297 lines
31 KiB
XML
297 lines
31 KiB
XML
<document id="2DC03004A6E0A800D16A999BD7D7A435" ID-DOI="10.1163/18759866-BjA10025" ID-GBIF-Dataset="6e337125-efbe-40e5-8add-e40f426af30a" ID-Zenodo-Dep="6147303" approvedBy="jonas" checkinTime="1645204469354" checkinUser="jonas" docAuthor="Páll-Gergely, Barna, Jochum, Adrienne, J. Vermeulen, Jaap, Anker, Katja, Hunyadi, András, Örstan, Aydin, Szabó, Ábel, Dányi, László & Schilthuizen, Menno" docDate="2022" docId="35368F2DFFB8FF9752DCF9C7FC58FEE3" docLanguage="en" docName="ContribZool.91.62-78.pdf" docOrigin="Contributions To Zoology 91" docTitle="Angustopila coprologos Pall-Gergely, Jochum & Hunyadi 2022, n. sp." docType="treatment" docVersion="5" lastPageNumber="69" masterDocId="C90FF755FFBCFF905059FF80FFB9FF9A" masterDocTitle="The world’s tiniest land snails from Laos and Vietnam (Gastropoda, Pulmonata, Hypselostomatidae)" masterLastPageNumber="78" masterPageNumber="62" pageNumber="66" updateTime="1694674530954" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="267C42FBCDA0944EDA1FD18E901379A3">The world’s tiniest land snails from Laos and Vietnam (Gastropoda, Pulmonata, Hypselostomatidae)</mods:title>
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<mods:namePart id="5EDC588066FB431C8B807375495F48A5">Páll-Gergely, Barna</mods:namePart>
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<mods:affiliation id="8A674491B5B92C42F0FEABFDD02528D6">Plant Protection Institute, Centre for Agricultural Research, ELKH, Herman Ottó út 15, 1022 Budapest, Hungary</mods:affiliation>
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<mods:nameIdentifier id="D9E6DCF554AF0EC7609B5A8CC02D6455" type="email">pall-gergely.barna@atk.hu & pallgergely2@gmail.com</mods:nameIdentifier>
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<mods:namePart id="38A5C38E62741751EBB4E16CD84A6DFF">Jochum, Adrienne</mods:namePart>
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<mods:affiliation id="8BD6F3B18A2E33C1D17B30B75D7C5047">Naturhistorisches Museum der Burgergemeinde Bern, 3005 Bern, Switzerland & Institute of Ecology and Evolution, University of Bern, 3012 Bern, Switzerland & Senckenberg Forschungsinstitut und Naturmuseum, 60325 Frankfurt am Main, Germany</mods:affiliation>
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<mods:nameIdentifier id="A04C7F8594A817FAF66EDDB80A6B218C" type="email">adrienne.jochum@senckenberg.de</mods:nameIdentifier>
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<mods:namePart id="5C5AC5C03C7FE32DA9FF195F5DCB7332">J. Vermeulen, Jaap</mods:namePart>
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<mods:affiliation id="B730542D44C663BA441EC1B8D4D4E062">JK Art and Science, Lauwerbes 8, 2318 AT Leiden, The Netherlands</mods:affiliation>
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<mods:nameIdentifier id="87EE1A90EE5293552BF15E4E0D838B36" type="email">jk.artandscience@gmail.com</mods:nameIdentifier>
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<mods:namePart id="99A429612ADBA39B2CA10D63E86AC2B4">Anker, Katja</mods:namePart>
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<mods:nameIdentifier id="60BE877FBBED6E90E5EE49BFBAC78B97" type="ORCID">0000-0002-5350-0979</mods:nameIdentifier>
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<mods:affiliation id="9878673EE4092B903862985464C9F44D">JK Art and Science, Lauwerbes 8, 2318 AT Leiden, The Netherlands</mods:affiliation>
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<mods:namePart id="0DAB11BC8CED421B158C6328CA54FC4F">Hunyadi, András</mods:namePart>
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<mods:affiliation id="78923F67C061AB73B087BACE5FCA4343">Adria sétány 10 G 2 / 5, 1148 Budapest, Hungary</mods:affiliation>
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<mods:nameIdentifier id="C3EA30EB7691F86F54B6238FA172644C" type="email">hunand@freemail.hu</mods:nameIdentifier>
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<mods:namePart id="669D6C923E5D413E0D3E24A5607309EB">Örstan, Aydin</mods:namePart>
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<mods:nameIdentifier id="6F55908ACF8B0545607AFB25F7F21246" type="ORCID">0000-0002-2167-5694</mods:nameIdentifier>
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<mods:affiliation id="468712049109C9B0E1D039B27454B681">Section of Mollusks, Carnegie Museum of Natural History, Pittsburgh, PA 15213, USA</mods:affiliation>
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<mods:nameIdentifier id="D3324D35E057FFDCCA09E017DB6C07AB" type="email">anothersnail@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="6859B8451E754FA6A397AE5B679C79AB">Szabó, Ábel</mods:namePart>
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<mods:nameIdentifier id="C3707109CE2F9E4281D084F38BBC715B" type="ORCID">0000-0003-0646-2639</mods:nameIdentifier>
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<mods:affiliation id="93D4EA76FB195B94162AF93A798453E0">Lithosphere Fluid Research Laboratory, Institute of Geography and Earth Sciences, Eötvös University, Pázmány p. sétány 1 / C, 1117 Budapest, Hungary</mods:affiliation>
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<mods:nameIdentifier id="C9ECB7506BFD14352AD91C0DC009BACB" type="email">abel.szabo@ttk.elte.hu</mods:nameIdentifier>
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<mods:namePart id="B23A2495ED7D872587DB9AEAA94C3895">Dányi, László</mods:namePart>
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<mods:affiliation id="90A9707B215C1C27679B566E37AB5D7D">Institute for Soil Sciences and Agricultural Chemistry, Centre for Agricultural Research, ELKH, Herman Ottó út 15, 1022 Budapest, Hungary</mods:affiliation>
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<mods:nameIdentifier id="E16266A464A59D4ED1B79DB809E5F94C" type="email">laszlodanyi@gmail.com</mods:nameIdentifier>
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<mods:namePart id="11B69326F80C751ECF5B56BB279819EE">Schilthuizen, Menno</mods:namePart>
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<mods:nameIdentifier id="3C62988DDFDEF9A08E0FC034127451B6" type="ORCID">0000-0001-6229-0347</mods:nameIdentifier>
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<mods:affiliation id="609FC80DA17934700743EB7F77B6F737">Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, The Netherlands & Institute for Biology Leiden, Leiden University, Sylviusweg 72, 2333 BE Leiden, The Netherlands & Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Jalan UMS, 88400 Kota Kinabalu, Sabah, Malaysia</mods:affiliation>
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<mods:nameIdentifier id="A73EB4A4E85B3FE369D96C49B6F5DA4D" type="email">menno.schilthuizen@naturalis.nl</mods:nameIdentifier>
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<mods:title id="A22CA692C30BBFCC24A27BA1704AD35F">Contributions To Zoology</mods:title>
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<mods:part id="707E7D01751BDE0CDF88ED5174DC2AAD">
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<mods:date id="E870905BD4742DE943CE1CA57C2CE02E">2022</mods:date>
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<mods:number id="BCB1090B7CCBBDD8E8EEBC446127814E">2022-01-05</mods:number>
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<mods:number id="27113D322D70A0E8386720EA21727980">91</mods:number>
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<mods:identifier id="E5AADBFB5AB3798B8C1F38F2E9A4A7B8" type="DOI">10.1163/18759866-BjA10025</mods:identifier>
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<treatment id="35368F2DFFB8FF9752DCF9C7FC58FEE3" ID-DOI="http://doi.org/10.5281/zenodo.6313874" ID-GBIF-Taxon="193401707" ID-Zenodo-Dep="6313874" LSID="urn:lsid:plazi:treatment:35368F2DFFB8FF9752DCF9C7FC58FEE3" httpUri="http://treatment.plazi.org/id/35368F2DFFB8FF9752DCF9C7FC58FEE3" lastPageId="7" lastPageNumber="69" pageId="4" pageNumber="66">
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<subSubSection id="F5856DB0FFB8FF9452DCF9C7FCFAF91C" pageId="4" pageNumber="66" type="nomenclature">
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<emphasis id="8FEBE229FFB8FF9452DCF9C7FCFAF91C" bold="true" pageId="4" pageNumber="66">
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<taxonomicName id="7A9F45B8FFB8FF9452DCF9C7FCBBF91F" ID-CoL="8V29J" authority="Pall-Gergely, Jochum & Hunyadi" authorityName="Pall-Gergely, Jochum & Hunyadi" authorityYear="2022" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="66" phylum="Mollusca" rank="species" species="coprologos" status="n. sp.">
