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<document id="29F90771DA2C9508C4324CD7F8C14AA5" ID-DOI="10.21248/contrib.entomol.71.1.029-085" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742999" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638270695310" checkinUser="felipe" docAuthor="Irmler, Ulrich" docDate="2021" docId="3A6C87E6FF934C7FFCD1DB8D01860711" docLanguage="en" docName="ContribEntomol.71.1.29-85.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 71 (1)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Lithocharodes denticulata Irmler 2021, spec. nov." docType="treatment" docVersion="3" lastPageNumber="41" masterDocId="C655FF9EFF984C73FFEEDB7E00260230" masterDocTitle="The Neotropical species of the genus Lithocharodes SHARP, 1876 (Coleoptera: Staphylinidae: Staphylininae: Xantholinini)" masterLastPageNumber="85" masterPageNumber="29" pageNumber="40" updateTime="1699437350205" updateUser="plazi">
<mods:mods id="D88A56E094A8963FF882A5B2195B9BE7" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B56AE29D9AF5146B733CBCF8BEAAE95E">
<mods:title id="2B4C2B3084E8984F97EA19AEA306A0D4">The Neotropical species of the genus Lithocharodes SHARP, 1876 (Coleoptera: Staphylinidae: Staphylininae: Xantholinini)</mods:title>
</mods:titleInfo>
<mods:name id="2BE50497683CAD61582F9BE3DF1F1073" type="personal">
<mods:role id="2D30FB96BB78F13EB6BF21B42F19EC24">
<mods:roleTerm id="4786A1CA80AD7F22A50C782300A8D005">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EEF0B5DB0B75DD629C5C48AF40AC1A89">Irmler, Ulrich</mods:namePart>
</mods:name>
<mods:typeOfResource id="D098AE6B0DACA602052519EB7AEB8004">text</mods:typeOfResource>
<mods:relatedItem id="F336A793B6CD5ED939CD468EC2E868AE" type="host">
<mods:titleInfo id="57DE141499F458E8B43903B4090B816B">
<mods:title id="FE99DC87BEE2F9D6BFC12C4BF37FE6E1">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
</mods:titleInfo>
<mods:part id="608CC838BA8785AFDB798CE90E88F9A5">
<mods:date id="27C62470EBF95A5DD2E0D87F2C343980">2021</mods:date>
<mods:detail id="2AD6A06302D4917F756D43A817A77D2F" type="pubDate">
<mods:number id="78C63D01C466287CBF82018F54647B09">2021-06-30</mods:number>
</mods:detail>
<mods:detail id="902F01569D2C221D39C2829A7E59DE6B" type="volume">
<mods:number id="B269DD50EC7E8FEC4DD0626C038C14EB">71</mods:number>
</mods:detail>
<mods:detail id="A224C5F62074152B51F767DA5CAA95A1" type="issue">
<mods:number id="02D79F89A851402F1740D6E89DDFA405">1</mods:number>
</mods:detail>
<mods:extent id="FCE0FB067C1E0A5DD7F099F535B3773F" unit="page">
<mods:start id="C04305FB7820EA227E7E749CF3D43DDB">29</mods:start>
<mods:end id="FE524CC2136725A25F6F87FCFE0DAD44">85</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="78C51876089F7FFAAAB3ED662DC3AB7B">journal article</mods:classification>
<mods:identifier id="52F7434660D7EE440B6B11A86CCD28E4" type="DOI">10.21248/contrib.entomol.71.1.029-085</mods:identifier>
<mods:identifier id="E85656FBEFF6BCC42733C8740C7A6F21" type="ISSN">0005-805X</mods:identifier>
<mods:identifier id="8945AAC9BB036FF4F055C495BE02015D" type="Zenodo-Dep">5742999</mods:identifier>
</mods:mods>
<treatment id="3A6C87E6FF934C7FFCD1DB8D01860711" LSID="urn:lsid:plazi:treatment:3A6C87E6FF934C7FFCD1DB8D01860711" httpUri="http://treatment.plazi.org/id/3A6C87E6FF934C7FFCD1DB8D01860711" lastPageId="12" lastPageNumber="41" pageId="11" pageNumber="40">
<subSubSection id="FADF657BFF934C78FCD1DB8D051B031F" pageId="11" pageNumber="40" type="nomenclature">
<paragraph id="B27A36F0FF934C78FCD1DB8D049F033E" blockId="11.[831,1341,243,335]" box="[831,1209,243,270]" pageId="11" pageNumber="40">
<heading id="E932819CFF934C78FCD1DB8D049F033E" box="[831,1209,243,270]" fontSize="12" level="2" pageId="11" pageNumber="40" reason="2">
<taxonomicName id="75C54D73FF934C78FCD1DB8D046C033E" ID-CoL="8Y63D" authority="Irmler, 2021" authorityName="Irmler" authorityYear="2021" box="[831,1098,243,270]" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="40" phylum="Arthropoda" rank="species" species="denticulata" status="sp. nov.">
<emphasis id="80B1EAE2FF934C78FCD1DB8D046C033E" box="[831,1098,243,270]" italics="true" pageId="11" pageNumber="40">Lithocharodes denticulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9B825799FF934C78FBBEDB8D049F033E" box="[1104,1209,243,270]" pageId="11" pageNumber="40" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="B27A36F0FF934C78FCD1DA65051B031F" blockId="11.[831,1341,243,335]" box="[831,1341,283,303]" pageId="11" pageNumber="40">
<uri id="C6543AF2FF934C78FCD1DA65051B031F" box="[831,1341,283,303]" pageId="11" pageNumber="40">
