282 lines
31 KiB
XML
282 lines
31 KiB
XML
<document ID-CLB-Dataset="24300" ID-DOI="http://dx.doi.org/10.3897/zookeys.905.39592" ID-GBIF-Dataset="ee5a0817-7cfc-4c21-ba7d-3beabb06cccd" ID-PMC="PMC7004242" ID-Pensoft-Pub="1313-2970-905-1" ID-Pensoft-UUID="4C7E0CB428DF56BDB15B7AC8D6707961" ID-PubMed="32063726" ID-ZooBank="F4C766E4633A41039FFDE952718F41FB" ModsDocID="1313-2970-905-1" checkinTime="1579539902851" checkinUser="pensoft" docAuthor="Salata, Sebastian & Fisher, Brian L." docDate="2020" docId="33F9FAC726FA966693005925208B4642" docLanguage="en" docName="ZooKeys 905: 1-235" docOrigin="ZooKeys 905" docSource="http://dx.doi.org/10.3897/zookeys.905.39592" docTitle="Pheidole vohemarensis Salata & Fisher 2020, sp. nov." docType="treatment" docUuid="55119B1A-806B-491E-B551-8711DE650ACE" docUuidSource="ZooBank" docVersion="7" id="4C7E0CB428DF56BDB15B7AC8D6707961" lastPageNumber="1" masterDocId="4C7E0CB428DF56BDB15B7AC8D6707961" masterDocTitle="Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups" masterLastPageNumber="235" masterPageNumber="1" pageNumber="1" updateTime="1732649601836" updateUser="ExternalLinkService">
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<mods:titleInfo id="8C785DC7628C180E282562D86F23C901">
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<mods:title id="39026B705B15C6C9086A5903814D4A0C">Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups</mods:title>
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<mods:namePart id="0D08E21DE0920B105363BD841D916AE3">Salata, Sebastian</mods:namePart>
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<mods:namePart id="3E829A7CC96EEF6626949643DA8F8148">Fisher, Brian L.</mods:namePart>
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<mods:typeOfResource id="0605D2EF3BA61D34253E979FD39E44D3">text</mods:typeOfResource>
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<mods:title id="FDE7B21967264EBE7E1678AA689E0F45">ZooKeys</mods:title>
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<mods:part id="E82401E9CAF63B3C51F47B73471BAD0B">
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<mods:date id="B45C7FE9865C4BAE7D9AEB827DB6EFDD">2020</mods:date>
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<mods:detail id="F838F8A3F4DC9B7FDA612A9547DE2A81" type="volume">
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<mods:number id="F6A9EDE2FC6B213BE0AB5C1A7235DBC0">905</mods:number>
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<mods:start id="7627970769569D51A73DC293C433FEFD">1</mods:start>
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<mods:end id="6FB09B08264DCE5C230FCFE7E13BA892">235</mods:end>
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<mods:url id="4252034DB75BED37C2CF6C56994A5FEF">http://dx.doi.org/10.3897/zookeys.905.39592</mods:url>
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<mods:classification id="7887AC7D86EFADAF8B298968D20DA0BA">journal article</mods:classification>
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<mods:identifier id="61069CA5FFE6036BCF3E0FA09A074A2E" type="DOI">http://dx.doi.org/10.3897/zookeys.905.39592</mods:identifier>
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<mods:identifier id="F0F36B5CDFA8D0B000FA0CCFED7BBF58" type="Pensoft-Pub">1313-2970-905-1</mods:identifier>
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<treatment id="33F9FAC726FA966693005925208B4642" ID-GBIF-Taxon="161777003" LSID="urn:lsid:zoobank.org:act:55119B1A-806B-491E-B551-8711DE650ACE" httpUri="http://treatment.plazi.org/id/33F9FAC726FA966693005925208B4642" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Formicidae" scope_order="Hymenoptera">
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<subSubSection id="5A597ADA9DE327FDE1245EFA6CCCDDAD" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="5E33013DF6B7E7AE830A5C7DAF582A6E" pageId="0" pageNumber="1">
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<taxonomicName id="8FE7AF30902E15DFFA5F714EF1C5DA4B" ID-CoL="CMM3W" LSID="http://zoobank.org/55119B1A-806B-491E-B551-8711DE650ACE" authority="Salata & Fisher, 2020" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole vohemarensis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="vohemarensis" status="sp. nov.">Pheidole vohemarensis</taxonomicName>
