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<document id="F17079F5370077C3C9E2C362A00E7AAB" ID-CLB-Dataset="4642" ID-DOI="10.5281/zenodo.5320906" ID-GBIF-Dataset="f33526c4-a22f-4c32-9fd9-34a1255f44eb" ID-ISSN="0374-1036" ID-Zenodo-Dep="5320906" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630136926086" checkinUser="felipe" docAuthor="Ito, Noboru" docDate="2009" docId="352E879FFF90FFCAFEADFDFEFB9DFD14" docLanguage="en" docName="ActaEntMusNatPra.2009.49.2.599-606.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 49 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Trichotichnus (Pseudotrichotichnus) jelineki Ito 2009, sp. nov." docType="treatment" docVersion="7" lastPageNumber="603" masterDocId="C917FFE7FF91FFCEFF99FF9AFFBBFFC7" masterDocTitle="Note on the subgenus Pseudotrichotichnus of the genus Trichotichnus from Southeast Asia (Coleoptera: Carabidae: Harpalini)" masterLastPageNumber="606" masterPageNumber="599" pageNumber="600" updateTime="1699168252378" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="B4073C50491441BC337C58B1D6F699AA" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="66DA0AA800299FB4DCBD487071B3763A">
<mods:title id="EBA520B3B4CB7C5EE4C20B269D49C44A">Note on the subgenus Pseudotrichotichnus of the genus Trichotichnus from Southeast Asia (Coleoptera: Carabidae: Harpalini)</mods:title>
</mods:titleInfo>
<mods:name id="FBA635D7B1703A277736B180E077F4EC" type="personal">
<mods:role id="2895FBF75638681FC40070881271ACAD">
<mods:roleTerm id="6B554E206096459CF3330390558BB55A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6FBC46C0F9AEE08CBE25A498DB31B558">Ito, Noboru</mods:namePart>
</mods:name>
<mods:typeOfResource id="B9D2F968FD7758C4D7A8E44EB9E358E9">text</mods:typeOfResource>
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<mods:title id="F33D0E313590E72949AE742620C93FE8">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="A2581DF0715CF6A59B34CAA48094F964">
<mods:date id="19D2CA86CFC84BC51112DC9040B7993F">2009</mods:date>
<mods:detail id="D05F9F60F194D10DBC1DE87279CB3DD4" type="pubDate">
<mods:number id="8A3E2F59EF1D4365B1618DECAD563B66">2009-12-15</mods:number>
</mods:detail>
<mods:detail id="91AA269424E78C707DB596D079A170B3" type="volume">
<mods:number id="854E4290F18C377323B10CBAC7B98A3B">49</mods:number>
</mods:detail>
<mods:detail id="75BE8EFF078A3185B29B2EA574DC6EFD" type="issue">
<mods:number id="7034E1F7FCC39C12FE2BFA60B3AECBEB">2</mods:number>
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<mods:extent id="61FD768870AF96EA603AE5E9C4E37559" unit="page">
<mods:start id="7560B0D9799B8D65B6B8150C32492024">599</mods:start>
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</mods:extent>
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<mods:classification id="4F6BE82699BF7CEE1CF73E6575E0E5A7">journal article</mods:classification>
<mods:identifier id="E8040843DA65AE7C1763781E83DFA7B0" type="CLB-Dataset">4642</mods:identifier>
<mods:identifier id="EB1DB215DC8D4017888FC1F63481EF47" type="DOI">10.5281/zenodo.5320906</mods:identifier>
<mods:identifier id="382C2A4C1A221065242C730DDD58371F" type="GBIF-Dataset">f33526c4-a22f-4c32-9fd9-34a1255f44eb</mods:identifier>
<mods:identifier id="CD81FA51C82610E078CE54607DDB293E" type="ISSN">0374-1036</mods:identifier>
<mods:identifier id="D4AAA5B1EE1B87B06EB67A0BBDB0D884" type="Zenodo-Dep">5320906</mods:identifier>
</mods:mods>
<treatment id="352E879FFF90FFCAFEADFDFEFB9DFD14" ID-DOI="http://doi.org/10.5281/zenodo.5342754" ID-GBIF-Taxon="185202416" ID-Zenodo-Dep="5342754" LSID="urn:lsid:plazi:treatment:352E879FFF90FFCAFEADFDFEFB9DFD14" httpUri="http://treatment.plazi.org/id/352E879FFF90FFCAFEADFDFEFB9DFD14" lastPageId="4" lastPageNumber="603" pageId="1" pageNumber="600">
<subSubSection id="F59D6502FF90FFCFFEADFDFEFCC5FDBA" box="[308,894,612,637]" pageId="1" pageNumber="600" type="nomenclature">
<paragraph id="BD383689FF90FFCFFEADFDFEFCC5FDBA" blockId="1.[308,894,612,637]" box="[308,894,612,637]" pageId="1" pageNumber="600">
<heading id="E67081E5FF90FFCFFEADFDFEFCC5FDBA" bold="true" box="[308,894,612,637]" centered="true" fontSize="8" level="2" pageId="1" pageNumber="600" reason="3">
<emphasis id="8FF3EA9BFF90FFCFFEADFDFEFCC5FDBA" bold="true" box="[308,894,612,637]" pageId="1" pageNumber="600">
<taxonomicName id="7A874D0AFF90FFCFFEADFDFEFC99FDBA" ID-CoL="58LGZ" authority="Ito, 2009" authorityName="Ito" authorityYear="2009" box="[308,802,612,637]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="600" phylum="Arthropoda" rank="species" species="jelineki" status="sp. nov." subGenus="Pseudotrichotichnus">
<emphasis id="8FF3EA9BFF90FFCFFEADFDFEFE74FDBA" bold="true" box="[308,463,612,637]" italics="true" pageId="1" pageNumber="600">Trichotichnus</emphasis>
(
<emphasis id="8FF3EA9BFF90FFCFFE47FDFEFD7AFDBA" bold="true" box="[478,705,612,637]" italics="true" pageId="1" pageNumber="600">Pseudotrichotichnus</emphasis>
)
<emphasis id="8FF3EA9BFF90FFCFFD48FDFEFC99FDBA" bold="true" box="[721,802,612,637]" italics="true" pageId="1" pageNumber="600">jelineki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="94C057E0FF90FFCFFCB0FDFFFCC5FDBA" box="[809,894,613,637]" pageId="1" pageNumber="600" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F59D6502FF90FFCAFDB3FD19FD45FDD4" lastPageId="4" lastPageNumber="603" pageId="1" pageNumber="600" type="description">
<paragraph id="BD383689FF90FFCFFDB3FD19FE61FD32" blockId="1.[113,1090,643,758]" pageId="1" pageNumber="600">
(
<figureCitation id="25BC2A0CFF90FFCFFDA9FD19FD3AFD51" box="[560,641,643,662]" captionStart="Figs" captionStartId="2.[153,192,1499,1518]" captionTargetBox="[168,1126,250,1426]" captionTargetId="figure-10@2.[151,1131,219,1426]" captionTargetPageId="2" captionText="Figs. 1–5. 1–2 – Trichotichnus (Pseudtrichotichnus) jelineki sp. nov., holotype (1 – habitus; 2 – male genitalia); 3 – T. (P.) vietnamensis sp. nov.; 4 – T. (P.) tenuitibialis sp. nov.; 5 – T. (P.) niger Louwerens, 1951. Scale bar: 1 mm. Abbreviations: l – lateral aspect, d – dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.5320908" httpUri="https://zenodo.org/record/5320908/files/figure.png" pageId="1" pageNumber="600">Figs. 1–2</figureCitation>
)
<emphasis id="8FF3EA9BFF90FFCFFFE8FD37FF4EFD07" bold="true" box="[113,245,684,704]" pageId="1" pageNumber="600">Type material.</emphasis>
<typeStatus id="623C882BFF90FFCFFF63FD37FEF6FD78" box="[250,333,685,704]" pageId="1" pageNumber="600" type="holotype">HOLOTYPE</typeStatus>
: J, ‘Nong Lom (lake) env., N 15˚02′ E 106˚35′, alt.
<quantity id="7A7F9B6CFF90FFCFFC91FD37FC80FD07" box="[776,827,685,704]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="600" unit="m" value="800.0">800m</quantity>
,
<quantity id="7A7F9B6CFF90FFCFFCDAFD37FCC3FD07" box="[835,888,685,704]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="600" unit="km" value="15.0">15 km</quantity>
SE of Ban Houaykong, Bolaven Plateau,
<collectingRegion id="7F43F86BFF90FFCFFE9EFD52FECDFD1C" box="[263,374,712,731]" country="Laos" name="Attapu" pageId="1" pageNumber="600">Attapu prov.</collectingRegion>
, South
<collectingCountry id="C5907619FF90FFCFFE2CFD52FE5AFD1C" box="[437,481,712,731]" name="Laos" pageId="1" pageNumber="600">Laos</collectingCountry>
,
<date id="C9391049FF90FFCFFE73FD52FDD6FD1C" box="[490,621,712,731]" pageId="1" pageNumber="600" value="1999-04-18" valueMax="1999-04-30" valueMin="1999-04-18">18-30.IV. 1999</date>
, E. Jendek &amp; O. Šauša leg.’ (OMNH).
<typeStatus id="623C882BFF90FFCFFC24FD52FBAEFD1E" box="[957,1045,712,731]" pageId="1" pageNumber="600" type="paratype">PARATYPES</typeStatus>
: 1 J
<specimenCount id="AB81FD00FF90FFCFFFE8FD78FF10FD32" box="[113,171,732,757]" pageId="1" pageNumber="600" type="female">10 ♀♀</specimenCount>
, same data as the
<typeStatus id="623C882BFF90FFCFFEDEFD78FE29FD32" box="[327,402,738,757]" pageId="1" pageNumber="600" type="holotype">holotype</typeStatus>
(NIKJ).
</paragraph>
<paragraph id="BD383689FF90FFCFFFE8FC97FE7BFCA2" blockId="1.[113,1089,781,1573]" pageId="1" pageNumber="600">
<emphasis id="8FF3EA9BFF90FFCFFFE8FC97FF47FCE2" bold="true" box="[113,252,781,805]" pageId="1" pageNumber="600">Description.</emphasis>
Body oblong, pitchy black, very shiny, with iridescent lustre on elytra; labial and maxillary palpi, antennae and tarsi light reddish brown, tibiae and femora dark reddish brown, labrum blackish brown.
</paragraph>
<paragraph id="BD383689FF90FFCFFF09FCF7FE40FB22" blockId="1.[113,1089,781,1573]" pageId="1" pageNumber="600">Head gently elevated on vertex, large, 0.75–0.78 times as wide as pronotum, with sparsely scattered, minute punctures; labrum subtrapezoidal, triangularly emarginate at apex; clypeus gently emarginate apically, with several obscure rugosities near each lateral seta; clypeal suture almost straight and deep throughout; frontal impressions also deep till supraorbital grooves, linearly oblique; eyes large, fairly prominent; temples short, steeply contracted behind; space between buccal fissure and genuine ventral margins of eyes fairly wide; antennae short, not surpassing elytral base, segment 3 pubescent in apical three fourths, as long as segment 4 and 1.8 times as long as segment 2; palpi slender, segment 3 of labial palpus 0.9 times as long as segment 2; ligula narrow, parallel-sided, with apical angles rectangular; paraglossae narrowly fan-shaped; mentum sharply toothed at middle of apex, epilobes not widened apicad; microsculpture visible as isodiametric meshes in apical half of clypeus, partly as vague transverse meshes on the other parts.</paragraph>
<paragraph id="BD383689FF90FFCFFF09FB74FCF6F9E2" blockId="1.[113,1089,781,1573]" pageId="1" pageNumber="600">Pronotum transversely subquadrate, widest at apical two fifths, 1.44–1.46 times as wide as long, sides arcuate in apical third, thence linearly oblique to base; surface moderately convex, largely smooth, sparsely covered with minute and somewhat coarse punctures only on basal foveae; anterior margin shallowly and obtrapezoidally concave, with thin border throughout; base 1.2 times as wide as apex, hardly bisinuate, with no border in middle; apical angles widely rounded, slightly protruding; basal angles obtuse, angularly rounded; lateral furrows impressed in a line throughout; basal foveae shallow, each with short groove at inner side; front and hind transverse impressions almost obsolete; median line thin, not reaching both anterior margin and base, with several minute punctures; microsculpture consisting of fine transverse meshes on disc and of isodiametric meshes in basal areas.</paragraph>
<caption id="E9F86601FF93FFCCFF00FA41FDC8F9E3" ID-DOI="http://doi.org/10.5281/zenodo.5320908" ID-Zenodo-Dep="5320908" httpUri="https://zenodo.org/record/5320908/files/figure.png" pageId="2" pageNumber="601" startId="2.[153,192,1499,1518]" targetBox="[168,1126,250,1426]" targetPageId="2">
<paragraph id="BD383689FF93FFCCFF00FA41FDC8F9E3" blockId="2.[153,1129,1499,1572]" pageId="2" pageNumber="601">
Figs. 1–5. 1–2 –
<taxonomicName id="7A874D0AFF93FFCCFEB7FA46FD18FA29" authority="Ito, 2009" authorityName="Ito" authorityYear="2009" box="[302,675,1499,1518]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="601" phylum="Arthropoda" rank="species" species="jelineki" status="sp. nov." subGenus="Pseudtrichotichnus">
<emphasis id="8FF3EA9BFF93FFCCFEB7FA46FE18FA29" box="[302,419,1500,1518]" italics="true" pageId="2" pageNumber="601">Trichotichnus</emphasis>
(
<emphasis id="8FF3EA9BFF93FFCCFE28FA46FDEDFA29" box="[433,598,1500,1518]" italics="true" pageId="2" pageNumber="601">Pseudtrichotichnus</emphasis>
)
<emphasis id="8FF3EA9BFF93FFCCFDFDFA46FD18FA29" box="[612,675,1500,1518]" italics="true" pageId="2" pageNumber="601">jelineki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="94C057E0FF93FFCCFD33FA41FD4BFA29" box="[682,752,1499,1518]" pageId="2" pageNumber="601" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (1 – habitus; 2 – male genitalia); 3 –
<taxonomicName id="7A874D0AFF93FFCCFF23FA6DFED9F9CE" authority="Ito, 2009" authorityName="Ito" authorityYear="2009" box="[186,354,1526,1545]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="601" phylum="Arthropoda" rank="species" species="vietnamensis" status="sp. nov." subGenus="Pseudotrichotichnus">
<emphasis id="8FF3EA9BFF93FFCCFF23FA6DFF70F9CE" box="[186,203,1527,1545]" italics="true" pageId="2" pageNumber="601">T.</emphasis>
(
<emphasis id="8FF3EA9BFF93FFCCFF4EFA6DFF53F9CE" box="[215,232,1527,1545]" italics="true" pageId="2" pageNumber="601">P.</emphasis>
)
<emphasis id="8FF3EA9BFF93FFCCFF6DFA6DFED9F9CE" box="[244,354,1527,1545]" italics="true" pageId="2" pageNumber="601">vietnamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="94C057E0FF93FFCCFEF1FA6CFE15F9CE" box="[360,430,1526,1545]" pageId="2" pageNumber="601" rank="species">sp. nov.</taxonomicNameLabel>
; 4 –
<taxonomicName id="7A874D0AFF93FFCCFE41FA6DFDC7F9CE" authority="Ito, 2009" authorityName="Ito" authorityYear="2009" box="[472,636,1526,1545]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="602" phylum="Arthropoda" rank="species" species="tenuitibialis" status="sp. nov." subGenus="Pseudotrichotichnus">
<emphasis id="8FF3EA9BFF93FFCCFE41FA6DFE5FF9CE" box="[472,484,1527,1545]" italics="true" pageId="2" pageNumber="601">T</emphasis>
. (
<emphasis id="8FF3EA9BFF93FFCCFE6FFA6DFDB8F9CE" box="[502,515,1527,1545]" italics="true" pageId="2" pageNumber="601">P</emphasis>
.)
<emphasis id="8FF3EA9BFF93FFCCFD8FFA6DFDC7F9CE" box="[534,636,1527,1545]" italics="true" pageId="2" pageNumber="601">tenuitibialis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="94C057E0FF93FFCCFD18FA6CFD7CF9CE" box="[641,711,1526,1545]" pageId="2" pageNumber="601" rank="species">sp. nov.</taxonomicNameLabel>
; 5 –
<taxonomicName id="7A874D0AFF93FFCCFD68FA6DFC42F9CF" authority="Louwerens, 1951" authorityName="Louwerens" authorityYear="1951" box="[753,1017,1525,1545]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="602" phylum="Arthropoda" rank="species" species="niger" subGenus="Pseudotrichotichnus">
<emphasis id="8FF3EA9BFF93FFCCFD68FA6DFD46F9CE" box="[753,765,1527,1545]" italics="true" pageId="2" pageNumber="601">T</emphasis>
. (
<emphasis id="8FF3EA9BFF93FFCCFC96FA6DFCA7F9CE" box="[783,796,1527,1545]" italics="true" pageId="2" pageNumber="601">P</emphasis>
.)
<emphasis id="8FF3EA9BFF93FFCCFCB6FA6DFCE7F9CE" box="[815,860,1527,1545]" italics="true" pageId="2" pageNumber="601">niger</emphasis>
Louwerens, 1951
</taxonomicName>
. Scale bar: 1 mm. Abbreviations: l – lateral aspect, d – dorsal aspect.
</paragraph>
</caption>
<caption id="E9F86601FF92FFCDFFF6FBAAFEFEFBBE" ID-DOI="http://doi.org/10.5281/zenodo.5320912" ID-Zenodo-Dep="5320912" httpUri="https://zenodo.org/record/5320912/files/figure.png" pageId="3" pageNumber="602" startId="3.[111,150,1072,1091]" targetBox="[139,1068,228,1054]" targetPageId="3">
<paragraph id="BD383689FF92FFCDFFF6FBAAFEFEFBBE" blockId="3.[111,1090,1072,1145]" pageId="3" pageNumber="602">
Figs. 6–9. Male genitalia. 6 –
<taxonomicName id="7A874D0AFF92FFCDFEF5FBABFCADFB84" authority="Ito, 2009" authorityName="Ito" authorityYear="2009" box="[364,790,1072,1091]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="602" phylum="Arthropoda" rank="species" species="vietnamensis" status="sp. nov." subGenus="Pseudotrichotichnus">
<emphasis id="8FF3EA9BFF92FFCDFEF5FBABFE5AFB84" box="[364,481,1073,1091]" italics="true" pageId="3" pageNumber="602">Trichotichnus</emphasis>
(
<emphasis id="8FF3EA9BFF92FFCDFE74FBABFD27FB84" box="[493,668,1073,1091]" italics="true" pageId="3" pageNumber="602">Pseudotrichotichnus</emphasis>
)
<emphasis id="8FF3EA9BFF92FFCDFD31FBABFCADFB84" box="[680,790,1073,1091]" italics="true" pageId="3" pageNumber="602">vietnamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="94C057E0FF92FFCDFC82FBAAFCDBFB84" box="[795,864,1072,1091]" pageId="3" pageNumber="602" rank="species">sp. nov.</taxonomicNameLabel>
; 7 –
<taxonomicName id="7A874D0AFF92FFCDFC1FFBABFB98FB84" authority="Ito, 2009" authorityName="Ito" authorityYear="2009" box="[902,1059,1072,1091]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="602" phylum="Arthropoda" rank="species" species="tenuitibialis" status="sp. nov." subGenus="Pseudotrichotichnus">
<emphasis id="8FF3EA9BFF92FFCDFC1FFBABFC2CFB84" box="[902,919,1073,1091]" italics="true" pageId="3" pageNumber="602">T.</emphasis>
(
<emphasis id="8FF3EA9BFF92FFCDFC3BFBABFC08FB84" box="[930,947,1073,1091]" italics="true" pageId="3" pageNumber="602">P.</emphasis>
)
<emphasis id="8FF3EA9BFF92FFCDFC24FBABFB98FB84" box="[957,1059,1073,1091]" italics="true" pageId="3" pageNumber="602">tenuitibialis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="94C057E0FF92FFCDFBB1FBAAFF2CFB99" pageId="3" pageNumber="602" rank="species">sp. nov.</taxonomicNameLabel>
; 8 –
<taxonomicName id="7A874D0AFF92FFCDFF26FBD1FE7BFB9A" authority="Louwerens, 1951" authorityName="Louwerens" authorityYear="1951" box="[191,448,1098,1118]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="602" phylum="Arthropoda" rank="species" species="niger" subGenus="Pseudotrichotichnus">
<emphasis id="8FF3EA9BFF92FFCDFF26FBD1FF6BFB9A" box="[191,208,1099,1117]" italics="true" pageId="3" pageNumber="602">T.</emphasis>
(
<emphasis id="8FF3EA9BFF92FFCDFF42FBD1FF57FB9A" box="[219,236,1099,1117]" italics="true" pageId="3" pageNumber="602">P.</emphasis>
)
<emphasis id="8FF3EA9BFF92FFCDFF6EFBD1FE9FFB9A" box="[247,292,1099,1117]" italics="true" pageId="3" pageNumber="602">niger</emphasis>
Louwerens, 1951
</taxonomicName>
; 9 –
<taxonomicName id="7A874D0AFF92FFCDFE70FBD1FD74FB99" authority="Habu, 1980" authorityName="Habu" authorityYear="1980" box="[489,719,1098,1118]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="602" phylum="Arthropoda" rank="species" species="miyakei" subGenus="Pseudotrichotichnus">
<emphasis id="8FF3EA9BFF92FFCDFE70FBD1FE41FB9A" box="[489,506,1099,1117]" italics="true" pageId="3" pageNumber="602">T.</emphasis>
(
<emphasis id="8FF3EA9BFF92FFCDFD9CFBD1FDADFB9A" box="[517,534,1099,1117]" italics="true" pageId="3" pageNumber="602">P.</emphasis>
)
<emphasis id="8FF3EA9BFF92FFCDFDB8FBD1FDD8FB9A" box="[545,611,1099,1117]" italics="true" pageId="3" pageNumber="602">miyakei</emphasis>
Habu, 1980
</taxonomicName>
. Scale bar: 1 mm. Abbreviations: l – lateral aspect, d – dorsal aspect.
</paragraph>
</caption>
<paragraph id="BD383689FF92FFCDFF09FB2FFE12FA0A" blockId="3.[113,1089,1205,1613]" pageId="3" pageNumber="602">Elytra oblong-oval, 1.25 times as wide as pronotum, nearly 1.5 times as long as wide, gently convex, impunctate; sides gently curved behind humeri, very weakly arcuate around midlength, rather steeply arcuate in apical one fifth; apices widely arcuate at sides, narrowly rounded at tips which are close to each other; base of each elytron very shallowly emarginate, rounded at humeral angles; striae deep, rather wide, and clearly crenulate, scutellar striole moderately long; intervals barely convex on disc, gradually becoming more convex apicad and basad, interval 3 with setiferous pore between apical third and two-fifths; marginal series consisting of (7–8) + (10–11) umbilicate pores; microsculpture observable as vague transverse lines. Hind wings developed.</paragraph>
<paragraph id="BD383689FF92FFCDFF09FA4FFE25F98A" blockId="3.[113,1089,1205,1613]" pageId="3" pageNumber="602">Ventral surface mostly smooth, sparsely punctate on mesepisterna; metepisterna elongate, 1.5 times as long as wide; metepisterna elongate, 1.55–1.60 times as long as wide; abdominal sternite 6 with two setae on each side in both sexes, truncate or feebly emarginate in male and fairly arcuate in female.</paragraph>
<paragraph id="BD383689FF95FFCAFF2CFF46FEE7FE54" blockId="4.[151,1127,219,724]" pageId="4" pageNumber="603">Legs moderately long; fore tibiae more or less dilated apicad, clearly sulcate in basal four-fifths, with two or three spines near apex along each external margin; hind tarsi short, 0.78–0.80 times as long as the width of head including eyes in male and 0.74–0.76 times as long in female, segment 1 0.92 times as long as segments 2 and 3 taken together, segment 3 0.85 times as long as segment 2 and 1.2 times as long as segment 4, all claws with two setae on ventral margin.</paragraph>
<paragraph id="BD383689FF95FFCAFF2CFE01FE41FE34" blockId="4.[151,1127,219,724]" pageId="4" pageNumber="603">
Aedeagus (
<figureCitation id="25BC2A0CFF95FFCAFEB6FE01FECAFE73" box="[303,369,411,436]" captionStart="Figs" captionStartId="2.[153,192,1499,1518]" captionTargetBox="[168,1126,250,1426]" captionTargetId="figure-10@2.[151,1131,219,1426]" captionTargetPageId="2" captionText="Figs. 1–5. 1–2 – Trichotichnus (Pseudtrichotichnus) jelineki sp. nov., holotype (1 – habitus; 2 – male genitalia); 3 – T. (P.) vietnamensis sp. nov.; 4 – T. (P.) tenuitibialis sp. nov.; 5 – T. (P.) niger Louwerens, 1951. Scale bar: 1 mm. Abbreviations: l – lateral aspect, d – dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.5320908" httpUri="https://zenodo.org/record/5320908/files/figure.png" pageId="4" pageNumber="603">Fig. 2</figureCitation>
) robust, clearly arcuate dorsally, thinned at apex, knob-shaped at tip; apical orifice wide, inner sac armed with small spines near basal bulb; apical lobe elongate, about 1.50 times as long as wide.
</paragraph>
<paragraph id="BD383689FF95FFCAFF2CFE61FD45FDD4" blockId="4.[151,1127,219,724]" box="[181,766,507,532]" pageId="4" pageNumber="603">
Body length:
<quantity id="7A7F9B6CFF95FFCAFEDEFE66FE5EFDD4" box="[327,485,508,532]" metricMagnitude="-2" metricUnit="m" metricValue="1.065" metricValueMax="1.09" metricValueMin="1.04" pageId="4" pageNumber="603" unit="mm" value="10.65" valueMax="10.9" valueMin="10.4">10.4–10.9 mm</quantity>
; body width: 3.8–4.0 mm.
</paragraph>
</subSubSection>
<subSubSection id="F59D6502FF95FFCAFF0EFD81FC46FDB3" pageId="4" pageNumber="603" type="diagnosis">
<paragraph id="BD383689FF95FFCAFF0EFD81FC46FDB3" blockId="4.[151,1127,219,724]" pageId="4" pageNumber="603">
<emphasis id="8FF3EA9BFF95FFCAFF0EFD81FE2AFDF4" bold="true" box="[151,401,539,563]" pageId="4" pageNumber="603">Differential diagnosis.</emphasis>
This new species is closely allied to
<taxonomicName id="7A874D0AFF95FFCAFC83FD87FC09FDF3" authorityName="Habu" authorityYear="1980" box="[794,946,540,564]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="603" phylum="Arthropoda" rank="species" species="miyakei" subGenus="Pseudotrichotichnus">
<emphasis id="8FF3EA9BFF95FFCAFC83FD87FC95FDF3" box="[794,814,541,564]" italics="true" pageId="4" pageNumber="603">T.</emphasis>
(
<emphasis id="8FF3EA9BFF95FFCAFCA4FD87FCEAFDF3" box="[829,849,541,564]" italics="true" pageId="4" pageNumber="603">P.</emphasis>
)
<emphasis id="8FF3EA9BFF95FFCAFCC6FD87FC09FDF3" box="[863,946,541,564]" italics="true" pageId="4" pageNumber="603">miyakei</emphasis>
</taxonomicName>
, but is distinguishable by the pronotal foveae more clearly depressed, the hind tarsi longer and the aedeagus armed with three small sclerites (
<figureCitation id="25BC2A0CFF95FFCAFE60FDC1FD81FDB3" box="[505,570,603,628]" captionStart="Figs" captionStartId="2.[153,192,1499,1518]" captionTargetBox="[168,1126,250,1426]" captionTargetId="figure-10@2.[151,1131,219,1426]" captionTargetPageId="2" captionText="Figs. 1–5. 1–2 – Trichotichnus (Pseudtrichotichnus) jelineki sp. nov., holotype (1 – habitus; 2 – male genitalia); 3 – T. (P.) vietnamensis sp. nov.; 4 – T. (P.) tenuitibialis sp. nov.; 5 – T. (P.) niger Louwerens, 1951. Scale bar: 1 mm. Abbreviations: l – lateral aspect, d – dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.5320908" httpUri="https://zenodo.org/record/5320908/files/figure.png" pageId="4" pageNumber="603">Fig. 2</figureCitation>
), which are missing in
<taxonomicName id="7A874D0AFF95FFCAFCABFDC7FC1BFDB3" authorityName="Habu" authorityYear="1980" box="[818,928,605,628]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="603" phylum="Arthropoda" rank="species" species="miyakei">
<emphasis id="8FF3EA9BFF95FFCAFCABFDC7FC1BFDB3" box="[818,928,605,628]" italics="true" pageId="4" pageNumber="603">T. miyakei</emphasis>
</taxonomicName>
(
<figureCitation id="25BC2A0CFF95FFCAFC37FDC1FC4BFDB3" box="[942,1008,603,628]" captionStart="Figs" captionStartId="3.[111,150,1072,1091]" captionTargetBox="[139,1068,228,1054]" captionTargetId="figure-14@3.[131,1070,219,1069]" captionTargetPageId="3" captionText="Figs. 6–9. Male genitalia. 6 – Trichotichnus (Pseudotrichotichnus) vietnamensis sp. nov.; 7 – T. (P.) tenuitibialis sp. nov.; 8 – T. (P.) niger Louwerens, 1951; 9 – T. (P.) miyakei Habu, 1980. Scale bar: 1 mm. Abbreviations: l – lateral aspect, d – dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.5320912" httpUri="https://zenodo.org/record/5320912/files/figure.png" pageId="4" pageNumber="603">Fig. 9</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F59D6502FF95FFCAFF0EFDE6FE4DFD74" pageId="4" pageNumber="603" type="etymology">
<paragraph id="BD383689FF95FFCAFF0EFDE6FE4DFD74" blockId="4.[151,1127,219,724]" pageId="4" pageNumber="603">
<emphasis id="8FF3EA9BFF95FFCAFF0EFDE6FEA3FD53" bold="true" box="[151,280,636,660]" pageId="4" pageNumber="603">Etymology.</emphasis>
This species is dedicated to the eminent Czech coleopterist Josef Jelínek at the occasion of his 70th anniversary.
</paragraph>
</subSubSection>
<subSubSection id="F59D6502FF95FFCAFF0EFD21FB9DFD14" box="[151,1062,699,724]" pageId="4" pageNumber="603" type="distribution">
<paragraph id="BD383689FF95FFCAFF0EFD21FB9DFD14" blockId="4.[151,1127,219,724]" box="[151,1062,699,724]" pageId="4" pageNumber="603">
<emphasis id="8FF3EA9BFF95FFCAFF0EFD21FE91FD14" bold="true" box="[151,298,699,723]" pageId="4" pageNumber="603">Distribution.</emphasis>
Known only from the
<typeStatus id="623C882BFF95FFCAFD84FD26FDF0FD13" box="[541,587,700,724]" pageId="4" pageNumber="603">type</typeStatus>
locality in southern
<collectingCountry id="C5907619FF95FFCAFCBFFD26FCE1FD14" box="[806,858,700,723]" name="Laos" pageId="4" pageNumber="603">Laos</collectingCountry>
(
<collectingRegion id="7F43F86BFF95FFCAFCF1FD26FBA0FD14" box="[872,1051,699,724]" country="Laos" name="Attapu" pageId="4" pageNumber="603">Attapu province</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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