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<document id="C8BF0CE86AB5F9AC9109DEB1C06B65E3" ID-CLB-Dataset="31852" ID-DOI="10.11646/zootaxa.4335.1.1" ID-GBIF-Dataset="f2ad3cb1-0bad-4be1-9e36-2170a6a08aaf" ID-ISSN="1175-5326" ID-Zenodo-Dep="1010014" ID-ZooBank="3429592D-23A8-4EBF-8412-BD57B3BC4314" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1507877826645" checkinUser="plazi" docAuthor="Azevedo, Celso O." docDate="2017" docId="3B3687C07C429B17FF67F7489B5FFE2A" docLanguage="en" docName="zootaxa.4335.1.1.pdf" docOrigin="Zootaxa 4335 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Dissomphalus sartus Azevedo 2017, sp. nov." docType="treatment" docVersion="15" lastPageNumber="52" masterDocId="C70FFFB87C739B24FFF0F23F9F65FFDF" masterDocTitle="Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species" masterLastPageNumber="72" masterPageNumber="1" pageNumber="50" updateTime="1698455438235" updateUser="plazi">
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<mods:title id="B0141ADEC5C58F9C61928899A22F7518">Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species</mods:title>
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<mods:namePart id="FAD6B2A3D6446C11812D2D5C4F01B13A">Azevedo, Celso O.</mods:namePart>
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<mods:date id="DA3C56405BA8D960358E4E6E08F940E7">2017</mods:date>
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<mods:number id="33013A5A899AFE186209B1C552693B4F">2017-10-13</mods:number>
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<mods:identifier id="E4CAAA9012FD9E4906BACA7C4388340D" type="DOI">10.11646/zootaxa.4335.1.1</mods:identifier>
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<treatment id="3B3687C07C429B17FF67F7489B5FFE2A" ID-DOI="http://doi.org/10.5281/zenodo.5020472" ID-GBIF-Taxon="135435261" ID-Zenodo-Dep="5020472" LSID="urn:lsid:plazi:treatment:3B3687C07C429B17FF67F7489B5FFE2A" httpUri="http://treatment.plazi.org/id/3B3687C07C429B17FF67F7489B5FFE2A" lastPageId="51" lastPageNumber="52" pageId="49" pageNumber="50">
<subSubSection id="FB85655D7C429B15FF67F7489EA4FA6C" pageId="49" pageNumber="50" type="nomenclature">
<paragraph id="B32036D67C429B15FF67F7489DBDFA4D" blockId="49.[151,728,1399,1459]" box="[151,728,1399,1426]" pageId="49" pageNumber="50">
<heading id="E86881BA7C429B15FF67F7489DBDFA4D" bold="true" box="[151,728,1399,1426]" fontSize="11" level="1" pageId="49" pageNumber="50" reason="1">
<emphasis id="81EBEAC47C429B15FF67F7489DBDFA4D" bold="true" box="[151,728,1399,1426]" pageId="49" pageNumber="50">
<taxonomicName id="749F4D557C429B15FF67F7489D08FA4D" authority="Brito &amp; Azevedo" authorityName="Azevedo" authorityYear="2017" box="[151,621,1399,1426]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="sartus" status="sp. nov.">
<emphasis id="81EBEAC47C429B15FF67F7489EF7FA4E" bold="true" box="[151,402,1399,1425]" italics="true" pageId="49" pageNumber="50">Dissomphalus sartus</emphasis>
Brito &amp; Azevedo
</taxonomicName>
,
<taxonomicNameLabel id="9AD857BF7C429B15FD8AF7479DBDFA4D" box="[634,728,1400,1426]" pageId="49" pageNumber="50" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="B32036D67C429B15FF67F7A59EA4FA6C" blockId="49.[151,728,1399,1459]" box="[151,449,1434,1459]" pageId="49" pageNumber="50">
(
<figureCitation id="2BA42A537C429B15FF6FF7A59F90FA6C" box="[159,245,1434,1459]" captionStart="FIGURES 13 – 24" captionStartId="11.[151,264,1615,1638]" captionTargetBox="[151,1436,306,1594]" captionTargetId="figure@11.[151,1436,303,1594]" captionTargetPageId="11" captionText="FIGURES 13 – 24. Dissomphalus spp., ♂, head, dorsal view. 13. D. racarus sp. nov. 14. D. teraus sp. nov. 15. D. urazius sp. nov. 16. D. afestus sp. nov. 17. D. arbeius sp. nov. 18. D. raziogus sp. nov. 19. D. areius sp. nov. 20. D. aretisus sp. nov. 21. D. citus sp. nov. 22. D. sartus sp. nov. 23. D. teobus sp. nov. 24. D. terbius sp. nov.. Scale bars = 20 µm, except figs 14, 20 = 10 µm and fig. 15 = 50 µm." httpUri="https://zenodo.org/record/1010018/files/figure.png" pageId="49" pageNumber="50">Figs 22</figureCitation>
,
<figureCitation id="2BA42A537C429B15FEF1F7A59E7AFA6C" box="[257,287,1434,1459]" captionStart="FIGURES 45 – 56" captionStartId="14.[151,264,1544,1567]" captionTargetBox="[151,1436,193,1524]" captionTargetId="figure@14.[151,1436,193,1524]" captionTargetPageId="14" captionText="FIGURES 45 – 56. Dissomphalus spp., ♂, metasomal tergal process, dorsal view. 45. D. teraus sp. nov. 46. D. urazius sp. nov. 47. D. afestus sp. nov. 48. D. arbeius sp. nov. 49. D. raziogus sp. nov. 50. D. areius sp. nov. 51. D. aretisus sp. nov. 52. D. citus sp. nov. 53. D. sartus sp. nov. 54. D. teobus sp. nov. 55. D. terbius sp. nov. 56. D. abarus sp. nov.. Scale bars = 20 µm except, figs 45 and 46 = 50 µm." httpUri="https://zenodo.org/record/1010024/files/figure.png" pageId="49" pageNumber="50">53</figureCitation>
