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<document id="C9C2E93A4EA2400A0D56AF0914A44AEF" ID-CLB-Dataset="123002" ID-DOI="10.11646/zootaxa.5175.2.1" ID-GBIF-Dataset="86116a08-1bf6-48c3-b318-c54c8fd0928c" ID-ISSN="1175-5326" ID-Zenodo-Dep="7003344" ID-ZooBank="C4601ABE-4C6F-48BA-BA24-3339A44DC15C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1660735833844" checkinUser="plazi" docAuthor="Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong &amp; Watts, Chris H. S." docDate="2022" docId="3B0087B3145AFF9EFF3AFF0CFEDD4344" docLanguage="en" docName="zootaxa.5175.2.1.pdf" docOrigin="Zootaxa 5175 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Copelatus nakamurai Gueorguiev 1970" docType="treatment" docVersion="7" lastPageNumber="175" masterDocId="C739FFCB1442FF86FFADFF9BFFD0464A" masterDocTitle="An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae)" masterLastPageNumber="205" masterPageNumber="151" pageNumber="175" updateTime="1699367278572" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="912FFB9A682B71EA4656EF20DD274807">An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae)</mods:title>
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<mods:namePart id="BABA63D272C3A76AC4466150AE1E1137">Alarie, Yves</mods:namePart>
<mods:affiliation id="6A7C3BE43C7C8A29C344197C43E721E1">Department of Biology, Laurentian University, Ramsey Lake Road, Sudbury, Ontario, P 3 E 2 C 6, Canada</mods:affiliation>
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<mods:namePart id="44ABAAF229B96946B240755FB80486CF">Michat, Mariano C.</mods:namePart>
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<mods:affiliation id="FBA77898436C09FBC1BD4A8581CE5EDB">University of Buenos Aires, Faculty of Exact and Natural Sciences, Department of Biodiversity and Experimental and Applied Biology (IBBEA), Buenos Aires, Argentina. https: // orcid. org / 0000 - 0002 - 1962 - 7976</mods:affiliation>
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<mods:namePart id="8107811ADCA958F0E044F532F4CF4176">Watanabe, Kohei</mods:namePart>
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<mods:affiliation id="F95FAB50CDA0B073DCB021878F917B66">Ishikawa Insect Museum, Inu- 3, Yawata-machi, Hakusan-shi, 920 - 2113 Japan. https: // orcid. org / 0000 - 0002 - 8761 - 232 X Naturhistorisches Museum Wien, Burgring 7, 1010 Vienna, Austria. https: // orcid. org / 0000 - 0001 - 5034 - 7342</mods:affiliation>
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<mods:namePart id="7A1CD2FF8BDC0F9CF2991A9A1F3AEB58">Shaverdo, Helena</mods:namePart>
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<mods:namePart id="37FCA749D30F7FC14267B21A80437488">Wang, Liang-Jong</mods:namePart>
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<mods:affiliation id="BFB1B64E1D0FBB96ED33A69DF6F3B903">Division of Forest Protection, Taiwan Forestry Research Institute, Taipei, Taiwan. https: // orcid. org / 0000 - 0001 - 5540 - 2021</mods:affiliation>
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<mods:namePart id="A30886BD12A4BCD4E9D0AFFAA68F4EAC">Watts, Chris H. S.</mods:namePart>
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<treatment id="3B0087B3145AFF9EFF3AFF0CFEDD4344" ID-DOI="http://doi.org/10.5281/zenodo.7003370" ID-GBIF-Taxon="199499363" ID-Zenodo-Dep="7003370" LSID="urn:lsid:plazi:treatment:3B0087B3145AFF9EFF3AFF0CFEDD4344" httpUri="http://treatment.plazi.org/id/3B0087B3145AFF9EFF3AFF0CFEDD4344" lastPageNumber="175" pageId="24" pageNumber="175">
