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<document ID-CLB-Dataset="21746" ID-DOI="http://dx.doi.org/10.3897/zookeys.938.51614" ID-GBIF-Dataset="5fcad13c-6e87-4afa-8c82-e12c2125ce2a" ID-PMC="PMC7289897" ID-Pensoft-Pub="1313-2970-938-87" ID-Pensoft-UUID="1F1A43237863590DA1C89D0B8CE995EB" ID-PubMed="32550786" ID-ZooBank="AF97861A141B44EF8A34AFEE6E792EF1" ModsDocID="1313-2970-938-87" checkinTime="1591376754349" checkinUser="pensoft" docAuthor="Hoshina, Hideto" docDate="2020" docId="3551D951E4325E6E9E51D6C99630554A" docLanguage="en" docName="ZooKeys 938: 87-95" docOrigin="ZooKeys 938" docSource="http://dx.doi.org/10.3897/zookeys.938.51614" docTitle="Triarthron itoi Hoshina 2020, sp. nov." docType="treatment" docUuid="198DA238-E24B-4329-A2B8-7A197EAAE909" docUuidSource="ZooBank" docVersion="6" id="1F1A43237863590DA1C89D0B8CE995EB" lastPageNumber="87" masterDocId="1F1A43237863590DA1C89D0B8CE995EB" masterDocTitle="Discovery of a new species of the genus Triarthron Maerkel, 1840 (Coleoptera, Leiodidae) with a key to Japanese species of the tribe Sogdini" masterLastPageNumber="95" masterPageNumber="87" pageNumber="87" updateTime="1732856901604" updateUser="ExternalLinkService">
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<mods:title id="3FB4B9E90CB19E548C06D851360C9867">Discovery of a new species of the genus Triarthron Maerkel, 1840 (Coleoptera, Leiodidae) with a key to Japanese species of the tribe Sogdini</mods:title>
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<mods:namePart id="3AAEEAE0E6F0830D7298EB12661B4F74">Hoshina, Hideto</mods:namePart>
<mods:affiliation id="90AFEB947120CD732A8E83A9DD5D147E">Faculty of Education, University of Fukui, Fukui City, 910 - 8507, Japan</mods:affiliation>
<mods:nameIdentifier id="9ED64ECB08A476E47518184D57819832" type="email">hhoshina@f-edu.u-fukui.ac.jp</mods:nameIdentifier>
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<mods:title id="DB649022340935202C8B16434812F0CA">ZooKeys</mods:title>
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<mods:date id="955793B735C5CB6DB0941062AD0FAB91">2020</mods:date>
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<mods:identifier id="49C7E107E9BEFB7E3DE3BE085E1CC4AC" type="DOI">http://dx.doi.org/10.3897/zookeys.938.51614</mods:identifier>
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<treatment id="3551D951E4325E6E9E51D6C99630554A" ID-GBIF-Taxon="164479809" LSID="urn:lsid:zoobank.org:act:198DA238-E24B-4329-A2B8-7A197EAAE909" httpUri="http://treatment.plazi.org/id/3551D951E4325E6E9E51D6C99630554A" lastPageNumber="87" pageId="0" pageNumber="87" scope_family="Leiodidae" scope_order="Coleoptera" scope_tribe="Sogdini">
<subSubSection id="26A17910614367F9757C903EA08B22BF" pageId="0" pageNumber="87" type="nomenclature">
<paragraph id="BACE5C0D6F0EE33403C158FCF76B1453" pageId="0" pageNumber="87">
<taxonomicName id="93D07DE3D7BB54C6B7AEB465640455DE" ID-CoL="8YBXZ" LSID="3551D951-E432-5E6E-9E51-D6C99630554A" authority="Hoshina" authorityName="Hoshina" authorityYear="2020" class="Insecta" family="Leiodidae" genus="Triarthron" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Triarthron itoi" order="Coleoptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="itoi" status="sp. nov.">Triarthron itoi Hoshina</taxonomicName>
<taxonomicNameLabel id="8C7AA8DC9CEAB38C5276D05CADBC8988" pageId="0" pageNumber="87">sp. nov.</taxonomicNameLabel>
<figureCitation id="D514796B278E82F4B206F735ACB16F1C" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitat of Triarthron itoi Hoshina, sp. nov. Scale bar: 1 mm." figureDoi="10.3897/zookeys.938.51614.figure1" httpUri="https://binary.pensoft.net/fig/418208" pageId="0" pageNumber="87">Figures 1</figureCitation>
<figureCitation id="05529BE5DB9E0CC2B5FF261C06CBA702" captionStart="Figures 27" captionStartId="F2" captionText="Figures 2 - 7. Triarthron itoi Hoshina, sp. nov. (2 - 5). 2 Body 3 right antennomere, dorsal view 4 left mesofemur, ventral view 5 left metafemur, ventral view. T. maerkelii Maerkel (6, 7). 6 Left mesofemur, ventral view 7 left metafemur, ventral view. Scale bar: A: 1 mm (2), B: 0.2 mm (3), C, D: 0.5 mm (4 - 7)." figureDoi="10.3897/zookeys.938.51614.figures2-7" httpUri="https://binary.pensoft.net/fig/418209" pageId="0" pageNumber="87">, 2-5</figureCitation>
<figureCitation id="F3A25F914D0BAAF70FB269EC05CD7ADF" captionStart="Figures 810" captionStartId="F3" captionText="Figures 8 - 10. Triarthron itoi Hoshina sp. nov. (8, 9). 8 Aedeagus, dorsal view 9 ditto, lateral view. T. maerkelii Maerkel (10) aedeagus, dorsal view. Scale bar: 0.5 mm (8 - 10)." figureDoi="10.3897/zookeys.938.51614.figures8-10" httpUri="https://binary.pensoft.net/fig/418210" pageId="0" pageNumber="87">, 8</figureCitation>
<figureCitation id="286044E5A42583FCB1D61E79F9A1A484" captionStart="Figures 810" captionStartId="F3" captionText="Figures 8 - 10. Triarthron itoi Hoshina sp. nov. (8, 9). 8 Aedeagus, dorsal view 9 ditto, lateral view. T. maerkelii Maerkel (10) aedeagus, dorsal view. Scale bar: 0.5 mm (8 - 10)." figureDoi="10.3897/zookeys.938.51614.figures8-10" httpUri="https://binary.pensoft.net/fig/418210" pageId="0" pageNumber="87">, 9</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="BF8C33C99199D07CC6F7EB531E821604" pageId="0" pageNumber="87" type="type locality">
<paragraph id="48725FAF3F9E3D1C884B91624DC24317" pageId="0" pageNumber="87">Type locality.</paragraph>
<paragraph id="0972AEA525108092629A5D9EBE7DE863" pageId="0" pageNumber="87">
Japan, Honshu: Nara Prefecture, Nara City, Nara Park,
<geoCoordinate id="5CB1D889913E9B2D4535C71A12AAC6BB" degrees="34" direction="north" minutes="41" orientation="latitude" precision="15" seconds="4" value="34.684444">34°41'4&quot;N</geoCoordinate>
,
<geoCoordinate id="BAD2995EB5C0BE9C7E74ADA1E5C5EF2A" degrees="135" direction="east" minutes="50" orientation="longitude" precision="15" seconds="36" value="135.84332">135°50'36&quot;E</geoCoordinate>
(DMS).
</paragraph>
</subSubSection>
<subSubSection id="7FC98CDFE2B393231469AC5574FB43B3" pageId="0" pageNumber="87" type="materials_examined">
<paragraph id="03FE0A4E9A6B0179A14F0B1D1897FDD7" pageId="0" pageNumber="87">Material examined.</paragraph>
<paragraph id="7945428997EEB91E47A6F440C851CB96" pageId="0" pageNumber="87">
<emphasis id="393DEC4400722B9993C958777B8FE714" bold="true" italics="true" pageId="0" pageNumber="87">Holotype</emphasis>
, ♂ (OSAKA. Type No.: OMNH TI 528): Japan, Honshu, Nara Prefecture, Nara City, Nara Park,
<geoCoordinate id="D2550C47CB777A67D9E6AA32D8E97E04" degrees="34" direction="north" minutes="41" orientation="latitude" precision="15" seconds="4" value="34.684444">34°41'4&quot;N</geoCoordinate>
,
<geoCoordinate id="B12142087F87F89F5AE969D11B49A2D7" degrees="135" direction="east" minutes="50" orientation="longitude" precision="15" seconds="36" value="135.84332">135°50'36&quot;E</geoCoordinate>
(DMS), 16.xii.2018, F.
<normalizedToken id="FFE85A6C764299A25DDFCDB431CDA73F" originalValue="Itô">Ito</normalizedToken>
leg. (OSAKA).
</paragraph>
</subSubSection>
<subSubSection id="270D215C4BEFFB99DCA50F4087DC87A7" pageId="0" pageNumber="87" type="materials_examined">
<paragraph id="F8D751CB0120FE40F91F356CD1F12518" pageId="0" pageNumber="87">Specimens examined of related species.</paragraph>
<paragraph id="41686204848AC7462CEFEABB04C0254A" pageId="0" pageNumber="87">
<taxonomicName id="36A96C5EE796CF4E1817EEAABF53E886" class="Insecta" family="Leiodidae" genus="Triarthron" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Triarthron maerkelii" order="Coleoptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="maerkelii">
<emphasis id="9239EC98D4543BCCB423E9E95FAE89AF" italics="true" pageId="0" pageNumber="87">Triarthron maerkelii</emphasis>
</taxonomicName>
<normalizedToken id="9AE1B7FE31701AC9D02F37E6C973B6D5" originalValue="Märkel">Maerkel</normalizedToken>
, 1840. 3 ♂♂, 1 ♀ (EUMJ), Japan - Honshu, Gunma Prefecture, Tsumagoi Village, Mt. Asashiki, 24.vii.1979, K.
<normalizedToken id="B6AA252A05813CD5FF4726055A52B7A2" originalValue="Itô">Ito</normalizedToken>
leg.; 1 ♂ (FU), Japan - Hokkaido, Ebetsu City, Nopporo Forest Park, 29.vi.2000, S. Hori leg.; 1 ♂ (FU), Sweden, SM.
<normalizedToken id="56383EF9AD62F7C63EBE208ADCB86F89" originalValue="Hornsö">Hornsoe</normalizedToken>
, 27.vi.1998, B. Andersson leg.
</paragraph>
</subSubSection>
<subSubSection id="0F7BF0DCA0222A6095768569F4150441" pageId="0" pageNumber="87" type="diagnosis">
<paragraph id="9641243E548955725F28A895D61694A5" pageId="0" pageNumber="87">Diagnosis.</paragraph>
<paragraph id="C82CF3BDAC9928406A7C37BC0873464B" pageId="0" pageNumber="87">Body length about 5 mm. Dorsum is almost concolorous, brown. Both mandibles were sharply curved inwardly at about apical 1/4. Mesofemur bearing five small teeth at dorsal lamina of posterior margin. Metafemur strongly expanded anteriorly at about half of the antero-apical margin and bearing a relatively long tooth and two tiny teeth at the dorsal lamina of the posterior margin. The median lobe of aedeagus weakly curved at lateral margins in dorsal view.</paragraph>
</subSubSection>
<subSubSection id="8FF8A7C981E89E8FA986F1123C4660A6" pageId="0" pageNumber="87" type="description">
<paragraph id="71860E4CC874D5FA62F923A2BA376073" pageId="0" pageNumber="87">Description.</paragraph>
<paragraph id="BD9D7E74BF7B7BC7A0AC419BAD3B87B9" pageId="0" pageNumber="87">
<emphasis id="A2A2E3BA6CE8EED6D14B65847F0F5E02" bold="true" italics="true" pageId="0" pageNumber="87">Measurement of holotype</emphasis>
. Body 4.8 mm in length; head 1.1 mm in length (from the front margin of the clypeus to base) and 1.3 mm in width; pronotum 1.2 mm in length and 2.0 mm in width; elytra 2.8 mm in length and 2.2 mm in width.
</paragraph>
<paragraph id="DA4129D60EDC11FC64E348A3BCAD9318" pageId="0" pageNumber="87">
<emphasis id="22804B8AD51A28407517E6498115167F" bold="true" italics="true" pageId="0" pageNumber="87">Coloration.</emphasis>
Dorsum shining and almost concolorous, brown (Fig.
<figureCitation id="3ECB94D16F2E21FC98BA687BC0D476BB" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitat of Triarthron itoi Hoshina, sp. nov. Scale bar: 1 mm." figureDoi="10.3897/zookeys.938.51614.figure1" httpUri="https://binary.pensoft.net/fig/418208" pageId="0" pageNumber="87">1</figureCitation>
); clypeus and labrum light brown; antennae brown and terminal three antennomeres slightly lighter than others; legs brown in general, but all trochanters and about basal 2/5 of metafemora blackish brown; mesoventrite, metaventrite, and abdominal ventrites light brown.
</paragraph>
<caption id="E204EE74BBF2DDA2C37AC58E57C3FB79" doi="10.3897/zookeys.938.51614.figure1" httpUri="https://binary.pensoft.net/fig/418208" pageId="0" pageNumber="87" start="Figure 1" startId="F1">
<paragraph id="289A7068839B8B188151462A33FE4BC1" pageId="0" pageNumber="87">
<emphasis id="162F19373D2AD4F6E3CE7A590FFDF8C6" bold="true" pageId="0" pageNumber="87">Figure 1.</emphasis>
Habitat of
<taxonomicName id="8DA02EFAB14C1AB4C8FB42D782ECDC40" authorityName="Hoshina" authorityYear="2020" class="Insecta" family="Leiodidae" genus="Triarthron" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Triarthron itoi" order="Coleoptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="itoi">
<emphasis id="688E0FB0D4686588ED86AD9F7491D023" italics="true" pageId="0" pageNumber="87">Triarthron itoi</emphasis>
</taxonomicName>
Hoshina, sp. nov. Scale bar: 1 mm.
</paragraph>
</caption>
<paragraph id="09BDB1AC9D84589368F392CD2DF094EA" pageId="0" pageNumber="87">
<emphasis id="B69C1BA5E58C734AE8F4C25C1FB8092C" bold="true" italics="true" pageId="0" pageNumber="87">Body</emphasis>
approximately 2.2 times as long as wide.
</paragraph>
<paragraph id="8A3FCC7380720725FC1941FA1DCCFE8A" pageId="0" pageNumber="87">
<emphasis id="C97FD854EE761344882A14333B72D3E9" bold="true" italics="true" pageId="0" pageNumber="87">Head</emphasis>
almost smooth, minutely, and densely punctate (Fig.
<figureCitation id="E1C0717C29C93C73409C437F79F4F767" captionStart="Figures 27" captionStartId="F2" captionText="Figures 2 - 7. Triarthron itoi Hoshina, sp. nov. (2 - 5). 2 Body 3 right antennomere, dorsal view 4 left mesofemur, ventral view 5 left metafemur, ventral view. T. maerkelii Maerkel (6, 7). 6 Left mesofemur, ventral view 7 left metafemur, ventral view. Scale bar: A: 1 mm (2), B: 0.2 mm (3), C, D: 0.5 mm (4 - 7)." figureDoi="10.3897/zookeys.938.51614.figures2-7" httpUri="https://binary.pensoft.net/fig/418209" pageId="0" pageNumber="87">2</figureCitation>
), and bearing a few short and very fine setae near each eye and a few long and fine setae near each lateral-basal corner of the clypeus; both mandibles sharply curved inwardly at about apical 1/4 and lacking large teeth at internal margins; antennomeres 1-3 longer than wide; antennomeres 4 and 11 about as long as wide; remaining antennomeres wider than long (Fig.
<figureCitation id="FAAE1ABE8FDB023EBEB3D0139F5FAFFB" captionStart="Figures 27" captionStartId="F2" captionText="Figures 2 - 7. Triarthron itoi Hoshina, sp. nov. (2 - 5). 2 Body 3 right antennomere, dorsal view 4 left mesofemur, ventral view 5 left metafemur, ventral view. T. maerkelii Maerkel (6, 7). 6 Left mesofemur, ventral view 7 left metafemur, ventral view. Scale bar: A: 1 mm (2), B: 0.2 mm (3), C, D: 0.5 mm (4 - 7)." figureDoi="10.3897/zookeys.938.51614.figures2-7" httpUri="https://binary.pensoft.net/fig/418209" pageId="0" pageNumber="87">3</figureCitation>
).
</paragraph>
<paragraph id="6070E31CF823FBA727B8B8D086A04D46" pageId="0" pageNumber="87">
<emphasis id="866432B2E63C0007612DE2527041CA7E" bold="true" italics="true" pageId="0" pageNumber="87">Pronotum</emphasis>
almost smooth and glabrous, widest at about basal 2/5 of lateral margins, minutely and densely punctate as head (Fig.
<figureCitation id="781820F66914C01A867C02CA6AD5F5B1" captionStart="Figures 27" captionStartId="F2" captionText="Figures 2 - 7. Triarthron itoi Hoshina, sp. nov. (2 - 5). 2 Body 3 right antennomere, dorsal view 4 left mesofemur, ventral view 5 left metafemur, ventral view. T. maerkelii Maerkel (6, 7). 6 Left mesofemur, ventral view 7 left metafemur, ventral view. Scale bar: A: 1 mm (2), B: 0.2 mm (3), C, D: 0.5 mm (4 - 7)." figureDoi="10.3897/zookeys.938.51614.figures2-7" httpUri="https://binary.pensoft.net/fig/418209" pageId="0" pageNumber="87">2</figureCitation>
), and with a transverse fine groove along the basal margin, which is interrupted at the central part (Fig.
<figureCitation id="2074E73334CFBAD56F0C716CED1D57B8" captionStart="Figures 27" captionStartId="F2" captionText="Figures 2 - 7. Triarthron itoi Hoshina, sp. nov. (2 - 5). 2 Body 3 right antennomere, dorsal view 4 left mesofemur, ventral view 5 left metafemur, ventral view. T. maerkelii Maerkel (6, 7). 6 Left mesofemur, ventral view 7 left metafemur, ventral view. Scale bar: A: 1 mm (2), B: 0.2 mm (3), C, D: 0.5 mm (4 - 7)." figureDoi="10.3897/zookeys.938.51614.figures2-7" httpUri="https://binary.pensoft.net/fig/418209" pageId="0" pageNumber="87">2</figureCitation>
).
</paragraph>
<paragraph id="95D4968540F1C3FFA6D6334E9BA93875" pageId="0" pageNumber="87">
<emphasis id="7F42E08124CCFA04D35F3912C8517F91" bold="true" italics="true" pageId="0" pageNumber="87">Scutellum</emphasis>
almost smooth and distinctly punctate (Fig.
<figureCitation id="7193B452218A6B1300EDD883E6A98ECA" captionStart="Figures 27" captionStartId="F2" captionText="Figures 2 - 7. Triarthron itoi Hoshina, sp. nov. (2 - 5). 2 Body 3 right antennomere, dorsal view 4 left mesofemur, ventral view 5 left metafemur, ventral view. T. maerkelii Maerkel (6, 7). 6 Left mesofemur, ventral view 7 left metafemur, ventral view. Scale bar: A: 1 mm (2), B: 0.2 mm (3), C, D: 0.5 mm (4 - 7)." figureDoi="10.3897/zookeys.938.51614.figures2-7" httpUri="https://binary.pensoft.net/fig/418209" pageId="0" pageNumber="87">2</figureCitation>
).
</paragraph>
<paragraph id="A1BA44781B153859C60FFF9F51DCF5FA" pageId="0" pageNumber="87">
<emphasis id="E2AC2175D327108542BAD79A88311DC6" bold="true" italics="true" pageId="0" pageNumber="87">Elytra</emphasis>
almost smooth and glabrous except for very sparse and fine setae along lateral margins, widest at about basal 1/3 of lateral margins (Fig.
<figureCitation id="C3D5AA8FF4937EF4F0F6533BF82FCA1C" captionStart="Figures 27" captionStartId="F2" captionText="Figures 2 - 7. Triarthron itoi Hoshina, sp. nov. (2 - 5). 2 Body 3 right antennomere, dorsal view 4 left mesofemur, ventral view 5 left metafemur, ventral view. T. maerkelii Maerkel (6, 7). 6 Left mesofemur, ventral view 7 left metafemur, ventral view. Scale bar: A: 1 mm (2), B: 0.2 mm (3), C, D: 0.5 mm (4 - 7)." figureDoi="10.3897/zookeys.938.51614.figures2-7" httpUri="https://binary.pensoft.net/fig/418209" pageId="0" pageNumber="87">2</figureCitation>
); each elytron bearing nine rows of punctures and ninth row present along lateral-downside margins and invisible in dorsal view (Fig.
<figureCitation id="9B0CBB4ECA4A91B6728C3A4B7CFD1814" captionStart="Figures 27" captionStartId="F2" captionText="Figures 2 - 7. Triarthron itoi Hoshina, sp. nov. (2 - 5). 2 Body 3 right antennomere, dorsal view 4 left mesofemur, ventral view 5 left metafemur, ventral view. T. maerkelii Maerkel (6, 7). 6 Left mesofemur, ventral view 7 left metafemur, ventral view. Scale bar: A: 1 mm (2), B: 0.2 mm (3), C, D: 0.5 mm (4 - 7)." figureDoi="10.3897/zookeys.938.51614.figures2-7" httpUri="https://binary.pensoft.net/fig/418209" pageId="0" pageNumber="87">2</figureCitation>
); punctures comprising nine rows of punctures distinct and larger than those of head and pronotum (Fig.
<figureCitation id="C54F092EC0133162526FFBA876F0DF4E" captionStart="Figures 27" captionStartId="F2" captionText="Figures 2 - 7. Triarthron itoi Hoshina, sp. nov. (2 - 5). 2 Body 3 right antennomere, dorsal view 4 left mesofemur, ventral view 5 left metafemur, ventral view. T. maerkelii Maerkel (6, 7). 6 Left mesofemur, ventral view 7 left metafemur, ventral view. Scale bar: A: 1 mm (2), B: 0.2 mm (3), C, D: 0.5 mm (4 - 7)." figureDoi="10.3897/zookeys.938.51614.figures2-7" httpUri="https://binary.pensoft.net/fig/418209" pageId="0" pageNumber="87">2</figureCitation>
); punctures between rows of punctures dense and minute (Fig.
<figureCitation id="29F5CA3F50F7EC4012C6B67F281D048A" captionStart="Figures 27" captionStartId="F2" captionText="Figures 2 - 7. Triarthron itoi Hoshina, sp. nov. (2 - 5). 2 Body 3 right antennomere, dorsal view 4 left mesofemur, ventral view 5 left metafemur, ventral view. T. maerkelii Maerkel (6, 7). 6 Left mesofemur, ventral view 7 left metafemur, ventral view. Scale bar: A: 1 mm (2), B: 0.2 mm (3), C, D: 0.5 mm (4 - 7)." figureDoi="10.3897/zookeys.938.51614.figures2-7" httpUri="https://binary.pensoft.net/fig/418209" pageId="0" pageNumber="87">2</figureCitation>
).
</paragraph>
<paragraph id="761CE130D0C6E17CFE12935773CDDDA5" pageId="0" pageNumber="87">
<emphasis id="2499DE577C33351B6FB1F99503722696" bold="true" italics="true" pageId="0" pageNumber="87">Hind wings</emphasis>
fully developed.
</paragraph>
<paragraph id="9E427419ECCC56966CF592F75DC69A8F" pageId="0" pageNumber="87">
<emphasis id="31F81E7AF609CF4391F1FA2405E2A2F3" bold="true" italics="true" pageId="0" pageNumber="87">Mesoventrite</emphasis>
weakly microreticulate and almost glabrous; metaventrite and abdominal ventrites distinctly microreticulate, and densely and finely pubescent.
</paragraph>
<paragraph id="7679E5221BC5DE16EB831A23B5E76A18" pageId="0" pageNumber="87">
<emphasis id="B15A6730343BBFEBE2CDF28FE9FBF623" bold="true" italics="true" pageId="0" pageNumber="87">Legs</emphasis>
with many small spines as other species of the genus
<taxonomicName id="C6B42A05BFCE5B4C8F2715694729BAFF" class="Insecta" family="Leiodidae" genus="Triarthron" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Triarthron" order="Coleoptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="genus">
<emphasis id="F06D5E6769A6DD3CD96EEB82EFD66C3F" italics="true" pageId="0" pageNumber="87">Triarthron</emphasis>
</taxonomicName>
; mesofemur approximately 2.8 times as wide as long, weakly expanded anteriorly at about half of antero-apical margin, and bearing five small teeth at dorsal lamina of posterior margin (Fig.
<figureCitation id="3C58B717B9EAFC90AA7FA780CB52FBA6" captionStart="Figures 27" captionStartId="F2" captionText="Figures 2 - 7. Triarthron itoi Hoshina, sp. nov. (2 - 5). 2 Body 3 right antennomere, dorsal view 4 left mesofemur, ventral view 5 left metafemur, ventral view. T. maerkelii Maerkel (6, 7). 6 Left mesofemur, ventral view 7 left metafemur, ventral view. Scale bar: A: 1 mm (2), B: 0.2 mm (3), C, D: 0.5 mm (4 - 7)." figureDoi="10.3897/zookeys.938.51614.figures2-7" httpUri="https://binary.pensoft.net/fig/418209" pageId="0" pageNumber="87">4</figureCitation>
); mesotibia weakly curved inwardly; metafemur approximately 2.8 times as wide as long, strongly expanded anteriorly at about half of antero-apical margin, and bearing a relatively long tooth and two tiny ones at dorsal lamina of posterior margin (Fig.
<figureCitation id="806E8E6AFA6BD1566EE188ADB75FA9AD" captionStart="Figures 27" captionStartId="F2" captionText="Figures 2 - 7. Triarthron itoi Hoshina, sp. nov. (2 - 5). 2 Body 3 right antennomere, dorsal view 4 left mesofemur, ventral view 5 left metafemur, ventral view. T. maerkelii Maerkel (6, 7). 6 Left mesofemur, ventral view 7 left metafemur, ventral view. Scale bar: A: 1 mm (2), B: 0.2 mm (3), C, D: 0.5 mm (4 - 7)." figureDoi="10.3897/zookeys.938.51614.figures2-7" httpUri="https://binary.pensoft.net/fig/418209" pageId="0" pageNumber="87">5</figureCitation>
); metatibia almost straight.
</paragraph>
<paragraph id="C5470A9E01C37BE1BAA101CB77799506" pageId="0" pageNumber="87">
<emphasis id="5E2398F2B1F88AFB00B0D8276AFB27FB" bold="true" italics="true" pageId="0" pageNumber="87">Aedeagus</emphasis>
slender in general (Figs
<figureCitation id="9F5205BAF0B0E1FB892822EE0701CC18" captionStart="Figures 810" captionStartId="F3" captionText="Figures 8 - 10. Triarthron itoi Hoshina sp. nov. (8, 9). 8 Aedeagus, dorsal view 9 ditto, lateral view. T. maerkelii Maerkel (10) aedeagus, dorsal view. Scale bar: 0.5 mm (8 - 10)." figureDoi="10.3897/zookeys.938.51614.figures8-10" httpUri="https://binary.pensoft.net/fig/418210" pageId="0" pageNumber="87">8</figureCitation>
,
<figureCitation id="DCB9CDBD2012D1C8E47025A76C69A72B" captionStart="Figures 810" captionStartId="F3" captionText="Figures 8 - 10. Triarthron itoi Hoshina sp. nov. (8, 9). 8 Aedeagus, dorsal view 9 ditto, lateral view. T. maerkelii Maerkel (10) aedeagus, dorsal view. Scale bar: 0.5 mm (8 - 10)." figureDoi="10.3897/zookeys.938.51614.figures8-10" httpUri="https://binary.pensoft.net/fig/418210" pageId="0" pageNumber="87">9</figureCitation>
); median lobe weakly and simply curved at lateral margins, round at apex in dorsal view (Fig.
<figureCitation id="A078AF0428B86F45E7DF9B62445CE16C" captionStart="Figures 810" captionStartId="F3" captionText="Figures 8 - 10. Triarthron itoi Hoshina sp. nov. (8, 9). 8 Aedeagus, dorsal view 9 ditto, lateral view. T. maerkelii Maerkel (10) aedeagus, dorsal view. Scale bar: 0.5 mm (8 - 10)." figureDoi="10.3897/zookeys.938.51614.figures8-10" httpUri="https://binary.pensoft.net/fig/418210" pageId="0" pageNumber="87">8</figureCitation>
) and almost straight in lateral view (Fig.
<figureCitation id="72DB8AD8F6940D1246E42BBB1AB9676D" captionStart="Figures 810" captionStartId="F3" captionText="Figures 8 - 10. Triarthron itoi Hoshina sp. nov. (8, 9). 8 Aedeagus, dorsal view 9 ditto, lateral view. T. maerkelii Maerkel (10) aedeagus, dorsal view. Scale bar: 0.5 mm (8 - 10)." figureDoi="10.3897/zookeys.938.51614.figures8-10" httpUri="https://binary.pensoft.net/fig/418210" pageId="0" pageNumber="87">9</figureCitation>
); both parameres almost symmetrical and round at apex (Fig.
<figureCitation id="21A1936F9D0A74D03513F8B696AE15FE" captionStart="Figures 810" captionStartId="F3" captionText="Figures 8 - 10. Triarthron itoi Hoshina sp. nov. (8, 9). 8 Aedeagus, dorsal view 9 ditto, lateral view. T. maerkelii Maerkel (10) aedeagus, dorsal view. Scale bar: 0.5 mm (8 - 10)." figureDoi="10.3897/zookeys.938.51614.figures8-10" httpUri="https://binary.pensoft.net/fig/418210" pageId="0" pageNumber="87">8</figureCitation>
) and almost straight in lateral view (Fig.
<figureCitation id="0F7BE6C332EC6B00F95E9C8C515A5F0F" captionStart="Figures 810" captionStartId="F3" captionText="Figures 8 - 10. Triarthron itoi Hoshina sp. nov. (8, 9). 8 Aedeagus, dorsal view 9 ditto, lateral view. T. maerkelii Maerkel (10) aedeagus, dorsal view. Scale bar: 0.5 mm (8 - 10)." figureDoi="10.3897/zookeys.938.51614.figures8-10" httpUri="https://binary.pensoft.net/fig/418210" pageId="0" pageNumber="87">9</figureCitation>
); each paramere bearing three apical setae (Figs
<figureCitation id="951425487B4976759A75A2980E07EDC9" captionStart="Figures 810" captionStartId="F3" captionText="Figures 8 - 10. Triarthron itoi Hoshina sp. nov. (8, 9). 8 Aedeagus, dorsal view 9 ditto, lateral view. T. maerkelii Maerkel (10) aedeagus, dorsal view. Scale bar: 0.5 mm (8 - 10)." figureDoi="10.3897/zookeys.938.51614.figures8-10" httpUri="https://binary.pensoft.net/fig/418210" pageId="0" pageNumber="87">8</figureCitation>
,
<figureCitation id="384716210262D440D8A1D41F03C09EF0" captionStart="Figures 810" captionStartId="F3" captionText="Figures 8 - 10. Triarthron itoi Hoshina sp. nov. (8, 9). 8 Aedeagus, dorsal view 9 ditto, lateral view. T. maerkelii Maerkel (10) aedeagus, dorsal view. Scale bar: 0.5 mm (8 - 10)." figureDoi="10.3897/zookeys.938.51614.figures8-10" httpUri="https://binary.pensoft.net/fig/418210" pageId="0" pageNumber="87">9</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B7564A995109FB2B368655E0575200BB" pageId="0" pageNumber="87" type="etymology">
<paragraph id="073E5AF6FC319436C77B7FA6FC00F3B4" pageId="0" pageNumber="87">Etymology.</paragraph>
<paragraph id="6D84C700A18957AAD7908B3EF073F45A" pageId="0" pageNumber="87">
The specific name is dedicated to Mr. Fukuo
<normalizedToken id="278EAEEB6D69BF8E9D936B36F964A1FB" originalValue="Itô">Ito</normalizedToken>
, the collector of the holotype.
</paragraph>
</subSubSection>
<subSubSection id="CF10BCBF5554D0AC465EB49534AF0B8F" pageId="0" pageNumber="87" type="distribution">
<paragraph id="26F9084EA0E2A36F9B2B28E9E51E6207" pageId="0" pageNumber="87">Distribution.</paragraph>
<paragraph id="D95037D1F45794686E192F369DEEF945" pageId="0" pageNumber="87">Japan: Honshu (Nara Prefecture).</paragraph>
</subSubSection>
<subSubSection id="3F518EE059AD3925BF58485FA95D1AE8" pageId="0" pageNumber="87" type="differential diagnosis">
<paragraph id="5B83BCF84A13EDA93A99A1C182ADA540" pageId="0" pageNumber="87">Differential diagnosis.</paragraph>
<paragraph id="1DB86AC4C21963713233191AE6D07A79" pageId="0" pageNumber="87">
Collecting the tribe
<taxonomicName id="FFF49A591459E18452C43CFF0320FDD4" authorityName="Lopatin" authorityYear="1961" lsidName="" pageId="0" pageNumber="87" rank="tribe" tribe="Sogdini">Sogdini</taxonomicName>
is generally not easy in Japan and identified Japanese specimens of the tribe are very small in quantity.
<taxonomicName id="1485B95C872DDBD9690E9DC0DA016D0A" authorityName="Hoshina" authorityYear="2020" class="Insecta" family="Leiodidae" genus="Triarthron" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Triarthron itoi" order="Coleoptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="itoi">
<emphasis id="9F6F26A16FA13529BA6ADB65596DF7A1" italics="true" pageId="0" pageNumber="87">Triarthron itoi</emphasis>
</taxonomicName>
Hoshina, sp. nov. is described based on only one specimen; therefore, the degree of individual variation cannot be determined in this species. However, I found some morphological features on that specimen that are clearly different from two known species of
<taxonomicName id="CA2A3DB7783E94840D22965ACC6B9F97" class="Insecta" family="Leiodidae" genus="Triarthron" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Triarthron" order="Coleoptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="genus">
<emphasis id="F300ABF41EC0A9E791FBD84FFD3E3C3C" italics="true" pageId="0" pageNumber="87">Triarthron</emphasis>
</taxonomicName>
, and recognize it as a new member of the genus.
<taxonomicName id="B299435F0C5DDA808601F3539A0CBDC6" authorityName="Hoshina" authorityYear="2020" class="Insecta" family="Leiodidae" genus="Triarthron" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Triarthron itoi" order="Coleoptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="itoi">
<emphasis id="71808C262A6680205175D0977868FFDB" italics="true" pageId="0" pageNumber="87">Triarthron itoi</emphasis>
</taxonomicName>
sp. nov. can be distinguished from
<taxonomicName id="A1AD1A5858FAAD9DCA3879349F730465" lsidName="T. maerkelii" pageId="0" pageNumber="87" rank="species" species="maerkelii">
<emphasis id="BBAA829575192D9898F8CD415A1CD84D" italics="true" pageId="0" pageNumber="87">T. maerkelii</emphasis>
</taxonomicName>
<normalizedToken id="BCE96C31BFDC3BFB23F3A255B647DA17" originalValue="Märkel">Maerkel</normalizedToken>
, 1840 by the following features: it has a large body whose length is 4.8 mm, mesofemur with relatively large teeth at the dorsal lamina of the posterior margin (Fig.
<figureCitation id="588F105C1BCE195EC047AED0B7F30B4F" captionStart="Figures 27" captionStartId="F2" captionText="Figures 2 - 7. Triarthron itoi Hoshina, sp. nov. (2 - 5). 2 Body 3 right antennomere, dorsal view 4 left mesofemur, ventral view 5 left metafemur, ventral view. T. maerkelii Maerkel (6, 7). 6 Left mesofemur, ventral view 7 left metafemur, ventral view. Scale bar: A: 1 mm (2), B: 0.2 mm (3), C, D: 0.5 mm (4 - 7)." figureDoi="10.3897/zookeys.938.51614.figures2-7" httpUri="https://binary.pensoft.net/fig/418209" pageId="0" pageNumber="87">4</figureCitation>
), metafemur strongly expanded anteriorly at about half of the antero-apical margin (Fig.
<figureCitation id="C29A592E9D17E7113CDB979DE9C24AEF" captionStart="Figures 27" captionStartId="F2" captionText="Figures 2 - 7. Triarthron itoi Hoshina, sp. nov. (2 - 5). 2 Body 3 right antennomere, dorsal view 4 left mesofemur, ventral view 5 left metafemur, ventral view. T. maerkelii Maerkel (6, 7). 6 Left mesofemur, ventral view 7 left metafemur, ventral view. Scale bar: A: 1 mm (2), B: 0.2 mm (3), C, D: 0.5 mm (4 - 7)." figureDoi="10.3897/zookeys.938.51614.figures2-7" httpUri="https://binary.pensoft.net/fig/418209" pageId="0" pageNumber="87">5</figureCitation>
), and median lobe of aedeagus weakly and simply curved at lateral margins in dorsal view (Fig.
<figureCitation id="4011B9D57B29CAE11E8BA606B95503BC" captionStart="Figures 810" captionStartId="F3" captionText="Figures 8 - 10. Triarthron itoi Hoshina sp. nov. (8, 9). 8 Aedeagus, dorsal view 9 ditto, lateral view. T. maerkelii Maerkel (10) aedeagus, dorsal view. Scale bar: 0.5 mm (8 - 10)." figureDoi="10.3897/zookeys.938.51614.figures8-10" httpUri="https://binary.pensoft.net/fig/418210" pageId="0" pageNumber="87">8</figureCitation>
). In contrast,
<taxonomicName id="B336D6D7DE4D5FFC237859732FF7FFCA" lsidName="T. maerkelii" pageId="0" pageNumber="87" rank="species" species="maerkelii">
<emphasis id="3C461F22A2DD0426304AF568FC3C50CF" italics="true" pageId="0" pageNumber="87">T. maerkelii</emphasis>
</taxonomicName>
differs in the following ways: it has a relatively small body whose length is 2.5-3.8 mm (
<bibRefCitation id="61D4C11CA7F513E2B9BCE2AE0B9CA4FA" author="Daffner, H" journalOrPublisher="Folia Entomologica Hungarica" pageId="0" pageNumber="87" pagination="9 - 163" refId="B2" refString="Daffner, H, 1983. Revision der palaerktischen Arten der Tribus Leiodini Leach (Coleoptera, Leiodidae). Folia Entomologica Hungarica 44: 9 - 163" title="Revision der palaerktischen Arten der Tribus Leiodini Leach (Coleoptera, Leiodidae)." volume="44" year="1983">Daffner 1983</bibRefCitation>
), mesofemur with teeth less-visible because of being hidden by ventral side of mesofemur (Fig.
<figureCitation id="DDD648D36FDEEA4B803C0ACE0F01DD32" captionStart="Figures 27" captionStartId="F2" captionText="Figures 2 - 7. Triarthron itoi Hoshina, sp. nov. (2 - 5). 2 Body 3 right antennomere, dorsal view 4 left mesofemur, ventral view 5 left metafemur, ventral view. T. maerkelii Maerkel (6, 7). 6 Left mesofemur, ventral view 7 left metafemur, ventral view. Scale bar: A: 1 mm (2), B: 0.2 mm (3), C, D: 0.5 mm (4 - 7)." figureDoi="10.3897/zookeys.938.51614.figures2-7" httpUri="https://binary.pensoft.net/fig/418209" pageId="0" pageNumber="87">6</figureCitation>
), metafemur relatively weakly expanded anteriorly at about half of antero-apical margin (Fig.
<figureCitation id="2025E3D1FB461BE742AC79D66F25FF1C" captionStart="Figures 27" captionStartId="F2" captionText="Figures 2 - 7. Triarthron itoi Hoshina, sp. nov. (2 - 5). 2 Body 3 right antennomere, dorsal view 4 left mesofemur, ventral view 5 left metafemur, ventral view. T. maerkelii Maerkel (6, 7). 6 Left mesofemur, ventral view 7 left metafemur, ventral view. Scale bar: A: 1 mm (2), B: 0.2 mm (3), C, D: 0.5 mm (4 - 7)." figureDoi="10.3897/zookeys.938.51614.figures2-7" httpUri="https://binary.pensoft.net/fig/418209" pageId="0" pageNumber="87">7</figureCitation>
), and median lobe sharply narrowed from about apical 2/5 towards apex (Fig.
<figureCitation id="07863124F63179C52E6D03F2630E3561" captionStart="Figures 810" captionStartId="F3" captionText="Figures 8 - 10. Triarthron itoi Hoshina sp. nov. (8, 9). 8 Aedeagus, dorsal view 9 ditto, lateral view. T. maerkelii Maerkel (10) aedeagus, dorsal view. Scale bar: 0.5 mm (8 - 10)." figureDoi="10.3897/zookeys.938.51614.figures8-10" httpUri="https://binary.pensoft.net/fig/418210" pageId="0" pageNumber="87">10</figureCitation>
).
</paragraph>
<caption id="6F6F61225F700354D31DA7AA51835D89" doi="10.3897/zookeys.938.51614.figures2-7" httpUri="https://binary.pensoft.net/fig/418209" pageId="0" pageNumber="87" start="Figures 27" startId="F2">
<paragraph id="8A2CBFA3FCAE53A2728C946A84E901A5" pageId="0" pageNumber="87">
<emphasis id="98BEB6DBE40DBCF5C419B1F629C17D32" bold="true" pageId="0" pageNumber="87">Figures 2-7.</emphasis>
<taxonomicName id="3FE5B3859AF5CFABBD44988BEACF0767" authorityName="Hoshina" authorityYear="2020" class="Insecta" family="Leiodidae" genus="Triarthron" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Triarthron itoi" order="Coleoptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="itoi">
<emphasis id="E99F93058543C100FF2DB9B57702C0A1" italics="true" pageId="0" pageNumber="87">Triarthron itoi</emphasis>
</taxonomicName>
Hoshina, sp. nov. (
<emphasis id="B09C58EAB8D91B82C52927FEE450726A" bold="true" pageId="0" pageNumber="87">2-5</emphasis>
).
<emphasis id="07B783F7CF7661E2C075B985D63DE894" bold="true" pageId="0" pageNumber="87">2</emphasis>
Body
<emphasis id="4021B66D5AAF2AF639F01400207496E6" bold="true" pageId="0" pageNumber="87">3</emphasis>
right antennomere, dorsal view
<emphasis id="112B96B1C17E8D91136F09AF9C952A80" bold="true" pageId="0" pageNumber="87">4</emphasis>
left mesofemur, ventral view
<emphasis id="4DCA5CF9D6188099E3B44EF2602F91FC" bold="true" pageId="0" pageNumber="87">5</emphasis>
left metafemur, ventral view.
<taxonomicName id="B596AA390E0B5D834E323A1290E140CB" lsidName="T. maerkelii" pageId="0" pageNumber="87" rank="species" species="maerkelii">
<emphasis id="EBD197EAE57F20760DAEB2C536EEDFFA" italics="true" pageId="0" pageNumber="87">T. maerkelii</emphasis>
</taxonomicName>
<normalizedToken id="038AE8322DB485D50EC4510E90ACE825" originalValue="Märkel">Maerkel</normalizedToken>
(
<emphasis id="BC39185E0FF1C5794D760621A0F630A6" bold="true" pageId="0" pageNumber="87">6, 7)</emphasis>
.
<emphasis id="0CF676194A1D417996AB5B3BE955B262" bold="true" pageId="0" pageNumber="87">6</emphasis>
Left mesofemur, ventral view
<emphasis id="575E2DE0982D581A9D85E7F8EF0332D3" bold="true" pageId="0" pageNumber="87">7</emphasis>
left metafemur, ventral view. Scale bar: A: 1 mm (
<emphasis id="E4D94FA07BD158639E019474036EC045" bold="true" pageId="0" pageNumber="87">2</emphasis>
), B: 0.2 mm (
<emphasis id="8420C3B30F258F8C7048724F27472EA4" bold="true" pageId="0" pageNumber="87">3</emphasis>
), C, D: 0.5 mm (
<emphasis id="10392276B99E400FCF7D10754718EB95" bold="true" pageId="0" pageNumber="87">4-7</emphasis>
).
</paragraph>
</caption>
<paragraph id="1136F10F025AEFADD305BDF7701080CB" pageId="0" pageNumber="87">
Moreover,
<taxonomicName id="B43FD1FB13D81B76E48B8A4E4341D0C2" lsidName="T. itoi" pageId="0" pageNumber="87" rank="species" species="itoi">
<emphasis id="4F937432F54B467EE6DAE7E31F0F3ECE" italics="true" pageId="0" pageNumber="87">T. itoi</emphasis>
</taxonomicName>
sp. nov. can be separated from
<taxonomicName id="82316A4C5F14B6D0518EEDE149384B53" lsidName="T. lecontei" pageId="0" pageNumber="87" rank="species" species="lecontei">
<emphasis id="C9641C0506BEADC3F8E42523EF58BAC8" italics="true" pageId="0" pageNumber="87">T. lecontei</emphasis>
</taxonomicName>
Horn, 1868 by having both mandibles sharply curved inwardly at about apical 1/4 and lacking large teeth at internal margins, and metafemur strongly expanded at the anterior margin (Fig.
<figureCitation id="D7742E5AC1047996613FD71CEC116CD0" captionStart="Figures 27" captionStartId="F2" captionText="Figures 2 - 7. Triarthron itoi Hoshina, sp. nov. (2 - 5). 2 Body 3 right antennomere, dorsal view 4 left mesofemur, ventral view 5 left metafemur, ventral view. T. maerkelii Maerkel (6, 7). 6 Left mesofemur, ventral view 7 left metafemur, ventral view. Scale bar: A: 1 mm (2), B: 0.2 mm (3), C, D: 0.5 mm (4 - 7)." figureDoi="10.3897/zookeys.938.51614.figures2-7" httpUri="https://binary.pensoft.net/fig/418209" pageId="0" pageNumber="87">5</figureCitation>
). In contrast,
<taxonomicName id="C4A6B392E4BEE34FD7D6765828C5F100" lsidName="T. lecontei" pageId="0" pageNumber="87" rank="species" species="lecontei">
<emphasis id="FCCD29111F1CABA1577FA2FCE9FD33EF" italics="true" pageId="0" pageNumber="87">T. lecontei</emphasis>
</taxonomicName>
has both mandibles relatively weakly curved inwardly at internal margins, right one with an elongated sub-apical tooth, and metafemur almost straight at the anterior margin (
<bibRefCitation id="557F376009540501D4691037416CAE35" author="Hatch, MH" journalOrPublisher="University of Washington Press, Seattle, Washington" pageId="0" pageNumber="87" refId="B3" refString="Hatch, MH, 1957. The beetles of the Pacific Northwest. Part II: Staphyliniformia. University of Washington Publications in Biology 16. University of Washington Press, Seattle, Washington" title="The beetles of the Pacific Northwest. Part II: Staphyliniformia. University of Washington Publications in Biology 16." year="1957">Hatch 1957</bibRefCitation>
;
<bibRefCitation id="663040434BB679837CE3D0843A1384E5" DOI="https://doi.org/10.11646/zootaxa.2102.1.1" author="Peck, SB" journalOrPublisher="Zootaxa" pageId="0" pageNumber="87" pagination="1 - 74" refId="B16" refString="Peck, SB, Cook, J, 2009. Review of the Sogdini of North and Central America (Coleoptera: Leiodidae: Leiodinae) with descriptions of fourteen new species and three new genera. Zootaxa 2102: 1 - 74, DOI: https://doi.org/10.11646/zootaxa.2102.1.1" title="Review of the Sogdini of North and Central America (Coleoptera: Leiodidae: Leiodinae) with descriptions of fourteen new species and three new genera." url="https://doi.org/10.11646/zootaxa.2102.1.1" volume="2102" year="2009">Peck and Cook 2009</bibRefCitation>
).
</paragraph>
<caption id="4B764AC602BEEBB80FE21FB36A23E406" doi="10.3897/zookeys.938.51614.figures8-10" httpUri="https://binary.pensoft.net/fig/418210" pageId="0" pageNumber="87" start="Figures 810" startId="F3">
<paragraph id="8359CE29A778A6A86293EB61FF50D28C" pageId="0" pageNumber="87">
<emphasis id="DEA37344312D4CBC2E2B1842F1F52190" bold="true" pageId="0" pageNumber="87">Figures 8-10.</emphasis>
<taxonomicName id="845B93C2B5F50ED7EC3B281C5076C7B2" authorityName="Hoshina" authorityYear="2020" class="Insecta" family="Leiodidae" genus="Triarthron" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Triarthron itoi" order="Coleoptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="itoi">
<emphasis id="B37F38689AC5465C6A893F6521DFF925" italics="true" pageId="0" pageNumber="87">Triarthron itoi</emphasis>
</taxonomicName>
Hoshina sp. nov. (
<emphasis id="8FF6BE921203DF382F87FC556248C664" bold="true" pageId="0" pageNumber="87">8, 9</emphasis>
).
<emphasis id="8F6B3FE48E4D6FBE50D938152B6B6B65" bold="true" pageId="0" pageNumber="87">8</emphasis>
Aedeagus, dorsal view
<emphasis id="69439866ECE233502AB8B01E5F59EB7C" bold="true" pageId="0" pageNumber="87">9</emphasis>
ditto, lateral view.
<taxonomicName id="ACFBFA2B0525E03F3E2A5A2CEC7D4DEF" lsidName="T. maerkelii" pageId="0" pageNumber="87" rank="species" species="maerkelii">
<emphasis id="5978DDD7123288C495B27B9A8924E5A9" italics="true" pageId="0" pageNumber="87">T. maerkelii</emphasis>
</taxonomicName>
<normalizedToken id="35A5BCF59BB25F24461FAE74657CC480" originalValue="Märkel">Maerkel</normalizedToken>
(
<emphasis id="6BB8AA64ACB352DEF627CDB6036139FE" bold="true" pageId="0" pageNumber="87">10</emphasis>
) aedeagus, dorsal view. Scale bar: 0.5 mm (
<emphasis id="3890CB097CB04742AA89DFF16A89C0B9" bold="true" pageId="0" pageNumber="87">8-10</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="4976C1942BA1B803C8B53BEB62BAD5CD" pageId="0" pageNumber="87" type="natural history">
<paragraph id="7FD8C54278AA5A38B6DC47312A8FDA93" pageId="0" pageNumber="87">Natural history.</paragraph>
<paragraph id="DF3DA7ECA05101318FC7C6238FF6B91A" pageId="0" pageNumber="87">
Life history of
<taxonomicName id="138316CB321C65E8D49FA0DCB474FB7F" authorityName="Hoshina" authorityYear="2020" class="Insecta" family="Leiodidae" genus="Triarthron" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Triarthron itoi" order="Coleoptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="itoi">
<emphasis id="E0EDAE03B41EE79C8DBEA490361DAF60" italics="true" pageId="0" pageNumber="87">Triarthron itoi</emphasis>
</taxonomicName>
Hoshina, sp. nov. is not known.
</paragraph>
</subSubSection>
</treatment>
</document>