,
<figureCitation id="2BA42A537C429B15FEDBF7A49E2FFA6C" box="[299,330,1435,1459]" captionStart="FIGURES 76 – 87" captionStartId="17.[151,264,1595,1618]" captionTargetBox="[159,1404,219,1523]" captionTargetId="figure@17.[147,1432,209,1558]" captionTargetPageId="17" captionText="FIGURES 76 – 87. Dissomphalus spp., ♂, hypopygium, internal view. 76. D. teraus sp. nov. 77. D. urazius sp. nov. 78. D. afestus sp. nov. 79. D. arbeius sp. nov. 80. D. raziogus sp. nov. 81. D. areius sp. nov. 82. D. aretisus sp. nov. 83. D. citus sp. nov. 84. D. sartus sp. nov. 85. D. teobus sp. nov. 86. D. terbius sp. nov. 87. D. abarus sp. nov.. Scale bars = 125 µm." httpUri="https://zenodo.org/record/1010030/files/figure.png" pageId="49" pageNumber="50">84</figureCitation>
,
<figureCitation id="2BA42A537C429B15FEA5F7A59EDDFA6C" box="[341,440,1434,1459]" captionStart="FIGURES 184 – 197" captionStartId="50.[151,264,1612,1635]" captionTargetBox="[159,1430,206,1544]" captionTargetId="figure@50.[151,1436,193,1575]" captionTargetPageId="50" captionText="FIGURES 184 – 197. Dissomphalus spp., ♂, genitalia, except otherwise indicated. 184 – 185. D. sartus sp. nov. 184. Dorsal view. 185. Ventral view. 186 – 187. D. teobus sp. nov. 186. Dorsal view. 187. Ventral view. 188 – 189. D. terbius sp. nov. 188. Dorsal view. 189. Ventral view. 190 – 194. D. unitus 190. Dorsal view. 191. Ventral view. 192. Outer lobe of aedeagal dorsal body, dorsal view. 193. Inner lobe of aedeagal dorsal body, dorsal view. 194. Apical lobe of aedeagal dorsal body, lateral view. 195 – 196. D. dilatatus 195. Dorsal view. 196. Ventral view. 197. D. vallensis. Dorsal view. Scale bars = 250 µm" httpUri="https://zenodo.org/record/1010048/files/figure.png" pageId="49" pageNumber="50">184–185</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C429B15FF67F7DD9E54F811" pageId="49" pageNumber="50" type="description">
<paragraph id="B32036D67C429B15FF67F7DD9C1EF858" blockId="49.[151,1437,1506,1999]" pageId="49" pageNumber="50">
<emphasis id="81EBEAC47C429B15FF67F7DD9E4CFA24" bold="true" box="[151,297,1506,1531]" pageId="49" pageNumber="50">Description.</emphasis>
Male. Body length
<quantity id="74679B337C429B15FDE6F7DD9DE4FA25" box="[534,641,1506,1531]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" pageId="49" pageNumber="50" unit="mm" value="1.85">1.85 mm</quantity>
. Color: head and mesosoma dark castaneous; metasoma castaneous. Head (
<figureCitation id="2BA42A537C429B15FF16F4399E5BF9C0" box="[230,318,1542,1567]" captionStart="FIGURES 13 – 24" captionStartId="11.[151,264,1615,1638]" captionTargetBox="[151,1436,306,1594]" captionTargetId="figure@11.[151,1436,303,1594]" captionTargetPageId="11" captionText="FIGURES 13 – 24. Dissomphalus spp., ♂, head, dorsal view. 13. D. racarus sp. nov. 14. D. teraus sp. nov. 15. D. urazius sp. nov. 16. D. afestus sp. nov. 17. D. arbeius sp. nov. 18. D. raziogus sp. nov. 19. D. areius sp. nov. 20. D. aretisus sp. nov. 21. D. citus sp. nov. 22. D. sartus sp. nov. 23. D. teobus sp. nov. 24. D. terbius sp. nov.. Scale bars = 20 µm, except figs 14, 20 = 10 µm and fig. 15 = 50 µm." httpUri="https://zenodo.org/record/1010018/files/figure.png" pageId="49" pageNumber="50">Fig. 22</figureCitation>
). Mandible with two distal teeth. Median clypeal lobe subtrapezoidal, with one angulate tooth; median clypeal carina low in profile, incomplete apically, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest straight. Pronotal disc coriaceous; anterior margin ecarinate. Metasoma (
<figureCitation id="2BA42A537C429B15FACCF4719AEAF9B8" box="[1340,1423,1614,1639]" captionStart="FIGURES 45 – 56" captionStartId="14.[151,264,1544,1567]" captionTargetBox="[151,1436,193,1524]" captionTargetId="figure@14.[151,1436,193,1524]" captionTargetPageId="14" captionText="FIGURES 45 – 56. Dissomphalus spp., ♂, metasomal tergal process, dorsal view. 45. D. teraus sp. nov. 46. D. urazius sp. nov. 47. D. afestus sp. nov. 48. D. arbeius sp. nov. 49. D. raziogus sp. nov. 50. D. areius sp. nov. 51. D. aretisus sp. nov. 52. D. citus sp. nov. 53. D. sartus sp. nov. 54. D. teobus sp. nov. 55. D. terbius sp. nov. 56. D. abarus sp. nov.. Scale bars = 20 µm except, figs 45 and 46 = 50 µm." httpUri="https://zenodo.org/record/1010024/files/figure.png" pageId="49" pageNumber="50">Fig. 53</figureCitation>
). Tergal process with lateral, shallow, subcircular and small depression, with small, high and dorsad tubercle placed on center of depression, pit small, with short and dense tuft of setae. Posterior hypopygeal margin straight or nearly so (
<figureCitation id="2BA42A537C429B15FF4EF4859E77F90C" box="[190,274,1722,1747]" captionStart="FIGURES 76 – 87" captionStartId="17.[151,264,1595,1618]" captionTargetBox="[159,1404,219,1523]" captionTargetId="figure@17.[147,1432,209,1558]" captionTargetPageId="17" captionText="FIGURES 76 – 87. Dissomphalus spp., ♂, hypopygium, internal view. 76. D. teraus sp. nov. 77. D. urazius sp. nov. 78. D. afestus sp. nov. 79. D. arbeius sp. nov. 80. D. raziogus sp. nov. 81. D. areius sp. nov. 82. D. aretisus sp. nov. 83. D. citus sp. nov. 84. D. sartus sp. nov. 85. D. teobus sp. nov. 86. D. terbius sp. nov. 87. D. abarus sp. nov.. Scale bars = 125 µm." httpUri="https://zenodo.org/record/1010030/files/figure.png" pageId="49" pageNumber="50">Fig. 84</figureCitation>
). Genitalia (
<figureCitation id="2BA42A537C429B15FE6AF4859D5CF90C" box="[410,569,1722,1747]" captionStart="FIGURES 184 – 197" captionStartId="50.[151,264,1612,1635]" captionTargetBox="[159,1430,206,1544]" captionTargetId="figure@50.[151,1436,193,1575]" captionTargetPageId="50" captionText="FIGURES 184 – 197. Dissomphalus spp., ♂, genitalia, except otherwise indicated. 184 – 185. D. sartus sp. nov. 184. Dorsal view. 185. Ventral view. 186 – 187. D. teobus sp. nov. 186. Dorsal view. 187. Ventral view. 188 – 189. D. terbius sp. nov. 188. Dorsal view. 189. Ventral view. 190 – 194. D. unitus 190. Dorsal view. 191. Ventral view. 192. Outer lobe of aedeagal dorsal body, dorsal view. 193. Inner lobe of aedeagal dorsal body, dorsal view. 194. Apical lobe of aedeagal dorsal body, lateral view. 195 – 196. D. dilatatus 195. Dorsal view. 196. Ventral view. 197. D. vallensis. Dorsal view. Scale bars = 250 µm" httpUri="https://zenodo.org/record/1010048/files/figure.png" pageId="49" pageNumber="50">Figs 184–185</figureCitation>
): paramere with dorsal margin wide basally; apical margin truncate. Aedeagal ventral ramus as long as dorsal body, wide, progressively narrowing apicad; cross section laminar; surface subhorizontal; inner margin straight; outer margin slightly sinuous; apex long, simple, slightly curved inward; additional inner ramus absent; basal stub short and very narrow. Aedeagal dorsal body with pair of apical lobe; outer lobe short, horizontal and very narrow, with apical margin abruptly acute and ventrad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring.
</paragraph>
<paragraph id="B32036D67C429B15FF37F5AD9E54F811" blockId="49.[151,1437,1506,1999]" pageId="49" pageNumber="50">
<emphasis id="81EBEAC47C429B15FF37F5AD9E22F874" bold="true" box="[199,327,1938,1963]" pageId="49" pageNumber="50">Variations.</emphasis>
Depression of tergal process very shallow, tubercle low; apical lobe of aedeagal dorsal body longer and narrower.
</paragraph>
</subSubSection>
<caption id="E7E0665E7C419B16FF67F4739A61F902" httpUri="https://zenodo.org/record/1010048/files/figure.png" pageId="50" pageNumber="51" targetBox="[159,1430,206,1544]" targetPageId="50">
<paragraph id="B32036D67C419B16FF67F4739A61F902" blockId="50.[151,1436,1612,1757]" pageId="50" pageNumber="51">
<emphasis id="81EBEAC47C419B16FF67F4739E15F9BE" bold="true" box="[151,368,1612,1635]" pageId="50" pageNumber="51">FIGURES 184–197.</emphasis>
<taxonomicName id="749F4D557C419B16FE88F4739D6DF9BD" authorityName="Ashmead" authorityYear="1893" box="[376,520,1612,1634]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="81EBEAC47C419B16FE88F4739D6DF9BD" box="[376,520,1612,1634]" italics="true" pageId="50" pageNumber="51">Dissomphalus</emphasis>
</taxonomicName>
spp., ♂, genitalia, except otherwise indicated. 184–185.
<taxonomicName id="749F4D557C419B16FBADF4729BD8F9BD" authority="Azevedo, 2017" authorityName="Azevedo" authorityYear="2017" box="[1117,1213,1613,1634]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="sartus" status="sp. nov.">
<emphasis id="81EBEAC47C419B16FBADF4729BD8F9BD" box="[1117,1213,1613,1634]" italics="true" pageId="50" pageNumber="51">D. sartus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9AD857BF7C419B16FB36F4729A7CF9BD" box="[1222,1305,1613,1634]" pageId="50" pageNumber="51" rank="species">
<emphasis id="81EBEAC47C419B16FB36F4729A7CF9BD" bold="true" box="[1222,1305,1613,1634]" pageId="50" pageNumber="51">sp. nov.</emphasis>
</taxonomicNameLabel>
184. Dorsal view. 185. Ventral view.186–187.
<taxonomicName id="749F4D557C419B16FE07F4539D3FF95E" authority="Azevedo, 2017" authorityName="Azevedo" authorityYear="2017" box="[503,602,1643,1665]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="teobus" status="sp. nov.">
<emphasis id="81EBEAC47C419B16FE07F4539D3FF95E" box="[503,602,1643,1665]" italics="true" pageId="50" pageNumber="51">D. teobus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9AD857BF7C419B16FD92F4549DD1F95F" box="[610,692,1643,1664]" pageId="50" pageNumber="51" rank="species">
<emphasis id="81EBEAC47C419B16FD92F4549DD1F95F" bold="true" box="[610,692,1643,1664]" pageId="50" pageNumber="51">sp. nov.</emphasis>