urn: lsid:zoobank. org:act:
<uuid id="C6630C25FF934C78FC1EDA65051B031F" box="[1008,1341,283,303]" pageId="11" pageNumber="40">9B194FB8-EFFE-4301-9681-174E24B19557</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="FADF657BFF934C78FCD1DA4503C9037F" box="[831,1007,315,335]" pageId="11" pageNumber="40" type="description">
<paragraph id="B27A36F0FF934C78FCD1DA4503C9037F" blockId="11.[831,1341,243,335]" box="[831,1007,315,335]" pageId="11" pageNumber="40">
<figureCitation id="2AFE2A75FF934C78FCD1DA450353037F" box="[831,885,315,335]" captionStart="Fig" captionStartId="41.[151,184,1966,1988]" captionTargetBox="[174,1383,251,1941]" captionTargetId="figure-110@41.[151,1398,235,1956]" captionTargetPageId="41" captionText="Fig. 13–18: L.-simillima-group:Lithocharodes triangula (13), L. simillima(14), L. surinamensis (15), L. brooksi(16), L. germana (17), L. denticulata (18); aedeagus (a), paramere (b), sternite VII of male (c), tergite VII of male (d); scale bare: A–D: 0.2 mm; B: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743011" httpUri="https://zenodo.org/record/5743011/files/figure.png" pageId="11" pageNumber="40">Figs 18</figureCitation>
a-d, 44 M, 49 A
</paragraph>
</subSubSection>
<subSubSection id="FADF657BFF934C78FCF4DA1703D50711" pageId="11" pageNumber="40" type="materials_examined">
<paragraph id="B27A36F0FF934C78FCF4DA1703D50711" blockId="11.[794,1399,361,1313]" pageId="11" pageNumber="40">
<emphasis id="80B1EAE2FF934C78FCF4DA17039E03B1" bold="true" box="[794,952,361,385]" pageId="11" pageNumber="40">Type material</emphasis>
:
<materialsCitation id="02AD3CADFF934C78FC22DA1703A70031" collectingDate="1999-02-13" collectionCode="KNHM" collectorName="F. Genier" country="Bolivia" elevation="300" latitude="-17.109167" location="Univ. de San Simon" longLatPrecision="21" longitude="-64.797775" pageId="11" pageNumber="40" specimenCount="1" stateProvince="Cochabamba" typeStatus="holotype">
male,
<typeStatus id="6D7E8852FF934C78FBFDDA17045203B1" box="[1043,1140,361,385]" pageId="11" pageNumber="40" type="holotype">holotype</typeStatus>
:
<collectingCountry id="CAD27660FF934C78FB6BDA1704F403B1" box="[1157,1234,361,385]" name="Bolivia" pageId="11" pageNumber="40">Bolivia</collectingCountry>
,
<collectingRegion id="7001F812FF934C78FB0DDA17055403B1" box="[1251,1394,361,385]" country="Bolivia" name="Cochabamba" pageId="11" pageNumber="40">Cochabamba</collectingRegion>
,
<collectingRegion id="7001F812FF934C78FCF4DAF7038F0391" box="[794,937,393,417]" country="Bolivia" name="Cochabamba" pageId="11" pageNumber="40">Cochabamba</collectingRegion>
,
<locationDeviation id="A3C82FD2FF934C78FC5DDAF705120391" box="[947,1332,393,417]" location="Est. Biol. Valle de Sajita" pageId="11" pageNumber="40">
<quantity id="753D9B15FF934C78FC5DDAF7042F0391" box="[947,1033,393,417]" metricMagnitude="4" metricUnit="m" metricValue="6.75" pageId="11" pageNumber="40" unit="km" value="67.5">67.5 km</quantity>
NE Est. Biol. Valle de Sajita
</locationDeviation>
,
<location id="B71A602BFF934C78FAD0DAF7038A03F1" LSID="urn:lsid:plazi:treatment:3A6C87E6FF934C7FFCD1DB8D01860711:B71A602BFF934C78FAD0DAF7038A03F1" country="Bolivia" latitude="-17.109167" longLatPrecision="21" longitude="-64.797775" name="Univ. de San Simon" pageId="11" pageNumber="40" stateProvince="Cochabamba">Univ. de San Simon</location>
(
<geoCoordinate id="D7F15037FF934C78FC54DAD7041103F1" box="[954,1079,425,449]" degrees="64" direction="west" minutes="47" orientation="longitude" pageId="11" pageNumber="40" precision="15" seconds="52" value="-64.797775">64°47'52&quot;W</geoCoordinate>
,
<geoCoordinate id="D7F15037FF934C78FBACDAD7048103F1" box="[1090,1191,425,449]" degrees="17" direction="south" minutes="6" orientation="latitude" pageId="11" pageNumber="40" precision="15" seconds="33" value="-17.109167">17°6'33&quot;S</geoCoordinate>
),
<quantity id="753D9B15FF934C78FB54DAD704DB03F1" box="[1210,1277,425,449]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="40" unit="m" value="300.0">
<elevation id="39E8D1C3FF934C78FB54DAD704DB03F1" box="[1210,1277,425,449]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="40" unit="m" value="300.0">300 m</elevation>
</quantity>
elev.,
<collectingMethod id="6B844EE7FF934C78FAD0DAD7038C03D1" pageId="11" pageNumber="40">flight intercept trap</collectingMethod>
, 9.-
<date id="C67B1030FF934C78FC26DAB7041003D1" box="[968,1078,457,481]" pageId="11" pageNumber="40" value="1999-02-13">
<collectingDate id="D63FE9D8FF934C78FC26DAB7041003D1" box="[968,1078,457,481]" pageId="11" pageNumber="40" value="1999-02-13">13.2.1999</collectingDate>
</date>
, leg.
<collectorName id="1F305326FF934C78FB86DAB404EF03D1" box="[1128,1225,457,481]" pageId="11" pageNumber="40">F. Genier</collectorName>
, #BOL1G99 069 (
<collectionCode id="D4D4AE35FF934C78FCCFDA97035F0031" box="[801,889,489,513]" country="Kuwait" httpUri="http://grbio.org/cool/k9cv-41mj" name="The Educational Science Museum [=Kuwait Natural History Museum?]" pageId="11" pageNumber="40">KNHM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="02AD3CADFF934C78FC7DDA94044B0011" collectingDate="1999-02-13" collectionCode="KNHM" collectorName="F. Genier" country="Bolivia" elevation="300" latitude="-17.109167" location="Univ. de San Simon" longLatPrecision="21" longitude="-64.797775" pageId="11" pageNumber="40" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Cochabamba" typeStatus="paratype">
<typeStatus id="6D7E8852FF934C78FC7DDA9403DB0032" box="[915,1021,490,514]" pageId="11" pageNumber="40" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="CAD27660FF934C78FBE1DA97047B0031" box="[1039,1117,489,513]" name="Bolivia" pageId="11" pageNumber="40">Bolivia</collectingCountry>
:
<specimenCount id="A4C3FD79FF934C78FB81DA9704E40031" box="[1135,1218,489,513]" pageId="11" pageNumber="40" type="male">3 males</specimenCount>
,
<specimenCount id="A4C3FD79FF934C78FB3ADA9705110031" box="[1236,1335,489,513]" pageId="11" pageNumber="40" type="female">3 females</specimenCount>
with same data as holotype (
<collectionCode id="D4D4AE35FF934C78FBE1D97704400011" box="[1039,1126,521,545]" country="Kuwait" httpUri="http://grbio.org/cool/k9cv-41mj" name="The Educational Science Museum [=Kuwait Natural History Museum?]" pageId="11" pageNumber="40">KNHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="02AD3CADFF934C78FB96D97405550051" collectingDate="1999-02-13" collectionCode="KNHM, UIC" collectorName="R. Hanley" country="Bolivia" elevation="300" latitude="-17.109167" location="Univ. de San Simon" longLatPrecision="21" longitude="-64.797775" pageId="11" pageNumber="40" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Cochabamba" typeStatus="holotype">
same location, date, and collecting method as
<typeStatus id="6D7E8852FF934C78FBFED95704480071" box="[1040,1134,553,577]" pageId="11" pageNumber="40" type="holotype">holotype</typeStatus>
but
<specimenCount id="A4C3FD79FF934C78FB44D95704DB0071" box="[1194,1277,553,577]" pageId="11" pageNumber="40" type="male">2 males</specimenCount>
,
<specimenCount id="A4C3FD79FF934C78FAE3D95705550071" box="[1293,1395,553,577]" pageId="11" pageNumber="40" type="female">2 females</specimenCount>
, leg.
<collectorName id="1F305326FF934C78FCBDD934039B0051" box="[851,957,585,609]" pageId="11" pageNumber="40">R. Hanley</collectorName>
, #BOL1H99 078 (3
<collectionCode id="D4D4AE35FF934C78FB54D93405280051" box="[1210,1294,586,609]" country="Kuwait" httpUri="http://grbio.org/cool/k9cv-41mj" name="The Educational Science Museum [=Kuwait Natural History Museum?]" pageId="11" pageNumber="40">KNHM</collectionCode>
, 1
<collectionCode id="D4D4AE35FF934C78FAD7D937054D0051" box="[1337,1387,585,609]" pageId="11" pageNumber="40">UIC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="02AD3CADFF934C78FCF4D914046D00D1" collectingDate="2000-10-21" collectionCode="KNHM" collectorName="R. Brooks" country="Peru" latitude="-12.003612" location="Cocha Salvador" longLatPrecision="21" longitude="-71.52667" municipality="Reserved Zone Manu National Park" pageId="11" pageNumber="40" specimenCount="4" specimenCount-male="4" stateProvince="Madre de Dios" typeStatus="holotype">
Peru:
<collectingRegion id="7001F812FF934C78FCB3D917042100B1" box="[861,1031,617,641]" country="Peru" name="Madre de Dios" pageId="11" pageNumber="40">Madre de Dios</collectingRegion>
,
<location id="B71A602BFF934C78FBF8D91704E500B1" LSID="urn:lsid:plazi:treatment:3A6C87E6FF934C7FFCD1DB8D01860711:B71A602BFF934C78FBF8D91704E500B1" box="[1046,1219,617,641]" country="Peru" latitude="-12.003612" longLatPrecision="21" longitude="-71.52667" municipality="Reserved Zone Manu National Park" name="Cocha Salvador" pageId="11" pageNumber="40" stateProvince="Madre de Dios">Cocha Salvador</location>
,
<collectingMunicipality id="521EAC8AFF934C78FB3CD917042D0091" pageId="11" pageNumber="40">Reserved Zone Manu National Park</collectingMunicipality>
(
<geoCoordinate id="D7F15037FF934C78FBCBD9F704840091" box="[1061,1186,649,673]" degrees="71" direction="west" minutes="31" orientation="longitude" pageId="11" pageNumber="40" precision="15" seconds="36" value="-71.52667">71°31'36&quot;W</geoCoordinate>
,
<geoCoordinate id="D7F15037FF934C78FB57D9F705390091" box="[1209,1311,649,673]" degrees="12" direction="south" minutes="0" orientation="latitude" pageId="11" pageNumber="40" precision="15" seconds="13" value="-12.003612">12°0'13&quot;S</geoCoordinate>
),
<collectingMethod id="6B844EE7FF934C78FAD0D9F7039400F1" pageId="11" pageNumber="40">flight intercept trap</collectingMethod>
,
<specimenCount id="A4C3FD79FF934C78FC2DD9D7043000F1" box="[963,1046,681,705]" pageId="11" pageNumber="40" type="male">4 males</specimenCount>
, 20.-
<date id="C67B1030FF934C78FBA5D9D704E300F1" box="[1099,1221,681,705]" pageId="11" pageNumber="40" value="2000-10-21">
<collectingDate id="D63FE9D8FF934C78FBA5D9D704E300F1" box="[1099,1221,681,705]" pageId="11" pageNumber="40" value="2000-10-21">21.10.2000</collectingDate>
</date>
, leg.