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<emphasis id="8FEBE229FFB8FF9452DCF9C7FC28F9F8" bold="true" box="[645,913,1607,1634]" italics="true" pageId="4" pageNumber="66">Angustopila coprologos</emphasis>
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Páll-Gergely, Jochum & Hunyadi
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</taxonomicName>
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<taxonomicNameLabel id="94D85F52FFB8FF945351F9ECFCFAF91C" box="[776,835,1644,1670]" pageId="4" pageNumber="66" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="F5856DB0FFB8FF9452DCF90FFC1EF954" pageId="4" pageNumber="66" type="multiple">
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</subSubSection>
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<caption id="E9E06EB3FFB9FF9550D8F930FDD2F95C" ID-DOI="http://doi.org/10.5281/zenodo.6147310" ID-Zenodo-Dep="6147310" box="[129,619,1712,1734]" httpUri="https://zenodo.org/record/6147310/files/figure.png" pageId="5" pageNumber="67" startId="5.[129,196,1712,1733]" targetBox="[131,1187,164,1691]" targetPageId="5">
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<paragraph id="BD203E3BFFB9FF9550D8F930FDD2F95C" blockId="5.[129,619,1712,1734]" box="[129,619,1712,1734]" pageId="5" pageNumber="67">
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<pageTitle id="FD00E65CFFB9FF9550D8F930FF76F95F" box="[129,207,1712,1733]" pageId="5" pageNumber="67">FIGURE 1</pageTitle>
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<taxonomicName id="7A9F45B8FFB9FF9550A0F930FE72F95F" authority="Páll-Gergely & Jochum & J.Vermeulen & Anker & Hunyadi & Örstan & Szabó, 2022" authorityName="Páll-Gergely & Jochum & J.Vermeulen & Anker & Hunyadi & Örstan & Szabó" authorityYear="2022" box="[249,459,1712,1733]" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="67" phylum="Mollusca" rank="species" species="coprologos" status="n. sp.">
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<emphasis id="8FEBE229FFB9FF9550A0F930FE72F95F" box="[249,459,1712,1733]" italics="true" pageId="5" pageNumber="67">Angustopila coprologos</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="94D85F52FFB9FF955189F930FDB9F95F" box="[464,512,1712,1733]" pageId="5" pageNumber="67" rank="species">n. sp.</taxonomicNameLabel>
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(holotype).
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</paragraph>
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</caption>
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<caption id="E9E06EB3FFBAFF96503BFC4BFCBEFC65" ID-DOI="http://doi.org/10.5281/zenodo.6147316" ID-Zenodo-Dep="6147316" httpUri="https://zenodo.org/record/6147316/files/figure.png" pageId="6" pageNumber="68" startId="6.[98,165,971,992]" targetBox="[193,1062,164,952]" targetPageId="6">
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<paragraph id="BD203E3BFFBAFF96503BFC4BFCBEFC65" blockId="6.[98,1082,971,1023]" pageId="6" pageNumber="68">
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<heading id="E6688957FFBAFF96503BFC4BFF0DFC7A" allCaps="true" box="[98,180,971,992]" fontSize="9" level="1" pageId="6" pageNumber="68" reason="3">FIGURE 2</heading>
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Granules on the shell surface of
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<taxonomicName id="7A9F45B8FFBAFF965258FC4CFDC1FC7A" authority="Páll-Gergely & Jochum & J.Vermeulen & Anker & Hunyadi & Örstan & Szabó, 2022" authorityName="Páll-Gergely & Jochum & J.Vermeulen & Anker & Hunyadi & Örstan & Szabó" authorityYear="2022" box="[513,632,971,993]" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="68" phylum="Mollusca" rank="species" species="coprologos" status="n. sp.">
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<emphasis id="8FEBE229FFBAFF965258FC4CFDC1FC7A" box="[513,632,971,993]" italics="true" pageId="6" pageNumber="68">A. coprologos</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="94D85F52FFBAFF965224FC4CFD15FC7B" box="[637,684,972,993]" pageId="6" pageNumber="68" rank="species">n. sp.</taxonomicNameLabel>
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shells. A–B shows a shell with brown ‘mud’, whereas C–D shows a shell with white (calcareous) granules.
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</paragraph>
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</caption>
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<subSubSection id="F5856DB0FFBAFF96503BFBAEFF6FFA11" pageId="6" pageNumber="68" type="materials_examined">
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<paragraph id="BD203E3BFFBAFF96503BFBAEFF6FFA11" blockId="6.[98,610,1069,1741]" pageId="6" pageNumber="68">
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<materialsCitation id="0DF73466FFBAFF96503BFBAEFE9AFA85" ID-GBIF-Occurrence="3468252306" collectedFrom="limestone gorge" collectingDate="2019-10-02" collectionCode="HNHM" collectorName="A. Hunyadi" country="Laos" elevation="510" latitude="18.1091" location="Phu Phako" longLatPrecision="1" longitude="105.062965" municipality="Lak Sao" pageId="6" pageNumber="68" specimenCode="HNHM 104885" specimenCount="1" stateProvince="Bolikhamsai Province" typeStatus="holotype">
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<emphasis id="8FEBE229FFBAFF96503BFBAEFEB8FBDD" box="[98,257,1069,1095]" italics="true" pageId="6" pageNumber="68">Type material:</emphasis>
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<collectingCountry id="C5887EABFFBAFF965150FBAEFE87FBD2" box="[265,318,1070,1096]" name="Laos" pageId="6" pageNumber="68" value="Laos">Laos</collectingCountry>
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,
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<collectingRegion id="7F5BF0D9FFBAFF965110FBADFDF9FBDD" box="[329,576,1069,1095]" pageId="6" pageNumber="68">Bolikhamsai Province</collectingRegion>
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,
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<locationDeviation id="AC922719FFBAFF965215FBADFF3FFBF1" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="68" unit="km" value="15.0">15 km</locationDeviation>
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</quantity>
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southeast +
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northeast (on a side road) towards centre of
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<collectingMunicipality id="5D44A441FFBAFF965123FBF5FE6DFB15" box="[378,468,1141,1167]" pageId="6" pageNumber="68">Lak Sao</collectingMunicipality>
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,
|
||
<location id="B84068E0FFBAFF9651B8FBF5FDE4FB15" LSID="urn:lsid:plazi:treatment:35368F2DFFB8FF9752DCF9C7FC58FEE3:B84068E0FFBAFF9651B8FBF5FDE4FB15" box="[481,605,1141,1167]" country="Laos" latitude="18.1091" longLatPrecision="1" longitude="105.062965" municipality="Lak Sao" name="Phu Phako" pageId="6" pageNumber="68" stateProvince="Bolikhamsai Province">Phu Phako</location>
|
||
,
|
||
<collectedFrom id="0E837DC6FFBAFF96503BFB19FE98FB29" box="[98,289,1177,1203]" pageId="6" pageNumber="68">limestone gorge</collectedFrom>
|
||
,
|
||
<quantity id="7A6793DEFFBAFF96516FFB19FE3BFB29" box="[310,386,1177,1204]" metricMagnitude="2" metricUnit="m" metricValue="5.1" pageId="6" pageNumber="68" unit="m" value="510.0">
|
||
<elevation id="36B2D908FFBAFF96516FFB19FE3BFB29" box="[310,386,1177,1204]" metricMagnitude="2" metricUnit="m" metricValue="5.1" pageId="6" pageNumber="68" unit="m" value="510.0">510 m</elevation>
|
||
</quantity>
|
||
a.s.l.,
|
||
<geoCoordinate id="D8AB58FCFFBAFF965180FB19FDE7FB2E" box="[473,606,1177,1204]" degrees="18" direction="north" minutes="06.546" orientation="latitude" pageId="6" pageNumber="68" precision="1" value="18.1091">18°06.546’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="D8AB58FCFFBAFF96503BFB3DFF57FB42" box="[98,238,1213,1240]" degrees="105" direction="east" minutes="03.778" orientation="longitude" pageId="6" pageNumber="68" precision="1" value="105.062965">105°03.778’E</geoCoordinate>
|
||
, (locality code: 2019/110), leg.
|
||
<collectorName id="106A5BEDFFBAFF9650CEFB61FE99FB61" box="[151,288,1249,1275]" pageId="6" pageNumber="68">A. Hunyadi</collectorName>
|
||
,
|
||
<date id="C92118FBFFBAFF96516BFB61FE52FB61" box="[306,491,1249,1275]" pageId="6" pageNumber="68" value="2019-10-02">
|
||
<collectingDate id="D965E113FFBAFF96516BFB61FE52FB61" box="[306,491,1249,1275]" pageId="6" pageNumber="68" value="2019-10-02">2 October 2019</collectingDate>
|
||
</date>
|
||
,
|
||
<typeStatus id="62248099FFBAFF9651A7FB61FDDBFB61" box="[510,610,1249,1275]" pageId="6" pageNumber="68">holotype</typeStatus>
|
||
(
|
||
<specimenCode id="ED399640FFBAFF965035FA85FEA5FA85" box="[108,284,1284,1311]" pageId="6" pageNumber="68">HNHM 104885</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="0DF73466FFBAFF965163FA85FDE4FA85" ID-GBIF-Occurrence="3468252301" box="[314,605,1284,1311]" collectionCode="CUMZ" country="Laos" pageId="6" pageNumber="68" specimenCode="CUMZ 7435" specimenCount="2" stateProvince="Bolikhamsai Province" typeStatus="paratype">
|
||
<specimenCode id="ED399640FFBAFF965163FA85FE7FFA85" box="[314,454,1284,1311]" pageId="6" pageNumber="68">CUMZ 7435</specimenCode>
|
||
/
|
||
<specimenCount id="AB99F5B2FFBAFF965196FA84FE62FA85" box="[463,475,1284,1311]" pageId="6" pageNumber="68" type="generic">2</specimenCount>
|
||
<typeStatus id="62248099FFBAFF9651B7FA85FDE4FA85" box="[494,605,1285,1311]" pageId="6" pageNumber="68">paratypes</typeStatus>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="0DF73466FFBAFF96503BFAA9FE2FFAD9" ID-GBIF-Occurrence="3468252307" box="[98,406,1320,1347]" collectionCode="SMF" country="Laos" pageId="6" pageNumber="68" specimenCode="SMF 365053" specimenCount="2" stateProvince="Bolikhamsai Province" typeStatus="paratype">
|
||
<specimenCode id="ED399640FFBAFF96503BFAA9FF43FAD9" box="[98,250,1320,1347]" pageId="6" pageNumber="68">SMF 365053</specimenCode>
|
||
/
|
||
<specimenCount id="AB99F5B2FFBAFF96515AFAA8FEB6FAD9" box="[259,271,1320,1347]" pageId="6" pageNumber="68" type="generic">2</specimenCount>
|
||
<typeStatus id="62248099FFBAFF96517EFAA9FE2FFAD9" box="[295,406,1321,1347]" pageId="6" pageNumber="68">paratypes</typeStatus>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="0DF73466FFBAFF9651E8FAA9FF26FAFD" ID-GBIF-Occurrence="3468252310" collectionCode="VNMN" country="Laos" pageId="6" pageNumber="68" specimenCount="2" stateProvince="Bolikhamsai Province" typeStatus="paratype">
|
||
<collectionCode id="DB8EA6FEFFBAFF9651E8FAA9FE41FAD9" box="[433,504,1321,1347]" collectionName="VNMN" pageId="6" pageNumber="68">VNMN</collectionCode>
|
||
/
|
||
<specimenCount id="AB99F5B2FFBAFF96525BFAA9FDA9FAD9" box="[514,528,1321,1347]" pageId="6" pageNumber="68" type="generic">2</specimenCount>
|
||
<typeStatus id="62248099FFBAFF96527EFAA9FF26FAFD" pageId="6" pageNumber="68">paratypes</typeStatus>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="0DF73466FFBAFF9650EFFACCFE4EFAFD" ID-GBIF-Occurrence="3468252308" box="[182,503,1356,1383]" collectionCode="ZRC, MOL" country="Laos" pageId="6" pageNumber="68" specimenCount="2" stateProvince="Bolikhamsai Province" typeStatus="paratype">
|
||
<collectionCode id="DB8EA6FEFFBAFF9650EFFACCFF5BFAFC" box="[182,226,1356,1382]" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection" pageId="6" pageNumber="68">ZRC</collectionCode>
|
||
.
|
||
<collectionCode id="DB8EA6FEFFBAFF9650B0FACCFEA4FAFC" box="[233,285,1356,1382]" collectionName="MOL" pageId="6" pageNumber="68">MOL</collectionCode>
|
||
.23217/
|
||
<specimenCount id="AB99F5B2FFBAFF965133FACCFECFFAFC" box="[362,374,1356,1382]" pageId="6" pageNumber="68" type="generic">2</specimenCount>
|
||
<typeStatus id="62248099FFBAFF9651D1FACDFE4EFAFD" box="[392,503,1357,1383]" pageId="6" pageNumber="68">paratypes</typeStatus>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="0DF73466FFBAFF965254FACDFF6BFA11" ID-GBIF-Occurrence="3468252305" country="Laos" pageId="6" pageNumber="68" specimenCount="697" stateProvince="Bolikhamsai Province" typeStatus="paratype">
|
||
HA/
|
||
<specimenCount id="AB99F5B2FFBAFF965261FACCFDD8FAFC" box="[568,609,1356,1382]" pageId="6" pageNumber="68" type="generic">697</specimenCount>
|
||
<typeStatus id="62248099FFBAFF96503BFAF1FF6BFA11" box="[98,210,1393,1419]" pageId="6" pageNumber="68">paratypes</typeStatus>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F5856DB0FFBAFF9650D8FA15FE66F9F9" pageId="6" pageNumber="68" type="diagnosis">
|
||
<paragraph id="BD203E3BFFBAFF9650D8FA15FE66F9F9" blockId="6.[98,610,1069,1741]" pageId="6" pageNumber="68">
|
||
<emphasis id="8FEBE229FFBAFF9650D8FA15FF4AFA34" box="[129,243,1429,1454]" italics="true" pageId="6" pageNumber="68">Diagnosis:</emphasis>
|
||
A strongly depressed-globular
|
||
<taxonomicName id="7A9F45B8FFBAFF96503BFA38FEFCFA48" box="[98,325,1464,1490]" pageId="6" pageNumber="68">
|
||
<emphasis id="8FEBE229FFBAFF96503BFA38FF5EFA48" box="[98,231,1464,1490]" italics="true" pageId="6" pageNumber="68">Angustopila</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
with a wide umbilicus, strong spiral sculpture consisting of a series of coarse elevations (flat-topped beads) in a chain-like pattern and four well-developed teeth (1 parietal, 2 palatal, 1 basal).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F5856DB0FFBAFF9750D8F9ECFED9FC41" lastPageId="7" lastPageNumber="69" pageId="6" pageNumber="68" type="description">
|
||
<paragraph id="BD203E3BFFBAFF9750D8F9ECFE07FCEA" blockId="6.[98,610,1069,1741]" lastBlockId="7.[128,640,171,1742]" lastPageId="7" lastPageNumber="69" pageId="6" pageNumber="68">
|
||
<emphasis id="8FEBE229FFBAFF9650D8F9ECFEB0F91F" box="[129,265,1644,1669]" italics="true" pageId="6" pageNumber="68">Description:</emphasis>
|
||
Shell tiny, whitish, strongly depressed-globular, dorsal part domeshaped, body whorl widest in standard apertural view; protoconch with slightly less than 1.5 whorls and a minute and very uneven pattern of polygonal, rather sharpcrested ridges locally forming minute, conical peaks where ridges split (the structure also reminiscent of cross-sectional trabecular bone); this surface structure forms raised spiral striae on last half of protoconch whorl, and gradually becomes stronger towards first teleoconch whorl; teleoconch bears similar basal sculpture as that of protoconch, strongly ornamented by a pattern of elevated, equidistantly-arranged rows of elegantly linked flat-topped beads; some fine, irregular radial sculpture also visible, but none especially conspicuous due to prominent bead-like striations; whorls 3.25–3.5, separated by deep suture; whorls slightly shouldered; aperture oblique to shell axis in lateral view; umbilicus wide, ca. one third of shell width, in living specimens containing several mud granules; aperture narrow, semilunar, sinulus relatively wide, rather strongly separated due to strong parietal tooth; peristome expanded, not reflected; parietal callus strongly protruding, remarkably visible in lateral perspective as projecting beyond peristome and nearly in line with shell periphery, detached from penultimate whorl; parietal tooth strongly developed, long, curved, pointing towards upper palatal tooth; palatal wall with an upper, smaller tooth situated at peristome edge, and a slightly larger, lower tooth situated somewhat deeper, both blunt; columellar/basal, tooth blunt with wide base, positioned on lower part of columellar area, located at similar distance from peristome edge as lower palatal tooth.
|
||
</paragraph>
|
||
<paragraph id="BD203E3BFFBBFF9750C6FCF9FE12FC09" blockId="7.[128,640,171,1742]" box="[159,427,889,916]" pageId="7" pageNumber="69">Soft anatomy unknown.</paragraph>
|
||
<paragraph id="BD203E3BFFBBFF9750C6FC1EFED9FC41" blockId="7.[128,640,171,1742]" pageId="7" pageNumber="69">
|
||
<emphasis id="8FEBE229FFBBFF9750C6FC1EFE1CFC22" box="[159,421,926,952]" italics="true" pageId="7" pageNumber="69">Measurements (in mm):</emphasis>
|
||
SW = 0.66–0.76; SH = 0.49–0.58 (n = 19).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F5856DB0FFBBFF9750C6FC65FEE0FB15" pageId="7" pageNumber="69" type="diagnosis">
|
||
<paragraph id="BD203E3BFFBBFF9750C6FC65FEE0FB15" blockId="7.[128,640,171,1742]" pageId="7" pageNumber="69">
|
||
<emphasis id="8FEBE229FFBBFF9750C6FC65FF00FBB9" italics="true" pageId="7" pageNumber="69">
|
||
Differential diagnosis:
|
||
<taxonomicName id="7A9F45B8FFBBFF9751FFFC65FF00FBB9" authority="Páll-Gergely & Jochum & J.Vermeulen & Anker & Hunyadi & Örstan & Szabó, 2022" authorityName="Páll-Gergely & Jochum & J.Vermeulen & Anker & Hunyadi & Örstan & Szabó" authorityYear="2022" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="69" phylum="Mollusca" rank="species" species="coprologos" status="n. sp.">Angustopila coprologos</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel id="94D85F52FFBBFF97509AFB8AFEB9FBBE" box="[195,256,1034,1060]" pageId="7" pageNumber="69" rank="species">n. sp.</taxonomicNameLabel>
|
||
can be easily distinguished from all other
|
||
<taxonomicName id="7A9F45B8FFBBFF9750BFFBADFE7DFBDD" box="[230,452,1069,1095]" pageId="7" pageNumber="69">
|
||
<emphasis id="8FEBE229FFBBFF9750BFFBADFED2FBDD" box="[230,363,1069,1095]" italics="true" pageId="7" pageNumber="69">Angustopila</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
by its depressed shell, the four apertural denticles and the pronounced sculpture.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F5856DB0FFBBFF9750C6FB19FF46FAD8" pageId="7" pageNumber="69" type="etymology">
|
||
<paragraph id="BD203E3BFFBBFF9750C6FB19FF46FAD8" blockId="7.[128,640,171,1742]" pageId="7" pageNumber="69">
|
||
<emphasis id="8FEBE229FFBBFF9750C6FB19FEA1FB29" box="[159,280,1177,1203]" italics="true" pageId="7" pageNumber="69">Etymology:</emphasis>
|
||
The specific epithet coprologos (Ancient Greek for dung gatherer) refers to the mud granules (here assumed to be faeces) placed on this species’ shell. Used as a noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F5856DB0FFBBFF9750C6FACCFE13FA6C" pageId="7" pageNumber="69" type="distribution">
|
||
<paragraph id="BD203E3BFFBBFF9750C6FACCFE13FA6C" blockId="7.[128,640,171,1742]" pageId="7" pageNumber="69">
|
||
<emphasis id="8FEBE229FFBBFF9750C6FACCFE91FAFC" box="[159,296,1356,1382]" italics="true" pageId="7" pageNumber="69">Distribution:</emphasis>
|
||
This species is known only from the
|
||
<typeStatus id="62248099FFBBFF975155FAF1FE84FA11" box="[268,317,1393,1419]" pageId="7" pageNumber="69">type</typeStatus>
|
||
locality in Bolikhamsay Province, Laos: ca.
|
||
<quantity id="7A6793DEFFBBFF975129FA14FE01FA34" box="[368,440,1428,1454]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="69" unit="km" value="13.0">13 km</quantity>
|
||
southeast (in a straight line) from the town Lak Sao (see also supplementary
|
||
<figureCitation id="25A422BEFFBBFF975169FA5CFE34FA6C" box="[304,397,1500,1526]" captionStart="FIGURE 1" captionStartId="5.[129,196,1712,1733]" captionTargetBox="[131,1187,164,1691]" captionTargetId="figure-3@5.[129,1187,164,1691]" captionText="FIGURE 1 Angustopila coprologos n. sp.(holotype)." figureDoi="http://doi.org/10.5281/zenodo.6147310" httpUri="https://zenodo.org/record/6147310/files/figure.png" pageId="7" pageNumber="69">figs S1–S</figureCitation>
|
||
<figureCitation id="25A422BEFFBBFF9751D4FA5CFE23FA6C" box="[397,410,1500,1526]" captionStart="FIGURE 2" captionStartId="6.[98,165,971,992]" captionTargetBox="[193,1062,164,952]" captionTargetId="figure-233@6.[193,1062,164,952]" captionText="FIGURE 2 Granules on the shell surface of A.coprologos n. sp. shells.A–B shows a shell with brown ‘mud’, whereas C–D shows a shell with white (calcareous) granules." figureDoi="http://doi.org/10.5281/zenodo.6147316" httpUri="https://zenodo.org/record/6147316/files/figure.png" pageId="7" pageNumber="69">2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F5856DB0FFBBFF9750C6F980FC58FEE3" pageId="7" pageNumber="69" type="biology_ecology">
|
||
<paragraph id="BD203E3BFFBBFF9750C6F980FC58FEE3" blockId="7.[128,640,171,1742]" lastBlockId="7.[676,1187,171,377]" pageId="7" pageNumber="69">
|
||
<emphasis id="8FEBE229FFBBFF9750C6F980FF40F980" box="[159,249,1536,1562]" italics="true" pageId="7" pageNumber="69">Habitat:</emphasis>
|
||
Since no living individuals were found (only shells of freshly dead specimens), no observations about the specific habitat of
|
||
<taxonomicName id="7A9F45B8FFBBFF9750D8F9EBFE3AF91F" authority="Páll-Gergely & Jochum & J.Vermeulen & Anker & Hunyadi & Örstan & Szabó, 2022" authorityName="Páll-Gergely & Jochum & J.Vermeulen & Anker & Hunyadi & Örstan & Szabó" authorityYear="2022" box="[129,387,1643,1669]" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="69" phylum="Mollusca" rank="species" species="coprologos" status="n. sp.">
|
||
<emphasis id="8FEBE229FFBBFF9750D8F9EBFE3AF91F" box="[129,387,1643,1669]" italics="true" pageId="7" pageNumber="69">Angustopila coprologos</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="94D85F52FFBBFF9751D2F9ECFE71F91C" box="[395,456,1644,1670]" pageId="7" pageNumber="69" rank="species">n. sp.</taxonomicNameLabel>
|
||
could be made. The living snails presumably live deep in limestone crevices close to (or on) root systems. The summer of 2019 was very wet and rainy with extensive flooding throughout the flood plains of the Mekong River. This might have played a role in the vast accumulation of empty shells at the base of rock formations (collection in
|
||
<date id="C92118FBFFBBFF975366FEDFFC6FFEE3" box="[831,982,351,377]" pageId="7" pageNumber="69" value="2019-10">October 2019</date>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |