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<mods:title id="AF229C47E64F9210D6FC1A39D7CBE46C">Some ascidians from the southern coast of Madagascar collected during the “ AtimoVatae ” survey</mods:title>
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<paragraph id="B8435009FF83FF8271A5CCF3FE4F6BEC" blockId="19.[151,497,151,210]" box="[151,497,151,177]" pageId="19" pageNumber="20">
<heading id="E30BE765FF83FF8271A5CCF3FE4F6BEC" bold="true" box="[151,497,151,177]" fontSize="11" level="1" pageId="19" pageNumber="20" reason="1">
<taxonomicName id="7FFC2B8AFF83FF8271A5CCF3FE4F6BEC" authority="Kott, 1992" authorityName="Kott" authorityYear="1992" box="[151,497,151,177]" class="Ascidiacea" family="Ritterellidae" genus="Ritterella" kingdom="Animalia" order="Enterogona" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="tokioka">
<emphasis id="8A888C1BFF83FF8271A5CCF3FE4F6BEC" bold="true" box="[151,497,151,177]" pageId="19" pageNumber="20">
<emphasis id="8A888C1BFF83FF8271A5CCF3FED96BEC" bold="true" box="[151,359,151,177]" italics="true" pageId="19" pageNumber="20">Ritterella tokioka</emphasis>
<bibRefCitation id="DC6D2DF8FF83FF82705DCCF3FE4F6BEC" author="Kott" box="[367,497,151,177]" pageId="19" pageNumber="20" refString="Kott, P. (1992) The Australian Ascidiacea. Part 3, Aplousobranchia (2). Memoirs of the Queensland Museum, 32 (2), 375 - 620." type="journal article" year="1992">Kott, 1992</bibRefCitation>
</emphasis>
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</heading>
</paragraph>
<paragraph id="B8435009FF83FF8271A5CCDEFEA86B8F" blockId="19.[151,497,151,210]" box="[151,278,186,210]" pageId="19" pageNumber="20">
(
<figureCitation id="20C74C8CFF83FF8271ACCCDEFEB36B8F" box="[158,269,186,210]" captionStart="FIGURE 16" captionStartId="19.[151,250,1467,1490]" captionTargetBox="[151,1436,901,1443]" captionTargetId="figure@19.[151,1436,901,1446]" captionTargetPageId="19" captionText="FIGURE 16. Ritterella tokioka: A, parts of colony; B, branchial sac. Scale bar A = 1 cm." httpUri="https://zenodo.org/record/246198/files/figure.png" pageId="19" pageNumber="20">Figure 16</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="F0E60382FF83FF8271A5CD66FBFD69C1" pageId="19" pageNumber="20" type="reference_group">
<paragraph id="B8435009FF83FF8271A5CD66FDFD6A6A" blockId="19.[151,1436,258,311]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="98EC7727FF83FF8271A5CD66FDFD6A6A" pageId="19" pageNumber="20">
<taxonomicName id="7FFC2B8AFF83FF8271A5CD66FE4A6A45" authority="Kott, 1992: 402" authorityName="Kott" authorityPageNumber="402" authorityYear="1992" box="[151,500,258,280]" class="Ascidiacea" family="Ritterellidae" genus="Ritterella" kingdom="Animalia" order="Enterogona" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="tokioka">
<emphasis id="8A888C1BFF83FF8271A5CD66FEF66A45" box="[151,328,258,280]" italics="true" pageId="19" pageNumber="20">Ritterella tokioka</emphasis>
<treatmentCitation id="395D7618FF83FF82707DCD67FE4A6A45" author="Kott" box="[335,500,259,280]" page="402" pageId="19" pageNumber="20" year="1992">
<bibRefCitation id="DC6D2DF8FF83FF82707DCD67FE7E6A45" author="Kott" box="[335,448,259,280]" pageId="19" pageNumber="20" refString="Kott, P. (1992) The Australian Ascidiacea. Part 3, Aplousobranchia (2). Memoirs of the Queensland Museum, 32 (2), 375 - 620." type="journal article" year="1992">Kott, 1992</bibRefCitation>
: 402
</treatmentCitation>
</taxonomicName>
,
<collectingCountry id="C0EB1099FF83FF8270CCCD67FD846A45" box="[510,570,259,280]" name="Japan" pageId="19" pageNumber="20">Japan</collectingCountry>
, nom. nov. for
<taxonomicName id="7FFC2B8AFF83FF8273E5CD67FBAE6A45" authority="Tokioka, 1953" authorityName="Tokioka" authorityYear="1953" box="[727,1040,258,280]" class="Ascidiacea" family="Ritterellidae" genus="Ritterella" kingdom="Animalia" order="Enterogona" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="pedunculata">
<emphasis id="8A888C1BFF83FF8273E5CD67FCCF6A45" box="[727,881,258,280]" italics="true" pageId="19" pageNumber="20">R. pedunculata</emphasis>
<bibRefCitation id="DC6D2DF8FF83FF82724BCD66FBAE6A45" author="Tokioka" box="[889,1040,258,280]" pageId="19" pageNumber="20" refString="Tokioka, T. (1953) Ascidians of Sagami Bay, 1 - 313, In Hattori H. (ed) Iwanami Shoten, Tokyo." type="book chapter" year="1953">Tokioka, 1953</bibRefCitation>
</taxonomicName>
. Monniot &amp; Monniot 1997 (1999) and synonymy,
<collectingCountry id="C0EB1099FF83FF827009CD45FE286A6A" box="[315,406,289,311]" name="Tanzania" pageId="19" pageNumber="20">Tanzania</collectingCountry>
&amp;
<collectingCountry id="C0EB1099FF83FF827085CD45FDFE6A6A" box="[439,576,289,311]" name="Mozambique" pageId="19" pageNumber="20">Mozambique</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="B8435009FF83FF8271A5CD07FDFF6A21" blockId="19.[151,1436,355,847]" box="[151,577,355,380]" pageId="19" pageNumber="20">
<emphasis id="8A888C1BFF83FF8271A5CD07FF426A21" bold="true" box="[151,252,355,380]" pageId="19" pageNumber="20">Stations.</emphasis>
TA 23 (MNHN: A1 RIT 33)
</paragraph>
<paragraph id="B8435009FF83FF8271F5CDE3FCD36AC2" blockId="19.[151,1436,355,847]" box="[199,877,391,415]" pageId="19" pageNumber="20">
<collectingCountry id="C0EB1099FF83FF8271F5CDE3FEEE6AC2" box="[199,336,391,415]" name="Madagascar" pageId="19" pageNumber="20">Madagascar</collectingCountry>
, Tulear, Vasseur col. 1969 (MNHN A1 RIT 29).
</paragraph>
<paragraph id="B8435009FF83FF8271F5CDC8FBFD69C1" blockId="19.[151,1436,355,847]" pageId="19" pageNumber="20">
The colonies are sandy lobes arising from a common base (
<figureCitation id="20C74C8CFF83FF827268CDC8FC156A99" box="[858,939,428,452]" captionStart="FIGURE 16" captionStartId="19.[151,250,1467,1490]" captionTargetBox="[151,1436,901,1443]" captionTargetId="figure@19.[151,1436,901,1446]" captionTargetPageId="19" captionText="FIGURE 16. Ritterella tokioka: A, parts of colony; B, branchial sac. Scale bar A = 1 cm." httpUri="https://zenodo.org/record/246198/files/figure.png" pageId="19" pageNumber="20">Fig. 16</figureCitation>
A). The widened head of the lobes is slightly transparent as the sand grains are less dense. The zooids correspond to the description and figures by
<bibRefCitation id="DC6D2DF8FF83FF82742CCDABFEF36951" author="Monniot" pageId="19" pageNumber="20" refString="Monniot, F. &amp; Monniot, C. (1999) Ascidians collected in Tanzania. Journal of East African Natural History, 86, 1 - 35." type="journal article" year="1999">Monniot &amp; Monniot (1999)</bibRefCitation>
. The branchial sac (
<figureCitation id="20C74C8CFF83FF827302CD90FD3F6951" box="[560,641,500,524]" captionStart="FIGURE 16" captionStartId="19.[151,250,1467,1490]" captionTargetBox="[151,1436,901,1443]" captionTargetId="figure@19.[151,1436,901,1446]" captionTargetPageId="19" captionText="FIGURE 16. Ritterella tokioka: A, parts of colony; B, branchial sac. Scale bar A = 1 cm." httpUri="https://zenodo.org/record/246198/files/figure.png" pageId="19" pageNumber="20">Fig. 16</figureCitation>
B) counts 10 to 11 rows of stigmata with parastigmatic vessels incomplete or only present on some rows, according to the zooids. The stomach is always long with about 10 longitudinal folds. There is an annular post-stomach and a caecum at the base of the rectum. The ovary is placed very close to the gut loop followed by a series of testis vesicles limited to the anterior part of the post-abdomen. No larvae have been found. The zooids from the south of
<collectingCountry id="C0EB1099FF83FF82735FCEE0FD4A69C1" box="[621,756,644,668]" name="Madagascar" pageId="19" pageNumber="20">Madagascar</collectingCountry>
remained yellow in formalin.
</paragraph>
</subSubSection>
<subSubSection id="F0E60382FF83FF8271FBCECCFAB9685A" pageId="19" pageNumber="20" type="discussion">
<paragraph id="B8435009FF83FF8271FBCECCFAB9685A" blockId="19.[151,1436,355,847]" pageId="19" pageNumber="20">
<emphasis id="8A888C1BFF83FF8271FBCECCFE83699D" bold="true" box="[201,317,680,704]" pageId="19" pageNumber="20">Remarks.</emphasis>
The colonies of
<collectingCountry id="C0EB1099FF83FF8270CACEC3FD3E69E2" box="[504,640,679,703]" name="Madagascar" pageId="19" pageNumber="20">Madagascar</collectingCountry>
are identified according to the description of the species given by
<taxonomicName id="7FFC2B8AFF83FF827455CEC3FEA569B9" class="Ascidiacea" family="Ritterellidae" genus="Ritterella" kingdom="Animalia" order="Enterogona" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="tokioka">
<bibRefCitation id="DC6D2DF8FF83FF827455CEC3FEA569B9" author="Tokioka" pageId="19" pageNumber="20" refString="Tokioka, T. (1953) Ascidians of Sagami Bay, 1 - 313, In Hattori H. (ed) Iwanami Shoten, Tokyo." type="book chapter" year="1953">Tokioka (1953)</bibRefCitation>
</taxonomicName>
for the Japanese material. The geographic distance between the populations suggests a convergence more than a specific identity, the identification remains in absence of obvious anatomical differences.
</paragraph>
</subSubSection>
<subSubSection id="F0E60382FF83FF8271F5CF70FC266812" pageId="19" pageNumber="20" type="reference_group">
<paragraph id="B8435009FF83FF8271F5CF70FC266812" blockId="19.[151,1436,355,847]" pageId="19" pageNumber="20">
Another
<taxonomicName id="7FFC2B8AFF83FF827018CF70FE2D6871" box="[298,403,788,812]" class="Ascidiacea" family="Ritterellidae" genus="Ritterella" kingdom="Animalia" order="Enterogona" pageId="19" pageNumber="20" phylum="Chordata" rank="genus">
<emphasis id="8A888C1BFF83FF827018CF70FE2D6871" box="[298,403,788,812]" italics="true" pageId="19" pageNumber="20">Ritterella</emphasis>
</taxonomicName>
species:
<taxonomicName id="7FFC2B8AFF83FF827330CF71FD426871" authority="Kott, 1957" authorityName="Kott" authorityYear="1957" box="[514,764,788,812]" class="Ascidiacea" family="Ritterellidae" genus="Ritterella" kingdom="Animalia" order="Enterogona" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="dispar">
<emphasis id="8A888C1BFF83FF827330CF71FDD46871" box="[514,618,788,812]" italics="true" pageId="19" pageNumber="20">R. dispar</emphasis>
(
<bibRefCitation id="DC6D2DF8FF83FF82734BCF70FD4A6871" author="Kott" box="[633,756,788,812]" pageId="19" pageNumber="20" refString="Kott, P. (1957) The ascidians of Australia. II. Aplousobranchiata Lahille: Clavelinidae Forbes &amp; Hanley and Polyclinidae Verrill. Australian Journal of marine &amp; freshwater Research, 8 (1), 64 - 110." type="journal article" year="1957">Kott, 1957</bibRefCitation>
)
</taxonomicName>
is also recorded from
<collectingCountry id="C0EB1099FF83FF8272C9CF70FB316871" box="[1019,1167,788,812]" name="Mozambique" pageId="19" pageNumber="20">Mozambique</collectingCountry>
and
<collectingCountry id="C0EB1099FF83FF8275F4CF70FA956871" box="[1222,1323,788,812]" name="Tanzania" pageId="19" pageNumber="20">Tanzania</collectingCountry>
(Monniot &amp; Monniot
<date id="CC4276C9FF83FF827029CF53FE206812" box="[283,414,823,847]" pageId="19" pageNumber="20" value="1997" valueMax="1999">19971999</date>
). It differs in having only 5 rows of stigmata.
</paragraph>
</subSubSection>
</treatment>
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