<subSubSection id="FBB3652E145AFF9EFF3AFF0CFDAF46FB" box="[151,639,150,178]" pageId="24" pageNumber="175" type="nomenclature">
<paragraph id="B31636A5145AFF9EFF3AFF0CFDAF46FB" blockId="24.[151,639,150,178]" box="[151,639,150,178]" pageId="24" pageNumber="175">
<heading id="E85E81C9145AFF9EFF3AFF0CFDAF46FB" bold="true" box="[151,639,150,178]" fontSize="11" level="1" pageId="24" pageNumber="175" reason="1">
<taxonomicName id="74A94D26145AFF9EFF3AFF0CFDAF46FB" ID-CoL="Y3CN" authority="Gueorguiev, 1970" authorityName="Gueorguiev" authorityYear="1970" box="[151,639,150,178]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="175" phylum="Arthropoda" rank="species" species="nakamurai">
<emphasis id="81DDEAB7145AFF9EFF3AFF0CFDAF46FB" bold="true" box="[151,639,150,178]" pageId="24" pageNumber="175">
<emphasis id="81DDEAB7145AFF9EFF3AFF0CFE4D46F8" bold="true" box="[151,413,151,178]" italics="true" pageId="24" pageNumber="175">Copelatus nakamurai</emphasis>
Guéorguiev, 1970
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="FBB3652E145AFF9EFF3AFF44FAC34757" pageId="24" pageNumber="175" type="materials_examined">
<paragraph id="B31636A5145AFF9EFF3AFF44FAC34757" blockId="24.[151,1437,223,1294]" pageId="24" pageNumber="175">
<emphasis id="81DDEAB7145AFF9EFF3AFF44FEAC46B3" bold="true" box="[151,380,223,249]" pageId="24" pageNumber="175">Source of material.</emphasis>
<materialsCitation id="03C13CF8145AFF9EFE2EFF44FAC34757" ID-GBIF-Occurrence="3889729301" collectingDate="2018-05-04" collectorName="Y. Yoshimura" country="Japan" county="The" location="Mudoji" municipality="Kasamatshu-cho" pageId="24" pageNumber="175" specimenCount="1" stateProvince="Gifu">
<collectingCounty id="5A774E29145AFF9EFE2EFF44FE7F46B3" box="[387,431,223,249]" pageId="24" pageNumber="175">The</collectingCounty>
two instar II and five instar III larvae studied were reared
<emphasis id="81DDEAB7145AFF9EFBEEFF7BFB5B46B3" box="[1091,1163,224,249]" italics="true" pageId="24" pageNumber="175">ex ovo</emphasis>
from adults collected at the following locality:
<emphasis id="81DDEAB7145AFF9EFE3AFE98FE374757" bold="true" box="[407,487,259,285]" pageId="24" pageNumber="175">
<collectingCountry id="CBBE7635145AFF9EFE3AFE98FE324757" box="[407,482,259,285]" name="Japan" pageId="24" pageNumber="175">Japan</collectingCountry>
.
</emphasis>
<collectingMunicipality id="5372ACDF145AFF9EFE43FE98FD7C4757" box="[494,684,259,285]" pageId="24" pageNumber="175">Kasamatshu-cho</collectingMunicipality>
,
<location id="B676607E145AFF9EFD1AFE98FCDC4757" LSID="urn:lsid:plazi:treatment:3B0087B3145AFF9EFF3AFF0CFEDD4344:B676607E145AFF9EFD1AFE98FCDC4757" box="[695,780,259,285]" country="Japan" county="The" municipality="Kasamatshu-cho" name="Mudoji" pageId="24" pageNumber="175" stateProvince="Gifu">Mudoji</location>
,
<collectingRegion id="716DF847145AFF9EFCBAFE98FC184757" box="[791,968,259,285]" country="Japan" name="Gifu" pageId="24" pageNumber="175">Gifu Prefecture</collectingRegion>
,
<date id="C7171065145AFF9EFC7EFE98FBEB4757" box="[979,1083,259,285]" pageId="24" pageNumber="175" value="2018-05-04">
<collectingDate id="D753E98D145AFF9EFC7EFE98FBEB4757" box="[979,1083,259,285]" pageId="24" pageNumber="175" value="2018-05-04">4.V.2018</collectingDate>
</date>
.
