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<document id="33BA20837FD6AC1385F7E98D1BB2CE34" ID-CLB-Dataset="4642" ID-DOI="10.5281/zenodo.5320906" ID-GBIF-Dataset="f33526c4-a22f-4c32-9fd9-34a1255f44eb" ID-ISSN="0374-1036" ID-Zenodo-Dep="5320906" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630136926086" checkinUser="felipe" docAuthor="Ito, Noboru" docDate="2009" docId="352E879FFF95FFCBFEA2FD61FED0FB54" docLanguage="en" docName="ActaEntMusNatPra.2009.49.2.599-606.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 49 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Trichotichnus (Pseudotrichotichnus) vietnamensis Ito 2009, sp. nov." docType="treatment" docVersion="7" lastPageNumber="604" masterDocId="C917FFE7FF91FFCEFF99FF9AFFBBFFC7" masterDocTitle="Note on the subgenus Pseudotrichotichnus of the genus Trichotichnus from Southeast Asia (Coleoptera: Carabidae: Harpalini)" masterLastPageNumber="606" masterPageNumber="599" pageNumber="603" updateTime="1699168252378" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="549DB2C645F4129F23BB691B7BCF1059">Note on the subgenus Pseudotrichotichnus of the genus Trichotichnus from Southeast Asia (Coleoptera: Carabidae: Harpalini)</mods:title>
</mods:titleInfo>
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<mods:namePart id="BC2F9731C5273015BC6857F2CB1C5B3D">Ito, Noboru</mods:namePart>
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<mods:typeOfResource id="255E3CA578725AFA3BA3701C8712A771">text</mods:typeOfResource>
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<mods:title id="9EF35D625E3D244D45102C799FD8A84A">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="D12F637E004695B6A38696BF7598BA71">
<mods:date id="725C966A0EAB059CC31ACDD17F6CEB30">2009</mods:date>
<mods:detail id="F0C7BABE94D08A1A91408B106DF1F679" type="pubDate">
<mods:number id="CE0BE5E9BE10CBE6B442AC83B47632EA">2009-12-15</mods:number>
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<mods:number id="366344DA4E52CC1190D16E0F528F3D39">49</mods:number>
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<mods:number id="1F57C5E88CDB4A26500592D8A146A727">2</mods:number>
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<mods:classification id="2273F1886CC165B878732EC3F99A4890">journal article</mods:classification>
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<treatment id="352E879FFF95FFCBFEA2FD61FED0FB54" ID-DOI="http://doi.org/10.5281/zenodo.5342762" ID-GBIF-Taxon="185202414" ID-Zenodo-Dep="5342762" LSID="urn:lsid:plazi:treatment:352E879FFF95FFCBFEA2FD61FED0FB54" httpUri="http://treatment.plazi.org/id/352E879FFF95FFCBFEA2FD61FED0FB54" lastPageId="5" lastPageNumber="604" pageId="4" pageNumber="603">
<subSubSection id="F59D6502FF95FFCAFEA2FD61FC79FCD3" box="[315,962,763,788]" pageId="4" pageNumber="603" type="nomenclature">
<paragraph id="BD383689FF95FFCAFEA2FD61FC79FCD3" blockId="4.[315,962,763,788]" box="[315,962,763,788]" pageId="4" pageNumber="603">
<heading id="E67081E5FF95FFCAFEA2FD61FC79FCD3" bold="true" box="[315,962,763,788]" centered="true" fontSize="8" level="2" pageId="4" pageNumber="603" reason="3">
<emphasis id="8FF3EA9BFF95FFCAFEA2FD61FC79FCD3" bold="true" box="[315,962,763,788]" pageId="4" pageNumber="603">
<taxonomicName id="7A874D0AFF95FFCAFEA2FD61FCDDFCD4" ID-CoL="7D4HX" authority="Ito, 2009" authorityName="Ito" authorityYear="2009" box="[315,870,763,788]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="603" phylum="Arthropoda" rank="species" species="vietnamensis" status="sp. nov." subGenus="Pseudotrichotichnus">
<emphasis id="8FF3EA9BFF95FFCAFEA2FD61FE6DFCD3" bold="true" box="[315,470,763,788]" italics="true" pageId="4" pageNumber="603">Trichotichnus</emphasis>
(
<emphasis id="8FF3EA9BFF95FFCAFE7FFD61FD72FCD3" bold="true" box="[486,713,763,788]" italics="true" pageId="4" pageNumber="603">Pseudotrichotichnus</emphasis>
)
<emphasis id="8FF3EA9BFF95FFCAFD41FD61FCDDFCD4" bold="true" box="[728,870,763,787]" italics="true" pageId="4" pageNumber="603">vietnamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="94C057E0FF95FFCAFCF4FD66FC79FCD3" box="[877,962,764,788]" pageId="4" pageNumber="603" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F59D6502FF95FFCAFDC9FC80FD15FCEA" box="[592,686,794,813]" pageId="4" pageNumber="603" type="description">
<paragraph id="BD383689FF95FFCAFDC9FC80FD15FCEA" blockId="4.[151,1127,794,988]" box="[592,686,794,813]" pageId="4" pageNumber="603">
(
<figureCitation id="25BC2A0CFF95FFCAFDCFFC80FD29FCEA" box="[598,658,794,813]" captionStart="Figs" captionStartId="2.[153,192,1499,1518]" captionTargetBox="[168,1126,250,1426]" captionTargetId="figure-10@2.[151,1131,219,1426]" captionTargetPageId="2" captionText="Figs. 15. 12 Trichotichnus (Pseudtrichotichnus) jelineki sp. nov., holotype (1 habitus; 2 male genitalia); 3 T. (P.) vietnamensis sp. nov.; 4 T. (P.) tenuitibialis sp. nov.; 5 T. (P.) niger Louwerens, 1951. Scale bar: 1 mm. Abbreviations: l lateral aspect, d dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.5320908" httpUri="https://zenodo.org/record/5320908/files/figure.png" pageId="4" pageNumber="603">Figs. 3</figureCitation>
,
<figureCitation id="25BC2A0CFF95FFCAFD05FC80FD1CFCEA" box="[668,679,794,813]" captionStart="Figs" captionStartId="3.[111,150,1072,1091]" captionTargetBox="[139,1068,228,1054]" captionTargetId="figure-14@3.[131,1070,219,1069]" captionTargetPageId="3" captionText="Figs. 69. Male genitalia. 6 Trichotichnus (Pseudotrichotichnus) vietnamensis sp. nov.; 7 T. (P.) tenuitibialis sp. nov.; 8 T. (P.) niger Louwerens, 1951; 9 T. (P.) miyakei Habu, 1980. Scale bar: 1 mm. Abbreviations: l lateral aspect, d dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.5320912" httpUri="https://zenodo.org/record/5320912/files/figure.png" pageId="4" pageNumber="603">6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="F59D6502FF95FFCAFF0EFCDEFE59FC1B" pageId="4" pageNumber="603" type="materials_examined">
<paragraph id="BD383689FF95FFCAFF0EFCDEFE59FC1B" blockId="4.[151,1127,794,988]" pageId="4" pageNumber="603">
<emphasis id="8FF3EA9BFF95FFCAFF0EFCDEFEA6FC90" bold="true" box="[151,285,835,855]" pageId="4" pageNumber="603">Type material.</emphasis>
<materialsCitation id="0DEF3CD4FF95FFCAFEBDFCDEFF5DFCB6" ID-GBIF-Occurrence="3339964305" collectingDate="1999-05-23" collectionCode="NMNS" collectorName="S. Ueno" country="Vietnam" elevation="1200" location="Mt. Pia Oac" pageId="4" pageNumber="603" specimenCount="1" stateProvince="Cao Bang" typeStatus="holotype">
<typeStatus id="623C882BFF95FFCAFEBDFCDEFECCFC92" box="[292,375,836,855]" pageId="4" pageNumber="603" type="holotype">HOLOTYPE</typeStatus>
: J,
<location id="B8586052FF95FFCAFE3DFCDEFDA9FC90" LSID="urn:lsid:plazi:treatment:352E879FFF95FFCBFEA2FD61FED0FB54:B8586052FF95FFCAFE3DFCDEFDA9FC90" box="[420,530,836,855]" country="Vietnam" name="Mt. Pia Oac" pageId="4" pageNumber="603" stateProvince="Cao Bang">Mt. Pia Oac</location>
, alt.
<quantity id="7A7F9B6CFF95FFCAFDA6FCDEFD39FC90" box="[575,642,836,855]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="603" unit="m" value="1200.0">
<elevation id="36AAD1BAFF95FFCAFDA6FCDEFD39FC90" box="[575,642,836,855]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="603" unit="m" value="1200.0">1,200m</elevation>
</quantity>
,
<collectingRegion id="7F43F86BFF95FFCAFD12FCDEFCA3FC90" box="[651,792,836,855]" country="Vietnam" name="Cao Bang" pageId="4" pageNumber="603">Cao Bang Prov.</collectingRegion>
, N.
