordinate>
],
<collectingDate id="D765E9FE7C519B06FE78F3739D40FEBA" box="[392,549,332,357]" pageId="34" pageNumber="35" value="2000-06-25" valueMax="2000-06-06" valueMin="2000-06-25">
<date id="C72110167C519B06FE78F3739D40FEBA" box="[392,549,332,357]" pageId="34" pageNumber="35" value="2000-06-25" valueMax="2000-06-06" valueMin="2000-06-25">25–6.VI.2000</date>
</collectingDate>
, F[light] I[nterception] T[rap],
<collectorName id="1E6A53007C519B06FC7BF3729B35FEBA" box="[907,1104,332,357]" pageId="34" pageNumber="35">H. &amp; A. Howden</collectorName>
[col.] (
<collectionCode id="D58EAE137C519B06FB55F3739B8BFEBB" box="[1189,1262,332,356]" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="34" pageNumber="35">CNCI</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="03F73C8B7C519B06FAF3F3729B57FE57" ID-GBIF-Occurrence="1671715807" collectingDate="1982-06-30" collectingDateMax="1982-07-14" collectingDateMin="1982-06-30" collectionCode="PMAE" collectorName="B. Gill" country="Switzerland" latitude="7.5166664" location="Ojo de Agua" longLatPrecision="1303" longitude="-80.416664" pageId="34" pageNumber="35" specimenCount="2" specimenCount-male="2" stateProvince="Los Santos" typeStatus="holotype">
<emphasis id="81EBEAC47C519B06FAF3F3729AF1FEBB" box="[1283,1428,333,356]" italics="true" pageId="34" pageNumber="35">
[
<collectingRegion id="715BF8347C519B06FAFDF3729AECFEBB" box="[1293,1417,333,356]" country="Panama" name="Los Santos" pageId="34" pageNumber="35">Los Santos</collectingRegion>
]
</emphasis>
:
<specimenCount id="A599FD5F7C519B06FF67F34F9FD9FE57" box="[151,188,368,392]" pageId="34" pageNumber="35" type="male">2♂</specimenCount>
,
<location id="B640600D7C519B06FF38F3509E3CFE57" LSID="urn:lsid:plazi:treatment:3B3687C07C529B06FF67F1989B7BFE73:B640600D7C519B06FF38F3509E3CFE57" box="[200,345,367,392]" country="Switzerland" latitude="7.5166664" longLatPrecision="1303" longitude="-80.416664" name="Ojo de Agua" pageId="34" pageNumber="35" stateProvince="Los Santos">Ojo de Agua</location>
, [
<geoCoordinate id="D6AB50117C519B06FE9CF34F9EA8FE57" box="[364,461,368,392]" direction="north" orientation="latitude" pageId="34" pageNumber="35" precision="925" value="7.5166664">07°31'N</geoCoordinate>
,
<geoCoordinate id="D6AB50117C519B06FE29F3509D25FE57" box="[473,576,367,392]" direction="west" orientation="longitude" pageId="34" pageNumber="35" precision="925" value="-80.416664">80°25'W</geoCoordinate>
],
<collectingDate id="D765E9FE7C519B06FDA3F34F9C48FE57" box="[595,813,368,392]" pageId="34" pageNumber="35" value="1982-06-30" valueMax="1982-07-14" valueMin="1982-06-30">
<date id="C72110167C519B06FDA3F34F9C48FE57" box="[595,813,368,392]" pageId="34" pageNumber="35" value="1982-06-30" valueMax="1982-07-14" valueMin="1982-06-30">30.VI–14.VII.1982</date>
</collectingDate>
,
<collectorName id="1E6A53007C519B06FCC8F34F9CE0FE57" box="[824,901,367,392]" pageId="34" pageNumber="35">B. Gill</collectorName>
[col.] (
<collectionCode id="D58EAE137C519B06FC26F34F9B4FFE57" box="[982,1066,368,392]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15193" name="Royal Alberta Museum" pageId="34" pageNumber="35">PMAE</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C519B06FF37F3AB9B7BFE73" box="[199,1054,404,429]" pageId="34" pageNumber="35" type="distribution">
<paragraph id="B32036D67C519B06FF37F3AB9B7BFE73" blockId="34.[151,1437,151,429]" box="[199,1054,404,429]" pageId="34" pageNumber="35">
<emphasis id="81EBEAC47C519B06FF37F3AB9E3CFE72" bold="true" box="[199,345,404,429]" pageId="34" pageNumber="35">Distribution</emphasis>
(
<figureCitation id="2BA42A537C519B06FE98F3AB9EAEFE73" box="[360,459,404,429]" captionStart="FIGURES 244 – 249" captionStartId="68.[151,264,1212,1235]" captionTargetBox="[156,1430,201,1176]" captionTargetId="figure@68.[148,1433,193,1184]" captionTargetPageId="68" captionText="FIGURES 244 – 249. Known distribution of Dissomphalus in Panama. 244. D. rufipalpis, D. teraus sp. nov. and D. vallensis. 245. D. sartus sp. nov. and D. trisus sp. nov.. 246. D. teobus sp. nov. and D. urazius sp. nov.. 247. D. terbius sp. nov. and D. unitus. 248. D. ubracus sp. nov.. 249. D. rettenmeyeri." httpUri="https://zenodo.org/record/1010062/files/figure.png" pageId="34" pageNumber="35">Fig. 244</figureCitation>
).
<collectingRegion id="715BF8347C519B06FE2EF3AA9D52FE73" box="[478,567,405,428]" country="Panama" name="Panama" pageId="34" pageNumber="35">
<collectingCountry id="CB8876467C519B06FE2EF3AA9D52FE73" box="[478,567,405,428]" name="Panama" pageId="34" pageNumber="35">Panama</collectingCountry>
</collectingRegion>
(
<collectingRegion id="715BF8347C519B06FDB6F3AB9D96FE73" box="[582,755,404,429]" country="Panama" name="Bocas del Toro" pageId="34" pageNumber="35">Bocas del Toro</collectingRegion>
,
<collectingRegion id="715BF8347C519B06FD0EF3AB9C3EFE72" box="[766,859,404,429]" country="Panama" name="Chiriqui" pageId="34" pageNumber="35">Chiriqui</collectingRegion>
and
<collectingRegion id="715BF8347C519B06FC62F3AA9B77FE73" box="[914,1042,404,428]" country="Panama" name="Los Santos" pageId="34" pageNumber="35">Los Santos</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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