<collectorName id="1E5C5373145AFF9EFBE9FE9FFB0F4757" box="[1092,1247,259,286]" pageId="24" pageNumber="175">Y. Yoshimura</collectorName>
, leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="FBB3652E145AFF9EFF6AFEBCFC564477" pageId="24" pageNumber="175" type="diagnosis">
<paragraph id="B31636A5145AFF9EFF6AFEBCFC564477" blockId="24.[151,1437,223,1294]" pageId="24" pageNumber="175">
<emphasis id="81DDEAB7145AFF9EFF6AFEBCFE1C470B" bold="true" box="[199,460,295,321]" pageId="24" pageNumber="175">Diagnosis (instar III).</emphasis>
The third instar of
<taxonomicName id="74A94D26145AFF9EFD02FEBCFC72470B" authorityName="Gueorguiev" authorityYear="1970" box="[687,930,295,321]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="175" phylum="Arthropoda" rank="species" species="nakamurai">
<emphasis id="81DDEAB7145AFF9EFD02FEBCFC72470B" box="[687,930,295,321]" italics="true" pageId="24" pageNumber="175">Copelatus nakamurai</emphasis>
</taxonomicName>
can easily be distinguished from that of the other species studied in this paper by the following combination of characters: HL =
<quantity id="74519B40145AFF9EFBE2FED0FB66472C" box="[1103,1206,331,358]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="24" pageNumber="175" unit="mm" value="0.8">0.80 mm</quantity>
; L3 &lt;2.00 mm; U &lt;
<quantity id="74519B40145AFF9EFF3AFEF4FF2E47C0" box="[151,254,367,394]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="24" pageNumber="175" unit="mm" value="0.4">0.40 mm</quantity>
; head capsule subquadrate, strongly constricted at level of occipital region (
<figureCitation id="2B922A20145AFF9EFBE0FEF4FB7047C3" box="[1101,1184,367,393]" captionStart="FIGURES 2829" captionStartId="25.[151,264,1943,1967]" captionTargetBox="[242,1341,196,1903]" captionTargetId="figure-18@25.[229,1357,181,1919]" captionTargetPageId="25" captionText="FIGURES 2829.Copelatus nakamurai Guéorguiev, 1970, instar III:(28) headcapsule, dorsal aspect (scale-likemicrosculptures not represented); (29) abdominal segment VIII, dorsal aspect. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7003392" httpUri="https://zenodo.org/record/7003392/files/figure.png" pageId="24" pageNumber="175">Fig. 28</figureCitation>
), scale-like sculpticels lacking over frontoclypeus, present over parietale; anterior margin of frontoclypeus narrowly convex, extending mesally well beyond adnasalia (
<figureCitation id="2B922A20145AFF9EFE50FE2CFD81479B" box="[509,593,439,465]" captionStart="FIGURES 2829" captionStartId="25.[151,264,1943,1967]" captionTargetBox="[242,1341,196,1903]" captionTargetId="figure-18@25.[229,1357,181,1919]" captionTargetPageId="25" captionText="FIGURES 2829.Copelatus nakamurai Guéorguiev, 1970, instar III:(28) headcapsule, dorsal aspect (scale-likemicrosculptures not represented); (29) abdominal segment VIII, dorsal aspect. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7003392" httpUri="https://zenodo.org/record/7003392/files/figure.png" pageId="24" pageNumber="175">Fig. 28</figureCitation>
); adnasalia margined with elongate acute teeth (
<figureCitation id="2B922A20145AFF9EFBC4FE2CFB6D479B" box="[1129,1213,439,465]" captionStart="FIGURE 64" captionStartId="44.[151,250,1943,1967]" captionTargetBox="[229,1366,195,1901]" captionTargetId="figure-20@44.[210,1377,181,1919]" captionTargetPageId="44" captionText="FIGURE 64. Anterior margin of left adnasalia of selected species of Copelatinae; magnification of 500x. Species codes as in Table 1." figureDoi="http://doi.org/10.5281/zenodo.7003434" httpUri="https://zenodo.org/record/7003434/files/figure.png" pageId="24" pageNumber="175">Fig. 64</figureCitation>
); MP2/MP1&gt; 1.50; GA/MP1&gt; 1.50; 1.30 &lt;LP2/LP1 &lt;1.90; inner margin of stipes lacking a dorsal linear row of spinulae; profemur with less than 3 PV and 4 AV secondary setae; metafemur with less than 3 AV secondary setae; urogomphus composed of one urogomphomere; U/HW &lt;0.40; U/LAS &lt;0.50; Palaearctic.
</paragraph>
</subSubSection>
<subSubSection id="FBB3652E145AFF9EFF6AFDDCFEDD4344" pageId="24" pageNumber="175" type="description">
<paragraph id="B31636A5145AFF9EFF6AFDDCFE12442B" blockId="24.[151,1437,223,1294]" box="[199,450,583,609]" pageId="24" pageNumber="175">
<emphasis id="81DDEAB7145AFF9EFF6AFDDCFE12442B" bold="true" box="[199,450,583,609]" pageId="24" pageNumber="175">Description, instar II</emphasis>
</paragraph>
<paragraph id="B31636A5145AFF9EFF6AFDF0FB7044CF" blockId="24.[151,1437,223,1294]" box="[199,1184,619,646]" pageId="24" pageNumber="175">
<emphasis id="81DDEAB7145AFF9EFF6AFDF0FEDF44CF" bold="true" box="[199,271,619,645]" pageId="24" pageNumber="175">Body:</emphasis>
Measurements and ratios aimed to characterize body shape are shown in
<tableCitation id="FE2B031E145AFF9EFBEBFDF0FB4C44CF" box="[1094,1180,619,645]" captionStart="TABLE 4" captionStartId="5.[151,239,151,175]" captionTargetPageId="5" captionText="TABLE 4. Measurements and ratios for the instar II of selected species of Copelatinae (species codes as in Table 1)." httpUri="http://table.plazi.org/id/E7D6662D1447FF83FF3AFF0CFAA046FA" pageId="24" pageNumber="175" tableUuid="E7D6662D1447FF83FF3AFF0CFAA046FA">Table 4</tableCitation>
.
</paragraph>
<paragraph id="B31636A5145AFF9EFF6AFD14FB50455F" blockId="24.[151,1437,223,1294]" pageId="24" pageNumber="175">
<emphasis id="81DDEAB7145AFF9EFF6AFD14FEC044E3" bold="true" box="[199,272,655,681]" pageId="24" pageNumber="175">Head:</emphasis>
Head capsule subquadrate, strongly constricted at level of occiput; anterior margin of frontoclypeus narrowly convex, extending well beyond level of adnasalia; scale-like sculpticels lacking over frontoclypeus, welldeveloped over parietale; adnasalia margined with elongate acute teeth (see
<figureCitation id="2B922A20145AFF9EFC44FD4CFBEC44BB" box="[1001,1084,727,753]" captionStart="FIGURE 64" captionStartId="44.[151,250,1943,1967]" captionTargetBox="[229,1366,195,1901]" captionTargetId="figure-20@44.[210,1377,181,1919]" captionTargetPageId="44" captionText="FIGURE 64. Anterior margin of left adnasalia of selected species of Copelatinae; magnification of 500x. Species codes as in Table 1." figureDoi="http://doi.org/10.5281/zenodo.7003434" httpUri="https://zenodo.org/record/7003434/files/figure.png" pageId="24" pageNumber="175">Fig. 64</figureCitation>
); HL =
<quantity id="74519B40145AFF9EFB3FFD4CFB2944B8" box="[1170,1273,727,754]" metricMagnitude="-4" metricUnit="m" metricValue="6.1" pageId="24" pageNumber="175" unit="mm" value="0.61">0.61 mm</quantity>
; A/MP = 1.49; MP/LP = 2.06; MP2/MP1 = 1.60; GA/MP1 = 1.91; LP2/LP1 = 2.24; MNL/MNW = 3.11.
</paragraph>
<paragraph id="B31636A5145AFF9EFF6AFC84FE084570" blockId="24.[151,1437,223,1294]" box="[199,472,799,826]" pageId="24" pageNumber="175">
<emphasis id="81DDEAB7145AFF9EFF6AFC84FEF94573" bold="true" box="[199,297,799,825]" pageId="24" pageNumber="175">Thorax:</emphasis>
L3 =
<quantity id="74519B40145AFF9EFEC0FC84FE044570" box="[365,468,799,826]" metricMagnitude="-3" metricUnit="m" metricValue="1.23" pageId="24" pageNumber="175" unit="mm" value="1.23">1.23 mm</quantity>
.
</paragraph>
<paragraph id="B31636A5145AFF9EFF6AFCD8FA9245C8" blockId="24.[151,1437,223,1294]" pageId="24" pageNumber="175">
<emphasis id="81DDEAB7145AFF9EFF6AFCD8FE954517" bold="true" box="[199,325,835,861]" pageId="24" pageNumber="175">Abdomen:</emphasis>
LAS =
<quantity id="74519B40145AFF9EFE02FCD8FDCB4514" box="[431,539,835,862]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="24" pageNumber="175" unit="mm" value="0.45">0.45 mm</quantity>
; LAS subcylindrical, slightly constricted posteriorly at point of insertion of urogomphi. Urogomphus: U =
<quantity id="74519B40145AFF9EFE5EFCFCFD8945C8" box="[499,601,871,898]" metricMagnitude="-4" metricUnit="m" metricValue="2.6" pageId="24" pageNumber="175" unit="mm" value="0.26">0.26 mm</quantity>
, composed of one urogomphomere; U/HW = 0.41; U/LAS = 0.58.
</paragraph>
<paragraph id="B31636A5145AFF9EFF6AFC10FF214580" blockId="24.[151,1437,223,1294]" pageId="24" pageNumber="175">
<emphasis id="81DDEAB7145AFF9EFF6AFC10FE8945EF" bold="true" box="[199,345,907,933]" pageId="24" pageNumber="175">Chaetotaxy:</emphasis>
Parietale with 8 temporal spines; position and number of secondary setae on legs are shown in
<tableCitation id="FE2B031E145AFF9EFF3AFC34FF3C4580" box="[151,236,943,970]" captionStart="TABLE 5" captionStartId="6.[151,239,151,175]" captionTargetPageId="6" captionText="TABLE 5. Number of secondary setae(including additional setae) on the legs of instar II of selected species of Copelatinae (species codes as in Table 1). A = anterior, AV = anteroventral, CO = coxa, D = dorsal, FE = femur, PV = posteroventral, n = number of specimens studied, Total = total number of secondary setae on segment (including additional setae) (species codes as in Table 1)." httpUri="http://table.plazi.org/id/E7D6662D1444FF80FF3AFF0CFEA14756" pageId="24" pageNumber="175" tableUuid="E7D6662D1444FF80FF3AFF0CFEA14756">Table 5</tableCitation>
.
</paragraph>
<paragraph id="B31636A5145AFF9EFF6AFC48FBB945A7" blockId="24.[151,1437,223,1294]" box="[199,1129,979,1006]" pageId="24" pageNumber="175">
<emphasis id="81DDEAB7145AFF9EFF6AFC48FD4645A7" bold="true" box="[199,662,979,1005]" pageId="24" pageNumber="175">
Description, instar III (
<figureCitation id="2B922A20145AFF9EFE76FC48FD8D45A7" box="[475,605,979,1005]" captionStart="FIGURES 2829" captionStartId="25.[151,264,1943,1967]" captionTargetBox="[242,1341,196,1903]" captionTargetId="figure-18@25.[229,1357,181,1919]" captionTargetPageId="25" captionText="FIGURES 2829.Copelatus nakamurai Guéorguiev, 1970, instar III:(28) headcapsule, dorsal aspect (scale-likemicrosculptures not represented); (29) abdominal segment VIII, dorsal aspect. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7003392" httpUri="https://zenodo.org/record/7003392/files/figure.png" pageId="24" pageNumber="175">Figs 2829</figureCitation>
,
<figureCitation id="2B922A20145AFF9EFDC7FC48FD5845A7" box="[618,648,979,1005]" captionStart="FIGURE 64" captionStartId="44.[151,250,1943,1967]" captionTargetBox="[229,1366,195,1901]" captionTargetId="figure-20@44.[210,1377,181,1919]" captionTargetPageId="44" captionText="FIGURE 64. Anterior margin of left adnasalia of selected species of Copelatinae; magnification of 500x. Species codes as in Table 1." figureDoi="http://doi.org/10.5281/zenodo.7003434" httpUri="https://zenodo.org/record/7003434/files/figure.png" pageId="24" pageNumber="175">64</figureCitation>
).
</emphasis>
As second-instar larva except as follows:
</paragraph>
<paragraph id="B31636A5145AFF9EFF6AFC6CFB70425B" blockId="24.[151,1437,223,1294]" box="[199,1184,1015,1042]" pageId="24" pageNumber="175">
<emphasis id="81DDEAB7145AFF9EFF6AFC6CFEDF425B" bold="true" box="[199,271,1015,1041]" pageId="24" pageNumber="175">Body:</emphasis>
Measurements and ratios aimed to characterize body shape are shown in
<tableCitation id="FE2B031E145AFF9EFBEBFC6CFB4C425B" box="[1094,1180,1015,1041]" captionStart="TABLE 6" captionStartId="7.[151,239,151,175]" captionTargetPageId="7" captionText="TABLE 6. Measurements and ratios for the instar III of selected species of Copelatinae (species codes as in Table 1)." pageId="24" pageNumber="175">Table 6</tableCitation>
.
</paragraph>
<paragraph id="B31636A5145AFF9EFF6AFB80FC6F4210" blockId="24.[151,1437,223,1294]" pageId="24" pageNumber="175">
<emphasis id="81DDEAB7145AFF9EFF6AFB80FE60427F" bold="true" box="[199,432,1051,1077]" pageId="24" pageNumber="175">
Head (
<figureCitation id="2B922A20145AFF9EFEB5FB80FEA2427F" box="[280,370,1051,1077]" captionStart="FIGURES 2829" captionStartId="25.[151,264,1943,1967]" captionTargetBox="[242,1341,196,1903]" captionTargetId="figure-18@25.[229,1357,181,1919]" captionTargetPageId="25" captionText="FIGURES 2829.Copelatus nakamurai Guéorguiev, 1970, instar III:(28) headcapsule, dorsal aspect (scale-likemicrosculptures not represented); (29) abdominal segment VIII, dorsal aspect. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7003392" httpUri="https://zenodo.org/record/7003392/files/figure.png" pageId="24" pageNumber="175">Figs 28</figureCitation>
,
<figureCitation id="2B922A20145AFF9EFE2CFB80FE71427F" box="[385,417,1051,1077]" captionStart="FIGURE 64" captionStartId="44.[151,250,1943,1967]" captionTargetBox="[229,1366,195,1901]" captionTargetId="figure-20@44.[210,1377,181,1919]" captionTargetPageId="44" captionText="FIGURE 64. Anterior margin of left adnasalia of selected species of Copelatinae; magnification of 500x. Species codes as in Table 1." figureDoi="http://doi.org/10.5281/zenodo.7003434" httpUri="https://zenodo.org/record/7003434/files/figure.png" pageId="24" pageNumber="175">64</figureCitation>
):
</emphasis>
HL =
<quantity id="74519B40145AFF9EFE52FB80FD77427C" box="[511,679,1051,1078]" metricMagnitude="-4" metricUnit="m" metricValue="8.25" metricValueMax="8.4" metricValueMin="8.1" pageId="24" pageNumber="175" unit="mm" value="0.825" valueMax="0.84" valueMin="0.81">0.810.84 mm</quantity>
; A/MP = 1.421.46; MP/LP = 2.022.25; MP2/MP1 = 1.511.74; GA/MP1 = 1.751.99; LP2/LP1 = 1.551.78; MNL/MNW = 2.852.98.
</paragraph>
<paragraph id="B31636A5145AFF9EFF6AFBF8FDC74234" blockId="24.[151,1437,223,1294]" box="[199,535,1123,1150]" pageId="24" pageNumber="175">
<emphasis id="81DDEAB7145AFF9EFF6AFBF8FEF94237" bold="true" box="[199,297,1123,1149]" pageId="24" pageNumber="175">Thorax:</emphasis>
L3 =
<quantity id="74519B40145AFF9EFEC0FBF8FDC34234" box="[365,531,1123,1150]" metricMagnitude="-3" metricUnit="m" metricValue="1.69" metricValueMax="1.72" metricValueMin="1.66" pageId="24" pageNumber="175" unit="mm" value="1.69" valueMax="1.72" valueMin="1.66">1.661.72 mm</quantity>
.
</paragraph>
<paragraph id="B31636A5145AFF9EFF6AFB1CFDB4428C" blockId="24.[151,1437,223,1294]" pageId="24" pageNumber="175">
<emphasis id="81DDEAB7145AFF9EFF6AFB1CFE7542EB" bold="true" box="[199,421,1159,1185]" pageId="24" pageNumber="175">
Abdomen (
<figureCitation id="2B922A20145AFF9EFEE5FB1CFE4642EB" box="[328,406,1159,1185]" captionStart="FIGURES 2829" captionStartId="25.[151,264,1943,1967]" captionTargetBox="[242,1341,196,1903]" captionTargetId="figure-18@25.[229,1357,181,1919]" captionTargetPageId="25" captionText="FIGURES 2829.Copelatus nakamurai Guéorguiev, 1970, instar III:(28) headcapsule, dorsal aspect (scale-likemicrosculptures not represented); (29) abdominal segment VIII, dorsal aspect. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7003392" httpUri="https://zenodo.org/record/7003392/files/figure.png" pageId="24" pageNumber="175">Fig 29</figureCitation>
):
</emphasis>
LAS =
<quantity id="74519B40145AFF9EFE5AFB1CFD4B42E8" box="[503,667,1159,1186]" metricMagnitude="-4" metricUnit="m" metricValue="6.75" metricValueMax="6.8999999999999995" metricValueMin="6.6" pageId="24" pageNumber="175" unit="mm" value="0.675" valueMax="0.69" valueMin="0.66">0.660.69 mm</quantity>
; Urogomphus, U =
<quantity id="74519B40145AFF9EFCDDFB1CFBC242E8" box="[880,1042,1159,1186]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" metricValueMax="3.3" metricValueMin="3.1" pageId="24" pageNumber="175" unit="mm" value="0.32" valueMax="0.33" valueMin="0.31">0.310.33 mm</quantity>
, composed of one urogomphomere; U/HW = 0.350.38; U/LAS = 0.450.49.
</paragraph>
<paragraph id="B31636A5145AFF9EFF6AFB54FEDD4344" blockId="24.[151,1437,223,1294]" pageId="24" pageNumber="175">
<emphasis id="81DDEAB7145AFF9EFF6AFB54FE8942A3" bold="true" box="[199,345,1231,1257]" pageId="24" pageNumber="175">Chaetotaxy:</emphasis>
Parietale with 910 temporal spines; position and number of secondary setae on legs are shown in
<tableCitation id="FE2B031E145AFF9EFF1EFB68FED94344" box="[179,265,1267,1294]" captionStart="TABLE 7" captionStartId="9.[151,239,566,592]" captionTargetPageId="9" captionText="TABLE 7. Number of secondary setae (including additional setae) on the legs of instar III of selected species of Copelatinae (species codes as in Table 1). A = anterior, AV = anteroventral, CO = coxa, D = dorsal, FE = femur, Pr = proximal, PV = posteroventral, TR = trochanter, n = number of specimens studied, Total = total number of secondary setae on segment." httpUri="http://table.plazi.org/id/E7D6662D144BFF8FFF3AFDADFEC944D2" pageId="24" pageNumber="175" tableUuid="E7D6662D144BFF8FFF3AFDADFEC944D2">Table 7</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>