</taxonomicNameLabel>
186. Dorsal view. 187. Ventral view. 188–189.
<taxonomicName id="749F4D557C419B16FB55F4539A69F95E" authority="Azevedo, 2017" authorityName="Azevedo" authorityYear="2017" box="[1189,1292,1643,1665]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="terbius" status="sp. nov.">
<emphasis id="81EBEAC47C419B16FB55F4539A69F95E" box="[1189,1292,1643,1665]" italics="true" pageId="50" pageNumber="51">D. terbius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9AD857BF7C419B16FAE5F4549A02F95F" box="[1301,1383,1643,1664]" pageId="50" pageNumber="51" rank="species">
<emphasis id="81EBEAC47C419B16FAE5F4549A02F95F" bold="true" box="[1301,1383,1643,1664]" pageId="50" pageNumber="51">sp. nov.</emphasis>
</taxonomicNameLabel>
188. Dorsal view. 189. Ventral view. 190–194.
<taxonomicName id="749F4D557C419B16FDBCF4B59DCEF940" authorityName="Azevedo. Seven" authorityYear="1999" box="[588,683,1674,1695]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="unitus">
<emphasis id="81EBEAC47C419B16FDBCF4B59DCEF940" box="[588,683,1674,1695]" italics="true" pageId="50" pageNumber="51">D. unitus</emphasis>
</taxonomicName>
190. Dorsal view. 191. Ventral view. 192. Outer lobe of aedeagal dorsal body, dorsal view. 193. Inner lobe of aedeagal dorsal body, dorsal view. 194. Apical lobe of aedeagal dorsal body, lateral view. 195– 196.
<taxonomicName id="749F4D557C419B16FEF2F4F79E1FF902" authorityName="Azevedo" authorityYear="1999" box="[258,378,1735,1757]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="dilatatus">
<emphasis id="81EBEAC47C419B16FEF2F4F79E1FF902" box="[258,378,1735,1757]" italics="true" pageId="50" pageNumber="51">D. dilatatus</emphasis>
</taxonomicName>
195. Dorsal view. 196. Ventral view. 197.
<taxonomicName id="749F4D557C419B16FCDAF4F79CC7F902" authorityName="Evans. Eighteen" authorityYear="1979" box="[810,930,1735,1757]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="vallensis">
<emphasis id="81EBEAC47C419B16FCDAF4F79CC7F902" box="[810,930,1735,1757]" italics="true" pageId="50" pageNumber="51">D. vallensis</emphasis>
</taxonomicName>
. Dorsal view. Scale bars = 250 µm
</paragraph>
</caption>
<subSubSection id="FB85655D7C419B16FF37F5369B7EF809" pageId="50" pageNumber="51" type="discussion">
<paragraph id="B32036D67C419B16FF37F5369B7EF809" blockId="50.[151,1436,1801,2006]" pageId="50" pageNumber="51">
<emphasis id="81EBEAC47C419B16FF37F5369E5EF8FD" bold="true" box="[199,315,1801,1826]" pageId="50" pageNumber="51">Remarks.</emphasis>
This species is similar to
<taxonomicName id="749F4D557C419B16FD72F5349D95F8FD" authority="Azevedo, 2017" authorityName="Azevedo" authorityYear="2017" box="[642,752,1803,1826]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="areius" status="sp. nov.">
<emphasis id="81EBEAC47C419B16FD72F5349D95F8FD" box="[642,752,1803,1826]" italics="true" pageId="50" pageNumber="51">D. areius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9AD857BF7C419B16FD0DF5369C38F8FD" box="[765,861,1801,1826]" pageId="50" pageNumber="51" rank="species">
<emphasis id="81EBEAC47C419B16FD0DF5369C38F8FD" bold="true" box="[765,861,1801,1826]" pageId="50" pageNumber="51">sp. nov.</emphasis>
</taxonomicNameLabel>
by having the paramere excavated apically and expanded basally and the aedeagal dorsal body with three narrow and long filaments which connect the apical lobes of the aedeagal dorsal body to the basal portion of the aedeagal dorsal body. However, this species has the apex of apical lobes of the aedeagal dorsal body abruptly acute and the aedeagal ventral ramus with the apex toothed, whereas
<taxonomicName id="749F4D557C419B16FEABF5A49EA4F86D" authority="Azevedo, 2017" authorityName="Azevedo" authorityYear="2017" box="[347,449,1947,1970]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="areius" status="sp. nov.">
<emphasis id="81EBEAC47C419B16FEABF5A49EA4F86D" box="[347,449,1947,1970]" italics="true" pageId="50" pageNumber="51">D. areius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9AD857BF7C419B16FE38F5A69D44F86D" box="[456,545,1945,1970]" pageId="50" pageNumber="51" rank="species">
<emphasis id="81EBEAC47C419B16FE38F5A69D44F86D" bold="true" box="[456,545,1945,1970]" pageId="50" pageNumber="51">sp. nov.</emphasis>
</taxonomicNameLabel>
has the apex of the apical lobes of the aedeagal dorsal body with a small laterad projection and the aedeagal ventral ramus with the apex translucent and smooth.
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C409B17FF37F2A89F92FE0F" pageId="51" pageNumber="52" type="materials_examined">
<paragraph id="B32036D67C409B17FF37F2A89F92FE0F" blockId="51.[151,1436,151,501]" pageId="51" pageNumber="52">
<materialsCitation id="03F73C8B7C409B17FF37F2A89C30FF0B" ID-GBIF-Occurrence="1671715716" collectingDate="2000-11-16" country="Panama" county="Material" elevation="80" latitude="8.0" location="Pirre" longLatPrecision="1302" longitude="-77.75" municipality="Darien Pr" pageId="51" pageNumber="52" specimenCount="1" specimenCount-male="1" stateProvince="PANAMA" typeStatus="holotype">
<emphasis id="81EBEAC47C409B17FF37F2A89EDCFF6F" bold="true" box="[199,441,151,176]" pageId="51" pageNumber="52">
<collectingCounty id="5A414E5A7C409B17FF37F2A89E4AFF6F" box="[199,303,151,176]" pageId="51" pageNumber="52">Material</collectingCounty>
examined.