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<taxonomicNameLabel id="F68120AD6B2FD5E6941ED65D1CA8BD14" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation id="3514D1AB31F0DC4A648B00E81AC981CC" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Pheidole vohamarensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0107947) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0107946)." figureDoi="10.3897/zookeys.905.39592.figure38" httpUri="https://binary.pensoft.net/fig/373085" pageId="0" pageNumber="1">Figs 38A-F, 85AA, 88K</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="2323A180A702303FD328C1F3033F0F44" pageId="0" pageNumber="1" type="type material">
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<paragraph id="BB0C8B850046B23BB398A807F6E52A27" pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph id="943DD75A0EE6C414DC2C7F52EED07D07" pageId="0" pageNumber="1">
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<emphasis id="22A5BB2A2E5E3479F864AC5EFD8DAFC7" bold="true" pageId="0" pageNumber="1">
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<emphasis id="0F9FC1609F50196EF3DA7607D0666739" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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.
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</emphasis>
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Madagascar. •1 major worker; Antsiranana; Ambondrobe, 41.1 km 175° Vohemar;
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<geoCoordinate id="05021390D5AF91DC7DADF462853841D5" degrees="13.71533" direction="south" orientation="latitude" precision="1" value="-13.71533">-13.71533</geoCoordinate>
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,
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<geoCoordinate id="94EFA29EFD49B23CC6B28ED3A3C54A83" degrees="50.10167" direction="east" orientation="longitude" precision="1" value="50.10167">50.10167</geoCoordinate>
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; alt. 10 m; 1 Dec 2004; Fisher et al. leg.; BLF11264, CASENT0107946 (CASC).
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<emphasis id="9C8EBD21C814AFAF5F280496ED4EC905" bold="true" pageId="0" pageNumber="1">
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<emphasis id="B9AE43F05AC951FA12A02DFBAC439742" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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.
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</emphasis>
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Madagascar. •2w., 1q.; same data as for holotype; CASENT0107947, CASENT0217986, CASENT0872084 (CASC).
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</paragraph>
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</subSubSection>
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<subSubSection id="E4A4478FEE250A9CE7A411FBA772FD7A" pageId="0" pageNumber="1" type="materials_examined">
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<paragraph id="6775841CC8A64D995104A0118DC09E40" pageId="0" pageNumber="1">Other material.</paragraph>
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<paragraph id="CDF337F212334C75695A445F54DB4F70" pageId="0" pageNumber="1">
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Madagascar. -
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<emphasis id="1E16D75142D305AE8D0B73ADFFEB7BD4" bold="true" italics="true" pageId="0" pageNumber="1">Antsiranana</emphasis>
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: •1s.; Binara Forest;
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<geoCoordinate id="DDB745BEA0AB9D5910B0447975BF47B3" degrees="13.26207" direction="south" orientation="latitude" precision="1" value="-13.26207">-13.26207</geoCoordinate>
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,
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<geoCoordinate id="440021EB61C2355B9155AF4993F3C08F" degrees="49.60505" direction="east" orientation="longitude" precision="1" value="49.60505">49.60505</geoCoordinate>
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; alt. 692 m; 20 Oct 2013; B.L. Fisher et al. leg.; CASENT0353324 (CASC). •2w., 1s.;
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<normalizedToken id="4AAF78903953550D2414CFAA733227A2" originalValue="Forêt">Foret</normalizedToken>