<collectorName id="1F305326FF934C78FAE6D9D4055400F1" box="[1288,1394,681,705]" pageId="11" pageNumber="40">R. Brooks</collectorName>
, #
<collectingCountry id="CAD27660FF934C78FCC9D9B7034A00D1" box="[807,876,713,737]" name="Peru" pageId="11" pageNumber="40">PERU</collectingCountry>
1B00 070 (
<collectionCode id="D4D4AE35FF934C78FC03D9B7046200D1" box="[1005,1092,713,737]" country="Kuwait" httpUri="http://grbio.org/cool/k9cv-41mj" name="The Educational Science Museum [=Kuwait Natural History Museum?]" pageId="11" pageNumber="40">KNHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="02AD3CADFF934C78FB8FD9B703A60151" collectingDate="2000-10-26" collectionCode="KNHM" collectorName="R. Brooks" country="Peru" county="River" elevation="400" latitude="-12.656111" location="Pentiacolla Lodge" longLatPrecision="21" longitude="-71.23194" municipality="Alto" pageId="11" pageNumber="40" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Madre de Dios" typeStatus="holotype">
<collectingRegion id="7001F812FF934C78FB8FD9B7053300D1" box="[1121,1301,713,737]" country="Peru" name="Madre de Dios" pageId="11" pageNumber="40">Madre de Dios</collectingRegion>
,
<location id="B71A602BFF934C78FAC4D9B703B80131" LSID="urn:lsid:plazi:treatment:3A6C87E6FF934C7FFCD1DB8D01860711:B71A602BFF934C78FAC4D9B703B80131" country="Peru" county="River" latitude="-12.656111" longLatPrecision="21" longitude="-71.23194" municipality="Alto" name="Pentiacolla Lodge" pageId="11" pageNumber="40" stateProvince="Madre de Dios">Pentiacolla Lodge</location>
,
<collectingMunicipality id="521EAC8AFF934C78FC5ED99703F90131" box="[944,991,745,769]" pageId="11" pageNumber="40">Alto</collectingMunicipality>
<collectingRegion id="7001F812FF934C78FC02D99704BF0131" box="[1004,1177,745,769]" country="Peru" name="Madre de Dios" pageId="11" pageNumber="40">Madre de Dios</collectingRegion>
<collectingCounty id="5B1B4E7CFF934C78FB49D99704C60131" box="[1191,1248,745,769]" pageId="11" pageNumber="40">River</collectingCounty>
(
<geoCoordinate id="D7F15037FF934C78FB18D99705550131" box="[1270,1395,745,769]" degrees="71" direction="west" minutes="13" orientation="longitude" pageId="11" pageNumber="40" precision="15" seconds="55" value="-71.23194">71°13'55&quot;W</geoCoordinate>
,
<geoCoordinate id="D7F15037FF934C78FCF4D87703AB0111" box="[794,909,777,801]" degrees="12" direction="south" minutes="39" orientation="latitude" pageId="11" pageNumber="40" precision="15" seconds="22" value="-12.656111">12°39'22&quot;S</geoCoordinate>
),
<quantity id="753D9B15FF934C78FC48D87703CF0111" box="[934,1001,777,801]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="11" pageNumber="40" unit="m" value="400.0">
<elevation id="39E8D1C3FF934C78FC48D87703CF0111" box="[934,1001,777,801]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="11" pageNumber="40" unit="m" value="400.0">400 m</elevation>
</quantity>
elev.,
<collectingMethod id="6B844EE7FF934C78FBD7D877053F0111" box="[1081,1305,777,801]" pageId="11" pageNumber="40">flight intercept trap</collectingMethod>
,
<specimenCount id="A4C3FD79FF934C78FAC4D87705550111" box="[1322,1395,777,801]" pageId="11" pageNumber="40" type="male">1 male</specimenCount>
,
<specimenCount id="A4C3FD79FF934C78FCF4D85703500171" box="[794,886,809,833]" pageId="11" pageNumber="40" type="female">1 female</specimenCount>
, 23.-
<date id="C67B1030FF934C78FC4CD857043B0171" box="[930,1053,809,833]" pageId="11" pageNumber="40" value="2000-10-26">
<collectingDate id="D63FE9D8FF934C78FC4CD857043B0171" box="[930,1053,809,833]" pageId="11" pageNumber="40" value="2000-10-26">26.10.2000</collectingDate>
</date>
, leg.
<collectorName id="1F305326FF934C78FBBCD854049A0171" box="[1106,1212,809,833]" pageId="11" pageNumber="40">R. Brooks</collectorName>
, #
<collectingCountry id="CAD27660FF934C78FB3DD857053E0171" box="[1235,1304,809,833]" name="Peru" pageId="11" pageNumber="40">PERU</collectingCountry>
1B00 099 (
<collectionCode id="D4D4AE35FF934C78FCCFD837035E0151" box="[801,888,841,865]" country="Kuwait" httpUri="http://grbio.org/cool/k9cv-41mj" name="The Educational Science Museum [=Kuwait Natural History Museum?]" pageId="11" pageNumber="40">KNHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="02AD3CADFF934C78FC65D834053A01D1" collectingDate="2011-06-13" collectionCode="KNHM, UIC" collectorName="Chaboo" country="Peru" elevation="295" latitude="-12.5523" longLatPrecision="7" longitude="-70.1098" pageId="11" pageNumber="40" specimenCount="28" specimenCount-female="23" specimenCount-male="5" stateProvince="Madre de Dios" typeStatus="holotype">
same region, CICRA field station, ~
<quantity id="753D9B15FF934C78FAE2D83705650151" box="[1292,1347,841,865]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="40" unit="km" value="2.0">2 km</quantity>
NW of cafeteria, research plot (
<geoCoordinate id="D7F15037FF934C78FBDAD817048D01B1" box="[1076,1195,873,897]" degrees="70.1098" direction="west" orientation="longitude" pageId="11" pageNumber="40" precision="5" value="-70.1098">70.1098°W</geoCoordinate>
,
<geoCoordinate id="D7F15037FF934C78FB5BD817050401B1" box="[1205,1314,873,897]" degrees="12.5523" direction="south" orientation="latitude" pageId="11" pageNumber="40" precision="5" value="-12.5523">12.5523°S</geoCoordinate>
),
<quantity id="753D9B15FF934C78FADAD817055101B1" box="[1332,1399,873,897]" metricMagnitude="2" metricUnit="m" metricValue="2.95" pageId="11" pageNumber="40" unit="m" value="295.0">
<elevation id="39E8D1C3FF934C78FADAD817055101B1" box="[1332,1399,873,897]" metricMagnitude="2" metricUnit="m" metricValue="2.95" pageId="11" pageNumber="40" unit="m" value="295.0">295 m</elevation>
</quantity>
elev.,
<collectingMethod id="6B844EE7FF934C78FCBDD8F704070191" box="[851,1057,905,929]" pageId="11" pageNumber="40">flight intercept trap</collectingMethod>
,
<collectingMethod id="6B844EE7FF934C78FBC7D8F704B80191" box="[1065,1182,905,929]" pageId="11" pageNumber="40">pit fall trap</collectingMethod>
,
<specimenCount id="A4C3FD79FF934C78FB48D8F704DE0191" box="[1190,1272,905,929]" pageId="11" pageNumber="40" type="male">5 males</specimenCount>
,
<specimenCount id="A4C3FD79FF934C78FAEED8F705550191" box="[1280,1395,905,929]" pageId="11" pageNumber="40" type="female">23 females</specimenCount>
, 7.-9.6., 9.-11.6., 11.-13.6., 12.-
<date id="C67B1030FF934C78FB82D8D704FC01F1" box="[1132,1242,937,961]" pageId="11" pageNumber="40" value="2011-06-13">
<collectingDate id="D63FE9D8FF934C78FB82D8D704FC01F1" box="[1132,1242,937,961]" pageId="11" pageNumber="40" value="2011-06-13">13.6.2011</collectingDate>
</date>
, leg.
<collectorName id="1F305326FF934C78FACCD8D7055101F1" box="[1314,1399,937,961]" pageId="11" pageNumber="40">Chaboo</collectorName>
team, #PER-11-FIT-019 (25
<collectionCode id="D4D4AE35FF934C78FB8DD8B4049101D1" box="[1123,1207,970,993]" country="Kuwait" httpUri="http://grbio.org/cool/k9cv-41mj" name="The Educational Science Museum [=Kuwait Natural History Museum?]" pageId="11" pageNumber="40">KNHM</collectionCode>
, 3
<collectionCode id="D4D4AE35FF934C78FB0CD8B7053201D1" box="[1250,1300,969,993]" pageId="11" pageNumber="40">UIC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="02AD3CADFF934C78FADAD8B705540651" collectingDate="2001-05-24" collectionCode="KNHM" collectorName="D. J. Bennett &amp; E Razuri" country="Peru" elevation="555" latitude="-12.8925" location="Dept." longLatPrecision="7" longitude="-71.4191" municipality="Villa Carmen" pageId="11" pageNumber="40" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Cusco" typeStatus="holotype">
<collectingRegion id="7001F812FF934C78FADAD8B7055001D1" box="[1332,1398,969,993]" country="Peru" name="Cusco" pageId="11" pageNumber="40">Cusco</collectingRegion>
<location id="B71A602BFF934C78FCF4D89403710631" LSID="urn:lsid:plazi:treatment:3A6C87E6FF934C7FFCD1DB8D01860711:B71A602BFF934C78FCF4D89403710631" box="[794,855,1002,1025]" country="Peru" latitude="-12.8925" longLatPrecision="7" longitude="-71.4191" municipality="Villa Carmen" name="Dept." pageId="11" pageNumber="40" stateProvince="Cusco">Dept.</location>
,
<collectingMunicipality id="521EAC8AFF934C78FC8CD89703D40631" box="[866,1010,1001,1025]" pageId="11" pageNumber="40">Villa Carmen</collectingMunicipality>
field station, ~
<quantity id="753D9B15FF934C78FB7AD89704F80631" box="[1172,1246,1001,1025]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="11" pageNumber="40" unit="km" value="1.7">1.7 km</quantity>