</emphasis>
<typeStatus id="6C2488747C409B17FE38F2A89D56FF6F" box="[456,563,151,176]" pageId="51" pageNumber="52" type="holotype">Holotype</typeStatus>
,
<specimenCount id="A599FD5F7C409B17FDB7F2A79D3BFF70" box="[583,606,152,175]" pageId="51" pageNumber="52" type="male">♂</specimenCount>
,
<collectingRegion id="715BF8347C409B17FD83F2A79D88FF6F" box="[627,749,152,176]" country="Panama" name="Panama" pageId="51" pageNumber="52">
<collectingCountry id="CB8876467C409B17FD83F2A79D88FF6F" box="[627,749,152,176]" name="Panama" pageId="51" pageNumber="52">PANAMA</collectingCountry>
</collectingRegion>
,
<emphasis id="81EBEAC47C409B17FCF0F2A79C88FF70" box="[768,1005,152,175]" italics="true" pageId="51" pageNumber="52">
<collectingMunicipality id="5344ACAC7C409B17FCF0F2A79C1EFF70" box="[768,891,152,175]" pageId="51" pageNumber="52">
<collectingRegion id="715BF8347C409B17FCF0F2A79C2AFF70" box="[768,847,152,175]" country="Panama" name="Darien" pageId="51" pageNumber="52">Darién</collectingRegion>
<collectingRegion id="715BF8347C409B17FCAFF2A79C1EFF70" box="[863,891,152,175]" country="Brazil" name="Para" pageId="51" pageNumber="52">Pr</collectingRegion>
</collectingMunicipality>
[rovincia]
</emphasis>
: P[arque] N[acional]
<collectingRegion id="715BF8347C409B17FB0BF2A89A2FFF6F" box="[1275,1354,151,176]" country="Panama" name="Darien" pageId="51" pageNumber="52">Darién</collectingRegion>
,
<location id="B640600D7C409B17FAAEF2A89AFDFF6F" LSID="urn:lsid:plazi:treatment:3B3687C07C429B17FF67F7489B5FFE2A:B640600D7C409B17FAAEF2A89AFDFF6F" box="[1374,1432,151,176]" country="Panama" county="Material" latitude="8.0" longLatPrecision="1302" longitude="-77.75" municipality="Darien Pr" name="Pirre" pageId="51" pageNumber="52" stateProvince="PANAMA">Pirre</location>
,
<location id="B640600D7C409B17FF67F2839FD8FF0A" LSID="urn:lsid:plazi:treatment:3B3687C07C429B17FF67F7489B5FFE2A:B640600D7C409B17FF67F2839FD8FF0A" box="[151,189,188,213]" country="Panama" county="Material" latitude="8.0" longLatPrecision="1302" longitude="-77.75" municipality="Darien Pr" name="Est" pageId="51" pageNumber="52" stateProvince="PANAMA">Est</location>
[ación]
<location id="B640600D7C409B17FEE2F2839EC5FF0A" LSID="urn:lsid:plazi:treatment:3B3687C07C429B17FF67F7489B5FFE2A:B640600D7C409B17FEE2F2839EC5FF0A" box="[274,416,188,213]" country="Panama" county="Material" latitude="8.0" longLatPrecision="1302" longitude="-77.75" municipality="Darien Pr" name="Rancho Frio" pageId="51" pageNumber="52" stateProvince="PANAMA">Rancho Frio</location>
,
<elevation id="38B2D1E57C409B17FE5CF2839E8DFF0B" box="[428,488,188,212]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="51" pageNumber="52" unit="m" value="80.0">
<quantity id="74679B337C409B17FE5CF2839E8DFF0B" box="[428,488,188,212]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="51" pageNumber="52" unit="m" value="80.0">80 m</quantity>
</elevation>
, [
<geoCoordinate id="D6AB50117C409B17FE0BF2839D39FF0B" box="[507,604,188,212]" direction="north" orientation="latitude" pageId="51" pageNumber="52" precision="925" value="8.0">08°00'N</geoCoordinate>
,
<geoCoordinate id="D6AB50117C409B17FD98F2839DB4FF0A" box="[616,721,188,213]" direction="west" orientation="longitude" pageId="51" pageNumber="52" precision="925" value="-77.75">77°45'W</geoCoordinate>
],
<collectingDate id="D765E9FE7C409B17FD13F2839C30FF0B" box="[739,853,188,213]" pageId="51" pageNumber="52" value="2000-11-16">
<date id="C72110167C409B17FD13F2839C30FF0B" box="[739,853,188,213]" pageId="51" pageNumber="52" value="2000-11-16">16 nov 00</date>
</collectingDate>
</materialsCitation>
–
<date id="C72110167C409B17FC95F2839C95FF0B" box="[869,1008,188,212]" pageId="51" pageNumber="52" value="2001-01-17">17 ene 2001</date>
, Malaise, R. Cambra, A. Santos [col.] (MIUP).
<materialsCitation id="03F73C8B7C409B17FEF4F2DF9BEDFEC3" ID-GBIF-Occurrence="1671715712" collectingDate="1999-10-17" collectingDateMax="1999-10-24" collectingDateMin="1999-10-17" collectionCode="MIUP" collectorName="Tr &amp; Malaise, A &amp; Santos" country="Panama" latitude="9.406944" location="Teribe" longLatPrecision="21" longitude="-82.93889" municipality="Wekso" pageId="51" pageNumber="52" specimenCount="3" specimenCount-male="3" stateProvince="Bocas del Toro" typeStatus="paratype">
<typeStatus id="6C2488747C409B17FEF4F2DF9E10FF27" box="[260,373,224,248]" pageId="51" pageNumber="52" type="paratype">Paratypes</typeStatus>
:
<collectingRegion id="715BF8347C409B17FE76F2DE9D5FFF27" box="[390,570,223,248]" country="Panama" name="Bocas del Toro" pageId="51" pageNumber="52">
<emphasis id="81EBEAC47C409B17FE76F2DE9D5FFF27" box="[390,570,223,248]" italics="true" pageId="51" pageNumber="52">Bocas del Toro</emphasis>
</collectingRegion>
:
<specimenCount id="A599FD5F7C409B17FDBFF2DF9D11FF27" box="[591,628,224,248]" pageId="51" pageNumber="52" type="male">3♂</specimenCount>
,
<collectingMunicipality id="5344ACAC7C409B17FD76F2E09DB0FF27" box="[646,725,223,248]" pageId="51" pageNumber="52">Wekso</collectingMunicipality>
,
<location id="B640600D7C409B17FD16F2E09C55FF27" LSID="urn:lsid:plazi:treatment:3B3687C07C429B17FF67F7489B5FFE2A:B640600D7C409B17FD16F2E09C55FF27" box="[742,816,223,248]" country="Panama" latitude="9.406944" longLatPrecision="21" longitude="-82.93889" municipality="Wekso" name="Teribe" pageId="51" pageNumber="52" stateProvince="Bocas del Toro">Teribe</location>
, P[arque] I[nternacional] L[a] A[mistad], 50 msnm, [
<geoCoordinate id="D6AB50117C409B17FF6EF33B9E46FEC2" box="[158,291,260,285]" direction="north" orientation="latitude" pageId="51" pageNumber="52" precision="15" value="9.406944">09º24'25&quot;N</geoCoordinate>
,
<geoCoordinate id="D6AB50117C409B17FEC1F33B9EDBFEC2" box="[305,446,260,285]" direction="west" orientation="longitude" pageId="51" pageNumber="52" precision="15" value="-82.93889">82º56'20&quot;W</geoCoordinate>
],
<collectingDate id="D765E9FE7C409B17FE21F33B9DE9FEC3" box="[465,652,260,284]" pageId="51" pageNumber="52" value="1999-10-17" valueMax="1999-10-24" valueMin="1999-10-17">
<date id="C72110167C409B17FE21F33B9DE9FEC3" box="[465,652,260,284]" pageId="51" pageNumber="52" value="1999-10-17" valueMax="1999-10-24" valueMin="1999-10-17">17–24 oct. 1999</date>
</collectingDate>
,
<collectorName id="1E6A53007C409B17FD68F33A9DD1FEC3" box="[664,692,261,284]" pageId="51" pageNumber="52">Tr</collectorName>
[ampa]
<collectorName id="1E6A53007C409B17FCF9F33B9CECFEC3" box="[777,905,260,285]" pageId="51" pageNumber="52">Malaise, A</collectorName>
.
<collectorName id="1E6A53007C409B17FC64F33B9CBAFEC3" box="[916,991,260,284]" pageId="51" pageNumber="52">Santos</collectorName>
[col.] (
<collectionCode id="D58EAE137C409B17FBC3F33B9B1AFEC3" box="[1075,1151,260,284]" pageId="51" pageNumber="52">MIUP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="03F73C8B7C409B17FB64F33B9B14FEBB" ID-GBIF-Occurrence="1671715702" collectingDate="1999-09-26" collectingDateMax="1999-09-30" collectingDateMin="1999-09-26" collectionCode="MIUP" collectorName="El Cope &amp; de Division &amp; Omar Torrijos H &amp; Tr &amp; Malaise, A &amp; Santos, P" country="Panama" county="El Cope" elevation="620" latitude="8.698889" location="Nac" longLatPrecision="21" longitude="-80.654724" municipality="Par" pageId="51" pageNumber="52" specimenCount="1" specimenCount-male="1" stateProvince="Cocle Pr" typeStatus="paratype">
<emphasis id="81EBEAC47C409B17FB64F33B9A07FEC3" box="[1172,1378,260,285]" italics="true" pageId="51" pageNumber="52">
<collectingRegion id="715BF8347C409B17FB64F33B9B9CFEC3" box="[1172,1273,260,285]" isEnumeration="true" pageId="51" pageNumber="52">
<collectingRegion id="715BF8347C409B17FB64F33B9BB0FEC2" box="[1172,1237,260,285]" country="Panama" name="Cocle" pageId="51" pageNumber="52">Coclé</collectingRegion>
<collectingRegion id="715BF8347C409B17FB2DF33A9B9CFEC3" box="[1245,1273,261,284]" country="Brazil" name="Para" pageId="51" pageNumber="52">Pr</collectingRegion>
</collectingRegion>
[ovincia]
</emphasis>
:
<specimenCount id="A599FD5F7C409B17FA82F33B9AF3FEC3" box="[1394,1430,260,284]" pageId="51" pageNumber="52" type="male">1♂</specimenCount>
,
<collectingCounty id="5A414E5A7C409B17FF67F3189F9EFE9F" box="[151,251,295,320]" pageId="51" pageNumber="52">
<collectorName id="1E6A53007C409B17FF67F3189F9EFE9F" box="[151,251,295,320]" pageId="51" pageNumber="52">El Copé</collectorName>
</collectingCounty>
,
<collectingMunicipality id="5344ACAC7C409B17FEFCF3179E50FEE0" box="[268,309,296,319]" pageId="51" pageNumber="52">Par</collectingMunicipality>
[que]
<location id="B640600D7C409B17FE89F3189ECDFE9F" LSID="urn:lsid:plazi:treatment:3B3687C07C429B17FF67F7489B5FFE2A:B640600D7C409B17FE89F3189ECDFE9F" box="[377,424,295,320]" country="Panama" county="El Cope" latitude="8.698889" longLatPrecision="21" longitude="-80.654724" municipality="Par" name="Nac" pageId="51" pageNumber="52" stateProvince="Cocle Pr">Nac</location>
[ional
<location id="B640600D7C409B17FE05F3189D83FE9F" LSID="urn:lsid:plazi:treatment:3B3687C07C429B17FF67F7489B5FFE2A:B640600D7C409B17FE05F3189D83FE9F" box="[501,742,295,320]" country="Panama" county="El Cope" latitude="8.698889" longLatPrecision="21" longitude="-80.654724" municipality="Par" name="General de Division" pageId="51" pageNumber="52" stateProvince="Cocle Pr">
General
<collectorName id="1E6A53007C409B17FDAAF3189D83FE9F" box="[602,742,295,320]" pageId="51" pageNumber="52">de División</collectorName>
</location>
]
<collectorName id="1E6A53007C409B17FD09F3179CA5FEE0" box="[761,960,295,320]" pageId="51" pageNumber="52">Omar Torrijos H</collectorName>
[errera],
<elevation id="38B2D1E57C409B17FBD7F3189BD4FEE0" box="[1063,1201,295,320]" metricMagnitude="2" metricUnit="m" metricValue="6.2" metricValueMax="6.7" metricValueMin="5.7" pageId="51" pageNumber="52" unit="m" value="620.0" valueMax="670.0" valueMin="570.0">
<quantity id="74679B337C409B17FBD7F3189BD4FEE0" box="[1063,1201,295,320]" metricMagnitude="2" metricUnit="m" metricValue="6.2" metricValueMax="6.7" metricValueMin="5.7" pageId="51" pageNumber="52" unit="m" value="620.0" valueMax="670.0" valueMin="570.0">570–670 m</quantity>
</elevation>
.s.n.m., [
<geoCoordinate id="D6AB50117C409B17FAE1F3189AFDFE9F" box="[1297,1432,295,320]" direction="north" orientation="latitude" pageId="51" pageNumber="52" precision="15" value="8.698889">08º41'56&quot;N</geoCoordinate>
,
<geoCoordinate id="D6AB50117C409B17FF67F3739E41FEBB" box="[151,292,332,356]" direction="west" orientation="longitude" pageId="51" pageNumber="52" precision="15" value="-80.654724">80º39'17&quot;W</geoCoordinate>
],
<collectingDate id="D765E9FE7C409B17FEC5F3739E8AFEBB" box="[309,495,332,357]" pageId="51" pageNumber="52" value="1999-09-26" valueMax="1999-09-30" valueMin="1999-09-26">
<date id="C72110167C409B17FEC5F3739E8AFEBB" box="[309,495,332,357]" pageId="51" pageNumber="52" value="1999-09-26" valueMax="1999-09-30" valueMin="1999-09-26">26–30 sep. 1999</date>
</collectingDate>
,
<collectorName id="1E6A53007C409B17FE0AF3729D73FEBB" box="[506,534,333,356]" pageId="51" pageNumber="52">Tr</collectorName>
[ampa]
<collectorName id="1E6A53007C409B17FD99F3739D82FEBB" box="[617,743,332,357]" pageId="51" pageNumber="52">Malaise, A</collectorName>
.
<collectorName id="1E6A53007C409B17FD01F3739C3CFEBB" box="[753,857,332,356]" pageId="51" pageNumber="52">Santos, P</collectorName>
. González [col.] (
<collectionCode id="D58EAE137C409B17FBEDF3739B0CFEBB" box="[1053,1129,332,356]" pageId="51" pageNumber="52">MIUP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="03F73C8B7C409B17FB8CF3729EACFE73" ID-GBIF-Occurrence="1671715711" collectingDate="1999-08-10" collectingDateMax="1999-08-14" collectingDateMin="1999-08-10" collectionCode="MIUP" collectorName="Tr &amp; Malaise, A &amp; Santos, P &amp; Gonzalez" country="Panama" location="Altos de Campana" longitude="-80.02862" municipality="Nac" pageId="51" pageNumber="52" specimenCount="1" specimenCount-male="1" stateProvince="Panama" typeStatus="paratype">
<emphasis id="81EBEAC47C409B17FB8CF3729A01FEBB" box="[1148,1380,333,356]" italics="true" pageId="51" pageNumber="52">
<collectingRegion id="715BF8347C409B17FB8CF3729BBCFEBB" box="[1148,1241,333,356]" country="Panama" name="Panama" pageId="51" pageNumber="52">
<collectingCountry id="CB8876467C409B17FB8CF3729BBCFEBB" box="[1148,1241,333,356]" name="Panama" pageId="51" pageNumber="52">Panama</collectingCountry>
</collectingRegion>
<collectingRegion id="715BF8347C409B17FB2FF3729B9EFEBB" box="[1247,1275,333,356]" country="Brazil" name="Para" pageId="51" pageNumber="52">Pr</collectingRegion>
[ovincia]
</emphasis>
:
<specimenCount id="A599FD5F7C409B17FA82F3739AF3FEBB" box="[1394,1430,332,356]" pageId="51" pageNumber="52" type="male">1♂</specimenCount>
, P[arque]
<collectingMunicipality id="5344ACAC7C409B17FF0EF3509E4BFE57" box="[254,302,367,392]" pageId="51" pageNumber="52">Nac</collectingMunicipality>
[ional]
<location id="B640600D7C409B17FE8CF3509D28FE57" LSID="urn:lsid:plazi:treatment:3B3687C07C429B17FF67F7489B5FFE2A:B640600D7C409B17FE8CF3509D28FE57" box="[380,589,367,392]" country="Panama" longitude="-80.02862" municipality="Nac" name="Altos de Campana" pageId="51" pageNumber="52" stateProvince="Panama">Altos de Campana</location>
, [
<geoCoordinate id="D6AB50117C409B17FDAFF3509D8DFE57" box="[607,744,367,392]" direction="west" orientation="longitude" pageId="51" pageNumber="52" precision="15" value="-8.720833">08º43'15&quot;W</geoCoordinate>
,
<geoCoordinate id="D6AB50117C409B17FD04F34F9CE5FE57" box="[756,896,368,392]" direction="west" orientation="longitude" pageId="51" pageNumber="52" precision="15" value="-80.02862">80º01'43&quot;W</geoCoordinate>
],
<collectingDate id="D765E9FE7C409B17FC62F34F9B2CFE57" box="[914,1097,368,392]" pageId="51" pageNumber="52" value="1999-08-10" valueMax="1999-08-14" valueMin="1999-08-10">
<date id="C72110167C409B17FC62F34F9B2CFE57" box="[914,1097,368,392]" pageId="51" pageNumber="52" value="1999-08-10" valueMax="1999-08-14" valueMin="1999-08-10">10–14 ago 1999</date>
</collectingDate>
,
<collectorName id="1E6A53007C409B17FBA3F34F9B0AFE58" box="[1107,1135,368,391]" pageId="51" pageNumber="52">Tr</collectorName>
[ampa]
<collectorName id="1E6A53007C409B17FB32F3509A25FE57" box="[1218,1344,367,392]" pageId="51" pageNumber="52">Malaise, A</collectorName>
.
<collectorName id="1E6A53007C409B17FABBF34F9FC2FE73" pageId="51" pageNumber="52">Santos, P</collectorName>
.
<collectorName id="1E6A53007C409B17FF44F3AB9E7BFE72" box="[180,286,404,429]" pageId="51" pageNumber="52">González</collectorName>
[col.] (
<collectionCode id="D58EAE137C409B17FE85F3AB9EA4FE73" box="[373,449,404,428]" pageId="51" pageNumber="52">MIUP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="03F73C8B7C409B17FE27F3AB9C63FE72" ID-GBIF-Occurrence="1671715719" box="[471,774,404,429]" collectingDate="2000-11-16" country="Panama" county="Material" elevation="80" latitude="8.0" location="Pirre" longLatPrecision="1302" longitude="-77.75" municipality="Darien Pr" pageId="51" pageNumber="52" specimenCount="3" specimenCount-male="3" stateProvince="PANAMA" typeStatus="paratype">
<specimenCount id="A599FD5F7C409B17FE27F3AB9E9FFE73" box="[471,506,404,428]" pageId="51" pageNumber="52" type="male">3♂</specimenCount>
same data as holotype
</materialsCitation>
; 1♂, same of holotype except
<quantity id="74679B337C409B17FB9BF3AB9BA1FE73" box="[1131,1220,404,428]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="51" pageNumber="52" unit="m" value="1000.0">1000 m</quantity>
,
<date id="C72110167C409B17FB23F3AB9A24FE72" box="[1235,1345,404,429]" pageId="51" pageNumber="52" value="1921-03-04">21 mar–4</date>
ab 2000 (MIUP).
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C409B17FF37F3E39B5FFE2A" box="[199,1082,476,501]" pageId="51" pageNumber="52" type="distribution">
<paragraph id="B32036D67C409B17FF37F3E39B5FFE2A" blockId="51.[151,1436,151,501]" box="[199,1082,476,501]" pageId="51" pageNumber="52">
<emphasis id="81EBEAC47C409B17FF37F3E39E3CFE2A" bold="true" box="[199,345,476,501]" pageId="51" pageNumber="52">Distribution</emphasis>
(
<figureCitation id="2BA42A537C409B17FE98F3E39EAEFE2A" box="[360,459,476,501]" captionStart="FIGURES 244 – 249" captionStartId="68.[151,264,1212,1235]" captionTargetBox="[156,1430,201,1176]" captionTargetId="figure@68.[148,1433,193,1184]" captionTargetPageId="68" captionText="FIGURES 244 – 249. Known distribution of Dissomphalus in Panama. 244. D. rufipalpis, D. teraus sp. nov. and D. vallensis. 245. D. sartus sp. nov. and D. trisus sp. nov.. 246. D. teobus sp. nov. and D. urazius sp. nov.. 247. D. terbius sp. nov. and D. unitus. 248. D. ubracus sp. nov.. 249. D. rettenmeyeri." httpUri="https://zenodo.org/record/1010062/files/figure.png" pageId="51" pageNumber="52">Fig. 245</figureCitation>
).
<collectingRegion id="715BF8347C409B17FE2EF3E29D52FE2B" box="[478,567,477,500]" country="Panama" name="Panama" pageId="51" pageNumber="52">
<collectingCountry id="CB8876467C409B17FE2EF3E29D52FE2B" box="[478,567,477,500]" name="Panama" pageId="51" pageNumber="52">Panama</collectingCountry>
</collectingRegion>
(
<collectingRegion id="715BF8347C409B17FDB6F3E39D96FE2B" box="[582,755,476,501]" country="Panama" name="Bocas del Toro" pageId="51" pageNumber="52">Bocas del Toro</collectingRegion>
,
<collectingRegion id="715BF8347C409B17FD0EF3E39C27FE2A" box="[766,834,476,501]" country="Panama" name="Cocle" pageId="51" pageNumber="52">Coclé</collectingRegion>
,
<collectingRegion id="715BF8347C409B17FCBEF3E29CC2FE2B" box="[846,935,477,500]" country="Panama" name="Panama" pageId="51" pageNumber="52">
<collectingCountry id="CB8876467C409B17FCBEF3E29CC2FE2B" box="[846,935,477,500]" name="Panama" pageId="51" pageNumber="52">Panama</collectingCountry>
</collectingRegion>
and
<collectingRegion id="715BF8347C409B17FC2DF3E39B4BFE2A" box="[989,1070,476,501]" country="Panama" name="Darien" pageId="51" pageNumber="52">Darién</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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