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<normalizedToken id="954E03522D72E32B5D87C4B67A11F077" originalValue="d’Analabe">d'Analabe</normalizedToken>
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, 30.0 km 72°ENE Daraina;
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<geoCoordinate id="121C2F4A2C0E57DBD1C99266AAB846E8" degrees="13.08333" direction="south" orientation="latitude" precision="1" value="-13.08333">-13.08333</geoCoordinate>
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,
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<geoCoordinate id="C193615A67AE1A114C83B5A1DD669A14" degrees="49.90833" direction="east" orientation="longitude" precision="1" value="49.90833">49.90833</geoCoordinate>
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; alt. 30 m; 27 Nov 2003; B.L. Fisher et al. leg.; CASENT0041430, CASENT0041449, CASENT0048846 (CASC). •1w., 2s.;
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<normalizedToken id="16A62935B00B947BF3115E6E7EC32F59" originalValue="Forêt">Foret</normalizedToken>
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de Bekaraoka, 6.8 km 60°ENE Daraina; -13.16667, 49.71; alt. 150 m; 7 Dec 2003; B.L. Fisher et al. leg.; CASENT0044312, CASENT0044422, CASENT0044438 (CASC). •2w., 2s.;
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<normalizedToken id="0174E09A843C4AE4EC33C200603181D6" originalValue="Forêt">Foret</normalizedToken>
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de Binara, 7.5 km 230°SW Daraina;
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<geoCoordinate id="89211480BE3E229709A0EC6F5D22C4BD" degrees="13.255" direction="south" orientation="latitude" precision="55" value="-13.255">-13.255</geoCoordinate>
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,
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<geoCoordinate id="8474C0D8A4283BC952511CC2ADCFC637" degrees="49.61667" direction="east" orientation="longitude" precision="1" value="49.61667">49.61667</geoCoordinate>
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; alt. 375 m; 1 Dec 2003; B.L. Fisher et al. leg.; CASENT0041832, CASENT0041833, CASENT0041890, CASENT0041895 (CASC). •1s.; Parc National de Marojejy, Manantenina River, 27.6 km 35°NE Andapa, 9.6 km 327°NNW Manantenina; -14.435, 49.76; alt. 775 m; 15 Nov 2003; B.L. Fisher et al. leg.; CASENT0045241 (CASC). •1w.;
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<normalizedToken id="D96E1EA59598C43E5EEC14FDAA2949A7" originalValue="Rés">Res</normalizedToken>
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. Analamerana, 28.4 km 99° Anivorano-Nord;
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<geoCoordinate id="63464580CB4FDEE86FF661B8EA12113A" degrees="12.74667" direction="south" orientation="latitude" precision="1" value="-12.74667">-12.74667</geoCoordinate>
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,
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<geoCoordinate id="ED9C9AB89E2785948FE554FDB5C4CB1B" degrees="49.49483" direction="east" orientation="longitude" precision="1" value="49.49483">49.49483</geoCoordinate>
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; alt. 60 m; 5 Dec 2004; B.L. Fisher et al. leg.; CASENT0054163 (CASC).
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</paragraph>
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</subSubSection>
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<subSubSection id="B13094A30C2E561DDA4F12F79D39979E" pageId="0" pageNumber="1" type="diagnosis">
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<paragraph id="9DFC847124EA768CEA9F6DF822991F16" pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph id="A6784BADBE96416931D720CECF1ED85C" pageId="0" pageNumber="1">
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Head, in full-face view, cordate, widened posteriorly; sides of the head with dense, long, suberect to erect pilosity; antennal scrobes present, impressed, and indistinctly delimited ventrally and posteriorly by carinulae; scrobe surface foveolate, with distinct, thick, moderately sparse, longitudinal rugae; propodeal spines long (PSL: 0.19-0.23 (0.21)); gaster shagreened; inner hypostomal teeth distinct, high, narrow, and triangular, closely spaced, with rounded apex; outer hypostomal teeth slightly lower and thinner than inner hypostomal teeth, lobe-like.
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<emphasis id="C3651930FD1D8A2C8A5CA4C51F304E46" bold="true" pageId="0" pageNumber="1">
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<emphasis id="E7F8191E3850E13884827CB62C05398C" italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
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.
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</emphasis>
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Head foveolate, genae with smooth notch; propodeal spines long (PSL: 0.12-0.14 (0.13)); promesonotum low, slightly convex, short, with posterior declivity relatively steep; mesosoma foveolate, katepisternum and mesonotum with smooth notches.
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</paragraph>
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</subSubSection>
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<subSubSection id="1D7BFF921DE34AAE85FE133C6C0BA79A" pageId="0" pageNumber="1" type="description">
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<paragraph id="D01F8CFF3E7812E51B08AF841D3F5E2A" pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph id="351B07DDE90B9D24906C4CFBD1022A0F" pageId="0" pageNumber="1">
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<emphasis id="E271B29E827426B8E1FAA16D3F1778B6" bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
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Measurements (
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<emphasis id="A8D4780BD8CCFD864505C45BBBDF354E" italics="true" pageId="0" pageNumber="1">N</emphasis>
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= 10): HL: 1.03-1.11 (1.08); HW: 0.99-1.1 (1.05); SL: 0.49-0.57 (0.52); EL: 0.12-0.14 (0.13); WL: 0.83-0.91 (0.87); PSL: 0.19-0.23 (0.21); MTL: 0.48-0.54 (0.51); PNW: 0.48-0.57 (0.53); PTW: 0.13-0.17 (0.15); PPW: 0.4-0.51 (0.47); CI: 95.1-99.3 (98.0); SI: 47.1-53.1 (49.7); PSLI: 18.2-20.8 (19.1); PPI: 29.1-36.1 (33.1); PNI: 45.9-52.9 (49.9); MTI: 44.8-50.8 (48.8).
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<emphasis id="809A636E4CEA96A01AE3B98B6C0A6398" bold="true" pageId="0" pageNumber="1">
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<emphasis id="E2BAFD460E4C4696E3A5982F96455BD7" italics="true" pageId="0" pageNumber="1">Head</emphasis>
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.
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</emphasis>
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In full-face view cordate, slightly longer than wide, anterior of eyes slightly convex, posterior of eyes convex, occipital margins of lobes convex (Fig.
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<figureCitation id="3736505610161A7ADD70271241A0DC58" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Pheidole vohamarensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0107947) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0107946)." figureDoi="10.3897/zookeys.905.39592.figure38" httpUri="https://binary.pensoft.net/fig/373085" pageId="0" pageNumber="1">38B</figureCitation>
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). In lateral view sub-oval; ventral and dorsal faces convex; dorsal face not depressed posteriorly; inner hypostomal teeth visible. Sides of the head with dense, long, suberect to erect pilosity; whole head with dense, short, suberect to erect pilosity. Antennal scrobes impressed and indistinctly delimited ventrally and posteriorly by carinulae; scrobe surface foveolate, with distinct, thick, moderately sparse, longitudinal rugae. Occipital lobes shiny, with thick, sparse, irregular rugae, rugae weakening posteriorly, interspaces with fine and dense rugulae; frons with moderately sparse, thick, longitudinal rugae, on the posterior part rugae longitudinal to irregular, interspaces with fine and dense rugulae, sometimes rugulae fading on the central part of frons; genae with dense, moderately thick, and irregular rugulae, interspaces with dense and fine rugulae; malar area with thick, sparse, longitudinal rugae, interspaces smooth to indistinctly rugulose. Centre of clypeus shiny and with indistinct, short rugulae, lateral sides with longitudinal rugae; median notch present, moderately wide and shallow; median longitudinal carina present; lateral longitudinal carinae present. Scape, when laid back, slightly exceeding the midlength of head; pilosity suberect to erect (Fig.
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<figureCitation id="1A732E53D00875D359328DB5A5BD5E8A" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Pheidole vohamarensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0107947) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0107946)." figureDoi="10.3897/zookeys.905.39592.figure38" httpUri="https://binary.pensoft.net/fig/373085" pageId="0" pageNumber="1">38B, D</figureCitation>
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). Inner hypostomal teeth distinct, high, narrow, and triangular, closely spaced, with rounded apex; outer hypostomal teeth slightly lower and thinner than inner hypostomal teeth, lobe-like (Fig. 85AA).
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<emphasis id="1B64467F828AA6F6719C2940CFED3EAD" bold="true" pageId="0" pageNumber="1">
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<emphasis id="3C8BC3F6B92B9885BDA3ABBD46E74E7A" italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
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.
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</emphasis>
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In lateral view, promesonotum relatively low and arched, posterior mesonotum relatively steep, without tubercle-like projections; promesonotal groove absent; metanotal groove absent; propodeal spines long, triangular, thin, with sharp apex and moderately narrow base; humeral area laterally weakly produced (Fig.
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<figureCitation id="F2227487234D2408CBE06AFA64AF5C5C" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Pheidole vohamarensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0107947) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0107946)." figureDoi="10.3897/zookeys.905.39592.figure38" httpUri="https://binary.pensoft.net/fig/373085" pageId="0" pageNumber="1">38D</figureCitation>
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). Surface foveolate, with additional sparse to dense, moderately thick rugoreticulation; dorsoventral propodeum with fading sculpture. Pilosity moderately sparse, long, erect (Fig.
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<figureCitation id="900FEC449AB94EDAB09B2C0494CE1216" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Pheidole vohamarensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0107947) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0107946)." figureDoi="10.3897/zookeys.905.39592.figure38" httpUri="https://binary.pensoft.net/fig/373085" pageId="0" pageNumber="1">38D, F</figureCitation>
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||
).
|
||
<emphasis id="1A60B5394863E7EEB465C724A3154EEC" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="336B5DF04FA992163D75B7C1CD60CAAD" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
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.
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</emphasis>
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Shiny; peduncle moderately long, foveolate, with very distinct, wide, horizontal lobes on its basal part; node relatively high, thin, with rounded apex, in rear view node with deep dorsoventral concavity; pilosity long and erect (Fig.
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<figureCitation id="72E524AB18B343D1623D0AA4592E6033" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Pheidole vohamarensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0107947) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0107946)." figureDoi="10.3897/zookeys.905.39592.figure38" httpUri="https://binary.pensoft.net/fig/373085" pageId="0" pageNumber="1">38D, F</figureCitation>
|
||
).
|
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<emphasis id="C983108504E0AADC2FE424670407D73C" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="B4ED84D61254AEF1939F8924C3EF66AE" italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
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.
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</emphasis>
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Shiny and foveolate; in dorsal view sides with long, wide, acute, and triangular projections; pilosity long and erect (Fig.
|
||
<figureCitation id="F020BE99B6128485020E259EF36E719A" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Pheidole vohamarensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0107947) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0107946)." figureDoi="10.3897/zookeys.905.39592.figure38" httpUri="https://binary.pensoft.net/fig/373085" pageId="0" pageNumber="1">38D, F</figureCitation>
|
||
).
|
||
<emphasis id="FC9136A3FE425D696C23998032EC32B3" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="589696F2E082CA92CD2DA96AFF6222ED" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||
.
|
||
</emphasis>
|
||
Shiny and shagreened; pilosity dense, long, and erect (Fig.
|
||
<figureCitation id="77F6CC67726D4F88A20B4EE83CA8E97A" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Pheidole vohamarensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0107947) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0107946)." figureDoi="10.3897/zookeys.905.39592.figure38" httpUri="https://binary.pensoft.net/fig/373085" pageId="0" pageNumber="1">38D, F</figureCitation>
|
||
).
|
||
<emphasis id="9BADC1A0ACAB7D28E5B6D38CB7DA8C7B" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="76327ADB433926BCC38803BEF4976AEF" italics="true" pageId="0" pageNumber="1">Colour</emphasis>
|
||
.
|
||
</emphasis>
|
||
Bright brown to dark brown, sometimes head with brighter colouration; legs yellow (Fig.
|
||
<figureCitation id="21D59E0A0FC9E9A90C03CD116E2541CF" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Pheidole vohamarensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0107947) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0107946)." figureDoi="10.3897/zookeys.905.39592.figure38" httpUri="https://binary.pensoft.net/fig/373085" pageId="0" pageNumber="1">38D, F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="B89BDEEDA64B345E5A80DD1EAD38B12E" doi="10.3897/zookeys.905.39592.figure38" httpUri="https://binary.pensoft.net/fig/373085" pageId="0" pageNumber="1" start="Figure 38" startId="F38">
|
||
<paragraph id="4F07ED8085544D149D028B1BD3B21F63" pageId="0" pageNumber="1">
|
||
<emphasis id="DC57437B00FD3C4E096F1E1544C3CA0D" bold="true" pageId="0" pageNumber="1">Figure 38.</emphasis>
|
||
<taxonomicName id="7E5E39E437A6F188B5A059B205433D5E" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole vohamarensis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="vohamarensis">
|
||
<emphasis id="AE61E9554824E0365A597FE3A6235761" italics="true" pageId="0" pageNumber="1">Pheidole vohamarensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., full-face view (
|
||
<emphasis id="CEC2E0D00E6C0938CE70DCF1C7FEF211" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
), profile (
|
||
<emphasis id="0DFCB62BE459C9235AF98339E5D8A29D" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
), and dorsal view (
|
||
<emphasis id="1F7BB77541BC54374851328BCFA19D72" bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||
) of paratype minor worker (CASENT0107947) and full-face view (
|
||
<emphasis id="4C94E91D838EBB80633FDA4ABB6D3A22" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
), profile (
|
||
<emphasis id="0EBA8D586C05EE0BC5D342E36468B454" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
), and dorsal view (
|
||
<emphasis id="25B67970CC1E832C054AED1F6DBACAEC" bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||
) of holotype major worker (CASENT0107946).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="402318C46FDB31B2AA68AB3AD7A37F77" pageId="0" pageNumber="1">
|
||
<emphasis id="679CC97EBBD59B19A8B03455FD9BD283" bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
|
||
Measurements (
|
||
<emphasis id="7A151212DCFE36E6E541BA02EA6892C4" italics="true" pageId="0" pageNumber="1">N</emphasis>
|
||
= 10): HL: 0.44-0.53 (0.49); HW: 0.4-0.45 (0.43); SL: 0.47-0.51 (0.5); EL: 0.08-0.11 (0.1); WL: 0.58-0.63 (0.6); PSL: 0.12-0.14 (0.13); MTL: 0.35-0.4 (0.37); PNW: 0.28-0.33 (0.3); PTW: 0.07-0.09 (0.08); PPW: 0.1-0.13 (0.11); CI: 82.6-94.3 (86.7); SI: 110.2-122.0 (115.8); PSLI: 23.4-30.9 (26.5); PPI: 60.4-82.9 (71.7); PNI: 67.2-75.3 (71.0); MTI: 81.0-90.9 (86.7).
|
||
<emphasis id="DA4B29F7E5C4B7A8822EF6FAAD7A7607" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="29407350B5E538658FA04545C307BC67" italics="true" pageId="0" pageNumber="1">Head</emphasis>
|
||
.
|
||
</emphasis>
|
||
Occipital margin straight or indistinctly concave; occipital carina indistinct (Fig.
|
||
<figureCitation id="4B2FCE5814A3DE5AF81909668EAC8F53" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Pheidole vohamarensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0107947) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0107946)." figureDoi="10.3897/zookeys.905.39592.figure38" httpUri="https://binary.pensoft.net/fig/373085" pageId="0" pageNumber="1">38A</figureCitation>
|
||
). Pilosity moderately dense, long, suberect. Head foveolate; genae with smooth notch. Clypeus foveolate; median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, surpassing the posterior head margin by two-fifths of its length; pilosity erect (Fig.
|
||
<figureCitation id="327C22D4F7366053F9A3273F5E982737" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Pheidole vohamarensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0107947) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0107946)." figureDoi="10.3897/zookeys.905.39592.figure38" httpUri="https://binary.pensoft.net/fig/373085" pageId="0" pageNumber="1">38A, C</figureCitation>
|
||
).
|
||
<emphasis id="EB7216DD8798121941CD44D66B7983F4" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="2BE7ADC3221BA7A4CEA2562ACFBA2D73" italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
|
||
.
|
||
</emphasis>
|
||
In lateral view, promesonotum low, slightly convex, short, with posterior declivity relatively steep; promesonotal groove absent; metanotal groove present; propodeal spines long, triangular, with acute apex and narrow base (Fig.
|
||
<figureCitation id="3A1974E12A5BE1150B2A4CAF7EC702F0" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Pheidole vohamarensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0107947) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0107946)." figureDoi="10.3897/zookeys.905.39592.figure38" httpUri="https://binary.pensoft.net/fig/373085" pageId="0" pageNumber="1">38C</figureCitation>
|
||
). Sculpture foveolate; katepisternum and mesonotum with smooth notches. Pilosity moderately sparse, short, and erect (Fig.
|
||
<figureCitation id="5647F7934A7CB5AA39FFCC41DB6998DA" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Pheidole vohamarensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0107947) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0107946)." figureDoi="10.3897/zookeys.905.39592.figure38" httpUri="https://binary.pensoft.net/fig/373085" pageId="0" pageNumber="1">38C, E</figureCitation>
|
||
).
|
||
<emphasis id="F2ED6EBA69C96133250C767FAA670662" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="EFA903776A21ECAB358CB4F9C5024D0D" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||
.
|
||
</emphasis>
|
||
Peduncle moderately short and thin; with few short, erect setae (Fig.
|
||
<figureCitation id="BB19880185D0680F8BCDE9A71D05B5EC" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Pheidole vohamarensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0107947) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0107946)." figureDoi="10.3897/zookeys.905.39592.figure38" httpUri="https://binary.pensoft.net/fig/373085" pageId="0" pageNumber="1">38C, E</figureCitation>
|
||
).
|
||
<emphasis id="B84F3CC9F9B2108D6A66946B5A66994D" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="849D135281C90B994CC64BC698C406FD" italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
|
||
.
|
||
</emphasis>
|
||
Short, low, and convex; with few short, erect setae (Fig.
|
||
<figureCitation id="C2DCE36110A0349C4D605DEC9140AE86" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Pheidole vohamarensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0107947) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0107946)." figureDoi="10.3897/zookeys.905.39592.figure38" httpUri="https://binary.pensoft.net/fig/373085" pageId="0" pageNumber="1">38C, E</figureCitation>
|
||
).
|
||
<emphasis id="E175327F142BB83913D88ECECB7037E6" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="E8C280E38CEFDD5C5FE9AEE84673F6B7" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||
.
|
||
</emphasis>
|
||
With moderately sparse, erect pilosity (Fig.
|
||
<figureCitation id="BF928A3A36AE699669F8B5D2696482CB" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Pheidole vohamarensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0107947) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0107946)." figureDoi="10.3897/zookeys.905.39592.figure38" httpUri="https://binary.pensoft.net/fig/373085" pageId="0" pageNumber="1">38C, E</figureCitation>
|
||
).
|
||
<emphasis id="7BF68BFE03E58377BE7BA2E4F0AD4AA6" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="468C6E2C37C9D2E7EDBCA581FC06A562" italics="true" pageId="0" pageNumber="1">Colour</emphasis>
|
||
.
|
||
</emphasis>
|
||
Yellow (Fig.
|
||
<figureCitation id="0EEF9EA57C9FE4B6658CF5B8571442E9" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Pheidole vohamarensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0107947) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0107946)." figureDoi="10.3897/zookeys.905.39592.figure38" httpUri="https://binary.pensoft.net/fig/373085" pageId="0" pageNumber="1">38C, E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D01E8A876E9783DA0EF3556394F11A87" pageId="0" pageNumber="1" type="etymology">
|
||
<paragraph id="6BDAE4383586D017D60268FD9D242148" pageId="0" pageNumber="1">Etymology.</paragraph>
|
||
<paragraph id="34655009F222570F1639F6BBD6BD22A6" pageId="0" pageNumber="1">From type locality.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1420A49DF5A959EE0232A1B334A80F6F" pageId="0" pageNumber="1" type="biology_ecology">
|
||
<paragraph id="E2E4A773F50CB72D8192E623046CF7F8" pageId="0" pageNumber="1">Biology.</paragraph>
|
||
<paragraph id="E1291155AB00A527AC58F31286DDAFF4" pageId="0" pageNumber="1">The species was collected between 10-775 m in elevation, in tropical dry forest, littoral rainforest, and rainforest. Nests were located in rotten logs.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8147B175A6A5877FFF0F7CE20881CB92" pageId="0" pageNumber="1" type="comments">
|
||
<paragraph id="9F66B982568CAE899F98CB9965B23EB1" pageId="0" pageNumber="1">Comments.</paragraph>
|
||
<paragraph id="08F0980ABA5DBD308E6A16EBA4B0684D" pageId="0" pageNumber="1">
|
||
<taxonomicName id="9C45B652F69FF61792B3E2953B3C987B" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole vohemarensis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="vohemarensis">
|
||
<emphasis id="BC9949DD875FC7206C8BB94A9634BD77" italics="true" pageId="0" pageNumber="1">Pheidole vohemarensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is most similar to
|
||
<taxonomicName id="4AD4B037361C5408F8FE2E6EB6D0B803" lsidName="P. manantenensis" pageId="0" pageNumber="1" rank="species" species="manantenensis">
|
||
<emphasis id="964265878DDB5F433CCFCD9F7668FA7D" italics="true" pageId="0" pageNumber="1">P. manantenensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis id="F3F3AA219D7B2B9147E44D6CE95DF72D" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="975BDE91F9C5E7069CDE1BD4AB22DDF8" italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
|
||
.
|
||
</emphasis>
|
||
It differs from
|
||
<taxonomicName id="0C8AAC7CFDAB3372A40927CBBA1EB6EF" lsidName="P. manantenensis" pageId="0" pageNumber="1" rank="species" species="manantenensis">
|
||
<emphasis id="8F62A874012EE19335EF5498A4FDA4BB" italics="true" pageId="0" pageNumber="1">P. manantenensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. in presence of longitudinal rugae on antennal scrobes, frons never with foveolae, higher promesonotum and petiolar peduncle with wide and distinct horizontal lobes on its basal part.
|
||
<emphasis id="B313F27526380042BB6067FD27893673" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="E87802CB23EF4EA8B5E5170DCDCDED11" italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
|
||
.
|
||
</emphasis>
|
||
It differs from
|
||
<taxonomicName id="7B1507A0A5B9EC0581B82087A0893239" lsidName="P. vohemarensis" pageId="0" pageNumber="1" rank="species" species="vohemarensis">
|
||
<emphasis id="B42064519422834EF01841EFB5E9E22B" italics="true" pageId="0" pageNumber="1">P. vohemarensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. in promesonotum with posterior declivity relatively steep and katepisternum and mesonotum with smooth notches.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |