556 lines
71 KiB
XML
556 lines
71 KiB
XML
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<mods:title id="67371CE09E52D39895E26F9936B0C7FB">Contributions of a small collection of terrestrial microsnails (Pupilloidea, Hypselostomatidae) from Myanmar with description of three new species</mods:title>
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<mods:affiliation id="86A5F194E47430E6694A084325184280">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand & Department of Biology, Faculty of Science, Khon Kaen University, Khon Kaen 40002, Thailand</mods:affiliation>
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<taxonomicName id="0A16F6821749528249192AC06A49AC46" LSID="35DDA19D-812C-590B-8914-D6A9EB13387B" authority="(Theobald, 1871)" baseAuthorityName="Theobald" baseAuthorityYear="1871" class="Gastropoda" family="Hypselostomatidae" genus="Bensonella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bensonella salwiniana" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="species" species="salwiniana">Bensonella salwiniana (Theobald, 1871)</taxonomicName>
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<figureCitation id="1C4513EBDDC831D2AAD4725193EB4AFA" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Bensonella salwiniana A specimen NHMUK 1912.4.16.66 from Burma B, C specimen CUMZ 14377 from Pyigyidagun Township, Mandalay Region D specimen CUMZ 14376 from Taunggyi District, Shan State E specimen CUMZ 14375 from Taunggyi District, Shan State F specimen CUMZ 14393 from Taunggyi District, Shan State. Abbreviations: a: angular lamella, b: basal plica, c: columellar lamella, ip: infraparietal lamella, lpl: lower palatal plica, p: parietal lamella, upl: upper palatal plica." figureDoi="10.3897/zookeys.1195.112112.figure4" httpUri="https://binary.pensoft.net/fig/1002053" pageId="0" pageNumber="157">Figs 4</figureCitation>
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<figureCitation id="55B59DE5D32ACB97A56738EE207518D9" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Bensonella salwiniana, specimen CUMZ 14397 from Taunggyi District, Shan State A genitalia and a small inset of penial appendix B preserved specimen with mantle edge, body and foot C snail showing colour in life D radula morphology: blue highlighting indicates central teeth and numbers indicate tooth order from lateral to marginal end. Abbreviations: at: atrium, e: epiphallus, fo: free oviduct, gd: gametolytic duct, gs: gametolytic sac, mc: mantle collar; p: penis, pa: penial appendix, pn: pneumostome; pr: penial retractor muscle, v: vagina, vd: vas deferens." figureDoi="10.3897/zookeys.1195.112112.figure5" httpUri="https://binary.pensoft.net/fig/1002054" pageId="0" pageNumber="157">, 5</figureCitation>
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<figureCitation id="E0CF7D67F5FBB50A63FAC49FF3FE9C40" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A Bensonella salwiniana, specimen CUMZ 14393 from Taunggyi District, Shan State B Bensonella anguloobtusa, specimen CUMZ 14401 from Taunggyi District, Shan State. Abbreviations: a: angular lamella, c: columellar lamella, itpl: interpalatal plica, lpl: lower palatal plica, p: parietal lamella, pt: palatal tubercle, sbc: subcolumellar lamella, upl: upper palatal plica." figureDoi="10.3897/zookeys.1195.112112.figure6" httpUri="https://binary.pensoft.net/fig/1002055" pageId="0" pageNumber="157">, 6A</figureCitation>
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<figureCitation id="4C28A9F58D7E945C87FA593AD7D78ACC" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Synoptic view of representatives of hypselostomatid snails recorded from Myanmar A Acinolaemus dayanum, specimen HA Collection B Acinolaemus cryptidentatus, specimen CUMZ 14413 C Anauchen eotvosi, holotype NHMUK 1903.7.1.1227.1 (after Pall-Gergely 2023 a: fig. 1 c) D Angustopila occidentalis, holotype HNHM 103483 (after Pall-Gergely et al. 2023: fig. 70 a) E Angustopila elevata, specimen HNHM 103484 (after Pall-Gergely et al. 2023: fig. 13 a) F Bensonella salwiniana, specimen CUMZ 14393 G Bensonella anguloobtusa, specimen CUMZ 14400 H Bensonella taiyaiorum sp. nov., paratype CUMZ 14403 I Bensonella lophiodera sp. nov., paratype CUMZ 14405 J Clostophis sankeyi, holotype UMZC I. 103320 (after Preece et al. 2022: fig. 70 e) K Clostophis thinbowguensis, holotype MNHN-IM- 2000 - 38057 (after Pall-Gergely et al. 2022: fig. 9 c) L Gyliotrachela bensonianum, syntype NHMUK 20191141 (after Gojsina et al. 2022: fig. 4 a) M Gyliotrachela hungerfordiana, specimen NHMUK collection from Perak, Malaysia N Gyliotrachela tianxingqiaoensis, holotype IZCAS TM 025075 (after Gojsina et al. 2022: fig. 3 a) O Gyliotrachela muangon, specimen CUMZ 14410 P Gyliotrachela aunglini sp. nov., paratype CUMZ 14412 Q Hypselostoma tubiferum, specimen NHMUK 1888.12.4.17 - 22 ex. Theobald collection from Tonduong, Burma." figureDoi="10.3897/zookeys.1195.112112.figure13" httpUri="https://binary.pensoft.net/fig/1002062" pageId="0" pageNumber="157">, 13F</figureCitation>
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</paragraph>
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<taxonomicName id="2B2A54865266956FEF03B75B1FC6F51E" authorityName="Tongkerd & Lwin & Páll-Gergely & Chanabun & Pholyotha & Prasankok & Seesamut & Siriwut & Srisonchai & Sutcharit & Panha" authorityYear="2024" baseAuthorityName="Tongkerd & Lwin & Páll-Gergely & Chanabun & Pholyotha & Prasankok & Seesamut & Siriwut & Srisonchai & Sutcharit & Panha" baseAuthorityYear="2024" class="Gastropoda" family="Hypselostomatidae" genus="Hypselostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypselostoma" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="genus">Hypselostoma</taxonomicName>
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<emphasis id="E157A65A629F75621FC552AC743334E2" italics="true" pageId="0" pageNumber="157">Pupa salwiniana</emphasis>
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Theobald, 1871: 400. Type locality: Shan States.
|
||
<bibRefCitation id="E7C30CCFF43153BC112351315B947004" author="Hanley, SC" journalOrPublisher="Taylor and Francis Group, New York" pageId="0" pageNumber="157" publicationUrl="https://www.biodiversitylibrary.org/page/46204402" refId="B27" refString="Hanley, SC, Theobald, W, 1870 1874 1876. [1870-1876] Conchologia Indica, being Illustrations of the Land and Freshwater Shells of British India. L. Reeve & Co., London,65 pp. 160 pls. [pp. 1-18, pls 1-40 (1870); pp. 35-48, pls 81-120 (1874); pp. 35-48, pls 81-120 (1874); pp. 57-65, pls 141-160 (1876)] https://www.biodiversitylibrary.org/page/46204402" title="[1870 - 1876] Conchologia Indica, being Illustrations of the Land and Freshwater Shells of British India. L. Reeve & Co., London, 65 pp. 160 pls. [pp. 1 - 18, pls 1 - 40 (1870); pp. 35 - 48, pls 81 - 120 (1874); pp. 35 - 48, pls 81 - 120 (1874); pp. 57 - 65, pls 141 - 160 (1876)]" url="https://www.biodiversitylibrary.org/page/46204402" year="1870">Hanley and Theobald 1874</bibRefCitation>
|
||
: 40, pl. 100, fig. 9.
|
||
<bibRefCitation id="5FB5F3AE0FB8DEDD484056FE7271F338" author="Sowerby, GB II" journalOrPublisher="Malacologia" pageId="0" pageNumber="157" publicationUrl="https://www.biodiversitylibrary.org/page/8229973" refId="B80" refString="Sowerby, GB II, 1877. [1875-1877] Monograph of the genus Cylindrella. In: Conchologia Iconica, or Illustrations of the Shells of Molluscous Animals, volume 20, pls 1-16 and unpaginated text. L. Reeve, London. [pls. 2-8 (1875); pls 1, 9-16 (1877)]. https://www.biodiversitylibrary.org/page/8229973" title="[1875 - 1877] Monograph of the genus Cylindrella. In: Conchologia Iconica, or Illustrations of the Shells of Molluscous Animals, volume 20, pls 1 - 16 and unpaginated text. L. Reeve, London. [pls. 2 - 8 (1875); pls 1, 9 - 16 (1877)]." url="https://www.biodiversitylibrary.org/page/8229973" year="1877">Sowerby 1877</bibRefCitation>
|
||
:
|
||
<emphasis id="724F3DAB0355649E328C20F2315E8910" italics="true" pageId="0" pageNumber="157">Pupa</emphasis>
|
||
pl. 16, fig. 150.