west cafeteria research transect (
<geoCoordinate id="D7F15037FF934C78FC06DF7704780611" box="[1000,1118,1033,1057]" degrees="71.4191" direction="west" orientation="longitude" pageId="11" pageNumber="40" precision="5" value="-71.4191">71.4191°W</geoCoordinate>
,
<geoCoordinate id="D7F15037FF934C78FB81DF7704FA0611" box="[1135,1244,1033,1057]" degrees="12.8925" direction="south" orientation="latitude" pageId="11" pageNumber="40" precision="5" value="-12.8925">12.8925°S</geoCoordinate>
),
<quantity id="753D9B15FF934C78FB1DDF7705100611" box="[1267,1334,1033,1057]" metricMagnitude="2" metricUnit="m" metricValue="5.55" pageId="11" pageNumber="40" unit="m" value="555.0">
<elevation id="39E8D1C3FF934C78FB1DDF7705100611" box="[1267,1334,1033,1057]" metricMagnitude="2" metricUnit="m" metricValue="5.55" pageId="11" pageNumber="40" unit="m" value="555.0">555 m</elevation>
</quantity>
elev.,
<collectingMethod id="6B844EE7FF934C78FCF4DF5703CC0671" box="[794,1002,1065,1089]" pageId="11" pageNumber="40">flight intercept trap</collectingMethod>
,
<specimenCount id="A4C3FD79FF934C78FC1ADF57041B0671" box="[1012,1085,1065,1089]" pageId="11" pageNumber="40" type="male">1 male</specimenCount>
,
<specimenCount id="A4C3FD79FF934C78FBA9DF57048B0671" box="[1095,1197,1065,1089]" pageId="11" pageNumber="40" type="female">3 females</specimenCount>
, 22.-
<date id="C67B1030FF934C78FB35DF57056E0671" box="[1243,1352,1065,1089]" pageId="11" pageNumber="40" value="2001-05-24">
<collectingDate id="D63FE9D8FF934C78FB35DF57056E0671" box="[1243,1352,1065,1089]" pageId="11" pageNumber="40" value="2001-05-24">24.5.2001</collectingDate>
</date>
, leg.
<collectorName id="1F305326FF934C78FCF4DF3403BC0651" box="[794,922,1098,1121]" pageId="11" pageNumber="40">D.J. Bennett</collectorName>
&amp;
<collectorName id="1F305326FF934C78FC23DF34040D0651" box="[973,1067,1097,1121]" pageId="11" pageNumber="40">E Razuri</collectorName>
, #PER-11-FIT-009 (
<collectionCode id="D4D4AE35FF934C78FAFDDF37054C0651" box="[1299,1386,1097,1121]" country="Kuwait" httpUri="http://grbio.org/cool/k9cv-41mj" name="The Educational Science Museum [=Kuwait Natural History Museum?]" pageId="11" pageNumber="40">KNHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="02AD3CADFF934C78FCF4DF17056B06F1" collectingDate="1997-06-03" collectionCode="KNHM" collectorName="J. Ashe &amp; R. Brooks" country="French Guiana" elevation="220" latitude="3.662778" location="Les Eaux Claires" longLatPrecision="21" longitude="-53.221947" pageId="11" pageNumber="40" specimenCount="1" typeStatus="holotype">
<collectingCountry id="CAD27660FF934C78FCF4DF1703F606B1" box="[794,976,1129,1153]" name="French Guiana" pageId="11" pageNumber="40">French Guiana</collectingCountry>
: SaüL.
<quantity id="753D9B15FF934C78FBC0DF14044506B1" box="[1070,1123,1129,1153]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="11" pageNumber="40" unit="km" value="7.0">7 km</quantity>
N,
<location id="B71A602BFF934C78FB70DF14055106B1" LSID="urn:lsid:plazi:treatment:3A6C87E6FF934C7FFCD1DB8D01860711:B71A602BFF934C78FB70DF14055106B1" box="[1182,1399,1129,1153]" country="French Guiana" latitude="3.662778" longLatPrecision="21" longitude="-53.221947" name="Les Eaux Claires" pageId="11" pageNumber="40">Les Eaux Claires</location>
(
<geoCoordinate id="D7F15037FF934C78FCCFDFF703B80691" box="[801,926,1161,1185]" degrees="53" direction="west" minutes="13" orientation="longitude" pageId="11" pageNumber="40" precision="15" seconds="19" value="-53.221947">53°13'19&quot;W</geoCoordinate>
,
<geoCoordinate id="D7F15037FF934C78FC49DFF704320691" box="[935,1044,1161,1185]" degrees="3" direction="north" minutes="39" orientation="latitude" pageId="11" pageNumber="40" precision="15" seconds="46" value="3.662778">3°39'46&quot;N</geoCoordinate>
),
<quantity id="753D9B15FF934C78FBCBDFF704410691" box="[1061,1127,1161,1185]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="11" pageNumber="40" unit="m" value="220.0">
<elevation id="39E8D1C3FF934C78FBCBDFF704410691" box="[1061,1127,1161,1185]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="11" pageNumber="40" unit="m" value="220.0">220 m</elevation>
</quantity>
elev.,
<collectingMethod id="6B844EE7FF934C78FB4BDFF705550691" box="[1189,1395,1161,1185]" pageId="11" pageNumber="40">flight intercept trap</collectingMethod>
, male, 31.5.-
<date id="C67B1030FF934C78FC7FDFD703D606F1" box="[913,1008,1193,1217]" pageId="11" pageNumber="40" value="1997-06-03">
<collectingDate id="D63FE9D8FF934C78FC7FDFD703D606F1" box="[913,1008,1193,1217]" pageId="11" pageNumber="40" value="1997-06-03">3.6.1997</collectingDate>
</date>
, leg.
<collectorName id="1F305326FF934C78FBCADFD4044906F1" box="[1060,1135,1193,1217]" pageId="11" pageNumber="40">J. Ashe</collectorName>
,
<collectorName id="1F305326FF934C78FB97DFD404C706F1" box="[1145,1249,1193,1217]" pageId="11" pageNumber="40">R. Brooks</collectorName>
(
<collectionCode id="D4D4AE35FF934C78FB01DFD7056006F1" box="[1263,1350,1193,1217]" country="Kuwait" httpUri="http://grbio.org/cool/k9cv-41mj" name="The Educational Science Museum [=Kuwait Natural History Museum?]" pageId="11" pageNumber="40">KNHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="02AD3CADFF934C78FAB9DFD403D60711" collectingDate="1997-06-07" collectionCode="KNHM" collectorName="J. Ashe &amp; R. Brooks" country="French Guiana" elevation="740" latitude="3.621667" location="Mt Galbao" longLatPrecision="21" longitude="-53.27833" pageId="11" pageNumber="40" specimenCount="2" specimenCount-female="2" typeStatus="holotype">
<location id="B71A602BFF934C78FAB9DFD4034106D1" LSID="urn:lsid:plazi:treatment:3A6C87E6FF934C7FFCD1DB8D01860711:B71A602BFF934C78FAB9DFD4034106D1" country="French Guiana" latitude="3.621667" longLatPrecision="21" longitude="-53.27833" name="Mt Galbao" pageId="11" pageNumber="40">Mt Galbao</location>
summit (
<geoCoordinate id="D7F15037FF934C78FC32DFB7047F06D1" box="[988,1113,1225,1249]" degrees="53" direction="west" minutes="16" orientation="longitude" pageId="11" pageNumber="40" precision="15" seconds="42" value="-53.27833">53°16'42&quot;W</geoCoordinate>
,
<geoCoordinate id="D7F15037FF934C78FB85DFB704FE06D1" box="[1131,1240,1225,1249]" degrees="3" direction="north" minutes="37" orientation="latitude" pageId="11" pageNumber="40" precision="15" seconds="18" value="3.621667">3°37'18&quot;N</geoCoordinate>
),
<quantity id="753D9B15FF934C78FB1CDFB7051306D1" box="[1266,1333,1225,1249]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="11" pageNumber="40" unit="m" value="740.0">
<elevation id="39E8D1C3FF934C78FB1CDFB7051306D1" box="[1266,1333,1225,1249]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="11" pageNumber="40" unit="m" value="740.0">740 m</elevation>
</quantity>
elev.,
<collectingMethod id="6B844EE7FF934C78FCF4DF9703D40731" box="[794,1010,1257,1281]" pageId="11" pageNumber="40">flight intercept trap</collectingMethod>
,
<specimenCount id="A4C3FD79FF934C78FBEEDF9704400731" box="[1024,1126,1257,1281]" pageId="11" pageNumber="40" type="female">2 females</specimenCount>
, 5.-
<date id="C67B1030FF934C78FB67DF9704CD0731" box="[1161,1259,1257,1281]" pageId="11" pageNumber="40" value="1997-06-07">
<collectingDate id="D63FE9D8FF934C78FB67DF9704CD0731" box="[1161,1259,1257,1281]" pageId="11" pageNumber="40" value="1997-06-07">7.6.1997</collectingDate>
</date>
, leg.
<collectorName id="1F305326FF934C78FAC6DF9405540731" box="[1320,1394,1257,1281]" pageId="11" pageNumber="40">J. Ashe</collectorName>
,
<collectorName id="1F305326FF934C78FCF4DE7403A40711" box="[794,898,1289,1313]" pageId="11" pageNumber="40">R. Brooks</collectorName>
(
<collectionCode id="D4D4AE35FF934C78FC7EDE7703CE0711" box="[912,1000,1289,1313]" country="Kuwait" httpUri="http://grbio.org/cool/k9cv-41mj" name="The Educational Science Museum [=Kuwait Natural History Museum?]" pageId="11" pageNumber="40">KNHM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="FADF657BFF934C78FCF4DE3703FB0452" pageId="11" pageNumber="40" type="diagnosis">
<paragraph id="B27A36F0FF934C78FCF4DE3703FB0452" blockId="11.[794,1399,1353,1634]" pageId="11" pageNumber="40">
<emphasis id="80B1EAE2FF934C78FCF4DE3703A10751" bold="true" box="[794,903,1353,1377]" pageId="11" pageNumber="40">Diagnosis</emphasis>
: The species is characterised by the apical row of large teeth of the endophallus. In addition, males can be identified by the specific shape of sternite VII with large lateral prominences and a wide medial prominence. Females are much more difficult to separate from similar species such as
<taxonomicName id="75C54D73FF934C78FC0CDE9404420431" authorityName="Irmler" authorityYear="2021" box="[994,1124,1514,1537]" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="40" phylum="Arthropoda" rank="species" species="nigerrima">
<emphasis id="80B1EAE2FF934C78FC0CDE9404420431" box="[994,1124,1514,1537]" italics="true" pageId="11" pageNumber="40">L. nigerrima</emphasis>
</taxonomicName>
. They are separated from similarly large species by the black colouration in combination with the sparse punctation of the posterior vertex, and the large eyes.
</paragraph>
</subSubSection>
<subSubSection id="FADF657BFF934C7FFCF4DDF701550691" lastPageId="12" lastPageNumber="41" pageId="11" pageNumber="40" type="description">
<paragraph id="B27A36F0FF934C78FCF4DDF703DD04F1" blockId="11.[794,1399,1673,2081]" pageId="11" pageNumber="40">
<emphasis id="80B1EAE2FF934C78FCF4DDF703BA0491" bold="true" box="[794,924,1673,1697]" pageId="11" pageNumber="40">Description</emphasis>
: Length:
<quantity id="753D9B15FF934C78FC10DDF704740491" box="[1022,1106,1673,1697]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="11" pageNumber="40" unit="mm" value="4.4">4.4 mm</quantity>
. Colouration: Blackish; legs and antennae brown.
</paragraph>
<paragraph id="B27A36F0FF934C7FFCF4DDB701550691" blockId="11.[794,1399,1673,2081]" lastBlockId="12.[189,795,233,1185]" lastPageId="12" lastPageNumber="41" pageId="11" pageNumber="40">
Head:
<quantity id="753D9B15FF934C78FC8CDDB703E704D1" box="[866,961,1737,1761]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="11" pageNumber="40" unit="mm" value="0.53">0.53 mm</quantity>
long,
<quantity id="753D9B15FF934C78FBE9DDB7044004D1" box="[1031,1126,1737,1761]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" pageId="11" pageNumber="40" unit="mm" value="0.52">0.52 mm</quantity>
wide; eyes large; slightly prominent; postocular sides slightly divergent; 1.8 times as long as eyes; interantennal furrows reaching anterior third of eyes; setiferous punctation moderately deep and dense; on average, interstices as wide as diameter of punctures; on anterior vertex interstices slightly shorter, on posterior vertex slightly wider; surface without microsculpture; shiny. Antennae with first antennomere slightly longer than half length of head; second and third antennomere conical; longer than wide; subsequent antennomeres wider than long and increasing in width; fourth antennomere 1.5 times as wide as long; tenth antennomere 2.2 times as wide as long; all antennomeres densely pubescent. Pronotum:
<quantity id="753D9B15FF944C7FFE95DA5701FC0371" box="[379,474,297,321]" metricMagnitude="-4" metricUnit="m" metricValue="6.7" pageId="12" pageNumber="41" unit="mm" value="0.67">0.67 mm</quantity>
long,
<quantity id="753D9B15FF944C7FFDC8DA5702A30371" box="[550,645,297,321]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" pageId="12" pageNumber="41" unit="mm" value="0.44">0.44 mm</quantity>
wide; widest in anterior third; anteriad, conically narrowed toward neck; posteriad, slightly narrowed in middle third and nearly parallel in posterior third; posterior angles widely rounded; posterior margin convex; setiferous punctation as dense and deep as on posterior vertex; wide midline impunctate; surface without microsculpture; shiny. Elytra:
<quantity id="753D9B15FF944C7FFEFED97701490011" box="[272,367,521,545]" metricMagnitude="-4" metricUnit="m" metricValue="6.2" pageId="12" pageNumber="41" unit="mm" value="0.62">0.62 mm</quantity>
long,
<quantity id="753D9B15FF944C7FFE51D977023B0011" box="[447,541,521,545]" metricMagnitude="-4" metricUnit="m" metricValue="5.6" pageId="12" pageNumber="41" unit="mm" value="0.56">0.56 mm</quantity>
wide; humeral angles obtuse; sides posteriad nearly parallel; posterior angles sub-rectangular; posterior margin slightly convex with wide incision at suture; setiferous punctation finer than on pronotum but slightly denser; surface with extremely fine isodiametric microsculpture; shiny. Abdomen with denser and finer setiferous punctation than on fore-body; microsculpture denser and deeper than on elytra; surface less shiny; sternite VII of male with long curved lateral teeth; posterior margin deeply emarginate on each side of middle; central prominence convex; tergite VII of male with straight posterior margin. Meso- and metatibia with three subapical ctenidia. Aedeagus long-oval with sub-rectangular anterior angles; posterior part of endophallus covered by minute teeth; in anterior part with row of conspicuously large teeth; paramere short and broad; bilobed; outer lobe broad; triangular and half as long as inner lobe; inner lobe with straight inner face and slightly curved outer face; sparsely covered by fine setae; setae on inner face denser; basal setae slightly thicker and longer than apical setae.
</paragraph>
</subSubSection>
<subSubSection id="FADF657BFF944C7FFF53DFB701860711" pageId="12" pageNumber="41" type="etymology">
<paragraph id="B27A36F0FF944C7FFF53DFB701860711" blockId="12.[189,793,1225,1313]" pageId="12" pageNumber="41">
<emphasis id="80B1EAE2FF944C7FFF53DFB7011206D1" bold="true" box="[189,308,1225,1249]" pageId="12" pageNumber="41">Etymology</emphasis>
: The species name is derived from the same Latin word (meaning toothed) and refers to the shape of the male sternite VII.
</paragraph>
</subSubSection>
</treatment>
</document>
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