<collectingCountry id="C5907619FF95FFCAFCA5FCDEFC3DFC90" box="[828,902,836,855]" name="Vietnam" pageId="4" pageNumber="603">Vietnam</collectingCountry>
,
<date id="C9391049FF95FFCAFC08FCDEFC51FC90" box="[913,1002,836,855]" pageId="4" pageNumber="603" value="1999-05-23">
<collectingDate id="D97DE9A1FF95FFCAFC08FCDEFC51FC90" box="[913,1002,836,855]" pageId="4" pageNumber="603" value="1999-05-23">23.V.1999</collectingDate>
</date>
,
<collectorName id="1072535FFF95FFCAFC6DFCDEFB81FC90" box="[1012,1082,836,855]" pageId="4" pageNumber="603">S. Uéno</collectorName>
leg. (
<collectionCode id="DB96AE4CFF95FFCAFF04FCC4FF64FCB6" box="[157,223,862,881]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="4" pageNumber="603">NMNS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="0DEF3CD4FF95FFCAFF77FCC5FDD0FCB6" ID-GBIF-Occurrence="3339964314" box="[238,619,856,882]" collectingDate="1999-05-23" collectionCode="NMNS" collectorName="S. Ueno" country="Vietnam" elevation="1200" location="Mt. Pia Oac" pageId="4" pageNumber="603" specimenCount="2" specimenCount-female="2" stateProvince="Cao Bang" typeStatus="paratype">
<typeStatus id="623C882BFF95FFCAFF77FCC5FEFDFCB7" box="[238,326,863,882]" pageId="4" pageNumber="603" type="paratype">PARATYPES</typeStatus>
:
<specimenCount id="AB81FD00FF95FFCAFEC8FCC4FE3AFCB6" box="[337,385,856,881]" pageId="4" pageNumber="603" type="female">2 ♀♀</specimenCount>
, same data as the holotype
</materialsCitation>
;
<materialsCitation id="0DEF3CD4FF95FFCAFDEDFCC4FEB2FC4B" ID-GBIF-Occurrence="3339964315" collectingDate="1995-05-10" collectorName="S. Ueno" country="Vietnam" elevation="1550" location="Sa Pa" pageId="4" pageNumber="603" specimenCount="1" specimenCount-female="1" stateProvince="Lao Cai" typeStatus="paratype">
2 JJ
<specimenCount id="AB81FD00FF95FFCAFD30FCC4FD72FCB6" box="[681,713,856,881]" pageId="4" pageNumber="603" type="female">1 ♀</specimenCount>
,
<location id="B8586052FF95FFCAFD41FCC4FCB6FCB5" LSID="urn:lsid:plazi:treatment:352E879FFF95FFCBFEA2FD61FED0FB54:B8586052FF95FFCAFD41FCC4FCB6FCB5" box="[728,781,862,882]" country="Vietnam" name="Sa Pa" pageId="4" pageNumber="603" stateProvince="Lao Cai">Sa Pa</location>
, alt.
<quantity id="7A7F9B6CFF95FFCAFCAEFCC4FC3BFCB5" box="[823,896,862,882]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="4" pageNumber="603" unit="m" value="1550.0">
<elevation id="36AAD1BAFF95FFCAFCAEFCC4FC3BFCB5" box="[823,896,862,882]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="4" pageNumber="603" unit="m" value="1550.0">1,550 m</elevation>
</quantity>
,
<collectingRegion id="7F43F86BFF95FFCAFC11FCC5FBB9FCB5" box="[904,1026,862,882]" country="Vietnam" name="Lao Cai" pageId="4" pageNumber="603">Lao Cai Prov.</collectingRegion>
,
<date id="C9391049FF95FFCAFB93FCC4FBD8FCB6" box="[1034,1123,862,881]" pageId="4" pageNumber="603" value="1995-05-10">
<collectingDate id="D97DE9A1FF95FFCAFB93FCC4FBD8FCB6" box="[1034,1123,862,881]" pageId="4" pageNumber="603" value="1995-05-10">10.V.1995</collectingDate>
</date>
,
<collectorName id="1072535FFF95FFCAFF0EFCE3FF66FC4B" box="[151,221,889,908]" pageId="4" pageNumber="603">S. Uéno</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="0DEF3CD4FF95FFCAFE8EFCE3FE80FC61" ID-GBIF-Occurrence="3339964310" collectingDate="1995-04-22" collectingDateMax="1995-05-15" collectingDateMin="1995-04-22" collectionCode="NMNS, NIKJ" collectorName="S. Ueno" country="Vietnam" elevation="960" location="Tam Dao" pageId="4" pageNumber="603" specimenCount="1" specimenCount-female="1" stateProvince="Lao Cai" typeStatus="paratype">
1 J,
<location id="B8586052FF95FFCAFEDEFCE3FE26FC4B" LSID="urn:lsid:plazi:treatment:352E879FFF95FFCBFEA2FD61FED0FB54:B8586052FF95FFCAFEDEFCE3FE26FC4B" box="[327,413,889,908]" country="Vietnam" name="Tam Dao" pageId="4" pageNumber="603">Tam Dao</location>
, alt.
<quantity id="7A7F9B6CFF95FFCAFE5EFCE3FDBAFC4B" box="[455,513,889,908]" metricMagnitude="2" metricUnit="m" metricValue="9.6" pageId="4" pageNumber="603" unit="m" value="960.0">
<elevation id="36AAD1BAFF95FFCAFE5EFCE3FDBAFC4B" box="[455,513,889,908]" metricMagnitude="2" metricUnit="m" metricValue="9.6" pageId="4" pageNumber="603" unit="m" value="960.0">960 m</elevation>
</quantity>
, N.
<collectingCountry id="C5907619FF95FFCAFDBCFCE3FDD4FC4B" box="[549,623,889,908]" name="Vietnam" pageId="4" pageNumber="603">Vietnam</collectingCountry>
,
<date id="C9391049FF95FFCAFDE0FCE3FD61FC4B" box="[633,730,889,908]" pageId="4" pageNumber="603" value="1995-04-22">
<collectingDate id="D97DE9A1FF95FFCAFDE0FCE3FD61FC4B" box="[633,730,889,908]" pageId="4" pageNumber="603" value="1995-04-22">22.IV.1995</collectingDate>
</date>
,
<collectorName id="1072535FFF95FFCAFD7AFCE3FC92FC4B" box="[739,809,889,908]" pageId="4" pageNumber="603">S. Uéno</collectorName>
leg. (
<collectionCode id="DB96AE4CFF95FFCAFCF8FCE3FC19FC4B" box="[865,930,889,908]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="4" pageNumber="603">NMNS</collectionCode>
,
<collectionCode id="DB96AE4CFF95FFCAFC35FCE3FC66FC4B" box="[940,989,889,908]" pageId="4" pageNumber="603">NIKJ</collectionCode>
),
<specimenCount id="AB81FD00FF95FFCAFC74FCE3FBB5FC4B" box="[1005,1038,883,908]" pageId="4" pageNumber="603" type="female">1 ♀</specimenCount>
, the same locality,
<date id="C9391049FF95FFCAFF7BFC09FE80FC61" box="[226,315,915,934]" pageId="4" pageNumber="603" value="1995-05-15">
<collectingDate id="D97DE9A1FF95FFCAFF7BFC09FE80FC61" box="[226,315,915,934]" pageId="4" pageNumber="603" value="1995-05-15">15.V.1995</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="0DEF3CD4FF95FFCAFEDCFC0EFC83FC60" ID-GBIF-Occurrence="3339964301" box="[325,824,910,936]" collectingDate="1995-04-22" collectingDateMax="1995-05-15" collectingDateMin="1995-04-22" collectionCode="NMNS, NIKJ" collectorName="S. Ueno" country="Vietnam" elevation="960" location="Tam Dao" pageId="4" pageNumber="603" specimenCount="3" specimenCount-female="3" stateProvince="Lao Cai" typeStatus="paratype">
2 JJ
<specimenCount id="AB81FD00FF95FFCAFEE5FC0EFE17FC60" box="[380,428,910,935]" pageId="4" pageNumber="603" type="female">3 ♀♀</specimenCount>
, the same locality,
<date id="C9391049FF95FFCAFDC0FC09FD06FC61" box="[601,701,915,934]" pageId="4" pageNumber="603" value="1995-04-22">
<collectingDate id="D97DE9A1FF95FFCAFDC0FC09FD06FC61" box="[601,701,915,934]" pageId="4" pageNumber="603" value="1995-04-22">22.IV.1995</collectingDate>
</date>
,
<collectorName id="1072535FFF95FFCAFD5EFC0EFCB7FC60" box="[711,780,916,935]" pageId="4" pageNumber="603">S. Uéno</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="0DEF3CD4FF95FFCAFCDFFC0EFF47FC06" ID-GBIF-Occurrence="3339964316" collectingDate="1995-04-21" collectingDateMax="1995-05-15" collectingDateMin="1995-04-22" collectionCode="NMNS, NIKJ" collectorName="S. Ueno" country="Vietnam" elevation="930" location="Tam Dao" pageId="4" pageNumber="603" specimenCount="1" specimenCount-female="1" stateProvince="Lao Cai" typeStatus="paratype">
<specimenCount id="AB81FD00FF95FFCAFCDFFC0EFCDDFC60" box="[838,870,910,935]" pageId="4" pageNumber="603" type="female">1 ♀</specimenCount>
, the same locality, alt.
<quantity id="7A7F9B6CFF95FFCAFBA9FC0EFBD8FC60" box="[1072,1123,916,935]" metricMagnitude="2" metricUnit="m" metricValue="9.3" pageId="4" pageNumber="603" unit="m" value="930.0">
<elevation id="36AAD1BAFF95FFCAFBA9FC0EFBD8FC60" box="[1072,1123,916,935]" metricMagnitude="2" metricUnit="m" metricValue="9.3" pageId="4" pageNumber="603" unit="m" value="930.0">930m</elevation>
</quantity>
,
<date id="C9391049FF95FFCAFF0EFC34FF47FC06" box="[151,252,942,961]" pageId="4" pageNumber="603" value="1995-04-21">
<collectingDate id="D97DE9A1FF95FFCAFF0EFC34FF47FC06" box="[151,252,942,961]" pageId="4" pageNumber="603" value="1995-04-21">21.IV.1995</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="0DEF3CD4FF95FFCAFE9EFC34FC25FC06" ID-GBIF-Occurrence="3339964302" box="[263,926,936,962]" collectingDate="1991-04-16" collectingDateMax="1991-04-23" collectingDateMin="1991-04-16" collectionCode="HNHM" collectorName="Strnad Jan" country="Vietnam" elevation="900" location="Tam Dao" pageId="4" pageNumber="603" specimenCount="2" specimenCount-female="2" stateProvince="Lao Cai" typeStatus="paratype">
<specimenCount id="AB81FD00FF95FFCAFE9EFC34FE8FFC06" box="[263,308,936,961]" pageId="4" pageNumber="603" type="female">2 ♀♀</specimenCount>
, the same locality, alt.
<quantity id="7A7F9B6CFF95FFCAFE64FC34FD94FC06" box="[509,559,942,961]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="603" unit="m" value="900.0">
<elevation id="36AAD1BAFF95FFCAFE64FC34FD94FC06" box="[509,559,942,961]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="603" unit="m" value="900.0">900m</elevation>
</quantity>
,
<date id="C9391049FF95FFCAFDA1FC34FD06FC06" box="[568,701,942,961]" pageId="4" pageNumber="603" value="1991-04-16" valueMax="1991-04-23" valueMin="1991-04-16">
<collectingDate id="D97DE9A1FF95FFCAFDA1FC34FD06FC06" box="[568,701,942,961]" pageId="4" pageNumber="603" value="1991-04-16" valueMax="1991-04-23" valueMin="1991-04-16">16.23.IV.1991</collectingDate>
</date>
,
<collectorName id="1072535FFF95FFCAFD5CFC34FCA5FC05" box="[709,798,942,962]" pageId="4" pageNumber="603">Strnad Jan</collectorName>
leg. (
<collectionCode id="DB96AE4CFF95FFCAFCCBFC34FC23FC06" box="[850,920,942,961]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="4" pageNumber="603">HNHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="0DEF3CD4FF95FFCAFC3FFC34FE65FC1B" ID-GBIF-Occurrence="3339964311" collectingDate="1989-04-20" collectionCode="DWBG" collectorName="F. Schmidt" country="Vietnam" location="Da Lat" pageId="4" pageNumber="603" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="AB81FD00FF95FFCAFC3FFC34FC7FFC06" box="[934,964,936,961]" pageId="4" pageNumber="603" type="female">1 ♀</specimenCount>
,
<location id="B8586052FF95FFCAFC4BFC34FBA9FC05" LSID="urn:lsid:plazi:treatment:352E879FFF95FFCBFEA2FD61FED0FB54:B8586052FF95FFCAFC4BFC34FBA9FC05" box="[978,1042,942,962]" country="Vietnam" name="Da Lat" pageId="4" pageNumber="603">Da Lat</location>
,
<collectingCountry id="C5907619FF95FFCAFB80FC34FBD8FC06" box="[1049,1123,942,961]" name="Vietnam" pageId="4" pageNumber="603">Vietnam</collectingCountry>
,
<date id="C9391049FF95FFCAFF0EFC53FF4CFC1B" box="[151,247,969,988]" pageId="4" pageNumber="603" value="1989-04-20">
<collectingDate id="D97DE9A1FF95FFCAFF0EFC53FF4CFC1B" box="[151,247,969,988]" pageId="4" pageNumber="603" value="1989-04-20">20.IV.1989</collectingDate>
</date>
,
<collectorName id="1072535FFF95FFCAFE99FC53FEE7FC1B" box="[256,348,969,988]" pageId="4" pageNumber="603">F. Schmidt</collectorName>
leg. (
<collectionCode id="DB96AE4CFF95FFCAFE0BFC53FE63FC1B" box="[402,472,969,988]" pageId="4" pageNumber="603">DWBG</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F59D6502FF95FFCBFF0EFC6EFD70FC34" lastPageId="5" lastPageNumber="604" pageId="4" pageNumber="603" type="description">
<paragraph id="BD383689FF95FFCAFF0EFC6EFDF7FB8B" blockId="4.[151,1127,1012,1613]" pageId="4" pageNumber="603">
<emphasis id="8FF3EA9BFF95FFCAFF0EFC6EFE9CFBCB" bold="true" box="[151,295,1012,1036]" pageId="4" pageNumber="603">Description.</emphasis>
Body oblong, fairly convex, pitchy black though a little lighter than in
<taxonomicName id="7A874D0AFF95FFCAFF0EFB8CFEBDFBEA" authorityName="Habu" authorityYear="1980" box="[151,262,1045,1069]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="603" phylum="Arthropoda" rank="species" species="miyakei">
<emphasis id="8FF3EA9BFF95FFCAFF0EFB8CFF1DFBEA" box="[151,166,1046,1069]" italics="true" pageId="4" pageNumber="603">T</emphasis>
.
<emphasis id="8FF3EA9BFF95FFCAFF2AFB8CFEBDFBEA" box="[179,262,1046,1069]" italics="true" pageId="4" pageNumber="603">miyakei</emphasis>
</taxonomicName>
, strongly shiny, iridescent on elytra; palpi, antennae, tarsi and tibiae light reddish brown, femora brown to blackish brown.
</paragraph>
<paragraph id="BD383689FF95FFCAFF2CFBCEFD6EFA6B" blockId="4.[151,1127,1012,1613]" pageId="4" pageNumber="603">Head rather large, 0.710.74 times as wide as pronotum, gently convex, very sparsely and microscopically punctate; labrum subquadrate, triangularly emarginate apically; clypeus transversely elevated, shallowly emarginate at apex; clypeal suture very weakly curved, deep even in middle, more strongly deepened at sides; frontal impressions as deep as clypeal suture at sides, obliquely linear; eyes large, well convex; temples steeply inclined, rather long, one-third the eye length; genuine ventral margin of eyes widely isolated from buccal fissure; antennae short, hardly reaching pronotal base, segment 3 pubescent in apical half, as long as segment 4 and 1.5 times as long as segment 2; labial palpi moderately thick, segment 3 as long as segment 2; ligula slim, slightly thickened apicad; paraglossae narrow, prolonged forwards beyond ligula; mentum with median tooth small and triangular, epilobes not widened apicad; microsculpture mostly invisible in 80× magnification.</paragraph>
<paragraph id="BD383689FF95FFCBFF2CFA2FFEE8FE94" blockId="4.[151,1127,1012,1613]" lastBlockId="5.[113,1089,219,1171]" lastPageId="5" lastPageNumber="604" pageId="4" pageNumber="603">Pronotum almost quadrate, widest at apical third, 1.4 times as wide as long, so widely convex that lateral furrows are linearly impressed; sides weakly curved between anterior margin and midlength, obliquely linear elsewhere; anterior margin weakly emarginate, straight in middle, without border in narrow middle area; base with broken border; anterior angles widely arcuate; basal angles obtuse, very narrowly rounded; basal foveae flattened, each with shallow groove; front transverse impression very shallow, hind one also shallow; disc smooth, punctation absent or sparse anteriorly, coarse and moderately dense in basal foveae; microsculpture largely consisting of transverse meshes, and of isodiametric meshes near basal punctures.</paragraph>
<paragraph id="BD383689FF94FFCBFF09FEC1FB87FD94" blockId="5.[113,1089,219,1171]" pageId="5" pageNumber="604">
Elytra oblong, weakly oval, 1.30 times as wide as pronotum, a little less than 1.50 times as long as wide, evenly convex, very sparsely and microscopically punctate; sides subparallel near midlength, abruptly and arcuately curved posteriorly; apices widely rounded at sides, with tips close to each other; base of each elytron shallowly emarginate, forming rounded angle with lateral margin; striae as in
<emphasis id="8FF3EA9BFF94FFCBFE50FE47FE63FE33" box="[457,472,477,500]" italics="true" pageId="5" pageNumber="604">T</emphasis>
.
<taxonomicName id="7A874D0AFF94FFCBFE7FFE46FDDEFE33" authority="Ito, 2009" authorityName="Ito" authorityYear="2009" box="[486,613,476,500]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="604" phylum="Arthropoda" rank="species" species="jelineki" status="sp. nov." subGenus="Pseudtrichotichnus">
(
<emphasis id="8FF3EA9BFF94FFCBFE76FE47FE44FE33" box="[495,511,477,500]" italics="true" pageId="5" pageNumber="604">P</emphasis>
.)
<emphasis id="8FF3EA9BFF94FFCBFD8FFE47FDDEFE33" box="[534,613,477,500]" italics="true" pageId="5" pageNumber="604">jelineki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="94C057E0FF94FFCBFDF5FE46FD78FE34" box="[620,707,476,499]" pageId="5" pageNumber="604" rank="species">sp. nov.</taxonomicNameLabel>
; intervals gently convex, becoming more convex towards apex and base, discal pore of interval 3 situated between apical third and two-fifths; marginal series widely interrupted medially, composed of 8 + 10 umbilicate pores; microsculpture consisting of fine and obscure transverse lines. Hind wings well developed.
</paragraph>
<paragraph id="BD383689FF94FFCBFF09FDC1FD50FD14" blockId="5.[113,1089,219,1171]" pageId="5" pageNumber="604">Ventral surface largely impunctate, very sparsely and minutely punctate on lateral areas of metasternum; metepisterna convergent posteriorly, nearly 1.5 times as long as wide; abdominal sternite 6 sparsely pubescent, with two setae at each side in both sexes, apex truncate or feebly emarginate in male and narrowly rounded in female.</paragraph>
<paragraph id="BD383689FF94FFCBFF09FD46FB84FCB4" blockId="5.[113,1089,219,1171]" pageId="5" pageNumber="604">
Fore tibiae a little more widened apicad than in
<taxonomicName id="7A874D0AFF94FFCBFD06FD47FCABFD33" authorityName="Habu" authorityYear="1980" box="[671,784,732,756]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="604" phylum="Arthropoda" rank="species" species="miyakei">
<emphasis id="8FF3EA9BFF94FFCBFD06FD47FD15FD33" box="[671,686,733,756]" italics="true" pageId="5" pageNumber="604">T</emphasis>
.
<emphasis id="8FF3EA9BFF94FFCBFD24FD47FCABFD33" box="[701,784,733,756]" italics="true" pageId="5" pageNumber="604">miyakei</emphasis>
</taxonomicName>
, sulcate in basal four-fifths, terminal spur expanded in basal third; hind tarsi 0.700.74 times as long as the width of head in male and 0.650.69 times as long in female, segment 1 0.9 times as long as segments 2 and 3 combined and 1.65 times as long as segment 2, segment 3 1.5 times as long as segment 4, all claws with two setae on outer margin and three setae on inner margin of ventral surface.
</paragraph>
<paragraph id="BD383689FF94FFCBFF09FCE1FC20FC14" blockId="5.[113,1089,219,1171]" pageId="5" pageNumber="604">
Aedeagus (
<figureCitation id="25BC2A0CFF94FFCBFE90FCE1FEF1FC54" box="[265,330,891,915]" captionStart="Figs" captionStartId="3.[111,150,1072,1091]" captionTargetBox="[139,1068,228,1054]" captionTargetId="figure-14@3.[131,1070,219,1069]" captionTargetPageId="3" captionText="Figs. 69. Male genitalia. 6 Trichotichnus (Pseudotrichotichnus) vietnamensis sp. nov.; 7 T. (P.) tenuitibialis sp. nov.; 8 T. (P.) niger Louwerens, 1951; 9 T. (P.) miyakei Habu, 1980. Scale bar: 1 mm. Abbreviations: l lateral aspect, d dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.5320912" httpUri="https://zenodo.org/record/5320912/files/figure.png" pageId="5" pageNumber="604">Fig. 6</figureCitation>
) similar in shape to
<taxonomicName id="7A874D0AFF94FFCBFDB3FCE7FD2CFC53" authority="Ito, 2009" authorityName="Ito" authorityYear="2009" box="[554,663,892,916]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="604" phylum="Arthropoda" rank="species" species="jelineki" status="sp. nov.">
<emphasis id="8FF3EA9BFF94FFCBFDB3FCE7FD82FC53" box="[554,569,893,916]" italics="true" pageId="5" pageNumber="604">T</emphasis>
.
<emphasis id="8FF3EA9BFF94FFCBFDD1FCE7FD2CFC53" box="[584,663,893,916]" italics="true" pageId="5" pageNumber="604">jelineki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="94C057E0FF94FFCBFD07FCE6FD4FFC54" box="[670,756,892,915]" pageId="5" pageNumber="604" rank="species">sp. nov.</taxonomicNameLabel>
, though less stout, with a little smaller basal bulb; apical orifice widely opened, inner sac not bearing spinose sclerites but with clusters of microtrichia near middle; apical lobe short, thickened at tip.
</paragraph>
<paragraph id="BD383689FF94FFCBFF09FC41FD70FC34" blockId="5.[113,1089,219,1171]" box="[144,715,987,1012]" pageId="5" pageNumber="604">
Body length:
<quantity id="7A7F9B6CFF94FFCBFEB8FC46FE09FC34" box="[289,434,988,1012]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.2" metricValueMin="8.8" pageId="5" pageNumber="604" unit="mm" value="9.5" valueMax="10.2" valueMin="8.8">8.810.2 mm</quantity>
; body width:
<quantity id="7A7F9B6CFF94FFCBFDDAFC41FD7CFC34" box="[579,711,987,1011]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" metricValueMax="4.2" metricValueMin="3.6" pageId="5" pageNumber="604" unit="mm" value="3.9" valueMax="4.2" valueMin="3.6">3.64.2 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="F59D6502FF94FFCBFFE8FC61FE47FB94" pageId="5" pageNumber="604" type="diagnosis">
<paragraph id="BD383689FF94FFCBFFE8FC61FE47FB94" blockId="5.[113,1089,219,1171]" pageId="5" pageNumber="604">
<emphasis id="8FF3EA9BFF94FFCBFFE8FC61FED6FBD4" bold="true" box="[113,365,1019,1043]" pageId="5" pageNumber="604">Differential diagnosis.</emphasis>
This new species is similar to
<taxonomicName id="7A874D0AFF94FFCBFD58FC67FC89FBD3" authorityName="Habu" authorityYear="1980" box="[705,818,1020,1044]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="604" phylum="Arthropoda" rank="species" species="miyakei">
<emphasis id="8FF3EA9BFF94FFCBFD58FC67FD6BFBD3" box="[705,720,1021,1044]" italics="true" pageId="5" pageNumber="604">T</emphasis>
.
<emphasis id="8FF3EA9BFF94FFCBFD46FC67FC89FBD3" box="[735,818,1021,1044]" italics="true" pageId="5" pageNumber="604">miyakei</emphasis>
</taxonomicName>
, but the body is smaller, the legs are paler and the aedeagus bears smaller basal bulb and narrow apical orifice armed with clusters of microtrichia (
<figureCitation id="25BC2A0CFF94FFCBFE34FBA1FE54FB94" box="[429,495,1083,1107]" captionStart="Figs" captionStartId="3.[111,150,1072,1091]" captionTargetBox="[139,1068,228,1054]" captionTargetId="figure-14@3.[131,1070,219,1069]" captionTargetPageId="3" captionText="Figs. 69. Male genitalia. 6 Trichotichnus (Pseudotrichotichnus) vietnamensis sp. nov.; 7 T. (P.) tenuitibialis sp. nov.; 8 T. (P.) niger Louwerens, 1951; 9 T. (P.) miyakei Habu, 1980. Scale bar: 1 mm. Abbreviations: l lateral aspect, d dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.5320912" httpUri="https://zenodo.org/record/5320912/files/figure.png" pageId="5" pageNumber="604">Fig. 6</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F59D6502FF94FFCBFFE8FBC6FCDEFBB4" box="[113,869,1115,1140]" pageId="5" pageNumber="604" type="etymology">
<paragraph id="BD383689FF94FFCBFFE8FBC6FCDEFBB4" blockId="5.[113,1089,219,1171]" box="[113,869,1115,1140]" pageId="5" pageNumber="604">
<emphasis id="8FF3EA9BFF94FFCBFFE8FBC6FF49FBB3" bold="true" box="[113,242,1116,1140]" pageId="5" pageNumber="604">Etymology.</emphasis>
The species name is derived from country of
<typeStatus id="623C882BFF94FFCBFD42FBC6FCB2FBB3" box="[731,777,1116,1140]" pageId="5" pageNumber="604">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="F59D6502FF94FFCBFFE8FBE1FED0FB54" box="[113,363,1147,1171]" pageId="5" pageNumber="604" type="distribution">
<paragraph id="BD383689FF94FFCBFFE8FBE1FED0FB54" blockId="5.[113,1089,219,1171]" box="[113,363,1147,1171]" pageId="5" pageNumber="604">
<emphasis id="8FF3EA9BFF94FFCBFFE8FBE1FEBFFB54" bold="true" box="[113,260,1147,1171]" pageId="5" pageNumber="604">Distribution.</emphasis>
<collectingCountry id="C5907619FF94FFCBFE90FBE1FEDCFB54" box="[265,359,1147,1171]" name="Vietnam" pageId="5" pageNumber="604">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>