|
||
<bibRefCitation id="B24D9DFD89B55BC11E1985AB7486382C" author="Pfeiffer, L" journalOrPublisher="Tome 1, Cassel: Th. Fischer" pageId="0" pageNumber="157" publicationUrl="https://www.biodiversitylibrary.org/page/11248776" refId="B69" refString="Pfeiffer, L, 1877. [1876-1877] Monographia Heliceorum viventium. Sistens descriptiones systematicas et criticas omnium huius familiae generum et specierum hodie cognitarum. Volumen 8. Lipsiae: F.A. Brockhaus, 729 pp. [pp. 1-480 (1876)] https://www.biodiversitylibrary.org/page/11248776" title="[1876 - 1877] Monographia Heliceorum viventium. Sistens descriptiones systematicas et criticas omnium huius familiae generum et specierum hodie cognitarum. Volumen 8. Lipsiae: F. A. Brockhaus, 729 pp. [pp. 1 - 480 (1876)]" url="https://www.biodiversitylibrary.org/page/11248776" year="1877">Pfeiffer 1877</bibRefCitation>
|
||
: 403.
|
||
</paragraph>
|
||
<paragraph id="F7B0A6DF1E7158F04DBC034FEBF40138" pageId="0" pageNumber="157">
|
||
<taxonomicName id="FCE6C3807F1859246922B3BAFDEE6FC5" authorityName="Tongkerd & Lwin & Páll-Gergely & Chanabun & Pholyotha & Prasankok & Seesamut & Siriwut & Srisonchai & Sutcharit & Panha" authorityYear="2024" baseAuthorityName="Tongkerd & Lwin & Páll-Gergely & Chanabun & Pholyotha & Prasankok & Seesamut & Siriwut & Srisonchai & Sutcharit & Panha" baseAuthorityYear="2024" class="Gastropoda" family="Hypselostomatidae" genus="Hypselostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypselostoma" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="genus">Hypselostoma</taxonomicName>
|
||
<emphasis id="300E655AF416A20748865A65D0B3FD47" italics="true" pageId="0" pageNumber="157">Pupa (Scopelophila) salwiniana</emphasis>
|
||
.
|
||
<bibRefCitation id="5CFA854DBCC74F97B1FE677A4D25AF06" author="Nevill, G" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="0" pageNumber="157" pagination="14 - 41" publicationUrl="https://www.biodiversitylibrary.org/page/35548100" refId="B43" refString="Nevill, G, 1877. List of the Mollusca brought back to Mr. Anderson from Yunnan and upper Burma, with descriptions of new species. Journal of the Asiatic Society of Bengal 46: 14 - 41, https://www.biodiversitylibrary.org/page/35548100" title="List of the Mollusca brought back to Mr. Anderson from Yunnan and upper Burma, with descriptions of new species." url="https://www.biodiversitylibrary.org/page/35548100" volume="46" year="1877">Nevill 1877</bibRefCitation>
|
||
: 23.
|
||
</paragraph>
|
||
<paragraph id="5F7FB68D8D3F3A775A8798A6B93E510D" pageId="0" pageNumber="157">
|
||
<taxonomicName id="F89244D615D35CA36935D3FBB11A9EFD" authorityName="Tongkerd & Lwin & Páll-Gergely & Chanabun & Pholyotha & Prasankok & Seesamut & Siriwut & Srisonchai & Sutcharit & Panha" authorityYear="2024" baseAuthorityName="Tongkerd & Lwin & Páll-Gergely & Chanabun & Pholyotha & Prasankok & Seesamut & Siriwut & Srisonchai & Sutcharit & Panha" baseAuthorityYear="2024" class="Gastropoda" family="Hypselostomatidae" genus="Hypselostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypselostoma" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="genus">Hypselostoma</taxonomicName>
|
||
<emphasis id="BF0DF0FDADEE23CE286501DEF8D141F5" italics="true" pageId="0" pageNumber="157">Pupa (Pupilla) salwiniana</emphasis>
|
||
.
|
||
<bibRefCitation id="49A62B69302594D3D36A3CEEDEBD2BF0" author="Pfeiffer, L" journalOrPublisher="Fischer, Cassel" pageId="0" pageNumber="157" refId="B70" refString="Pfeiffer, L, 1880. [1878-1881] Nomenclator heliceorum viventium. Fischer, Cassel" title="[1878 - 1881] Nomenclator heliceorum viventium." year="1880">Pfeiffer 1880</bibRefCitation>
|
||
: 355.
|
||
</paragraph>
|
||
<paragraph id="D5EA9EF10AACA875CA6FBEDC98F07DE8" pageId="0" pageNumber="157">
|
||
<taxonomicName id="3BC0E3ABB8FAC98B8020CDC5745252FD" authorityName="Tongkerd & Lwin & Páll-Gergely & Chanabun & Pholyotha & Prasankok & Seesamut & Siriwut & Srisonchai & Sutcharit & Panha" authorityYear="2024" baseAuthorityName="Tongkerd & Lwin & Páll-Gergely & Chanabun & Pholyotha & Prasankok & Seesamut & Siriwut & Srisonchai & Sutcharit & Panha" baseAuthorityYear="2024" class="Gastropoda" family="Hypselostomatidae" genus="Hypselostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypselostoma" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="genus">Hypselostoma</taxonomicName>
|
||
<emphasis id="FEFF3FBAF1A71C5DBCA576063770CE65" italics="true" pageId="0" pageNumber="157">Pupa salwinieana</emphasis>
|
||
[sic].
|
||
<bibRefCitation id="E2E2FF00890461A8834CDD8BD874B595" DOI="https://doi.org/10.1111/j.1469-7998.1888.tb06701.x" author="Godwin-Austen, HH" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="157" pagination="240 - 245" refId="B23" refString="Godwin-Austen, HH, 1888. On some land-mollusks from Burmah, with descriptions of some new species. Proceedings of the Zoological Society of London 56 (2): 240 - 245, DOI: https://doi.org/10.1111/j.1469-7998.1888.tb06701.x" title="On some land-mollusks from Burmah, with descriptions of some new species." url="https://doi.org/10.1111/j.1469-7998.1888.tb06701.x" volume="56" year="1888">Godwin-Austen 1888</bibRefCitation>
|
||
: 244.
|
||
</paragraph>
|
||
<paragraph id="A34334F1A72438EBFF2D121FD1DE5A65" pageId="0" pageNumber="157">
|
||
<taxonomicName id="3E9CFB0F4996F0903B62BA20FC546EB3" baseAuthorityName="Theobald" baseAuthorityYear="1870" class="Gastropoda" family="Hypselostomatidae" genus="Boysidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Boysidia salwiniana" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="species" species="salwiniana">Boysidia salwiniana</taxonomicName>
|
||
.
|
||
<bibRefCitation id="6CE632B7EF3A8339FF8CCA01C80B183F" DOI="https://doi.org/10.5962/bhl.title.12891" author="Gude, GK" journalOrPublisher="Taylor & Francis, London" pageId="0" pageNumber="157" refId="B25" refString="Gude, GK, 1914. Mollusca-II. Trochomorphidae-Janellidae. In: Shipley AE, Marshall GAK (Eds) The Fauna of British India, Including Ceylon and Burma. Taylor & Francis, London, DOI: https://doi.org/10.5962/bhl.title.12891" title="Mollusca-II. Trochomorphidae-Janellidae. In: Shipley AE, Marshall GAK (Eds) The Fauna of British India, Including Ceylon and Burma." url="https://doi.org/10.5962/bhl.title.12891" year="1914">Gude 1914</bibRefCitation>
|
||
: 295, 296.
|
||
<bibRefCitation id="36DDF712C1CA6CF45226264F9DB17FB4" author="Pilsbry, HA" journalOrPublisher="ZooKeys" pageId="0" pageNumber="157" refId="B74" refString="Pilsbry, HA, 1917 1918. [1916-1918] Manual of Conchology, Structural and Systematic, with Illustrations of the Species. Series 2, Pulmonata (Vol. 24): Pupillidae (Gastrocoptinae). Academy of Natural Sciences of Philadelphia 380 pp. [pp. 113-256, pls. 14-38 (1917); pp. 257-380, pls 39-49 (1918)]" title="[1916 - 1918] Manual of Conchology, Structural and Systematic, with Illustrations of the Species. Series 2, Pulmonata (Vol. 24): Pupillidae (Gastrocoptinae). Academy of Natural Sciences of Philadelphia 380 pp. [pp. 113 - 256, pls. 14 - 38 (1917); pp. 257 - 380, pls 39 - 49 (1918)]" year="1917">Pilsbry 1917</bibRefCitation>
|
||
: 206-208, pl. 33, fig. 11.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1CF88222E679C3A4D634C806AE7D5D5E" pageId="0" pageNumber="157" type="materials_examined">
|
||
<paragraph id="0A0A7BB6EA90AB829958909A28E46FD8" pageId="0" pageNumber="157">Material examined.</paragraph>
|
||
<paragraph id="33CDD454B8D640FE1CD91BBCE031F343" pageId="0" pageNumber="157">
|
||
<materialsCitation id="96B9A2B96916823D84BC63FB29EC5DD7" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" country="Myanmar" location="Shan State" specimenCount="1" stateProvince="Shan State">
|
||
Burma [
|
||
<collectingCountry id="8B688449B8ADD9F96ED9F8D502AEC8B4" name="Myanmar">Myanmar</collectingCountry>
|
||
]: NHMUK 1912.4.16.66 (2 shells; Fig.
|
||
<figureCitation id="E022663CA808B3DC2565163B45AAD9BD" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Bensonella salwiniana A specimen NHMUK 1912.4.16.66 from Burma B, C specimen CUMZ 14377 from Pyigyidagun Township, Mandalay Region D specimen CUMZ 14376 from Taunggyi District, Shan State E specimen CUMZ 14375 from Taunggyi District, Shan State F specimen CUMZ 14393 from Taunggyi District, Shan State. Abbreviations: a: angular lamella, b: basal plica, c: columellar lamella, ip: infraparietal lamella, lpl: lower palatal plica, p: parietal lamella, upl: upper palatal plica." figureDoi="10.3897/zookeys.1195.112112.figure4" httpUri="https://binary.pensoft.net/fig/1002053" pageId="0" pageNumber="157">4A</figureCitation>
|
||
) ex. Beddome collection.
|
||
<collectingRegion id="0829A6BB7351DD88A48E39446C342F7A" country="Myanmar" name="Shan">Shan State</collectingRegion>
|
||
</materialsCitation>
|
||
:
|
||
<materialsCitation id="AD0A8C2BE1864C5F4FB3C27228DA32EF" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" country="Myanmar" location="Taunggyi District" municipality="Ywangan Township" specimenCount="1" stateProvince="Shan State">
|
||
SMF 227428/2 (2 shells). Monastery,
|
||
<collectingMunicipality id="DF721747F134D544BC143152DE2EBE3B">Ywangan Township</collectingMunicipality>
|
||
,
|
||
<location id="7F114B53C1088B38F66929C925132ABB" LSID="urn:lsid:plazi:treatment:35DDA19D812C590B8914D6A9EB13387B:7F114B53C1088B38F66929C925132ABB" country="Myanmar" municipality="Ywangan Township" name="Taunggyi District" stateProvince="Shan State">Taunggyi District</location>
|
||
,
|
||
<collectingRegion id="C5F28E49FCAD42D842BAB1E887DFBB37" country="Myanmar" name="Shan">Shan State</collectingRegion>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="FD8884E845123C33F4A9335D53379E5A" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" country="Myanmar" county="Fig." latitude="21.228695" location="Dragon Rock" longLatPrecision="1" longitude="96.555336" municipality="Figs" specimenCount="1" stateProvince="Shan State">
|
||
<collectingCountry id="1D22D855D5EB4A9E50EC61C16791F590" name="Myanmar">Myanmar</collectingCountry>
|
||
(locality code SH1;
|
||
<geoCoordinate id="1F84BD91AF9DC814A625FDC263198221" degrees="21" direction="north" minutes="13" orientation="latitude" precision="1" seconds="43.3" value="21.228695">21°13'43.3"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="E607519E4F98D6CD84E8B4204363110B" degrees="96" direction="east" minutes="33" orientation="longitude" precision="1" seconds="19.2" value="96.555336">96°33'19.2"E</geoCoordinate>
|
||
): CUMZ 14375 (1 shell;
|
||
<collectingCounty id="FFA09878B502B7E5F37FE4D263404F23">Fig.</collectingCounty>
|
||
<figureCitation id="F70747682930C3C3478716423E7C5F9E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Bensonella salwiniana A specimen NHMUK 1912.4.16.66 from Burma B, C specimen CUMZ 14377 from Pyigyidagun Township, Mandalay Region D specimen CUMZ 14376 from Taunggyi District, Shan State E specimen CUMZ 14375 from Taunggyi District, Shan State F specimen CUMZ 14393 from Taunggyi District, Shan State. Abbreviations: a: angular lamella, b: basal plica, c: columellar lamella, ip: infraparietal lamella, lpl: lower palatal plica, p: parietal lamella, upl: upper palatal plica." figureDoi="10.3897/zookeys.1195.112112.figure4" httpUri="https://binary.pensoft.net/fig/1002053" pageId="0" pageNumber="157">4E</figureCitation>
|
||
); CUMZ 14391 (2 shells); CUMZ 14392 (11 shells; measured); CUMZ 14393 (2 shells;
|
||
<collectingMunicipality id="A18285AC2A696D8D04CC852EA1FFAAFF">Figs</collectingMunicipality>
|
||
<figureCitation id="845764CFE128D821F4FCAB79DC32D2EB" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Bensonella salwiniana A specimen NHMUK 1912.4.16.66 from Burma B, C specimen CUMZ 14377 from Pyigyidagun Township, Mandalay Region D specimen CUMZ 14376 from Taunggyi District, Shan State E specimen CUMZ 14375 from Taunggyi District, Shan State F specimen CUMZ 14393 from Taunggyi District, Shan State. Abbreviations: a: angular lamella, b: basal plica, c: columellar lamella, ip: infraparietal lamella, lpl: lower palatal plica, p: parietal lamella, upl: upper palatal plica." figureDoi="10.3897/zookeys.1195.112112.figure4" httpUri="https://binary.pensoft.net/fig/1002053" pageId="0" pageNumber="157">4F</figureCitation>
|
||
,
|
||
<figureCitation id="69DEDD376AC60CD166AB66B43F578C05" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A Bensonella salwiniana, specimen CUMZ 14393 from Taunggyi District, Shan State B Bensonella anguloobtusa, specimen CUMZ 14401 from Taunggyi District, Shan State. Abbreviations: a: angular lamella, c: columellar lamella, itpl: interpalatal plica, lpl: lower palatal plica, p: parietal lamella, pt: palatal tubercle, sbc: subcolumellar lamella, upl: upper palatal plica." figureDoi="10.3897/zookeys.1195.112112.figure6" httpUri="https://binary.pensoft.net/fig/1002055" pageId="0" pageNumber="157">6A</figureCitation>
|
||
,
|
||
<figureCitation id="CB5B7C2A00F3D636BBD6CA4E0B80C9CD" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Synoptic view of representatives of hypselostomatid snails recorded from Myanmar A Acinolaemus dayanum, specimen HA Collection B Acinolaemus cryptidentatus, specimen CUMZ 14413 C Anauchen eotvosi, holotype NHMUK 1903.7.1.1227.1 (after Pall-Gergely 2023 a: fig. 1 c) D Angustopila occidentalis, holotype HNHM 103483 (after Pall-Gergely et al. 2023: fig. 70 a) E Angustopila elevata, specimen HNHM 103484 (after Pall-Gergely et al. 2023: fig. 13 a) F Bensonella salwiniana, specimen CUMZ 14393 G Bensonella anguloobtusa, specimen CUMZ 14400 H Bensonella taiyaiorum sp. nov., paratype CUMZ 14403 I Bensonella lophiodera sp. nov., paratype CUMZ 14405 J Clostophis sankeyi, holotype UMZC I. 103320 (after Preece et al. 2022: fig. 70 e) K Clostophis thinbowguensis, holotype MNHN-IM- 2000 - 38057 (after Pall-Gergely et al. 2022: fig. 9 c) L Gyliotrachela bensonianum, syntype NHMUK 20191141 (after Gojsina et al. 2022: fig. 4 a) M Gyliotrachela hungerfordiana, specimen NHMUK collection from Perak, Malaysia N Gyliotrachela tianxingqiaoensis, holotype IZCAS TM 025075 (after Gojsina et al. 2022: fig. 3 a) O Gyliotrachela muangon, specimen CUMZ 14410 P Gyliotrachela aunglini sp. nov., paratype CUMZ 14412 Q Hypselostoma tubiferum, specimen NHMUK 1888.12.4.17 - 22 ex. Theobald collection from Tonduong, Burma." figureDoi="10.3897/zookeys.1195.112112.figure13" httpUri="https://binary.pensoft.net/fig/1002062" pageId="0" pageNumber="157">13F</figureCitation>
|
||
).
|
||
<location id="69C55DBA52B9F990C00B1E5821B3410B" LSID="urn:lsid:plazi:treatment:35DDA19D812C590B8914D6A9EB13387B:69C55DBA52B9F990C00B1E5821B3410B" country="Myanmar" county="Fig." latitude="21.228695" longLatPrecision="1" longitude="96.555336" municipality="Figs" name="Dragon Rock" stateProvince="Shan State">Dragon Rock</location>
|
||
,
|
||
<location id="D01A823E3AE55D989A793A8CFD075470" LSID="urn:lsid:plazi:treatment:35DDA19D812C590B8914D6A9EB13387B:D01A823E3AE55D989A793A8CFD075470" country="Myanmar" county="Fig." latitude="21.228695" longLatPrecision="1" longitude="96.555336" municipality="Figs" name="Pindaya Township" stateProvince="Shan State">Pindaya Township</location>
|
||
,
|
||
<location id="3F1C0903F974C33642D95E1491D5504E" LSID="urn:lsid:plazi:treatment:35DDA19D812C590B8914D6A9EB13387B:3F1C0903F974C33642D95E1491D5504E" country="Myanmar" county="Fig." latitude="21.228695" longLatPrecision="1" longitude="96.555336" municipality="Figs" name="Taunggyi District" stateProvince="Shan State">Taunggyi District</location>
|
||
,
|
||
<collectingRegion id="980E5AC91F9DDDE626C0DDEE8C409059" country="Myanmar" name="Shan">Shan State</collectingRegion>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="AA887E827CA846D27125BE361A382426" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" country="Myanmar" county="Blue Diamond Co." latitude="20.925417" location="Ltd." longLatPrecision="1" longitude="96.65034" municipality="Fig." specimenCount="1" stateProvince="Mandalay Region">
|
||
<collectingCountry id="F153275EC06507ACF67C8FA39069E843" name="Myanmar">Myanmar</collectingCountry>
|
||
(locality code SH5;
|
||
<geoCoordinate id="2A51D46B3FF8D51A74228331199C89BE" degrees="20" direction="north" minutes="55" orientation="latitude" precision="1" seconds="31.5" value="20.925417">20°55'31.5"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="B827F8F4321738760A4986DB17568FF6" degrees="96" direction="east" minutes="39" orientation="longitude" precision="1" seconds="01.2" value="96.65034">96°39'01.2"E</geoCoordinate>
|
||
): CUMZ 14376 (1 shell;
|
||
<collectingMunicipality id="8837DBD123156486D7E8D41698E368B4">Fig.</collectingMunicipality>
|
||
<figureCitation id="86B78277F82902CEA259C1E91C41F26F" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Bensonella salwiniana A specimen NHMUK 1912.4.16.66 from Burma B, C specimen CUMZ 14377 from Pyigyidagun Township, Mandalay Region D specimen CUMZ 14376 from Taunggyi District, Shan State E specimen CUMZ 14375 from Taunggyi District, Shan State F specimen CUMZ 14393 from Taunggyi District, Shan State. Abbreviations: a: angular lamella, b: basal plica, c: columellar lamella, ip: infraparietal lamella, lpl: lower palatal plica, p: parietal lamella, upl: upper palatal plica." figureDoi="10.3897/zookeys.1195.112112.figure4" httpUri="https://binary.pensoft.net/fig/1002053" pageId="0" pageNumber="157">4D</figureCitation>
|
||
); CUMZ 14394 (4 shells).
|
||
<collectingCounty id="5581FA69FB35E119109628E98491BE3C">Blue Diamond Co.</collectingCounty>
|
||
,
|
||
<location id="44D148CE05C2A56A55802C55694D70CE" LSID="urn:lsid:plazi:treatment:35DDA19D812C590B8914D6A9EB13387B:44D148CE05C2A56A55802C55694D70CE" country="Myanmar" county="Blue Diamond Co." latitude="20.925417" longLatPrecision="1" longitude="96.65034" municipality="Fig." name="Ltd." stateProvince="Mandalay Region">Ltd.</location>
|
||
,
|
||
<location id="98AA461A8A932710E87BBF7B8E373196" LSID="urn:lsid:plazi:treatment:35DDA19D812C590B8914D6A9EB13387B:98AA461A8A932710E87BBF7B8E373196" country="Myanmar" county="Blue Diamond Co." latitude="20.925417" longLatPrecision="1" longitude="96.65034" municipality="Fig." name="Pyigyidagun Township" stateProvince="Mandalay Region">Pyigyidagun Township</location>
|
||
,
|
||
<collectingRegion id="43EF2CAD10B561E641A214842767E82A" country="Myanmar" name="Mandalay">Mandalay Region</collectingRegion>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="0ED8A50DC5385B426433A866C3AC77D6" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" country="Myanmar" county="Fig." latitude="21.903444" location="Taunggyi District" longLatPrecision="1" longitude="96.077446" municipality="Aik Kham Cave" specimenCount="1" stateProvince="Shan State">
|
||
<collectingCountry id="B908834A8FFD1A7ABC5EEDFEDF78649A" name="Myanmar">Myanmar</collectingCountry>
|
||
(locality code MD1;
|
||
<geoCoordinate id="22564BE9D6FEA54E60B8193733F4C1D8" degrees="21" direction="north" minutes="54" orientation="latitude" precision="1" seconds="12.4" value="21.903444">21°54'12.4"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="987B11B2DD5C2001841550E9872BF657" degrees="96" direction="east" minutes="04" orientation="longitude" precision="1" seconds="38.8" value="96.077446">96°04'38.8"E</geoCoordinate>
|
||
): CUMZ 14377 (1 shell;
|
||
<collectingCounty id="31C945B5EC89105A1A9862AAF80B59E3">Fig.</collectingCounty>
|
||
<figureCitation id="28DCDF1AD8A00634078748B9E86DBF49" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Bensonella salwiniana A specimen NHMUK 1912.4.16.66 from Burma B, C specimen CUMZ 14377 from Pyigyidagun Township, Mandalay Region D specimen CUMZ 14376 from Taunggyi District, Shan State E specimen CUMZ 14375 from Taunggyi District, Shan State F specimen CUMZ 14393 from Taunggyi District, Shan State. Abbreviations: a: angular lamella, b: basal plica, c: columellar lamella, ip: infraparietal lamella, lpl: lower palatal plica, p: parietal lamella, upl: upper palatal plica." figureDoi="10.3897/zookeys.1195.112112.figure4" httpUri="https://binary.pensoft.net/fig/1002053" pageId="0" pageNumber="157">4B, C</figureCitation>
|
||
); CUMZ 14395 (1 shell).
|
||
<collectingMunicipality id="D3714840E7230E358C840CB922D44A90">Aik Kham Cave</collectingMunicipality>
|
||
,
|
||
<location id="00485B36247DC68BDCDA6BDC08FE4846" LSID="urn:lsid:plazi:treatment:35DDA19D812C590B8914D6A9EB13387B:00485B36247DC68BDCDA6BDC08FE4846" country="Myanmar" county="Fig." latitude="21.903444" longLatPrecision="1" longitude="96.077446" municipality="Aik Kham Cave" name="Taunggyi District" stateProvince="Shan State">Taunggyi District</location>
|
||
,
|
||
<collectingRegion id="F55A6C1DB54BF998D60529C6DBA8F35B" country="Myanmar" name="Shan">Shan State</collectingRegion>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="AB08DB765275898F1A25C5295504F7A3" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" country="Myanmar" latitude="20.818611" location="Taunggyi District" longLatPrecision="1" longitude="97.22833" municipality="Montawa Cave" specimenCount="1" stateProvince="Shan State">
|
||
<collectingCountry id="EA9D96E708358A81429B024F04958563" name="Myanmar">Myanmar</collectingCountry>
|
||
(locality code SH10;
|
||
<geoCoordinate id="0A1B1126BE17C03F35A3376F62DAE597" degrees="20" direction="north" minutes="49" orientation="latitude" precision="1" seconds="07.0" value="20.818611">20°49'07.0"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="13A8B46D76B9E79069CAE7F3D3FC2852" degrees="97" direction="east" minutes="13" orientation="longitude" precision="1" seconds="42.0" value="97.22833">97°13'42.0"E</geoCoordinate>
|
||
): CUMZ 14396 (22 shells).
|
||
<collectingMunicipality id="9BC2DDC060BB10F912E7DDDE40314155">Montawa Cave</collectingMunicipality>
|
||
,
|
||
<location id="6E97C018538ADC14677B91EDBCB002AC" LSID="urn:lsid:plazi:treatment:35DDA19D812C590B8914D6A9EB13387B:6E97C018538ADC14677B91EDBCB002AC" country="Myanmar" latitude="20.818611" longLatPrecision="1" longitude="97.22833" municipality="Montawa Cave" name="Taunggyi District" stateProvince="Shan State">Taunggyi District</location>
|
||
,
|
||
<collectingRegion id="336277262975C63B0A8EE35FB77E2C2E" country="Myanmar" name="Shan">Shan State</collectingRegion>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="83E5449A60B79C73E7730AB3E26CD136" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" country="Myanmar" latitude="20.754389" location="Fig." longLatPrecision="1" longitude="97.01753" specimenCount="85">
|
||
<collectingCountry id="FDED863AA3843875640A451FC6584069" name="Myanmar">Myanmar</collectingCountry>
|
||
(locality code SH11;
|
||
<geoCoordinate id="6B43C9F24C1476427D426BBEA1C95973" degrees="20" direction="north" minutes="45" orientation="latitude" precision="1" seconds="15.8" value="20.754389">20°45'15.8"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="A462D76ED2C9FC030FA11553F9498B85" degrees="97" direction="east" minutes="01" orientation="longitude" precision="1" seconds="03.1" value="97.01753">97°01'03.1"E</geoCoordinate>
|
||
): CUMZ 14397 (
|
||
<specimenCount id="52290AE5E200E717E5AB5274FC87C8B6" count="85" type="generic">85 specimens</specimenCount>
|
||
in ethanol;
|
||
<location id="D9840337C13D64FBBDFD90D114E5E501" LSID="urn:lsid:plazi:treatment:35DDA19D812C590B8914D6A9EB13387B:D9840337C13D64FBBDFD90D114E5E501" country="Myanmar" latitude="20.754389" longLatPrecision="1" longitude="97.01753" name="Fig.">Fig.</location>
|
||
<figureCitation id="F334FB2B8EF1C59C9939F1533F4325F8" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Bensonella salwiniana, specimen CUMZ 14397 from Taunggyi District, Shan State A genitalia and a small inset of penial appendix B preserved specimen with mantle edge, body and foot C snail showing colour in life D radula morphology: blue highlighting indicates central teeth and numbers indicate tooth order from lateral to marginal end. Abbreviations: at: atrium, e: epiphallus, fo: free oviduct, gd: gametolytic duct, gs: gametolytic sac, mc: mantle collar; p: penis, pa: penial appendix, pn: pneumostome; pr: penial retractor muscle, v: vagina, vd: vas deferens." figureDoi="10.3897/zookeys.1195.112112.figure5" httpUri="https://binary.pensoft.net/fig/1002054" pageId="0" pageNumber="157">5</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="7DCBE87DEEDD638A7CC8FB7612AEEEDE" doi="10.3897/zookeys.1195.112112.figure4" httpUri="https://binary.pensoft.net/fig/1002053" pageId="0" pageNumber="157" start="Figure 4" startId="F4">
|
||
<paragraph id="028D09F0C7461D0DD66ED17F807BF97A" pageId="0" pageNumber="157">
|
||
<emphasis id="1F86533F29DCF90435460B849EBB91BB" bold="true" pageId="0" pageNumber="157">Figure 4.</emphasis>
|
||
<taxonomicName id="1D637DD110AE24BCFAC15F8083547031" baseAuthorityName="Theobald" baseAuthorityYear="1871" class="Gastropoda" family="Hypselostomatidae" genus="Bensonella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bensonella salwiniana" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="species" species="salwiniana">
|
||
<emphasis id="A7E834D6B9E20E7A77BEDEB439E113A3" italics="true" pageId="0" pageNumber="157">Bensonella salwiniana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B8007057362FDABB86B78DFC1A083513" bold="true" pageId="0" pageNumber="157">A</emphasis>
|
||
specimen NHMUK 1912.4.16.66 from
|
||
<collectingCountry id="991B243E2108442750791AB003ADA6CE" name="Myanmar">Burma</collectingCountry>
|
||
<emphasis id="94A5F789F5D68077CA674177DD976C68" bold="true" pageId="0" pageNumber="157">B, C</emphasis>
|
||
specimen CUMZ 14377 from Pyigyidagun Township,
|
||
<collectingRegion id="11C5D28306651909F78B271A4AB850B9" country="Myanmar" name="Mandalay">Mandalay Region</collectingRegion>
|
||
<emphasis id="5977B3E8743304A70A59104CBA84E5DA" bold="true" pageId="0" pageNumber="157">D</emphasis>
|
||
specimen CUMZ 14376 from Taunggyi District,
|
||
<collectingRegion id="5FDCEF42A7D539169DD72D6F0460E07A" country="Myanmar" name="Shan">Shan State</collectingRegion>
|
||
<emphasis id="2A3C391B3C3336ADA28C5633F6C49F40" bold="true" pageId="0" pageNumber="157">E</emphasis>
|
||
specimen CUMZ 14375 from Taunggyi District,
|
||
<collectingRegion id="74E7BCA8C528C6FB2BB3D976EEEBBD0F" country="Myanmar" name="Shan">Shan State</collectingRegion>
|
||
<emphasis id="DB3E29A00462A9ABB9ADC6418A39EF9D" bold="true" pageId="0" pageNumber="157">F</emphasis>
|
||
specimen CUMZ 14393 from Taunggyi District,
|
||
<collectingRegion id="AA117A599E04C6D68541DA51C8B15DAF" country="Myanmar" name="Shan">Shan State</collectingRegion>
|
||
. Abbreviations:
|
||
<emphasis id="B3013B98C4A1540F2FA85529D925C3C1" bold="true" pageId="0" pageNumber="157">a</emphasis>
|
||
: angular lamella,
|
||
<emphasis id="A67087649B19B54AF6AC1D69E1F4E8A4" bold="true" pageId="0" pageNumber="157">b</emphasis>
|
||
: basal plica,
|
||
<emphasis id="CAE0C16ADC55E05FADA561ECCED6CA39" bold="true" pageId="0" pageNumber="157">c</emphasis>
|
||
: columellar lamella,
|
||
<emphasis id="0A6D18917DAF3C2178642FB2A2F863CB" bold="true" pageId="0" pageNumber="157">ip</emphasis>
|
||
: infraparietal lamella,
|
||
<emphasis id="60A95345047A44529EF1C3932A67FD0C" bold="true" pageId="0" pageNumber="157">lpl</emphasis>
|
||
: lower palatal plica,
|
||
<emphasis id="06A26D15479118F472BCC5336AE5B574" bold="true" pageId="0" pageNumber="157">p</emphasis>
|
||
: parietal lamella,
|
||
<emphasis id="FEA067419C33337655AF9CAA933BCD9A" bold="true" pageId="0" pageNumber="157">upl</emphasis>
|
||
: upper palatal plica.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="A66CBAB14003106F7186BE246A9CDDC4" doi="10.3897/zookeys.1195.112112.figure5" httpUri="https://binary.pensoft.net/fig/1002054" pageId="0" pageNumber="157" start="Figure 5" startId="F5">
|
||
<paragraph id="613236C47A5C2BA38454A485CDD2149B" pageId="0" pageNumber="157">
|
||
<emphasis id="E7646DF57DE27B001777DA918B9B690C" bold="true" pageId="0" pageNumber="157">Figure 5.</emphasis>
|
||
<taxonomicName id="F4EF1663F52CBA892321E2EDF56657C2" baseAuthorityName="Theobald" baseAuthorityYear="1871" class="Gastropoda" family="Hypselostomatidae" genus="Bensonella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bensonella salwiniana" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="species" species="salwiniana">
|
||
<emphasis id="BD5417C523B82CA59EF35E54DE87B715" italics="true" pageId="0" pageNumber="157">Bensonella salwiniana</emphasis>
|
||
</taxonomicName>
|
||
, specimen CUMZ 14397 from Taunggyi District,
|
||
<collectingRegion id="0CF940F5DD7E192FA4BD430AFE1911EA" country="Myanmar" name="Shan">Shan State</collectingRegion>
|
||
<emphasis id="E377F3EFCAE5D7F01008FCEA6BF2442F" bold="true" pageId="0" pageNumber="157">A</emphasis>
|
||
genitalia and a small inset of penial appendix
|
||
<emphasis id="209609F8ABC85D1C0E18200A82740EEE" bold="true" pageId="0" pageNumber="157">B</emphasis>
|
||
preserved specimen with mantle edge, body and foot
|
||
<emphasis id="662F52F7E9B0440E0149BDBF6DBD14D9" bold="true" pageId="0" pageNumber="157">C</emphasis>
|
||
snail showing colour in life
|
||
<emphasis id="6372F9289F708491E91B1BB636D5BC72" bold="true" pageId="0" pageNumber="157">D</emphasis>
|
||
radula morphology: blue highlighting indicates central teeth and numbers indicate tooth order from lateral to marginal end. Abbreviations:
|
||
<emphasis id="D9C3091F5BF8B88DF9153C359223A63D" bold="true" pageId="0" pageNumber="157">at</emphasis>
|
||
: atrium,
|
||
<emphasis id="C5859D433C2C7BBAA6A8FBDBA5E32A09" bold="true" pageId="0" pageNumber="157">e</emphasis>
|
||
: epiphallus,
|
||
<emphasis id="56C786F42A1BF03A9FD0048B1DE7E722" bold="true" pageId="0" pageNumber="157">fo</emphasis>
|
||
: free oviduct,
|
||
<emphasis id="74F27B6D033C999ABD40AAB97EE80CE3" bold="true" pageId="0" pageNumber="157">gd</emphasis>
|
||
: gametolytic duct,
|
||
<emphasis id="11CABAAC3297C9CCE0047E7EEAF56F6A" bold="true" pageId="0" pageNumber="157">gs</emphasis>
|
||
: gametolytic sac,
|
||
<emphasis id="0163DF063F0BEDF8EFDC562F6FCA7278" bold="true" pageId="0" pageNumber="157">mc</emphasis>
|
||
: mantle collar;
|
||
<emphasis id="65E3041CE9172352DCC8DC64871352B6" bold="true" pageId="0" pageNumber="157">p</emphasis>
|
||
: penis,
|
||
<emphasis id="FF20BC65B6D39E549B7D37A8A7F45BE4" bold="true" pageId="0" pageNumber="157">pa</emphasis>
|
||
: penial appendix,
|
||
<emphasis id="D78F5C7A37FA98F391BE363AA84928B6" bold="true" pageId="0" pageNumber="157">pn</emphasis>
|
||
: pneumostome;
|
||
<emphasis id="D8F6FBDBDF3FAEC4CCD4849A337E1F60" bold="true" pageId="0" pageNumber="157">pr</emphasis>
|
||
: penial retractor muscle,
|
||
<emphasis id="CCE72ADE0947C81FA9A0D9CACBFEFC11" bold="true" pageId="0" pageNumber="157">v</emphasis>
|
||
: vagina,
|
||
<emphasis id="2DF5C5F4ACF9F7B34AFAB082A38060E3" bold="true" pageId="0" pageNumber="157">vd</emphasis>
|
||
: vas deferens.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="57FFAA056E31498504406965B89AB2E6" doi="10.3897/zookeys.1195.112112.figure6" httpUri="https://binary.pensoft.net/fig/1002055" pageId="0" pageNumber="157" start="Figure 6" startId="F6">
|
||
<paragraph id="F8217E883C17F821586618C015C810FB" pageId="0" pageNumber="157">
|
||
<emphasis id="B173A7EC0B7B491245FC1DD7C2F1B469" bold="true" pageId="0" pageNumber="157">Figure 6.</emphasis>
|
||
<emphasis id="F82FB7F25016FF1B8A0EF7D704BB304E" bold="true" pageId="0" pageNumber="157">A</emphasis>
|
||
<taxonomicName id="A612E6437543804188DBEB81B909A074" baseAuthorityName="Theobald" baseAuthorityYear="1871" class="Gastropoda" family="Hypselostomatidae" genus="Bensonella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bensonella salwiniana" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="species" species="salwiniana">
|
||
<emphasis id="BD3869822BC8CA34D69191FAC7B184ED" italics="true" pageId="0" pageNumber="157">Bensonella salwiniana</emphasis>
|
||
</taxonomicName>
|
||
, specimen CUMZ 14393 from Taunggyi District,
|
||
<collectingRegion id="726DC3CD5B81503A244D1C7605C48612" country="Myanmar" name="Shan">Shan State</collectingRegion>
|
||
<emphasis id="BDC84C58A3F967A50785F915F6A3901A" bold="true" pageId="0" pageNumber="157">B</emphasis>
|
||
<taxonomicName id="C05C648AE93212D88A45577CCEB06A94" baseAuthorityName="Inkhavilay & Panha" baseAuthorityYear="2016" class="Gastropoda" family="Hypselostomatidae" genus="Bensonella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bensonella anguloobtusa" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="species" species="anguloobtusa">
|
||
<emphasis id="9075819FC436AABB86E14F752C0D3BF5" italics="true" pageId="0" pageNumber="157">Bensonella anguloobtusa</emphasis>
|
||
</taxonomicName>
|
||
, specimen CUMZ 14401 from Taunggyi District,
|
||
<collectingRegion id="4596EB86C0706EFBE4A6B8775FB10475" country="Myanmar" name="Shan">Shan State</collectingRegion>
|
||
. Abbreviations:
|
||
<emphasis id="08F30043FC9E4E98B690F157DE507220" bold="true" pageId="0" pageNumber="157">a</emphasis>
|
||
: angular lamella,
|
||
<emphasis id="83009D7F28493758BD15B90AD27F4805" bold="true" pageId="0" pageNumber="157">c</emphasis>
|
||
: columellar lamella,
|
||
<emphasis id="E5D6BEC4DB09AFF5A94D7AEC4DD12D73" bold="true" pageId="0" pageNumber="157">itpl</emphasis>
|
||
: interpalatal plica,
|
||
<emphasis id="06091CFE0FD33EF12E8F092D45B789C5" bold="true" pageId="0" pageNumber="157">lpl</emphasis>
|
||
: lower palatal plica,
|
||
<emphasis id="19BC494C1A3F707D739ADFB9D6E9C9B0" bold="true" pageId="0" pageNumber="157">p</emphasis>
|
||
: parietal lamella,
|
||
<emphasis id="88ED5EB14AE43D1F09A6C69071F1D9B1" bold="true" pageId="0" pageNumber="157">pt</emphasis>
|
||
: palatal tubercle,
|
||
<emphasis id="DB0E96F6F0D49EE43EC43D6906F27C8F" bold="true" pageId="0" pageNumber="157">sbc</emphasis>
|
||
: subcolumellar lamella,
|
||
<emphasis id="F950300EF01E5FC9DE996120BD0F50C2" bold="true" pageId="0" pageNumber="157">upl</emphasis>
|
||
: upper palatal plica.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="CE3BA79678831E3619A5D79B09642DAF" pageId="0" pageNumber="157" type="description">
|
||
<paragraph id="0466B23FE2C17EB358947317E87BE00C" pageId="0" pageNumber="157">Description.</paragraph>
|
||
<paragraph id="476A7D2A5E30843C364EAA12949B32E4" pageId="0" pageNumber="157">
|
||
Shell ovate-conical, high spire, yellowish brown, with 6-
|
||
<normalizedToken id="83A255C4BB6B7D01C26A682478E70174" originalValue="6½">61/2</normalizedToken>
|
||
convex whorls. Shell height 5.6-6.3 mm and shell width 3.3-3.7 mm. Apex blunt; protoconch ~
|
||
<normalizedToken id="278A50630B354B361F817168F10CBB5C" originalValue="1½">11/2</normalizedToken>
|
||
whorls, sculptured with radial wrinkles. Teleoconch with smooth, irregular growth lines; suture well impressed and deep. Last whorl large and rounded. Peristome thickened and broadly expanded; lip whitish. Aperture rounded-subquadrate with six or seven dentitions. Parietal lamella large, long, strong, broadly blunt, and located slightly deeper inside aperture; infraparietal lamella very small and sometimes absent. Angular lamella blunt and reaching peristomal lip. Palatal tubercle inconspicuous. Upper palatal plica long, contacting peristome and sometimes elevated in middle; lower palatal plica strong and large. Basal plica weak to strong nodule shape. Columellar lamella strong and large, tubercle-like. Umbilicus very narrowly perforate, rounded and deep.
|
||
</paragraph>
|
||
<paragraph id="F371B79FE37F12C0B8C6B546526EA0FF" pageId="0" pageNumber="157">
|
||
<emphasis id="90F39B96D4DBB1D525C22161D4D200B0" bold="true" pageId="0" pageNumber="157">
|
||
<emphasis id="D384ABF6CECEBF957E0BE66E47DDFA4F" italics="true" pageId="0" pageNumber="157">Genital system</emphasis>
|
||
.
|
||
</emphasis>
|
||
Atrium short and slightly enlarged. Penis very long, thin tube and slightly enlarged at both ends; penial appendix short. Penial retractor muscle large and inserted between penis and epiphallus junction. Epiphallus slender tube, almost same length as penis, and ~ 2/3 of its length enlarged with similar diameter to free oviduct. Vas deferens very thin tube, connected between epiphallus and free oviduct, and held in position with thin connective tissue near epiphallus-vas deferens junction to atrium.
|
||
</paragraph>
|
||
<paragraph id="13BB279D708D662DBECEA3254A521FD6" pageId="0" pageNumber="157">Vagina slender tube, short and ~ 1/3 of penis length. Gametolytic organ strongly developed; gametolytic duct slender tube, almost same length as vagina+free oviduct; gametolytic sac enlarged and bulbous. Free oviduct larger in diameter than vagina, and approximately same length as vagina.</paragraph>
|
||
<paragraph id="65A840714DA9D0A586226E92FC260A79" pageId="0" pageNumber="157">
|
||
<emphasis id="7E57FE25CE304DD59CE8CEBFCFD0F919" bold="true" pageId="0" pageNumber="157">
|
||
<emphasis id="944D546A7DCB4A9C3585C75AFB6D08F8" italics="true" pageId="0" pageNumber="157">Animal</emphasis>
|
||
.
|
||
</emphasis>
|
||
Preserved specimen with blackish to greyish reticulated skin, and mantle collar well-developed and whitish. Pneumostome wide and situated in the bay of angular lamella and upper palatal plicae. Foot short, holopodal, unipartite, and sole of foot blackish to greyish in colour. Living snails possess blackish tentacles: upper tentacles short and stout, and lower tentacles very short.
|
||
</paragraph>
|
||
<paragraph id="480D5CBA2D654EC5642E9F8C3E0B4819" pageId="0" pageNumber="157">
|
||
<emphasis id="6B0ED3EE5659723172C3F26A2CA68F13" bold="true" pageId="0" pageNumber="157">
|
||
<taxonomicName id="CDD18504CDFFE95C828186CBB862F952" class="Bivalvia" family="Limidae" genus="Radula" kingdom="Animalia" lsidName="Radula" order="Limida" pageId="0" pageNumber="157" phylum="Mollusca" rank="genus">
|
||
<emphasis id="2CA4F4566147123B376ED9B06190465D" italics="true" pageId="0" pageNumber="157">Radula</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Teeth arranged in nearly straight row with formula 15-(6, 5)-1-(5, 6)-11+. Central tooth unicuspid, with long triangular shape and pointed tip. Lateral teeth bicuspid, inner and outer cusps clearly separated at base, situated next to each other, and nearly aligned in transverse row. Inner and outer cusps long and triangular, and outer cusp comparatively smaller than inner cusp. Marginal teeth starting at approximately tooth number 5 or 6, inner and outer cusps of marginals joined at base. Innermost marginals bicuspid, similar to lateral teeth; outermost teeth multicuspid, cusps small with almost same size and shape, and situated on same base.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6C31FCC00785C3EA11011E546940529C" pageId="0" pageNumber="157" type="distribution">
|
||
<paragraph id="68BC3448E22F8ABAD185A466A7E27F44" pageId="0" pageNumber="157">Distribution.</paragraph>
|
||
<paragraph id="AE44351209D83484F6412C0F5B1E79F2" pageId="0" pageNumber="157">
|
||
This species is known from several localities in northern Myanmar: Bhamo [Bhamo District, Kachin State]; Pingoung [Pingku Hills, Muse District, Shan State]; Shan Hills (
|
||
<bibRefCitation id="883AAC81931872217F1283092DCF4B7D" DOI="https://doi.org/10.5962/bhl.title.12891" author="Gude, GK" journalOrPublisher="Taylor & Francis, London" pageId="0" pageNumber="157" refId="B25" refString="Gude, GK, 1914. Mollusca-II. Trochomorphidae-Janellidae. In: Shipley AE, Marshall GAK (Eds) The Fauna of British India, Including Ceylon and Burma. Taylor & Francis, London, DOI: https://doi.org/10.5962/bhl.title.12891" title="Mollusca-II. Trochomorphidae-Janellidae. In: Shipley AE, Marshall GAK (Eds) The Fauna of British India, Including Ceylon and Burma." url="https://doi.org/10.5962/bhl.title.12891" year="1914">Gude 1914</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="97B07C61C882B30481B284AE0DF773D0" author="Pilsbry, HA" journalOrPublisher="ZooKeys" pageId="0" pageNumber="157" refId="B74" refString="Pilsbry, HA, 1917 1918. [1916-1918] Manual of Conchology, Structural and Systematic, with Illustrations of the Species. Series 2, Pulmonata (Vol. 24): Pupillidae (Gastrocoptinae). Academy of Natural Sciences of Philadelphia 380 pp. [pp. 113-256, pls. 14-38 (1917); pp. 257-380, pls 39-49 (1918)]" title="[1916 - 1918] Manual of Conchology, Structural and Systematic, with Illustrations of the Species. Series 2, Pulmonata (Vol. 24): Pupillidae (Gastrocoptinae). Academy of Natural Sciences of Philadelphia 380 pp. [pp. 113 - 256, pls. 14 - 38 (1917); pp. 257 - 380, pls 39 - 49 (1918)]" year="1917">Pilsbry 1917</bibRefCitation>
|
||
). In this study, several specimens were also collected from Shan State and the Mandalay Region.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="68001AF6F2AAD5D0A0B15A3E2A44D92B" pageId="0" pageNumber="157" type="differential diagnosis">
|
||
<paragraph id="DE17B7DF819BBD445A7E21FD31771838" pageId="0" pageNumber="157">Differential diagnosis.</paragraph>
|
||
<paragraph id="CE8CE8D7D2507FEFE43F9F51CED5AF69" pageId="0" pageNumber="157">
|
||
<taxonomicName id="EF28130BF59504F1AE44B1ADD9E62845" baseAuthorityName="Theobald" baseAuthorityYear="1871" class="Gastropoda" family="Hypselostomatidae" genus="Bensonella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bensonella salwiniana" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="species" species="salwiniana">
|
||
<emphasis id="AFD331AFB62D93424164EA5ABE468E58" italics="true" pageId="0" pageNumber="157">Bensonella salwiniana</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="EE22CCB17F55359F16D98F37DEB428DE" authorityName="Pilsbry & Vanatta" authorityYear="1900" class="Gastropoda" family="Hypselostomatidae" genus="Bensonella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bensonella" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="genus">
|
||
<emphasis id="AD27B079581CFDDB1A95857402EE65CD" italics="true" pageId="0" pageNumber="157">Bensonella</emphasis>
|
||
</taxonomicName>
|
||
species reported from Thailand and Laos in having a much larger (height 4-6 mm) and ovate conical shell, 6-7 whorls, inconspicuous palatal tubercle, and generally with six apertural dentitions. In contrast,
|
||
<taxonomicName id="B6170A1A32BEC958BCF17D0494BB39BD" lsidName="B. nabhitabhatai" pageId="0" pageNumber="157" rank="species" species="nabhitabhatai">
|
||
<emphasis id="B6A5B89B903CAC4D8499B6B3F843A518" italics="true" pageId="0" pageNumber="157">B. nabhitabhatai</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EB864700B68E9A7A798529ACD45142A8" lsidName="B. tamphathai" pageId="0" pageNumber="157" rank="species" species="tamphathai">
|
||
<emphasis id="37D46D7BA636431620DE860F01DA98FC" italics="true" pageId="0" pageNumber="157">B. tamphathai</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="8A6811470C378E19A401975EEEF7D804" lsidName="B. pangmapaensis" pageId="0" pageNumber="157" rank="species" species="pangmapaensis">
|
||
<emphasis id="5FA34825C8D4A9CDA5AFEB6526D01F82" italics="true" pageId="0" pageNumber="157">B. pangmapaensis</emphasis>
|
||
</taxonomicName>
|
||
tend to have smaller shells (height 2-4 mm), 4-5 whorls, a strong palatal tubercle, and generally bears 8-10 apertural dentitions (
|
||
<bibRefCitation id="A8B3E00DCAC5F813E232C309840AFB26" author="Panha, S" journalOrPublisher="Walkerana" pageId="0" pageNumber="157" pagination="107 - 112" refId="B59" refString="Panha, S, Burch, JB, 2000. A new species of Paraboysidia from Thailand (Pulmonata: Pupillidae). Walkerana 11 (25): 107 - 112" title="A new species of Paraboysidia from Thailand (Pulmonata: Pupillidae)." volume="11" year="2000">Panha and Burch 2000</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="E25E10453DC17C0818EDC80F305DBF42" author="Panha, S" journalOrPublisher="Walkerana" pageId="0" pageNumber="157" refId="B61" refString="Panha, S, Burch, JB, 2002b. The pupillid genus Paraboysidia in Thailand (Pulmonata: Stylommatophora). Walkerana 12[2001](28): 77-94. [Date of publication follow Jirapatrasilp et al. (2023)]" title="The pupillid genus Paraboysidia in Thailand (Pulmonata: Stylommatophora). Walkerana 12 [2001] (28): 77 - 94. [Date of publication follow Jirapatrasilp et al. (2023)]" year="2002 b">2002b</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="BA3350EDE3263029ACCAE182FE70860D" pageId="0" pageNumber="157">
|
||
For further comparison, the three species from Laos, namely
|
||
<taxonomicName id="260E2FBB8CFD1C14A45ADCFC38B82F84" lsidName="B. wangviangensis" pageId="0" pageNumber="157" rank="species" species="wangviangensis">
|
||
<emphasis id="360B96DA41A96AB9B054AE2A28685D1B" italics="true" pageId="0" pageNumber="157">B. wangviangensis</emphasis>
|
||
</taxonomicName>
|
||
(Panha & Tongkerd, 2003),
|
||
<taxonomicName id="D27E3A1A0E9BDD2A63E265624AE04C1E" lsidName="B. paralella" pageId="0" pageNumber="157" rank="species" species="paralella">
|
||
<emphasis id="2B1EFF8471BD9DF3931E5417CC5E0FB8" italics="true" pageId="0" pageNumber="157">B. paralella</emphasis>
|
||
</taxonomicName>
|
||
(Inkhavilay & Panha, 2016), and
|
||
<taxonomicName id="94EAC0C9B9D8D780F2446DEF98E6401A" lsidName="B. anguloobtusa" pageId="0" pageNumber="157" rank="species" species="anguloobtusa">
|
||
<emphasis id="E9DA86BC9B8BAA12DE9E1ADB5BB2CEDE" italics="true" pageId="0" pageNumber="157">B. anguloobtusa</emphasis>
|
||
</taxonomicName>
|
||
(Inkhavilay & Panha, 2016) possess a smaller shell (height 1-3 mm), a strongly developed palatal tubercle and generally bears 3-4 whorls (
|
||
<bibRefCitation id="8B6779BDF7BA7FF9F48D2414F1535A02" author="Panha, S" journalOrPublisher="Walkerana" pageId="0" pageNumber="157" refId="B61" refString="Panha, S, Burch, JB, 2002b. The pupillid genus Paraboysidia in Thailand (Pulmonata: Stylommatophora). Walkerana 12[2001](28): 77-94. [Date of publication follow Jirapatrasilp et al. (2023)]" title="The pupillid genus Paraboysidia in Thailand (Pulmonata: Stylommatophora). Walkerana 12 [2001] (28): 77 - 94. [Date of publication follow Jirapatrasilp et al. (2023)]" year="2002 b">Panha and Burch 2002b</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="79AD8B716DEDF0B0AFB5185D58173707" author="Panha, S" journalOrPublisher="Walkerana" pageId="0" pageNumber="157" refId="B63" refString="Panha, S, Burch, JB, 2005. An introduction to the microsnails of Thailand. Malacological Review 37/38(2004-2005): 1-155. [Date of publication follow Jirapatrasilp et al. (2023)]" title="An introduction to the microsnails of Thailand. Malacological Review 37 / 38 (2004 - 2005): 1 - 155. [Date of publication follow Jirapatrasilp et al. (2023)]" year="2005">2005</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="18CFE97BB8263CF84605DCBF7433C122" author="Panha, S" journalOrPublisher="Walkerana" pageId="0" pageNumber="157" refId="B64" refString="Panha, S, Tongkerd, P, Sutcharit, C, Tumpeesuwan, S, Vongsombath, C, 2003. A new species of Paraboysidia (Pupillidae: Gastrocoptinae) from Laos. Walkerana 13[2002](29/30): 123-128." title="A new species of Paraboysidia (Pupillidae: Gastrocoptinae) from Laos. Walkerana 13 [2002] (29 / 30): 123 - 128." year="2003">Panha et al. 2003</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="536CA4A0D62D2DFF50062307C2CE8FA4" author="Inkhavilay, K" journalOrPublisher="Journal of Conchology" pageId="0" pageNumber="157" pagination="213 - 232" refId="B28" refString="Inkhavilay, K, Sutcharit, C, Tongkerd, P, Panha, S, 2016. New species of micro snails from Laos (Pulmonata: Vertiginidae and Diapheridae). Journal of Conchology 42 (4): 213 - 232" title="New species of micro snails from Laos (Pulmonata: Vertiginidae and Diapheridae)." volume="42" year="2016">Inkhavilay et al. 2016</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="652DB56D806745DDCD632FE56B8A057B" pageId="0" pageNumber="157">
|
||
Although this species is very similar to
|
||
<taxonomicName id="8A3E8562A2A7DBCBBAADD66792ACA4D1" lsidName="B. gittenbergeri" pageId="0" pageNumber="157" rank="species" species="gittenbergeri">
|
||
<emphasis id="A0F3C015C600C56AC521E1DBA034FE9E" italics="true" pageId="0" pageNumber="157">B. gittenbergeri</emphasis>
|
||
</taxonomicName>
|
||
(Maassen, 2008) from Luang Namtha Province, Laos, in shell shape and size, it differs by having a lower palatal plica not extended to an expanded lip, a very narrow umbilicus, and an unextended palato-basal wall on the anterior side. In contrast,
|
||
<taxonomicName id="E2552DF5E8FAC9D3717961BF270CF6CE" lsidName="B. gittenbergeri" pageId="0" pageNumber="157" rank="species" species="gittenbergeri">
|
||
<emphasis id="6212CC641D0D9011E3218F8083F97583" italics="true" pageId="0" pageNumber="157">B. gittenbergeri</emphasis>
|
||
</taxonomicName>
|
||
possesses a long lower palatal plica that extends to an expanded lip, has a wider umbilicus, and its palato-basal wall is extended anteriorly (
|
||
<bibRefCitation id="23E1600BCECE086405F63BBF36AB8309" author="Maassen, WJM" journalOrPublisher="Basteria" pageId="0" pageNumber="157" pagination="233 - 240" refId="B35" refString="Maassen, WJM, 2008. Remarks on a small collection of terrestrial molluscs from north-west Laos, with descriptions of three new species (Mollusca: Pulmonata: Streptaxidae, Vertiginidae). Basteria 72: 233 - 240" title="Remarks on a small collection of terrestrial molluscs from north-west Laos, with descriptions of three new species (Mollusca: Pulmonata: Streptaxidae, Vertiginidae)." volume="72" year="2008">Maassen 2008</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="D0418F2BC3448070D73BDA37080F968B" author="Inkhavilay, K" journalOrPublisher="Journal of Conchology" pageId="0" pageNumber="157" pagination="213 - 232" refId="B28" refString="Inkhavilay, K, Sutcharit, C, Tongkerd, P, Panha, S, 2016. New species of micro snails from Laos (Pulmonata: Vertiginidae and Diapheridae). Journal of Conchology 42 (4): 213 - 232" title="New species of micro snails from Laos (Pulmonata: Vertiginidae and Diapheridae)." volume="42" year="2016">Inkhavilay et al. 2016</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D70FD890715DBBA267E1B9448E03BA22" pageId="0" pageNumber="157">
|
||
<taxonomicName id="6A16F80535A8BC17BE8604D6BE9F17F2" baseAuthorityName="Theobald" baseAuthorityYear="1871" class="Gastropoda" family="Hypselostomatidae" genus="Bensonella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bensonella salwiniana" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="species" species="salwiniana">
|
||
<emphasis id="78E5CE3B51A5786DDA6570E444C9581C" italics="true" pageId="0" pageNumber="157">Bensonella salwiniana</emphasis>
|
||
</taxonomicName>
|
||
clearly differs from the two newly described species,
|
||
<taxonomicName id="97A07AF3E593607B8052DC41B4C5B832" lsidName="B. lophiodera" pageId="0" pageNumber="157" rank="species" species="lophiodera">
|
||
<emphasis id="E95039ABEF35F5383FD28E56C93580A4" italics="true" pageId="0" pageNumber="157">B. lophiodera</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. and
|
||
<taxonomicName id="E5FE6666A4069D227473F72E91320DB1" lsidName="B. taiyaiorum" pageId="0" pageNumber="157" rank="species" species="taiyaiorum">
|
||
<emphasis id="956009EB08366D4EA311EA7777EEF529" italics="true" pageId="0" pageNumber="157">B. taiyaiorum</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., in having a larger shell (height 5-6 mm), and a broadly expanded peristome without cervical crest and lacking supra- and subcolumellar lamellae. Moreover, the latter two species possess smaller shells (height ~ 3 mm), a slightly expanded lip, and a conspicuous cervical crest. In addition,
|
||
<taxonomicName id="119B0D379CDECF7851935FEA04A9040E" lsidName="B. taiyaiorum" pageId="0" pageNumber="157" rank="species" species="taiyaiorum">
|
||
<emphasis id="DF277B6D5C8FA5E0F8535B8A9D628CF4" italics="true" pageId="0" pageNumber="157">B. taiyaiorum</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. has two upper palatal plicae, palatal tubercle strongly developed, supra-, subcolumellar lamellae and u-shaped plica present on the parietal wall, and a cervical crest far from the apertural lip. In contrast,
|
||
<taxonomicName id="3B2766D7544A476FB8E5285ADF29ED5E" lsidName="B. lophiodera" pageId="0" pageNumber="157" rank="species" species="lophiodera">
|
||
<emphasis id="B4AECECD3B474D219F9BD4B68B5C34BC" italics="true" pageId="0" pageNumber="157">B. lophiodera</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. possesses a weakly expanded lip, subcolumellar lamella present and cervical crest close to apertural lip.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5D0EF8E6A62FE5A7A97DC5330D51EE2B" pageId="0" pageNumber="157" type="remarks">
|
||
<paragraph id="7DF2EFA1D5B67D1B9823BD67036F2FF1" pageId="0" pageNumber="157">Remarks.</paragraph>
|
||
<paragraph id="11283847E1EAFFF6F39EA01547A73BE0" pageId="0" pageNumber="157">
|
||
The species was described based on specimens received from F. Fedden (1839-1887). The specimens lacked a precise type locality, and only 'Shan
|
||
<normalizedToken id="3F0A1EC23D4547A82331D6A761D1F1DA" originalValue="States’">States'</normalizedToken>
|
||
was stated in the original description. Although the type specimen could not be located, several recently collected and historical specimens were examined. This species possesses strongly developed parietal and angular lamellae, and lacks a tuba, which are the diagnostic characters of
|
||
<taxonomicName id="DA8BC3E292890ACD00D3F87BC9A7B85C" authorityName="Pilsbry & Vanatta" authorityYear="1900" class="Gastropoda" family="Hypselostomatidae" genus="Bensonella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bensonella" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="genus">
|
||
<emphasis id="808423C4F9576444C43E8760A9A9A46E" italics="true" pageId="0" pageNumber="157">Bensonella</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="816AF9E87B3DABE6851BEFD2B84DE61C" pageId="0" pageNumber="157">The examined individuals either possess or lack an infraparietal lamella.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |